211 lines
29 KiB
XML
211 lines
29 KiB
XML
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<mods:title id="55551BBFDD19D28F9E8B4E40721BBEBA">Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida)</mods:title>
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<mods:date id="343003D2F4521B926BEC047660129794">2021</mods:date>
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<treatment id="5038879DCE28550290A12B40FDD2FCE6" ID-DOI="http://doi.org/10.5281/zenodo.5714978" ID-GBIF-Taxon="190310266" ID-Zenodo-Dep="5714978" LSID="urn:lsid:plazi:treatment:5038879DCE28550290A12B40FDD2FCE6" httpUri="http://treatment.plazi.org/id/5038879DCE28550290A12B40FDD2FCE6" lastPageId="4" lastPageNumber="576" pageId="3" pageNumber="575">
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<subSubSection id="908B6500CE28550590A12B40FCA9F8B6" box="[151,796,1859,1886]" pageId="3" pageNumber="575" type="nomenclature">
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<paragraph id="D82E368BCE28550590A12B40FCA9F8B6" blockId="3.[151,796,1859,1921]" box="[151,796,1859,1886]" pageId="3" pageNumber="575">
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<heading id="836681E7CE28550590A12B40FCA9F8B6" bold="true" box="[151,796,1859,1886]" fontSize="11" level="1" pageId="3" pageNumber="575" reason="1">
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<emphasis id="EAE5EA99CE28550590A12B40FCA9F8B6" bold="true" box="[151,796,1859,1886]" pageId="3" pageNumber="575">
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<taxonomicName id="1F914D08CE28550590A12B40FD3DF8B6" authority="(Attems, 1936)" authorityName="Golovatch" authorityYear="2021" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[151,648,1859,1886]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="permutatus" status="comb. nov.">
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<emphasis id="EAE5EA99CE28550590A12B40FE7CF8B5" bold="true" box="[151,457,1859,1886]" italics="true" pageId="3" pageNumber="575">Pyrgodesmus permutatus</emphasis>
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(
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<bibRefCitation id="BC004B7ACE28550591EE2B40FDCAF8B6" author="Attems, C." box="[472,639,1859,1885]" pageId="3" pageNumber="575" pagination="133 - 323" refId="ref2812" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="F1D657E2CE28550592A32B40FCA9F8B6" box="[661,796,1859,1885]" pageId="3" pageNumber="575" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="908B6500CE28550590A12B64FEB3F86A" box="[151,262,1895,1921]" pageId="3" pageNumber="575" type="description">
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<paragraph id="D82E368BCE28550590A12B64FEB3F86A" blockId="3.[151,796,1859,1921]" box="[151,262,1895,1921]" pageId="3" pageNumber="575">
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<figureCitation id="40AA2A0ECE28550590A12B64FEB3F86A" box="[151,262,1895,1921]" captionStart-0="FIGURES 9–18" captionStart-1="FIGURES 19–30" captionStartId-0="3.[151,264,1717,1741]" captionStartId-1="4.[151,264,1812,1836]" captionTargetBox-0="[156,1434,817,1690]" captionTargetBox-1="[176,1412,804,1775]" captionTargetId-0="figure-306@3.[151,1436,813,1694]" captionTargetId-1="figure-231@4.[159,1429,803,1775]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." captionText-1="FIGURES 19–30. Pyrgodesmus permutatus (Attems, 1936), comb. nov., male syntype. 19. Head, dorsal view. 20. Anterior part of body, ventral view. 21. Antenna, lateral view. 22. Gnathochilarium, ventral view. 23. Anteromedial part of gnathochilarium, ventral view. 24. Right paraterga 11 and 12, dorsal view. 25. Cross-section of body ring 8, posterior view. 26. Posterior part of body, ventral view. 27. Fine structure of limbus, dorsal view. 28. Leg of body ring 8. 29. Left gonopod, mesal view. 30. Telopodite of right gonopod, mesal view. All after Silvestri (1920), drawn not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5709648" figureDoi-1="http://doi.org/10.5281/zenodo.5709650" httpUri-0="https://zenodo.org/record/5709648/files/figure.png" httpUri-1="https://zenodo.org/record/5709650/files/figure.png" pageId="3" pageNumber="575">Figs 9–30</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="908B6500CE28550290A12BAFFE8EFF26" lastPageId="4" lastPageNumber="576" pageId="3" pageNumber="575" type="reference_group">
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<paragraph id="D82E368BCE28550590A12BAFFB63F82F" blockId="3.[151,1422,1964,2019]" box="[151,1238,1964,1988]" pageId="3" pageNumber="575">
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<treatmentCitationGroup id="F88111A5CE28550590A12BAFFB63F82F" box="[151,1238,1964,1988]" pageId="3" pageNumber="575">
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<taxonomicName id="1F914D08CE28550590A12BAFFE35F82F" ID-CoL="4QT8C" authorityName="Pocock" authorityYear="1892" box="[151,384,1964,1988]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="obscurus">
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<emphasis id="EAE5EA99CE28550590A12BAFFE35F82F" box="[151,384,1964,1988]" italics="true" pageId="3" pageNumber="575">Pyrgodesmus obscurus</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="5930109ACE28550591AD2BAFFDD6F82F" author="Silvestri, F." box="[411,611,1964,1988]" page="120" pageId="3" pageNumber="575" year="1920">
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<bibRefCitation id="BC004B7ACE28550591AD2BAFFDD6F82F" author="Silvestri, F." box="[411,611,1964,1988]" pageId="3" pageNumber="575" pagination="117 - 135" refId="ref3455" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri, 1920: 120</bibRefCitation>
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</treatmentCitation>
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, figs 1–3 (D), non
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<treatmentCitation id="5930109ACE28550593152BAFFB66F82F" author="Pocock, R. I." box="[803,1235,1964,1988]" page="155" pageId="3" pageNumber="575" year="1892">
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<taxonomicName id="1F914D08CE28550593152BAFFB66F82F" ID-CoL="4QT8C" authority="Pocock, 1892: 155" authorityName="Pocock" authorityPageNumber="155" authorityYear="1892" box="[803,1235,1964,1988]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="obscurus">
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<emphasis id="EAE5EA99CE28550593152BAFFBBEF82F" box="[803,1035,1964,1988]" italics="true" pageId="3" pageNumber="575">Pyrgodesmus obscurus</emphasis>
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<bibRefCitation id="BC004B7ACE28550594242BAFFB66F82F" author="Pocock, R. I." box="[1042,1235,1964,1988]" pageId="3" pageNumber="575" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock, 1892: 155</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D82E368BCE28550590A12BC9FA3BF809" blockId="3.[151,1422,1964,2019]" box="[151,1422,1994,2019]" pageId="3" pageNumber="575">
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<treatmentCitationGroup id="F88111A5CE28550590A12BC9FA3BF809" box="[151,1422,1994,2019]" pageId="3" pageNumber="575">
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<treatmentCitation id="5930109ACE28550590A12BC9FDC6F809" author="Attems, C." box="[151,627,1994,2019]" page="246" pageId="3" pageNumber="575" year="1936">
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||
<taxonomicName id="1F914D08CE28550590A12BC9FDC6F809" ID-CoL="3R9M2" authority="Attems, 1936: 246" authorityName="Attems" authorityPageNumber="246" authorityYear="1936" box="[151,627,1994,2019]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="permutatus" status="nom. nov.">
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<emphasis id="EAE5EA99CE28550590A12BC9FE05F809" box="[151,432,1994,2018]" italics="true" pageId="3" pageNumber="575">Klimakodesmus permutatus</emphasis>
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||
<bibRefCitation id="BC004B7ACE28550591832BC8FDC6F809" author="Attems, C." box="[437,627,1994,2019]" pageId="3" pageNumber="575" pagination="133 - 323" refId="ref2812" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133 - 323." type="journal article" year="1936">Attems, 1936: 246</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
,
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||
<taxonomicNameLabel id="F1D657E2CE285505924B2BC8FD56F809" box="[637,739,1995,2018]" pageId="3" pageNumber="575">nom. nov.</taxonomicNameLabel>
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||
pro
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||
<taxonomicName id="1F914D08CE28550593262BC9FC4CF809" ID-CoL="4QT8C" authorityName="Pocock" authorityYear="1892" box="[784,1017,1994,2018]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="obscurus">
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||
<emphasis id="EAE5EA99CE28550593262BC9FC4CF809" box="[784,1017,1994,2018]" italics="true" pageId="3" pageNumber="575">Pyrgodesmus obscurus</emphasis>
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</taxonomicName>
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||
in the sense of
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||
<treatmentCitation id="5930109ACE28550594AE2BC9FAEBF809" author="Silvestri, F." box="[1176,1374,1994,2018]" page="118" pageId="3" pageNumber="575" year="1920">
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<bibRefCitation id="BC004B7ACE28550594AE2BC9FAEBF809" author="Silvestri, F." box="[1176,1374,1994,2018]" pageId="3" pageNumber="575" pagination="117 - 135" refId="ref3455" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri, 1920: 118</bibRefCitation>
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(D).
|
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</paragraph>
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<paragraph id="D82E368BCE2F550290A12C94FE8EFF26" blockId="4.[151,1437,151,205]" pageId="4" pageNumber="576">
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<treatmentCitationGroup id="F88111A5CE2F550290A12C94FE8EFF26" pageId="4" pageNumber="576">
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<taxonomicName id="1F914D08CE2F550290A12C94FE1AFF45" ID-CoL="3R9M2" authorityName="Attems" authorityYear="1936" box="[151,431,151,175]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="permutatus">
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||
<emphasis id="EAE5EA99CE2F550290A12C94FE1AFF45" box="[151,431,151,175]" italics="true" pageId="4" pageNumber="576">Klimakodesmus permutatus</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="5930109ACE2F550291FC2C94FD30FF44" author="Attems, C." box="[458,645,151,175]" page="271" pageId="4" pageNumber="576" year="1940">
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<bibRefCitation id="BC004B7ACE2F550291FC2C94FD30FF44" author="Attems, C." box="[458,645,151,175]" pageId="4" pageNumber="576" pagination="1 - 577" refId="ref2838" refString="Attems, C. (1940) Myriopoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaeriotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich, 70, i - xxxii + 1 - 577. https: // doi. org / 10.1515 / 9783111609645" type="book chapter" year="1940">Attems, 1940: 271</bibRefCitation>
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</treatmentCitation>
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||
, figs 384–386 (D, R);
|
||
<treatmentCitation id="5930109ACE2F550293502C94FBC5FF44" ID-DOI="http://doi.org/10.5281/zenodo.6059180" ID-Zenodo-Dep="6059180" author="De Zoysa, H. K. S. & Nguyen, D. A. & Wickramasinghe, S." box="[870,1136,151,175]" httpUri="http://treatment.plazi.org/id/1E1387ACFFD11B68FF6F403C1D84FB4C" page="477" pageId="4" pageNumber="576" year="2016">
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<bibRefCitation id="BC004B7ACE2F550293502C94FBC5FF44" author="De Zoysa, H. K. S. & Nguyen, D. A. & Wickramasinghe, S." box="[870,1136,151,175]" pageId="4" pageNumber="576" pagination="451 - 482" refId="ref2941" refString="De Zoysa, H. K. S., Nguyen, D. A. & Wickramasinghe, S. (2016) Annotated checklist of millipedes (Myriapoda: Diplopoda) of Sri Lanka. Zootaxa, 4061 (5), 451 - 482. https: // doi. org / 10.11646 / zootaxa. 4061.5.1" type="journal article" year="2016">
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||
De Zoysa
|
||
<emphasis id="EAE5EA99CE2F550293FA2C94FC4AFF44" box="[972,1023,151,175]" italics="true" pageId="4" pageNumber="576">et al.</emphasis>
|
||
, 2016: 477
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(R, M);
|
||
<treatmentCitation id="5930109ACE2F550294F32C94FF58FF26" author="Aswathy, M. D. & Golovatch, S. I. & Sudhikumar, A. V." page="374" pageId="4" pageNumber="576" year="2021">
|
||
<bibRefCitation id="BC004B7ACE2F550294F32C94FF58FF26" author="Aswathy, M. D. & Golovatch, S. I. & Sudhikumar, A. V." pageId="4" pageNumber="576" pagination="373 - 382" refId="ref2712" refString="Aswathy, M. D., Golovatch, S. I. & Sudhikumar, A. V. (2021) The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae). Zootaxa, 4980 (2), 373 - 382. https: // doi. org / 10.11646 / zootaxa. 4980.2.8" type="journal article" year="2021">
|
||
Aswathy
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||
<emphasis id="EAE5EA99CE2F550295132C94FAE2FF44" box="[1317,1367,151,175]" italics="true" pageId="4" pageNumber="576">et al.</emphasis>
|
||
, 2021: 374
|
||
</bibRefCitation>
|
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</treatmentCitation>
|
||
(D, R).
|
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="908B6500CE2F550290A12CFFFE4FFD02" pageId="4" pageNumber="576" type="discussion">
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<paragraph id="D82E368BCE2F550290A12CFFFE95FEB5" blockId="4.[151,1437,251,781]" pageId="4" pageNumber="576">
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||
<emphasis id="EAE5EA99CE2F550290A12CFFFEB5FEFD" bold="true" box="[151,256,252,278]" pageId="4" pageNumber="576">Remark.</emphasis>
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||
As noted above, the original description and illustrations of this species were most detailed (
|
||
<bibRefCitation id="BC004B7ACE2F5502950B2CF8FF66FED1" author="Silvestri, F." pageId="4" pageNumber="576" pagination="117 - 135" refId="ref3455" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri 1920</bibRefCitation>
|
||
), albeit misidentified as belonging to
|
||
<taxonomicName id="1F914D08CE2F550292BD2D1CFC25FED2" authorityName="Pocock" authorityYear="1892" box="[651,912,287,313]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="EAE5EA99CE2F550292BD2D1CFC25FED2" box="[651,912,287,313]" italics="true" pageId="4" pageNumber="576">Pyrgodesmus obscurus</emphasis>
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||
</taxonomicName>
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||
. Silvestri’s iconography is reproduced in full (
|
||
<figureCitation id="40AA2A0ECE2F550290A92D40FEA1FEB5" box="[159,276,323,350]" captionStart-0="FIGURES 9–18" captionStart-1="FIGURES 19–30" captionStartId-0="3.[151,264,1717,1741]" captionStartId-1="4.[151,264,1812,1836]" captionTargetBox-0="[156,1434,817,1690]" captionTargetBox-1="[176,1412,804,1775]" captionTargetId-0="figure-306@3.[151,1436,813,1694]" captionTargetId-1="figure-231@4.[159,1429,803,1775]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." captionText-1="FIGURES 19–30. Pyrgodesmus permutatus (Attems, 1936), comb. nov., male syntype. 19. Head, dorsal view. 20. Anterior part of body, ventral view. 21. Antenna, lateral view. 22. Gnathochilarium, ventral view. 23. Anteromedial part of gnathochilarium, ventral view. 24. Right paraterga 11 and 12, dorsal view. 25. Cross-section of body ring 8, posterior view. 26. Posterior part of body, ventral view. 27. Fine structure of limbus, dorsal view. 28. Leg of body ring 8. 29. Left gonopod, mesal view. 30. Telopodite of right gonopod, mesal view. All after Silvestri (1920), drawn not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5709648" figureDoi-1="http://doi.org/10.5281/zenodo.5709650" httpUri-0="https://zenodo.org/record/5709648/files/figure.png" httpUri-1="https://zenodo.org/record/5709650/files/figure.png" pageId="4" pageNumber="576">Figs 9–30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D82E368BCE2F550290F12D64FC2DFE4D" blockId="4.[151,1437,251,781]" pageId="4" pageNumber="576">
|
||
<emphasis id="EAE5EA99CE2F550290F12D64FE04FE6A" bold="true" box="[199,433,359,385]" pageId="4" pageNumber="576">Brief redescription.</emphasis>
|
||
Body length
|
||
<emphasis id="EAE5EA99CE2F5502927B2D6BFDD2FE6A" box="[589,615,360,385]" italics="true" pageId="4" pageNumber="576">ca</emphasis>
|
||
<quantity id="1F699B6ECE2F550292462D64FD62FE69" box="[624,727,359,386]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="576" unit="mm" value="11.5">11.5 mm</quantity>
|
||
. Coloration mostly black-brown, regardless of tegument being heavily coated with an earth crust (
|
||
<figureCitation id="40AA2A0ECE2F550292292D88FD27FE4D" box="[543,658,395,422]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1717,1741]" captionTargetBox="[156,1434,817,1690]" captionTargetId="figure-306@3.[151,1436,813,1694]" captionTargetPageId="3" captionText="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="4" pageNumber="576">Figs 9–12</figureCitation>
|
||
) or clean (
|
||
<figureCitation id="40AA2A0ECE2F5502933E2D88FC3EFE4D" box="[776,907,395,422]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1717,1741]" captionTargetBox="[156,1434,817,1690]" captionTargetId="figure-306@3.[151,1436,813,1694]" captionTargetPageId="3" captionText="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="4" pageNumber="576">Figs 13–18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D82E368BCE2F550290A12DACFCB8FE22" blockId="4.[151,1437,251,781]" box="[151,781,431,458]" pageId="4" pageNumber="576">
|
||
All main characters as in
|
||
<taxonomicName id="1F914D08CE2F550291832DB3FD80FE22" authorityName="Pocock" authorityYear="1892" box="[437,565,431,457]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="EAE5EA99CE2F550291832DB3FD80FE22" box="[437,565,431,457]" italics="true" pageId="4" pageNumber="576">P. obscurus</emphasis>
|
||
</taxonomicName>
|
||
, except as follows.
|
||
</paragraph>
|
||
<paragraph id="D82E368BCE2F550290F12DD0FEE3FD95" blockId="4.[151,1437,251,781]" pageId="4" pageNumber="576">
|
||
Paramedian tubercles/crests (
|
||
<emphasis id="EAE5EA99CE2F550292272DD7FD88FE05" bold="true" box="[529,573,468,494]" pageId="4" pageNumber="576">PM</emphasis>
|
||
) slightly less strongly developed, high, largely fully separated and not fused medially even at base (except for
|
||
<emphasis id="EAE5EA99CE2F5502922F2DFBFDF0FDF9" bold="true" box="[537,581,504,530]" pageId="4" pageNumber="576">PM</emphasis>
|
||
19), inclined anteriad on collum and on rings 2 and 3 (
|
||
<figureCitation id="40AA2A0ECE2F5502948A2DF4FA87FDF9" box="[1212,1330,503,530]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1717,1741]" captionTargetBox="[156,1434,817,1690]" captionTargetId="figure-306@3.[151,1436,813,1694]" captionTargetPageId="3" captionText="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="4" pageNumber="576">Figs 9–18</figureCitation>
|
||
). Surface below
|
||
<emphasis id="EAE5EA99CE2F550290D72E1FFEB8FDDD" bold="true" box="[225,269,540,566]" pageId="4" pageNumber="576">PM</emphasis>
|
||
regularly microgranulate, with neither
|
||
<emphasis id="EAE5EA99CE2F550292892E1FFD53FDDD" bold="true" box="[703,742,540,566]" pageId="4" pageNumber="576">DL</emphasis>
|
||
nor
|
||
<emphasis id="EAE5EA99CE2F550293212E18FCAAFDDE" bold="true" box="[791,799,539,565]" pageId="4" pageNumber="576">i</emphasis>
|
||
, nor
|
||
<emphasis id="EAE5EA99CE2F550293602E18FC37FDDE" bold="true" box="[854,898,539,565]" pageId="4" pageNumber="576">Am</emphasis>
|
||
, nor
|
||
<emphasis id="EAE5EA99CE2F5502938C2E18FC53FDDE" bold="true" box="[954,998,539,565]" pageId="4" pageNumber="576">Cm</emphasis>
|
||
discernible beneath an earth crust (
|
||
<figureCitation id="40AA2A0ECE2F5502955C2E18FF66FDB1" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1717,1741]" captionTargetBox="[156,1434,817,1690]" captionTargetId="figure-306@3.[151,1436,813,1694]" captionTargetPageId="3" captionText="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="4" pageNumber="576">Figs 9–12</figureCitation>
|
||
), but readily visible in a clean animal (
|
||
<figureCitation id="40AA2A0ECE2F5502924B2E3CFD4AFDB1" box="[637,767,575,602]" captionStart="FIGURES 9–18" captionStartId="3.[151,264,1717,1741]" captionTargetBox="[156,1434,817,1690]" captionTargetId="figure-306@3.[151,1436,813,1694]" captionTargetPageId="3" captionText="FIGURES 9–18. Pyrgodesmus permutatus (Attems, 1936), comb. nov., habituses of two male syntypes, one coated with earth (10–12) and the other clean (13–18). All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="4" pageNumber="576">Figs 13–18</figureCitation>
|
||
). Tip of epiproct invisible from above between basally fused
|
||
<emphasis id="EAE5EA99CE2F550290A12E67FF76FD95" bold="true" box="[151,195,612,638]" pageId="4" pageNumber="576">PM</emphasis>
|
||
19 (
|
||
<figureCitation id="40AA2A0ECE2F550290C32E60FEFCFD95" box="[245,329,611,638]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[176,1412,804,1775]" captionTargetId="figure-231@4.[159,1429,803,1775]" captionTargetPageId="4" captionText="FIGURES 19–30. Pyrgodesmus permutatus (Attems, 1936), comb. nov., male syntype. 19. Head, dorsal view. 20. Anterior part of body, ventral view. 21. Antenna, lateral view. 22. Gnathochilarium, ventral view. 23. Anteromedial part of gnathochilarium, ventral view. 24. Right paraterga 11 and 12, dorsal view. 25. Cross-section of body ring 8, posterior view. 26. Posterior part of body, ventral view. 27. Fine structure of limbus, dorsal view. 28. Leg of body ring 8. 29. Left gonopod, mesal view. 30. Telopodite of right gonopod, mesal view. All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709650" httpUri="https://zenodo.org/record/5709650/files/figure.png" pageId="4" pageNumber="576">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D82E368BCE2F550290F12E84FE4FFD02" blockId="4.[151,1437,251,781]" pageId="4" pageNumber="576">
|
||
Gonopods (
|
||
<figureCitation id="40AA2A0ECE2F5502917C2E84FE51FD49" box="[330,484,647,674]" captionStart="FIGURES 19–30" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[176,1412,804,1775]" captionTargetId="figure-231@4.[159,1429,803,1775]" captionTargetPageId="4" captionText="FIGURES 19–30. Pyrgodesmus permutatus (Attems, 1936), comb. nov., male syntype. 19. Head, dorsal view. 20. Anterior part of body, ventral view. 21. Antenna, lateral view. 22. Gnathochilarium, ventral view. 23. Anteromedial part of gnathochilarium, ventral view. 24. Right paraterga 11 and 12, dorsal view. 25. Cross-section of body ring 8, posterior view. 26. Posterior part of body, ventral view. 27. Fine structure of limbus, dorsal view. 28. Leg of body ring 8. 29. Left gonopod, mesal view. 30. Telopodite of right gonopod, mesal view. All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709650" httpUri="https://zenodo.org/record/5709650/files/figure.png" pageId="4" pageNumber="576">Figs 29 & 30</figureCitation>
|
||
) much as in
|
||
<taxonomicName id="1F914D08CE2F5502924F2E8BFD4EFD4A" authorityName="Pocock" authorityYear="1892" box="[633,763,647,673]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="EAE5EA99CE2F5502924F2E8BFD4EFD4A" box="[633,763,647,673]" italics="true" pageId="4" pageNumber="576">P. obscurus</emphasis>
|
||
</taxonomicName>
|
||
, but gonocoel slightly less deep, coxites apparently neither conspicuously microsetose nor micropapillate, telopodites/solenomeres more clearly exposed, but stout, their tips being somewhat curved dorsad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="908B6500CE2F550290F12EF0FDD2FCE6" box="[199,615,755,781]" pageId="4" pageNumber="576" type="distribution">
|
||
<paragraph id="D82E368BCE2F550290F12EF0FDD2FCE6" blockId="4.[151,1437,251,781]" box="[199,615,755,781]" pageId="4" pageNumber="576">
|
||
<emphasis id="EAE5EA99CE2F550290F12EF0FED4FCE6" bold="true" box="[199,353,755,781]" pageId="4" pageNumber="576">Distribution.</emphasis>
|
||
Paradenyia,
|
||
<collectingCountry id="A086761BCE2F550291C42EF0FDD1FCE6" box="[498,612,755,781]" name="Sri Lanka" pageId="4" pageNumber="576">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |