treatments-xml/data/5A/F1/6E/5AF16EEF611F5A5A9AD1421032313815.xml

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<mods:title id="575CAEC73DD0313F27C161A70882CA88">A new species of Nitokra Boeck, 1865 (Copepoda, Harpacticoida, Ameiridae) from the Caribbean coast of Colombia</mods:title>
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<mods:namePart id="2BC4F099E0E69FBDB0473E3EADD56FC9">Fuentes-Reines, Juan M.</mods:namePart>
<mods:affiliation id="A7D30EE171CC5B9BC1BDD2B4456BA25E">Universidad del Magdalena, Grupo de Investigacion en Biodiversidad y Ecologia Aplicada, A. A 731 Santa Marta, Magdalena, Colombia</mods:affiliation>
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<mods:namePart id="115156CD50B3DEC2E6BDC86E32B8DC56">Suarez-Morales, Eduardo</mods:namePart>
<mods:nameIdentifier id="9152F242DD43BECDFF099B7CCE12CDCA" type="ORCID">https://orcid.org/0000-0003-2051-8707</mods:nameIdentifier>
<mods:affiliation id="F80E22B84D878113854A2B8C035BA2CD">El Colegio de la Frontera Sur (ECOSUR), A. P. 424, 77014 Chetumal, Quintana Roo, Mexico</mods:affiliation>
<mods:nameIdentifier id="CDB4AFD99988488587168454BD5DEE4E" type="email">esuarez@ecosur.mx</mods:nameIdentifier>
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<mods:namePart id="DD224CEE2EEBB75E63C616E3EC04523A">Silva-Briano, Marcelo</mods:namePart>
<mods:affiliation id="F35D1C3AA464F67343282F8B81F069D0">Departamento de Biologia, Universidad Autonoma de Aguascalientes (UAA), 20131 Aguascalientes, Mexico</mods:affiliation>
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<mods:date id="CDB9247B0179586FAA596198320E8B3B">2022</mods:date>
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<treatment id="5AF16EEF611F5A5A9AD1421032313815" ID-GBIF-Taxon="204393653" LSID="urn:lsid:zoobank.org:act:90D3530F-D17B-48E7-BC9A-90F41ED5C0E5" httpUri="http://treatment.plazi.org/id/5AF16EEF611F5A5A9AD1421032313815" lastPageNumber="111" pageId="0" pageNumber="111" scope_class="Copepoda" scope_family="Ameiridae" scope_order="Harpacticoida" scope_subFamily="Ameirinae">
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<paragraph id="326A5905308916BF1276C7B6A6A753EA" pageId="0" pageNumber="111">
<taxonomicName id="F2E7589AFA701CCF23BB2DDB45A60B54" LSID="https://zoobank.org/90D3530F-D17B-48E7-BC9A-90F41ED5C0E5" authority="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano, 2022" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis" status="sp. nov.">Nitokra puebloviejensis</taxonomicName>
<taxonomicNameLabel id="E7B7D00F1A892243E7B447E71A772329" pageId="0" pageNumber="111">sp. nov.</taxonomicNameLabel>
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<paragraph id="A5209AE3ECD1A961E2745DD5D394ED40" pageId="0" pageNumber="111">
<figureCitation id="3CDA3F33DDC9328ECCCEEC0318189354" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">Figs 1</figureCitation>
<figureCitation id="D3090EB235D23A78BA97E64A82E10517" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">, 2</figureCitation>
<figureCitation id="8DD0A541F0CA206CEA5596847C708A8B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">, 3</figureCitation>
<figureCitation id="C3B490ADCC5FD4B0B6DA527232D43B43" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">, 4</figureCitation>
<figureCitation id="889B2FC8A360600B303969B18CDEDE95" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">, 5</figureCitation>
<figureCitation id="686D52A60391F32115BD625DFF18B64C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0CD5EC2E4AA04CE396B1D755ED8B789F" pageId="0" pageNumber="111" type="materials_examined">
<paragraph id="EF8D7E87760A65AF607D706080166696" pageId="0" pageNumber="111">Material examined.</paragraph>
<paragraph id="36E9D32AC7D4A60300FB8793E98B1963" pageId="0" pageNumber="111">
<materialsCitation id="DED1DF2C23A879163FC3A578C8312145" accessionNumber="INV10139" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" location="Adult" specimenCount="1" typeStatus="holotype">
<location id="132BFCE6EEF9A02CC8D468A9C3A0403F" LSID="urn:lsid:plazi:treatment:5AF16EEF611F5A5A9AD1421032313815:132BFCE6EEF9A02CC8D468A9C3A0403F" name="Adult">Adult</location>
female 
<typeStatus id="E8139E7D93868EC1B19E61D37C4BB1E4">holotype</typeStatus>
(
<accessionNumber id="D0624CA4F2CF79A1D6304E345592812B" httpUri="http://www.ncbi.nlm.nih.gov/protein/INV10139">INV10139</accessionNumber>
)
</materialsCitation>
, 
<materialsCitation id="8CC0C73F179F7268872E76812CDCDB0D" accessionNumber="CRU10140" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="J. M. Fuentes-Reines" country="Colombia" location="Cienaga Grande de Santa Marta" longitude="-74.325455" specimenCount="1" typeStatus="allotype">
ethanol-preserved, vial, 
<location id="6B4B999EAFF5D1E7178F8B43D64810CF" LSID="urn:lsid:plazi:treatment:5AF16EEF611F5A5A9AD1421032313815:6B4B999EAFF5D1E7178F8B43D64810CF" country="Colombia" longitude="-74.325455" name="Cienaga Grande de Santa Marta">
<normalizedToken id="46F68B6C911AF73325420CEA57AB0099" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta
</location>
, 
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(10°5211.25&quot;N, 
<geoCoordinate id="62B3538991FDE08C6B2D6268E0BF2396" degrees="74" direction="west" minutes="19" orientation="longitude" precision="1" seconds="31.64" value="-74.325455">74°19'31.64&quot;W</geoCoordinate>
), littoral plankton, coll. 
<collectorName id="E9FD322B6322717470E68805D0C30814">
J.M. 
<normalizedToken id="29E650D11647F7A7A033D03E65F7EE5B" originalValue="Fuentes-Reinés">Fuentes-Reines</normalizedToken>
</collectorName>
; adult male 
<typeStatus id="8350AC5B330EB236FAFC1FBF00E005DD">allotype</typeStatus>
(INV 
<accessionNumber id="138DA9F0727FA93D31BC00838A3E0C0B" httpUri="http://www.ncbi.nlm.nih.gov/protein/CRU10140">CRU10140</accessionNumber>
)
</materialsCitation>
, 
<materialsCitation id="5B15D7E2DDB82BACD5DA92873BF2C386" accessionNumber="INV10139" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" location="Adult" specimenCount="1" typeStatus="Paratypes">
ethanol-preserved, vial, same sampling data as holotype. 
<typeStatus id="91AD92F146B3B62BD687AFA80FFAA8E5">Paratypes</typeStatus>
</materialsCitation>
: 
<materialsCitation id="8DE625A0AFC274B6363D497A643A63F5" accessionNumber="CRU10141, CRU10142" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" location="Adult" specimenCount="four" specimenCount-female="1" specimenCount-male="1" typeStatus="allotype">
<specimenCount id="70F607F08317B6A0D1D5EB87C2EDE3C0" type="female">four females</specimenCount>
(INV 
<accessionNumber id="9B7C861308047EF95D2001B9BEB8C6DD" httpUri="http://www.ncbi.nlm.nih.gov/protein/CRU10141">CRU10141</accessionNumber>
), and 
<specimenCount id="C04B69FF2FDC2328210B1ED82D839377" type="male">three males</specimenCount>
(INV 
<accessionNumber id="3D436145D8DA80429131F664185BA5A4" httpUri="http://www.ncbi.nlm.nih.gov/protein/CRU10142">CRU10142</accessionNumber>
), same sampling data as holotype and 
<typeStatus id="80870DDD316E1AAA0A7009585CAE56B8">allotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DAC2E770F490F2B183D7D2254E6BB6E4" pageId="0" pageNumber="111" type="materials_examined">
<paragraph id="B80AE60352AE39145AEE5B948C0D5564" pageId="0" pageNumber="111">Additional material.</paragraph>
<paragraph id="57089C9884F68C5B88CA4F157C1A00BB" pageId="0" pageNumber="111">
Six adult females, 
<specimenCount id="90A335B6D8BB99609F0ACE6605575B1D" type="adult">four adult</specimenCount>
males in first 
<normalizedToken id="AF59F0DAC8D6E1E28C2CA24EDE4C273B" originalValue="authors’">authors'</normalizedToken>
collection. One female and 
<specimenCount id="563A4E07C69E365E32D773788CEC0BBA" type="male">two male</specimenCount>
individuals prepared for SEM analysis.
</paragraph>
</subSubSection>
<subSubSection id="464392FFF72C6AE450086C11E7B6E8E4" pageId="0" pageNumber="111" type="type locality">
<paragraph id="E9DDFB30310D4ADD872443E98A3ED801" pageId="0" pageNumber="111">Type locality.</paragraph>
<paragraph id="9456020D6297FA922C1B7CD3AC5B0EA0" pageId="0" pageNumber="111">
Puebloviejo, 
<normalizedToken id="E7133A79F583F063F3CCE1E65D10D463" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, northern Colombia (
<geoCoordinate id="611B8282CEB50F356C74D2380137C534" degrees="10" direction="north" minutes="52" orientation="latitude" precision="1" seconds="11.25" value="10.869792">10°52'11.25&quot;N</geoCoordinate>
, 
<geoCoordinate id="601726A675FEFB7FA43E2891BF96D12B" degrees="74" direction="west" minutes="19" orientation="longitude" precision="1" seconds="31.64" value="-74.325455">74°19'31.64&quot;W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="5DF644E99BA09A9DEE3943988CE75E47" pageId="0" pageNumber="111" type="etymology">
<paragraph id="B82CACB4823BB00508D20CF0FA9BCB01" pageId="0" pageNumber="111">Etymology.</paragraph>
<paragraph id="FF5C53924C9BC4A3909AD94DD0ED5734" pageId="0" pageNumber="111">The new species is named in reference to the type locality of the new species by adding the toponimic suffix in singular. The gender of the species suffix is feminine to match that of the genus.</paragraph>
</subSubSection>
<subSubSection id="D12E7B341749F1A7A5B7D874F1F7ED71" pageId="0" pageNumber="111" type="differential diagnosis">
<paragraph id="2E312BF6CFC4722F7AB08637ECB0673A" pageId="0" pageNumber="111">Differential diagnosis.</paragraph>
<paragraph id="FFE52EE67CB4A6F5098EBC05646A938D" pageId="0" pageNumber="111">
<taxonomicName id="41FD32984A9D179C57D247F7228B776E" authorityName="Boeck" authorityYear="1865" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="047E7552071DE25BE18019037B1E777F" italics="true" pageId="0" pageNumber="111">Nitokra</emphasis>
</taxonomicName>
with 1 inner seta and 5 setae on P1EXP2 and EXP3, respectively and 455 and 777 elements on P2-P4ENP3 and P2-P4EXP3, respectively, plus 111 inner setae on P2-P4ENP1, respectively. Female rostrum hourglass-shaped, distal segment of mandibular palp with 6 setal elements. P1ENP1 almost reaching distal margin of P1EXP3. Modified, club-shaped inner basipodal seta on leg1. Maxilliped with distinctive group of 5 setae inserted medially on the syncoxa.
</paragraph>
</subSubSection>
<subSubSection id="88E0FBDD23473C445905F6A10DC2A724" pageId="0" pageNumber="111" type="description">
<paragraph id="0D469CA52A0C864CCB310A33EDDB45CE" pageId="0" pageNumber="111">Description of female.</paragraph>
<paragraph id="1511278AAA6A872588132CB76F5311A7" pageId="0" pageNumber="111">
Body subcylindrical, tapering posteriorly (Fig. 
<figureCitation id="BC5A6DE0CE743E45DAD0F31BC14ACBB3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1A</figureCitation>
), total body length 728-784 
<normalizedToken id="78244DD2B65241ED8FA09DF6F16CD3B8" originalValue="μm">μm</normalizedToken>
(average = 768 
<normalizedToken id="58B819DC3A93F0DEA4DE0E757D091F9A" originalValue="μm">μm</normalizedToken>
, 
<emphasis id="9B0AFB6D4BBA36EC7D793E0F6CCD6F58" italics="true" pageId="0" pageNumber="111">n</emphasis>
= 6; holotype length = 784 
<normalizedToken id="6395F90089722E99BA357A54DEAB66A1" originalValue="μm">μm</normalizedToken>
).
</paragraph>
<caption id="9A9B023BE18A70F8B12F7BE8BAE4931B" doi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111" start="Figure 1" startId="F1">
<paragraph id="7014474769EBF657D53221C316C4F5D5" pageId="0" pageNumber="111">
<emphasis id="88F6F0E669010CC401B63C8517E2F9C0" bold="true" pageId="0" pageNumber="111">Figure 1.</emphasis>
<taxonomicName id="3FD49620BBCDAC8A3AF978A9B04C2D2A" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="91D3B6AA8C8F9FE7CC559198EFA77104" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="322BADA86C617422C6D59220E7179C0D" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia, digital photos. 
<emphasis id="10A9187733C21EC633BA6DAF5D44988B" bold="true" pageId="0" pageNumber="111">A</emphasis>
holotype female, habitus, lateral view 
<emphasis id="DCAFD513A259ECFB95C3C6541CDDDCBF" bold="true" pageId="0" pageNumber="111">B</emphasis>
rostrum, ventral view 
<emphasis id="4AE128382B5C31CDB7E5FDE2187D5E58" bold="true" pageId="0" pageNumber="111">C</emphasis>
labrum, ventral view 
<emphasis id="6667B96C6B6E2F2596D1DEA0C1B0374F" bold="true" pageId="0" pageNumber="111">D</emphasis>
urosomites and caudal rami, ventral view 
<emphasis id="585F3B0AC68EAB4C3513A8174F4F2B0D" bold="true" pageId="0" pageNumber="111">E</emphasis>
anal somite with anal operculum and spine ornamentation, dorsal view 
<emphasis id="C5BBAFBF23CF50A0D764D81B7AFA5F5D" bold="true" pageId="0" pageNumber="111">F</emphasis>
anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 
<normalizedToken id="CE765710D4C6223F2FE3D246E1EBE6D9" originalValue="μm">μm</normalizedToken>
(
<emphasis id="62E5149688B3DFFAD13B607CA0955DF1" bold="true" pageId="0" pageNumber="111">A, C</emphasis>
); 10 
<normalizedToken id="441DF5EA3EC71CB7610E33264E3B5AC3" originalValue="μm">μm</normalizedToken>
(
<emphasis id="0816C4A1C5897AE8A9A0A51503359B3E" bold="true" pageId="0" pageNumber="111">B</emphasis>
); 20 
<normalizedToken id="D47DBF94492E05910CF348513D2453A7" originalValue="μm">μm</normalizedToken>
(
<emphasis id="3DA7C2808A2D140990A9A9104510DCA5" bold="true" pageId="0" pageNumber="111">E, F</emphasis>
).
</paragraph>
</caption>
<paragraph id="2BB39354E904C529161905A072615D1A" pageId="0" pageNumber="111">
Rostrum small, slightly protruding, discernible in dorsal view; roughly hourglass-shaped, with flat tip; rostrum furnished apically with two pairs of short slender sensilla (Figs 
<figureCitation id="14CF7DF1842E6400D2099F82595B7CDA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1B</figureCitation>
, 
<figureCitation id="07C5D0475F302FA038A194084B891DCC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2A</figureCitation>
). Labrum subtriangular, strong, heavily chitinized, with rugose edge apically (Figs 
<figureCitation id="4081445B959CD9F022348182CF478BA0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1C</figureCitation>
, 
<figureCitation id="C4A0BD54056A81DAA95922413A2CF3D9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2B</figureCitation>
). Urosome short, thick (Figs 
<figureCitation id="24625C6552216BF1E0C52C8DA793FF41" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1D</figureCitation>
, 
<figureCitation id="ECDD5CA683F168BD8B068D494091137C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3D-F</figureCitation>
), comprising fifth pedigerous somite, genital double-somite and 3 free abdominal somites. Anal somite furnished with row of spinules on proximal ventral surface and along insertion of caudal rami (Figs 
<figureCitation id="44AC3C16EC5AE0F96E6C5E23551420BA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1F</figureCitation>
, 
<figureCitation id="60927761E80F716817A96B7C57F5A4BF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3D-F</figureCitation>
, 
<figureCitation id="F9B8852B8C28D90CDE0CB97B6A32B9BD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4A</figureCitation>
). Anal operculum semicircular, ornamented with 5 or 6 spines (Figs 
<figureCitation id="B380CCED4AD8BDEBC45A3AE0C3A49CB3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1E</figureCitation>
, 
<figureCitation id="6B1A0CA9222D75B36199D7A3F1D81454" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4A</figureCitation>
). Caudal ramus short, subquadrate, armed with 7 caudal setae (Figs 
<figureCitation id="BF4F0007F59D90CE556CF45F97F586B0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1F</figureCitation>
, 
<figureCitation id="BB37229E49655FD6EBBAE71A14BD590A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4A</figureCitation>
), caudal seta I shorter than ramus; seta II about 2.6 
<normalizedToken id="959CCDF1F89EBE662A84F3ADAEAE20FB" originalValue="×">x</normalizedToken>
as long as I, seta III on distal outer position, slightly longer than seta II. Setae IV and V thick, long, the former being longest; seta VI about 1.6 
<normalizedToken id="902FB128E2A8BF2294AAFC592F9FEC63" originalValue="×">x</normalizedToken>
as long as seta I. Dorsal seta (VII) simple, about as long as caudal ramus, shorter than seta VI (Figs 
<figureCitation id="72C51677B4F20C6D7BCE28F936276AB1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia, digital photos. A holotype female, habitus, lateral view B rostrum, ventral view C labrum, ventral view D urosomites and caudal rami, ventral view E anal somite with anal operculum and spine ornamentation, dorsal view F anal somite and caudal rami showing caudal setae I-VII, ventral view. Scale bars: 50 μm (A, C); 10 μm (B); 20 μm (E, F)." figureDoi="10.3897/zookeys.1128.86210.figure1" httpUri="https://binary.pensoft.net/fig/766921" pageId="0" pageNumber="111">1F</figureCitation>
, 
<figureCitation id="8C638F1607E527E0BF2C9FD8EC1618B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4A</figureCitation>
).
</paragraph>
<paragraph id="2708A0864B63C32BD4B70DEBBE273DC5" pageId="0" pageNumber="111">
<emphasis id="4883F9487FC23D39E4F590D1A2AC0EC2" bold="true" italics="true" pageId="0" pageNumber="111">Antennule</emphasis>
8-segmented, tapering distally (Fig. 
<figureCitation id="D432ADF1E118FA14C2490BE545536936" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4E</figureCitation>
), first segment robust, subquadrate, unornamented, with single seta. Aesthetasc on fourth segment long, reaching beyond distal end of terminal segment. Fourth segment as long as first. Segmental armature as follows (s = seta, ae = aesthetasc): 1(1s),2(4s),3(6s),4(6+1ae).5(3s),6(2),7(4),8(7s+1ae).
</paragraph>
<paragraph id="3C066F9A2B4D9A9BE6212C84706D37CC" pageId="0" pageNumber="111">
<emphasis id="966DF9EA2E3580954745354AE3CB2317" bold="true" italics="true" pageId="0" pageNumber="111">Antenna</emphasis>
(Figs 
<figureCitation id="FDBD8DECA2EA66466F5FFB66E2FD3E8F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2E</figureCitation>
, 
<figureCitation id="9960414890F0960EEA98CA58D9AFADBF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3A</figureCitation>
). Coxa short, subquadrate, smooth. Basis subrectangular, lacking abexopodal seta, armed with short distal spine and row of minute spinules at spine insertion. First endopodal segment subrectangular, smooth, second endopodal segment longer than first, furnished with subdistal row of spinules on inner margin (Fig. 
<figureCitation id="EBEBA089D0006AE20CA3FC1DA1A1EB8E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3A</figureCitation>
), with 2 lateral inner spines and 6 apical setal elements, outermost 2 basally fused at insertion. EXP 1-segmented, cylindrical, armed with 3 subequal setal elements (Figs 
<figureCitation id="BD507E148383D8A58B72FC1756A78F29" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2E</figureCitation>
, 
<figureCitation id="B133C35EBBCEB77416B49C98D048E805" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3A</figureCitation>
, 
<figureCitation id="AAAB43A46F855428D62FC6CA85FCEA3A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6C, D</figureCitation>
).
</paragraph>
<caption id="5179FD09F096400994B1F3773075B409" doi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111" start="Figure 2" startId="F2">
<paragraph id="8002A07498F562C8F68BD8781DC9A6B8" pageId="0" pageNumber="111">
<emphasis id="822F14D2E444383587FF8D85191309A2" bold="true" pageId="0" pageNumber="111">Figure 2.</emphasis>
<taxonomicName id="F8C05D50FAB6FDB5CA0F48CB695EBCEC" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="46AEC8A218ED3B1583F3C2DBF7B054FD" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="051E07E2CB7BA2D2C9C12C8AED2A6B86" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia. Adult female holotype. 
<emphasis id="8B12BCD64DB615675ABBFA967591C360" bold="true" pageId="0" pageNumber="111">A</emphasis>
rostrum 
<emphasis id="4B27B123F2F89DFDFCE833701554E603" bold="true" pageId="0" pageNumber="111">B</emphasis>
labrum 
<emphasis id="095A842F80ECD806C383963ECE0DE0C0" bold="true" pageId="0" pageNumber="111">C</emphasis>
mandible with palp 
<emphasis id="C1F28B14DBCADB7890D257B2404365D5" bold="true" pageId="0" pageNumber="111">D</emphasis>
gnathal blade with teeth and dorsal seta 
<emphasis id="79154B47761623EFC2FD1B7791F68DCF" bold="true" pageId="0" pageNumber="111">E</emphasis>
antenna (some setae cut short) 
<emphasis id="7C267F5DA2C625EF53896059BBE2B40D" bold="true" pageId="0" pageNumber="111">F</emphasis>
maxillule showing armature of lobes 
<emphasis id="14FA0CA0CB22D1F639B52779248B160D" bold="true" pageId="0" pageNumber="111">G</emphasis>
maxilla 
<emphasis id="B10FC4C462FEDAA0C304DC7C49ACC67C" bold="true" pageId="0" pageNumber="111">H</emphasis>
maxilliped showing row of setal elements on syncoxa 
<emphasis id="D33B2E3583F62493ED9B97D8EAE5EA43" bold="true" pageId="0" pageNumber="111">I</emphasis>
detail of accessory seta of maxilliped ENP 
<emphasis id="C28394E0AEBF3FF2F0310FB4E2DC0312" bold="true" pageId="0" pageNumber="111">J</emphasis>
antennule showing segmentation. Scale bars: 15 
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(
<emphasis id="CD0754704FC22C8303A6EEF39E194017" bold="true" pageId="0" pageNumber="111">A-D, I</emphasis>
); 20 
<normalizedToken id="7A89DF0CCE0B65E5321136071F1A332B" originalValue="μm">μm</normalizedToken>
(
<emphasis id="2242A44C4A92F2DC32A3D56BE95D6AE5" bold="true" pageId="0" pageNumber="111">D-F</emphasis>
); 25 
<normalizedToken id="54C63828D8E86868428D2013F918EC7F" originalValue="μm">μm</normalizedToken>
(
<emphasis id="BC032CE2FDEDCE316BBCF75AA3390BBF" bold="true" pageId="0" pageNumber="111">G, H, J</emphasis>
).
</paragraph>
</caption>
<caption id="16CF859F53AE450457DC4B27AA3800C0" doi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111" start="Figure 3" startId="F3">
<paragraph id="89D5550FE85EBB9E52FA2D99B71DCDC6" pageId="0" pageNumber="111">
<emphasis id="2DF3984033B446D56511B7FBECC75371" bold="true" pageId="0" pageNumber="111">Figure 3.</emphasis>
<taxonomicName id="D670B3D2C0387FD7CAF1A2B620E3E747" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="9593465FB34541701795A44D14BE86C6" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="F343D5E18A180713F6FB03967CACDF2D" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia. SEM-prepared adult female. 
<emphasis id="A271095398FBEE9ECE46B61D3B597AD6" bold="true" pageId="0" pageNumber="111">A</emphasis>
antenna showing EXP and ENP 
<emphasis id="8A59FCA5013E91E3D82241F1B13319D2" bold="true" pageId="0" pageNumber="111">B</emphasis>
posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view 
<emphasis id="B566D8C3E0CCD414DEC2F5F292C27591" bold="true" pageId="0" pageNumber="111">C</emphasis>
fifth leg exopods (at arrow), ventral view 
<emphasis id="4F55B167392FFF5C53ACF308C936536C" bold="true" pageId="0" pageNumber="111">D</emphasis>
anal somite and caudal rami, ventral view 
<emphasis id="E73298C84FC5708A7AA038BB592800E7" bold="true" pageId="0" pageNumber="111">E</emphasis>
anal somite, ventral view 
<emphasis id="0B3BE69F5501B18CFEAED31DE83D2A86" bold="true" pageId="0" pageNumber="111">F</emphasis>
urosome, ventral view.
</paragraph>
</caption>
<paragraph id="AA09633D35281A38EC0F238588B00E82" pageId="0" pageNumber="111">
<emphasis id="4015530C79F955BE171D7B0F6871B542" bold="true" italics="true" pageId="0" pageNumber="111">Mandible</emphasis>
(Fig. 
<figureCitation id="C954757D10F10A308ADBBA9A54B94212" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2C, D</figureCitation>
). Gnathal blade armed with 13 teeth, 6 large, 7 small, 1 long spinule, and long dorsal seta ornamented with short spinules (Fig. 
<figureCitation id="D572ABB51D26FD2C107CCC9C1E780E98" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2D</figureCitation>
). Mandibular palp 2-segmented, basal segment short, subquadrate, with short robust seta (Fig. 
<figureCitation id="AF546413BFB9CDBC540C3B225694617B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2C</figureCitation>
). Endopodal segment subrectangular, armed with 1 short lateral and 5 apical setae (Fig. 
<figureCitation id="061C66FED10106781C229E3649BE8003" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2C</figureCitation>
).
</paragraph>
<paragraph id="276927ADCCA578E3C626BE040C503A7A" pageId="0" pageNumber="111">
<emphasis id="2255FB6B991DAE4B770EEC348CEACD94" bold="true" italics="true" pageId="0" pageNumber="111">Maxillule</emphasis>
(Fig. 
<figureCitation id="30F972D32B7F3C45FAE6B844E30C3290" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
). With large praecoxa; precoxal arthrite (pca in Fig. 
<figureCitation id="43B9F1554113E7861D5565DEFCF751CA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
) rectangular, unornamented, armed with group of 4 apical and 2 subapical spiniform elements. Coxal endite shorter than precoxal arthrite, armed with 1 curved serrate spiniform element and 2 smooth setae (cxe in Fig. 
<figureCitation id="CB438B2494FE48CB943E7C214A6516B9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
). Basis (bas in Fig. 
<figureCitation id="96F48CCAB9B963DF7C545A2C4E3E9DDC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
) shorter than coxal endite, seemingly with five subequal apical and subapical setae; exopod reduced, represented by 1 seta; endopod (enp in Fig. 
<figureCitation id="BF83A9E4222AF319EF32AB59D7E2AA14" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
) 1-segmented, armed with 2 subequal plumose setae inserted apically (Fig. 
<figureCitation id="21F08461F0DD3CE0B78D7C389756C1D3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2F</figureCitation>
).
</paragraph>
<paragraph id="10DC027396DBEB17CD4F5EF8DF7A3DC3" pageId="0" pageNumber="111">
<emphasis id="47662392CD8A16DAE5F6CA2F17EC06D5" bold="true" italics="true" pageId="0" pageNumber="111">Maxilla</emphasis>
(Fig. 
<figureCitation id="B6136589198C61B9FE5B285E5226BCE4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2G</figureCitation>
). Syncoxa unornamented, with 2 endites, proximalmost armed with short spiniform element and slender modified seta furnished with distal tuft of setules; second endite with 2 slender apical setae. Allobasis produced into strong serrate claw with short, curved adjacent spiniform element and slender seta. ENP 1-segmented, armed with 2 setae.
</paragraph>
<paragraph id="F1046406D669852A309D86FAB48D38E6" pageId="0" pageNumber="111">
<emphasis id="1C5DF201A7F6BF4FF5CC6BABD7960546" bold="true" italics="true" pageId="0" pageNumber="111">Maxilliped</emphasis>
(Fig. 
<figureCitation id="507B4E5E6DD3DAF796D1277B86984F86" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2H, I</figureCitation>
). Subchelate. Syncoxa with single setulated seta on inner distal corner and distinctive group of 4 or 5 short slender elements inserted medially on the outer margin of the syncoxa, as in male (Fig. 
<figureCitation id="5F11E6EA3D7EDBE9BEF1D9FD587A3FC0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2H</figureCitation>
, arrowhead in Fig. 
<figureCitation id="1623B7E95DEFAF8ACA5D040D19195608" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6E</figureCitation>
). ENP represented by long, slender claw with short accessory seta (Fig. 
<figureCitation id="80F42AA297E42DA04055AE9770E7E28A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype. A rostrum B labrum C mandible with palp D gnathal blade with teeth and dorsal seta E antenna (some setae cut short) F maxillule showing armature of lobes G maxilla H maxilliped showing row of setal elements on syncoxa I detail of accessory seta of maxilliped ENP J antennule showing segmentation. Scale bars: 15 μm (A-D, I); 20 μm (D-F); 25 μm (G, H, J)." figureDoi="10.3897/zookeys.1128.86210.figure2" httpUri="https://binary.pensoft.net/fig/766922" pageId="0" pageNumber="111">2I</figureCitation>
).
</paragraph>
<paragraph id="EABF5C5318D278B04B8EA24B25B3A732" pageId="0" pageNumber="111">
P1 (Fig. 
<figureCitation id="7CFF3CDBD197417BC5CF5259C0306F5D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">5A</figureCitation>
). Intercoxal sclerite smooth. Coxa with transverse row of spinules proximally and spinules row distally. Basis with spinules bordering insertion of exopodal and endopodal rami, inner basipodal spine short, reaching proximal 1/3 of length of first ENP segment. Outer basipodal spine short. EXP and ENP 3-segmented. Exopodal ramus shorter than endopod, reaching about the margin end of second exopodal segment. EXP1lacking inner setae, EXP2 with inner seta; EXP3 with 3 outer spines and 2 geniculate apical setae. ENP 3-segmented; ENP 2 with inner seta, ENP1 subrectangular. ENP3 with 1 apical spiniform seta, 1 geniculate apical seta, and short plumose inner seta.
</paragraph>
<paragraph id="811CE560C1D5CC77FF285208F43D0825" pageId="0" pageNumber="111">
P2 (Fig. 
<figureCitation id="27E8FD8084C11BE39849CA4A6AD24B8E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">5B-D</figureCitation>
). Intercoxal sclerite with transverse rows of long spinules (Fig. 
<figureCitation id="D5AA8CC88A5B9A4D67D81FBFC6B63E0E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6C</figureCitation>
). Coxa with transverse row of long spinules plus 2 rows of minute spinules (Fig. 
<figureCitation id="55A2C589065682A6CAD0DCCDFF5DF329" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6D</figureCitation>
). Outer basipodal spine short (
<normalizedToken id="411A9BD47C27E754E71454017DABCA0F" originalValue="“obs”">&quot;obs&quot;</normalizedToken>
in Fig. 
<figureCitation id="04362D50C4370C41F03BFEB877930B0D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6B</figureCitation>
). EXP and ENP 3-segmented. EXP1 lacking inner seta, EXP2 with long, slender inner seta, EXP3 with 3 outer spines, 1 apical and 3 inner setae. ENP shorter than EXP, reaching slightly beyond halfway of EXP3. ENP-3 with 2 inner setae (Fig. 
<figureCitation id="F3410686BE641248A8DFF7E487D72087" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">5B</figureCitation>
).
</paragraph>
<paragraph id="482D06118C2C879F03A814AB8E7C6BA8" pageId="0" pageNumber="111">
P3 (Fig. 
<figureCitation id="97D82EE77909892887EA736DFA0632EC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">5E</figureCitation>
). Intercoxal sclerite smooth. Coxa and basis as in P2. EXP and ENP 3-segmented. ENP slightly shorter than EXP. ENP as in P2 except for additional inner plumose seta on ENP3.
</paragraph>
<paragraph id="D06124FB5D9849E42EC16BD02C5961D8" pageId="0" pageNumber="111">
P4 (Fig. 
<figureCitation id="556567F221106579552F144B8798C13C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. Adult female holotype A leg 1 B leg 2 C leg 2 intercoxal sclerite D leg 2 coxa and basipod ornamentation, anterior view E leg 3 F leg 4. Scale bars: 20 μm (A, B, D-F); 10 μm (C)." figureDoi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111">5F</figureCitation>
). Intercoxal sclerite, coxa and basis as in P2 and P3. EXP and ENP 3-segmented. ENP shorter than EXP, barely reaching 1/3 of EXP3. EXP as in P3 except for thinner outer spines on EXP3. ENP as in P3.
</paragraph>
<paragraph id="4F51AD82203B5EFBC0C0906C6FDDDAAB" pageId="0" pageNumber="111">
P5 (Fig. 
<figureCitation id="D2BA02719529331078CAF58B8F3185C7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult female. A antenna showing EXP and ENP B posterior end of anal somite and insertion of caudal rami showing ornamentation, semi-lateral view C fifth leg exopods (at arrow), ventral view D anal somite and caudal rami, ventral view E anal somite, ventral view F urosome, ventral view." figureDoi="10.3897/zookeys.1128.86210.figure3" httpUri="https://binary.pensoft.net/fig/766923" pageId="0" pageNumber="111">3C</figureCitation>
at arrow, 4C). EXP and baseoendopod not fused, baseoendopod subtriangular, reaching EXP midlength, segment bearing 5 setae, apical being longest. EXP subquadrate, with spinules row along inner margin; EXP armed with 5 unequally long setae (Fig. 
<figureCitation id="70FA708A812CEDD3A696DEC5335A7B77" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4C</figureCitation>
).
</paragraph>
<caption id="9F258A7BA7EE185078BC8696451163A0" doi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111" start="Figure 4" startId="F4">
<paragraph id="F76EA71188D32600E70E3C772F641026" pageId="0" pageNumber="111">
<emphasis id="B2D09F78A1DBDF1679223CA0BC83EA2C" bold="true" pageId="0" pageNumber="111">Figure 4.</emphasis>
<taxonomicName id="A6DECD0A438B771EE5E1F0DC9A5E1CEC" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="BD41FC9598A0EB84F4D988F37AD22FF5" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="F14954AF4F2C0C6BAF00577A3DC912D4" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia. 
<emphasis id="6ACCC4AC08D794176F7CECCD8D4DE2AD" bold="true" pageId="0" pageNumber="111">A</emphasis>
adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII 
<emphasis id="044DB4A7ABE65AA8BE6C35D6429793DC" bold="true" pageId="0" pageNumber="111">B</emphasis>
female anal and preanal urosomites, ventral view 
<emphasis id="390199ECDFB7F65A02732A7A3FF5EE19" bold="true" pageId="0" pageNumber="111">C</emphasis>
female P5 
<emphasis id="1E04B92023FC78E3240F8AD6141498C2" bold="true" pageId="0" pageNumber="111">D</emphasis>
female genital field and P6 with setae (arrowed) 
<emphasis id="C0F6A6E23FAEBDE65DDA2004798F4987" bold="true" pageId="0" pageNumber="111">E</emphasis>
male geniculated antennule 
<emphasis id="7320051F6D6371DEBBA83112433A7268" bold="true" pageId="0" pageNumber="111">F</emphasis>
modified, club-shaped basipodal spine of male P1 
<emphasis id="D4DA36AD8F398CECEB47615E6162BC7B" bold="true" pageId="0" pageNumber="111">G</emphasis>
male leg 6 
<emphasis id="CA18E424A9AC7CA1E50839EBB16DE235" bold="true" pageId="0" pageNumber="111">H</emphasis>
male P3, anterior view 
<emphasis id="ED52C458CABEC126E2456AAE5370592E" bold="true" pageId="0" pageNumber="111">I</emphasis>
male P5, ventral view. Scale bars: 25 
<normalizedToken id="FA5F87B6D2F368554EAFEEEF8EDA5656" originalValue="μm">μm</normalizedToken>
(
<emphasis id="CA3859E3F157334AD090C3F753A4809B" bold="true" pageId="0" pageNumber="111">A-D, I</emphasis>
); 10 
<normalizedToken id="16429E06BADED482BE967364E2EE4318" originalValue="μm">μm</normalizedToken>
(
<emphasis id="25EE3E9B7EBA8267F05203558FDEB6FA" bold="true" pageId="0" pageNumber="111">E-G</emphasis>
); 20 
<normalizedToken id="CCB49D6C0E6EF64CB9A040EF6B7D6555" originalValue="μm">μm</normalizedToken>
(
<emphasis id="DEAE24FC3A73BA2595E7D2B1E362CEA5" bold="true" pageId="0" pageNumber="111">H</emphasis>
).
</paragraph>
</caption>
<paragraph id="791DEEC5F71EC97B1499C378CDBED64E" pageId="0" pageNumber="111">
P6 (Fig. 
<figureCitation id="45A6176274EC1592B2FBA310FB8D0F9A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4D</figureCitation>
). Represented by narrow transverse plate with subdistal lobe-like processes marked by a rounded notch. Plate bearing a small seta on each side (Fig. 
<figureCitation id="24A937DE34C91C54287B40F2523A3B47" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4D</figureCitation>
at arrows).
</paragraph>
<caption id="5FB2A3017A26FC97CDD0A8C76D260BCC" doi="10.3897/zookeys.1128.86210.figure5" httpUri="https://binary.pensoft.net/fig/766925" pageId="0" pageNumber="111" start="Figure 5" startId="F5">
<paragraph id="546199758D780BC78C49A21D623317BC" pageId="0" pageNumber="111">
<emphasis id="395A6CD848272F5BC7646A7780287D14" bold="true" pageId="0" pageNumber="111">Figure 5.</emphasis>
<taxonomicName id="C56403ED239CEFAB98BF6DCA478ED753" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="CB2FEDE27FF44574BBF2C9FA7BF33E4B" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="F39DEDD427BF743024746B08AAF0415D" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia. Adult female holotype 
<emphasis id="0D4A6F629D1023DC293C620C3591E9DC" bold="true" pageId="0" pageNumber="111">A</emphasis>
leg 1 
<emphasis id="B1B7C09C33F907141C1E33DF012B6269" bold="true" pageId="0" pageNumber="111">B</emphasis>
leg 2 
<emphasis id="52D8AE28965116C79AC25B3AA776B2F0" bold="true" pageId="0" pageNumber="111">C</emphasis>
leg 2 intercoxal sclerite 
<emphasis id="C7E6D84F14DE75F6BF5F0DE1856A20F4" bold="true" pageId="0" pageNumber="111">D</emphasis>
leg 2 coxa and basipod ornamentation, anterior view 
<emphasis id="B15CD976A672C40D7B47124EDE0D9F7C" bold="true" pageId="0" pageNumber="111">E</emphasis>
leg3 
<emphasis id="C45727DE016A4AC12DD027FA3C74291D" bold="true" pageId="0" pageNumber="111">F</emphasis>
leg 4. Scale bars: 20 
<normalizedToken id="3C3B75973777AA38AA5021212646364B" originalValue="μm">μm</normalizedToken>
(
<emphasis id="28ACDB4210352CE3FC2284538CE9C788" bold="true" pageId="0" pageNumber="111">A, B, D-F</emphasis>
); 10 
<normalizedToken id="58B1D83AF3E9BD1F20157D9D36992839" originalValue="μm">μm</normalizedToken>
(
<emphasis id="5845C8DC81ADF49F8D9A359BB80DB6D6" bold="true" pageId="0" pageNumber="111">C</emphasis>
).
</paragraph>
</caption>
<paragraph id="8A556F98CC3A5910F08078144530E012" pageId="0" pageNumber="111">Armature formula of female P1-P5 as follows:</paragraph>
<paragraph id="9971DD7E1905744653276D1D4B251BA9" pageId="0" pageNumber="111">
<table id="7FB40E9D32C2EC0FEDCD2F2FC14B49D6" pageId="0" pageNumber="111">
<tr id="92A080B0D58F918F92DEB08CBEF898A2" pageId="0" pageNumber="111">
<th id="85958CFFF9A61C6747B00C9280EC145D" colspan="1" isEmpty="true" pageId="0" pageNumber="111" rowspan="1">-</th>
<th id="E18511CDDAEDBE3E2EDBF157C8C7E044" colspan="1" pageId="0" pageNumber="111" rowspan="1">Exopod</th>
<th id="4EF1B3E6CF8F6ED41E082AE3C3AE3886" colspan="1" pageId="0" pageNumber="111" rowspan="1">Endopod</th>
</tr>
<tr id="6817991E6AE281EF40F861658FF65409" pageId="0" pageNumber="111">
<td id="DDE592E7A4680E735683A9DE0A0EC697" colspan="1" pageId="0" pageNumber="111" rowspan="1">P1</td>
<td id="057CC9C1FF77680D89E8A00EB7AF05D1" colspan="1" pageId="0" pageNumber="111" rowspan="1">I-0; I-1; III,2,0</td>
<td id="1FCF03028F3E89129F01F1B0671DEE0E" colspan="1" pageId="0" pageNumber="111" rowspan="1">0-1;0-1;I,I,2,0</td>
</tr>
<tr id="A8F981C4A933A36AA8195E7E32B3D1B6" pageId="0" pageNumber="111">
<td id="41491308962F509AC49673A0BB07B2BE" colspan="1" pageId="0" pageNumber="111" rowspan="1">P2</td>
<td id="4E300C5098AE025A77AFB477945CC34A" colspan="1" pageId="0" pageNumber="111" rowspan="1">I-0; I-1;III,2,2</td>
<td id="4585E3412CFA6635FEC875E7162E2FB5" colspan="1" pageId="0" pageNumber="111" rowspan="1">0-1;0-1; I,2,1</td>
</tr>
<tr id="B1D38C107E4F6828AF4C6C8BF6108165" pageId="0" pageNumber="111">
<td id="62A2FBCBAA25614CCC10D185592C9F3A" colspan="1" pageId="0" pageNumber="111" rowspan="1">P3</td>
<td id="E7E602C3912FC8DF72930CC452118D88" colspan="1" pageId="0" pageNumber="111" rowspan="1">I-0; I-1;III,2,2</td>
<td id="6C1332A5A3D36F5A38AA070212B194D5" colspan="1" pageId="0" pageNumber="111" rowspan="1">0-1;0-1; I,2,2</td>
</tr>
<tr id="3C7CAB074E16E45ED374A0C474AD63D8" pageId="0" pageNumber="111">
<td id="D3FA7679653C298FA89A4F2BDC4F1674" colspan="1" pageId="0" pageNumber="111" rowspan="1">P4</td>
<td id="0800995B3FAB3DCD7B398A5BE4DDC8A6" colspan="1" pageId="0" pageNumber="111" rowspan="1">I-0;I-1;III,2,2</td>
<td id="0C991311EDC25D477E018CC0B72181A0" colspan="1" pageId="0" pageNumber="111" rowspan="1">0-1;0-1;I,2,2</td>
</tr>
<tr id="5FFED94C60B90C5EF45BDD8A6542D091" pageId="0" pageNumber="111">
<td id="52B644852AB4245B34B04410295E69C8" colspan="1" pageId="0" pageNumber="111" rowspan="1">P5</td>
<td id="CD717BE1E9A522644E160D02EE80703A" colspan="1" pageId="0" pageNumber="111" rowspan="1">5</td>
<td id="2A0F2E1A5CACEA3ECC01FD951DC9DC9C" colspan="1" pageId="0" pageNumber="111" rowspan="1">5</td>
</tr>
</table>
</paragraph>
<paragraph id="71904C2E99FE07BC12C5EC124409A08C" pageId="0" pageNumber="111">
<emphasis id="14CB89D8D9983AF497F4B878C609B984" bold="true" pageId="0" pageNumber="111">Male.</emphasis>
Smaller than female, total body length 578-588 
<normalizedToken id="81A4CA614F6E8515FE59D49F54B7DE6F" originalValue="μm">μm</normalizedToken>
(average length = 578 
<normalizedToken id="A7DF3D07D0394A33CB851B599B32575D" originalValue="μm">μm</normalizedToken>
, 
<emphasis id="422C1F2BCF37C48FC9FF4B0B84456495" italics="true" pageId="0" pageNumber="111">n</emphasis>
= 3; allotype specimen length = 588 
<normalizedToken id="6A28993D6A21B6079F5BABC4A5791136" originalValue="μm">μm</normalizedToken>
).
</paragraph>
<paragraph id="4EF2A5CF9C6378DAD28F521100E5091E" pageId="0" pageNumber="111">
Anal operculum, rostrum, antennae (Fig. 
<figureCitation id="4682F3AEF98319627F7FB314551E3DFA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6C, D</figureCitation>
), and mouthparts as in female. Sexual dimorphism expressed in the antennule, urosome, P1basis, distal inner seta of P3ENP3, P5, and P6.
</paragraph>
<caption id="FD9D57A83FD3217D07C1067A8DEEDCF8" doi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111" start="Figure 6" startId="F6">
<paragraph id="96EA84185344D64386820759056081E1" pageId="0" pageNumber="111">
<emphasis id="4BF73280DDAD2FE8AC10754C53C1A6C2" bold="true" pageId="0" pageNumber="111">Figure 6.</emphasis>
<taxonomicName id="6BBAAEF36AE5A530913CBD452D762CFD" authorityName="Fuentes-Reinés &amp; Suárez-Morales &amp; Silva-Briano" authorityYear="2022" class="Copepoda" family="Ameiridae" genus="Nitokra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nitokra puebloviejensis" order="Harpacticoida" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="puebloviejensis">
<emphasis id="5733C78C63FE9CA9FB760A6E7FDD18FB" italics="true" pageId="0" pageNumber="111">Nitokra puebloviejensis</emphasis>
</taxonomicName>
sp. nov. from Puebloviejo, 
<normalizedToken id="FBBA64FC892AAEC001757BE164F06B88" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta, Colombia. SEM-prepared adult male individual 
<emphasis id="42FE61503581977748C7ABD2834A0156" bold="true" pageId="0" pageNumber="111">A</emphasis>
antennules, semi-lateral view 
<emphasis id="8C03D56CBDFB97FF07253454160629D6" bold="true" pageId="0" pageNumber="111">B</emphasis>
same, showing detail of proximal segments 
<emphasis id="9F2F1A3708330F4BB22E0224207199E9" bold="true" pageId="0" pageNumber="111">C</emphasis>
antenna, posterior view 
<emphasis id="2F88492CE1C461AE5BF01E74D91EC125" bold="true" pageId="0" pageNumber="111">D</emphasis>
same showing exopodal ramus and endopodal segments 
<emphasis id="353363541CB96157ACCFC070036EE534" bold="true" pageId="0" pageNumber="111">E</emphasis>
maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) 
<emphasis id="A7E49020F5DB90A5E7AB6D0DC010083F" bold="true" pageId="0" pageNumber="111">F</emphasis>
leg 1 showing modified, club-shaped basipodal spine (at arrow).
</paragraph>
</caption>
<paragraph id="FB6B2BFE8464E448F458F56748F809B9" pageId="0" pageNumber="111">
<emphasis id="58DDB5355FDD5FA8600180EA8FC92BCC" bold="true" italics="true" pageId="0" pageNumber="111">Antennule</emphasis>
(Fig. 
<figureCitation id="F903B093F233482665FBA451667F19E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6A, B</figureCitation>
) haplocer, 10-segmented; armature formula difficult to discern, purportedly as follows: 1(1s), 2(9s), 3(6s), 4(2s), 5(9+ae), 6(1s),7(3s), 8(1s),9(2s),10(7+ae).
</paragraph>
<paragraph id="3672EA793CFB1472A97DC179117F3B8A" pageId="0" pageNumber="111">
<emphasis id="A44C70EBCB24F74DB90C3706FFCFC55D" bold="true" italics="true" pageId="0" pageNumber="111">Antenna</emphasis>
(Fig. 
<figureCitation id="15BF197B5C8AE8ED7D58264587CA1835" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6C, D</figureCitation>
), maxilliped (Fig. 
<figureCitation id="32A0212A40812F54A0BD3758F840D9B8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6E</figureCitation>
), mouthparts, and P1-P4 as in female. Ventral ornamentation of urosomites as in female except urosomite 2. Posterior margins of urosomites with row of small spinules.
</paragraph>
<paragraph id="D11509C60D2CC52FEE4A95B37DD981FF" pageId="0" pageNumber="111">
P1 basis and P3ENP3. P1 basis with modified, club-shaped inner spine (Fig. 
<figureCitation id="FA599680F70BCE9ACF23A0C819063C8E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4F</figureCitation>
, arrowheads in Fig. 
<figureCitation id="4566840524C0DE261665DE3B68944F72" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. SEM-prepared adult male individual A antennules, semi-lateral view B same, showing detail of proximal segments C antenna, posterior view D same showing exopodal ramus and endopodal segments E maxilliped, ventral view showing row of setal elements on syncoxal medial surface (arrowhead) F leg 1 showing modified, club-shaped basipodal spine (at arrow)." figureDoi="10.3897/zookeys.1128.86210.figure6" httpUri="https://binary.pensoft.net/fig/766926" pageId="0" pageNumber="111">6F</figureCitation>
). P3ENP3 with distal inner spine thinner than in female.
</paragraph>
<paragraph id="63721369E660BC56F59813B514319BEE" pageId="0" pageNumber="111">
P5 (Fig. 
<figureCitation id="31DC0740F3866A2C655F47E55B6F3802" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nitokra puebloviejensis sp. nov. from Puebloviejo, Cienaga Grande de Santa Marta, Colombia. A adult female anal and preanal urosomites, dorsal view showing anal operculum and caudal setae I-VII B female anal and preanal urosomites, ventral view C female P 5 D female genital field and P 6 with setae (arrowed) E male geniculated antennule F modified, club-shaped basipodal spine of male P 1 G male leg 6 H male P 3, anterior view I male P 5, ventral view. Scale bars: 25 μm (A-D, I); 10 μm (E-G); 20 μm (H)." figureDoi="10.3897/zookeys.1128.86210.figure4" httpUri="https://binary.pensoft.net/fig/766924" pageId="0" pageNumber="111">4I</figureCitation>
). EXP and baseoendopod separated. The former subquadrate, with 5 or 6 setae, Baseoendopod reaching about proximal 1/3 of EXP, armed with 4 elements.
</paragraph>
<paragraph id="AB352FC96E2E119F053E8A2E2564E73B" pageId="0" pageNumber="111">
P6. With 2 unequal setae, inner one about 3 
<normalizedToken id="841A70DC51D009E14489DAFEFF1AD335" originalValue="×">x</normalizedToken>
as long as outer seta. Caudal rami as in female.
</paragraph>
</subSubSection>
<subSubSection id="D561B2276695581A3D7D239E8F52CE96" pageId="0" pageNumber="111" type="variability">
<paragraph id="D6F8EDEC22A9305CFE81BF1BA2E9BC5E" pageId="0" pageNumber="111">Variability.</paragraph>
<paragraph id="B25C439979C0DA4DD13129F88F18C24E" pageId="0" pageNumber="111">One male with 3 setal elements (instead of 4) on P5ENP. Another male was observed to possess 6 setae instead of 5 on P5EXP.</paragraph>
</subSubSection>
<subSubSection id="C13B05BF2BA438A50C308950787EAD6E" pageId="0" pageNumber="111" type="habitat">
<paragraph id="EF698DE138280586EA0243D9B8E91C5A" pageId="0" pageNumber="111">Habitat.</paragraph>
<paragraph id="E043C552E2D5B85DEC6F6B192F8291FB" pageId="0" pageNumber="111">
The new species is known only from the type locality, Puebloviejo, 
<normalizedToken id="25341DAEFA4D2597288DEAA34A3F5F90" originalValue="Ciénaga">Cienaga</normalizedToken>
Grande de Santa Marta (northern Colombia). The site where it was collected is a shallow mangrove area, 0.7 m deep, with water temperature 26-31 °C; local salinity was 15-20 PSU, and pH values was 7.5-8.1.
</paragraph>
</subSubSection>
</treatment>
</document>