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<document ID-CLB-Dataset="8072" ID-DOI="http://dx.doi.org/10.3897/zookeys.1019.59881" ID-GBIF-Dataset="c81147b7-467a-4477-a36e-76f3fd8358c7" ID-PMC="PMC7921074" ID-Pensoft-Pub="1313-2970-1019-93" ID-Pensoft-UUID="68E025484FB95656A3666768A4D97EF1" ID-PubMed="33688849" ID-ZooBank="740EFFB93ADA4B2ABD23A839AAE71FB2" ModsDocID="1313-2970-1019-93" checkinTime="1614092297092" checkinUser="pensoft" docAuthor="Giron, Jennifer C. &amp; Short, Andrew Edward Z." docDate="2021" docId="5429D85BD30B57C3BB2AE8D7FC4BE1AF" docLanguage="en" docName="ZooKeys 1019: 93-140" docOrigin="ZooKeys 1019" docPubDate="2021-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1019.59881" docTitle="Tobochares anthonyae Girón &amp; Short 2021, sp. nov." docType="treatment" docUuid="DBADDDCB-34F7-4608-810E-D488FBBD3ABE" docUuidSource="ZooBank" docVersion="7" id="68E025484FB95656A3666768A4D97EF1" lastPageNumber="93" masterDocId="68E025484FB95656A3666768A4D97EF1" masterDocTitle="Review of the Neotropical water scavenger beetle genus Tobochares Short &amp; Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records" masterLastPageNumber="140" masterPageNumber="93" pageNumber="93" updateTime="1732683750272" updateUser="ExternalLinkService">
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<mods:title id="C1E2C8AC3D04E815DC47BB7FDEB99226">Review of the Neotropical water scavenger beetle genus Tobochares Short &amp; Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records</mods:title>
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<mods:namePart id="C104928CEFF796B25AD6D460F782656A">Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier id="BE9951EBCD7D73E4C0B7039BD8DB83BB" type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation id="0FF613D2F9BF35954F011486F46B1B33">Department of Entomology, Purdue University, West Lafayette, IN 47907, USA &amp; Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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<mods:namePart id="0BA626C97E8EAFEB7856630160749C1B">Short, Andrew Edward Z.</mods:namePart>
<mods:affiliation id="50EEC4E295710FD12DAF83C7C9D64C30">Department of Ecology and Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
<mods:nameIdentifier id="9F944363D1E4EBB70F69F4FC143ACD8F" type="email">aezshort@ku.edu</mods:nameIdentifier>
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<mods:typeOfResource id="3485BAFBBF0377BA04574504EE739A4A">text</mods:typeOfResource>
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<mods:date id="74AE9B1405E73143B2A934912C853335">2021</mods:date>
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<mods:number id="3EF9D163B51A0C9B0CFCE6B30B7560B2">2021-02-22</mods:number>
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<mods:number id="D016418420CF5B72F380850451244745">1019</mods:number>
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<mods:classification id="616D4E04CF789493012F0B6816355D94">journal article</mods:classification>
<mods:identifier id="B4D5C4F6915293952BF3D76F933BCD42" type="DOI">http://dx.doi.org/10.3897/zookeys.1019.59881</mods:identifier>
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<treatment id="5429D85BD30B57C3BB2AE8D7FC4BE1AF" ID-GBIF-Taxon="179905048" LSID="urn:lsid:zoobank.org:act:DBADDDCB-34F7-4608-810E-D488FBBD3ABE" httpUri="http://treatment.plazi.org/id/5429D85BD30B57C3BB2AE8D7FC4BE1AF" lastPageNumber="93" pageId="0" pageNumber="93" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_subFamily="Acidocerinae">
<subSubSection id="9F3B8C0B03E155CDCB723C1A3F22BBD2" pageId="0" pageNumber="93" type="nomenclature">
<paragraph id="8179CE81195D7340D8A063C479E758D9" pageId="0" pageNumber="93">
<taxonomicName id="0303D422E3317733FFA6324433877C03" ID-CoL="8YBWN" LSID="http://zoobank.org/DBADDDCB-34F7-4608-810E-D488FBBD3ABE" authority="Girón &amp; Short, 2021" authorityName="Girón &amp; Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares anthonyae" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="anthonyae" status="sp. nov.">Tobochares anthonyae</taxonomicName>
<taxonomicNameLabel id="A4649077B99F5A283ED5C4276DBBFE3C" pageId="0" pageNumber="93">sp. nov.</taxonomicNameLabel>
<figureCitation id="415F7E48BA4033439DC9A34975304591" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">Figs 7A-C</figureCitation>
<figureCitation id="8C7AE38A7D21870E7E6EBA24AF0DCB78" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">, 11F</figureCitation>
<figureCitation id="40ECB8747CA980635385B841D40F8839" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Tobochares spp." figureDoi="10.3897/zookeys.1019.59881.figure13" httpUri="https://binary.pensoft.net/fig/512611" pageId="0" pageNumber="93">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8AEAF187D30084AA8F02424BDE00EF19" pageId="0" pageNumber="93" type="type material">
<paragraph id="624DCDC579272FD0913951DCF512AC85" pageId="0" pageNumber="93">Type material examined.</paragraph>
<paragraph id="A26E8DA521437ED0F311F679AD5D89AB" pageId="0" pageNumber="93">
<emphasis id="F454EACD57BDFC193818FF0310BBDD02" bold="true" pageId="0" pageNumber="93">
<emphasis id="6983D8F0A21B0B8C125C070170929D8A" italics="true" pageId="0" pageNumber="93">Holotype</emphasis>
(male)
</emphasis>
: &quot;Venezuela:
<normalizedToken id="348B9DCE35761AC0D4DD0C081C202839" originalValue="Bolívar">Bolivar</normalizedToken>
:
<geoCoordinate id="D27EB68C9E1C73E9EDAD5FD992C3F98F" degrees="6" direction="north" minutes="13" orientation="latitude" precision="1" seconds="4.6" value="6.2179446">6°13'4.6&quot;N</geoCoordinate>
,
<geoCoordinate id="03F44387B500E07F71F6156D5B3D4185" degrees="67" direction="west" minutes="14" orientation="longitude" precision="1" seconds="26.4" value="-67.24066">67°14'26.4&quot;W</geoCoordinate>
; 60 m; ca. 25 Km E of El Burro; rocky morichal; 12.i.2009; leg. Short et al.; VZ09-0113-01X&quot; (MIZA).
<emphasis id="99C6A2D9BC412D983224CF5018B5698C" bold="true" pageId="0" pageNumber="93">
<emphasis id="EDA2649F5D09E9E8B56DF359C315C4B0" italics="true" pageId="0" pageNumber="93">Paratypes</emphasis>
(3 exs.): Venezuela:
<normalizedToken id="A786DE0E29801FA1FFF3CEC59D9F1ED4" originalValue="Bolívar">Bolivar</normalizedToken>
</emphasis>
: &quot;
<geoCoordinate id="0A91ED4C58FF61CD9F166FF14120FE29" degrees="6" direction="north" minutes="13" orientation="latitude" precision="1" seconds="4.6" value="6.2179446">6°13'4.6&quot;N</geoCoordinate>
,
<geoCoordinate id="B0419B90E7949D39D35D4B7AAECD9D5B" degrees="67" direction="west" minutes="14" orientation="longitude" precision="1" seconds="26.4" value="-67.24066">67°14'26.4&quot;W</geoCoordinate>
; 60 m; ca. 25 Km E of El Burro; rocky morichal; 7.viii.2008; leg. Short,
<normalizedToken id="2054D6A4D7996B32BF8FCD6552B73449" originalValue="García">Garcia</normalizedToken>
, Joly; AS-08-077&quot; (1, SEMC); same data as holotype (2, SEMC).
</paragraph>
</subSubSection>
<subSubSection id="9314B1093B6C66F745D3572591BE3340" pageId="0" pageNumber="93" type="differential diagnosis">
<paragraph id="7B6C5AC46A7E7F5C68E2215A777A989D" pageId="0" pageNumber="93">Differential diagnosis.</paragraph>
<paragraph id="EEC6A4CDB11194AE673ED5539B1A56FE" pageId="0" pageNumber="93">
The general habitus and coloration of
<taxonomicName id="45E3B62929096C35A86AF65D78E84D71" lsidName="T. anthonyae" pageId="0" pageNumber="93" rank="species" species="anthonyae">
<emphasis id="347A57225E6BD77C03CA8961CF99253B" italics="true" pageId="0" pageNumber="93">T. anthonyae</emphasis>
</taxonomicName>
is similar to that of several species in the
<taxonomicName id="89E78FF4CCD01E7B631D5709761F815E" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares communis" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="communis">
<emphasis id="B44DC57AF5DB2BA9E651C9B473B8B85D" italics="true" pageId="0" pageNumber="93">Tobochares communis</emphasis>
</taxonomicName>
group; nevertheless, the elytral punctation
<taxonomicName id="084DB87F4C84778FE9CA3A6E2583A700" lsidName="T. anthonyae" pageId="0" pageNumber="93" rank="species" species="anthonyae">
<emphasis id="D34DB76B34CC6F3DB461E2548809F9DB" italics="true" pageId="0" pageNumber="93">T. anthonyae</emphasis>
</taxonomicName>
is relatively distinct: all kinds of punctures are relatively large, similar in size and degree of impression, the serial punctures are aligned in rows and slightly impressed, forming shallow longitudinal grooves, and the interserial punctures are somewhat irregularly distributed in a single row (Fig.
<figureCitation id="711207B160DCF6DE0794D2451A15A888" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7A</figureCitation>
). The relatively large punctures, similar in size and degree of impression may resemble those of
<taxonomicName id="296AD13560DCAC3E3491ACCB2109CE3D" lsidName="T. communis" pageId="0" pageNumber="93" rank="species" species="communis">
<emphasis id="91E0A47EAAEB1C39DCA03521030ADA77" italics="true" pageId="0" pageNumber="93">T. communis</emphasis>
</taxonomicName>
, but in this species the serial punctures are not impressed to form grooves (Fig.
<figureCitation id="2CE1851DFA320F6AD99C7BF8198DFF53" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. communis A dorsal view B lateral view C ventral view D-F T. microps D dorsal view E lateral view F ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure6" httpUri="https://binary.pensoft.net/fig/512604" pageId="0" pageNumber="93">6A</figureCitation>
), as they are in
<taxonomicName id="099AD437981C47FFE77B1BC7ECC56500" lsidName="T. anthonyae" pageId="0" pageNumber="93" rank="species" species="anthonyae">
<emphasis id="B9D44D4CE1AB487D75C5996339305040" italics="true" pageId="0" pageNumber="93">T. anthonyae</emphasis>
</taxonomicName>
. In addition, the overall shape of the aedeagus, especially the shape of the median lobe of
<taxonomicName id="791D3556B85A6071DAC11E46F73B3CDE" lsidName="T. anthonyae" pageId="0" pageNumber="93" rank="species" species="anthonyae">
<emphasis id="8864AC3BDC13EF5BCF3D675DC6289836" italics="true" pageId="0" pageNumber="93">T. anthonyae</emphasis>
</taxonomicName>
is unique among members of the
<taxonomicName id="2FBAE1D853A8FB106F4C3AD84AE80518" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares communis" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="communis">
<emphasis id="FDC1451A98AED4FF30BA1895000A399B" italics="true" pageId="0" pageNumber="93">Tobochares communis</emphasis>
</taxonomicName>
group: the median lobe gradually and slightly narrows towards a broadly rounded apex, and the gonopore is located near the apex of the median lobe (Fig.
<figureCitation id="7C14DFC9559FBA8F3020E62405764516" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">11F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B419E1484ADF00EDD834A8995FD98DD3" pageId="0" pageNumber="93" type="description">
<paragraph id="B5473630422069B56A7F38F7ABA9D0A5" pageId="0" pageNumber="93">Description.</paragraph>
<paragraph id="7F282B22329BEADDB44143569D7AB629" pageId="0" pageNumber="93">
<emphasis id="E226E301103F8791642CFD8D55E5D5FA" bold="true" italics="true" pageId="0" pageNumber="93">Size and form</emphasis>
: Body length 1.8-2.0 mm. Body elongate oval, moderately convex (Fig.
<figureCitation id="3D17489E9382D07771B3729DB64B2D3C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7B</figureCitation>
).
<emphasis id="7DF055CFB1A37B4E15F7D087BF40FD47" bold="true" italics="true" pageId="0" pageNumber="93">Color and punctation</emphasis>
: Dorsal and ventral surfaces of body, dark brown, with slightly paler margins of pronotum (Fig.
<figureCitation id="145FE9882C12436DF3FE74F7F57BC168" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7A, B</figureCitation>
); mouthparts yellowish brown; antennae light brown; legs orange with yellow tarsi (Fig.
<figureCitation id="096D11DF931FC7F2E2C6FB87A0F08AB1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7C</figureCitation>
). Ground punctation on head, pronotum and elytra rather shallowly marked.
<emphasis id="B41555A85F9A627ED0229F2A43D18B81" bold="true" italics="true" pageId="0" pageNumber="93">Head</emphasis>
: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e.g., Fig.
<figureCitation id="4A5212D4662DF1DC4989BB018E9B43DF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2D</figureCitation>
), and outer margins slightly bulging from outline of head; in lateral view, eyes not anteriorly emarginate (e.g., Fig.
<figureCitation id="7EFAB7EC13EFD9B6D157A1989E742E32" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2E</figureCitation>
).
<emphasis id="85F04A0142318225FEAA8458874B0797" bold="true" italics="true" pageId="0" pageNumber="93">Thorax</emphasis>
: Elytra with all kinds of punctures similar in size and degree of impression (Fig.
<figureCitation id="23D1E1A6773F9366C1B428F54DF35A93" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7A</figureCitation>
); serial punctures aligned in rows, slightly impressed, forming shallow longitudinal grooves; interserial punctures somewhat irregularly distributed in a single row (Fig.
<figureCitation id="6748E2B4364E1368A2C6BD30A0EE89C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7A</figureCitation>
).
<taxonomicName id="701E813595D7B72FFB1000F8C0FE3DC8" genus="Metafemora" lsidName="Metafemora" pageId="0" pageNumber="93" rank="genus">Metafemora</taxonomicName>
mostly glabrous on anterior face, with hydrofuge pubescence along basal third of antero-dorsal margin (Fig.
<figureCitation id="D95EA3DA5F0A9D8EE6EF86D070217A8A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7C</figureCitation>
). Elevation of mesoventrite forming a low longitudinal bulge (Fig.
<figureCitation id="C25356B3642E20A71F76A2DAAE74D23B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7C</figureCitation>
). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig.
<figureCitation id="B25D1378651B8DE29AEF77B66778B40F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of Tobochares spp. in the Tobochares communis species group A-C T. anthonyae A dorsal view B lateral view C ventral view D-F T. atures D dorsal view E lateral view F ventral view G-I T. pemon G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1019.59881.figure7" httpUri="https://binary.pensoft.net/fig/512605" pageId="0" pageNumber="93">7C</figureCitation>
).
<emphasis id="BD92ECC28A20BDC0D73CC64DCEEFEB73" bold="true" italics="true" pageId="0" pageNumber="93">Abdomen</emphasis>
: Abdominal ventrites uniformly and very densely pubescent. Aedeagus (Fig.
<figureCitation id="6D22407EAA3BFDEAA901D60862E7BE4D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Tobochares spp. A T. benettii B T. goias C T. fusus D T. luteomargo E T. pemon F T. anthonyae G T. atures H T. communis I T. microps J T. romanoae K T. akoerio L T. kappel M T. arawak N T. canaima. Scale bars: 0.5 mm (A-D); 0.1 mm (E-N)." figureDoi="10.3897/zookeys.1019.59881.figure11" httpUri="https://binary.pensoft.net/fig/512609" pageId="0" pageNumber="93">11F</figureCitation>
). Basal piece 0.3
<normalizedToken id="131A94AEC5EBC3C2905C2061CF87881F" originalValue="×">x</normalizedToken>
the length of a paramere; greatest width of a paramere nearly 0.5
<normalizedToken id="5D241B6E35574356A689B41742EE8F88" originalValue="×">x</normalizedToken>
greatest width of median lobe; outer margins of parameres nearly straight, only slightly curved inwards along apical region; apex of paramere rounded; median lobe roughly triangular, widely rounded at apex; gonopore situated at apical fourth of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="6F664EA31D1FAEFE336C43F3618A876C" pageId="0" pageNumber="93" type="etymology">
<paragraph id="4D9378CFA0847469E800E767010E3298" pageId="0" pageNumber="93">Etymology.</paragraph>
<paragraph id="D32CF25028605D60EA413728C5A23056" pageId="0" pageNumber="93">Named after Becky Anthony, program and meetings manager at the Entomological Society of America (ESA), in recognition of all her hard work in service to the society and the entomological community.</paragraph>
</subSubSection>
<subSubSection id="1D63B15CEE08C46715767CBA1EC0EE82" pageId="0" pageNumber="93" type="distribution">
<paragraph id="C1C7730E9EFD85E0E1BC31B68458121D" pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph id="BC7A6C2681A3EFB70380336B1BF27A11" pageId="0" pageNumber="93">
Only known from a single locality just south of the Orinoco River along the northwestern edge of the Guiana Shield. See Fig.
<figureCitation id="34FF70B1C5B6FF8D6C1CE5F18BFC547C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Tobochares spp." figureDoi="10.3897/zookeys.1019.59881.figure13" httpUri="https://binary.pensoft.net/fig/512611" pageId="0" pageNumber="93">13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3204CCD4813A0EF0AF90943662D47E45" pageId="0" pageNumber="93" type="life history">
<paragraph id="45BB415B4E2E9FA39893D6A2DE5DDAEA" pageId="0" pageNumber="93">Life history.</paragraph>
<paragraph id="159D96D5186993F4E1748E54F02050E3" pageId="0" pageNumber="93">The specimens were collected along a stream that was flowing over exposed granite.</paragraph>
</subSubSection>
</treatment>
</document>