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<emphasis id="CA936E0EFFA5FF96DA8BF9DDFBF12AAC" bold="true" box="[923,1132,1650,1677]" italics="true" pageId="5" pageNumber="16">Calappa saheliensis</emphasis>
<bibRefCitation id="9C76CFEDFFA5FF96DD63F9DCFADD2AAD" author="VAN STRAELEN V." box="[1139,1344,1650,1677]" pageId="5" pageNumber="16" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937</bibRefCitation>
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(
<figureCitation id="60DCAE99FFA5FF96DD27F93DFBE82A8D" box="[1079,1141,1682,1708]" captionStart="FIG" captionStartId="6.[132,142,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-26@6.[698,1267,768,1324]" captionTargetPageId="6" captionText="FIG. 2. — Brachyurancrabs from thelate Messinian of Oran(Les Planteursquarry,Arambourg coll.):A-D,Calappa saheliensis Van Straelen,1937:A,holotype MNHN.F.R03768, dorsal view;B, paratype MNHN.F.A28898, dorsal view;C, paratype MNHN.F.A28899, dorsal view;D, paratype MNHN.F.A28902, dorsal view;E, F, Wahrania arambourgi (Van Straelen, 1937), n. comb. holotype MNHN.F.R03773, dorsal view and line drawing of carapace. Abbreviations: b, branchiocardiac groove; ca, cardiac region; i, intestinal region; ms, mesogastric region; mt, metagastric region; o, orbit; u, urogastric region; 1, pseudorostral spine (= rostrum); 2, postorbital spine; 3, hepatic spine; 3.1, accessorial spine of hepatic spine;4, epibranchial spine;5, mesobranchial spine;6, 7, metabranchial spines. Scale bars: 1 cm. Photographs: Lilian Cazes." figureDoi="http://doi.org/10.5281/zenodo.10566284" httpUri="https://zenodo.org/record/10566284/files/figure.png" pageId="5" pageNumber="16">Fig. 2</figureCitation>
A-D)
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<emphasis id="CA936E0EFFA5FF96DA3DF967FC7D2AC1" box="[813,992,1736,1760]" italics="true" pageId="5" pageNumber="16">Calappa saheliensis</emphasis>
<bibRefCitation id="9C76CFEDFFA5FF96DAF6F966FB492AC1" author="VAN STRAELEN V." box="[998,1236,1736,1760]" pageId="5" pageNumber="16" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937: 473</bibRefCitation>
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, 474, pl. 33, fig. 1. —
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<collectingCountry id="80F0F28CFFA5FF96DA3DF94DFC362ADB" box="[813,939,1762,1786]" name="Sint Martin" pageId="5" pageNumber="16">Saint Martin</collectingCountry>
&amp; Müller 1988: 251
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. —
<treatmentCitation id="7946940DFFA5FF96DDBDF94DFCCB2B34" author="MOISSETTE P. &amp; MULLER P." page="738" pageId="5" pageNumber="16" year="1990">
<bibRefCitation id="9C76CFEDFFA5FF96DDBDF94DFCCB2B34" author="MOISSETTE P. &amp; MULLER P." pageId="5" pageNumber="16" pagination="737 - 747" refId="ref11138" refString="MOISSETTE P. &amp; MULLER P. 1990. - Crustaces Decapodes des facies marno-diatomitique du Messinien d'Oranie (Algerie Occidentale). Geobios 23 (6): 737 - 747. https: // doi. org / 10.1016 / S 0016 - 6995 (06) 80339 - 6" type="journal article" year="1990">Moissette &amp; Müller 1990: 738</bibRefCitation>
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. —
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Mayoral
<emphasis id="CA936E0EFFA5FF96DAC9F951FB942B34" box="[985,1033,1789,1813]" italics="true" pageId="5" pageNumber="16">et al.</emphasis>
1998: 508
</bibRefCitation>
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, fig. 2.4. —
<treatmentCitation id="7946940DFFA5FF96DDE8F952FC082B11" author="DIAZ-MEDINA G. &amp; HYZNY M. &amp; KLOMPMAKER A. A." page="181" pageId="5" pageNumber="16" year="2017">
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Díaz-Medina
<emphasis id="CA936E0EFFA5FF96DC90F951FA2D2B34" box="[1408,1456,1789,1813]" italics="true" pageId="5" pageNumber="16">et al.</emphasis>
2017: 181
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. —
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, 371.
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<taxonomicName id="3FE7C99FFFA5FF96DA3DF8EBFC442B7D" authorityName="Van Straelen" authorityYear="1936" box="[813,985,1860,1884]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="sahelensis">
<emphasis id="CA936E0EFFA5FF96DA3DF8EBFC442B7D" box="[813,985,1860,1884]" italics="true" pageId="5" pageNumber="16">Calappa sahelensis</emphasis>
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[sic]
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Schweitzer
<emphasis id="CA936E0EFFA5FF96DD80F8EAFB222B7D" box="[1168,1215,1860,1884]" italics="true" pageId="5" pageNumber="16">et al.</emphasis>
2010: 83
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.
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<taxonomicName id="3FE7C99FFFA5FF96DA3DF8DEFC672BA8" ID-CoL="BSB7L" authorityName="Van Straelen" authorityYear="1937" box="[813,1018,1905,1929]" class="Malacostraca" family="Calappidae" genus="Calappa" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA5FF96DA3DF8DEFCE12BA8" box="[813,892,1905,1929]" italics="true" pageId="5" pageNumber="16">Calappa</emphasis>
cf.
<emphasis id="CA936E0EFFA5FF96DA8CF8DEFC672BA8" box="[924,1018,1905,1929]" italics="true" pageId="5" pageNumber="16">saheliensis</emphasis>
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Mayoral
<emphasis id="CA936E0EFFA5FF96DD79F8DEFB0A2BA8" box="[1129,1175,1905,1929]" italics="true" pageId="5" pageNumber="16">et al.</emphasis>
1998: 508
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, fig. 2.3.
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(
<emphasis id="CA936E0EFFA5FF96DC73F8DEFA1A2BA9" box="[1379,1415,1905,1928]" italics="true" pageId="5" pageNumber="16">non</emphasis>
<taxonomicName id="3FE7C99FFFA5FF96DC9CF8DEFB152B82" ID-CoL="BSB7L" authority="Van Straelen, 1936" authorityName="Van Straelen" authorityYear="1936" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA5FF96DC9CF8DEFC592B82" italics="true" pageId="5" pageNumber="16">Calappa saheliensis</emphasis>
Van Straelen, 1936
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA5FF94DA3DF817FEB02D2B" lastPageId="7" lastPageNumber="18" pageId="5" pageNumber="16" type="materials_examined">
<paragraph id="F858B21CFFA5FF96DA3DF817FA2C2BCB" blockId="5.[813,1457,1976,2026]" pageId="5" pageNumber="16">
<materialsCitation id="488FB841FFA5FF96DA3DF817FB632BF1" box="[813,1278,1976,2000]" collectionCode="MNHN" collectorName="Arambourg" country="Algeria" county="Sanaouber" location="Les Planteurs quarry" municipality="Sidi El Houari" pageId="5" pageNumber="16" specimenCode="MNHN.F.R03768" specimenCount="1" typeStatus="holotype">
<smallCapsWord id="FEBE24C0FFA5FF96DA3DF817FCC12BEF" baselines="1993,1994" box="[813,860,1976,1999]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="5" pageNumber="16">TYPE</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA5FF96DA73F812FC202BEF" baselines="1994" box="[867,957,1981,1998]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="5" pageNumber="16">MATERIAL</smallCapsWord>
. —
<typeStatus id="275C0CBEFFA5FF96DAFBF817FBD92BF1" box="[1003,1092,1976,2000]" pageId="5" pageNumber="16" type="holotype">Holotype</typeStatus>
<specimenCode id="A8411A67FFA5FF96DD5CF817FB632BF1" box="[1100,1278,1976,2000]" pageId="5" pageNumber="16">MNHN.F.R03768</specimenCode>
</materialsCitation>
and
<materialsCitation id="488FB841FFA5FF96DC3FF817FA2C2BCB" collectionCode="MNHN" collectorName="Arambourg" country="Algeria" county="Sanaouber" location="Les Planteurs quarry" municipality="Sidi El Houari" pageId="5" pageNumber="16" specimenCode="MNHN.F.A28898, A28899, A28900, A28901, A28902" specimenCount="5" typeStatus="paratype">
<specimenCount id="EEE17995FFA5FF96DC3FF817FACD2BF1" box="[1327,1360,1976,2000]" pageId="5" pageNumber="16" type="generic">five</specimenCount>
<typeStatus id="275C0CBEFFA5FF96DC47F817FA322BF1" box="[1367,1455,1976,2000]" pageId="5" pageNumber="16">paratypes</typeStatus>
<specimenCode id="A8411A67FFA5FF96DA3EF87CFC4F2BCB" box="[814,978,2002,2026]" pageId="5" pageNumber="16">MNHN.F.A28898</specimenCode>
,
<specimenCode id="A8411A67FFA5FF96DACBF87DFBBC2BCB" box="[987,1057,2002,2026]" pageId="5" pageNumber="16">A28899</specimenCode>
,
<specimenCode id="A8411A67FFA5FF96DD3AF87DFBED2BCB" box="[1066,1136,2002,2026]" pageId="5" pageNumber="16">A28900</specimenCode>
,
<specimenCode id="A8411A67FFA5FF96DD69F87DFB222BCB" box="[1145,1215,2002,2026]" pageId="5" pageNumber="16">A28901</specimenCode>
,
<specimenCode id="A8411A67FFA5FF96DDD8F87DFA8D2BCB" box="[1224,1296,2002,2026]" pageId="5" pageNumber="16">A28902</specimenCode>
(
<collectorName id="5512D7CAFFA5FF96DC07F87DFAE12BCB" box="[1303,1404,2002,2026]" pageId="5" pageNumber="16">Arambourg</collectorName>
coll.).
</materialsCitation>
</paragraph>
<caption id="AC98E294FFA6FF95D994F8D2FA302BC8" ID-DOI="http://doi.org/10.5281/zenodo.10566284" ID-Zenodo-Dep="10566284" httpUri="https://zenodo.org/record/10566284/files/figure.png" pageId="6" pageNumber="17" startId="6.[132,142,1917,1934]" targetBox="[132,1455,215,1878]" targetPageId="6" targetType="figure">
<paragraph id="F858B21CFFA6FF95D994F8D2FA302BC8" blockId="6.[132,1457,1917,2025]" pageId="6" pageNumber="17">
<smallCapsWord id="FEBE24C0FFA6FF95D994F8D2FF002BAF" baselines="1930,1931" box="[132,157,1917,1934]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="17">FIG</smallCapsWord>
. 2. — Brachyuran crabs from the late
<taxonomicName id="3FE7C99FFFA6FF95D8A9F8D2FD9A2BAF" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[441,519,1917,1934]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
of Oran(Les Planteurs quarry,Arambourg coll.):
<emphasis id="CA936E0EFFA6FF95DA62F8D2FC0E2BAF" bold="true" box="[882,915,1917,1934]" pageId="6" pageNumber="17">A-D</emphasis>
,
<taxonomicName id="3FE7C99FFFA6FF95DA8AF8D2FB5B2BAF" authority="Van Straelen, 1937" authorityName="Van Straelen" authorityYear="1937" box="[922,1222,1917,1934]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA6FF95DA8AF8D2FBAC2BAF" box="[922,1073,1917,1934]" italics="true" pageId="6" pageNumber="17">Calappa saheliensis</emphasis>
<bibRefCitation id="9C76CFEDFFA6FF95DD24F8D2FB5B2BAF" author="VAN STRAELEN V." box="[1076,1222,1917,1934]" pageId="6" pageNumber="17" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen,1937</bibRefCitation>
</taxonomicName>
:
<emphasis id="CA936E0EFFA6FF95DDDAF8D2FB4A2BAF" bold="true" box="[1226,1239,1917,1934]" pageId="6" pageNumber="17">A</emphasis>
,holotype MNHN.F.R03768, dorsal view;
<emphasis id="CA936E0EFFA6FF95D9F5F83BFF6F2B84" bold="true" box="[229,242,1940,1957]" pageId="6" pageNumber="17">B</emphasis>
, paratype MNHN.F.A28898, dorsal view;
<emphasis id="CA936E0EFFA6FF95DB28F83BFDDB2B84" bold="true" box="[568,582,1940,1957]" pageId="6" pageNumber="17">C</emphasis>
, paratype MNHN.F.A28899, dorsal view;
<emphasis id="CA936E0EFFA6FF95DA9BF83BFC042B84" bold="true" box="[907,921,1940,1957]" pageId="6" pageNumber="17">D</emphasis>
, paratype MNHN.F.A28902, dorsal view;
<emphasis id="CA936E0EFFA6FF95DDCEF83BFB602B84" bold="true" box="[1246,1277,1940,1957]" pageId="6" pageNumber="17">E, F</emphasis>
,
<taxonomicName id="3FE7C99FFFA6FF95DC15F83BFEB62B9D" authority="(Van Straelen, 1937)" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" baseAuthorityName="Van Straelen" baseAuthorityYear="1937" class="Malacostraca" family="Goneplacidae" genus="Wahrania" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="arambourgi" status="comb. nov.">
<emphasis id="CA936E0EFFA6FF95DC15F83BFA322B84" box="[1285,1455,1940,1957]" italics="true" pageId="6" pageNumber="17">Wahrania arambourgi</emphasis>
(
<bibRefCitation id="9C76CFEDFFA6FF95D998F804FEBB2B9D" author="VAN STRAELEN V." box="[136,294,1963,1980]" pageId="6" pageNumber="17" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="D1A0D375FFA6FF95D823F804FEE02B9D" box="[307,381,1963,1980]" pageId="6" pageNumber="17" rank="species">n. comb.</taxonomicNameLabel>
holotype MNHN.F.R03773, dorsal view and line drawing of carapace. Abbreviations:
<emphasis id="CA936E0EFFA6FF95DD26F804FBDC2B9D" bold="true" box="[1078,1089,1963,1980]" pageId="6" pageNumber="17">b</emphasis>
, branchiocardiac groove;
<emphasis id="CA936E0EFFA6FF95DC06F804FAB62B9D" bold="true" box="[1302,1323,1963,1980]" pageId="6" pageNumber="17">ca</emphasis>
, cardiac region;
<emphasis id="CA936E0EFFA6FF95D994F86DFF142BF2" bold="true" box="[132,137,1986,2003]" pageId="6" pageNumber="17">i</emphasis>
, intestinal region;
<emphasis id="CA936E0EFFA6FF95D80DF86DFEA52BF2" bold="true" box="[285,312,1986,2003]" pageId="6" pageNumber="17">ms</emphasis>
, mesogastric region;
<emphasis id="CA936E0EFFA6FF95D8F8F86DFE622BF2" bold="true" box="[488,511,1986,2003]" pageId="6" pageNumber="17">mt</emphasis>
, metagastric region;
<emphasis id="CA936E0EFFA6FF95DBBBF86DFD2B2BF2" bold="true" box="[683,694,1986,2003]" pageId="6" pageNumber="17">o</emphasis>
, orbit;
<emphasis id="CA936E0EFFA6FF95DBE1F86DFD612BF2" bold="true" box="[753,764,1986,2003]" pageId="6" pageNumber="17">u</emphasis>
, urogastric region;
<emphasis id="CA936E0EFFA6FF95DA8AF86DFC392BF2" bold="true" box="[922,932,1986,2003]" pageId="6" pageNumber="17">1</emphasis>
, pseudorostral spine (= rostrum);
<emphasis id="CA936E0EFFA6FF95DDA8F86DFB5F2BF2" bold="true" box="[1208,1218,1986,2003]" pageId="6" pageNumber="17">2</emphasis>
, postorbital spine;
<emphasis id="CA936E0EFFA6FF95DC4EF86DFAF52BF2" bold="true" box="[1374,1384,1986,2003]" pageId="6" pageNumber="17">3</emphasis>
, hepatic spine;
<emphasis id="CA936E0EFFA6FF95D9A9F877FF4F2BC8" bold="true" box="[185,210,2008,2025]" pageId="6" pageNumber="17">3.1</emphasis>
, accessorial spine of hepatic spine;
<emphasis id="CA936E0EFFA6FF95D8E4F877FE632BC8" bold="true" box="[500,510,2008,2025]" pageId="6" pageNumber="17">4</emphasis>
, epibranchial spine;
<emphasis id="CA936E0EFFA6FF95DBB3F877FD302BC8" bold="true" box="[675,685,2008,2025]" pageId="6" pageNumber="17">5</emphasis>
, mesobranchial spine;
<emphasis id="CA936E0EFFA6FF95DA77F877FCEC2BC8" bold="true" box="[871,881,2008,2025]" pageId="6" pageNumber="17">6</emphasis>
,
<emphasis id="CA936E0EFFA6FF95DA6BF877FC182BC8" bold="true" box="[891,901,2008,2025]" pageId="6" pageNumber="17">7</emphasis>
, metabranchial spines. Scale bars: 1 cm. Photographs: Lilian Cazes.
</paragraph>
</caption>
<paragraph id="F858B21CFFA7FF94D994FF77FEB02D2B" blockId="7.[132,775,215,266]" pageId="7" pageNumber="18">
<smallCapsWord id="FEBE24C0FFA7FF94D994FF77FF282CCC" baselines="233,233" box="[132,181,216,239]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="18">TYPE</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA7FF94D9ABFF72FE8B2CCF" baselines="234" box="[187,278,221,238]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="7" pageNumber="18">LOCALITY</smallCapsWord>
. —
<location id="FD38E4C7FFA7FF94D851FF77FD942CD1" LSID="urn:lsid:plazi:treatment:704E030AFFA5FF94DA8BF9DDFE8E2BCA:FD38E4C7FFA7FF94D851FF77FD942CD1" box="[321,521,215,240]" country="Algeria" county="Sanaouber" municipality="Sidi El Houari" name="Les Planteurs quarry" pageId="7" pageNumber="18">Les Planteurs quarry</location>
,
<collectingCounty id="1139CA90FFA7FF94DB02FF78FDE52CCE" box="[530,632,215,239]" pageId="7" pageNumber="18">Sanaouber</collectingCounty>
district of
<collectingMunicipality id="183C2866FFA7FF94DBF1FF78FF752D2B" pageId="7" pageNumber="18">Sidi El Houari</collectingMunicipality>
,
<collectingCountry id="80F0F28CFFA7FF94D9E2FF5DFEB42D2B" box="[242,297,242,266]" name="Algeria" pageId="7" pageNumber="18">Oran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA7FF94D994FE8DFDA12D1B" box="[132,572,290,314]" pageId="7" pageNumber="18" type="biology_ecology">
<paragraph id="F858B21CFFA7FF94D994FE8DFDA12D1B" blockId="7.[132,572,290,314]" box="[132,572,290,314]" pageId="7" pageNumber="18">
<smallCapsWord id="FEBE24C0FFA7FF94D994FE8DFF282D16" baselines="307,307" box="[132,181,290,313]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="18">
<typeStatus id="275C0CBEFFA7FF94D994FE8DFF282D16" box="[132,181,290,313]" pageId="7" pageNumber="18">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="FEBE24C0FFA7FF94D9ACFE88FF7C2D19" baselines="308" box="[188,225,295,312]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="age" pageId="7" pageNumber="18">AGE</smallCapsWord>
. — Late Miocene (late
<taxonomicName id="3FE7C99FFFA7FF94D8DAFE8DFDAF2D1B" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[458,562,290,314]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA7FF94D994FEF8FEC92F6E" pageId="7" pageNumber="18" type="description">
<paragraph id="F858B21CFFA7FF94D994FEF8FE882D4E" blockId="7.[131,776,343,847]" box="[132,277,343,369]" pageId="7" pageNumber="18">
<smallCapsWord id="FEBE24C0FFA7FF94D994FEF8FE882D4E" baselines="362,362" box="[132,277,343,369]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="18">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="F858B21CFFA7FF94D994FED8FF792DB0" blockId="7.[131,776,343,847]" box="[132,228,375,401]" pageId="7" pageNumber="18">
<emphasis id="CA936E0EFFA7FF94D994FED8FF792DB0" box="[132,228,375,401]" italics="true" pageId="7" pageNumber="18">Carapace</emphasis>
</paragraph>
<paragraph id="F858B21CFFA7FF94D994FE38FEC92F6E" blockId="7.[131,776,343,847]" pageId="7" pageNumber="18">Carapace subovate, strongly vaulted longitudinally and transversely, with spined posterolateral flanges; maximum width at level of third posterolateral tooth; front poorly preserved; orbits small, rounded; supraorbital margin entire; anterolateral margins strongly arcuate; posterolateral margins strongly convex; flange extensions with five triangular teeth: three teeth equal in size, followed by two teeth decreasing in size posteriorly; posterior margin short, slightly convex; carapace surface covered by small tubercles densely arranged uniformly; tubercles more scattered along posterior margin; central regions bounded longitudinally by deep two parallel grooves running from orbits to cardiac region; small tubercle between metagastric and urogastric regions.</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA7FF94D994FCDAFE8E2BCA" pageId="7" pageNumber="18" type="discussion">
<paragraph id="F858B21CFFA7FF94D994FCDAFE912FAC" blockId="7.[131,776,885,2027]" box="[132,268,885,911]" pageId="7" pageNumber="18">
<smallCapsWord id="FEBE24C0FFA7FF94D994FCDAFE912FAC" baselines="904,904" box="[132,268,885,911]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Discussion" pageId="7" pageNumber="18">DISCUSSION</smallCapsWord>
</paragraph>
<paragraph id="F858B21CFFA7FF94D994FC3AFECF28AF" blockId="7.[131,776,885,2027]" pageId="7" pageNumber="18">
<bibRefCitation id="9C76CFEDFFA7FF94D994FC3AFEC02F8E" author="VAN STRAELEN V." box="[132,349,917,943]" pageId="7" pageNumber="18" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen (1937)</bibRefCitation>
reported
<specimenCount id="EEE17995FFA7FF94D8D6FC3AFDAF2F8E" box="[454,562,917,943]" count="8" pageId="7" pageNumber="18" type="generic">eight type</specimenCount>
specimens partially decorticated. The check of the type material highlighted that
<specimenCount id="EEE17995FFA7FF94D9ACFC7AFEC52FCE" box="[188,344,981,1007]" count="2" pageId="7" pageNumber="18" type="generic" typeStatus="paratype">two paratypes</specimenCount>
seem to be lost. Though
<bibRefCitation id="9C76CFEDFFA7FF94DB6AFC7AFF4F282E" author="VAN STRAELEN V." pageId="7" pageNumber="18" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen (1937)</bibRefCitation>
did not provide a detailed description of this species, he assigned it to
<taxonomicName id="3FE7C99FFFA7FF94D861FBBBFE51280F" authorityName="Weber" authorityYear="1795" box="[369,460,1044,1070]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA7FF94D861FBBBFE51280F" box="[369,460,1044,1070]" italics="true" pageId="7" pageNumber="18">Calappa</emphasis>
</taxonomicName>
without justification. Based on
<bibRefCitation id="9C76CFEDFFA7FF94D9BAFB9BFD8E286F" author="SCHWEITZER C. E. &amp; FELDMANN R. M." box="[170,531,1076,1102]" pageId="7" pageNumber="18" pagination="1 - 8" refId="ref11872" refString="SCHWEITZER C. E. &amp; FELDMANN R. M. 2019. - Part R, Revised, Volume 1, Chapter 8 T 4: Systematic descriptions: Superfamily Calappoidea. Treatise Online 121: 1 - 8. https: // doi. org / 10.17161 / to. v 0 i 0.11742" type="journal article" year="2019">Schweitzer &amp; Feldmann (2019)</bibRefCitation>
, we concur with this systematic assignment for the presence of the spined posterolateral flanges.
</paragraph>
<paragraph id="F858B21CFFA7FF94D98BFB3BFF7B2A4C" blockId="7.[131,776,885,2027]" pageId="7" pageNumber="18">
Regarding the Miocene of the Mediterranean and Paratethys,
<taxonomicName id="3FE7C99FFFA7FF94D9C9FB1BFEA928EF" authorityName="Weber" authorityYear="1795" box="[217,308,1204,1230]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA7FF94D9C9FB1BFEA928EF" box="[217,308,1204,1230]" italics="true" pageId="7" pageNumber="18">Calappa</emphasis>
</taxonomicName>
is represented by three species,
<taxonomicName id="3FE7C99FFFA7FF94DBBCFB1BFEF628CF" authority="Brocchi, 1883" authorityName="Brocchi" authorityYear="1883" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="heberti">
<emphasis id="CA936E0EFFA7FF94DBBCFB1BFF5628CF" italics="true" pageId="7" pageNumber="18">Calappa heberti</emphasis>
Brocchi, 1883
</taxonomicName>
,
<taxonomicName id="3FE7C99FFFA7FF94D864FB7BFDC628CF" authority="Lorenthey" authorityName="Lorenthey" box="[372,603,1236,1262]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="praelata">
<emphasis id="CA936E0EFFA7FF94D864FB7BFE7B28CF" box="[372,486,1236,1262]" italics="true" pageId="7" pageNumber="18">C. praelata</emphasis>
Lőrenthey
</taxonomicName>
<emphasis id="CA936E0EFFA7FF94DB70FB7BFDEB28CC" box="[608,630,1236,1261]" italics="true" pageId="7" pageNumber="18">in</emphasis>
Lőrenthey &amp; Beurlen, 1929, and
<taxonomicName id="3FE7C99FFFA7FF94D870FB5BFE70292C" authorityName="Van Straelen" authorityYear="1937" box="[352,493,1267,1294]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D870FB5BFE70292C" box="[352,493,1267,1294]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
. As pointed out by
<bibRefCitation id="9C76CFEDFFA7FF94DBD8FB5BFEF1290F" author="DIAZ-MEDINA G. &amp; HYZNY M. &amp; KLOMPMAKER A. A." pageId="7" pageNumber="18" pagination="173 - 195" refId="ref9681" refString="DIAZ-MEDINA G., HYZNY M. &amp; KLOMPMAKER A. A. 2017. - A lithostratigraphic and palaeoenvironmental framework for the late Miocene El Caracolar section (Granada Basin, Betic Cordillera, Spain). Geobios 50: 173 - 195. https: // doi. org / 10.1016 / j. geobios. 2017.04.003" type="journal article" year="2017">
Díaz-Medina
<emphasis id="CA936E0EFFA7FF94D9F2FABBFE85290C" box="[226,280,1299,1325]" italics="true" pageId="7" pageNumber="18">et al.</emphasis>
(2017)
</bibRefCitation>
the last two species are morphologically very close to each other. However, the reappraisal of the
<typeStatus id="275C0CBEFFA7FF94D9A2FAFBFF7F294F" box="[178,226,1364,1390]" pageId="7" pageNumber="18">type</typeStatus>
series of
<taxonomicName id="3FE7C99FFFA7FF94D842FAFBFE79294C" authorityName="Van Straelen" authorityYear="1937" box="[338,484,1363,1390]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D842FAFBFE79294C" box="[338,484,1363,1390]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
has pointed out that the carapace regions are covered by small tubercles densely and uniformly arranged (
<emphasis id="CA936E0EFFA7FF94D87EFA3BFE31298C" box="[366,428,1428,1453]" italics="true" pageId="7" pageNumber="18">versus</emphasis>
large tubercles in these regions in
<taxonomicName id="3FE7C99FFFA7FF94D9B3FA1CFE8129EC" box="[163,284,1459,1485]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="praelata">
<emphasis id="CA936E0EFFA7FF94D9B3FA1CFE8129EC" box="[163,284,1459,1485]" italics="true" pageId="7" pageNumber="18">C. praelata</emphasis>
</taxonomicName>
).
<taxonomicName id="3FE7C99FFFA7FF94D825FA1CFE7E29EC" authorityName="Brocchi" authorityYear="1883" box="[309,483,1459,1485]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="heberti">
<emphasis id="CA936E0EFFA7FF94D825FA1CFE7E29EC" box="[309,483,1459,1485]" italics="true" pageId="7" pageNumber="18">Calappa heberti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="3FE7C99FFFA7FF94DB68FA1CFC9A29EC" authorityName="Van Straelen" authorityYear="1937" box="[632,775,1459,1485]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94DB68FA1CFC9A29EC" box="[632,775,1459,1485]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
in having large tubercles in the anterior half of the carapace and short squamous ridges in the posterior portion. Therefore, based on the different carapace ornamentations,
<taxonomicName id="3FE7C99FFFA7FF94D993F99CFE8E2A6D" authorityName="Van Straelen" authorityYear="1937" box="[131,275,1586,1613]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D993F99CFE8E2A6D" box="[131,275,1586,1613]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
can be considered as a valid species within
<taxonomicName id="3FE7C99FFFA7FF94D996F9FDFF422A4D" authorityName="Weber" authorityYear="1795" box="[134,223,1618,1644]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA7FF94D996F9FDFF422A4D" box="[134,223,1618,1644]" italics="true" pageId="7" pageNumber="18">Calappa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F858B21CFFA7FF94D98BF9DDFE8E2BCA" blockId="7.[131,776,885,2027]" pageId="7" pageNumber="18">
<bibRefCitation id="9C76CFEDFFA7FF94D98BF9DDFE112AAD" author="MAYORAL E. &amp; MULLER P. &amp; MUNIZ F." box="[155,396,1650,1676]" pageId="7" pageNumber="18" pagination="505 - 510" refId="ref10841" refString="MAYORAL E., MULLER P. &amp; MUNIZ F. 1998. - Lower Pliocene decapod crustaceans from the southwestern Iberian Peninsula (Guadalquivir Basin, Sevilla, Spain). Geobios 31 (4): 505 - 510. https: // doi. org / 10.1016 / S 0016 - 6995 (98) 80122 - 8" type="journal article" year="1998">
Mayoral
<emphasis id="CA936E0EFFA7FF94D811F9DCFEA52AAD" box="[257,312,1650,1676]" italics="true" pageId="7" pageNumber="18">et al.</emphasis>
(1998)
</bibRefCitation>
reported an incomplete calappid crab from the early Pliocene of Guadalquivir Basin (
<collectingCountry id="80F0F28CFFA7FF94DBAEF93DFD622A8D" box="[702,767,1682,1708]" name="Spain" pageId="7" pageNumber="18">Spain</collectingCountry>
) identified as
<taxonomicName id="3FE7C99FFFA7FF94D807F91DFD962AED" authorityName="Van Straelen" authorityYear="1937" box="[279,523,1714,1740]" class="Malacostraca" family="Calappidae" genus="Calappa" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D807F91DFEEF2AED" box="[279,370,1714,1740]" italics="true" pageId="7" pageNumber="18">Calappa</emphasis>
cf.
<emphasis id="CA936E0EFFA7FF94D88DF91DFD962AED" box="[413,523,1714,1740]" italics="true" pageId="7" pageNumber="18">saheliensis</emphasis>
</taxonomicName>
, based on the smooth ornamentation of the posterior part of the carapace. It is difficult, however, to assert that this poorly preserved specimen could be related to
<taxonomicName id="3FE7C99FFFA7FF94D8FAF8BDFD1F2B0D" authorityName="Van Straelen" authorityYear="1937" box="[490,642,1810,1836]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D8FAF8BDFD1F2B0D" box="[490,642,1810,1836]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
simply for the smooth ornamentation, lacking the main diagnostic characters of this species, such as orbits, antero- and posterolateral margins, urogastric and cardiac regions, and spined posterolateral flanges. In conclusion, we limit the stratigraphic range of
<taxonomicName id="3FE7C99FFFA7FF94D896F81DFD842BEA" authorityName="Van Straelen" authorityYear="1937" box="[390,537,1969,1996]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="saheliensis">
<emphasis id="CA936E0EFFA7FF94D896F81DFD842BEA" box="[390,537,1969,1996]" italics="true" pageId="7" pageNumber="18">C. saheliensis</emphasis>
</taxonomicName>
to the late Miocene (
<taxonomicName id="3FE7C99FFFA7FF94D99CF87EFE9A2BCA" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[140,263,2001,2027]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="13" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Messinian">Messinian</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>