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<mods:title id="03CAC23B578F127E4719153E9A41E1A4">Revision of the troglomorphic genus Troglostygnopsis Šilhavý, 1974 (Opiliones, Laniatores, Stygnopsidae)</mods:title>
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<mods:namePart id="67B2808E2ED9DF2DB7F495A8C1BE9B45">Cruz-López, Jesús A.</mods:namePart>
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<mods:affiliation id="58FF4A41D369CBAA561E9769D1E20FB3">Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. 3 er. Circuito exterior s / n. Apartado Postal 70 - 153, C. P. 04510, Ciudad Universitaria, Coyoacán, Mexico City, Mexico. &amp; Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Campo Experimental Valles Centrales, Oaxaca, Melchor Ocampo No. 7, C. P. 68200, Santo Domingo Barrio Bajo, Etla, Oaxaca, Mexico.</mods:affiliation>
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<mods:namePart id="C298D05925431CF852496B852360B42F">Francke, Oscar F.</mods:namePart>
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<mods:affiliation id="609651BB161DA8B10764E181B1D2AD28">Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. 3 er. Circuito exterior s / n. Apartado Postal 70 - 153, C. P. 04510, Ciudad Universitaria, Coyoacán, Mexico City, Mexico.</mods:affiliation>
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<treatment id="553D87E0034EC314BC3587B3FDC93388" ID-DOI="http://doi.org/10.5281/zenodo.5708140" ID-GBIF-Taxon="190303704" ID-Zenodo-Dep="5708140" LSID="urn:lsid:plazi:treatment:553D87E0034EC314BC3587B3FDC93388" httpUri="http://treatment.plazi.org/id/553D87E0034EC314BC3587B3FDC93388" lastPageId="11" lastPageNumber="85" pageId="6" pageNumber="80">
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<paragraph id="DD2B36F6034EC319BC3587B3FC49340D" blockId="6.[366,1222,1254,1351]" box="[602,985,1254,1281]" pageId="6" pageNumber="80">
<heading id="8663819A034EC319BC3587B3FC49340D" box="[602,985,1254,1281]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="80" reason="2">
<taxonomicName id="1A944D75034EC319BC3587B3FCE7340D" ID-CoL="BFXNT" authority="Cruz-Lopez &amp; Francke, 2021" box="[602,887,1254,1281]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="80" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE4034EC319BC3587B3FCE7340D" bold="true" box="[602,887,1254,1281]" italics="true" pageId="6" pageNumber="80">Troglostygnopsis kalebi</emphasis>
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<taxonomicNameLabel id="F4D3579F034EC319BD1087B2FC49340D" box="[895,985,1255,1281]" pageId="6" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="DD2B36F6034EC319BF01865FFB563428" blockId="6.[366,1222,1254,1351]" box="[366,1222,1290,1316]" pageId="6" pageNumber="80">
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="DD2B36F6034EC319BC888678FCDD344B" blockId="6.[366,1222,1254,1351]" box="[743,845,1325,1351]" pageId="6" pageNumber="80">
<figureCitation id="45AF2A73034EC319BC888678FCDD344B" box="[743,845,1325,1351]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[187,230,1989,2015]" captionStartId-1="8.[188,231,1928,1954]" captionStartId-2="9.[187,230,1153,1179]" captionStartId-3="9.[187,230,1963,1989]" captionTargetBox-0="[189,1398,414,1956]" captionTargetBox-1="[191,1400,596,1892]" captionTargetBox-2="[189,1398,317,1115]" captionTargetBox-3="[187,1396,1261,1929]" captionTargetId-0="figure-34@7.[189,1398,414,1956]" captionTargetId-1="figure-65@8.[191,1400,596,1892]" captionTargetId-2="figure-15@9.[189,1398,317,1115]" captionTargetId-3="figure-40@9.[187,1396,1261,1929]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="9" captionText-0="Fig. 4. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Dorsal view. B. Lateral view." captionText-1="Fig. 5. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalp, mesal view. D. Pedipalpal femur, mesal view. E. Pedipalpal tibia and tarsus, ventral view." captionText-2="Fig. 6. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Apical femur, patella and basal tibia IV. B. Patella IV." captionText-3="Fig. 7. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850), penis. A. Dorsal view. B. Lateral view. C. Ventral view. Scale bar in C applicable to all figures." figureDoi-0="http://doi.org/10.5281/zenodo.5708148" figureDoi-1="http://doi.org/10.5281/zenodo.5708150" figureDoi-2="http://doi.org/10.5281/zenodo.5708152" figureDoi-3="http://doi.org/10.5281/zenodo.5708154" httpUri-0="https://zenodo.org/record/5708148/files/figure.png" httpUri-1="https://zenodo.org/record/5708150/files/figure.png" httpUri-2="https://zenodo.org/record/5708152/files/figure.png" httpUri-3="https://zenodo.org/record/5708154/files/figure.png" pageId="6" pageNumber="80">Figs 47</figureCitation>
</paragraph>
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<paragraph id="DD2B36F6034EC319BED2863AFB933485" blockId="6.[189,1027,1390,1417]" box="[189,1027,1390,1417]" pageId="6" pageNumber="80">
<treatmentCitationGroup id="FD8411D8034EC319BED2863AFB933485" box="[189,1027,1390,1417]" pageId="6" pageNumber="80">
<taxonomicName id="1A944D75034EC319BED2863AFE393485" authorityName="Silhavy" authorityYear="1974" box="[189,425,1391,1417]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="80" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="EFE0EAE4034EC319BED2863AFE143485" box="[189,388,1391,1417]" italics="true" pageId="6" pageNumber="80">Troglostygnopsis</emphasis>
sp
</taxonomicName>
. CNANOp0050
<treatmentCitation id="5C3510E7034EC319BC02863AFC6F3485" author="Cruz-Lopez J. A. &amp; Francke O. F." box="[621,1023,1391,1417]" page="318" pageId="6" pageNumber="80" year="2017">
<bibRefCitation id="B9054B07034EC319BC02863AFC6F3485" author="Cruz-Lopez J. A. &amp; Francke O. F." box="[621,1023,1391,1417]" pageId="6" pageNumber="80" pagination="317 - 360" refId="ref5089" refString="Cruz-Lopez J. A. &amp; Francke O. F. 2017. Total evidence phylogeny of the North American harvestman family Stygnopsidae (Opiliones: Laniatores: Grassatores) reveals hidden diversity. Invertebrate Systematics 31: 317 - 360. https: // doi. org / 10.1071 / IS 16053" type="journal article" year="2017">Cruz-López &amp; Francke 2017: 318</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="958E657D034EC319BED286E5FE603733" pageId="6" pageNumber="80" type="diagnosis">
<paragraph id="DD2B36F6034EC319BED286E5FEA634C6" blockId="6.[189,310,1456,1482]" box="[189,310,1456,1482]" pageId="6" pageNumber="80">
<heading id="8663819A034EC319BED286E5FEA634C6" bold="true" box="[189,310,1456,1482]" fontSize="11" level="3" pageId="6" pageNumber="80" reason="3">
<emphasis id="EFE0EAE4034EC319BED286E5FEA634C6" bold="true" box="[189,310,1456,1482]" pageId="6" pageNumber="80">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034EC319BED2868AFE603733" blockId="6.[189,1399,1502,1599]" pageId="6" pageNumber="80">
Movable finger of chelicera with two teeth, low and wide (
<figureCitation id="45AF2A73034EC319BD1E868AFC5E34F5" box="[881,974,1502,1529]" captionStart="Fig" captionStartId="8.[188,231,1928,1954]" captionTargetBox="[191,1400,596,1892]" captionTargetId="figure-65@8.[191,1400,596,1892]" captionTargetPageId="8" captionText="Fig. 5. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalp, mesal view. D. Pedipalpal femur, mesal view. E. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708150" httpUri="https://zenodo.org/record/5708150/files/figure.png" pageId="6" pageNumber="80">Fig. 5B</figureCitation>
). Tubercles of leg IV large and very close each other, patella with many of these tubercles (
<figureCitation id="45AF2A73034EC319BD2D8557FC0F3711" box="[834,927,1538,1565]" captionStart="Fig" captionStartId="9.[187,230,1153,1179]" captionTargetBox="[189,1398,317,1115]" captionTargetId="figure-15@9.[189,1398,317,1115]" captionTargetPageId="9" captionText="Fig. 6. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Apical femur, patella and basal tibia IV. B. Patella IV." figureDoi="http://doi.org/10.5281/zenodo.5708152" httpUri="https://zenodo.org/record/5708152/files/figure.png" pageId="6" pageNumber="80">Fig. 6B</figureCitation>
). Penis with two pairs of MS D, MS E1 just above the level of E2.
</paragraph>
</subSubSection>
<subSubSection id="958E657D034EC319BED28532FB4637DE" pageId="6" pageNumber="80" type="etymology">
<paragraph id="DD2B36F6034EC319BED28532FED4378D" blockId="6.[189,324,1639,1665]" box="[189,324,1639,1665]" pageId="6" pageNumber="80">
<heading id="8663819A034EC319BED28532FED4378D" bold="true" box="[189,324,1639,1665]" fontSize="11" level="3" pageId="6" pageNumber="80" reason="3">
<emphasis id="EFE0EAE4034EC319BED28532FED4378D" bold="true" box="[189,324,1639,1665]" pageId="6" pageNumber="80">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034EC319BED285C0FB4637DE" blockId="6.[188,1398,1685,1747]" pageId="6" pageNumber="80">
Patronymic honoring our colleague and friend Kaleb Zárate, an enthusiastic speleologist who has helped us collecting troglomorphic arachnids during several expeditions in southeastern
<collectingCountry id="A5837666034EC319BA1B85EDFB4137DE" box="[1140,1233,1720,1746]" name="Mexico" pageId="6" pageNumber="80">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="958E657D034EC318BED285ACFEFE3066" lastPageId="7" lastPageNumber="81" pageId="6" pageNumber="80" type="materials_examined">
<paragraph id="DD2B36F6034EC319BED285ACFE3A361F" blockId="6.[189,426,1785,1811]" box="[189,426,1785,1811]" pageId="6" pageNumber="80">
<heading id="8663819A034EC319BED285ACFE3A361F" bold="true" box="[189,426,1785,1811]" fontSize="11" level="3" pageId="6" pageNumber="80" reason="3">
<emphasis id="EFE0EAE4034EC319BED285ACFE3A361F" bold="true" box="[189,426,1785,1811]" pageId="6" pageNumber="80">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034EC319BE85847DFB333685" blockId="6.[187,1398,1832,1929]" pageId="6" pageNumber="80">
<materialsCitation id="6DFC3CAB034EC319BE85847DFB333685" ID-GBIF-Occurrence="3407626304" collectingDate="2011-06-17" collectorName="O. Francke &amp; J. Cruz-Lopez &amp; R. Monjaraz-Ruedas &amp; G. Contreras &amp; K. Zarate" country="Mexico" latitude="16.300554" location="Cueva del Cipres" longLatPrecision="21" longitude="-92.318054" municipality="Las Rosas" pageId="6" pageNumber="80" specimenCode="CNAN-T0850" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="EFE0EAE4034EC319BE85847DFECC364E" bold="true" box="[234,348,1832,1858]" pageId="6" pageNumber="80">
<typeStatus id="022F8854034EC319BE85847DFECC364E" box="[234,348,1832,1858]" pageId="6" pageNumber="80" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="A5837666034EC319BED2841EFEA13669" box="[189,305,1867,1893]" name="Mexico" pageId="6" pageNumber="80">MEXICO</collectingCountry>
<specimenCount id="CB92FD7F034EC319BF298419FECE366A" box="[326,350,1868,1894]" pageId="6" pageNumber="80" type="male"></specimenCount>
;
<collectingRegion id="1F50F814034EC319BF05841EFE5B3669" box="[362,459,1867,1893]" country="Mexico" name="Chiapas" pageId="6" pageNumber="80">Chiapas</collectingRegion>
,
<collectingMunicipality id="3D4FAC8C034EC319BFB98419FDDF366A" box="[470,591,1868,1894]" pageId="6" pageNumber="80">Las Rosas</collectingMunicipality>
,
<location id="D84B602D034EC319BC36841EFCB43669" LSID="urn:lsid:plazi:treatment:553D87E0034EC314BC3587B3FDC93388:D84B602D034EC319BC36841EFCB43669" box="[601,804,1867,1893]" country="Mexico" latitude="16.300554" longLatPrecision="21" longitude="-92.318054" municipality="Las Rosas" name="Cueva del Cipres" pageId="6" pageNumber="80" stateProvince="Chiapas">Cueva del Ciprés</location>
;
<geoCoordinate id="B8A05031034EC319BD41841EFC21366A" box="[814,945,1866,1894]" degrees="16" direction="north" minutes="18" orientation="latitude" pageId="6" pageNumber="80" precision="15" seconds="2" value="16.300554">16º182″ N</geoCoordinate>
,
<geoCoordinate id="B8A05031034EC319BDD4841EFBD3366A" box="[955,1091,1866,1894]" degrees="92" direction="west" minutes="19" orientation="longitude" pageId="6" pageNumber="80" precision="15" seconds="5" value="-92.318054">92º195″ W</geoCoordinate>
;
<date id="A92A1036034EC319BA21841EFB743669" box="[1102,1252,1867,1894]" pageId="6" pageNumber="80" value="2011-06-17">
<collectingDate id="B96EE9DE034EC319BA21841EFB743669" box="[1102,1252,1867,1894]" pageId="6" pageNumber="80" value="2011-06-17">17 Jun. 2011</collectingDate>
</date>
;
<collectorName id="70615320034EC319BA81841EFAE33669" box="[1262,1395,1867,1893]" pageId="6" pageNumber="80">O. Francke</collectorName>
,
<collectorName id="70615320034EC319BED4843AFEF43685" box="[187,356,1903,1929]" pageId="6" pageNumber="80">J. Cruz-López</collectorName>
,
<collectorName id="70615320034EC319BF1F843AFDF63684" box="[368,614,1902,1929]" pageId="6" pageNumber="80">R. Monjaraz-Ruedas</collectorName>
,
<collectorName id="70615320034EC319BC1D843AFC983685" box="[626,776,1903,1929]" pageId="6" pageNumber="80">G. Contreras</collectorName>
and
<collectorName id="70615320034EC319BD2E843AFC3F3685" box="[833,943,1903,1929]" pageId="6" pageNumber="80">K. Zárate</collectorName>
leg.;
<specimenCode id="8D329E8D034EC319BD9E843BFB0F3685" box="[1009,1183,1902,1929]" pageId="6" pageNumber="80">CNAN-T0850</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD2B36F6034FC318BE85825CFEFE3066" blockId="7.[189,1398,265,362]" pageId="7" pageNumber="81">
<materialsCitation id="6DFC3CAB034FC318BE85825CFEFA3066" ID-GBIF-Occurrence="3407626303" collectingDate="2011-06-17" collectorName="O. Francke &amp; J. Cruz-Lopez &amp; R. Monjaraz-Ruedas &amp; G. Contreras &amp; K. Zarate" country="Mexico" latitude="16.300554" location="MEXICO" longLatPrecision="21" longitude="-92.318054" municipality="Las Rosas" pageId="7" pageNumber="81" specimenCode="CNAN-T0851" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="EFE0EAE4034FC318BE85825CFEF7302F" bold="true" box="[234,359,265,291]" pageId="7" pageNumber="81">
<typeStatus id="022F8854034FC318BE85825CFEF7302F" box="[234,359,265,291]" pageId="7" pageNumber="81" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="A5837666034FC318BED28279FEA1304A" box="[189,305,300,326]" name="Mexico" pageId="7" pageNumber="81">MEXICO</collectingCountry>
<specimenCount id="CB92FD7F034FC318BF208279FEEE304A" box="[335,382,300,326]" pageId="7" pageNumber="81" type="male">1 ♂</specimenCount>
,
<specimenCount id="CB92FD7F034FC318BFE08279FE2E304A" box="[399,446,300,326]" pageId="7" pageNumber="81" type="female">1 ♀</specimenCount>
; same collection data as for holotype; male is a DNA voucher CNANOp0050;
<specimenCode id="8D329E8D034FC318BED2821AFEFA3066" box="[189,362,335,362]" pageId="7" pageNumber="81">CNAN-T0851</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="89EB667E034FC318BED48490FAFF36D3" ID-DOI="http://doi.org/10.5281/zenodo.5708148" ID-Zenodo-Dep="5708148" box="[187,1391,1989,2016]" httpUri="https://zenodo.org/record/5708148/files/figure.png" pageId="7" pageNumber="81" startId="7.[187,230,1989,2015]" targetBox="[189,1398,414,1956]" targetPageId="7">
<paragraph id="DD2B36F6034FC318BED48490FAFF36D3" blockId="7.[187,1391,1989,2016]" box="[187,1391,1989,2016]" pageId="7" pageNumber="81">
<emphasis id="EFE0EAE4034FC318BED48490FE9936D3" bold="true" box="[187,265,1989,2015]" pageId="7" pageNumber="81">Fig. 4.</emphasis>
<taxonomicName id="1A944D75034FC318BF7F8490FDB736D3" box="[272,551,1989,2015]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="81" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE4034FC318BF7F8490FDB736D3" box="[272,551,1989,2015]" italics="true" pageId="7" pageNumber="81">Troglostygnopsis kalebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D3579F034FC318BC418493FD1B36EC" box="[558,651,1990,2016]" pageId="7" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (CNAN-T0850).
<emphasis id="EFE0EAE4034FC318BD9A8490FB9A36D3" bold="true" box="[1013,1034,1989,2015]" pageId="7" pageNumber="81">A</emphasis>
. Dorsal view.
<emphasis id="EFE0EAE4034FC318BADB8493FB5836EC" bold="true" box="[1204,1224,1990,2016]" pageId="7" pageNumber="81">B</emphasis>
. Lateral view.
</paragraph>
</caption>
<subSubSection id="958E657D0340C314BED2825DFC1F3318" lastPageId="11" lastPageNumber="85" pageId="8" pageNumber="82" type="description">
<paragraph id="DD2B36F60340C317BED2825DFEDE302E" blockId="8.[189,334,264,290]" box="[189,334,264,290]" pageId="8" pageNumber="82">
<heading id="8663819A0340C317BED2825DFEDE302E" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="8" pageNumber="82" reason="3">
<emphasis id="EFE0EAE40340C317BED2825DFEDE302E" bold="true" box="[189,334,264,290]" pageId="8" pageNumber="82">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F60340C317BE858262FDD5305E" blockId="8.[189,834,311,373]" box="[234,581,311,338]" pageId="8" pageNumber="82">
<emphasis id="EFE0EAE40340C317BE858262FEBA305D" bold="true" box="[234,298,311,337]" pageId="8" pageNumber="82">Male</emphasis>
(based on the
<typeStatus id="022F88540340C317BFBA8262FDAE305E" box="[469,574,311,338]" pageId="8" pageNumber="82" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="DD2B36F60340C317BED2820EFCD23078" blockId="8.[189,834,311,373]" box="[189,834,346,373]" pageId="8" pageNumber="82">MEASUREMENTS. Scutum length: 5.0, scutum width: 4.2.</paragraph>
<paragraph id="DD2B36F60340C317BED282F6FAE630EC" blockId="8.[189,1399,414,480]" pageId="8" pageNumber="82">
BODY (
<figureCitation id="45AF2A730340C317BF7F82F7FECB30B0" box="[272,347,418,444]" captionStart="Fig" captionStartId="7.[187,230,1989,2015]" captionTargetBox="[189,1398,414,1956]" captionTargetId="figure-34@7.[189,1398,414,1956]" captionTargetPageId="7" captionText="Fig. 4. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Dorsal view. B. Lateral view." figureDoi="http://doi.org/10.5281/zenodo.5708148" httpUri="https://zenodo.org/record/5708148/files/figure.png" pageId="8" pageNumber="82">Fig. 4</figureCitation>
). Scutum
<typeStatus id="022F88540340C317BFB882F6FD9A30B1" box="[471,522,419,445]" pageId="8" pageNumber="82">type</typeStatus>
zeta ‘ζ’, with both constrictions marked, specially C1, which marks the limits between prosoma and opisthosoma. Lateral margins of the scutum projected in lateral clear areas,
</paragraph>
<caption id="89EB667E0340C317BED384DDFD8536E4" ID-DOI="http://doi.org/10.5281/zenodo.5708150" ID-Zenodo-Dep="5708150" httpUri="https://zenodo.org/record/5708150/files/figure.png" pageId="8" pageNumber="82" startId="8.[188,231,1928,1954]" targetBox="[191,1400,596,1892]" targetPageId="8">
<paragraph id="DD2B36F60340C317BED384DDFD8536E4" blockId="8.[188,1401,1928,2025]" pageId="8" pageNumber="82">
<emphasis id="EFE0EAE40340C317BED384DDFE8136AE" bold="true" box="[188,273,1928,1954]" pageId="8" pageNumber="82">Fig. 5.</emphasis>
<taxonomicName id="1A944D750340C317BF4F84DDFDAE36AE" box="[288,574,1928,1954]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="82" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE40340C317BF4F84DDFDAE36AE" box="[288,574,1928,1954]" italics="true" pageId="8" pageNumber="82">Troglostygnopsis kalebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D3579F0340C317BC2384DCFD2036AF" box="[588,688,1929,1955]" pageId="8" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (CNAN-T0850).
<emphasis id="EFE0EAE40340C317BA5984DDFBDB36AE" bold="true" box="[1078,1099,1928,1954]" pageId="8" pageNumber="82">A</emphasis>
. Chelicera, mesal view.
<emphasis id="EFE0EAE40340C317BED284FEFF4136C9" bold="true" box="[189,209,1963,1989]" pageId="8" pageNumber="82">B</emphasis>
. Chelicera, frontal view.
<emphasis id="EFE0EAE40340C317BC6984FEFD8B36C9" bold="true" box="[518,539,1963,1989]" pageId="8" pageNumber="82">C</emphasis>
. Pedipalp, mesal view.
<emphasis id="EFE0EAE40340C317BD5384FEFCC136C9" bold="true" box="[828,849,1963,1989]" pageId="8" pageNumber="82">D</emphasis>
. Pedipalpal femur, mesal view.
<emphasis id="EFE0EAE40340C317BAB884FEFB7B36C9" bold="true" box="[1239,1259,1963,1989]" pageId="8" pageNumber="82">E</emphasis>
. Pedipalpal tibia and tarsus, ventral view.
</paragraph>
</caption>
<caption id="89EB667E0341C316BED487D4FE2D35B3" ID-DOI="http://doi.org/10.5281/zenodo.5708152" ID-Zenodo-Dep="5708152" httpUri="https://zenodo.org/record/5708152/files/figure.png" pageId="9" pageNumber="83" startId="9.[187,230,1153,1179]" targetBox="[189,1398,317,1115]" targetPageId="9">
<paragraph id="DD2B36F60341C316BED487D4FE2D35B3" blockId="9.[186,1398,1153,1215]" pageId="9" pageNumber="83">
<emphasis id="EFE0EAE40341C316BED487D4FE983597" bold="true" box="[187,264,1153,1179]" pageId="9" pageNumber="83">Fig. 6.</emphasis>
<taxonomicName id="1A944D750341C316BF6187D4FDB43597" box="[270,548,1153,1179]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="83" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE40341C316BF6187D4FDB43597" box="[270,548,1153,1179]" italics="true" pageId="9" pageNumber="83">Troglostygnopsis kalebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D3579F0341C316BC4587D4FD163597" box="[554,646,1153,1179]" pageId="9" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (CNAN-T0850).
<emphasis id="EFE0EAE40341C316BD8387D4FB913597" bold="true" box="[1004,1025,1153,1179]" pageId="9" pageNumber="83">A</emphasis>
. Apical femur, patella and basal tibia IV.
<emphasis id="EFE0EAE40341C316BF4F87F1FEA435B2" bold="true" box="[288,308,1188,1214]" pageId="9" pageNumber="83">B</emphasis>
. Patella IV.
</paragraph>
</caption>
<caption id="89EB667E0341C316BED484FEFC1D36E4" ID-DOI="http://doi.org/10.5281/zenodo.5708154" ID-Zenodo-Dep="5708154" httpUri="https://zenodo.org/record/5708154/files/figure.png" pageId="9" pageNumber="83" startId="9.[187,230,1963,1989]" targetBox="[187,1396,1261,1929]" targetPageId="9">
<paragraph id="DD2B36F60341C316BED484FEFC1D36E4" blockId="9.[187,1398,1963,2025]" pageId="9" pageNumber="83">
<emphasis id="EFE0EAE40341C316BED484FEFE9936C9" bold="true" box="[187,265,1963,1989]" pageId="9" pageNumber="83">Fig. 7.</emphasis>
<taxonomicName id="1A944D750341C316BF7F84FEFDB736C9" box="[272,551,1963,1989]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="83" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE40341C316BF7F84FEFDB736C9" box="[272,551,1963,1989]" italics="true" pageId="9" pageNumber="83">Troglostygnopsis kalebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D3579F0341C316BC4184F9FD1B36CA" box="[558,651,1964,1990]" pageId="9" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (CNAN-T0850), penis.
<emphasis id="EFE0EAE40341C316BA2D84FEFBC736C9" bold="true" box="[1090,1111,1963,1989]" pageId="9" pageNumber="83">A</emphasis>
. Dorsal view.
<emphasis id="EFE0EAE40341C316BB6E84FEFA8536C9" bold="true" box="[1281,1301,1963,1989]" pageId="9" pageNumber="83">B</emphasis>
. Lateral view.
<emphasis id="EFE0EAE40341C316BF6E849BFE8636E4" bold="true" box="[257,278,1998,2024]" pageId="9" pageNumber="83">C</emphasis>
. Ventral view. Scale bar in C applicable to all figures.
</paragraph>
</caption>
<paragraph id="DD2B36F60342C315BED2825CFC71304B" blockId="10.[189,1399,265,327]" pageId="10" pageNumber="84">large and rounded. Entire dorsum smooth, with few small tubercles in middle of each mesotergal area. Ocularium at the frontal margin of the prosoma, acute, without eyes.</paragraph>
<paragraph id="DD2B36F60342C315BED2823BFCCB30FD" blockId="10.[189,1400,365,498]" pageId="10" pageNumber="84">
CHELICERA (
<figureCitation id="45AF2A730342C315BF29823BFE1B3084" box="[326,395,365,392]" captionStart="Fig" captionStartId="8.[188,231,1928,1954]" captionTargetBox="[191,1400,596,1892]" captionTargetId="figure-65@8.[191,1400,596,1892]" captionTargetPageId="8" captionText="Fig. 5. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalp, mesal view. D. Pedipalpal femur, mesal view. E. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708150" httpUri="https://zenodo.org/record/5708150/files/figure.png" pageId="10" pageNumber="84">Fig. 5</figureCitation>
AB). Scutum/cheliceral hand ratio: 1.29. Basichelicerite large and smooth, with few small tubercles on ventral and apical regions. Cheliceral hand slightly widened but not swollen. Fixed finger with four teeth distributed evenly throughout, central triangular-shaped. Movable finger with two teeth, one basal and one subterminal, basalmost rounded.
</paragraph>
<paragraph id="DD2B36F60342C315BED2814CFA973226" blockId="10.[186,1399,536,810]" pageId="10" pageNumber="84">
PEDIPALPS (
<figureCitation id="45AF2A730342C315BF57814CFEEA333F" box="[312,378,536,563]" captionStart="Fig" captionStartId="8.[188,231,1928,1954]" captionTargetBox="[191,1400,596,1892]" captionTargetId="figure-65@8.[191,1400,596,1892]" captionTargetPageId="8" captionText="Fig. 5. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalp, mesal view. D. Pedipalpal femur, mesal view. E. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708150" httpUri="https://zenodo.org/record/5708150/files/figure.png" pageId="10" pageNumber="84">Fig. 5</figureCitation>
CE). Measurements: 2.31/1.10/1.80/1.80/1.10.Trochanter globose, with long spiniform setiferous tubercles on ventral and dorsal faces. Femur compressed laterally, armed with dorsal and ventral row of about 10 very large spiniform setiferous tubercles. Additionally, femur with mesal apical spiniform tubercle, near patella. Patella covered with spiniform tubercles on dorsal and mesal faces. Tibia with large main armature, relative sizes of SST on mesal side 1 = 3&gt; 2, and on ectal margin 3 = 2 = 1. Also on tibia three very small tubercles between SST 1, SST 2 and SST 3 on mesal side, and four small ones between SST 1 and SST 2, one between SST2 and SST 3, and one apically. Tarsus with six major tubercles on mesal side (1&gt; 2&gt; 3&gt; 4&gt; 5&gt; 6) and six on ectal side (1&gt; 2&gt; 3&gt; 4&gt; 5).
</paragraph>
<paragraph id="DD2B36F60342C315BED28004FBBA32BD" blockId="10.[189,1398,848,945]" pageId="10" pageNumber="84">
LEGS (
<figureCitation id="45AF2A730342C315BF698005FEDD3267" box="[262,333,848,875]" captionStart="Fig" captionStartId="9.[187,230,1153,1179]" captionTargetBox="[189,1398,317,1115]" captionTargetId="figure-15@9.[189,1398,317,1115]" captionTargetPageId="9" captionText="Fig. 6. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850). A. Apical femur, patella and basal tibia IV. B. Patella IV." figureDoi="http://doi.org/10.5281/zenodo.5708152" httpUri="https://zenodo.org/record/5708152/files/figure.png" pageId="10" pageNumber="84">Fig. 6</figureCitation>
). All segments long and slender, legs IIII without armature, ornate only with few disperse small tubercles. Ornamentation of femur IV formed by rounded and acuminate tubercles, which form longitudinal rows along entire femur, patella covered with many tubercles.
</paragraph>
<caption id="89EB667E0342C315BED385AEFD6B36E5" ID-DOI="http://doi.org/10.5281/zenodo.5708156" ID-Zenodo-Dep="5708156" httpUri="https://zenodo.org/record/5708156/files/figure.png" pageId="10" pageNumber="84" startId="10.[188,231,1787,1813]" targetBox="[188,1397,1077,1752]" targetPageId="10">
<paragraph id="DD2B36F60342C315BED385AEFD6B36E5" blockId="10.[187,1401,1787,2025]" pageId="10" pageNumber="84">
<emphasis id="EFE0EAE40342C315BED385AEFE823619" bold="true" box="[188,274,1787,1813]" pageId="10" pageNumber="84">Fig. 8.</emphasis>
Distribution of selected troglobitic genera of
<taxonomicName id="1A944D750342C315BD0A85AEFB433619" authority="Sorensen, 1932" authorityName=", Sorensen" authorityYear="1932" box="[869,1235,1787,1813]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="family">Stygnopsidae Sørensen, 1932</taxonomicName>
. Squares are distribution of
<taxonomicName id="1A944D750342C315BF01844BFD763634" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[366,742,1822,1851]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="EFE0EAE40342C315BF01844BFDA53634" box="[366,565,1822,1848]" italics="true" pageId="10" pageNumber="84">Troglostygnopsis</emphasis>
<bibRefCitation id="B9054B070342C315BC52844AFD763634" author="Silhavy V." box="[573,742,1822,1851]" pageId="10" pageNumber="84" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
: black =
<taxonomicName id="1A944D750342C315BD3A844AFB2E3634" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[853,1214,1822,1851]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE40342C315BD3A844AFB9E3634" box="[853,1038,1822,1848]" italics="true" pageId="10" pageNumber="84">T. anophthalma</emphasis>
<bibRefCitation id="B9054B070342C315BA79844AFB2E3634" author="Silhavy V." box="[1046,1214,1822,1851]" pageId="10" pageNumber="84" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
; red =
<taxonomicName id="1A944D750342C315BB7C844AFAE93634" box="[1299,1401,1822,1848]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="kalebi" status="sp. nov.">
<emphasis id="EFE0EAE40342C315BB7C844AFAE93634" box="[1299,1401,1822,1848]" italics="true" pageId="10" pageNumber="84">T. kalebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D3579F0342C315BED38417FE873650" box="[188,279,1858,1884]" pageId="10" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
Circles show distribution of
<emphasis id="EFE0EAE40342C315BC028417FD503657" box="[621,704,1858,1883]" italics="true" pageId="10" pageNumber="84">Brujita</emphasis>
Cruz-López, in press,
<emphasis id="EFE0EAE40342C315BDA38414FB8C3657" box="[972,1052,1857,1883]" italics="true" pageId="10" pageNumber="84">Toojah</emphasis>
Cruz-López, in press and the dubious record for
<taxonomicName id="1A944D750342C315BFF88430FC583673" authority="(Goodnight &amp; Goodnight, 1971)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[407,968,1893,1919]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="EFE0EAE40342C315BFF88430FDD73673" box="[407,583,1893,1919]" italics="true" pageId="10" pageNumber="84">Mictlana inops</emphasis>
(
<bibRefCitation id="B9054B070342C315BC398430FC2F3673" author="Goodnight C. J. &amp; Goodnight M. L." box="[598,959,1893,1919]" pageId="10" pageNumber="84" pagination="33 - 45" refId="ref5499" refString="Goodnight C. J. &amp; Goodnight M. L. 1971. Opilionids (Phalangida) of the family Phalangofidae from Mexican caves. Bulletin of the Association for Mexican Cave Studies 4: 33 - 45." type="journal article" year="1971">Goodnight &amp; Goodnight, 1971</bibRefCitation>
)
</taxonomicName>
reported by
<bibRefCitation id="B9054B070342C315BA368430FE9D36AE" author="Goodnight C. J. &amp; Goodnight M. L." pageId="10" pageNumber="84" pagination="83 - 96" refId="ref5538" refString="Goodnight C. J. &amp; Goodnight M. L. 1973. Opilionids (Phalangida) from Mexican caves. Bulletin of the Association for Mexican Cave Studies 5: 83 - 96." type="journal article" year="1973">Goodnight &amp; Goodnight (1973)</bibRefCitation>
: black =
<emphasis id="EFE0EAE40342C315BFEB84DCFDB536AE" box="[388,549,1928,1954]" italics="true" pageId="10" pageNumber="84">B. chapulapa</emphasis>
Cruz-López, in press; red =
<taxonomicName id="1A944D750342C315BDE184DCFB3736AE" authority="Cruz-Lopez" authorityName="Cruz-Lopez" box="[910,1191,1928,1954]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="cimutaa">
<emphasis id="EFE0EAE40342C315BDE184DCFB9D36AE" box="[910,1037,1929,1954]" italics="true" pageId="10" pageNumber="84">T. cimutaa</emphasis>
Cruz-López
</taxonomicName>
, in press; blue =
<taxonomicName id="1A944D750342C315BED384F9FE2336CA" authority="? Pentagons" authorityName="Pentagons" box="[188,435,1963,1990]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="EFE0EAE40342C315BED384F9FEB536C9" box="[188,293,1964,1989]" italics="true" pageId="10" pageNumber="84">M. inops</emphasis>
? Pentagons
</taxonomicName>
are the distribution of
<taxonomicName id="1A944D750342C315BCA684FEFB0D36CA" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[713,1181,1963,1990]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="EFE0EAE40342C315BCA684FEFCA336C9" box="[713,819,1963,1989]" italics="true" pageId="10" pageNumber="84">Mictlana</emphasis>
<bibRefCitation id="B9054B070342C315BD5584FEFB0D36CA" author="Cruz-Lopez J. A. &amp; Francke O. F." box="[826,1181,1963,1990]" pageId="10" pageNumber="84" pagination="827 - 891" refId="ref4973" refString="Cruz-Lopez J. A. &amp; Francke O. F. 2015. Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goonight, 1944 (Opiliones, Laniatores, Stygnopsidae). Zoological Journal of the Linnean Society 175: 827 - 891. https: // doi. org / 10.1111 / zoj. 12299" type="journal article" year="2015">Cruz-López &amp; Francke, 2015</bibRefCitation>
</taxonomicName>
: black =
<taxonomicName id="1A944D750342C315BB6284F9FDAD36E4" authority="(Goodnight &amp; Goodnight, 1973)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1973" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="EFE0EAE40342C315BB6284F9FAE936C9" box="[1293,1401,1963,1989]" italics="true" pageId="10" pageNumber="84">M. plana</emphasis>
(
<bibRefCitation id="B9054B070342C315BEAB849BFDA436E4" author="Goodnight C. J. &amp; Goodnight M. L." box="[196,564,1998,2024]" pageId="10" pageNumber="84" pagination="83 - 96" refId="ref5538" refString="Goodnight C. J. &amp; Goodnight M. L. 1973. Opilionids (Phalangida) from Mexican caves. Bulletin of the Association for Mexican Cave Studies 5: 83 - 96." type="journal article" year="1973">Goodnight &amp; Goodnight, 1973</bibRefCitation>
)
</taxonomicName>
; red =
<taxonomicName id="1A944D750342C315BCE2849AFD6536E4" box="[653,757,1999,2024]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="EFE0EAE40342C315BCE2849AFD6536E4" box="[653,757,1999,2024]" italics="true" pageId="10" pageNumber="84">M. inops</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="DD2B36F60343C314BED2825CFD8E30BC" blockId="11.[188,1401,265,432]" pageId="11" pageNumber="85">
PENIS (
<figureCitation id="45AF2A730343C314BF65825CFEC2302F" box="[266,338,265,291]" captionStart="Fig" captionStartId="9.[187,230,1963,1989]" captionTargetBox="[187,1396,1261,1929]" captionTargetId="figure-40@9.[187,1396,1261,1929]" captionTargetPageId="9" captionText="Fig. 7. Troglostygnopsis kalebi sp. nov., ♂, holotype (CNAN-T0850), penis. A. Dorsal view. B. Lateral view. C. Ventral view. Scale bar in C applicable to all figures." figureDoi="http://doi.org/10.5281/zenodo.5708154" httpUri="https://zenodo.org/record/5708154/files/figure.png" pageId="11" pageNumber="85">Fig. 7</figureCitation>
). Pars distalis wide, spoon-shaped. Follis multifolded, apices of dorsal bilobular projection acute, stylus arises from inside. Two pairs of MS D, lateral to base of follis, one pair very small and other formed by large acute MS. Five to six pairs of acute MS C forming curved rows on lateral sides. MS A+B formed by 1617 spatulate setae, latero-basally to pars distalis. MS E composed of two pairs, MS E1 just above level of E2.
</paragraph>
<paragraph id="DD2B36F60343C314BE858282FED530FD" blockId="11.[189,911,471,533]" box="[234,325,471,497]" pageId="11" pageNumber="85">
<emphasis id="EFE0EAE40343C314BE858282FED530FD" bold="true" box="[234,325,471,497]" pageId="11" pageNumber="85">Female</emphasis>
</paragraph>
<paragraph id="DD2B36F60343C314BED282AEFC1F3318" blockId="11.[189,911,471,533]" box="[189,911,506,533]" pageId="11" pageNumber="85">Very similar to male, without remarkable sexual dimorphism.</paragraph>
</subSubSection>
<subSubSection id="958E657D0343C314BED28169FDC93388" box="[189,601,572,645]" pageId="11" pageNumber="85" type="distribution">
<paragraph id="DD2B36F60343C314BED28169FEC6335A" blockId="11.[189,342,572,598]" box="[189,342,572,598]" pageId="11" pageNumber="85">
<heading id="8663819A0343C314BED28169FEC6335A" bold="true" box="[189,342,572,598]" fontSize="11" level="3" pageId="11" pageNumber="85" reason="3">
<emphasis id="EFE0EAE40343C314BED28169FEC6335A" bold="true" box="[189,342,572,598]" pageId="11" pageNumber="85">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F60343C314BED2813EFDC93388" blockId="11.[189,601,618,645]" box="[189,601,618,645]" pageId="11" pageNumber="85">
Known only from the
<typeStatus id="022F88540343C314BFAD813EFE653389" box="[450,501,619,645]" pageId="11" pageNumber="85">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>