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<document id="ADE5507D994ED56EFD82997D3B8D91CC" ID-DOI="10.5252/geodiversitas2024v46a2" ID-ISSN="1638-9395" ID-Zenodo-Dep="10566276" ID-ZooBank="urn:lsid:zoobank.org:pub:A141C894-DEC7-42FD-BC7C-2A8A692B0A31" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1706171845289" checkinUser="plazi" docAuthor="Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni &amp; Chény, Cédric" docDate="2024" docId="704E030AFFA8FF98DA4CF8BDFD622E11" docLanguage="en" docName="geodiversitas.2024.46.2.pdf" docOrigin="Geodiversitas 46 (2)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a2.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Parahyas oranensis Charbonnier &amp; Garassino &amp; Pasini &amp; Chény 2024, n. comb." docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="8C777B72FFA0FF93D910FFAFFF9D2C21" masterDocTitle="Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria" masterLastPageNumber="29" masterPageNumber="13" pageNumber="19" updateTime="1706559400298" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="AE8EB34CC48C919246534854644AC097">Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria</mods:title>
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<mods:namePart id="608952A18A9F68AF185B08ABCAC3EA92">Charbonnier, Sylvain</mods:namePart>
<mods:affiliation id="CEC5DE0E7EE82097B49927B536BFC2F4">Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), MNHN, CNRS, Sorbonne Université, Muséum national d’Histoire naturelle, 57 rue Cuvier, F- 75231 Paris, cedex 05 (France)</mods:affiliation>
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<mods:namePart id="5DF4AD3A25E83D3543C9C6BF212A0FD4">Garassino, Alessandro</mods:namePart>
<mods:affiliation id="408B4675605D206E254851022004AB18">Department of Earth and Biological Sciences, Loma Linda University, Loma Linda, 92350 CA (United States)</mods:affiliation>
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<mods:namePart id="2F90F2B6C3FD7A042EC1604E733DB5C6">Pasini, Giovanni</mods:namePart>
<mods:affiliation id="BF98C79E8B5AE6675297578CC074CFD4">Via Alessandro Volta, 16 22070 Appiano Gentile (Italy)</mods:affiliation>
<mods:nameIdentifier id="6AED6F440A4907390120F5005E004738" type="email">giovannialdopasini@gmail.com</mods:nameIdentifier>
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<mods:namePart id="17EE74104639ABB4D2A90A91F8E98F48">Chény, Cédric</mods:namePart>
<mods:affiliation id="A24143F455EBA09CEBDF23787F1D655E">Géosciences Rennes, UMR 6118, CNRS Université de Rennes, F- 35000 Rennes (France)</mods:affiliation>
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<mods:title id="0690744564A2A3D8779118A1D90BD226">Geodiversitas</mods:title>
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<mods:date id="5008DDA6BAC69C033364FA095381E82E">2024</mods:date>
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<mods:number id="6AFF38766A484192055AA0DB5A75B119">46</mods:number>
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<treatment id="704E030AFFA8FF98DA4CF8BDFD622E11" ID-DOI="http://doi.org/10.5281/zenodo.10566304" ID-Zenodo-Dep="10566304" LSID="urn:lsid:plazi:treatment:704E030AFFA8FF98DA4CF8BDFD622E11" httpUri="http://treatment.plazi.org/id/704E030AFFA8FF98DA4CF8BDFD622E11" lastPageId="11" lastPageNumber="22" pageId="8" pageNumber="19">
<subSubSection id="B0FDE197FFA8FF9BDA4CF8BDFA1D2B0C" box="[860,1408,1810,1837]" pageId="8" pageNumber="19" type="nomenclature">
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<heading id="A3100570FFA8FF9BDA4CF8BDFA1D2B0C" box="[860,1408,1810,1837]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="19" reason="2">
<taxonomicName id="3FE7C99FFFA8FF9BDA4CF8BDFA8E2B0C" authority="(Van Straelen, 1937)" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" baseAuthorityName="Van Straelen" baseAuthorityYear="1937" box="[860,1299,1810,1837]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="oranensis" status="comb. nov.">
<emphasis id="CA936E0EFFA8FF9BDA4CF8BDFBB62B0C" bold="true" box="[860,1067,1810,1837]" italics="true" pageId="8" pageNumber="19">Parahyas oranensis</emphasis>
(
<bibRefCitation id="9C76CFEDFFA8FF9BDD29F8BCFA962B0C" author="VAN STRAELEN V." box="[1081,1291,1811,1837]" pageId="8" pageNumber="19" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="D1A0D375FFA8FF9BDC0FF8BCFA1D2B0C" box="[1311,1408,1811,1837]" pageId="8" pageNumber="19" rank="species">n. comb.</taxonomicNameLabel>
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<paragraph id="F858B21CFFA8FF9BDD56F89CFB0B2B6C" blockId="8.[1094,1174,1843,1869]" box="[1094,1174,1843,1869]" pageId="8" pageNumber="19">
(
<figureCitation id="60DCAE99FFA8FF9BDD5FF89CFB102B6C" box="[1103,1165,1843,1869]" captionStart="FIG" captionStartId="9.[132,143,1703,1720]" captionTargetBox="[132,1306,215,1661]" captionTargetId="figure-111@9.[699,1268,757,1301]" captionTargetPageId="9" captionText="FIG. 3. — Parahyas oranensis (Van Straelen,1937),n. comb.from the late Messinian of Oran (Les Planteurs quarry,Arambourg coll.):A, B, holotype MNHN.F.R03767, dorsal view and line drawing of carapace; C, D, MNHN.FA28903 (ex paratype of Calappa saheliensis Van Straelen, 1937), dorsal view and line drawing of carapace. Abbreviations:b, branchiocardiac groove; br, branchial region;ca, cardiac region;i, intestinal region;mt, metabranchial region; ps, cup-shaped postorbital spine; u, urogastric region. Scale bars: 1 cm. Photographs: Lilian Cazes." figureDoi="http://doi.org/10.5281/zenodo.10566286" httpUri="https://zenodo.org/record/10566286/files/figure.png" pageId="8" pageNumber="19">Fig. 3</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="D8F79532FFA8FF9BDA3DF82AFB862BCA" pageId="8" pageNumber="19">
<treatmentCitation id="7946940DFFA8FF9BDA3DF82AFB392BBD" author="VAN STRAELEN V." box="[813,1188,1924,1948]" page="474" pageId="8" pageNumber="19" year="1937">
<taxonomicName id="3FE7C99FFFA8FF9BDA3DF82AFB392BBD" ID-CoL="BR9L3" authority="Van Straelen, 1937: 474" authorityName="Van Straelen" authorityPageNumber="474" authorityYear="1937" box="[813,1188,1924,1948]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="oranensis">
<emphasis id="CA936E0EFFA8FF9BDA3DF82AFC292BBD" box="[813,948,1925,1948]" italics="true" pageId="8" pageNumber="19">Hyas oranensis</emphasis>
<bibRefCitation id="9C76CFEDFFA8FF9BDAA8F82BFB392BBD" author="VAN STRAELEN V." box="[952,1188,1924,1948]" pageId="8" pageNumber="19" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937: 474</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 475, pl. 33, fig. 2. —
<treatmentCitation id="7946940DFFA8FF9BDC6CF82BFBA32B97" author="SAINT MARTIN J. - P. &amp; MULLER P." page="251" pageId="8" pageNumber="19" year="1988">
<bibRefCitation id="9C76CFEDFFA8FF9BDC6CF82BFBA32B97" author="SAINT MARTIN J. - P. &amp; MULLER P." pageId="8" pageNumber="19" pagination="251 - 257" refId="ref11818" refString="SAINT MARTIN J. - P. &amp; MULLER P. 1988. - Les Crustaces Decapodes du Messinien recifal d'Oranie (Algerie). Geobios 21 (2): 251 - 257. https: // doi. org / 10.1016 / S 0016 - 6995 (88) 80022 - 6" type="journal article" year="1988">
<collectingCountry id="80F0F28CFFA8FF9BDC6CF82BFCEF2B97" name="Sint Martin" pageId="8" pageNumber="19">Saint Martin</collectingCountry>
&amp; Müller 1988: 251
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="7946940DFFA8FF9BDD73F831FA132B96" author="MOISSETTE P. &amp; MULLER P." box="[1123,1422,1950,1975]" page="738" pageId="8" pageNumber="19" year="1990">
<bibRefCitation id="9C76CFEDFFA8FF9BDD73F831FA132B96" author="MOISSETTE P. &amp; MULLER P." box="[1123,1422,1950,1975]" pageId="8" pageNumber="19" pagination="737 - 747" refId="ref11138" refString="MOISSETTE P. &amp; MULLER P. 1990. - Crustaces Decapodes des facies marno-diatomitique du Messinien d'Oranie (Algerie Occidentale). Geobios 23 (6): 737 - 747. https: // doi. org / 10.1016 / S 0016 - 6995 (06) 80339 - 6" type="journal article" year="1990">Moissette &amp; Müller 1990: 738</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="7946940DFFA8FF9BDA3BF816FBB12BF0" author="SCHWEITZER C. E. &amp; FELDMANN R. M. &amp; GARASSINO A. &amp; KARASAWA H. &amp; SCHWEIGERT G." box="[811,1068,1977,2001]" page="97" pageId="8" pageNumber="19" year="2010">
<bibRefCitation id="9C76CFEDFFA8FF9BDA3BF816FBB12BF0" author="SCHWEITZER C. E. &amp; FELDMANN R. M. &amp; GARASSINO A. &amp; KARASAWA H. &amp; SCHWEIGERT G." box="[811,1068,1977,2001]" pageId="8" pageNumber="19" pagination="1 - 222" refId="ref11928" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. &amp; SCHWEIGERT G. 2010. - Systematic list of fossil decapod crustacean species. Crustaceana Monographs 10: 1 - 222. https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
Schweitzer
<emphasis id="CA936E0EFFA8FF9BDA8CF815FC562BF0" box="[924,971,1977,2001]" italics="true" pageId="8" pageNumber="19">et al.</emphasis>
2010: 97
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="7946940DFFA8FF9BDD47F816FAB72BF0" author="KATO H. &amp; NAKASHIMA R. &amp; YANAGISAWA Y." box="[1111,1322,1977,2001]" page="270" pageId="8" pageNumber="19" year="2013">
<bibRefCitation id="9C76CFEDFFA8FF9BDD47F816FAB72BF0" author="KATO H. &amp; NAKASHIMA R. &amp; YANAGISAWA Y." box="[1111,1322,1977,2001]" pageId="8" pageNumber="19" pagination="269 - 287" refId="ref10615" refString="KATO H., NAKASHIMA R. &amp; YANAGISAWA Y. 2013. - Two new Early Pliocene species of the crab genus Hyas Leach, 1814 (Majoidea, Oregoniidae) from northeast Japan, in FRAAIJE R. H. B., HYZNY M., JAGT J. W. M., KROBICKI M. &amp; VAN BAKEL B. W. M. (eds), Proceedings of the 5 th Symposium on Mesozoic and Cenozoic Decapod Crustaceans. Krakow, Poland, 2013: A tribute to Pal Mihaly Muller. Scripta Geologica 147: 269 - 287." type="journal article" year="2013">
Kato
<emphasis id="CA936E0EFFA8FF9BDD9DF815FB212BF0" box="[1165,1212,1977,2001]" italics="true" pageId="8" pageNumber="19">et al.</emphasis>
2013: 270
</bibRefCitation>
</treatmentCitation>
, 275-277. —
<treatmentCitation id="7946940DFFA8FF9BDA3DF87CFB8A2BCA" author="GUINOT D. &amp; CARBOT-CHANONA G. &amp; VEGA F. J." box="[813,1047,2003,2027]" page="307" pageId="8" pageNumber="19" year="2019">
<bibRefCitation id="9C76CFEDFFA8FF9BDA3DF87CFB8A2BCA" author="GUINOT D. &amp; CARBOT-CHANONA G. &amp; VEGA F. J." box="[813,1047,2003,2027]" pageId="8" pageNumber="19" pagination="285 - 322" refId="ref10254" refString="GUINOT D., CARBOT-CHANONA G. &amp; VEGA F. J. 2019. - Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new hypothesis on Lecythocaridae Schweitzer &amp; Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura). Geodiversitas 41 (7): 285 - 322. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 7. http: // geodiversitas. com / 41 / 7" type="journal article" year="2019">
Guinot
<emphasis id="CA936E0EFFA8FF9BDA6AF87BFC342BCA" box="[890,937,2003,2027]" italics="true" pageId="8" pageNumber="19">et al.</emphasis>
2019: 307
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="AC98E294FFA9FF9AD994F908FD7D2ADD" ID-DOI="http://doi.org/10.5281/zenodo.10566286" ID-Zenodo-Dep="10566286" httpUri="https://zenodo.org/record/10566286/files/figure.png" pageId="9" pageNumber="20" startId="9.[132,143,1703,1720]" targetBox="[132,1306,215,1661]" targetPageId="9" targetType="figure">
<paragraph id="F858B21CFFA9FF9AD994F908FD7D2ADD" blockId="9.[132,1456,1703,1788]" pageId="9" pageNumber="20">
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. 3. —
<taxonomicName id="3FE7C99FFFA9FF9AD9C1F908FD8B2A99" authority="(Van Straelen, 1937)" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" baseAuthorityName="Van Straelen" baseAuthorityYear="1937" box="[209,534,1703,1720]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="oranensis" status="comb. nov.">
<emphasis id="CA936E0EFFA9FF9AD9C1F908FEF02A99" box="[209,365,1703,1720]" italics="true" pageId="9" pageNumber="20">Parahyas oranensis</emphasis>
(
<bibRefCitation id="9C76CFEDFFA9FF9AD865F908FD8C2A99" author="VAN STRAELEN V." box="[373,529,1703,1720]" pageId="9" pageNumber="20" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen,1937</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="D1A0D375FFA9FF9ADB0CF908FDF82A99" box="[540,613,1703,1720]" pageId="9" pageNumber="20" rank="species">n. comb.</taxonomicNameLabel>
from the late
<taxonomicName id="3FE7C99FFFA9FF9ADBDFF908FCBF2A99" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[719,802,1703,1720]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
of Oran (Les Planteurs quarry,Arambourg coll.):
<emphasis id="CA936E0EFFA9FF9ADDB7F908FB572A99" bold="true" box="[1191,1226,1703,1720]" pageId="9" pageNumber="20">A, B</emphasis>
, holotype MNHN.F.R03767, dorsal view and line drawing of carapace;
<emphasis id="CA936E0EFFA9FF9AD8F4F911FD962AEE" bold="true" box="[484,523,1726,1743]" pageId="9" pageNumber="20">C, D</emphasis>
, MNHN.FA28903 (ex paratype of
<taxonomicName id="3FE7C99FFFA9FF9ADA35F911FBED2AEE" authority="Van Straelen, 1937" authorityName="Van Straelen" authorityYear="1937" box="[805,1136,1726,1743]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA9FF9ADA35F911FC542AEE" box="[805,969,1726,1743]" italics="true" pageId="9" pageNumber="20">Calappa saheliensis</emphasis>
<bibRefCitation id="9C76CFEDFFA9FF9ADADEF911FBED2AEE" author="VAN STRAELEN V." box="[974,1136,1726,1743]" pageId="9" pageNumber="20" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937</bibRefCitation>
</taxonomicName>
), dorsal view and line drawing of carapace. Abbreviations:
<emphasis id="CA936E0EFFA9FF9AD822F97AFEA02AC7" bold="true" box="[306,317,1749,1766]" pageId="9" pageNumber="20">b</emphasis>
, branchiocardiac groove;
<emphasis id="CA936E0EFFA9FF9ADB05F97AFDB52AC7" bold="true" box="[533,552,1749,1766]" pageId="9" pageNumber="20">br</emphasis>
, branchial region;
<emphasis id="CA936E0EFFA9FF9ADBAEF97AFD4E2AC7" bold="true" box="[702,723,1749,1766]" pageId="9" pageNumber="20">ca</emphasis>
, cardiac region;
<emphasis id="CA936E0EFFA9FF9ADA4CF97AFCFC2AC7" bold="true" box="[860,865,1749,1766]" pageId="9" pageNumber="20">i</emphasis>
, intestinal region;
<emphasis id="CA936E0EFFA9FF9ADAE6F97AFB902AC7" bold="true" box="[1014,1037,1749,1766]" pageId="9" pageNumber="20">mt</emphasis>
, metabranchial region;
<emphasis id="CA936E0EFFA9FF9ADDDFF97AFB792AC7" bold="true" box="[1231,1252,1749,1766]" pageId="9" pageNumber="20">ps</emphasis>
, cup-shaped postorbital spine;
<emphasis id="CA936E0EFFA9FF9AD9ACF944FF5A2ADD" bold="true" box="[188,199,1771,1788]" pageId="9" pageNumber="20">u</emphasis>
, urogastric region. Scale bars: 1 cm. Photographs: Lilian Cazes.
</paragraph>
</caption>
<subSubSection id="B0FDE197FFA9FF9AD994F89EFEB02B95" pageId="9" pageNumber="20" type="materials_examined">
<paragraph id="F858B21CFFA9FF9AD994F89EFE662B42" blockId="9.[132,775,1840,1891]" pageId="9" pageNumber="20">
<materialsCitation id="488FB841FFA9FF9AD994F89EFDE62B69" box="[132,635,1840,1864]" collectionCode="MNHN" collectorName="Arambourg" country="Algeria" county="Sanaouber" location="Les Planteurs quarry" municipality="Sidi El Houari" pageId="9" pageNumber="20" specimenCode="MNHN.F.R03767" specimenCount="1" typeStatus="holotype">
<smallCapsWord id="FEBE24C0FFA9FF9AD994F89EFF2B2B67" baselines="1858,1858" box="[132,182,1841,1864]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="9" pageNumber="20">TYPE</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA9FF9AD9AFF899FE822B66" baselines="1859" box="[191,287,1846,1863]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="9" pageNumber="20">MATERIAL</smallCapsWord>
. —
<typeStatus id="275C0CBEFFA9FF9AD840F89FFE322B69" box="[336,431,1840,1864]" pageId="9" pageNumber="20" type="holotype">Holotype</typeStatus>
<specimenCode id="A8411A67FFA9FF9AD8A9F89EFDE82B69" box="[441,629,1840,1864]" pageId="9" pageNumber="20">MNHN.F.R03767</specimenCode>
,
</materialsCitation>
<materialsCitation id="488FB841FFA9FF9ADB94F89EFE662B42" collectionCode="MNHN" collectorName="Arambourg" country="Algeria" county="Sanaouber" location="Les Planteurs quarry" municipality="Sidi El Houari" pageId="9" pageNumber="20" specimenCode="MNHN.F.A28903" specimenCount="1" typeStatus="paratype">
<specimenCount id="EEE17995FFA9FF9ADB94F89EFD352B69" box="[644,680,1841,1864]" pageId="9" pageNumber="20" type="generic">one</specimenCount>
<typeStatus id="275C0CBEFFA9FF9ADBA1F89EFC9A2B68" box="[689,775,1841,1865]" pageId="9" pageNumber="20">paratype</typeStatus>
<specimenCode id="A8411A67FFA9FF9AD996F8E4FEA02B42" box="[134,317,1867,1891]" pageId="9" pageNumber="20">MNHN.F.A28903</specimenCode>
(
<collectorName id="5512D7CAFFA9FF9AD85AF8E4FE212B42" box="[330,444,1867,1891]" pageId="9" pageNumber="20">Arambourg</collectorName>
coll.).
</materialsCitation>
</paragraph>
<paragraph id="F858B21CFFA9FF9AD994F82DFEB02B95" blockId="9.[132,775,1922,1972]" pageId="9" pageNumber="20">
<smallCapsWord id="FEBE24C0FFA9FF9AD994F82DFF282BB9" baselines="1939,1940" box="[132,181,1922,1945]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="9" pageNumber="20">TYPE</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA9FF9AD9ABF828FE8B2BB9" baselines="1940" box="[187,278,1927,1944]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="9" pageNumber="20">LOCALITY</smallCapsWord>
. —
<location id="FD38E4C7FFA9FF9AD851F82DFD942BBB" LSID="urn:lsid:plazi:treatment:704E030AFFA8FF98DA4CF8BDFD622E11:FD38E4C7FFA9FF9AD851F82DFD942BBB" box="[321,521,1922,1946]" country="Algeria" county="Sanaouber" municipality="Sidi El Houari" name="Les Planteurs quarry" pageId="9" pageNumber="20">Les Planteurs quarry</location>
,
<collectingCounty id="1139CA90FFA9FF9ADB02F82DFDE52BBB" box="[530,632,1922,1946]" pageId="9" pageNumber="20">Sanaouber</collectingCounty>
district of
<collectingMunicipality id="183C2866FFA9FF9ADBF1F82DFF752B95" pageId="9" pageNumber="20">Sidi El Houari</collectingMunicipality>
,
<collectingCountry id="80F0F28CFFA9FF9AD9E2F833FEB42B95" box="[242,297,1948,1972]" name="Algeria" pageId="9" pageNumber="20">Oran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA9FF9AD994F87BFDA12BCA" box="[132,572,2003,2027]" pageId="9" pageNumber="20" type="biology_ecology">
<paragraph id="F858B21CFFA9FF9AD994F87BFDA12BCA" blockId="9.[132,572,2003,2027]" box="[132,572,2003,2027]" pageId="9" pageNumber="20">
<smallCapsWord id="FEBE24C0FFA9FF9AD994F87BFF282BC8" baselines="2021,2021" box="[132,181,2004,2027]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="9" pageNumber="20">
<typeStatus id="275C0CBEFFA9FF9AD994F87BFF282BC8" box="[132,181,2004,2027]" pageId="9" pageNumber="20">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA9FF9AD9ACF876FF7C2BCB" baselines="2022" box="[188,225,2009,2026]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="age" pageId="9" pageNumber="20">AGE</smallCapsWord>
. — Late Miocene (late
<taxonomicName id="3FE7C99FFFA9FF9AD8DAF87CFDAF2BCA" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[458,562,2003,2027]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA9FF99DA3DF89DFB7F2B0D" lastPageId="10" lastPageNumber="21" pageId="9" pageNumber="20" type="description">
<paragraph id="F858B21CFFA9FF9ADA3DF89DFC202B6B" blockId="9.[813,1457,1842,2027]" box="[813,957,1842,1868]" pageId="9" pageNumber="20">
<smallCapsWord id="FEBE24C0FFA9FF9ADA3DF89DFC202B6B" baselines="1861,1861" box="[813,957,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="9" pageNumber="20">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="F858B21CFFA9FF9ADA3DF8FDFC102B4D" blockId="9.[813,1457,1842,2027]" box="[813,909,1874,1900]" pageId="9" pageNumber="20">
<emphasis id="CA936E0EFFA9FF9ADA3DF8FDFC102B4D" box="[813,909,1874,1900]" italics="true" pageId="9" pageNumber="20">Carapace</emphasis>
</paragraph>
<paragraph id="F858B21CFFA9FF99DA3DF8DEFB7F2B0D" blockId="9.[813,1457,1842,2027]" lastBlockId="10.[814,1458,1778,1836]" lastPageId="10" lastPageNumber="21" pageId="9" pageNumber="20">Carapace pyriform, convex both longitudinally and transversely, length slightly exceeding width; widest posteriorly; rostrum not preserved; orbit composed of supraorbital rim and widely cup-shaped postorbital spine bearing marginal spines; anterolateral margins poorly preserved; posterolateral margins rounded, bearing row of 6-7 wide tubercles extending along entire length of margin; posterior margin partially preserved; hepatic region depressed; proto-, meso-, and metagastric regions raised; diamond-shaped cardiac region strongly elevated; branchial regions inflated; urogastric and cardiac regions raised and limited laterally by deep branchiocardiac groove; intestinal region protruding distally beyond and under branchial regions; dorsal surface of carapace covered with small tubercles arranged uniformly.</paragraph>
</subSubSection>
<caption id="AC98E294FFAAFF99D994F9C2FDA52A8D" ID-DOI="http://doi.org/10.5281/zenodo.10566290" ID-Zenodo-Dep="10566290" httpUri="https://zenodo.org/record/10566290/files/figure.png" pageId="10" pageNumber="21" startId="10.[132,143,1645,1662]" targetBox="[132,1455,215,1603]" targetPageId="10" targetType="figure">
<paragraph id="F858B21CFFAAFF99D994F9C2FDA52A8D" blockId="10.[132,1456,1645,1708]" pageId="10" pageNumber="21">
<smallCapsWord id="FEBE24C0FFAAFF99D994F9C2FF002A5F" baselines="1658,1659" box="[132,157,1645,1662]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="21">FIG</smallCapsWord>
. 4. — Cancrid crabs from the late
<taxonomicName id="3FE7C99FFFAAFF99D8AEF9C2FD8F2A5F" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[446,530,1645,1662]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
of Oran (Les Planteurs quarry, Arambourg coll.):
<emphasis id="CA936E0EFFAAFF99DABBF9C2FC562A5F" bold="true" box="[939,971,1645,1662]" pageId="10" pageNumber="21">A-E</emphasis>
,
<taxonomicName id="3FE7C99FFFAAFF99DAC6F9C2FB462A5F" authority="Meyer, 1843" authorityName="Meyer" authorityYear="1843" box="[982,1243,1645,1662]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="sismondai">
<emphasis id="CA936E0EFFAAFF99DAC6F9C2FBF62A5F" box="[982,1131,1645,1662]" italics="true" pageId="10" pageNumber="21">Cancer sismondai</emphasis>
<bibRefCitation id="9C76CFEDFFAAFF99DD61F9C2FB462A5F" author="MEYER H. VON" box="[1137,1243,1645,1662]" pageId="10" pageNumber="21" pagination="579 - 590" refId="ref10902" refString="MEYER H. VON 1843. - Briefliche Mittheilungen an Prof. Bronn gerichtet. Neues Jahrbuch fur Mineralogie, Geognosie, Geologie und Petrefaktenkunde 1843: 579 - 590. https: // www. biodiversitylibrary. org / page / 36300129" type="journal article" year="1843">Meyer, 1843</bibRefCitation>
</taxonomicName>
: MNHN.F.A51761, dorsal and ventral views (
<emphasis id="CA936E0EFFAAFF99D80FF92BFEB12AB4" bold="true" box="[287,300,1668,1685]" pageId="10" pageNumber="21">A</emphasis>
,
<emphasis id="CA936E0EFFAAFF99D827F92BFED92AB4" bold="true" box="[311,324,1668,1685]" pageId="10" pageNumber="21">B</emphasis>
); MNHN.F.A51741, dorsal and ventral views (
<emphasis id="CA936E0EFFAAFF99DBAFF92BFD502AB4" bold="true" box="[703,717,1668,1685]" pageId="10" pageNumber="21">C</emphasis>
,
<emphasis id="CA936E0EFFAAFF99DBC7F92BFD782AB4" bold="true" box="[727,741,1668,1685]" pageId="10" pageNumber="21">D</emphasis>
); MNHN.F.A51740, ventral view (
<emphasis id="CA936E0EFFAAFF99DAEAF92BFB9B2AB4" bold="true" box="[1018,1030,1668,1685]" pageId="10" pageNumber="21">E</emphasis>
);
<emphasis id="CA936E0EFFAAFF99DD05F92BFBBD2AB4" bold="true" box="[1045,1056,1668,1685]" pageId="10" pageNumber="21">F</emphasis>
,
<taxonomicName id="3FE7C99FFFAAFF99DD3AF92BFB142AB4" box="[1066,1161,1668,1685]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="CA936E0EFFAAFF99DD3AF92BFBFB2AB4" box="[1066,1126,1668,1685]" italics="true" pageId="10" pageNumber="21">Cancer</emphasis>
sp.
</taxonomicName>
, MNHN.F.A51742, outer view of P1 chela. Scale bars: 2 cm. Photographs: Lilian Cazes.
</paragraph>
</caption>
<subSubSection id="B0FDE197FFAAFF98DA3DF8FDFD622E11" lastPageId="11" lastPageNumber="22" pageId="10" pageNumber="21" type="discussion">
<paragraph id="F858B21CFFAAFF99DA3DF8FDFC332B4B" blockId="10.[811,1455,1874,2027]" box="[813,942,1874,1900]" pageId="10" pageNumber="21">
<smallCapsWord id="FEBE24C0FFAAFF99DA3DF8FDFC332B4B" baselines="1893,1893" box="[813,942,1874,1900]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Discussion" pageId="10" pageNumber="21">DISCUSSION</smallCapsWord>
</paragraph>
<paragraph id="F858B21CFFAAFF98DA3DF8DDFEDF2D30" blockId="10.[811,1455,1874,2027]" lastBlockId="11.[130,776,215,561]" lastPageId="11" lastPageNumber="22" pageId="10" pageNumber="21">
<bibRefCitation id="9C76CFEDFFAAFF99DA3DF8DDFC662BAA" author="VAN STRAELEN V." box="[813,1019,1905,1932]" pageId="10" pageNumber="21" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen (1937)</bibRefCitation>
figured only the
<typeStatus id="275C0CBEFFAAFF99DDB6F8DEFA9F2BAA" box="[1190,1282,1905,1931]" pageId="10" pageNumber="21" type="holotype">holotype</typeStatus>
but the
<typeStatus id="275C0CBEFFAAFF99DC45F8DDFA322BAD" box="[1365,1455,1906,1932]" pageId="10" pageNumber="21" type="paratype">paratype</typeStatus>
is better preserved and shows some additional characters such as the wide cup-shaped postorbital spine, the well-developed intestinal region beyond and under the inflated branchial regions, and the posterolateral margins with a row of large aligned tubercles.
</paragraph>
<paragraph id="F858B21CFFABFF98D98BFEB7FD622E11" blockId="11.[130,776,215,561]" pageId="11" pageNumber="22">
Extant members of the
<taxonomicName id="3FE7C99FFFABFF98D89EFEB8FD8C2D10" authorityName="Garth" authorityYear="1958" box="[398,529,279,305]" class="Malacostraca" family="Oregoniidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="22" phylum="Arthropoda" rank="family">Oregoniidae</taxonomicName>
are relatively rare in the Mediterranean Basin and only
<taxonomicName id="3FE7C99FFFABFF98D8C2FE97FF212D50" authority="A. Milne-Edwards, 1882" authorityName="A. Milne-Edwards" authorityYear="1882" class="Malacostraca" family="Oregoniidae" genus="Ergasticus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFABFF98D8C2FE97FDA82D70" box="[466,565,312,337]" italics="true" pageId="11" pageNumber="22">Ergasticus</emphasis>
A. Milne-Edwards, 1882
</taxonomicName>
has been reported (
<bibRefCitation id="9C76CFEDFFABFF98D86EFEF8FE642D50" author="GBIF" box="[382,505,343,369]" pageId="11" pageNumber="22" pagination="2022" refId="ref10039" refString="GBIF 2022. - Ergasticus A. Milne-Edwards, 1882 in GBIF Secretariat (2022). GBIF Backbone Taxonomy. Checklist dataset https: // doi. org / 10.15468 / 39 omei accessed via GBIF. org on 2023 - 04 - 25." type="journal article" year="2022">GBIF 2022</bibRefCitation>
:https://www.gbif.org/species/4375129). In the fossil record,
<taxonomicName id="3FE7C99FFFABFF98D8F7FED8FF242D90" authority="Glaessner, 1928" authorityName="Glaessner" authorityYear="1928" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="CA936E0EFFABFF98D8F7FED8FD052DB0" box="[487,664,375,401]" italics="true" pageId="11" pageNumber="22">Hyas meridionalis</emphasis>
Glaessner, 1928
</taxonomicName>
was described from the Miocene of
<collectingCountry id="80F0F28CFFABFF98DB20FE39FDE32D91" box="[560,638,406,432]" name="Austria" pageId="11" pageNumber="22">Austria</collectingCountry>
, but the
<typeStatus id="275C0CBEFFABFF98DBC7FE38FC982D90" box="[727,773,407,433]" pageId="11" pageNumber="22">type</typeStatus>
specimen is poorly preserved and, even if it is surely a member of
<taxonomicName id="3FE7C99FFFABFF98D98FFE79FE982DD1" authorityName="Samouelle" authorityYear="1819" box="[159,261,470,496]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="29" phylum="Arthropoda" rank="superFamily" superFamily="Majoidea">Majoidea</taxonomicName>
, the generic assignment seems to be doubtful. In conclusion,
<taxonomicName id="3FE7C99FFFABFF98D815FE59FEC32E31" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[261,350,502,528]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CA936E0EFFABFF98D815FE59FEC32E31" box="[261,350,502,528]" italics="true" pageId="11" pageNumber="22">Parahyas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A0D375FFABFF98D875FE58FE322E30" box="[357,431,503,529]" pageId="11" pageNumber="22" rank="genus">n. gen.</taxonomicNameLabel>
is most likely the only Mediterranean representative of the
<taxonomicName id="3FE7C99FFFABFF98D8BCFDB9FDB22E11" authorityName="Garth" authorityYear="1958" box="[428,559,534,560]" class="Malacostraca" family="Oregoniidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="22" phylum="Arthropoda" rank="family">Oregoniidae</taxonomicName>
in the fossil record.
</paragraph>
</subSubSection>
</treatment>
</document>
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