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<document id="76B6088D7F6F17223B4D70ACAE57CB45" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626381872300" checkinUser="felipe" docAuthor="Oláh, János, Johanson, Kjell Arne &amp; Barnard, Peter C." docDate="2008" docId="627D87E1FFD1F735FF7EFE06FF097977" docLanguage="en" docName="zt01738p171.pdf" docOrigin="Zootaxa 1738" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cheumatopsyche congolana Olah &amp; Johanson" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="9E44FF99FFFEF71AFFE9FF9BFFD07D08" masterDocTitle="Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)" masterLastPageNumber="171" masterPageNumber="1" pageNumber="48" updateTime="1699011095215" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E386FF648B6C1F549A669C3D07FB7234">Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)</mods:title>
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<mods:namePart id="5748EDBB361158FFD9097495FDF38DEB">Oláh, János</mods:namePart>
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<mods:namePart id="A646537F57235CFF81D6BCA6B51D307D">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="F2E46BF355E27D745E178A49F9116957">Barnard, Peter C.</mods:namePart>
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<mods:title id="175C6E73D6A2FB399251AC34B6B038CC">Zootaxa</mods:title>
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<mods:date id="C6FA8B5C4627BD11E1317117B5D60A11">2008</mods:date>
<mods:detail id="0143E52EA48667CC15992AE8B21AD08E" type="pubDate">
<mods:number id="A1863F78E4A5B3B44CA9291AA99D090C">2008-03-31</mods:number>
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<mods:number id="5A9ED2C6F9CBAFB2EB3D4FA81CB09BEC">1738</mods:number>
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<mods:start id="7BA7CA9710F71FADF052F7D04EBB880E">1</mods:start>
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<treatment id="627D87E1FFD1F735FF7EFE06FF097977" ID-DOI="http://doi.org/10.5281/zenodo.5108474" ID-Zenodo-Dep="5108474" LSID="urn:lsid:plazi:treatment:627D87E1FFD1F735FF7EFE06FF097977" httpUri="http://treatment.plazi.org/id/627D87E1FFD1F735FF7EFE06FF097977" lastPageNumber="48" pageId="47" pageNumber="48">
<subSubSection id="A2CE657CFFD1F735FF7EFE06FC5F7CBF" box="[151,911,413,439]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="EA6B36F7FFD1F735FF7EFE06FC5F7CBF" blockId="47.[151,911,413,479]" box="[151,911,413,439]" pageId="47" pageNumber="48">
<heading id="B123819BFFD1F735FF7EFE06FC5F7CBF" bold="true" box="[151,911,413,439]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="D8A0EAE5FFD1F735FF7EFE06FC5F7CBF" bold="true" box="[151,911,413,439]" pageId="47" pageNumber="48">
<taxonomicName id="2DD44D74FFD1F735FF7EFE06FD017CBF" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" box="[151,721,413,439]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FF7EFE06FE3C7CBF" bold="true" box="[151,492,413,439]" italics="true" pageId="47" pageNumber="48">Cheumatopsyche congolana</emphasis>
Oláh &amp; Johanson
</taxonomicName>
, nomen novum
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFD1F735FF7EFE5EFEBD7CD7" box="[151,365,453,479]" pageId="47" pageNumber="48" type="description">
<paragraph id="EA6B36F7FFD1F735FF7EFE5EFEBD7CD7" blockId="47.[151,911,413,479]" box="[151,365,453,479]" pageId="47" pageNumber="48">
<figureCitation id="72EF2A72FFD1F735FF7EFE5EFF2A7CD7" box="[151,250,453,479]" captionStart="FIGURES 99–108" captionStartId="39.[151,269,1260,1284]" captionTargetBox="[173,1399,188,1225]" captionTargetId="figure-148@39.[151,1436,168,1236]" captionTargetPageId="39" captionText="FIGURES 99–108. Right forewing of Cheumatopsyche species in species groups 4–7, with shape and distribution of wing pattern. 99 — Cheumatopsyche camilla Malicky, 1997; 100 — Cheumatopsyche congolana Oláh &amp; Johanson, new name; 101 — Cheumatopsyche albofasciata (McLachlan, 1872); 102 — Cheumatopsyche doan Oláh &amp; Johanson, new species; 103 — Cheumatopsyche dubitans Mosely, 1942; 104 — Cheumatopsyche falcifera (Ulmer, 1930); 105 — Cheumatopsyche nohondoan Oláh &amp; Johanson, new species; 106 — Cheumatopsyche quadrifasciata Oláh &amp; Johanson, new species; 107 — Cheumatopsyche sexfasciata (Ulmer, 1904); 108 — Cheumatopsyche chirapali Oláh &amp; Johanson, new species." pageId="47" pageNumber="48">Fig. 100</figureCitation>
,
<figureCitation id="72EF2A72FFD1F735FEEEFE5EFEBD7CD7" box="[263,365,453,479]" captionStart="FIGURES 121–122" captionStartId="48.[151,269,1166,1190]" captionTargetBox="[528,1039,460,1133]" captionTargetId="figure-302@48.[467,1120,444,1158]" captionTargetPageId="48" captionText="FIGURES 121–122. Cheumatopsyche congolana Oláh &amp; Johanson, new name, holotype. 121 — genitalia, dorsal; 122 — gonocoxite, ventral." pageId="47" pageNumber="48">121–122</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFD1F735FF7EFD8EFD697F24" box="[151,697,533,556]" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="EA6B36F7FFD1F735FF7EFD8EFD697F24" blockId="47.[151,697,533,556]" box="[151,697,533,556]" pageId="47" pageNumber="48">
<treatmentCitationGroup id="CAC411D9FFD1F735FF7EFD8EFD697F24" box="[151,697,533,556]" pageId="47" pageNumber="48">
<treatmentCitation id="6B7510E6FFD1F735FF7EFD8EFD657F24" author="Jacquemart, S." box="[151,693,533,556]" page="45" pageId="47" pageNumber="48" year="1966">
<taxonomicName id="2DD44D74FFD1F735FF7EFD8EFD657F24" authority="Jacquemart, 1966: 45" authorityName="Jacquemart" authorityPageNumber="45" authorityYear="1966" box="[151,693,533,556]" class="Insecta" family="Hydropsychidae" genus="Hydropsychodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="D8A0EAE5FFD1F735FF7EFD8EFE167F24" box="[151,454,533,556]" italics="true" pageId="47" pageNumber="48">Hydropsychodes bimaculata</emphasis>
<bibRefCitation id="8E454B06FFD1F735FE24FD8EFD657F24" author="Jacquemart, S." box="[461,693,533,556]" pageId="47" pageNumber="48" pagination="35 - 56" refId="ref63282" refString="Jacquemart, S. (1966) Trichoptera. Exploration du Parc National de la Garamba. Mission H. de Saeger (1949 - 1952), 51 (2), 35 - 56." type="journal article" year="1966">Jacquemart, 1966: 45</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFD1F735FF7EFDC6FF097977" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="EA6B36F7FFD1F735FF7EFDC6FE5D7F7F" blockId="47.[151,1436,605,1151]" box="[151,397,605,631]" pageId="47" pageNumber="48">
<materialsCitation id="5ABC3CAAFFD1F735FF7EFDC6FE5A7F7F" box="[151,394,605,631]" country="Democratic Republic of the Congo" location="Zaire" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">
<emphasis id="D8A0EAE5FFD1F735FF7EFDC6FE907F7F" bold="true" box="[151,320,605,631]" pageId="47" pageNumber="48">
<typeStatus id="356F8855FFD1F735FF7EFDC6FF047F7F" box="[151,212,605,631]" pageId="47" pageNumber="48">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="92C37667FFD1F735FEA1FDC6FE5A7F7F" box="[328,394,605,631]" name="Democratic Republic of the Congo" pageId="47" pageNumber="48">Zaïre</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="EA6B36F7FFD1F735FF2FFD1EFF097977" blockId="47.[151,1436,605,1151]" pageId="47" pageNumber="48">
Because
<taxonomicName id="2DD44D74FFD1F735FEDBFD1EFE247F96" authorityName="Ulmer" authorityYear="1905" box="[306,500,645,670]" class="Insecta" family="Hydropsychidae" genus="Hydropsychodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="D8A0EAE5FFD1F735FEDBFD1EFE247F96" box="[306,500,645,670]" italics="true" pageId="47" pageNumber="48">Hydropsychodes</emphasis>
</taxonomicName>
is now in synonymy with
<emphasis id="D8A0EAE5FFD1F735FCD3FD1EFBD97F96" box="[826,1033,645,670]" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="2DD44D74FFD1F735FCD3FD1EFBD57F96" box="[826,1029,645,670]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">Cheumatopsyche</taxonomicName>
,
</emphasis>
<taxonomicName id="2DD44D74FFD1F735FBFAFD1EFE917FCF" authority="Jacquemart, 1966" authorityName="Jacquemart" authorityYear="1966" class="Insecta" family="Hydropsychidae" genus="Hydropsychodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="D8A0EAE5FFD1F735FBFAFD1EFAB37F96" box="[1043,1379,645,670]" italics="true" pageId="47" pageNumber="48">Hydropsychodes bimaculata</emphasis>
<bibRefCitation id="8E454B06FFD1F735FA84FD1EFE917FCF" author="Jacquemart, S." pageId="47" pageNumber="48" pagination="35 - 56" refId="ref63282" refString="Jacquemart, S. (1966) Trichoptera. Exploration du Parc National de la Garamba. Mission H. de Saeger (1949 - 1952), 51 (2), 35 - 56." type="journal article" year="1966">Jacquemart, 1966</bibRefCitation>
</taxonomicName>
is a junior homonym of
<taxonomicName id="2DD44D74FFD1F735FD85FD36FBA47FCF" authority="(Ulmer, 1930)" baseAuthorityName="Ulmer" baseAuthorityYear="1930" box="[620,1140,685,711]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="D8A0EAE5FFD1F735FD85FD36FC117FCE" box="[620,961,685,710]" italics="true" pageId="47" pageNumber="48">Cheumatopsyche bimaculata</emphasis>
(Ulmer, 1930)
</taxonomicName>
, and should be renamed, as suggested by
<bibRefCitation id="8E454B06FFD1F735FEBCFD4EFE3A7FE7" author="Scott, K. M. F." box="[341,490,725,751]" pageId="47" pageNumber="48" pagination="299 - 422" refId="ref65964" refString="Scott, K. M. F. (1983) On the Hydropsychidae (Trichoptera) of Southern Africa with keys to African genera of imagos, larvae and pupae and species lists. Annals of the Cape Province Museum (Natural History), 14 (8), 299 - 422." type="journal article" year="1983">Scott (1983)</bibRefCitation>
.
<taxonomicName id="2DD44D74FFD1F735FE1CFD4EFC907FE6" box="[501,832,725,750]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FE1CFD4EFC907FE6" box="[501,832,725,750]" italics="true" pageId="47" pageNumber="48">Cheumatopsyche congolana</emphasis>
</taxonomicName>
is here proposed as a new replacement name for
<taxonomicName id="2DD44D74FFD1F735FA96FD4EFE247E1F" authority="Jacquemart, 1966" authorityName="Jacquemart" authorityYear="1966" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="D8A0EAE5FFD1F735FA96FD4EFECB7E1E" italics="true" pageId="47" pageNumber="48">H. bimaculata</emphasis>
<bibRefCitation id="8E454B06FFD1F735FECBFD66FE247E1F" author="Jacquemart, S." box="[290,500,765,791]" pageId="47" pageNumber="48" pagination="35 - 56" refId="ref63282" refString="Jacquemart, S. (1966) Trichoptera. Exploration du Parc National de la Garamba. Mission H. de Saeger (1949 - 1952), 51 (2), 35 - 56." type="journal article" year="1966">Jacquemart, 1966</bibRefCitation>
</taxonomicName>
. Although
<taxonomicName id="2DD44D74FFD1F735FD9CFD66FCCB7E1E" authorityName="Jacquemart" authorityYear="1966" box="[629,795,765,790]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="D8A0EAE5FFD1F735FD9CFD66FCCB7E1E" box="[629,795,765,790]" italics="true" pageId="47" pageNumber="48">C. bimaculata</emphasis>
</taxonomicName>
was described from the female only, its special forewing venation, including a sessile fork I, differentiates it from
<taxonomicName id="2DD44D74FFD1F735FC90FCBEFA927E37" authority="Olah &amp; Johanson. Most" authorityName="Olah &amp; Johanson. Most" box="[889,1346,805,831]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FC90FCBEFBC97E36" box="[889,1049,805,830]" italics="true" pageId="47" pageNumber="48">C. congolana</emphasis>
Oláh &amp; Johanson. Most
</taxonomicName>
importantly however is the completely different pattern of the forewings. The male genitalia of
<taxonomicName id="2DD44D74FFD1F735FB55FCD6FA8B7E6E" box="[1212,1371,845,870]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FB55FCD6FA8B7E6E" box="[1212,1371,845,870]" italics="true" pageId="47" pageNumber="48">C. congolana</emphasis>
</taxonomicName>
were not illustrated, but the general similarity between genitalia of
<taxonomicName id="2DD44D74FFD1F735FC9CFCEEFBC57E86" box="[885,1045,885,910]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FC9CFCEEFBC57E86" box="[885,1045,885,910]" italics="true" pageId="47" pageNumber="48">C. congolana</emphasis>
</taxonomicName>
and
<taxonomicName id="2DD44D74FFD1F735FBB9FCEEFAC97E87" authority="(Mosely)" baseAuthorityName="Mosely" baseAuthorityYear="1935" box="[1104,1305,885,911]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="D8A0EAE5FFD1F735FBB9FCEEFB757E86" box="[1104,1189,885,910]" italics="true" pageId="47" pageNumber="48">C. afra</emphasis>
(Mosely)
</taxonomicName>
was stated by
<bibRefCitation id="8E454B06FFD1F735FF52FC06FE497EBF" author="Jacquemart, S." box="[187,409,925,951]" pageId="47" pageNumber="48" pagination="35 - 56" refId="ref63282" refString="Jacquemart, S. (1966) Trichoptera. Exploration du Parc National de la Garamba. Mission H. de Saeger (1949 - 1952), 51 (2), 35 - 56." type="journal article" year="1966">Jacquemart (1966)</bibRefCitation>
. However, examination of the genitalia of the
<typeStatus id="356F8855FFD1F735FC5FFC06FBCC7EBF" box="[950,1052,925,951]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CC5AE32FFD1F735FBC2FC06FB557EBF" box="[1067,1157,925,951]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="47" pageNumber="48" type="Museum">MRAC</collectionCode>
) mounted in permanent microscope slide and although very distorted, flattened and deformed, revealed that
<taxonomicName id="2DD44D74FFD1F735FB69FC5EFACF7ED6" box="[1152,1311,965,990]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FB69FC5EFACF7ED6" box="[1152,1311,965,990]" italics="true" pageId="47" pageNumber="48">C. congolana</emphasis>
</taxonomicName>
has rather short apicoventral setal lobes on segment X (
<figureCitation id="72EF2A72FFD1F735FD4EFC76FCDB790F" box="[679,779,1005,1031]" captionStart="FIGURES 121–122" captionStartId="48.[151,269,1166,1190]" captionTargetBox="[528,1039,460,1133]" captionTargetId="figure-302@48.[467,1120,444,1158]" captionTargetPageId="48" captionText="FIGURES 121–122. Cheumatopsyche congolana Oláh &amp; Johanson, new name, holotype. 121 — genitalia, dorsal; 122 — gonocoxite, ventral." pageId="47" pageNumber="48">Fig. 121</figureCitation>
) and belongs to the
<emphasis id="D8A0EAE5FFD1F735FC10FC76FB16790E" box="[1017,1222,1005,1030]" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="2DD44D74FFD1F735FC10FC76FBBA790E" authorityName="Gibbs" authorityYear="1973" box="[1017,1130,1005,1030]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="lestoni">C. lestoni</taxonomicName>
species
</emphasis>
group and not the
<taxonomicName id="2DD44D74FFD1F735FF7EFB8EFECE7926" authorityName="Mosely" authorityYear="1942" box="[151,286,1045,1070]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dubitans">
<emphasis id="D8A0EAE5FFD1F735FF7EFB8EFECE7926" box="[151,286,1045,1070]" italics="true" pageId="47" pageNumber="48">C. dubitans</emphasis>
</taxonomicName>
species group.
<taxonomicName id="2DD44D74FFD1F735FE3AFB8EFD037926" baseAuthorityName="Mosely" baseAuthorityYear="1935" box="[467,723,1045,1070]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="D8A0EAE5FFD1F735FE3AFB8EFD037926" box="[467,723,1045,1070]" italics="true" pageId="47" pageNumber="48">Cheumatopsyche afra</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="2DD44D74FFD1F735FC6FFB8EFBDD7926" authorityName="Mosely" authorityYear="1942" box="[902,1037,1045,1070]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dubitans">
<emphasis id="D8A0EAE5FFD1F735FC6FFB8EFBDD7926" box="[902,1037,1045,1070]" italics="true" pageId="47" pageNumber="48">C. dubitans</emphasis>
</taxonomicName>
species group due to the presence of longer apicoventral lobes.
<taxonomicName id="2DD44D74FFD1F735FE13FBA6FC98795E" box="[506,840,1085,1110]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="congolana">
<emphasis id="D8A0EAE5FFD1F735FE13FBA6FC98795E" box="[506,840,1085,1110]" italics="true" pageId="47" pageNumber="48">Cheumatopsyche congolana</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="2DD44D74FFD1F735FB4FFBA6FF057977" authority="Marlier, 1961" authorityName="Marlier" authorityYear="1961" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tenerima">
<emphasis id="D8A0EAE5FFD1F735FB4FFBA6FAE3795E" box="[1190,1331,1085,1110]" italics="true" pageId="47" pageNumber="48">C. tenerima</emphasis>
<bibRefCitation id="8E454B06FFD1F735FAD7FBA6FF057977" author="Marlier, G." pageId="47" pageNumber="48" pagination="158 - 212" refId="ref64100" refString="Marlier, G. (1961) Hydropsychidae du Kivu. Revue de Zoologie et Botanique de l'Afrique, 63 (1 - 2), 158 - 212." type="journal article" year="1961">Marlier, 1961</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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