treatments-xml/data/54/69/8C/54698C42DFF65A60820498366B6ACE5A.xml

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<mods:title id="0C79CED72CF7BBA7F0720E5DC6C01051">Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="B2FC28F7A0D77C5E9059D867C982D487">Linsky, Marek</mods:namePart>
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<mods:affiliation id="616F18670D6958E66BA6337B202E7AF6">Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<mods:namePart id="A4BFA4AEABD6A477D7CEF72FF87154A7">Ciamporova-Zatovicova, Zuzana</mods:namePart>
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<mods:namePart id="3BB4FD67FA2E225F7D841D2634893551">Lassova, Kristina</mods:namePart>
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<mods:affiliation id="8A3460D1357774856B1DDB91D569B5DC">Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<paragraph id="266ACBD7E1311BF579F825369B5B1D67" pageId="0" pageNumber="575">
3.2.5. 
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<taxonomicNameLabel id="643150FE24FEAC0797F282EFB79A6364" pageId="0" pageNumber="575">sp. nov.</taxonomicNameLabel>
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<paragraph id="E497542423BEFF1D4A2D646D398A5299" pageId="0" pageNumber="575">
<figureCitation id="F6AA92C9C35D4360F1665DED23F47C74" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">Figs 2c</figureCitation>
<figureCitation id="6F811703E8F9301768870B77C81CA17E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">, 3c</figureCitation>
<figureCitation id="D7E2503E16B1A97852B32C1DB0ACDA60" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">, 5c</figureCitation>
<figureCitation id="80FFB3738AF73941854AAA3DF7E80789" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">, 6c</figureCitation>
<figureCitation id="41D35C604453BD1AAD26269C38D58EAA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">, 7c</figureCitation>
<figureCitation id="FC2CF6C27B3CAD0534CE0DBA1FDF4ED6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">, 8c</figureCitation>
<figureCitation id="DDFAE42BA072D4568673B91025B8333D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">, 9c</figureCitation>
<figureCitation id="07A02AB9B194E559F2F04CD967BFDDC0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Rumilara obscura sp. nov. (a - d): a prosternum and mesosternum; b foreleg (without coxa); c midleg; d hindleg. Schematic illustrations: e midleg of Hexanchorus cordillierae (alp - anterior lateral pubescence; mst - mesotibial tubercle; plp - posterior lateral pubescence; vs - ventral seta); f aedeagus of Rumilara riberai sp. nov. (ap - apophyse; co - corona; df - dorsal fibula; en - endophallus; mi - fibula minor; mj - fibula major; ml - median lobe; pa - paramere; ph - phallobase). Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure10" httpUri="https://binary.pensoft.net/fig/770433" pageId="0" pageNumber="575">, 10f</figureCitation>
<figureCitation id="EB30236F72E26425A669D106E294FB05" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">, 13</figureCitation>
</paragraph>
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<paragraph id="BEED4208C679211CE363F7F0F02547BD" pageId="0" pageNumber="575">Material examined.</paragraph>
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(PUCE) &quot; 
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, 
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Santo Domingo de los 
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wide, with sand, gravel, boulders, 
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Jr &amp; 
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lgt.&quot;
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with the same data as the holotype
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, 
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<quantity id="A35F3FB5D714689E79073A4E494F3C45" metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">914m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="7D253155137F7F4516B61BC6EA55621E" value="2013-08-11">11.8.2013</collectingDate>
, very shallow stream running along a rock covered by periphyton, 
<normalizedToken id="1BA3C1F0B55EF9E47761FA9B94EBB304" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="D44FA62284B72C10BCB1A7CE15B5BCFF" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
<materialsCitation id="7A92A062C70E8B10C02E084E61C86C47" ID-GBIF-Occurrence="3968369302" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Ecuador" elevation="917" latitude="-0.33325" location="Rio Toachi" longLatPrecision="1" longitude="-78.94072" municipality="Otongachi" specimenCount="13" specimenCount-larva="13" stateProvince="Santo Domingo de los Tsachilas" typeStatus="Other material">
<emphasis id="E1C8453B1DF4103019AC8B8CC1284C26" bold="true" italics="true" pageId="0" pageNumber="575">
<typeStatus id="BA77F3BFE3D897C81587AD5D5ACA07A7" isOtherTypeStatus="true">Other material</typeStatus>
</emphasis>
: 
<specimenCount id="347CE5DBDF51802CF420ABC1D43C5182" type="larva">13 larvae</specimenCount>
with the same data as the holotype
</materialsCitation>
; 
<materialsCitation id="AE992713C0A20FBA4C92A679E3D4252D" ID-GBIF-Occurrence="3968369301" collectingDate="2013-08-11" country="Ecuador" elevation="914" latitude="-0.33069444" location="Rio Toachi" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" specimenCount="6" specimenCount-larva="6" stateProvince="Santo Domingo de los Tsachilas prov.">
<specimenCount id="17C8A32D65FEBEB6DF29DF5F54E0021E" type="larva">6 larvae</specimenCount>
&quot; 
<collectingCountry id="C4F8EF54BE948254970E4245D364C7DF" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="763FC797886D3CC710B8205357A582FE" country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken id="37D57BADDF718F75E2925419A1BDAF63" originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality id="E758706E794220627FA5CC3610F3EE35">Otongachi</collectingMunicipality>
env., next to the tributary of 
<location id="2BDF7D21F6C307E96522C2BA40E3BCD0" LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:2BDF7D21F6C307E96522C2BA40E3BCD0" country="Ecuador" latitude="-0.33069444" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken id="FA0B5D6F94A08249EB457DAA2415974D" originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate id="880D0B853CF5BE344755C3400F61CC26" degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="50.5" value="-0.33069444">00°19′50.5″ S</geoCoordinate>
, 
<geoCoordinate id="57587E171F3ADA5FFD522BE26070A660" degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="41.6" value="-78.94489">78°56′41.6″ W</geoCoordinate>
, 
<elevation id="972FE47928A879F44BC63C221F10884B" metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">
<quantity id="DF2A5EFF6DEE0D12E3EB2352237B83B4" metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">914m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="54171C8DB94DC002252425E51398C76F" value="2013-08-11">11.8.2013</collectingDate>
, very shallow stream running along a rock covered by periphyton, 
<normalizedToken id="C8E43D156FBE470F28D1DEDC0084F785" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="C67A98FF2E56E0639ABDA19123FBB45A" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
; 
<materialsCitation id="73E5BD070357BFA0E0BBF03A91642D08" ID-GBIF-Occurrence="3968369313" collectingDate="2013-08-11" country="Ecuador" elevation="852" latitude="-0.3261111" location="Rio Toachi" longLatPrecision="1" longitude="-78.94975" municipality="Otongachi" specimenCount="1" specimenCount-larva="1" stateProvince="Santo Domingo de los Tsachilas prov.">
<specimenCount id="F8833A5B7E3E137D61C83D16C21EE928" type="larva">1 larva</specimenCount>
&quot; 
<collectingCountry id="9ED6AB46A9236FDF7D6B1DEDCE5A428F" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="DADA11FD680079ECAD2BB60D21FDDFDF" country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken id="C10EE9CFB7E9D2301569C77662887166" originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality id="B29925C6C0FC51B5854A47461CC173E9">Otongachi</collectingMunicipality>
env., tributary of 
<location id="24772860EA6DDA9D1A35DC5B6A6462DC" LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:24772860EA6DDA9D1A35DC5B6A6462DC" country="Ecuador" latitude="-0.3261111" longLatPrecision="1" longitude="-78.94975" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken id="7F59AB90149DCCE2743521036CF0CCE3" originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate id="29871563ED944118B35E139457AFC382" degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="34.0" value="-0.3261111">00°19′34.0″ S</geoCoordinate>
, 
<geoCoordinate id="50ADF0CB933E5B3F455E52607CFD8153" degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="59.1" value="-78.94975">78°56′59.1″ W</geoCoordinate>
, 
<elevation id="E386340F181D452534542C829D7EEF40" metricMagnitude="2" metricUnit="m" metricValue="8.52" unit="m" value="852.0">
<quantity id="E2BFEE9EAACA827C798BC72326A346D6" metricMagnitude="2" metricUnit="m" metricValue="8.52" unit="m" value="852.0">852m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="4D0143D1E18CC1F2A3897EB5CE20BA31" value="2013-08-11">11.8.2013</collectingDate>
, wild river ca 
<quantity id="18E2437282BAE12103862A031FB5AF8F" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation id="2CDD872CE078CFC30FDDB84A0B73A14E" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20m</elevation>
</quantity>
wide, fast flowing, with large boulders, 
<normalizedToken id="5DD9F99DAE9A944C9AEF09717C9192AB" originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken id="64ABD8DA56EE161236D4643195B1F65F" originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
.
</paragraph>
<caption id="5933B808DF9236161A0F8838C5F97DCD" doi="10.3897/asp.80.e84013.figure15" httpUri="https://binary.pensoft.net/fig/770438" pageId="0" pageNumber="575" start="Figure 15" startId="F15">
<paragraph id="795297ECB7EF1800849710793BD57191" pageId="0" pageNumber="575">
<emphasis id="A87DC7352B0883ACEFCC75A67184850B" bold="true" pageId="0" pageNumber="575">Figure 15.</emphasis>
<taxonomicName id="6379519926648846A18A8CD290DD69E6" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Deleve" baseAuthorityYear="1968" class="Insecta" family="Elmidae" genus="Rumilara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rumilara leleupi" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="843C3FA807207AFA3F2D2940513E1F64" italics="true" pageId="0" pageNumber="575">Rumilara leleupi</emphasis>
</taxonomicName>
<emphasis id="53164824E0109A6FE5036B2B820BCC7A" bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
: 
<emphasis id="E453180E609E09AF37F8E75C59F558D5" bold="true" pageId="0" pageNumber="575">a</emphasis>
aedeagus, left lateral view; 
<emphasis id="3B0D0215BC5022752FB772BA945D7146" bold="true" pageId="0" pageNumber="575">b</emphasis>
aedeagus, ventral view; 
<emphasis id="13E82781A222CB6CACCD87F1798160FE" bold="true" pageId="0" pageNumber="575">c</emphasis>
aedeagus, right lateral view; 
<emphasis id="984AE1F7E052426892D451ECC4BB1D69" bold="true" pageId="0" pageNumber="575">d</emphasis>
male sternite VIII. Scale: 
<quantity id="17B3F33547BF55476F1F1BA8A40956EA" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="36851823CFED7DE08E0463495CE22073" pageId="0" pageNumber="575" type="adult diagnosis">
<paragraph id="65756C2385AAACEF3D20B279AD3654E4" pageId="0" pageNumber="575">Adult diagnosis.</paragraph>
<paragraph id="C72F3F20DCDDC4F98406EA1705328510" pageId="0" pageNumber="575">The species can be distinguished by a combination of following characters: 1) head only partially clothed by pubescence, clypeus bare; 2) femora bare or with a row of dark, long hairs on dorsal margin; 3) elytra without dense pubescence; 4) aedeagus with rod-like sclerite; 5) parameres only feebly asymmetrical.</paragraph>
</subSubSection>
<subSubSection id="29AFCFAC8E6E006D4A69CB2F16E2B47E" pageId="0" pageNumber="575" type="description">
<paragraph id="17B70AD70B236E601181957D44948D49" pageId="0" pageNumber="575">Adult diagnostic description.</paragraph>
<paragraph id="0735020BBDA236E09AC627E9EFFA9257" pageId="0" pageNumber="575">
Male. 
<emphasis id="B5F65A1FADEB044050E2D1B3662417B6" bold="true" italics="true" pageId="0" pageNumber="575">Colour</emphasis>
: Body (Figs 
<figureCitation id="D1F77377D70CDC8E4E0822D1EF3BF368" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2c</figureCitation>
, 
<figureCitation id="8885F43FFF1AA7276F160A5608C6820B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3c</figureCitation>
) black; lateral sides of femora and tibiae brown; tarsal claws pale brown with reddish tincture. 
<emphasis id="CA195D0F1DD3CDD8712C54BF7704E9FC" bold="true" italics="true" pageId="0" pageNumber="575">Pubescence</emphasis>
: Head behind eyes clothed with short, prone pubescence. Elytra very readily covered by short, recumbent pubescence, and with numerous moderately long, curved, semi-erect, hair-like setae. Femora with long, dark, densely set, hair-like setae near dorsal margin; pro- and metatibiae apically with a fringe of long, dark, hair-like setae on inner margin; mesotibiae without pubescence. Metaventrite readily clothed with short, prone pubescence. 
<emphasis id="422796C0EBE5F294AFF9C74ED167BC7A" bold="true" italics="true" pageId="0" pageNumber="575">Surface</emphasis>
: Head, clypeus and pronotum densely punctate. Elytra with rows of punctures moderately deeply impressed; third interval slightly elevated in anterior 1/5; remaining intervals nearly flat. Prosternal process (Fig. 
<figureCitation id="6A5B76E0FD2B23F979DC276A0A494BF0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7c</figureCitation>
) moderately wide, concave between procoxae, then subtriangular with rounded apex. 
<emphasis id="436F783DDD595BC48BB21FDDDF8AAD8F" bold="true" italics="true" pageId="0" pageNumber="575">Aedeagus</emphasis>
(Figs 
<figureCitation id="9C27C95CFAD0E854BB9DC2FDF959F614" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">13a-c</figureCitation>
) with long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one slightly longer. - Female externally similar to male, except slightly larger.
</paragraph>
</subSubSection>
<subSubSection id="AC519D82819338EE5DA300DA1CCDF435" pageId="0" pageNumber="575" type="description">
<paragraph id="B82C96AEA5524907AB7F4BC9587416E3" pageId="0" pageNumber="575">Measurements.</paragraph>
<paragraph id="A776C12F9684D76226C3E48BE3F3758F" pageId="0" pageNumber="575">♂ - CL: 2.16-2.53 mm; PL: 0.56-0.65 mm; PW: 0.71-0.79 mm; EL: 1.60-1.88 mm; EW: 0.83-0.97 mm. ♀ - CL: 2.68-2.69 mm; PL: 0.68-0.70 mm; PW: 0.82-0.85 mm; EL: 1.99-2.00 mm; EW: 1.04-1.06 mm.</paragraph>
</subSubSection>
<subSubSection id="C15AF7509FC074A969BFC3FACEDB8668" pageId="0" pageNumber="575" type="variation">
<paragraph id="C34A038BE7B3028BA07888F05D691893" pageId="0" pageNumber="575">Variation.</paragraph>
<paragraph id="9AA156BEC527655BB71B5BD462462948" pageId="0" pageNumber="575">The greatest differences were observed in distribution of pubescence (mainly on legs, around eyes, partly on pronotum and elytra) due to its often removal.</paragraph>
</subSubSection>
<subSubSection id="4431E72915544CEC89AA88BD8F2CD189" pageId="0" pageNumber="575" type="etymology">
<paragraph id="D033C76B12041D974FF3EF7D578EB29B" pageId="0" pageNumber="575">Etymology.</paragraph>
<paragraph id="2C3D06D79A2F6E9A181B4F09898F378D" pageId="0" pageNumber="575">
Named after Ignacio Ribera 
<normalizedToken id="C5929AC2BE16E1FB111BC65AC76327C4" originalValue="Galán">Galan</normalizedToken>
a great expert on systematics, phylogeny, evolution, biogeography, and conservation of water beetles, who untimely passed away.
</paragraph>
</subSubSection>
<subSubSection id="201E76C2BD1747486147E00632F2BA5C" pageId="0" pageNumber="575" type="larva">
<paragraph id="DDE502B49D1E40B4CF26A881C0CE1B88" pageId="0" pageNumber="575">Larva</paragraph>
<paragraph id="A05963906CC8F5D77E71710B8A2BA32C" pageId="0" pageNumber="575">
(Fig. 
<figureCitation id="16AE18265DA4B851B9A649C01EE212EE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">9c</figureCitation>
) with length 2.92-2.96 mm, greatest width 0.96-0.99 mm. Pleurites on abdominal segments 1-5. Abdominal segments 1-6 explanate. Surface around the frontal pair of pronotal gibbosities of the same colour as the rest of pronotum. Tergum of abdominal segments 1-7 with a pair of distinct, longitudinally arranged small gibbosities.
</paragraph>
</subSubSection>
<subSubSection id="C749B7C6D6245446AC861BFFC550A851" pageId="0" pageNumber="575" type="distribution">
<paragraph id="26C8E5B4288BCA003C1FDBCA8ABF6F44" pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph id="DBB866AC5271484B58788A1C2B9BACF9" pageId="0" pageNumber="575">
Known from three localities in Santo Domingo de los 
<normalizedToken id="73B617F55D79BD3588C39CC4413197A5" originalValue="Tsáchilas">Tsachilas</normalizedToken>
Province (Figs 
<figureCitation id="8A9E1BC601A35EC0F4125FAEE1D15B60" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling sites of Rumilara gen. nov. in Ecuador: a Cotopaxi prov., Otonga env., Rio Esmeralda; b Cotopaxi prov., Artos env., stream above confluence with Rio Esmeralda; c Santo Domingo de los Tsachilas prov., Otongachi env., tributary of Rio Toachi; d Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.; e Morona-Santiago prov., small stream near Indanza; f Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la Encanada; g fast-flowing stream in Napo prov.; h Morona-Santiago prov., tributary of Rio Tinajillas; i Morona-Santiago prov., tributary of Rio Tinajillas; (localities of R. obscura - a, b; R. riberai - c, d; R. paterna - e, f; R. suppressa - g - i)." figureDoi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575">16c-d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9141F06F3849536D4CE425F3D8F0A5C4" pageId="0" pageNumber="575" type="remarks">
<paragraph id="4B5E1B486F91C3895985FA4AA0DE177A" pageId="0" pageNumber="575">Remarks.</paragraph>
<paragraph id="D62E6C17055C3B2EFC9775A5794CB592" pageId="0" pageNumber="575">Several paratypes are markedly smaller than the rest (including holotype), however, they agree in all diagnostic characters.</paragraph>
</subSubSection>
</treatment>
</document>