285 lines
32 KiB
XML
285 lines
32 KiB
XML
<document id="DAC6C7048A1B7004093CC65EBBBCE5EB" ID-DOI="10.11646/zootaxa.3700.2.1" ID-GBIF-Dataset="3d28eeb6-88a2-4fb2-aebd-796c8b2c2287" ID-ISSN="1175-5326" ID-Zenodo-Dep="218268" ID-ZooBank="8119DAED-D7AF-4858-BC63-F082151A4DAE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460662093739" checkinUser="plazi" docAuthor="Zaragoza, Juan A. & Pérez, Toni" docDate="2013" docId="620687CFFFE9FF9CEED61AF7FDAE17CF" docLanguage="en" docName="zt03700p225.pdf" docOrigin="Zootaxa 3700 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chthonius Zaragoza & Pérez, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="213" masterDocId="9E3FFFB7FFE0FF90EE411F6BFF80144F" masterDocTitle="Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy" masterLastPageNumber="225" masterPageNumber="201" pageNumber="210" updateTime="1698347634734" updateUser="plazi">
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<mods:mods id="18E01D284426F6BF5EF462E5E8B9D047" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1D6D79F4FA903E9C3C723B7B14846148">
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<mods:title id="423739D9E06EFDAD16871807F7842877">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<mods:name id="B16225F5F38FDFE2ED6C386B51659645" type="personal">
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<mods:namePart id="80A6CDFC248C16F9A2D378618A15E031">Zaragoza, Juan A.</mods:namePart>
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<mods:name id="CCBECBBF2D8941B49434BE0E78C79866" type="personal">
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<mods:roleTerm id="36C1015D943945B5BDB17C0DCD1E5D78">Author</mods:roleTerm>
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<mods:namePart id="0F0370773A6D12796B89E798C4FDE6D7">Pérez, Toni</mods:namePart>
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<mods:typeOfResource id="3F5846995DF4BD4D6E771105CDB61CC8">text</mods:typeOfResource>
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<mods:title id="00700A7B244DBBE768080C246D8252B2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A880D9DA48115FB14B0770FF3ECF2040">
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<mods:date id="49F55FF015ABEDB67DE7FFC0B86CA4E7">2013</mods:date>
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<mods:detail id="B09BA7AB6E2A26386BC439D7C0EBC49A" type="volume">
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<mods:number id="1F7279947CC2303C4386D623CDD177A7">3700</mods:number>
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<mods:detail id="C690A93B36F7C07299F6F984DE17CCF6" type="issue">
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<mods:number id="E461361364C2CFB947EFE4343F404D1F">2</mods:number>
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<mods:classification id="A03AEB9F747C2916DF3DB193E7D6A004">journal article</mods:classification>
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<mods:identifier id="0F5C4B9873AA63C81C2F81D8F4FF51D8" type="DOI">10.11646/zootaxa.3700.2.1</mods:identifier>
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<mods:identifier id="6FE2E641B6347F5510D9D62ECC6209DE" type="GBIF-Dataset">3d28eeb6-88a2-4fb2-aebd-796c8b2c2287</mods:identifier>
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<mods:identifier id="AA216C0BD4C1233800616C0AD32A9631" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0410F5022D0D89A3B095F014BBA0159E" type="Zenodo-Dep">218268</mods:identifier>
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<mods:identifier id="2A645D5D62AA06F70CCBE59CD1D67519" type="ZooBank">8119DAED-D7AF-4858-BC63-F082151A4DAE</mods:identifier>
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<treatment id="620687CFFFE9FF9CEED61AF7FDAE17CF" ID-DOI="http://doi.org/10.5281/zenodo.6149888" ID-GBIF-Taxon="119574438" ID-Zenodo-Dep="6149888" LSID="urn:lsid:plazi:treatment:620687CFFFE9FF9CEED61AF7FDAE17CF" httpUri="http://treatment.plazi.org/id/620687CFFFE9FF9CEED61AF7FDAE17CF" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210">
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<subSubSection id="A2B56552FFE9FF99EED61AF7FEA71197" pageId="9" pageNumber="210" type="nomenclature">
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<paragraph id="EA1036D9FFE9FF99EED61AF7FDB811F8" blockId="9.[151,568,1436,1496]" box="[151,568,1436,1463]" pageId="9" pageNumber="210">
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<heading id="B15881B5FFE9FF99EED61AF7FDB811F8" bold="true" box="[151,568,1436,1463]" fontSize="11" level="1" pageId="9" pageNumber="210" reason="1">
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<emphasis id="D8DBEACBFFE9FF99EED61AF7FDB811F8" bold="true" box="[151,568,1436,1463]" pageId="9" pageNumber="210">
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<emphasis id="D8DBEACBFFE9FF99EED61AF7FE5411F9" bold="true" box="[151,468,1436,1462]" italics="true" pageId="9" pageNumber="210">
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<taxonomicName id="2DAF4D5AFFE9FF99EED61AF7FE9611F9" ID-CoL="62LNY" box="[151,278,1436,1462]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
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(E.)
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<taxonomicName id="2DAF4D5AFFE9FF99EF121AF7FE5411F9" box="[339,468,1436,1462]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov.">giennensis</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="C3E857B0FFE9FF99EF9A1AF6FDB811F8" box="[475,568,1437,1463]" pageId="9" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EA1036D9FFE9FF99EED61AD4FEA71197" blockId="9.[151,568,1436,1496]" box="[151,295,1471,1496]" pageId="9" pageNumber="210">
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(
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<figureCitation id="72942A5CFFE9FF99EEDE1AD4FE9F1197" box="[159,287,1471,1496]" captionStart-0="FIGURES 13 – 16" captionStart-1="FIGURES 17 – 20" captionStartId-0="10.[151,264,799,822]" captionStartId-1="11.[151,264,1952,1975]" captionTargetBox-0="[214,1364,209,778]" captionTargetBox-1="[192,1430,228,1917]" captionTargetId-0="figure@10.[207,1380,193,779]" captionTargetId-1="figure@11.[151,1436,193,1932]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." captionText-1="FIGURES 17 – 20. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 17, right pedipalp femur, dorsal view; 18, right patella, dorsal view; 19, right chela, lateral view; 20, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17 – 19)." httpUri-0="https://zenodo.org/record/218273/files/figure.png" httpUri-1="https://zenodo.org/record/218274/files/figure.png" pageId="9" pageNumber="210">Figs 13–20</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A2B56552FFE9FF99EED6196CFD271227" pageId="9" pageNumber="210" type="materials_examined">
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<paragraph id="EA1036D9FFE9FF99EED6196CFD271227" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
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<emphasis id="D8DBEACBFFE9FF99EED6196CFECF126F" bold="true" box="[151,335,1543,1568]" pageId="9" pageNumber="210">
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<typeStatus id="3514887BFFE9FF99EED6196CFF52126F" box="[151,210,1543,1568]" pageId="9" pageNumber="210">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="3514887BFFE9FF99EF1E196CFE47126F" box="[351,455,1543,1568]" pageId="9" pageNumber="210" type="holotype">Holotype</typeStatus>
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♂,
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<collectingCountry id="92B87649FFE9FF99EC44196CFDC8126F" box="[517,584,1543,1568]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
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, Andalusia, Jaén province, Peal de Becerro, Cueva de los Ladrones (37°51′2.39″N, 2°57′48.69″W;
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<quantity id="2D579B3CFFE9FF99EFB91947FDD4120A" box="[504,596,1580,1605]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="9" pageNumber="210" unit="m" value="1575.0">1575 m</quantity>
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.a.s.l.),
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<date id="9E111019FFE9FF99ECE11947FCA3120B" box="[672,803,1580,1604]" pageId="9" pageNumber="210" value="2012-06-03">03.VI.2012</date>
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, lgt. G.E.V. (DEUA: 20011).
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<typeStatus id="3514887BFFE9FF99EA2F1946FB50120A" box="[1134,1232,1581,1605]" pageId="9" pageNumber="210" type="paratype">Paratype</typeStatus>
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tritonymph, same details as
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<typeStatus id="3514887BFFE9FF99EF441924FEE71227" box="[261,359,1615,1640]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
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(MCNB:
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<date id="9E111019FFE9FF99EF981924FDD21227" box="[473,594,1615,1640]" pageId="9" pageNumber="210" value="2013" valueMax="2543">2013-2543</date>
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MZB).
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</paragraph>
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</subSubSection>
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<subSubSection id="A2B56552FFE9FF99EE86191FFDB912FF" pageId="9" pageNumber="210" type="etymology">
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<paragraph id="EA1036D9FFE9FF99EE86191FFDB912FF" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
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<emphasis id="D8DBEACBFFE9FF99EE86191FFECE12C2" bold="true" box="[199,334,1652,1677]" pageId="9" pageNumber="210">Etymology.</emphasis>
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Latin adjective derived from
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<emphasis id="D8DBEACBFFE9FF99ECDD191EFD7312C3" box="[668,755,1653,1676]" italics="true" pageId="9" pageNumber="210">Giennis</emphasis>
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, the Latin name for the Roman city of Jaén, which currently gives its name to the whole province.
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</paragraph>
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</subSubSection>
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<subSubSection id="A2B56552FFE9FF99EE8619D7FC98132B" pageId="9" pageNumber="210" type="diagnosis">
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<paragraph id="EA1036D9FFE9FF99EE8619D7FC98132B" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
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<emphasis id="D8DBEACBFFE9FF99EE8619D7FEC1129A" bold="true" box="[199,321,1724,1749]" pageId="9" pageNumber="210">Diagnosis.</emphasis>
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Facies weakly troglomorphic. No eyes or eye-spots. Chelicera with 6 setae on palm and 4–5 microsetae, movable cheliceral finger without isolated subapical tooth, male spinneret strongly reduced. Carapace with two macrosetae on posterior margin, without preocular microsetae. Pedipalp femur length
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<quantity id="2D579B3CFFE9FF99EAA1186FFAC61353" box="[1248,1350,1796,1821]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="210" unit="mm" value="0.61">0.61 mm</quantity>
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(5.8×); chela lateral length 0.87 (5.4×); fixed finger with 20 teeth, movable finger with 12 teeth and raised lamina with 10 rounded vestigial teeth; trichobothrium
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<emphasis id="D8DBEACBFFE9FF99EC151826FDEF132B" box="[596,623,1869,1892]" italics="true" pageId="9" pageNumber="210">ist</emphasis>
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level with
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<emphasis id="D8DBEACBFFE9FF99ECAA1827FC90132A" box="[747,784,1868,1893]" italics="true" pageId="9" pageNumber="210">esb</emphasis>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2B56552FFE9FF9CEE861804FBB61683" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210" type="description">
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||
<paragraph id="EA1036D9FFE9FF99EE861804FF5513BA" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
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||
<emphasis id="D8DBEACBFFE9FF99EE861804FED913C7" bold="true" box="[199,345,1903,1928]" pageId="9" pageNumber="210">Description.</emphasis>
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<emphasis id="D8DBEACBFFE9FF99EF231804FD8513C7" box="[354,517,1903,1928]" italics="true" pageId="9" pageNumber="210">
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Male
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<typeStatus id="3514887BFFE9FF99EFE51804FD8513C7" box="[420,517,1903,1928]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
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</emphasis>
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. Small, weakly troglomorphic hypogean species; integument depigmented, pale brown; marked hispid granulation on lateral surface of ocular and posterior areas of carapace, on cheliceral palm, on base of paraxial face of pedipalpal femur, on base of movable finger and on dorsodistal surface of pedipalpal hand.
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</paragraph>
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<caption id="BED06651FFEAFF9AEED61C74FF76173C" httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211" targetBox="[214,1364,209,778]" targetPageId="10">
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<paragraph id="EA1036D9FFEAFF9AEED61C74FF76173C" blockId="10.[151,1436,799,883]" pageId="10" pageNumber="211">
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<emphasis id="D8DBEACBFFEAFF9AEED61C74FED3177A" bold="true" box="[151,339,799,822]" pageId="10" pageNumber="211">FIGURES 13–16.</emphasis>
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<taxonomicName id="2DAF4D5AFFEAFF9AEF191C4BFC871779" box="[344,775,800,822]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov." subGenus="Ephippiochthonius">
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<emphasis id="D8DBEACBFFEAFF9AEF191C4BFC871779" box="[344,775,800,822]" italics="true" pageId="10" pageNumber="211">Chthonius (Ephippiochthonius) giennensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C3E857B0FFEAFF9AED4C1C4BFCDD177A" box="[781,861,800,821]" pageId="10" pageNumber="211" rank="species">
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<emphasis id="D8DBEACBFFEAFF9AED4C1C4BFCDD177A" bold="true" box="[781,861,800,821]" pageId="10" pageNumber="211">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, male holotype.
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<emphasis id="D8DBEACBFFEAFF9AEA441C4BFB9E177A" bold="true" box="[1029,1054,800,821]" pageId="10" pageNumber="211">13</emphasis>
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, carapace;
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<emphasis id="D8DBEACBFFEAFF9AEACE1C4BFB28177A" bold="true" box="[1167,1192,800,821]" pageId="10" pageNumber="211">14</emphasis>
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, partial view of anterior margin of carapace;
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<emphasis id="D8DBEACBFFEAFF9AEF2A1C54FE04171B" bold="true" box="[363,388,831,852]" pageId="10" pageNumber="211">15</emphasis>
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, left chelicera;
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<emphasis id="D8DBEACBFFEAFF9AEC661C54FDC0171B" bold="true" box="[551,576,831,852]" pageId="10" pageNumber="211">16</emphasis>
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, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14–16), 0.2 (Fig. 13).
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</paragraph>
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</caption>
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<paragraph id="EA1036D9FFEAFF9AEE861CCBFD26106B" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
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Carapace (
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<figureCitation id="72942A5CFFEAFF9AEF031CCBFE1617F7" box="[322,406,928,953]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Fig. 13</figureCitation>
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) slightly longer than broad, slightly constricted posteriorly, subquadrate; anterior margin (
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<figureCitation id="72942A5CFFEAFF9AEEDE1CAFFF711792" box="[159,241,964,989]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Fig. 14</figureCitation>
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) prominent, medially dentate and with rudimentary epistome; without eyes or eye-spots; 18 setae, without preocular microsetae, formula: 4:6:4:2:2, anteromedial seta
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<quantity id="2D579B3CFFEAFF9AED721C83FC15104F" box="[819,917,1000,1024]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="211" unit="mm" value="0.11">0.11 mm</quantity>
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long, ocular seta
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<quantity id="2D579B3CFFEAFF9AEA1B1C83FB40104F" box="[1114,1216,1000,1024]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="211" unit="mm" value="0.09">0.09 mm</quantity>
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||
. Five lyrifissures in anterior and ocular areas, one in posterior area.
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</paragraph>
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<paragraph id="EA1036D9FFEAFF9AEE861B5BFB041022" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
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Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
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<date id="9E111019FFEAFF9AEDA91B5AFB891007" box="[1000,1033,1072,1096]" pageId="10" pageNumber="211">I 3</date>
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||
+ 3 marginal microsetae,
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<date id="9E111019FFEAFF9AEB7F1B5AFAE91007" box="[1342,1385,1072,1096]" pageId="10" pageNumber="211">II 4</date>
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||
+ 9 bipinnate coxal spines,
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<date id="9E111019FFEAFF9AEFDF1B3EFE4F1022" box="[414,463,1108,1133]" pageId="10" pageNumber="211">III 5</date>
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+ 5 bipinnate coxal spines,
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<date id="9E111019FFEAFF9AED491B3EFCBD1022" box="[776,829,1108,1133]" pageId="10" pageNumber="211">IV 6</date>
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; intercoxal tubercle bisetose.
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</paragraph>
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<paragraph id="EA1036D9FFEAFF9AEE861B13FC591097" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 9 setae on sternite II; sternite III with genital opening flanked by 7 setae on each side, with 4+4 internal glandular setae and (3)8(3) marginal setae. Chaetotaxy of sternites IV–XI: (2)7(2):8:6:6:6:6:6:0. Anal cone 0+2 setae.</paragraph>
|
||
<paragraph id="EA1036D9FFEAFF9AEE861B8FFBF111C2" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
|
||
Chelicera (
|
||
<figureCitation id="72942A5CFFEAFF9AEF071B8FFE4B10B2" box="[326,459,1252,1277]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Figs 15–16</figureCitation>
|
||
) with 6 setae on palm and 4–5 microsetae laterally, seta
|
||
<emphasis id="D8DBEACBFFEAFF9AEA3D1B8FFB1710B2" box="[1148,1175,1252,1277]" italics="true" pageId="10" pageNumber="211">vb</emphasis>
|
||
short,
|
||
<quantity id="2D579B3CFFEAFF9AEAAF1B8EFAD810B3" box="[1262,1368,1253,1277]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="211" unit="mm" value="0.04">0.04 mm</quantity>
|
||
long, microsetae
|
||
<quantity id="2D579B3CFFEAFF9AEF581A63FE3F116F" box="[281,447,1288,1312]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="211" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.02–0.03 mm</quantity>
|
||
long; seta
|
||
<emphasis id="D8DBEACBFFEAFF9AEC7E1A63FDD5116E" box="[575,597,1288,1313]" italics="true" pageId="10" pageNumber="211">gl</emphasis>
|
||
0.52 from base of movable finger. Fixed finger with 14 teeth, two distal teeth distinctly larger than others, proximally decreasing in size. Movable finger without an isolated subapical tooth (
|
||
<emphasis id="D8DBEACBFFEAFF9AEEA41A3BFF7B1126" box="[229,251,1360,1385]" italics="true" pageId="10" pageNumber="211">di</emphasis>
|
||
), a large distal tooth and 9 contiguous teeth reduced in size proximally; spinneret strongly reduced. Rallum with 11 blades, serrulae exterior and interior with 16 and 14 blades respectively.
|
||
</paragraph>
|
||
<paragraph id="EA1036D9FFEAFF9AEE861AF3FA0A12BB" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
|
||
Pedipalp (
|
||
<figureCitation id="72942A5CFFEAFF9AEF7B1AF3FE3E11FF" box="[314,446,1432,1457]" captionStart="FIGURES 17 – 20" captionStartId="11.[151,264,1952,1975]" captionTargetBox="[192,1430,228,1917]" captionTargetId="figure@11.[151,1436,193,1932]" captionTargetPageId="11" captionText="FIGURES 17 – 20. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 17, right pedipalp femur, dorsal view; 18, right patella, dorsal view; 19, right chela, lateral view; 20, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17 – 19)." httpUri="https://zenodo.org/record/218274/files/figure.png" pageId="10" pageNumber="211">Figs 17–20</figureCitation>
|
||
) with femoral chaetotaxy 3:6:3:5:1, four lyrifissures: one antiaxial and three paraxial. Chelal hand markedly depressed at level of
|
||
<emphasis id="D8DBEACBFFEAFF9AECC01AD7FD41119A" box="[641,705,1468,1493]" italics="true" pageId="10" pageNumber="211">ib-isb</emphasis>
|
||
, with a rounded short hump distad of
|
||
<emphasis id="D8DBEACBFFEAFF9AEA2A1AD7FB2B119A" box="[1131,1195,1468,1493]" italics="true" pageId="10" pageNumber="211">ib-isb</emphasis>
|
||
and a distinct ventral hollow just before base of movable chelal finger; chaetotaxy 4:5:4. Fixed finger with 20 pointed teeth and one rounded proximal tooth, all with dental canals except the two small distal ones and proximal one; third distal tooth from tip modified, a little wider than the others; four microtubercles at base; tip of finger with an accessory tooth (
|
||
<emphasis id="D8DBEACBFFEAFF9AEEE11927FF36122A" box="[160,182,1612,1637]" italics="true" pageId="10" pageNumber="211">td</emphasis>
|
||
) on antiaxial face; five teeth at level of
|
||
<emphasis id="D8DBEACBFFEAFF9AEC341926FD2C122B" box="[629,684,1613,1636]" italics="true" pageId="10" pageNumber="211">est/it</emphasis>
|
||
occupying
|
||
<quantity id="2D579B3CFFEAFF9AED6E1926FC07122B" box="[815,903,1613,1637]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="211" unit="mm" value="0.1">0.1 mm</quantity>
|
||
, distance between apices
|
||
<quantity id="2D579B3CFFEAFF9AEAE61926FA9C122B" box="[1191,1308,1613,1637]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="211" unit="mm" value="0.024">0.024 mm</quantity>
|
||
. Distal half of movable finger with 12 pointed teeth with dental canals, two distal ones tiny and the proximal one reduced in size; proximal half of finger with slightly raised lamina with 10 rounded vestigial teeth reaching slightly proximad o trichobothrium
|
||
<emphasis id="D8DBEACBFFEAFF9AEF1D19D3FEF5129E" box="[348,373,1720,1745]" italics="true" pageId="10" pageNumber="211">sb</emphasis>
|
||
, level with coupled sensilla
|
||
<emphasis id="D8DBEACBFFEAFF9AECF719D2FD51129F" box="[694,721,1721,1744]" italics="true" pageId="10" pageNumber="211">pc</emphasis>
|
||
; basal apodeme long and apically indented. Trichobothria as in figs 18–19;
|
||
<emphasis id="D8DBEACBFFEAFF9AEF5D19B6FEB712BB" box="[284,311,1757,1780]" italics="true" pageId="10" pageNumber="211">ist</emphasis>
|
||
level with
|
||
<emphasis id="D8DBEACBFFEAFF9AEFF519B7FE5912BA" box="[436,473,1756,1781]" italics="true" pageId="10" pageNumber="211">esb</emphasis>
|
||
, both forming an obtuse angle with
|
||
<emphasis id="D8DBEACBFFEAFF9AED2E19B7FC0A12BA" box="[879,906,1756,1781]" italics="true" pageId="10" pageNumber="211">eb</emphasis>
|
||
; distance
|
||
<emphasis id="D8DBEACBFFEAFF9AEDBA19B7FBB512BA" box="[1019,1077,1756,1781]" italics="true" pageId="10" pageNumber="211">st–sb</emphasis>
|
||
2.1 longer than distance
|
||
<emphasis id="D8DBEACBFFEAFF9AEB0F19B7FA0312BA" box="[1358,1411,1756,1781]" italics="true" pageId="10" pageNumber="211">sb–b</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EA1036D9FFEAFF9AEE86186AFC491357" blockId="10.[151,1437,928,2033]" box="[199,969,1792,1817]" pageId="10" pageNumber="211">Leg IV. Tactile seta ratios: tibia 0.57, basitarsus 0.36 and tarsus 0.28.</paragraph>
|
||
<paragraph id="EA1036D9FFEAFF9AEE86184EFB0913BF" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
|
||
<emphasis id="D8DBEACBFFEAFF9AEE86184EFE731373" box="[199,499,1828,1853]" italics="true" pageId="10" pageNumber="211">Measurements and ratios.</emphasis>
|
||
Body 1.48. Carapace 0.48/0.44 (1.1). Chelicera: palm 0.41/0.19 (2.1), movable finger 0.21. Pedipalp: femur 0.61/0.11 (5.8); patella 0.27/0.13 (2.1); chela 0.87/0.17 (5.3), depth 0.16 (5.4); hand 0.34 (2.1), length/depth (2.1); movable finger 0.52; ratio movable finger/hand 1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.3; ratio chela/carapace 1.8. Leg I: femur 0.35/0.06 (5.5); patella 0.17/0.06 (2.9); tibia 0.20/ 0.05 (4.3); tarsus 0.39/0.04 (10.4); ratio femur/patella 2.1. Leg IV: femur+patella 0.54/0.22 (2.5); tibia 0.38/0.08 (4.7); basitarsus 0.19/0.06 (3.1); telotarsus 0.39/0.04 (10.1); ratio telotarsus/basitarsus 2.0.
|
||
</paragraph>
|
||
<caption id="BED06651FFEBFF9BEED618CBFE5E13BB" httpUri="https://zenodo.org/record/218274/files/figure.png" pageId="11" pageNumber="212" targetBox="[192,1430,228,1917]" targetPageId="11">
|
||
<paragraph id="EA1036D9FFEBFF9BEED618CBFE5E13BB" blockId="11.[151,1436,1952,2036]" pageId="11" pageNumber="212">
|
||
<emphasis id="D8DBEACBFFEBFF9BEED618CBFED513F9" bold="true" box="[151,341,1952,1975]" pageId="11" pageNumber="212">FIGURES 17–20.</emphasis>
|
||
<taxonomicName id="2DAF4D5AFFEBFF9BEF1D18CAFC8F13F8" box="[348,783,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov." subGenus="Ephippiochthonius">
|
||
<emphasis id="D8DBEACBFFEBFF9BEF1D18CAFC8F13F8" box="[348,783,1953,1975]" italics="true" pageId="11" pageNumber="212">Chthonius (Ephippiochthonius) giennensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C3E857B0FFEBFF9BED5618CAFCE813F9" box="[791,872,1953,1974]" pageId="11" pageNumber="212" rank="species">
|
||
<emphasis id="D8DBEACBFFEBFF9BED5618CAFCE813F9" bold="true" box="[791,872,1953,1974]" pageId="11" pageNumber="212">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis id="D8DBEACBFFEBFF9BEA5618CAFBB013F9" bold="true" box="[1047,1072,1953,1974]" pageId="11" pageNumber="212">17</emphasis>
|
||
, right pedipalp femur, dorsal view;
|
||
<emphasis id="D8DBEACBFFEBFF9BEED618ABFF30139A" bold="true" box="[151,176,1984,2005]" pageId="11" pageNumber="212">18</emphasis>
|
||
, right patella, dorsal view;
|
||
<emphasis id="D8DBEACBFFEBFF9BEF8C18ABFE66139A" bold="true" box="[461,486,1984,2005]" pageId="11" pageNumber="212">19</emphasis>
|
||
, right chela, lateral view;
|
||
<emphasis id="D8DBEACBFFEBFF9BECB618ABFC90139A" bold="true" box="[759,784,1984,2005]" pageId="11" pageNumber="212">20</emphasis>
|
||
, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17–19).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EA1036D9FFECFF9CEE861FFCFBB61683" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="213">
|
||
<emphasis id="D8DBEACBFFECFF9CEE861FFCFED714FF" bold="true" box="[199,343,151,176]" pageId="12" pageNumber="213">Tritonymph</emphasis>
|
||
(data coinciding with
|
||
<typeStatus id="3514887BFFECFF9CEC121FFCFD3514FF" box="[595,693,151,176]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
omitted). Carapace quadrate. Coxa
|
||
<date id="9E111019FFECFF9CEA061FF3FBE614FF" box="[1095,1126,152,176]" pageId="12" pageNumber="213">I 3</date>
|
||
+ 2 marginal microsetae,
|
||
<date id="9E111019FFECFF9CEBC81FF3FF25149B" pageId="12" pageNumber="213">II 4</date>
|
||
+ 7–8 bipinnate coxal spines,
|
||
<date id="9E111019FFECFF9CEC101FD6FD11149A" box="[593,657,188,213]" pageId="12" pageNumber="213">III 5</date>
|
||
+ 4 bipinnate coxal spines. Chaetotaxy of sternites II–XI: 5:(2)6(2):(1)6(1):6:6:6:6:6:7:0. Cheliceral palm with 5 setae and 4 microsetae laterally, seta
|
||
<emphasis id="D8DBEACBFFECFF9CEAF31FB4FB4814B7" box="[1202,1224,223,248]" italics="true" pageId="12" pageNumber="213">gl</emphasis>
|
||
0.61 from base of movable finger; fixed finger with 10 teeth, two distal teeth distinctly larger than the others; movable finger without an isolated subapical tooth (
|
||
<emphasis id="D8DBEACBFFECFF9CEF961E4CFE6C150F" box="[471,492,295,320]" italics="true" pageId="12" pageNumber="213">di</emphasis>
|
||
), a large distal tooth and 6 teeth reduced in size proximally; spinneret blunt toothshaped, more developed than in male
|
||
<typeStatus id="3514887BFFECFF9CEC091E27FD2E152A" box="[584,686,332,357]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
. Serrulae exterior and interior with 15 and 14 blades respectively. Chelal hand chaetotaxy 4–4–4; fixed finger with 14 pointed teeth and one rounded proximal tooth, all with dental canals except the two small distal ones and the proximal one; distal half of movable finger with 10 pointed teeth, two tiny distal ones and reduced proximal one without dental canals; proximal half of finger with slightly raised lamina with 9 rounded, vestigial teeth. Leg IV: tactile seta ratios: tibia 0.55, basitarsus 0.39, tarsus 0.28.
|
||
<emphasis id="D8DBEACBFFECFF9CEED61D6AFE361657" box="[151,438,511,536]" italics="true" pageId="12" pageNumber="213">Measurements and ratios</emphasis>
|
||
: Body 0.96. Carapace 0.33/0.33 (1.0). Chelicera: palm 0.30/0.14 (2.2), movable finger 0.14. Pedipalp: femur 0.38/0.09 (4.4); patella 0.19/0.10 (2.0); chela width and depth 0.62/0.12 (5.1); hand 0.25 (2.0), length/dept (2.0); movable finger 0.36; ratio movable finger/hand 1.5; ratio femur/movable finger 1.0; ratio femur/carapace 1.2, ratio chela/carapace 1.9. Leg I: femur 0.23/0.05 (4.5); patella 0.11/0.05 (2.4); tibia 0.12/0.04 (3.0); tarsus 0.25/0.03 (7.7); ratio femur/patella 2.0. Leg IV: femur+patella 0.36/0.14 (2.5); tibia 0.24/0.06 (4.0); basitarsus 0.12/0.05 (2.5); telotarsus 0.25/0.04 (7.1); ratio telotarsus/basitarsus 2.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2B56552FFECFF9CEE861DBCFC221712" pageId="12" pageNumber="213" type="discussion">
|
||
<paragraph id="EA1036D9FFECFF9CEE861DBCFC221712" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="213">
|
||
<emphasis id="D8DBEACBFFECFF9CEE861DBCFEBB16BF" bold="true" box="[199,315,727,752]" pageId="12" pageNumber="213">Remarks.</emphasis>
|
||
<emphasis id="D8DBEACBFFECFF9CEF031DB3FD8A16BF" box="[322,522,728,752]" italics="true" pageId="12" pageNumber="213">
|
||
C. (E.)
|
||
<taxonomicName id="2DAF4D5AFFECFF9CEFD51DB2FD8A16BF" box="[404,522,729,752]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov.">giennensis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="C3E857B0FFECFF9CEC501DBCFDEB16BF" box="[529,619,727,752]" pageId="12" pageNumber="213" rank="species">
|
||
<emphasis id="D8DBEACBFFECFF9CEC501DBCFDEB16BF" bold="true" box="[529,619,727,752]" pageId="12" pageNumber="213">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shares with
|
||
<emphasis id="D8DBEACBFFECFF9CECB61DB3FC3516BF" box="[759,949,727,752]" italics="true" pageId="12" pageNumber="213">
|
||
C. (E.)
|
||
<taxonomicName id="2DAF4D5AFFECFF9CED0B1DBCFC3516BF" box="[842,949,727,752]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="espanyoli">espanyoli</taxonomicName>
|
||
</emphasis>
|
||
and the other five species mentioned in the discussion of the latter the same main characters, but differs from all of them in having 4–5 microsetae on the cheliceral palm and in the position of trichobothrium
|
||
<emphasis id="D8DBEACBFFECFF9CECB11C4AFC8B1777" box="[752,779,801,824]" italics="true" pageId="12" pageNumber="213">ist</emphasis>
|
||
level with
|
||
<emphasis id="D8DBEACBFFECFF9CEDC81C74FC2E1777" box="[905,942,799,824]" italics="true" pageId="12" pageNumber="213">esb</emphasis>
|
||
, as opposed to 1 microsetae (rarely 0–2) on the cheliceral palm and
|
||
<emphasis id="D8DBEACBFFECFF9CEFE31C2EFE3D1713" box="[418,445,837,860]" italics="true" pageId="12" pageNumber="213">ist</emphasis>
|
||
markedly distal of
|
||
<emphasis id="D8DBEACBFFECFF9CECD71C2FFD3B1712" box="[662,699,836,861]" italics="true" pageId="12" pageNumber="213">esb</emphasis>
|
||
in the other species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2B56552FFECFF9CEE861C0CFDAE17CF" box="[199,558,871,896]" pageId="12" pageNumber="213" type="distribution">
|
||
<paragraph id="EA1036D9FFECFF9CEE861C0CFDAE17CF" blockId="12.[151,1437,151,896]" box="[199,558,871,896]" pageId="12" pageNumber="213">
|
||
<emphasis id="D8DBEACBFFECFF9CEE861C0CFEE017CF" bold="true" box="[199,352,871,896]" pageId="12" pageNumber="213">Distribution.</emphasis>
|
||
Andalusia,
|
||
<collectingCountry id="92B87649FFECFF9CEFA71C0CFDAA17CF" box="[486,554,871,896]" name="Spain" pageId="12" pageNumber="213">Spain</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |