Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida) Author Golovatch, Sergei I. text Zootaxa 2021 2021-11-18 5068 4 572 578 journal article 3390 10.11646/zootaxa.5068.4.6 79b5ca84-737c-4722-9994-9f5cdefbebb3 1175-5326 5709642 E4A3743F-14A9-4292-AC55-56B7E7F0905C Pyrgodesmus permutatus ( Attems, 1936 ) , comb. nov. Figs 9–30 Pyrgodesmus obscurus — Silvestri, 1920: 120 , figs 1–3 (D), non Pyrgodesmus obscurus Pocock, 1892: 155 . Klimakodesmus permutatus Attems, 1936: 246 , nom. nov. pro Pyrgodesmus obscurus in the sense of Silvestri, 1920: 118 (D). Klimakodesmus permutatus — Attems, 1940: 271 , figs 384–386 (D, R); De Zoysa et al. , 2016: 477 (R, M); Aswathy et al. , 2021: 374 (D, R). Remark. As noted above, the original description and illustrations of this species were most detailed ( Silvestri 1920 ), albeit misidentified as belonging to Pyrgodesmus obscurus . Silvestri’s iconography is reproduced in full ( Figs 9–30 ). Brief redescription. Body length ca 11.5 mm . Coloration mostly black-brown, regardless of tegument being heavily coated with an earth crust ( Figs 9–12 ) or clean ( Figs 13–18 ). All main characters as in P. obscurus , except as follows. Paramedian tubercles/crests ( PM ) slightly less strongly developed, high, largely fully separated and not fused medially even at base (except for PM 19), inclined anteriad on collum and on rings 2 and 3 ( Figs 9–18 ). Surface below PM regularly microgranulate, with neither DL nor i , nor Am , nor Cm discernible beneath an earth crust ( Figs 9–12 ), but readily visible in a clean animal ( Figs 13–18 ). Tip of epiproct invisible from above between basally fused PM 19 ( Fig. 26 ). Gonopods ( Figs 29 & 30 ) much as in P. obscurus , but gonocoel slightly less deep, coxites apparently neither conspicuously microsetose nor micropapillate, telopodites/solenomeres more clearly exposed, but stout, their tips being somewhat curved dorsad. Distribution. Paradenyia, Sri Lanka .