610 lines
68 KiB
XML
610 lines
68 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.947.51259" ID-GBIF-Dataset="b9b0ed44-7490-48f1-8cc1-cee431f5d1ff" ID-PMC="PMC7363713" ID-Pensoft-Pub="1313-2970-947-71" ID-Pensoft-UUID="8E93BC6985D85A52AD26A514847748FA" ID-PubMed="32733130" ID-ZooBank="3297FBE4111C48499533225A53F7DB3C" ModsDocID="1313-2970-947-71" checkinTime="1594285657262" checkinUser="pensoft" docAuthor="Hrivniak, Ľubos, Sroka, Pavel, Bojkova, Jindriska, Godunko, Roman J., Namin, Javid Imanpour, Bagheri, Samereh, Nejat, Farshad, Abdoli, Ashgar & Staniczek, Arnold H." docDate="2020" docId="622A6F8DBAD95BBAAC28119F4D70AD06" docLanguage="en" docName="ZooKeys 947: 71-102" docOrigin="ZooKeys 947" docSource="http://dx.doi.org/10.3897/zookeys.947.51259" docTitle="Epeorus (Caucasiron) alborzicus Hrivniak & Sroka 2020, sp. nov." docType="treatment" docUuid="F1721BB2-DC7C-4BBC-9AD2-8252A5D01EBF" docUuidSource="ZooBank" docVersion="4" id="8E93BC6985D85A52AD26A514847748FA" lastPageNumber="71" masterDocId="8E93BC6985D85A52AD26A514847748FA" masterDocTitle="Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) in Iran, with descriptions of three new species" masterLastPageNumber="102" masterPageNumber="71" pageNumber="71" updateTime="1668169075919" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) in Iran, with descriptions of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hrivniak, Ľubos</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Faculty of Sciences, University of South Bohemia, Branisovska 31, 37005 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="email">lubos.hrivniak@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sroka, Pavel</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4367-6564</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bojkova, Jindriska</mods:namePart>
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<mods:affiliation>Department of Botany and Zoology, Masaryk University, Kotlarska 2, 61137 Brno, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Department of Invertebrate Zoology and Hydrobiology, University of Lodz, Banacha 12 / 16, 90237 Lodz, Poland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-2196-3327</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Namin, Javid Imanpour</mods:namePart>
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<mods:affiliation>Department of Fishery, Faculty of Natural Resources, University of Guilan, POB 1144, Sowmehsara-Rasht, Iran</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bagheri, Samereh</mods:namePart>
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<mods:affiliation>Department of Fishery, Faculty of Natural Resources, University of Guilan, POB 1144, Sowmehsara-Rasht, Iran</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nejat, Farshad</mods:namePart>
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<mods:affiliation>Department of Biodiversity and Ecosystem Management, Environmental Sciences Research Institute, Shahid Beheshti University, Daneshjou Boulevard, 1983969411 Tehran, Iran</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Abdoli, Ashgar</mods:namePart>
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<mods:affiliation>Department of Biodiversity and Ecosystem Management, Environmental Sciences Research Institute, Shahid Beheshti University, Daneshjou Boulevard, 1983969411 Tehran, Iran</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Staniczek, Arnold H.</mods:namePart>
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<mods:affiliation>Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-137X</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>947</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>71</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.947.51259</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.947.51259</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-947-71</mods:identifier>
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<mods:identifier type="ZooBank">3297FBE4111C48499533225A53F7DB3C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8E93BC6985D85A52AD26A514847748FA</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164986641" LSID="urn:lsid:zoobank.org:act:F1721BB2-DC7C-4BBC-9AD2-8252A5D01EBF" httpUri="http://treatment.plazi.org/id/622A6F8DBAD95BBAAC28119F4D70AD06" lastPageNumber="71" pageId="0" pageNumber="71">
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<subSubSection pageId="0" pageNumber="71" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="71">
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||
<taxonomicName LSID="622A6F8D-BAD9-5BBA-AC28-119F4D70AD06" authority="Hrivniak & Sroka" authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) alborzicus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alborzicus" status="sp. nov." subGenus="Caucasiron">Epeorus (Caucasiron) alborzicus Hrivniak & Sroka</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">Figures 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="71" type="type material">
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<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
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: female mature larva: IRAN, Mazandaran Province, Panjab village, unnamed brook (LT of Haraz River);
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<geoCoordinate degrees="36" direction="north" minutes="05" orientation="latitude" precision="1" seconds="52.8" value="36.098">36°05'52.8"N</geoCoordinate>
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,
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<geoCoordinate degrees="052" direction="east" minutes="15" orientation="longitude" precision="1" seconds="16.0" value="52.254444">052°15'16.0"E</geoCoordinate>
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(locality no. 152); 955 m a.s.l.; J.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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, T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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, J. Imanpour Namin, S. Bagheri leg., 9.5.2018, SMNS_EPH_010056.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Paratypes</emphasis>
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: 38 female larvae (3 mounted on slide), 10 male larvae (2 mounted on slide): same data as holotype, SMNS_EPH_010056. DNA extracted from 1 female (code: IR11, stored in EtOH) and 2 males (codes: IR12 and IR14, both stored in EtOH).
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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33 female larvae, 24 male larvae: IRAN, Tehran Province, Zayegan village, Lalan River;
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<geoCoordinate degrees="35" direction="north" minutes="58" orientation="latitude" precision="1" seconds="39.2" value="35.977554">35°58'39.2"N</geoCoordinate>
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,
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<geoCoordinate degrees="051" direction="east" minutes="34" orientation="longitude" precision="1" seconds="56.5" value="51.58236">051°34'56.5"E</geoCoordinate>
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(locality no. 55); 2290 m a.s.l.; A. Staniczek, M. Pallmann, R. J. Godunko, F. Nejat leg., 8.5.2017, SMNS_EPH_007617.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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1 female larva: IRAN, Golestan Province, above Chah-e Ja village, unnamed brook (RT of river flowing to Fazelabad);
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<geoCoordinate degrees="36" direction="north" minutes="40" orientation="latitude" precision="1" seconds="22.8" value="36.673">36°40'22.8"N</geoCoordinate>
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,
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<geoCoordinate degrees="054" direction="east" minutes="46" orientation="longitude" precision="1" seconds="37.9" value="54.777195">054°46'37.9"E</geoCoordinate>
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(locality no. 104); 1450 m a.s.l.; J.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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, T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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, J. Imanpour Namin leg., 27.4.2018. DNA extracted specimen (code: IR13, stored in EtOH).
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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17 female larvae (3 mounted on slide), 6 male larvae: IRAN, Alborz Province, 2.5 km W of Asara village, Karaj River;
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<geoCoordinate degrees="36" direction="north" minutes="01" orientation="latitude" precision="1" seconds="52.1" value="36.03114">36°01'52.1"N</geoCoordinate>
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,
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<geoCoordinate degrees="051" direction="east" minutes="13" orientation="longitude" precision="1" seconds="10.0" value="51.219444">051°13'10.0"E</geoCoordinate>
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(locality no. 58); 1890 m a.s.l.; A. Staniczek, M. Pallmann, F. Nejat leg., 10.5.2017, SMNS_EPH_007627.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">The holotype and 50 paratypes are deposited in SMNS, 50 paratypes (including DNA extracted specimens) are deposited in IECA and 29 paratypes in MMTT_DOE.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="materials_examined">
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<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="71">8 larvae: same data as holotype, SMNS_EPH_010056; young instars or damaged specimens.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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13 larvae: IRAN, Mazandaran Province, NE of Kahrud village, unnamed brook (LT of Haraz River);
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<geoCoordinate degrees="36" direction="north" minutes="03" orientation="latitude" precision="1" seconds="42.7" value="36.06186">36°03'42.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="052" direction="east" minutes="15" orientation="longitude" precision="1" seconds="24.8" value="52.25689">052°15'24.8"E</geoCoordinate>
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(locality no. 153); 1020 m a.s.l.; J.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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, T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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, J. Imanpour Namin, S. Bagheri leg., 9.5.2018.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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2 larvae: IRAN, Mazandaran Province, 3.5 km E of Polour village, Lasem Rud (RT of Haraz River);
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<geoCoordinate degrees="35" direction="north" minutes="50" orientation="latitude" precision="1" seconds="09.4" value="35.83594">35°50'09.4"N</geoCoordinate>
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,
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<geoCoordinate degrees="052" direction="east" minutes="04" orientation="longitude" precision="1" seconds="38.4" value="52.07733">052°04'38.4"E</geoCoordinate>
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(locality no. 73); 2100 m a.s.l.; A. Staniczek, M. Pallmann, F. Nejat leg., 14.5.2017, SMNS_EPH_007680; 17 larvae: S. Bagheri leg., 16.4.2018.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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1 larva: IRAN, Mazandaran Province, 1.5 km S of Part Kola village, Shirin Rud (LT of Sefidrud);
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<geoCoordinate degrees="36" direction="north" minutes="9" orientation="latitude" precision="1" seconds="04.3" value="36.151196">36°9'04.3"N</geoCoordinate>
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,
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<geoCoordinate degrees="053" direction="east" minutes="20" orientation="longitude" precision="1" seconds="54.7" value="53.348526">053°20'54.7"E</geoCoordinate>
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(locality no. 63); 750 m a.s.l.; A. Staniczek, M. Pallmann, F. Nejat leg., 11.5.2017, SMNS_EPH_007641; 10 larvae: S. Bagheri leg., 5.4.2018.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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7 larvae: IRAN, Mazandaran Province, 3.5 km W of Razan village, Baladeh River;
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<geoCoordinate degrees="36" direction="north" minutes="11" orientation="latitude" precision="1" seconds="39.6" value="36.194336">36°11'39.6"N</geoCoordinate>
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,
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<geoCoordinate degrees="052" direction="east" minutes="8" orientation="longitude" precision="1" seconds="34.6" value="52.142944">052°8'34.6"E</geoCoordinate>
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(locality no. 73); 1360 m a.s.l.; A. Staniczek, M. Pallmann, F. Nejat leg., 14.5.2017, SMNS_EPH_007677.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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1 larva: IRAN, Tehran Province, Lalan village, Lalan River;
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<geoCoordinate degrees="35" direction="north" minutes="59" orientation="latitude" precision="1" seconds="50.3" value="35.997307">35°59'50.3"N</geoCoordinate>
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,
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<geoCoordinate degrees="051" direction="east" minutes="34" orientation="longitude" precision="1" seconds="51.0" value="51.580833">051°34'51.0"E</geoCoordinate>
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(locality no. 53); 2438 m a.s.l.; A. Staniczek, M. Pallmann, R. J. Godunko, F. Nejat leg., 8.5.2017, SMNS_EPH_007613.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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17 larvae: IRAN, Tehran Province, Igol village, Fasham River;
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<geoCoordinate degrees="35" direction="north" minutes="55" orientation="latitude" precision="1" seconds="11.2" value="35.91978">35°55'11.2"N</geoCoordinate>
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,
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<geoCoordinate degrees="051" direction="east" minutes="28" orientation="longitude" precision="1" seconds="51.3" value="51.48092">051°28'51.3"E</geoCoordinate>
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(locality no. 56); 2020 m a.s.l.; A. Staniczek, M. Pallmann, R. J. Godunko, F. Nejat leg., 8.5.2017, SMNS_EPH_007618.
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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10 larvae: IRAN, Alborz Province, 4 km NW of Shahrestanak village, Shahrestanak River;
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<geoCoordinate degrees="35" direction="north" minutes="59" orientation="latitude" precision="1" seconds="01.2" value="35.983665">35°59'01.2"N</geoCoordinate>
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,
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<geoCoordinate degrees="051" direction="east" minutes="19" orientation="longitude" precision="1" seconds="09.6" value="51.319332">051°19'09.6"E</geoCoordinate>
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(locality no. 57); 2100 m a.s.l.; A. Staniczek, M. Pallmann, F. Nejat leg., 10.5.2017, SMNS_EPH_007622.
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="71" type="etymology">
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<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="71">The species name refers to the type locality and distribution of the species in the Alborz mountain range.</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="71" type="localities and habitat preferences of larvae">
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<paragraph pageId="0" pageNumber="71">Localities and habitat preferences of larvae.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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Larvae inhabit small streams (2-8 m width, 20-50 cm depth) at high altitudes (six of eleven localities at approx. 2000 m a.s.l.) in the central Alborz (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Epeorus (Caucasiron) species in northern Iran. Colour of symbols shows the occurrence of species: red - E. (C.) alborzicus sp. nov., light blue - E. (C.) shargi sp. nov., violet - E. (C.) cf. znojkoi, black - E. (C.) iranicus, yellow - E. (C.) nigripilosus. The letter H shows the localities of holotypes. Black symbols with white cross show unrevised records of E. (C.) iranicus." figureDoi="10.3897/zookeys.947.51259.figure9" httpUri="https://binary.pensoft.net/fig/430257" pageId="0" pageNumber="71">9</figureCitation>
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). One larva was found in the eastern Alborz (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Epeorus (Caucasiron) species in northern Iran. Colour of symbols shows the occurrence of species: red - E. (C.) alborzicus sp. nov., light blue - E. (C.) shargi sp. nov., violet - E. (C.) cf. znojkoi, black - E. (C.) iranicus, yellow - E. (C.) nigripilosus. The letter H shows the localities of holotypes. Black symbols with white cross show unrevised records of E. (C.) iranicus." figureDoi="10.3897/zookeys.947.51259.figure9" httpUri="https://binary.pensoft.net/fig/430257" pageId="0" pageNumber="71">9</figureCitation>
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). Larvae were found only in cold and clear streams where they dwelled on large stones in riffles with very fast flow. All localities were situated in deep valleys with rivers draining high mountains. They were mostly treeless, only sometimes with sparse solitary shrubs and trees at the banks (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photos of the localities of the new Epeorus (Caucasiron) species described herein: A unnamed brook near Panjab village - type locality of E. (C.) alborzicus sp. nov. B unnamed brook near Kahrud-e Bala village - locality of E. (C.) alborzicus sp. nov. C unnamed river near Shirinabad village - type locality of E. (C.) shargi sp. nov. D unnamed brook near Chah-e Ja village - locality of E. (C.) shargi sp. nov. E Gamasiab River near Varayeneh village - type locality of E. (C.) zagrosicus sp. nov. F Yasuj fall near Yasuj village - locality of E. (C.) zagrosicus sp. nov." figureDoi="10.3897/zookeys.947.51259.figure10" httpUri="https://binary.pensoft.net/fig/430258" pageId="0" pageNumber="71">10A, B</figureCitation>
|
||
). Streams had a very coarse bed substrate with prevailing boulders and stones and a low share of fine sediments, and turbulent to strongly turbulent flow. They were characteristic of high fluctuation of discharge, with sudden peaks of discharge after spates on the mountains (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photos of the localities of the new Epeorus (Caucasiron) species described herein: A unnamed brook near Panjab village - type locality of E. (C.) alborzicus sp. nov. B unnamed brook near Kahrud-e Bala village - locality of E. (C.) alborzicus sp. nov. C unnamed river near Shirinabad village - type locality of E. (C.) shargi sp. nov. D unnamed brook near Chah-e Ja village - locality of E. (C.) shargi sp. nov. E Gamasiab River near Varayeneh village - type locality of E. (C.) zagrosicus sp. nov. F Yasuj fall near Yasuj village - locality of E. (C.) zagrosicus sp. nov." figureDoi="10.3897/zookeys.947.51259.figure10" httpUri="https://binary.pensoft.net/fig/430258" pageId="0" pageNumber="71">10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="description">
|
||
<paragraph pageId="0" pageNumber="71">Description of larva.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
General colouration of larvae yellowish brown with dark brown maculation. Body length of mature larvae: 13.3-15.8 mm (female), 10.3-11.3 mm (male). Length of cerci approximately 1.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
body length.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Head.</emphasis>
|
||
Shape trapezoidal; anterior and lateral margin rounded, posterior margin rounded in female, slightly rounded or nearly straight in male (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1D, E</figureCitation>
|
||
). Anterior margin with shallow concavity medially. Head dimensions of mature larvae: length 2.8-3.1 mm, width 4.0-4.6 mm (female); length 2.2-2.7 mm, width 3.2-3.7 mm (male). Head width/length ratio: 1.4-1.5 (both male and female). Dorso-medial part with pair of stripes. Pair of maculae located between ocelli (sometimes fused into single macula). Rounded maculae ventrolateral of lateral ocelli and blurred maculae near inner edges of compound eyes. Pale stripes extending horizontally from lateral ocelli to lateral edges of head. Pair of elongated, curved maculae located along coronal suture. Compound eyes grey to black in female, brownish or greyish and basally black in male mature larva. Ocelli blackish, basally paler. Antennae yellowish brown, scapus and pedicellus darkened. Anterior margin of head densely covered with hair-like setae extending to lateral margins and directed medio-dorsally. Dorsal surface of head covered with fine hair-like setae and sparsely distributed stick-like setae. Sparse longer and fine hair-like setae located posteriorly to eyes.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">Figure 1.</emphasis>
|
||
<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) alborzicus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alborzicus" subGenus="Caucasiron">Epeorus (Caucasiron) alborzicus</taxonomicName>
|
||
sp. nov., larva:
|
||
<emphasis bold="true" pageId="0" pageNumber="71">A</emphasis>
|
||
habitus in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">B</emphasis>
|
||
habitus in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">C</emphasis>
|
||
habitus in lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">D</emphasis>
|
||
head of male in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">E</emphasis>
|
||
head of female in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">F</emphasis>
|
||
middle leg in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">G</emphasis>
|
||
distal part of abdomen in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">H-J</emphasis>
|
||
colouration of abdominal terga
|
||
<emphasis bold="true" pageId="0" pageNumber="71">K-M</emphasis>
|
||
colouration of abdominal sterna.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Mouthparts</emphasis>
|
||
.
|
||
</emphasis>
|
||
Labrum (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2A</figureCitation>
|
||
) widened anteriorly, with anterior margin slightly rounded or nearly straight (in dorsal view). Lateral angles rounded (shape of labrum may vary among individual specimens). Dorsal surface (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2A</figureCitation>
|
||
, right half) sparsely covered with setae of different size; 4-6 longer bristle-like setae located antero-medially and two bristles antero-laterally. Epipharynx with longer, slightly plumose bristles situated along lateral to anterior margin (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2A</figureCitation>
|
||
, left half, range of setation figured as large black dots), and cluster of fine, hair-like setae medially (not figured). Posterior margin of labrum irregularly concave; group of 6-17 setae of various size located on ventral surface close to posterior margin. Outer incisors of both mandibles (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2B, C</figureCitation>
|
||
) with three apical teeth; outer tooth blunt in both mandibles. Inner incisor of left mandible with three apical teeth, right inner incisor bifurcated.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) alborzicus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alborzicus" subGenus="Caucasiron">Epeorus (Caucasiron) alborzicus</taxonomicName>
|
||
sp. nov., larva:
|
||
<emphasis bold="true" pageId="0" pageNumber="71">A</emphasis>
|
||
labrum (right half in dorsal view, left half in ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="71">B</emphasis>
|
||
incisors of right mandible in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">C</emphasis>
|
||
incisors of left mandible in ventral view (both flattened on slide)
|
||
<emphasis bold="true" pageId="0" pageNumber="71">D</emphasis>
|
||
setae on dorsal surface of femora
|
||
<emphasis bold="true" pageId="0" pageNumber="71">E</emphasis>
|
||
surface and posterior margin of abdominal tergum VII
|
||
<emphasis bold="true" pageId="0" pageNumber="71">F</emphasis>
|
||
gill I
|
||
<emphasis bold="true" pageId="0" pageNumber="71">G</emphasis>
|
||
gill III
|
||
<emphasis bold="true" pageId="0" pageNumber="71">H</emphasis>
|
||
abdominal segments VIII-X
|
||
<emphasis bold="true" pageId="0" pageNumber="71">I</emphasis>
|
||
sternum IX, female
|
||
<emphasis bold="true" pageId="0" pageNumber="71">J</emphasis>
|
||
sternum IX, male
|
||
<emphasis bold="true" pageId="0" pageNumber="71">K</emphasis>
|
||
gill VII (flattened on slide)
|
||
<emphasis bold="true" pageId="0" pageNumber="71">L-M</emphasis>
|
||
gill VII (in natural position from ventral view), variability in shape.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pronotum anteriorly narrowed, lateral edges nearly straight. Metanotum with slight postero-medial projection. Dorsal surface covered with fine, hair-like setae (as on abdominal terga and head); sparse longer, hair-like setae along pro-, meso- and metanotal suture.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Colour pattern of femora as in Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1F</figureCitation>
|
||
. Femora without medial hypodermal spot. Patella-tibial suture darkened; tarsi proximally and distally darkened. Coxal projections of fore- and hind legs pointed or bluntly pointed; in middle legs blunt. Trochanteres with spatulate setae as on dorsal surface of femora (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2D</figureCitation>
|
||
). Tibiae of forelegs 1.20-1.37
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
femur length, tibiae of middle legs 1.0-1.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
femur length, and tibiae of hind legs 0.92-1.08
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
femur length. Tarsi of all legs 0.26-0.34
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
tibia length. Dorsal surface of femora covered by short and sporadically elongated spatulate setae (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2D</figureCitation>
|
||
), hair-like setae, and sparsely distributed stick-like setae. Anterior margin of femora with short, pointed or bluntly pointed spine-like setae; posterior margin with row of long blade-like setae and sparse row of bluntly pointed, spine-like setae. Dorsal margin of tibiae and tarsi with row of long setae; ventral margin of both with irregular row of spine-like setae accumulated distally. Tarsal claws with 2-3 denticles.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Abdominal terga</emphasis>
|
||
.
|
||
</emphasis>
|
||
Colour pattern of abdominal terga (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1A, H-J</figureCitation>
|
||
) consists of transversal stripe along anterior margin of terga I-IX (X), medially extending to single blurred macula or pair of rounded maculae on terga II-IV and short triangular or nearly rectangular macula on terga V-IX. Terga VIII and IX (X) medially darkened. Pattern of abdominal terga sometimes poorly expressed, only with medially thickened transversal stripe along anterior margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
Lateral margins with oblique maculae on terga I-IX, sometimes dorso-posteriorly extended. Pair of sigilla sometimes coloured, in form of short stripes or spots located antero-laterally to medial macula. Denticles on posterior margin on terga of various size, irregular and pointed (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2E</figureCitation>
|
||
). Surface of terga covered with hair-like setae and sparsely with stick-like setae. Tergum X with distinct postero-lateral projections (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2H</figureCitation>
|
||
, arrow). Supra-tergalial projection (sensu
|
||
<bibRefCitation DOI="https://doi.org/10.1007/978-94-007-0872-3" author="Kluge, NJ" journalOrPublisher="Springer, Dordrecht" pageId="0" pageNumber="71" refId="B20" refString="Kluge, NJ, 2004. The phylogenetic system of Ephemeroptera. Springer, Dordrecht, DOI: https://doi.org/10.1007/978-94-007-0872-3" title="The phylogenetic system of Ephemeroptera." url="https://doi.org/10.1007/978-94-007-0872-3" year="2004">Kluge 2004</bibRefCitation>
|
||
) short and blunt. Longitudinal row of hair-like setae along abdominal terga present medially.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Abdominal sterna</emphasis>
|
||
.
|
||
</emphasis>
|
||
Yellowish, with distinct colour pattern in form of medial circular macula (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1B, G, K-M</figureCitation>
|
||
, best expressed on sterna II-VI). Medio-anterior sigilla partly pigmented, lateral sigilla not pigmented; medio-posterior sigilla in form of pale spots in intensively pigmented specimens. Nerve ganglia occasionally darkened. Intensity of colouration varies among individuals (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1K-M</figureCitation>
|
||
). Sternum IX with V-shaped medial emargination; surface covered by irregularly distributed short hair-like setae, and medially accumulated longer hair-like setae (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2I, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Gills</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal surface of gill plate I yellowish; of gill plates II-VII greyish on anterior half, brownish (sometimes reddish) on posterior half. Ventral margin of all gill plates yellowish. Projection of gill plate III well developed (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2G</figureCitation>
|
||
). Gill plate VII relatively wide (in natural position of ventral view, Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2L, M</figureCitation>
|
||
). Filaments of gills II-VI reaching 0.40-0.58
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of respective plate, filaments of gill VII reaching 0.18-0.24
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
(in late-instar larvae).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<taxonomicName genus="Cerci" lsidName="Cerci" pageId="0" pageNumber="71" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Cerci</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Yellowish brown, basally darkened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="subimago, imago and eggs">
|
||
<paragraph pageId="0" pageNumber="71">Subimago, imago and eggs.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="morphological diagnostics of larvae">
|
||
<paragraph pageId="0" pageNumber="71">Morphological diagnostics of larvae.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
The main larval diagnostic characters of
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. are as follows: (i) colour pattern of abdominal terga (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1A, H-J</figureCitation>
|
||
) and sterna (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1B, K-M</figureCitation>
|
||
), (ii) presence of distinct postero-lateral projections on tergum X (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2H</figureCitation>
|
||
), (iii) absence of medial hypodermal femur spot (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1F</figureCitation>
|
||
), (iv) gill plate VII relatively wide (in natural position from ventral view; Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Epeorus (Caucasiron) alborzicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K-M colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure1" httpUri="https://binary.pensoft.net/fig/430249" pageId="0" pageNumber="71">1G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2L, M</figureCitation>
|
||
), and (v) fine hair-like setae on surface of abdominal terga (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Epeorus (Caucasiron) alborzicus sp. nov., larva: A labrum (right half in dorsal view, left half in ventral view) B incisors of right mandible in ventral view C incisors of left mandible in ventral view (both flattened on slide) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H abdominal segments VIII-X I sternum IX, female J sternum IX, male K gill VII (flattened on slide) L-M gill VII (in natural position from ventral view), variability in shape." figureDoi="10.3897/zookeys.947.51259.figure2" httpUri="https://binary.pensoft.net/fig/430250" pageId="0" pageNumber="71">2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="affinities">
|
||
<paragraph pageId="0" pageNumber="71">Affinities.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
The combination of diagnostic characters mentioned above clearly distinguish larvae of
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. from all other
|
||
<taxonomicName authorityName="Kluge" authorityYear="1997" class="Insecta" family="Heptageniidae" genus="Caucasiron" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Caucasiron" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Caucasiron</emphasis>
|
||
</taxonomicName>
|
||
species known so far. However, some of the diagnostic characters occur also in other
|
||
<taxonomicName authorityName="Kluge" authorityYear="1997" class="Insecta" family="Heptageniidae" genus="Caucasiron" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Caucasiron" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Caucasiron</emphasis>
|
||
</taxonomicName>
|
||
species distributed in the Caucasus. The colour pattern of abdominal sterna in
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. is similar in
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="bicolliculatus" pageId="0" pageNumber="71" rank="species" species="bicolliculatus">bicolliculatus</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.3.2" author="Hrivniak, Ľ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="353 - 374" refId="B16" refString="Hrivniak, Ľ, Sroka, P, Godunko, RJ, Zurovcova, M, 2017. Mayflies of the genus Epeorus Eaton, 1881 s.l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey. Zootaxa 4341: 353 - 374, DOI: https://doi.org/10.11646/zootaxa.4341.3.2" title="Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey." url="https://doi.org/10.11646/zootaxa.4341.3.2" volume="4341" year="2017">Hrivniak et al. 2017</bibRefCitation>
|
||
: 356, fig. 8) and
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="alpestris" pageId="0" pageNumber="71" rank="species" species="alpestris">alpestris</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Braasch, D" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="71" pagination="283 - 294" refId="B6" refString="Braasch, D, 1979. Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae). Reichenbachia 17: 283 - 294" title="Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae)." volume="17" year="1979">Braasch 1979</bibRefCitation>
|
||
: 284, fig. 1d). Both species also lack a medial hypodermal femur spot.
|
||
<taxonomicName authorityName="Hrivniak, Sroka, Godunko & Zurovcova" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (C.) bicolliculatus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="bicolliculatus" subGenus="C.">Epeorus (C.) bicolliculatus</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. by (i) the presence of flattened setae on the surface of abdominal terga (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.3.2" author="Hrivniak, Ľ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="353 - 374" refId="B16" refString="Hrivniak, Ľ, Sroka, P, Godunko, RJ, Zurovcova, M, 2017. Mayflies of the genus Epeorus Eaton, 1881 s.l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey. Zootaxa 4341: 353 - 374, DOI: https://doi.org/10.11646/zootaxa.4341.3.2" title="Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey." url="https://doi.org/10.11646/zootaxa.4341.3.2" volume="4341" year="2017">Hrivniak et al. 2017</bibRefCitation>
|
||
: 359, fig. 23), (ii) the presence of paired postero-medial protuberances on terga II-IX (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.3.2" author="Hrivniak, Ľ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="353 - 374" refId="B16" refString="Hrivniak, Ľ, Sroka, P, Godunko, RJ, Zurovcova, M, 2017. Mayflies of the genus Epeorus Eaton, 1881 s.l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey. Zootaxa 4341: 353 - 374, DOI: https://doi.org/10.11646/zootaxa.4341.3.2" title="Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey." url="https://doi.org/10.11646/zootaxa.4341.3.2" volume="4341" year="2017">Hrivniak et al. 2017</bibRefCitation>
|
||
: 356, figs 10, 11; 360, figs 31, 32), and (iii) the absence of a postero-lateral projection on the tergum X.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<taxonomicName baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (C.) alpestris" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alpestris" subGenus="C.">Epeorus (C.) alpestris</taxonomicName>
|
||
differs by the characteristic colour pattern of abdominal terga (
|
||
<bibRefCitation author="Braasch, D" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="71" pagination="283 - 294" refId="B6" refString="Braasch, D, 1979. Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae). Reichenbachia 17: 283 - 294" title="Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae)." volume="17" year="1979">Braasch 1979</bibRefCitation>
|
||
: 294, fig. 1c) and the absence of postero-lateral projections on the tergum X.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
The presence of postero-lateral projections on the abdominal tergum X is characteristic for two species distributed in the Caucasus,
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="magnus" pageId="0" pageNumber="71" rank="species" species="magnus">magnus</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. (C.) nigripilosus" pageId="0" pageNumber="71" rank="species" species="nigripilosus" subGenus="C.">E. (C.) nigripilosus</taxonomicName>
|
||
, and sporadically also in
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="znojkoi" pageId="0" pageNumber="71" rank="species" species="znojkoi">znojkoi</taxonomicName>
|
||
.
|
||
<taxonomicName baseAuthorityName="Braasch" baseAuthorityYear="1978" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (C.) magnus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="magnus" subGenus="C.">Epeorus (C.) magnus</taxonomicName>
|
||
differs from
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. in the absence of colouration of abdominal sterna and the characteristic setation on the dorsal margin of labrum (numerous thickened bristle-like setae, Hrivniak et al. in prep.).
|
||
<taxonomicName baseAuthorityName="Sinitshenkova" baseAuthorityYear="1976" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (C.) nigripilosus" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="nigripilosus" subGenus="C.">Epeorus (C.) nigripilosus</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. by the presence of the distinct medial hypodermal femur spot and unique colour pattern of abdominal sterna (
|
||
<bibRefCitation author="Sinitshenkova, ND" journalOrPublisher="Entomologicheskoye obozreniye" pageId="0" pageNumber="71" pagination="853 - 862" refId="B32" refString="Sinitshenkova, ND, 1976. Mayflies of the Genus Iron Eaton (Ephemeroptera, Heptageniidae) in the Fauna of the Caucasus. Entomologicheskoye obozreniye 55: 853 - 862" title="Mayflies of the Genus Iron Eaton (Ephemeroptera, Heptageniidae) in the Fauna of the Caucasus." volume="55" year="1976">Sinitshenkova 1976</bibRefCitation>
|
||
: 89, fig. 28).
|
||
<taxonomicName baseAuthorityName="Tshernova" baseAuthorityYear="1938" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (C.) znojkoi" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="znojkoi" subGenus="C.">Epeorus (C.) znojkoi</taxonomicName>
|
||
can be clearly distinguished from
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. by the colour pattern of abdominal terga and conspicuous reddish colouration of abdominal sterna (
|
||
<bibRefCitation author="Braasch, D" journalOrPublisher="Entomologische Nachrichten" pageId="0" pageNumber="71" pagination="166 - 173" refId="B7" refString="Braasch, D, 1980. Beitrag zur Kenntnis der Gattung Iron Eaton (Heptageniidae, Ephemeroptera) im Kaukasus (UdSSR), 2. Entomologische Nachrichten 24: 166 - 173" title="Beitrag zur Kenntnis der Gattung Iron Eaton (Heptageniidae, Ephemeroptera) im Kaukasus (UdSSR), 2." volume="24" year="1980">Braasch 1980</bibRefCitation>
|
||
: 172, fig. 4b-c).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
Two species,
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="soldani" pageId="0" pageNumber="71" rank="species" species="soldani">soldani</taxonomicName>
|
||
and
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="sinitshenkovae" pageId="0" pageNumber="71" rank="species" species="sinitshenkovae">sinitshenkovae</taxonomicName>
|
||
, are lacking a medial hypodermal femur spot just like
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. Both can be separated from the latter by the absence of postero-lateral projections on tergum X, narrower gill plates VII (in natural position from ventral view), and the absence of a distinct colour pattern of abdominal sterna. Additionally,
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="soldani" pageId="0" pageNumber="71" rank="species" species="soldani">soldani</taxonomicName>
|
||
differs from
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. by the presence of flattened setae on the surface of abdominal terga (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.3.2" author="Hrivniak, Ľ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="353 - 374" refId="B16" refString="Hrivniak, Ľ, Sroka, P, Godunko, RJ, Zurovcova, M, 2017. Mayflies of the genus Epeorus Eaton, 1881 s.l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey. Zootaxa 4341: 353 - 374, DOI: https://doi.org/10.11646/zootaxa.4341.3.2" title="Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkey." url="https://doi.org/10.11646/zootaxa.4341.3.2" volume="4341" year="2017">Hrivniak et al. 2017</bibRefCitation>
|
||
: 359, fig. 25).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
Other
|
||
<taxonomicName authorityName="Kluge" authorityYear="1997" class="Insecta" family="Heptageniidae" genus="Caucasiron" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Caucasiron" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Caucasiron</emphasis>
|
||
</taxonomicName>
|
||
species distributed in the Caucasus and adjacent areas do not share important diagnostic characters with
|
||
<taxonomicName lsidName="E. (C.) alborzicus" pageId="0" pageNumber="71" rank="species" species="alborzicus" subGenus="C.">E. (C.) alborzicus</taxonomicName>
|
||
sp. nov. All of these species can be easily distinguished by the following combination of characters: (i) absence of the colour pattern of abdominal sterna and presence of the medial hypodermal femur spot in
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="turcicus" pageId="0" pageNumber="71" rank="species" species="turcicus">turcicus</taxonomicName>
|
||
,
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="longimaculatus" pageId="0" pageNumber="71" rank="species" species="longimaculatus">longimaculatus</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. (C.) shargi" pageId="0" pageNumber="71" rank="species" species="shargi" subGenus="C.">E. (C.) shargi</taxonomicName>
|
||
sp. nov. and (ii) colour pattern of abdominal terga and sterna in
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="caucasicus" pageId="0" pageNumber="71" rank="species" species="caucasicus">caucasicus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Braasch, D" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="71" pagination="283 - 294" refId="B6" refString="Braasch, D, 1979. Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae). Reichenbachia 17: 283 - 294" title="Beitrag zur Kenntnis der Gattung Iron Eaton im Kaukasus (UdSSR) (III) (Ephemeroptera, Heptageniidae)." volume="17" year="1979">Braasch 1979</bibRefCitation>
|
||
: fig. 3a),
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="iranicus" pageId="0" pageNumber="71" rank="species" species="iranicus">iranicus</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Braasch, D" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="71" pagination="261 - 272" refId="B9" refString="Braasch, D, Soldan, T, 1979. Neue Heptageniidae aus Asien (Ephemeroptera). Reichenbachia 17: 261 - 272" title="Neue Heptageniidae aus Asien (Ephemeroptera)." volume="17" year="1979">
|
||
Braasch and
|
||
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
|
||
1979
|
||
</bibRefCitation>
|
||
: fig. 12), and
|
||
<taxonomicName lsidName="E. (C.) zagrosicus" pageId="0" pageNumber="71" rank="species" species="zagrosicus" subGenus="C.">E. (C.) zagrosicus</taxonomicName>
|
||
sp. nov. (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) zagrosicus sp. nov., larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G distal part of abdomen in ventral view H-J colouration of abdominal terga K colouration of abdominal sterna." figureDoi="10.3897/zookeys.947.51259.figure5" httpUri="https://binary.pensoft.net/fig/430253" pageId="0" pageNumber="71">5A-C, G, H-K</figureCitation>
|
||
). The larva of
|
||
<taxonomicName genus="E." lsidName="E." pageId="0" pageNumber="71" rank="genus">E.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="0" pageNumber="71" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="insularis" pageId="0" pageNumber="71" rank="species" species="insularis">insularis</taxonomicName>
|
||
is currently not described.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.947.51259.figure3" httpUri="https://binary.pensoft.net/fig/430251" pageId="0" pageNumber="71" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) shargi" order="Ephemeroptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">Epeorus (Caucasiron) shargi</taxonomicName>
|
||
sp. nov., larva:
|
||
<emphasis bold="true" pageId="0" pageNumber="71">A</emphasis>
|
||
habitus in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">B</emphasis>
|
||
habitus in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">C</emphasis>
|
||
habitus in lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">D</emphasis>
|
||
head of male in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">E</emphasis>
|
||
head of female in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">F</emphasis>
|
||
middle leg in dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">G</emphasis>
|
||
distal part of abdomen in ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="71">H-K</emphasis>
|
||
colouration of abdominal terga
|
||
<emphasis bold="true" pageId="0" pageNumber="71">L</emphasis>
|
||
colouration of abdominal sterna.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |