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<document id="EABC9D847A44E967940534961EA36B17" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626381872300" checkinUser="felipe" docAuthor="Oláh, János, Johanson, Kjell Arne &amp; Barnard, Peter C." docDate="2008" docId="627D87E1FFB2F755FF7EF964FD247CFA" docLanguage="en" docName="zt01738p171.pdf" docOrigin="Zootaxa 1738" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cheumatopsyche sambava Olah &amp; Johanson 2008, new species" docType="treatment" docVersion="2" lastPageNumber="80" masterDocId="9E44FF99FFFEF71AFFE9FF9BFFD07D08" masterDocTitle="Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)" masterLastPageNumber="171" masterPageNumber="1" pageNumber="77" updateTime="1699011095215" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F1AAF16266C80E3A3487D8D2836D5880">Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)</mods:title>
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<mods:name id="90597638CA67D9DEAFB76D2C1E6F19C6" type="personal">
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<mods:namePart id="4F29CCBFC834BD5B3C6120B012C7F6EF">Oláh, János</mods:namePart>
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<mods:namePart id="E0415C9FF54F283EAECF34ACE6D4E8C3">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="580A54F30405F05F1A2624D37234C259">Barnard, Peter C.</mods:namePart>
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<mods:title id="5AB7ECA99B14A8351D484055CBD5DCDB">Zootaxa</mods:title>
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<mods:part id="8680EC829B85C33534EC55B55C628787">
<mods:date id="C676B76266D74E51B05134ABBDF8FDFC">2008</mods:date>
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<mods:number id="47A6DFE8342454478847B9E6B8AC967C">2008-03-31</mods:number>
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<mods:number id="20572A8A2E8E19D40417781BD10D3A3C">1738</mods:number>
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<mods:identifier id="D5DE53971821610A917B813977557B9C" type="ISSN">1175­5334</mods:identifier>
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<treatment id="627D87E1FFB2F755FF7EF964FD247CFA" LSID="urn:lsid:plazi:treatment:627D87E1FFB2F755FF7EF964FD247CFA" httpUri="http://treatment.plazi.org/id/627D87E1FFB2F755FF7EF964FD247CFA" lastPageId="79" lastPageNumber="80" pageId="76" pageNumber="77">
<subSubSection id="A2CE657CFFB2F756FF7EF964FC887A11" box="[151,856,1791,1817]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="EA6B36F7FFB2F756FF7EF964FC887A11" blockId="76.[151,856,1791,1858]" box="[151,856,1791,1817]" pageId="76" pageNumber="77">
<heading id="B123819BFFB2F756FF7EF964FC887A11" bold="true" box="[151,856,1791,1817]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="D8A0EAE5FFB2F756FF7EF964FC887A11" bold="true" box="[151,856,1791,1817]" pageId="76" pageNumber="77">
<taxonomicName id="2DD44D74FFB2F756FF7EF964FD6B7A11" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[151,699,1791,1817]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sambava" status="sp. nov.">
<emphasis id="D8A0EAE5FFB2F756FF7EF964FE077A11" bold="true" box="[151,471,1791,1817]" italics="true" pageId="76" pageNumber="77">Cheumatopsyche sambava</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="C393579EFFB2F756FD21F964FC887A11" box="[712,856,1791,1817]" pageId="76" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFB2F755FF7EF8B3FC687C5A" lastPageId="79" lastPageNumber="80" pageId="76" pageNumber="77" type="description">
<paragraph id="EA6B36F7FFB2F756FF7EF8B3FEE37A4A" blockId="76.[151,856,1791,1858]" box="[151,307,1832,1858]" pageId="76" pageNumber="77">
<figureCitation id="72EF2A72FFB2F756FF7EF8B3FEE37A4A" box="[151,307,1832,1858]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="76" pageNumber="77">Fig. 191194</figureCitation>
</paragraph>
<paragraph id="EA6B36F7FFB2F756FF7EF8E3FA4B7AEA" blockId="76.[151,1436,1912,2018]" pageId="76" pageNumber="77">
This species is similar to
<taxonomicName id="2DD44D74FFB2F756FE54F8E3FD9B7A99" authority="Oláh &amp; Johanson &amp; Barnard, 2008" authorityName="Oláh &amp; Johanson &amp; Barnard" authorityYear="2008" box="[445,587,1912,1937]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="ampanga" status="sp. nov.">
<emphasis id="D8A0EAE5FFB2F756FE54F8E3FD9B7A99" box="[445,587,1912,1937]" italics="true" pageId="76" pageNumber="77">C. ampanga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C393579EFFB2F756FDBBF8E3FD317A9A" box="[594,737,1912,1938]" pageId="76" pageNumber="77" rank="species">new species</taxonomicNameLabel>
, from which it is easily distinguished by its smaller size and the forewings withou pattern. Segment IX and dorsocaudal spiny lobe are both shorter, and apices of the apicoventral lobes are characteristically recurved mesad in dorsal view, and recurved dorsad in lateral view. The
</paragraph>
<paragraph id="EA6B36F7FFB3F757FF7EFF03FE147DD2" blockId="77.[151,1435,152,218]" pageId="77" pageNumber="78">harpagones are not sigmoid, but are nearly straight, with broad bases in ventral view; and the phallotheca is constricted at mid-length.</paragraph>
<caption id="BEAB667FFFB3F757FF7EF918FCBF7BB5" pageId="77" pageNumber="78" startId="77.[151,269,1667,1691]" targetBox="[175,1392,286,1600]" targetPageId="77">
<paragraph id="EA6B36F7FFB3F757FF7EF918FCBF7BB5" blockId="77.[151,1436,1667,1725]" pageId="77" pageNumber="78">
<emphasis id="D8A0EAE5FFB3F757FF7EF918FEA97B93" bold="true" box="[151,377,1667,1691]" pageId="77" pageNumber="78">FIGURES 187190.</emphasis>
<taxonomicName id="2DD44D74FFB3F757FE69F918FC917B92" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[384,833,1667,1690]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="rineta" status="sp. nov.">
<emphasis id="D8A0EAE5FFB3F757FE69F918FDAD7B92" box="[384,637,1667,1690]" italics="true" pageId="77" pageNumber="78">Cheumatopsyche rineta</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="C393579EFFB3F757FCA5F918FC007B92" box="[844,976,1667,1690]" pageId="77" pageNumber="78" rank="species">new species</taxonomicNameLabel>
, holotype. 187 — genitalia, lateral; 188 — genitalia, dorsal; 189 — gonocoxite, ventral; 190 — phallus, lateral.
</paragraph>
</caption>
<paragraph id="EA6B36F7FFB3F757FF2FF96AFF397ADB" blockId="77.[151,1436,1777,2003]" pageId="77" pageNumber="78">
Male. Body and wing membrane brown with light pubescence; wing without pattern. Maxillary palp segment I shortest, segment II longer, segments III and IV equally long, segment V as long as sum of segments I IV. Head dorsum dark brown, with 5 warts of same dark colour. Swollen setal wart absent on proepisternum. Protarsal claws asymmetrical, laterally flanked by setal bundle. Each protibiae with 2 spurs. Forewing length
<quantity id="2D2C9B12FFB3F757FF7EF80AFF287AA3" box="[151,248,1937,1963]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="77" pageNumber="78" unit="mm" value="4.1">4.1 mm</quantity>
, hind wing length
<quantity id="2D2C9B12FFB3F757FE08F80AFD927AA3" box="[481,578,1937,1963]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="77" pageNumber="78" unit="mm" value="3.4">3.4 mm</quantity>
. Forewing crossveins m-cu and cu almost touching. Hind wing fork I absent.
</paragraph>
<caption id="BEAB667FFFB0F754FF7EF9B8FC417B55" pageId="78" pageNumber="79" startId="78.[151,269,1571,1595]" targetBox="[182,1413,191,1534]" targetPageId="78">
<paragraph id="EA6B36F7FFB0F754FF7EF9B8FC417B55" blockId="78.[151,1436,1571,1629]" pageId="78" pageNumber="79">
<emphasis id="D8A0EAE5FFB0F754FF7EF9B8FEA97B33" bold="true" box="[151,377,1571,1595]" pageId="78" pageNumber="79">FIGURES 191194.</emphasis>
<taxonomicName id="2DD44D74FFB0F754FE69F9BFFCB17B33" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[384,865,1572,1595]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sambava" status="sp. nov.">
<emphasis id="D8A0EAE5FFB0F754FE69F9BFFD4D7B33" box="[384,669,1572,1595]" italics="true" pageId="78" pageNumber="79">Cheumatopsyche sambava</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="C393579EFFB0F754FC85F9BFFC217B33" box="[876,1009,1572,1595]" pageId="78" pageNumber="79" rank="species">new species</taxonomicNameLabel>
, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral.
</paragraph>
</caption>
<paragraph id="EA6B36F7FFB0F755FF2EF90AFC687C5A" blockId="78.[151,1436,1681,2027]" lastBlockId="79.[151,1436,152,498]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
Male genitalia. Segment IX fused annularly, generally short; tergum about one-third as long as sternum (
<figureCitation id="72EF2A72FFB0F754FF76F922FED87BDB" box="[159,264,1721,1747]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 191</figureCitation>
); anterior margins of segment IX regularly convex in lateral view, tergal part less excised, flatter than sternal; anterosternal margin produced anterad opposite to base of coxopodites (
<figureCitation id="72EF2A72FFB0F754FB92F97AFB317BF3" box="[1147,1249,1761,1787]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 191</figureCitation>
); apical lobe on posterolateral margins broadly triangular, short, located well above coxopodite bases, about at mid-height of segment; spine row on posterior margins of segment IX interrupted between dorsocaudal spiny lobe and apical lobe (
<figureCitation id="72EF2A72FFB0F754FEEAF8C2FEB97A7B" box="[259,361,1881,1907]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 191</figureCitation>
). Intersegmental area between segment IX and segment X deeply stepped. Segment X relatively short, with tuboid apicoventral setal lobes bending dorsad in lateral view (
<figureCitation id="72EF2A72FFB0F754FBBBF81AFB697A93" box="[1106,1209,1921,1947]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 191</figureCitation>
); curving mesad in dorsal view (
<figureCitation id="72EF2A72FFB0F754FED8F832FE4A7ACB" box="[305,410,1961,1987]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 192</figureCitation>
); lobes widely separated in dorsal view (
<figureCitation id="72EF2A72FFB0F754FC69F832FC387ACB" box="[896,1000,1961,1987]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 192</figureCitation>
). Transverse sutures crossing central part of segment X (
<figureCitation id="72EF2A72FFB0F754FE92F84AFE317AE3" box="[379,481,2001,2027]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="78" pageNumber="79">Fig. 191</figureCitation>
). Longitudinal sutures not visible. Setaless mesocaudal lobe forming nearly mar- ginally straight plate in dorsal view (
<figureCitation id="72EF2A72FFB1F755FDABFF03FD787DBA" box="[578,680,152,178]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="79" pageNumber="80">Fig. 192</figureCitation>
). Pair of lateral setose areas forming elevated warts, located well beyond transverse sutures at base of apicoventral setal lobes. Coxopodites exceeding beyond apex of segment X, almost straight, with dilated apex. Harpagones slender, finger-like, curving dorsad in lateral view (
<figureCitation id="72EF2A72FFB1F755FA85FF73FF167C22" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="79" pageNumber="80">Fig. 191</figureCitation>
); bases broad in ventral view (
<figureCitation id="72EF2A72FFB1F755FDC2FE8BFD427C22" box="[555,658,272,298]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="79" pageNumber="80">Fig. 193</figureCitation>
). Phallotheca (
<figureCitation id="72EF2A72FFB1F755FCD7FE8BFC727C22" box="[830,930,272,298]" captionStart="FIGURES 191194" captionStartId="78.[151,269,1571,1595]" captionTargetBox="[182,1413,191,1534]" captionTargetId="figure-189@78.[151,1436,158,1587]" captionTargetPageId="78" captionText="FIGURES 191194. Cheumatopsyche sambava Oláh &amp; Johanson, new species, holotype. 191 — genitalia, lateral; 192 — genitalia, dorsal; 193 — gonocoxite, ventral; 194 — phallus, lateral." pageId="79" pageNumber="80">Fig. 194</figureCitation>
) strongly constricted at mid-length; sclerotised endothecal process elongate; phallotremal sclerites small, oval.
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFB1F755FF2EFEC4FE0A7CAA" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="EA6B36F7FFB1F755FF2EFEC4FE0A7CAA" blockId="79.[151,1436,152,498]" pageId="79" pageNumber="80">
<materialsCitation id="5ABC3CAAFFB1F755FF2EFEC4FE0A7CAA" collectingDate="1959-11" collectionCode="MNHN" collectorName="P. Soga" country="Madagascar" location="Est de Sambava Reserve" municipality="Marojejy-Andrakata" pageId="79" pageNumber="80" specimenCount="1" typeStatus="holotype">
<emphasis id="D8A0EAE5FFB1F755FF2EFEC4FDA37C71" bold="true" box="[199,627,351,377]" pageId="79" pageNumber="80">
<typeStatus id="356F8855FFB1F755FF2EFEC4FEE97C71" box="[199,313,351,377]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="92C37667FFB1F755FE7BFEC4FDBE7C71" box="[402,622,351,377]" name="Madagascar" pageId="79" pageNumber="80">MADAGASCAR</collectingCountry>
:
</emphasis>
<location id="EF0B602CFFB1F755FD95FEFBFC737C72" LSID="urn:lsid:plazi:treatment:627D87E1FFB2F755FF7EF964FD247CFA:EF0B602CFFB1F755FD95FEFBFC737C72" box="[636,931,352,378]" country="Madagascar" municipality="Marojejy-Andrakata" name="Est de Sambava Reserve" pageId="79" pageNumber="80">Est de Sambava Reserve</location>
nat. XII,
<collectingMunicipality id="0A0FAC8DFFB1F755FBF2FEFBFADD7C72" box="[1051,1293,352,378]" pageId="79" pageNumber="80">Marojejy-Andrakata</collectingMunicipality>
,
<date id="9E6A1037FFB1F755FAF5FEFBFAA47C72" box="[1308,1396,352,378]" pageId="79" pageNumber="80" value="1959-11">
<collectingDate id="8E2EE9DFFFB1F755FAF5FEFBFAA47C72" box="[1308,1396,352,378]" pageId="79" pageNumber="80" value="1959-11">xi.1959</collectingDate>
</date>
[
<collectorName id="47215321FFB1F755FA6CFEFBFF047CAA" pageId="79" pageNumber="80">P. Soga</collectorName>
] (
<collectionCode id="8CC5AE32FFB1F755FF02FE13FE997CAA" box="[235,329,392,418]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
, in alcohol).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFB1F755FF2EFE34FDD57CC2" box="[199,517,431,458]" pageId="79" pageNumber="80" type="distribution">
<paragraph id="EA6B36F7FFB1F755FF2EFE34FDD57CC2" blockId="79.[151,1436,152,498]" box="[199,517,431,458]" pageId="79" pageNumber="80">
<emphasis id="D8A0EAE5FFB1F755FF2EFE34FEB97CC1" bold="true" box="[199,361,431,457]" pageId="79" pageNumber="80">Distribution:</emphasis>
<collectingCountry id="92C37667FFB1F755FE98FE2BFDD17CC2" box="[369,513,432,458]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFFB1F755FF2EFE4CFD247CFA" box="[199,756,471,498]" pageId="79" pageNumber="80" type="etymology">
<paragraph id="EA6B36F7FFB1F755FF2EFE4CFD247CFA" blockId="79.[151,1436,152,498]" box="[199,756,471,498]" pageId="79" pageNumber="80">
<emphasis id="D8A0EAE5FFB1F755FF2EFE4CFE877CF9" bold="true" box="[199,343,471,497]" pageId="79" pageNumber="80">Etymology:</emphasis>
from the name of the
<typeStatus id="356F8855FFB1F755FDB5FE43FD5E7CFA" box="[604,654,472,498]" pageId="79" pageNumber="80">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>