292 lines
42 KiB
XML
292 lines
42 KiB
XML
<document id="E378AC49E4E5859D7A68AF6BB7E30190" ID-CLB-Dataset="46195" ID-DOI="10.11646/zootaxa.3779.5.6" ID-GBIF-Dataset="b798f275-02d9-456e-98d0-f116607218b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="226678" ID-ZooBank="491EB308-B6CD-411C-AF74-78B37D8C526F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460398479612" checkinUser="plazi" docAuthor="Archangelsky, Miguel & Brand, Cecilia" docDate="2014" docId="4932878EFF9EFFAF66A0F9A6FBD0FE67" docLanguage="en" docName="zt03779p572.pdf" docOrigin="Zootaxa 3779 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luchoelmis kapenkemkensis" docType="treatment" docVersion="9" lastPageNumber="572" masterDocId="B50BFFF6FF9AFFA66637FF94FF95FFB7" masterDocTitle="A new species of Luchoelmis Spangler & Staines (Coleoptera: Elmidae) from Argentina and its probable larva" masterLastPageNumber="572" masterPageNumber="563" pageNumber="567" updateTime="1698298959420" updateUser="ExternalLinkService">
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<mods:titleInfo id="9B2A949A16CA2C677A72159DFDBE6C1C">
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<mods:title id="25530A678FD85E67129266EDB8383052">A new species of Luchoelmis Spangler & Staines (Coleoptera: Elmidae) from Argentina and its probable larva</mods:title>
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<mods:name id="431259B8794BF9154CF3566A9323F54E" type="personal">
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<mods:namePart id="E496A031BB8A8670E9912B3092959A5A">Archangelsky, Miguel</mods:namePart>
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<mods:namePart id="E6D1C76E94A7F1D6E53C04E431A82706">Brand, Cecilia</mods:namePart>
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<mods:title id="F600FD4878BD839501A83FB1E9830EFD">Zootaxa</mods:title>
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<mods:part id="2C149EAA5F362193E4B1EF96CE345487">
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<mods:date id="6C069014AAAC300AC6FF3A3973929417">2014</mods:date>
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<mods:number id="539748A6060274EEB7E8C238C7084D61">3779</mods:number>
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<mods:number id="BF3996E02AF48385CC62A85DA9384E36">5</mods:number>
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<mods:identifier id="0C4C0B190A9BF199D402E17D20C58DF4" type="DOI">10.11646/zootaxa.3779.5.6</mods:identifier>
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<treatment id="4932878EFF9EFFAF66A0F9A6FBD0FE67" ID-DOI="http://doi.org/10.5281/zenodo.6134051" ID-GBIF-Taxon="119418912" ID-Zenodo-Dep="6134051" LSID="urn:lsid:plazi:treatment:4932878EFF9EFFAF66A0F9A6FBD0FE67" httpUri="http://treatment.plazi.org/id/4932878EFF9EFFAF66A0F9A6FBD0FE67" lastPageId="9" lastPageNumber="572" pageId="4" pageNumber="567">
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<subSubSection id="89816513FF9EFFA266A0F9A6FD10F9FB" box="[151,645,1586,1612]" pageId="4" pageNumber="567" type="nomenclature">
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<paragraph id="C1243698FF9EFFA266A0F9A6FD10F9FB" blockId="4.[151,645,1586,1612]" box="[151,645,1586,1612]" pageId="4" pageNumber="567">
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<heading id="9A6C81F4FF9EFFA266A0F9A6FD10F9FB" bold="true" box="[151,645,1586,1612]" fontSize="10" level="2" pageId="4" pageNumber="567" reason="2">
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<emphasis id="F3EFEA8AFF9EFFA266A0F9A6FD10F9FB" bold="true" box="[151,645,1586,1612]" pageId="4" pageNumber="567">
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<taxonomicName id="069B4D1BFF9EFFA266A0F9A6FE76F9FC" ID-CoL="3WBCC" box="[151,483,1586,1611]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="kapenkemkensis">
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<emphasis id="F3EFEA8AFF9EFFA266A0F9A6FE76F9FC" bold="true" box="[151,483,1586,1611]" italics="true" pageId="4" pageNumber="567">Luchoelmis kapenkemkensis</emphasis>
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</taxonomicName>
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mature larva
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="89816513FF9EFFAF66A0F9EFFBD0FE67" lastPageId="9" lastPageNumber="572" pageId="4" pageNumber="567" type="description">
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<paragraph id="C1243698FF9EFFA266A0F9EFFED4F948" blockId="4.[151,1436,1659,2008]" pageId="4" pageNumber="567">
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<emphasis id="F3EFEA8AFF9EFFA266A0F9EFFEB4F923" bold="true" box="[151,289,1659,1684]" pageId="4" pageNumber="567">Description</emphasis>
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. Body (
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<figureCitation id="59A02A1DFF9EFFA26749F9EFFE43F923" box="[382,470,1659,1684]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="4" pageNumber="567">Figs 12</figureCitation>
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,
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<figureCitation id="59A02A1DFF9EFFA267D4F9E8FDBEF923" box="[483,555,1660,1684]" captionStart="FIGURES 22 – 26" captionStartId="8.[151,264,1847,1870]" captionTargetBox="[273,1343,217,1791]" captionTargetId="figure@8.[236,1358,193,1817]" captionTargetPageId="8" captionText="FIGURES 22 – 26. Mature larva of Luchoelmis kapenkemkensis: 22) Thorax, ventral view; 23) detail of first five abdominal segments, dorsal view; 24) fourth abdominal segment, lateral view; 25) operculum of ninth abdominal segment, ventral view; 26) hooks of tracheal chamber. Scale bars: Figs 22, 23: 0.5 mm; Figs 24 – 26: 0.1 mm." httpUri="https://zenodo.org/record/226684/files/figure.png" pageId="4" pageNumber="567">23, 24</figureCitation>
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) elongate, sides subparallel, widest at thorax, abdominal segments narrowing towards posterior end; body subtriangular in cross-section. Dorsal surface of thorax and abdomen strongly tuberculate, tubercles arranged in longitudinal rows. Color reddish-brown. Length:
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<quantity id="06639B7DFF9EFFA26273F957FB5AF96C" box="[1092,1231,1731,1756]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="4" pageNumber="567" unit="mm" value="3.7" valueMax="3.9" valueMin="3.5">3.5–3.9 mm</quantity>
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; maximum width:
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<quantity id="06639B7DFF9EFFA266A0F973FEA8F948" box="[151,317,1767,1792]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="4" pageNumber="567" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.70 mm</quantity>
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.
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</paragraph>
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<paragraph id="C1243698FF9EFFA266F0F89FFCF3F803" blockId="4.[151,1436,1659,2008]" pageId="4" pageNumber="567">
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Head capsule (
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<figureCitation id="59A02A1DFF9EFFA2675AF89FFE7AF893" box="[365,495,1803,1828]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="4" pageNumber="567">Figs 12, 13</figureCitation>
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) exposed, not covered by pronotum, anterior margin lacking tooth between base of antenna and clypeus. Surface with several large tubercles on disc, close to frontal lines. Coronal line very short, frontal lines long, extending to inner margin of antennal sockets. Fronto-clypeal suture present, feeble. Gula subtrapezoidal, narrower than maxillolabial complex; basal margin wider and concave, distal margin narrower and convex. Five stemmata on each side of head at about midlength.
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</paragraph>
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<paragraph id="C1243698FF9EFFA366F0F82BFBD3FF4F" blockId="4.[151,1436,1659,2008]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="568" pageId="4" pageNumber="567">
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Labrum (
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<figureCitation id="59A02A1DFF9EFFA26702F82BFE56F86F" box="[309,451,1983,2008]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="4" pageNumber="567">Figs 15, 16</figureCitation>
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) subrectangular, slightly wider in anterior third; anterior margin slightly concave, anterolateral margins rounded, each with a row of three strong dorsal ramose setae. Six strong ramose setae arranged in a transverse row in anterior third on dorsal surface. Ventral surface with anterior row of ramose setae, rest of ventral surface covered by short pubescence oriented mediad and posteriorly.
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</paragraph>
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<paragraph id="C1243698FF9FFFA366F0FE90FD3BFE30" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="568">
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Antennae (
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<figureCitation id="59A02A1DFF9FFFA3677FFE90FE43FEAB" box="[328,470,260,285]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="5" pageNumber="568">Figs 13, 14</figureCitation>
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) short, three-segmented, located on anterolateral corners of head capsule. Basal antennomere short and wide, apically with a crown of ramose setae. Second antennomere the longest, cylindrical, with a few distal setae, bearing a short sensorioum. Third antennomere the shortest, shorter than sensorium of second antennomere, bearing a long apical seta.
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</paragraph>
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<caption id="95E46610FF9FFFA366A0F8CEFED0F819" httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="5" pageNumber="568" targetBox="[296,1306,445,1877]" targetPageId="5">
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<paragraph id="C1243698FF9FFFA366A0F8CEFED0F819" blockId="5.[151,1436,1882,1966]" pageId="5" pageNumber="568">
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<emphasis id="F3EFEA8AFF9FFFA366A0F8CEFEC1F8C7" bold="true" box="[151,340,1882,1905]" pageId="5" pageNumber="568">FIGURES 12–16.</emphasis>
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Mature larva of
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<taxonomicName id="069B4D1BFF9FFFA367C8F8CFFC88F8C6" box="[511,797,1883,1905]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="kapenkemkensis">
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<emphasis id="F3EFEA8AFF9FFFA367C8F8CFFC88F8C6" box="[511,797,1883,1905]" italics="true" pageId="5" pageNumber="568">Luchoelmis kapenkemkensis</emphasis>
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</taxonomicName>
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: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14–16: 0.05 mm.
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</paragraph>
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</caption>
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<paragraph id="C1243698FF9CFFA066F0FF03FB7CFF4F" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
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Mandibles (
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<figureCitation id="59A02A1DFF9CFFA06779FF03FE47FF07" box="[334,466,151,176]" captionStart="FIGURES 17 – 21" captionStartId="7.[151,264,1640,1663]" captionTargetBox="[296,1260,357,1603]" captionTargetId="figure@7.[277,1275,337,1623]" captionTargetPageId="7" captionText="FIGURES 17 – 21. Mature larva of Luchoelmis kapenkemkensis: 17) mandible, ventral view; 18) mandible, dorsal view; 19) maxillolabial complex showing right maxilla and labium, ventral view; 20) mesothoracic leg; 21) detail of maxillolabial complex showing distal part of maxillae and labium. Scale bars, Figs 17 – 19: 0.05 mm; Fig. 20: 0.1 mm." httpUri="https://zenodo.org/record/226683/files/figure.png" pageId="6" pageNumber="569">Figs 17, 18</figureCitation>
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) symmetrical, apex with three blunt teeth. Dorsal surface with inner margin straight and sharp. Ventral surface with inner margin concave, with a comb of long stout submarginal setae. Inner margin of mandible with long setose prostheca; outer margin with two large ramose setae close to midlength.
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</paragraph>
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<paragraph id="C1243698FF9CFFA066F0FE90FC37FE42" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
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Maxillae (
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<figureCitation id="59A02A1DFF9CFFA0670CFE90FE54FEAB" box="[315,449,260,285]" captionStart="FIGURES 17 – 21" captionStartId="7.[151,264,1640,1663]" captionTargetBox="[296,1260,357,1603]" captionTargetId="figure@7.[277,1275,337,1623]" captionTargetPageId="7" captionText="FIGURES 17 – 21. Mature larva of Luchoelmis kapenkemkensis: 17) mandible, ventral view; 18) mandible, dorsal view; 19) maxillolabial complex showing right maxilla and labium, ventral view; 20) mesothoracic leg; 21) detail of maxillolabial complex showing distal part of maxillae and labium. Scale bars, Figs 17 – 19: 0.05 mm; Fig. 20: 0.1 mm." httpUri="https://zenodo.org/record/226683/files/figure.png" pageId="6" pageNumber="569">Figs 19, 21</figureCitation>
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): cardo narrow, irregularly suboval, transverse, with a stout seta close to outer margin. Stipes the largest segment, subrectangular, 2.6 times as long as wide; distal third with several setae distributed as follows: two long setae on outer margin (distal one ramose), inner margin with one short stout seta, distal margin with one ramose seta close to base of lacinia. Lacinia and galea well developed; lacinia subtriangular, fused to stipes, with inner margin bearing a group of stout setae; galea 1-segmented, shorter than lacinia, elongate, with several apical setae. Palpus four-segmented, first palpomere the shortest, remaining three palpomeres subequal in length, last palpomere bearing several short apical setae and sensoria.
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</paragraph>
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<paragraph id="C1243698FF9CFFA066F0FE6BFAF0FD1F" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
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Labium (
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<figureCitation id="59A02A1DFF9CFFA0671AFE6BFE39FDAF" box="[301,428,511,536]" captionStart="FIGURES 17 – 21" captionStartId="7.[151,264,1640,1663]" captionTargetBox="[296,1260,357,1603]" captionTargetId="figure@7.[277,1275,337,1623]" captionTargetPageId="7" captionText="FIGURES 17 – 21. Mature larva of Luchoelmis kapenkemkensis: 17) mandible, ventral view; 18) mandible, dorsal view; 19) maxillolabial complex showing right maxilla and labium, ventral view; 20) mesothoracic leg; 21) detail of maxillolabial complex showing distal part of maxillae and labium. Scale bars, Figs 17 – 19: 0.05 mm; Fig. 20: 0.1 mm." httpUri="https://zenodo.org/record/226683/files/figure.png" pageId="6" pageNumber="569">Figs 19, 21</figureCitation>
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) large, subdivided into a large postmentum and a short prementum, forming together with maxillae the maxilla-labial complex. Postmentum subrectangular, 1.6 times as long as wide, ventral surface with several short ramose setae at each side of middline; basal corners each with a large and stout seta; distal corners each with a large ramose seta. Prementum short, poorly sclerotized, wider than long, distal margins densely setose. Palps two-segmented basal palpomere slightly shorter, distal palpomere with several distal setae and sensoria.
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</paragraph>
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<paragraph id="C1243698FF9CFFA066F0FD20FF45FC5A" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
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Thorax (
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<figureCitation id="59A02A1DFF9CFFA06710FD20FEE8FD7B" box="[295,381,692,717]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="6" pageNumber="569">Figs 12</figureCitation>
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,
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<figureCitation id="59A02A1DFF9CFFA067BDFD20FE3DFD7B" box="[394,424,692,716]" captionStart="FIGURES 17 – 21" captionStartId="7.[151,264,1640,1663]" captionTargetBox="[296,1260,357,1603]" captionTargetId="figure@7.[277,1275,337,1623]" captionTargetPageId="7" captionText="FIGURES 17 – 21. Mature larva of Luchoelmis kapenkemkensis: 17) mandible, ventral view; 18) mandible, dorsal view; 19) maxillolabial complex showing right maxilla and labium, ventral view; 20) mesothoracic leg; 21) detail of maxillolabial complex showing distal part of maxillae and labium. Scale bars, Figs 17 – 19: 0.05 mm; Fig. 20: 0.1 mm." httpUri="https://zenodo.org/record/226683/files/figure.png" pageId="6" pageNumber="569">20</figureCitation>
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,
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<figureCitation id="59A02A1DFF9CFFA06783FD20FE44FD7B" box="[436,465,692,716]" captionStart="FIGURES 22 – 26" captionStartId="8.[151,264,1847,1870]" captionTargetBox="[273,1343,217,1791]" captionTargetId="figure@8.[236,1358,193,1817]" captionTargetPageId="8" captionText="FIGURES 22 – 26. Mature larva of Luchoelmis kapenkemkensis: 22) Thorax, ventral view; 23) detail of first five abdominal segments, dorsal view; 24) fourth abdominal segment, lateral view; 25) operculum of ninth abdominal segment, ventral view; 26) hooks of tracheal chamber. Scale bars: Figs 22, 23: 0.5 mm; Figs 24 – 26: 0.1 mm." httpUri="https://zenodo.org/record/226684/files/figure.png" pageId="6" pageNumber="569">22</figureCitation>
|
||
) strongly sclerotized; tergal plates with sagittal lines and with large tubercles arranged in longitudinal rows, those on pronotum sinuous. Prothorax the largest segment, pronotum subtrapezoidal, anterior and posterior corners rounded; ventral region with seven sclerites: one large and irregularly shaped transverse anterolateral pair, one small, triangular lateral pair, one large posterior pair and one small suboval sclerite between procoxae; coxal cavities open. Meso- and metathorax shorter than prothorax, twice as wide as long; each segment ventrally with five sclerites: one large anterior subpentagonal sclerite, and two smaller subrectangular sclerites on each side; coxal cavities open. Legs five-segmetned; coxa the largest segment, subtriangular; trochanter smaller, subtriangular; femur and tibia elongate, femur slightly longer and wider than tibia; claw stout, slightly shorter than tibia.
|
||
</paragraph>
|
||
<paragraph id="C1243698FF9CFFA066F0FC63FF4BFAE2" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
|
||
Abdomen (
|
||
<figureCitation id="59A02A1DFF9CFFA0677EFC63FE31FBA7" box="[329,420,1015,1040]" captionStart="FIGURES 12 – 16" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[296,1306,445,1877]" captionTargetId="figure@5.[296,1308,445,1879]" captionTargetPageId="5" captionText="FIGURES 12 – 16. Mature larva of Luchoelmis kapenkemkensis: 12) habitus, dorsolateral view; 13) head capsule, dorsal view; 14) antenna, dorsal view; 15) labrum, dorsal view; 16) labrum, ventral view. Scale bars, Fig. 12: 1 mm; Fig. 13: 0.1 mm; Figs 14 – 16: 0.05 mm." httpUri="https://zenodo.org/record/226682/files/figure.png" pageId="6" pageNumber="569">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation id="59A02A1DFF9CFFA06782FC63FE68FBA7" box="[437,509,1015,1040]" captionStart="FIGURES 22 – 26" captionStartId="8.[151,264,1847,1870]" captionTargetBox="[273,1343,217,1791]" captionTargetId="figure@8.[236,1358,193,1817]" captionTargetPageId="8" captionText="FIGURES 22 – 26. Mature larva of Luchoelmis kapenkemkensis: 22) Thorax, ventral view; 23) detail of first five abdominal segments, dorsal view; 24) fourth abdominal segment, lateral view; 25) operculum of ninth abdominal segment, ventral view; 26) hooks of tracheal chamber. Scale bars: Figs 22, 23: 0.5 mm; Figs 24 – 26: 0.1 mm." httpUri="https://zenodo.org/record/226684/files/figure.png" pageId="6" pageNumber="569">23–26</figureCitation>
|
||
) well sclerotized, composed of nine segments, tapering towards posterior end; segments I–VII with sagittal line, subequal in length; segment IX the longest. Tergal plates with setiferous tubercles arranged in eight longitudinal rows on segments I–VI, three rows above spiracle and one row of smaller tubercles below spiracle; segments VII–VIII with six longitudinal rows; segment IX with three rows of tubercles, one dorsal and two lateral; all tubercles bearing large comb-like setae. Pleural sclerites present on segments I–V; sterna of segments I–V subrectangular, wider than long. Segment IX elongate, 3.4 times as long as previous segment, bearing a dorsal keel, apex slightly emarginated; sternal area with apical gill chamber, operculum subtriangular, sharply pointed, covering a pair of strong distal hooks bearing an inner row of sharp spines on inner margin and several long setae on outer margin. Spiracles present on segments I–VIII, those of segments I–VI much larger.
|
||
</paragraph>
|
||
<paragraph id="C1243698FF9CFFA066F0FACBFC23F9C3" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
|
||
<emphasis id="F3EFEA8AFF9CFFA066F0FACBFE21FACF" bold="true" box="[199,436,1375,1400]" pageId="6" pageNumber="569">Comparative notes.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="069B4D1BFF9CFFA06403FAF5FC9CFACF" box="[564,777,1376,1400]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA06403FAF5FC9CFACF" box="[564,777,1376,1400]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
display several characters not found in the other known species of the genus, i.e. color pattern, abdominal plastron and cleaning fringes on meso- and metatibiae. These characters make
|
||
<taxonomicName id="069B4D1BFF9CFFA0676EFA3DFDB8FA77" box="[345,557,1448,1472]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA0676EFA3DFDB8FA77" box="[345,557,1448,1472]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
markedly different from the remaining species, but at the same time it also shows most of the generic diagnostic characters defining
|
||
<taxonomicName id="069B4D1BFF9CFFA06502FA58FC23FA53" box="[821,950,1484,1508]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF9CFFA06502FA58FC23FA53" box="[821,950,1484,1508]" italics="true" pageId="6" pageNumber="569">Luchoelmis</emphasis>
|
||
</taxonomicName>
|
||
(with the exception of the tibial cleaning fringes, but see below). Based on this we think, that
|
||
<taxonomicName id="069B4D1BFF9CFFA064C1FA65FC5CF9BF" box="[758,969,1520,1544]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA064C1FA65FC5CF9BF" box="[758,969,1520,1544]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
could represent a distinct species within
|
||
<taxonomicName id="069B4D1BFF9CFFA066A0F980FE8DF99B" box="[151,280,1556,1580]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF9CFFA066A0F980FE8DF99B" box="[151,280,1556,1580]" italics="true" pageId="6" pageNumber="569">Luchoelmis</emphasis>
|
||
</taxonomicName>
|
||
, something that could explain the strong differences found in the larvae. It is worth mentioning here that the
|
||
<taxonomicName id="069B4D1BFF9CFFA066C3F9ADFE11F9E7" box="[244,388,1592,1616]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9CFFA066C3F9ADFE11F9E7" box="[244,388,1592,1616]" italics="true" pageId="6" pageNumber="569">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
specimens we examined also have cleaning fringes on the meso- and metatibiae, something that contradicts the original description of
|
||
<bibRefCitation id="A50A4B69FF9CFFA06444F9C8FC27F9C3" author="Spangler" box="[627,946,1628,1653]" pageId="6" pageNumber="569" refString="Spangler, P. J. & Staines, C. L. (2002) Luchoelmis, a new genus of Elmidae (Coleoptera) from Chile and Argentina. Insecta Mundi, 16, 215 - 220." type="journal article" year="2002">Spangler and Staines (2002)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C1243698FF9CFFA066F0F9EBFB55F80E" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="569">
|
||
Larval differences between
|
||
<taxonomicName id="069B4D1BFF9CFFA067CBF915FD5BF92F" box="[508,718,1664,1688]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA067CBF915FD5BF92F" box="[508,718,1664,1688]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="069B4D1BFF9CFFA06534F915FC04F92F" box="[771,913,1664,1688]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9CFFA06534F915FC04F92F" box="[771,913,1664,1688]" italics="true" pageId="6" pageNumber="569">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
are very pronounced. This is something that at a first glance called our attention, and made us doubt of the association with adults of
|
||
<taxonomicName id="069B4D1BFF9CFFA0626AF931FABAF90B" box="[1117,1327,1700,1724]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA0626AF931FABAF90B" box="[1117,1327,1700,1724]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
, but after seven years of fieldwork no other elmid adults and larvae have appeared at this locality (
|
||
<taxonomicName id="069B4D1BFF9CFFA062B5F95DFA84F957" box="[1154,1297,1736,1760]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9CFFA062B5F95DFA84F957" box="[1154,1297,1736,1760]" italics="true" pageId="6" pageNumber="569">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
larvae were associated with adults by rearing). Looking into the literature, significant differences in larval morphology among species of the same genus have been reported in
|
||
<collectingCountry id="B98C7608FF9CFFA064F0F89BFCBBF89F" box="[711,814,1807,1832]" name="Australia" pageId="6" pageNumber="569">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="A50A4B69FF9CFFA06509F89BFC48F89F" author="Glaister" box="[830,989,1807,1832]" pageId="6" pageNumber="569" refString="Glaister, A. (1999) Guide to the identification of Australian Elmidae larvae (Insecta: Coleoptera). Cooperative Research Centre for Freshwater Ecology, Identification guide No. 21, 49 pp." type="book" year="1999">Glaister 1999</bibRefCitation>
|
||
). Within the genera
|
||
<taxonomicName id="069B4D1BFF9CFFA062F2F884FF5AF8FA" authority="Carter & Zeck" authorityName="Carter & Zeck" class="Insecta" family="Elmidae" genus="Notriolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF9CFFA062F2F884FABAF89F" box="[1221,1327,1808,1832]" italics="true" pageId="6" pageNumber="569">Notriolus</emphasis>
|
||
Carter & Zeck
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="069B4D1BFF9CFFA0673AF8A0FDBFF8FA" authority="Carter & Zeck" authorityName="Carter & Zeck" box="[269,554,1844,1869]" class="Insecta" family="Elmidae" genus="Simsonia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF9CFFA0673AF8A0FEE1F8FC" box="[269,372,1844,1867]" italics="true" pageId="6" pageNumber="569">Simsonia</emphasis>
|
||
Carter & Zeck
|
||
</taxonomicName>
|
||
very marked differences in thoracic and abdominal morphology have been detected (
|
||
<bibRefCitation id="A50A4B69FF9CFFA06733F8C3FE36F8C7" author="Glaister" box="[260,419,1879,1904]" pageId="6" pageNumber="569" refString="Glaister, A. (1999) Guide to the identification of Australian Elmidae larvae (Insecta: Coleoptera). Cooperative Research Centre for Freshwater Ecology, Identification guide No. 21, 49 pp." type="book" year="1999">Glaister 1999</bibRefCitation>
|
||
), and some of the differences found in these Australian genera are similar to those found between
|
||
<taxonomicName id="069B4D1BFF9CFFA066CBF8E9FE19F823" box="[252,396,1916,1940]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9CFFA066CBF8E9FE19F823" box="[252,396,1916,1940]" italics="true" pageId="6" pageNumber="569">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="069B4D1BFF9CFFA067F4F8E9FD00F823" box="[451,661,1916,1940]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9CFFA067F4F8E9FD00F823" box="[451,661,1916,1940]" italics="true" pageId="6" pageNumber="569">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
(e.g. number of pleural sclerites, presence and arrangement of tergal tubercles). Therefore we suggest that
|
||
<taxonomicName id="069B4D1BFF9CFFA0640BF834FD28F80F" box="[572,701,1952,1976]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF9CFFA0640BF834FD28F80F" box="[572,701,1952,1976]" italics="true" pageId="6" pageNumber="569">Luchoelmis</emphasis>
|
||
</taxonomicName>
|
||
could also show a diverse larval morphology.
|
||
</paragraph>
|
||
<paragraph id="C1243698FF9DFFA166F0FF0CFD35FEAB" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="570">
|
||
Larvae of
|
||
<taxonomicName id="069B4D1BFF9DFFA1670DFF0DFD9EFF07" box="[314,523,152,176]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9DFFA1670DFF0DFD9EFF07" box="[314,523,152,176]" italics="true" pageId="7" pageNumber="570">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from those of
|
||
<taxonomicName id="069B4D1BFF9DFFA165ABFF0DFBBEFF07" box="[924,1067,152,176]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9DFFA165ABFF0DFBBEFF07" box="[924,1067,152,176]" italics="true" pageId="7" pageNumber="570">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: 1) pleural sclerites on first five abdominal segments (on first four in
|
||
<taxonomicName id="069B4D1BFF9DFFA16277FF29FB47FF63" box="[1088,1234,188,212]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9DFFA16277FF29FB47FF63" box="[1088,1234,188,212]" italics="true" pageId="7" pageNumber="570">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
); 2) thoracic and abdominal terga with large tubercles arranged in longitudinal rows (absent in
|
||
<taxonomicName id="069B4D1BFF9DFFA1627EFF75FB4AFF4F" box="[1097,1247,224,248]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9DFFA1627EFF75FB4AFF4F" box="[1097,1247,224,248]" italics="true" pageId="7" pageNumber="570">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
); 3) operculum subtriangular (subpentagonal in
|
||
<taxonomicName id="069B4D1BFF9DFFA16437FE91FD05FEAB" box="[512,656,260,284]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="cekalovici">
|
||
<emphasis id="F3EFEA8AFF9DFFA16437FE91FD05FEAB" box="[512,656,260,284]" italics="true" pageId="7" pageNumber="570">L. cekalovici</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="95E46610FF9DFFA166A0F9FCFB20F90B" httpUri="https://zenodo.org/record/226683/files/figure.png" pageId="7" pageNumber="570" targetBox="[296,1260,357,1603]" targetPageId="7">
|
||
<paragraph id="C1243698FF9DFFA166A0F9FCFB20F90B" blockId="7.[151,1436,1640,1724]" pageId="7" pageNumber="570">
|
||
<emphasis id="F3EFEA8AFF9DFFA166A0F9FCFEC0F9C9" bold="true" box="[151,341,1640,1663]" pageId="7" pageNumber="570">FIGURES 17–21.</emphasis>
|
||
Mature larva of
|
||
<taxonomicName id="069B4D1BFF9DFFA16430F9FDFCB2F9C8" box="[519,807,1641,1663]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF9DFFA16430F9FDFCB2F9C8" box="[519,807,1641,1663]" italics="true" pageId="7" pageNumber="570">Luchoelmis kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
: 17) mandible, ventral view; 18) mandible, dorsal view; 19) maxillolabial complex showing right maxilla and labium, ventral view; 20) mesothoracic leg; 21) detail of maxillolabial complex showing distal part of maxillae and labium. Scale bars, Figs 17–19: 0.05 mm; Fig. 20: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="95E46610FF92FFAE66A0F8A3FC62F83C" httpUri="https://zenodo.org/record/226684/files/figure.png" pageId="8" pageNumber="571" targetBox="[273,1343,217,1791]" targetPageId="8">
|
||
<paragraph id="C1243698FF92FFAE66A0F8A3FC62F83C" blockId="8.[151,1436,1847,1931]" pageId="8" pageNumber="571">
|
||
<emphasis id="F3EFEA8AFF92FFAE66A0F8A3FEC3F8FB" bold="true" box="[151,342,1847,1870]" pageId="8" pageNumber="571">FIGURES 22–26.</emphasis>
|
||
Mature larva of
|
||
<taxonomicName id="069B4D1BFF92FFAE6430F8A3FCB2F8FA" box="[519,807,1847,1869]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF92FFAE6430F8A3FCB2F8FA" box="[519,807,1847,1869]" italics="true" pageId="8" pageNumber="571">Luchoelmis kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
: 22) Thorax, ventral view; 23) detail of first five abdominal segments, dorsal view; 24) fourth abdominal segment, lateral view; 25) operculum of ninth abdominal segment, ventral view; 26) hooks of tracheal chamber. Scale bars: Figs 22, 23: 0.5 mm; Figs 24–26: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C1243698FF93FFAF66F0FF0CFBD0FE67" blockId="9.[151,1436,151,464]" pageId="9" pageNumber="572">
|
||
Larvae of
|
||
<taxonomicName id="069B4D1BFF93FFAF670EFF0DFD9FFF07" box="[313,522,152,176]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="kapenkemkensis">
|
||
<emphasis id="F3EFEA8AFF93FFAF670EFF0DFD9FFF07" box="[313,522,152,176]" italics="true" pageId="9" pageNumber="572">L. kapenkemkensis</emphasis>
|
||
</taxonomicName>
|
||
could be mistaken with those of
|
||
<taxonomicName id="069B4D1BFF93FFAF654EFF0CFC7EFF07" box="[889,1003,152,176]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF654EFF0CFC7EFF07" box="[889,1003,152,176]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
due to the presence of pleural sclerites on the first five abdominal segments; actually they would key out as
|
||
<taxonomicName id="069B4D1BFF93FFAF65D8FF28FBF4FF63" box="[1007,1121,188,212]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF65D8FF28FBF4FF63" box="[1007,1121,188,212]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
in the keys of
|
||
<bibRefCitation id="A50A4B69FF93FFAF631DFF29FE10FF4F" author="Manzo" pageId="9" pageNumber="572" refString="Manzo, M. V. & Archangelsky, M. (2008) A key to the known larvae of South American Elmidae (Coleoptera: Byrrhoidea), with a description of the mature larva of Macrelmis saltensis Manzo. Annales de Limnologie, 44 (1), 63 - 74. http: // dx. doi. org / 10.1051 / limn: 2008023" type="journal article" year="2008">Manzo & Archangelsky (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="A50A4B69FF93FFAF678BFF4BFD78FF4F" box="[444,749,223,248]" pageId="9" pageNumber="572" refString="Archangelsky, M., Manzo, M. V., Michat, M. C. & Torres, P. L. M. (2009) Coleoptera. In: Dominguez, E. & Fernandez, H. R. (Eds.), Macroinvertebrados bentonicos sudamericanos. Sistematica y Biologia. Fundacion Miguel Lillo, Tucuman, Argentina, pp. 411 - 468." type="book chapter">
|
||
Archangelsky
|
||
<emphasis id="F3EFEA8AFF93FFAF6468FF75FD02FF4F" box="[607,663,224,248]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
. The following characters will easily separate larvae of these two genera: 1) anterior margin of head capsule lacking tooth between base of antenna and clypeus (present in
|
||
<taxonomicName id="069B4D1BFF93FFAF66A0FEBCFE9CFEF7" box="[151,265,296,320]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF66A0FEBCFE9CFEF7" box="[151,265,296,320]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
); 2) thoracic and abdominal terga with large tubercles arranged in longitudinal rows, bearing long comblike setae (larvae of
|
||
<taxonomicName id="069B4D1BFF93FFAF67B3FED8FE63FED3" box="[388,502,332,356]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF67B3FED8FE63FED3" box="[388,502,332,356]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
only have a row of tubercles bearing flat setae on the margins of the abdominal terga); 3) operculum subtriangular (subpentagonal in
|
||
<taxonomicName id="069B4D1BFF93FFAF6510FEE4FC0CFE3F" box="[807,921,368,392]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF6510FEE4FC0CFE3F" box="[807,921,368,392]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
); 4) color reddish-brown (dark brown in
|
||
<taxonomicName id="069B4D1BFF93FFAF66A0FE00FE9CFE1B" box="[151,265,404,428]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF66A0FE00FE9CFE1B" box="[151,265,404,428]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
); 5) abdominal terga I–VII with complete sagittal line (in
|
||
<taxonomicName id="069B4D1BFF93FFAF65A9FE00FB85FE1B" box="[926,1040,404,428]" class="Insecta" family="Elmidae" genus="Stethelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3EFEA8AFF93FFAF65A9FE00FB85FE1B" box="[926,1040,404,428]" italics="true" pageId="9" pageNumber="572">Stethelmis</emphasis>
|
||
</taxonomicName>
|
||
complete sagittal line present only on abdominal segments I–III, on segments IV–V or IV–VI sagittal line incomplete).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |