Revision of the troglomorphic genus Troglostygnopsis Šilhavý, 1974 (Opiliones, Laniatores, Stygnopsidae) Author Cruz-López, Jesús A. 7D2ECD16-DDC3-4DE6-9C15-EF95C478E572 Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. 3 er. Circuito exterior s / n. Apartado Postal 70 - 153, C. P. 04510, Ciudad Universitaria, Coyoacán, Mexico City, Mexico. & Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Campo Experimental Valles Centrales, Oaxaca, Melchor Ocampo No. 7, C. P. 68200, Santo Domingo Barrio Bajo, Etla, Oaxaca, Mexico. thelyphonidito@gmail.com Author Francke, Oscar F. 8E9540FA-BF4F-4C9F-A3D8-D94768BE6C8D Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. 3 er. Circuito exterior s / n. Apartado Postal 70 - 153, C. P. 04510, Ciudad Universitaria, Coyoacán, Mexico City, Mexico. offb@ib.unam.mx text European Journal of Taxonomy 2021 2021-02-18 735 74 88 journal article 3469 10.5852/ejt.2021.735.1241 b496729a-abd4-4212-af76-b212cb9c2f88 2118-9773 5708138 327CCB1E-31AC-4121-926E-354AA324991C Troglostygnopsis kalebi sp. nov. urn:lsid:zoobank.org:act: 46F6E6D4-2D3C-4E8A-B942-3B191904DF8E Figs 4–7 Troglostygnopsis sp . CNANOp0050 Cruz-López & Francke 2017: 318 . Diagnosis Movable finger of chelicera with two teeth, low and wide ( Fig. 5B ). Tubercles of leg IV large and very close each other, patella with many of these tubercles ( Fig. 6B ). Penis with two pairs of MS D, MS E1 just above the level of E2. Etymology Patronymic honoring our colleague and friend Kaleb Zárate, an enthusiastic speleologist who has helped us collecting troglomorphic arachnids during several expeditions in southeastern Mexico . Material examined Holotype MEXICO • ♂ ; Chiapas , Las Rosas , Cueva del Ciprés ; 16º18′2″ N , 92º19′5″ W ; 17 Jun. 2011 ; O. Francke , J. Cruz-López , R. Monjaraz-Ruedas , G. Contreras and K. Zárate leg.; CNAN-T0850 . Paratypes MEXICO • 1 ♂ , 1 ♀ ; same collection data as for holotype; male is a DNA voucher CNANOp0050; CNAN-T0851 . Fig. 4. Troglostygnopsis kalebi sp. nov. , ♂, holotype (CNAN-T0850). A . Dorsal view. B . Lateral view. Description Male (based on the holotype ) MEASUREMENTS. Scutum length: 5.0, scutum width: 4.2. BODY ( Fig. 4 ). Scutum type zeta ‘ζ’, with both constrictions marked, specially C1, which marks the limits between prosoma and opisthosoma. Lateral margins of the scutum projected in lateral clear areas, Fig. 5. Troglostygnopsis kalebi sp. nov. , ♂, holotype (CNAN-T0850). A . Chelicera, mesal view. B . Chelicera, frontal view. C . Pedipalp, mesal view. D . Pedipalpal femur, mesal view. E . Pedipalpal tibia and tarsus, ventral view. Fig. 6. Troglostygnopsis kalebi sp. nov. , ♂, holotype (CNAN-T0850). A . Apical femur, patella and basal tibia IV. B . Patella IV. Fig. 7. Troglostygnopsis kalebi sp. nov. , ♂, holotype (CNAN-T0850), penis. A . Dorsal view. B . Lateral view. C . Ventral view. Scale bar in C applicable to all figures. large and rounded. Entire dorsum smooth, with few small tubercles in middle of each mesotergal area. Ocularium at the frontal margin of the prosoma, acute, without eyes. CHELICERA ( Fig. 5 A–B). Scutum/cheliceral hand ratio: 1.29. Basichelicerite large and smooth, with few small tubercles on ventral and apical regions. Cheliceral hand slightly widened but not swollen. Fixed finger with four teeth distributed evenly throughout, central triangular-shaped. Movable finger with two teeth, one basal and one subterminal, basalmost rounded. PEDIPALPS ( Fig. 5 C–E). Measurements: 2.31/1.10/1.80/1.80/1.10.Trochanter globose, with long spiniform setiferous tubercles on ventral and dorsal faces. Femur compressed laterally, armed with dorsal and ventral row of about 10 very large spiniform setiferous tubercles. Additionally, femur with mesal apical spiniform tubercle, near patella. Patella covered with spiniform tubercles on dorsal and mesal faces. Tibia with large main armature, relative sizes of SST on mesal side 1 = 3> 2, and on ectal margin 3 = 2 = 1. Also on tibia three very small tubercles between SST 1, SST 2 and SST 3 on mesal side, and four small ones between SST 1 and SST 2, one between SST2 and SST 3, and one apically. Tarsus with six major tubercles on mesal side (1> 2> 3> 4> 5> 6) and six on ectal side (1> 2> 3> 4> 5). LEGS ( Fig. 6 ). All segments long and slender, legs I–III without armature, ornate only with few disperse small tubercles. Ornamentation of femur IV formed by rounded and acuminate tubercles, which form longitudinal rows along entire femur, patella covered with many tubercles. Fig. 8. Distribution of selected troglobitic genera of Stygnopsidae Sørensen, 1932 . Squares are distribution of Troglostygnopsis Šilhavý, 1974 : black = T. anophthalma Šilhavý, 1974 ; red = T. kalebi sp. nov. Circles show distribution of Brujita Cruz-López, in press, Toojah Cruz-López, in press and the dubious record for Mictlana inops ( Goodnight & Goodnight, 1971 ) reported by Goodnight & Goodnight (1973) : black = B. chapulapa Cruz-López, in press; red = T. cimutaa Cruz-López , in press; blue = M. inops ? Pentagons are the distribution of Mictlana Cruz-López & Francke, 2015 : black = M. plana ( Goodnight & Goodnight, 1973 ) ; red = M. inops . PENIS ( Fig. 7 ). Pars distalis wide, spoon-shaped. Follis multifolded, apices of dorsal bilobular projection acute, stylus arises from inside. Two pairs of MS D, lateral to base of follis, one pair very small and other formed by large acute MS. Five to six pairs of acute MS C forming curved rows on lateral sides. MS A+B formed by 16–17 spatulate setae, latero-basally to pars distalis. MS E composed of two pairs, MS E1 just above level of E2. Female Very similar to male, without remarkable sexual dimorphism. Distribution Known only from the type locality.