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<mods:title id="6CAB67859E3389FA3847B8D98A4AC0FA">Two new genera and two new species of troglobitic harvestmen of Stygnopsidae (Opiliones, Laniatores, Gonyleptoidea) from Oaxaca, Mexico, with notes on selected morphological characters</mods:title>
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<mods:affiliation id="DBA87C23FE5A2E08616BFB69C41F3C67">Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apartado postal 70 - 153, 04510, Ciudad de México (Mexico) and Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Centro Regional de Investigación Pacífico Sur (CIRPAS), Campo Experimental Valles Centrales, Melchor Ocampo No. 7, Santo Domingo Barrio Bajo, Villa de Etla, 68200 Oaxaca (Mexico) thelyphonidito @ gmail. com</mods:affiliation>
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<heading id="EF0B4016FFDA0F129270FAC8FAA33373" box="[884,1383,1012,1039]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="102" reason="2">
Genus 
<taxonomicName id="73FC8CF9FFDA0F1292C5FAC9FAA33373" ID-CoL="7PP7K" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[961,1383,1012,1039]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDA0F1292C5FAC9FBE43372" box="[961,1056,1012,1038]" italics="true" pageId="3" pageNumber="102">Mictlana</emphasis>
Cruz-López &amp; Francke, 2015
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<taxonomicName id="73FC8CF9FFDA0F129229FD10FB15333A" ID-CoL="7PP7K" authority="Cruz-Lopez &amp; Francke, 2015: 878" authorityName="Cruz-Lopez &amp; Francke" authorityPageNumber="878" authorityYear="2015" box="[813,1233,1069,1094]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDA0F129229FD10FC463339" box="[813,898,1069,1093]" italics="true" pageId="3" pageNumber="102">Mictlana</emphasis>
<bibRefCitation id="D06D8A8BFFDA0F129282FD10FB15333A" author="CRUZ- LOPEZ J. A. &amp; FRANCKE O. F." box="[902,1233,1069,1094]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="3" pageNumber="102" pagination="827 - 891" part="175" refId="ref6681" refString="CRUZ- LOPEZ J. A. &amp; FRANCKE O. F. 2015. - Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae). Zoological Journal of the Linnean Society 175: 827 - 891. https: // doi. org / 10.1111 / zoj. 12299" title="Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae)" type="journal article" year="2015">Cruz-López &amp; Francke, 2015: 878</bibRefCitation>
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. — 
<treatmentCitation id="355DD16BFFDA0F1295F2FD10FC08331C" pageId="3" pageNumber="102">
<bibRefCitation id="D06D8A8BFFDA0F1295F2FD10FC08331C" author="CRUZ- LOPEZ J. A." journalOrPublisher="Zootaxa" pageId="3" pageNumber="102" pagination="533 - 548" part="4378" refId="ref6567" refString="CRUZ- LOPEZ J. A. 2018 a. - A new leaf-litter harvestman species of the genus Karos (Opiliones: Stygnopsidae: Karosinae), with a reanalysis of the morphological phylogeny of the genus. Zootaxa 4378: 533 - 548. https: // doi. org / 10.11646 / zootaxa. 4378.4.5" title="A new leaf-litter harvestman species of the genus Karos (Opiliones: Stygnopsidae: Karosinae), with a reanalysis of the morphological phylogeny of the genus" type="journal article" year="2018">Cruz-López 2018a: 534; 2018b: 80</bibRefCitation>
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. — 
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<bibRefCitation id="D06D8A8BFFDA0F1292FDFD75FA8C331C" author="AGUINAGA M. A. &amp; CRUZ- LOPEZ J. A." box="[1017,1352,1096,1120]" journalOrPublisher="Revista Mexicana de Biodiversidad" pageId="3" pageNumber="102" pagination="902907" part="90" refId="ref6509" refString="AGUINAGA M. A. &amp; CRUZ- LOPEZ J. A. 2019. - Dos nuevas especies cavernicolas del genero Serrobunus (Opiliones: Stygnopsidae: Stygnopsinae) del noreste de Mexico. Revista Mexicana de Biodiversidad 90: e 902907. https: // doi. org / 10.22201 / ib. 20078706 e. 2019.90.2907" title="Dos nuevas especies cavernicolas del genero Serrobunus (Opiliones: Stygnopsidae: Stygnopsinae) del noreste de Mexico" type="journal article" year="2019">Aguiñaga &amp; Cruz-López 2019: 9</bibRefCitation>
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. — 
<treatmentCitation id="355DD16BFFDA0F129472FD75FBCF3306" author="CRUZ- LOPEZ J. A. &amp; FRANCKE O. F." page="286" pageId="3" pageNumber="102" year="2019">
<bibRefCitation id="D06D8A8BFFDA0F129472FD75FBCF3306" author="Cruz- Lopez" etAl="et al." firstAuthor="Cruz- Lopez" journalOrPublisher="Journal of Natural History" pageId="3" pageNumber="102" pagination="1451 - 1464" part="53" refId="ref6797" refString="CRUZ- LOPEZ J. A. &amp; FRANCKE O. F. 2019 a. - New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae), from Northern Oaxaca, Mexico, with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus. Journal of Natural History 53: 1451 - 1464. https: // doi. org / 10.1080 / 00222933.2019.1657194" title="New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae), from Northern Oaxaca, Mexico, with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus" type="journal article" year="2019">
Cruz-López 
<emphasis id="86882B68FFDA0F12926AFD5EFC593306" box="[878,925,1122,1146]" italics="true" pageId="3" pageNumber="102">et al.</emphasis>
2019: 286
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.
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<paragraph id="B443F77AFFDA0F129229FDAFFA7433D6" blockId="3.[813,1456,1170,1194]" box="[813,1456,1170,1194]" pageId="3" pageNumber="102">
<typeStatus id="6B4749D8FFDA0F129229FDAFFC9933D4" box="[813,861,1170,1193]" pageId="3" pageNumber="102">TYPE</typeStatus>
SPECIES. — 
<taxonomicName id="73FC8CF9FFDA0F1292D4FDAFFA6833D6" ID-CoL="849F9" authority="Goodnight &amp; Goodnight, 1971" authorityName="Goodnight &amp; Goodnight" authorityYear="1971" box="[976,1452,1170,1194]" class="Arachnida" family="Stygnopsidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="86882B68FFDA0F1292D4FDAFFBB733D6" box="[976,1139,1170,1194]" italics="true" pageId="3" pageNumber="102">Hoplobunus inops</emphasis>
Goodnight &amp; Goodnight, 1971
</taxonomicName>
.
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<paragraph id="B443F77AFFDA0F129229FDFFFA743204" blockId="3.[811,1456,1218,1400]" pageId="3" pageNumber="102">
EMENDED DIAGNOSIS. — Troglobitic karosins, eyeless, with two dorsal lobes on the prosoma, the small one located in the middle and the anterior one larger and rounded. Scutum rectangular, 
<typeStatus id="6B4749D8FFDA0F129439FDCAFAA33273" box="[1341,1383,1271,1295]" pageId="3" pageNumber="102">type</typeStatus>
iota (i) with the lateral clear areas well marked. Chelicera with heterogeneous dentition, movable finger with a basal triangular tooth. Penis with the pars distalis spear-shaped, with two short pairs of MS C and two pairs of MS A, both contiguous and forming a lateral row.
</paragraph>
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<subSubSection id="FCE6A4F1FFDA0F149229FC8EFC323190" lastPageId="5" lastPageNumber="104" pageId="3" pageNumber="102" type="discussion">
<paragraph id="B443F77AFFDA0F129229FC8EFC5532B7" blockId="3.[811,1457,1459,2028]" box="[813,913,1459,1485]" pageId="3" pageNumber="102">REMARKS</paragraph>
<paragraph id="B443F77AFFDA0F159229FCEEFA5A3050" blockId="3.[811,1457,1459,2028]" lastBlockId="4.[813,1457,1778,2028]" lastPageId="4" lastPageNumber="103" pageId="3" pageNumber="102">
Cruz-López &amp; Francke (2015) recovered 
<taxonomicName id="73FC8CF9FFDA0F1295DAFCE9FC2E3171" authority="Goodnight &amp; Goodnight, 1944" authorityName="Goodnight &amp; Goodnight" authorityYear="1944" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDA0F1295DAFCE9FAD33291" box="[1246,1303,1492,1517]" italics="true" pageId="3" pageNumber="102">Karos</emphasis>
Goodnight &amp; Goodnight, 1944
</taxonomicName>
genera-group clade (currently 
<taxonomicName id="73FC8CF9FFDA0F129438FCCEFA6C3172" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[1340,1448,1523,1550]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae</taxonomicName>
) with nine synapomorphies: mesotergal areas not completely covered with tubercles, ocularium in the middle of prosoma, ocularium small, cheliceral dentition homogeneous, no sexual dimorphism on chelicera size, males with chelicera small, follis of penis not on apical depression, pedipalpal femur with a mesodistal setiferous tubercle and pedipalpal patella with mesal armature. At that moment, Cruz-López &amp; Francke (2015) considered that the anterior rounded lobe on prosoma of 
<taxonomicName id="73FC8CF9FFDA0F12924FFE2FFC2D3057" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[843,1001,1810,1836]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="86882B68FFDA0F12924FFE2FFC2D3057" box="[843,1001,1810,1836]" italics="true" pageId="3" pageNumber="102">Mictlana inops</emphasis>
</taxonomicName>
was the ocularium due to the size of the structure, being in this way a reversion from the synapomorphy “ocularium in the middle” to the anterior position on prosoma in this species. Later, Cruz-López &amp; Francke (2017) postulated the possibility that the small lobe on the middle of the prosoma could be the ocularium by comparison of this structure with the true ocularium of other karosins, and then, the anterior lobe would be a developed frontal bulge, structure widely found in other karosins but poorly studied; without evidence of eyes position, this supposition is uncertain. Clear figures of the frontal bulge are available in the description of 
<taxonomicName id="73FC8CF9FFDD0F159018FE4FFDBF30F7" authority="Cruz-Lopez, 2018" authorityName="Cruz-Lopez" authorityYear="2018" box="[284,635,1905,1931]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="morronei">
<emphasis id="86882B68FFDD0F159018FE4FFE7230F7" box="[284,438,1906,1931]" italics="true" pageId="4" pageNumber="103">Karos morronei</emphasis>
Cruz-López, 2018
</taxonomicName>
(Cruz-López 2018a: figs 5-7). Similar problem determining the position of the true ocularium was detected by Cruz-López 
<emphasis id="86882B68FFDD0F159381FE8FFD7C30B7" box="[645,696,1969,1995]" italics="true" pageId="4" pageNumber="103">et al.</emphasis>
(2016) on the troglobitic Belizean 
<taxonomicName id="73FC8CF9FFDD0F1590B0FEECFC313070" authority="(Goodnight &amp; Goodnight, 1977)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1977" box="[436,1013,1778,2027]" class="Arachnida" family="Pyramidopidae" genus="Jarmilana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="86882B68FFDD0F1590B0FEECFD983097" box="[436,604,2001,2027]" italics="true" pageId="4" pageNumber="103">Jarmilana pecki</emphasis>
(Goodnight &amp; Goodnight, 1977)
</taxonomicName>
, a pyramidopid harvestman without eyes or retina evidence, and with two dorsal lobes on prosoma.
</paragraph>
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TABLE 1. — Troglobitic and troglophilic species of 
<taxonomicName id="73FC8CF9FFDD0F15931CF9E6FCD13791" authority="Sorensen, 1932" authorityName="SOrensen" authorityYear="1932" box="[536,789,219,237]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="family">Stygnopsidae Sørensen, 1932</taxonomicName>
, presenting their current subfamilial assignment, troglomorphisms and distribution.
</paragraph>
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<emphasis id="86882B68FFDD0F159182F800FE87362D" bold="true" box="[134,323,317,337]" pageId="4" pageNumber="103">Subfamily/Species</emphasis>
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<emphasis id="86882B68FFDD0F159372F800FC5F362D" bold="true" box="[630,923,317,337]" pageId="4" pageNumber="103">Habitat Troglomorphisms</emphasis>
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<emphasis id="86882B68FFDD0F1595C2F800FABE362D" bold="true" box="[1222,1402,317,337]" pageId="4" pageNumber="103">State and country</emphasis>
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<taxonomicName id="73FC8CF9FFDD0F159182F85CFE3F3609" authority="Cruz-Lopez &amp; Francke, 2017" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[134,507,352,373]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae Cruz-López &amp; Francke, 2017</taxonomicName>
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<tr id="0ACCF538FFDDF0EE9182F843FA6936EE" box="[134,1453,382,402]" gridrow="2" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F843FDAA36EE" box="[134,622,382,402]" gridcol="0" gridrow="2" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F843FDAA36EE" authority="(Goodnight &amp; Goodnight, 1971)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[149,622,382,402]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="86882B68FFDD0F159191F843FEFB36EE" box="[149,319,382,402]" italics="true" pageId="4" pageNumber="103">Huasteca gratiosa</emphasis>
(Goodnight &amp; Goodnight, 1971)
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F843FB7236EE" box="[630,1206,382,402]" gridcol="1" gridrow="2" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2F843FA6936EE" box="[1222,1453,382,402]" gridcol="2" gridrow="2" pageId="4" pageNumber="103">San Luís Potosí, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182F8ABFA6936D7" box="[134,1453,406,427]" gridrow="3" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F8ABFDAA36D7" box="[134,622,406,427]" gridcol="0" gridrow="3" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F8AAFDA036D7" authority="(Goodnight &amp; Goodnight, 1971)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[149,612,406,427]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="86882B68FFDD0F159191F8AAFEF136D7" box="[149,309,407,427]" italics="true" pageId="4" pageNumber="103">Huasteca rugosa</emphasis>
(Goodnight &amp; Goodnight, 1971)
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F8ABFB7236D7" box="[630,1206,406,427]" gridcol="1" gridrow="3" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2F8ABFA6936D7" box="[1222,1453,406,427]" gridcol="2" gridrow="3" pageId="4" pageNumber="103">Veracruz, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182F893FA6936BF" box="[134,1453,430,451]" gridrow="4" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F893FDAA36BF" box="[134,622,430,451]" gridcol="0" gridrow="4" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F892FD8A36BF" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[149,590,431,451]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="86882B68FFDD0F159191F892FEFC36BF" box="[149,312,431,451]" italics="true" pageId="4" pageNumber="103">Huasteca silhavyi</emphasis>
Cruz-López &amp; Francke, 2015
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F893FB7236BF" box="[630,1206,430,451]" gridcol="1" gridrow="4" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2F893FA6936BF" box="[1222,1453,430,451]" gridcol="2" gridrow="4" pageId="4" pageNumber="103">Veracruz, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182F8FAFA6936A7" box="[134,1453,455,475]" gridrow="5" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F8FAFDAA36A7" box="[134,622,455,475]" gridcol="0" gridrow="5" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F8FAFD8736A7" authority="Cruz-Lopez &amp; Francke, 2019" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2019" box="[149,579,455,475]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="kardia">
<emphasis id="86882B68FFDD0F159191F8FAFEE936A7" box="[149,301,455,475]" italics="true" pageId="4" pageNumber="103">Huasteca kardia</emphasis>
Cruz-López &amp; Francke, 2019
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F8FAFB7236A7" box="[630,1206,455,475]" gridcol="1" gridrow="5" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2F8FAFA6936A7" box="[1222,1453,455,475]" gridcol="2" gridrow="5" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182F8E2FA69368F" box="[134,1453,479,499]" gridrow="6" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F8E2FDAA368F" box="[134,622,479,499]" gridcol="0" gridrow="6" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F8E2FDF0368F" authority="Goodnight &amp; Goodnight, 1971" authorityName="Goodnight &amp; Goodnight" authorityYear="1971" box="[149,564,479,499]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="86882B68FFDD0F159191F8E2FED4368F" box="[149,272,479,499]" italics="true" pageId="4" pageNumber="103">Karos parvus</emphasis>
Goodnight &amp; Goodnight, 1971
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F8E2FB72368F" box="[630,1206,479,499]" gridcol="1" gridrow="6" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2F8E2FA69368F" box="[1222,1453,479,499]" gridcol="2" gridrow="6" pageId="4" pageNumber="103">Tamaulipas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182F8CAFA693577" box="[134,1453,503,523]" gridrow="7" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182F8CAFDAA3577" box="[134,622,503,523]" gridcol="0" gridrow="7" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191F8CAFD893577" authority="Goodnight &amp; Goodnight, 1971" authorityName="Goodnight &amp; Goodnight" authorityYear="1971" box="[149,589,503,523]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="projectus">
<emphasis id="86882B68FFDD0F159191F8CAFEED3577" box="[149,297,503,523]" italics="true" pageId="4" pageNumber="103">Karos projectus</emphasis>
Goodnight &amp; Goodnight, 1971
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372F8CAFA693577" box="[630,1453,503,523]" colspan="2" colspanRight="1" gridcol="1" gridrow="7" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented, legs slightly elongated San Luís Potosí, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB32FA693558" box="[134,1453,527,548]" gridrow="8" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FB32FDAA3558" box="[134,622,527,548]" gridcol="0" gridrow="8" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FB2DFDED3558" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[149,553,527,548]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="tersum">
<emphasis id="86882B68FFDD0F159191FB2DFED73558" box="[149,275,528,548]" italics="true" pageId="4" pageNumber="103">Karos tersum</emphasis>
Cruz-López &amp; Francke, 2015
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FB32FB723558" box="[630,1206,527,548]" gridcol="1" gridrow="8" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs slightly elongated,</td>
<td id="491D9C44FFDDF0EE95C2FB32FA693558" box="[1222,1453,527,548]" gridcol="2" gridrow="8" pageId="4" pageNumber="103">San Luís Potosí, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB14FA693541" box="[134,1453,553,573]" gridrow="9" pageId="4" pageNumber="103" rowspan-0="1" rowspan-2="1">
<td id="491D9C44FFDDF0EE9372FB14FB723541" box="[630,1206,553,573]" gridcol="1" gridrow="9" pageId="4" pageNumber="103">eyes reduced</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB7CFA693529" box="[134,1453,577,597]" gridrow="10" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FB7CFDAA3529" box="[134,622,577,597]" gridcol="0" gridrow="10" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FB7CFD8A3529" authority="(Goodnight &amp; Goodnight, 1971)" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[149,590,577,597]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="86882B68FFDD0F159191FB7CFEDB3529" box="[149,287,577,597]" italics="true" pageId="4" pageNumber="103">Mictlana inops</emphasis>
(Goodnight &amp; Goodnight, 1971)
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FB7CFB723529" box="[630,1206,577,597]" gridcol="1" gridrow="10" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, high tarsal</td>
<td id="491D9C44FFDDF0EE95C2FB7CFA693529" box="[1222,1453,577,597]" gridcol="2" gridrow="10" pageId="4" pageNumber="103">Tamaulipas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB66FA693513" box="[134,1453,603,623]" gridrow="11" pageId="4" pageNumber="103" rowspan-0="1" rowspan-2="1">
<td id="491D9C44FFDDF0EE9372FB66FB723513" box="[630,1206,603,623]" gridcol="1" gridrow="11" pageId="4" pageNumber="103">count, eyes absent</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB4EFA6935FB" box="[134,1453,627,647]" gridrow="12" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FB4EFDAA35FB" box="[134,622,627,647]" gridcol="0" gridrow="12" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FB4EFD8A35FB" authority="(Goodnight &amp; Goodnight, 1973)" authorityName="Cruz-López" authorityYear="2021" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1973" box="[149,590,627,647]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="86882B68FFDD0F159191FB4EFEDA35FB" box="[149,286,627,647]" italics="true" pageId="4" pageNumber="103">Mictlana plana</emphasis>
(
<bibRefCitation id="D06D8A8BFFDD0F15902DFB4EFD8D35FB" author="Goodnight &amp; Goodnight" box="[297,585,627,647]" firstAuthor="Goodnight" pageId="4" pageNumber="103" pagination="83 - 96" refId="ref7174" refString="GOODNIGHT C. J. &amp; GOODNIGHT M. L. 1973. - Opilionids (Phalangida) from Mexican caves. Bulletin of the Association for Mexican Cave Studies 5: 83 - 96." type="journal article" year="1973">Goodnight &amp; Goodnight, 1973</bibRefCitation>
)
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FB4EFB7235FB" box="[630,1206,627,647]" gridcol="1" gridrow="12" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, high tarsal</td>
<td id="491D9C44FFDDF0EE95C2FB4EFA6935FB" box="[1222,1453,627,647]" gridcol="2" gridrow="12" pageId="4" pageNumber="103">San Luís Potosí, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FBB6FA6935DC" box="[134,1453,651,672]" gridrow="13" pageId="4" pageNumber="103" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FBB6FDAA35DC" box="[134,622,651,672]" gridcol="0" gridrow="13" pageId="4" pageNumber="103">n. comb.</th>
<td id="491D9C44FFDDF0EE9372FBB6FB7235DC" box="[630,1206,651,672]" gridcol="1" gridrow="13" pageId="4" pageNumber="103">count, eyes absent</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB99FA6935C5" box="[134,1453,676,697]" gridrow="14" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FB99FDAA35C5" box="[134,622,676,697]" gridcol="0" gridrow="14" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FB98FDFD35C5" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[149,569,676,697]" class="Insecta" family="Pyralidae" genus="Potosa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="reddelli">
<emphasis id="86882B68FFDD0F159191FB98FEE635C5" box="[149,290,677,697]" italics="true" pageId="4" pageNumber="103">Potosa reddelli</emphasis>
Cruz-López &amp; Francke, 2015
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FB99FB7235C5" box="[630,1206,676,697]" gridcol="1" gridrow="14" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2FB99FA6935C5" box="[1222,1453,676,697]" gridcol="2" gridrow="14" pageId="4" pageNumber="103">Hidalgo, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FB81FA6935AD" box="[134,1453,700,721]" gridrow="15" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FB81FDAA35AD" box="[134,622,700,721]" gridcol="0" gridrow="15" pageId="4" pageNumber="103">
<emphasis id="86882B68FFDD0F159191FB80FEFB35AD" box="[149,319,701,721]" italics="true" pageId="4" pageNumber="103">‘Karos’ depressus</emphasis>
Goodnight &amp; Goodnight, 1971
</th>
<td id="491D9C44FFDDF0EE9372FB81FB7235AD" box="[630,1206,700,721]" gridcol="1" gridrow="15" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, eyes reduced</td>
<td id="491D9C44FFDDF0EE95C2FB81FA6935AD" box="[1222,1453,700,721]" gridcol="2" gridrow="15" pageId="4" pageNumber="103">Querétaro, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FBDAFA693580" box="[134,1453,743,764]" gridrow="16" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FBDAFA693580" box="[134,1453,743,764]" colspan="3" colspanRight="2" gridcol="0" gridrow="16" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159182FBDAFE673580" authority="Sorensen, 1932" authorityName="Sorensen" authorityYear="1932" box="[134,419,743,764]" pageId="4" pageNumber="106" rank="subFamily" subFamily="Stygnopsinae">Stygnopsinae Sørensen, 1932</taxonomicName>
</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA38FA693465" box="[134,1453,773,793]" gridrow="17" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FA38FDAA3465" box="[134,622,773,793]" gridcol="0" gridrow="17" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FA38FD9E3465" authority="Cokendolpher" box="[149,602,773,793]" class="Arachnida" family="Stygnopsidae" genus="Chinquipellobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="coahuilaensis">
<emphasis id="86882B68FFDD0F159191FA38FE083465" box="[149,460,773,793]" italics="true" pageId="4" pageNumber="103">Chinquipellobunus coahuilaensis</emphasis>
Cokendolpher
</taxonomicName>
,
</th>
<td id="491D9C44FFDDF0EE9372FA38FB723465" box="[630,1206,773,793]" gridcol="1" gridrow="17" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2FA38FA693465" box="[1222,1453,773,793]" gridcol="2" gridrow="17" pageId="4" pageNumber="103">Coahuila, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA20FA69344D" box="[134,1453,797,817]" gridrow="18" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FA20FDAA344D" box="[134,622,797,817]" gridcol="0" gridrow="18" pageId="4" pageNumber="103">2004</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA08FA693436" box="[134,1453,821,842]" gridrow="19" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FA08FDAA3436" box="[134,622,821,842]" gridcol="0" gridrow="19" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FA08FE553436" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1967" box="[149,401,821,842]" class="Arachnida" family="Stygnopsidae" genus="Chinquipellobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="madlae">
<emphasis id="86882B68FFDD0F159191FA08FE553436" box="[149,401,821,842]" italics="true" pageId="4" pageNumber="103">Chinquipellobunus madlae</emphasis>
</taxonomicName>
(Goodnight &amp;
</th>
<td id="491D9C44FFDDF0EE9372FA08FB723436" box="[630,1206,821,842]" gridcol="1" gridrow="19" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, eyes absent</td>
<td id="491D9C44FFDDF0EE95C2FA08FA693436" box="[1222,1453,821,842]" gridcol="2" gridrow="19" pageId="4" pageNumber="103">Texas, U.S.A.</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA70FA69341E" box="[134,1453,845,866]" gridrow="20" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FA70FDAA341E" box="[134,622,845,866]" gridcol="0" gridrow="20" pageId="4" pageNumber="103">Goodnight, 1967)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA58FA693406" box="[134,1453,869,890]" gridrow="21" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FA58FDAA3406" box="[134,622,869,890]" gridcol="0" gridrow="21" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FA5BFE753406" box="[149,433,870,890]" class="Arachnida" family="Stygnopsidae" genus="Chinquipellobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="86882B68FFDD0F159191FA5BFE753406" box="[149,433,870,890]" italics="true" pageId="4" pageNumber="103">Chinquipellobunus mexicanus</emphasis>
</taxonomicName>
(Goodnight &amp;
</th>
<td id="491D9C44FFDDF0EE9372FA58FB723406" box="[630,1206,869,890]" gridcol="1" gridrow="21" pageId="4" pageNumber="103">Data not available</td>
<td id="491D9C44FFDDF0EE95C2FA58FA693406" box="[1222,1453,869,890]" gridcol="2" gridrow="21" pageId="4" pageNumber="103">Nuevo León, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA43FA6934EE" box="[134,1453,894,914]" gridrow="22" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FA43FDAA34EE" box="[134,622,894,914]" gridcol="0" gridrow="22" pageId="4" pageNumber="103">Goodnight, 1971)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FAABFA6934D6" box="[134,1453,918,938]" gridrow="23" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FAABFDAA34D6" box="[134,622,918,938]" gridcol="0" gridrow="23" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FAABFDCE34D6" authority="Goodnight &amp;" box="[149,522,918,938]" class="Arachnida" family="Stygnopsidae" genus="Chinquipellobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="osorioi">
<emphasis id="86882B68FFDD0F159191FAABFE4834D6" box="[149,396,918,938]" italics="true" pageId="4" pageNumber="103">Chinquipellobunus osorioi</emphasis>
Goodnight &amp;
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FAABFB7234D6" box="[630,1206,918,938]" gridcol="1" gridrow="23" pageId="4" pageNumber="103">Data not available</td>
<td id="491D9C44FFDDF0EE95C2FAABFA6934D6" box="[1222,1453,918,938]" gridcol="2" gridrow="23" pageId="4" pageNumber="103">Nuevo León, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FA93FA6934BE" box="[134,1453,942,962]" gridrow="24" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FA93FDAA34BE" box="[134,622,942,962]" gridcol="0" gridrow="24" pageId="4" pageNumber="103">Goodnight, 1944</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FAFBFA6934A6" box="[134,1453,966,986]" gridrow="25" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FAFBFDAA34A6" box="[134,622,966,986]" gridcol="0" gridrow="25" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FAFBFE4934A6" baseAuthorityName="C.J.Goodnight &amp; M.L.Goodnight" baseAuthorityYear="1967" box="[149,397,966,986]" class="Arachnida" family="Stygnopsidae" genus="Chinquipellobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="russelli">
<emphasis id="86882B68FFDD0F159191FAFBFE4934A6" box="[149,397,966,986]" italics="true" pageId="4" pageNumber="103">Chinquipellobunus russelli</emphasis>
</taxonomicName>
(Goodnight &amp;
</th>
<td id="491D9C44FFDDF0EE9372FAFBFB7234A6" box="[630,1206,966,986]" gridcol="1" gridrow="25" pageId="4" pageNumber="103">Data not available</td>
<td id="491D9C44FFDDF0EE95C2FAFBFA6934A6" box="[1222,1453,966,986]" gridcol="2" gridrow="25" pageId="4" pageNumber="103">Texas, U.S.A.</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FAE3FA69348E" box="[134,1453,990,1010]" gridrow="26" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FAE3FDAA348E" box="[134,622,990,1010]" gridcol="0" gridrow="26" pageId="4" pageNumber="103">Goodnight, 1967)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FACBFA693376" box="[134,1453,1014,1034]" gridrow="27" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FACBFDAA3376" box="[134,622,1014,1034]" gridcol="0" gridrow="27" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FACBFDF23376" authority="Cruz-Lopez &amp; Francke, 2017" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[149,566,1014,1034]" class="Arachnida" family="Stygnopsidae" genus="Iztlina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="venefica">
<emphasis id="86882B68FFDD0F159191FACBFEE43376" box="[149,288,1014,1034]" italics="true" pageId="4" pageNumber="103">Iztlina venefica</emphasis>
Cruz-López &amp; Francke, 2017
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FACBFB723376" box="[630,1206,1014,1034]" gridcol="1" gridrow="27" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated, high</td>
<td id="491D9C44FFDDF0EE95C2FACBFA693376" box="[1222,1453,1014,1034]" gridcol="2" gridrow="27" pageId="4" pageNumber="103">Chiapas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD2DFA693358" box="[134,1453,1040,1060]" gridrow="28" pageId="4" pageNumber="103" rowspan-0="1" rowspan-2="1">
<td id="491D9C44FFDDF0EE9372FD2DFB723358" box="[630,1206,1040,1060]" gridcol="1" gridrow="28" pageId="4" pageNumber="103">tarsal count</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD18FA693340" box="[134,1453,1061,1084]" gridrow="29" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FD18FDAA3340" box="[134,622,1061,1084]" gridcol="0" gridrow="29" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FD15FE3A3340" authority="Silhavy, 1977" authorityName="Silhavy" authorityYear="1977" box="[149,510,1061,1084]" class="Arachnida" family="Stygnopsidae" genus="Mexotroglinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="sbordonii">
<emphasis id="86882B68FFDD0F159191FD15FEBD3340" box="[149,377,1064,1084]" italics="true" pageId="4" pageNumber="103">Mexotroglinus sbordonii</emphasis>
Šilhavý, 1977
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FD18FB723340" box="[630,1206,1061,1084]" gridcol="1" gridrow="29" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, eyes absent</td>
<td id="491D9C44FFDDF0EE95C2FD18FA693340" box="[1222,1453,1061,1084]" gridcol="2" gridrow="29" pageId="4" pageNumber="103">Chiapas, México</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD7DFA693328" box="[134,1453,1088,1108]" gridrow="30" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FD7DFDAA3328" box="[134,622,1088,1108]" gridcol="0" gridrow="30" pageId="4" pageNumber="103">
<emphasis id="86882B68FFDD0F159191FD7DFEE73328" box="[149,291,1088,1108]" italics="true" pageId="4" pageNumber="103">Minisge kanoni</emphasis>
Cruz-López, Monjaraz-Ruedas &amp;
</th>
<td id="491D9C44FFDDF0EE9372FD7DFB723328" box="[630,1206,1088,1108]" gridcol="1" gridrow="30" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, high tarsal</td>
<td id="491D9C44FFDDF0EE95C2FD7DFA693328" box="[1222,1453,1088,1108]" gridcol="2" gridrow="30" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD65FA693312" box="[134,1453,1112,1134]" gridrow="31" pageId="4" pageNumber="103" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FD65FDAA3312" box="[134,622,1112,1134]" gridcol="0" gridrow="31" pageId="4" pageNumber="103">Francke, 2019</th>
<td id="491D9C44FFDDF0EE9372FD65FB723312" box="[630,1206,1112,1134]" gridcol="1" gridrow="31" pageId="4" pageNumber="103">count, eyes reduced</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD4FFA6933FA" box="[134,1453,1138,1158]" gridrow="32" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FD4FFDAA33FA" box="[134,622,1138,1158]" gridcol="0" gridrow="32" pageId="4" pageNumber="103">
<emphasis id="86882B68FFDD0F159191FD4FFED233FA" box="[149,278,1138,1158]" italics="true" pageId="4" pageNumber="103">Minisge sagai</emphasis>
Cruz-López, Monjaraz-Ruedas &amp;
</th>
<td id="491D9C44FFDDF0EE9372FD4FFB7233FA" box="[630,1206,1138,1158]" gridcol="1" gridrow="32" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, high tarsal</td>
<td id="491D9C44FFDDF0EE95C2FD4FFA6933FA" box="[1222,1453,1138,1158]" gridcol="2" gridrow="32" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FDB7FA6933E3" box="[134,1453,1162,1183]" gridrow="33" pageId="4" pageNumber="103" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FDB7FDAA33E3" box="[134,622,1162,1183]" gridcol="0" gridrow="33" pageId="4" pageNumber="103">Francke, 2019</th>
<td id="491D9C44FFDDF0EE9372FDB7FB7233E3" box="[630,1206,1162,1183]" gridcol="1" gridrow="33" pageId="4" pageNumber="103">count, eyes reduced</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD9EFA6933C4" box="[134,1453,1187,1208]" gridrow="34" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FD9EFDAA33C4" box="[134,622,1187,1208]" gridcol="0" gridrow="34" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FD99FDD533C4" authority="Cruz-Lopez &amp; Francke, 2016" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2016" box="[149,529,1187,1208]" class="Arachnida" family="Stygnopsidae" genus="Philora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="izel">
<emphasis id="86882B68FFDD0F159191FD99FF3F33C4" box="[149,251,1188,1208]" italics="true" pageId="4" pageNumber="103">Philora izel</emphasis>
Cruz-López &amp; Francke, 2016
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FD9EFB7233C4" box="[630,1206,1187,1208]" gridcol="1" gridrow="34" pageId="4" pageNumber="103">Troglophilic?Cuticle depigmented, eyes reduced</td>
<td id="491D9C44FFDDF0EE95C2FD9EFA6933C4" box="[1222,1453,1187,1208]" gridcol="2" gridrow="34" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FD86FA6933AC" box="[134,1453,1211,1232]" gridrow="35" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FD86FDAA33AC" box="[134,622,1211,1232]" gridcol="0" gridrow="35" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FD81FDFA33AC" authority="Cruz-Lopez &amp; Francke, 2016" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2016" box="[149,574,1212,1232]" class="Arachnida" family="Stygnopsidae" genus="Philora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="nympha">
<emphasis id="86882B68FFDD0F159191FD81FEEC33AC" box="[149,296,1212,1232]" italics="true" pageId="4" pageNumber="103">Philora nympha</emphasis>
Cruz-López &amp; Francke, 2016
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FD86FB7233AC" box="[630,1206,1211,1232]" gridcol="1" gridrow="35" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FD86FA6933AC" box="[1222,1453,1211,1232]" gridcol="2" gridrow="35" pageId="4" pageNumber="103">Puebla, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FDECFA693394" box="[134,1453,1233,1256]" gridrow="36" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FDECFDAA3394" box="[134,622,1233,1256]" gridcol="0" gridrow="36" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FDE9FE113394" authority="Silhavy, 1977" authorityName="Silhavy" authorityYear="1977" box="[149,469,1233,1256]" class="Arachnida" family="Stygnopsidae" genus="Sbordonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="86882B68FFDD0F159191FDE9FE943394" box="[149,336,1236,1256]" italics="true" pageId="4" pageNumber="103">Sbordonia armigera</emphasis>
Šilhavý, 1977
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FDECFB723394" box="[630,1206,1233,1256]" gridcol="1" gridrow="36" pageId="4" pageNumber="103">Troglophilic?</td>
<td id="491D9C44FFDDF0EE95C2FDECFA693394" box="[1222,1453,1233,1256]" gridcol="2" gridrow="36" pageId="4" pageNumber="103">Chiapas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FDC4FA693271" box="[134,1453,1273,1293]" gridrow="37" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FDC4FDAA3271" box="[134,622,1273,1293]" gridcol="0" gridrow="37" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FDC4FDAC3271" authority="Goodnight &amp; Goodnight, 1942" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[149,616,1273,1293]" class="Arachnida" family="Phalangodidae" genus="Serrobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="86882B68FFDD0F159191FDC4FE813271" box="[149,325,1273,1293]" italics="true" pageId="4" pageNumber="103">Serrobunus boneti</emphasis>
Goodnight &amp; Goodnight, 1942
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FDC4FB723271" box="[630,1206,1273,1293]" gridcol="1" gridrow="37" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FDC4FA693271" box="[1222,1453,1273,1293]" gridcol="2" gridrow="37" pageId="4" pageNumber="103">San Luís Potosí, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FC21FA69324F" box="[134,1453,1308,1331]" gridrow="38" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FC21FDAA324F" box="[134,622,1308,1331]" gridcol="0" gridrow="38" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FC23FE33324E" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[149,503,1308,1331]" class="Arachnida" family="Phalangodidae" genus="Serrobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="queretarius">
<emphasis id="86882B68FFDD0F159191FC23FEB7324E" box="[149,371,1310,1330]" italics="true" pageId="4" pageNumber="103">Serrobunus queretarius</emphasis>
<bibRefCitation id="D06D8A8BFFDD0F15907DFC21FE33324E" author="Silhavy" box="[377,503,1308,1331]" firstAuthor="Silhavy" pageId="4" pageNumber="103" pagination="175 - 194" refId="ref7589" refString="SILHAVY V. 1974. - Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FC21FB72324F" box="[630,1206,1308,1331]" gridcol="1" gridrow="38" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FC21FA69324F" box="[1222,1453,1308,1331]" gridcol="2" gridrow="38" pageId="4" pageNumber="103">Querétaro, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FC7EFA693224" box="[134,1453,1347,1368]" gridrow="39" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FC7EFDAA3224" box="[134,622,1347,1368]" gridcol="0" gridrow="39" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FC7EFDAD3224" authority="Aguinaga &amp; Cruz-Lopez, 2019" authorityName="Aguinaga &amp; Cruz-Lopez" authorityYear="2019" box="[149,617,1347,1368]" class="Arachnida" family="Phalangodidae" genus="Serrobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="linares">
<emphasis id="86882B68FFDD0F159191FC7EFE823224" box="[149,326,1347,1368]" italics="true" pageId="4" pageNumber="103">Serrobunus linares</emphasis>
Aguiñaga &amp; Cruz-López, 2019
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FC7EFB723224" box="[630,1206,1347,1368]" gridcol="1" gridrow="39" pageId="4" pageNumber="103">Troglophilic Cuticle slightly pigmented</td>
<td id="491D9C44FFDDF0EE95C2FC7EFA693224" box="[1222,1453,1347,1368]" gridcol="2" gridrow="39" pageId="4" pageNumber="103">Nuevo León, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FC54FA693201" box="[134,1453,1385,1405]" gridrow="40" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FC54FDAA3201" box="[134,622,1385,1405]" gridcol="0" gridrow="40" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FC54FD993201" authority="Aguinaga &amp; Cruz-Lopez" box="[149,605,1385,1405]" class="Arachnida" family="Phalangodidae" genus="Serrobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="paulbryanti">
<emphasis id="86882B68FFDD0F159191FC54FEB63201" box="[149,370,1385,1405]" italics="true" pageId="4" pageNumber="103">Serrobunus paulbryanti</emphasis>
Aguiñaga &amp; Cruz-López
</taxonomicName>
,
</th>
<td id="491D9C44FFDDF0EE9372FC54FB723201" box="[630,1206,1385,1405]" gridcol="1" gridrow="40" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, eyes</td>
<td id="491D9C44FFDDF0EE95C2FC54FA693201" box="[1222,1453,1385,1405]" gridcol="2" gridrow="40" pageId="4" pageNumber="103">Tamaulipas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FCBCFA6932EA" box="[134,1453,1409,1430]" gridrow="41" pageId="4" pageNumber="103" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FCBCFDAA32EA" box="[134,622,1409,1430]" gridcol="0" gridrow="41" pageId="4" pageNumber="103">2019</th>
<td id="491D9C44FFDDF0EE9372FCBCFB7232EA" box="[630,1206,1409,1430]" gridcol="1" gridrow="41" pageId="4" pageNumber="103">absent</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FCA7FA6932D3" box="[134,1453,1434,1455]" gridrow="42" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FCA7FDAA32D3" box="[134,622,1434,1455]" gridcol="0" gridrow="42" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FCA7FEAC32D3" box="[149,360,1434,1455]" class="Arachnida" family="Stygnopsidae" genus="Stygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="apoalensis">
<emphasis id="86882B68FFDD0F159191FCA7FEAC32D3" box="[149,360,1434,1455]" italics="true" pageId="4" pageNumber="103">Stygnopsis apoalensis</emphasis>
</taxonomicName>
(Goodnight &amp; Goodnight,
</th>
<td id="491D9C44FFDDF0EE9372FCA7FB7232D3" box="[630,1206,1434,1455]" gridcol="1" gridrow="42" pageId="4" pageNumber="103">Troglophilic Legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FCA7FA6932D3" box="[1222,1453,1434,1455]" gridcol="2" gridrow="42" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FC8EFA6932BB" box="[134,1453,1459,1479]" gridrow="43" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FC8EFDAA32BB" box="[134,622,1459,1479]" gridcol="0" gridrow="43" pageId="4" pageNumber="103">1973)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FCF7FA6932A3" box="[134,1453,1482,1503]" gridrow="44" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FCF7FDAA32A3" box="[134,622,1482,1503]" gridcol="0" gridrow="44" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FCF6FEA932A3" box="[149,365,1483,1503]" class="Arachnida" family="Stygnopsidae" genus="Stygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="oaxacensis">
<emphasis id="86882B68FFDD0F159191FCF6FEA932A3" box="[149,365,1483,1503]" italics="true" pageId="4" pageNumber="103">Stygnopsis oaxacensis</emphasis>
</taxonomicName>
(Goodnight &amp; Goodnight,
</th>
<td id="491D9C44FFDDF0EE9372FCF7FB7232A3" box="[630,1206,1482,1503]" gridcol="1" gridrow="44" pageId="4" pageNumber="103">Troglophilic Legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FCF7FA6932A3" box="[1222,1453,1482,1503]" gridcol="2" gridrow="44" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FCDEFA69328B" box="[134,1453,1507,1527]" gridrow="45" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FCDEFDAA328B" box="[134,622,1507,1527]" gridcol="0" gridrow="45" pageId="4" pageNumber="103">1973)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FCC6FA693173" box="[134,1453,1531,1551]" gridrow="46" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FCC6FDAA3173" box="[134,622,1531,1551]" gridcol="0" gridrow="46" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FCC6FE883173" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[149,332,1531,1551]" class="Arachnida" family="Stygnopsidae" genus="Stygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="86882B68FFDD0F159191FCC6FE883173" box="[149,332,1531,1551]" italics="true" pageId="4" pageNumber="103">Stygnopsis robusta</emphasis>
</taxonomicName>
(Goodnight &amp; Goodnight,
</th>
<td id="491D9C44FFDDF0EE9372FCC6FB723173" box="[630,1206,1531,1551]" gridcol="1" gridrow="46" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FCC6FA693173" box="[1222,1453,1531,1551]" gridcol="2" gridrow="46" pageId="4" pageNumber="103">Veracruz, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FF2EFA69315B" box="[134,1453,1555,1575]" gridrow="47" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FF2EFDAA315B" box="[134,622,1555,1575]" gridcol="0" gridrow="47" pageId="4" pageNumber="103">1971)</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FF16FA693143" box="[134,1453,1579,1599]" gridrow="48" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FF16FDAA3143" box="[134,622,1579,1599]" gridcol="0" gridrow="48" pageId="4" pageNumber="103">
<emphasis id="86882B68FFDD0F159191FF16FE463143" box="[149,386,1579,1599]" italics="true" pageId="4" pageNumber="103">Tonalteca spinooculorum</emphasis>
(Goodnight &amp;
</th>
<td id="491D9C44FFDDF0EE9372FF16FB723143" box="[630,1206,1579,1599]" gridcol="1" gridrow="48" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FF16FA693143" box="[1222,1453,1579,1599]" gridcol="2" gridrow="48" pageId="4" pageNumber="103">Oaxaca, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FF7EFA69312B" box="[134,1453,1603,1623]" gridrow="49" pageId="4" pageNumber="103" rowspan-1="1" rowspan-2="1">
<th id="491D9C44FFDDF0EE9182FF7EFDAA312B" box="[134,622,1603,1623]" gridcol="0" gridrow="49" pageId="4" pageNumber="103">
<bibRefCitation id="D06D8A8BFFDD0F1591A0FF7EFE81312B" author="Goodnight" box="[164,325,1603,1623]" firstAuthor="Goodnight" pageId="4" pageNumber="103" pagination="83 - 96" refId="ref7174" refString="GOODNIGHT C. J. &amp; GOODNIGHT M. L. 1973. - Opilionids (Phalangida) from Mexican caves. Bulletin of the Association for Mexican Cave Studies 5: 83 - 96." type="journal article" year="1973">Goodnight, 1973</bibRefCitation>
)
</th>
</tr>
<tr id="0ACCF538FFDDF0EE9182FF64FA69310C" box="[134,1453,1625,1648]" gridrow="50" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FF64FDAA310C" box="[134,622,1625,1648]" gridcol="0" gridrow="50" pageId="4" pageNumber="103">
<taxonomicName id="73FC8CF9FFDD0F159191FF66FDFF3113" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[149,571,1625,1648]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="86882B68FFDD0F159191FF66FE723113" box="[149,438,1627,1647]" italics="true" pageId="4" pageNumber="103">Troglostygnopsis anophthalma</emphasis>
<bibRefCitation id="D06D8A8BFFDD0F1590B8FF64FDFF3113" author="Silhavy" box="[444,571,1625,1648]" firstAuthor="Silhavy" pageId="4" pageNumber="103" pagination="175 - 194" refId="ref7589" refString="SILHAVY V. 1974. - Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
</th>
<td id="491D9C44FFDDF0EE9372FF64FB72310C" box="[630,1206,1625,1648]" gridcol="1" gridrow="50" pageId="4" pageNumber="103">Troglobitic Cuticle depigmented, legs elongated, high tarsal</td>
<td id="491D9C44FFDDF0EE95C2FF64FA69310C" box="[1222,1453,1625,1648]" gridcol="2" gridrow="50" pageId="4" pageNumber="103">Chiapas, Mexico</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FF48FA6931F5" box="[134,1453,1653,1673]" gridrow="51" pageId="4" pageNumber="103" rowspan-0="1" rowspan-2="1">
<td id="491D9C44FFDDF0EE9372FF48FB7231F5" box="[630,1206,1653,1673]" gridcol="1" gridrow="51" pageId="4" pageNumber="103">count, eyes absent</td>
</tr>
<tr id="0ACCF538FFDDF0EE9182FFAFFA6931D5" box="[134,1453,1682,1705]" gridrow="52" pageId="4" pageNumber="103">
<th id="491D9C44FFDDF0EE9182FFAFFDAA31D5" box="[134,622,1682,1705]" gridcol="0" gridrow="52" pageId="4" pageNumber="103">
<emphasis id="86882B68FFDD0F159191FFA9FE9031D4" box="[149,340,1684,1704]" italics="true" pageId="4" pageNumber="103">‘Hoplobunus’ zullinii</emphasis>
Šilhavý, 1977
</th>
<td id="491D9C44FFDDF0EE9372FFAFFB7231D5" box="[630,1206,1682,1705]" gridcol="1" gridrow="52" pageId="4" pageNumber="103">Troglobitic Cuticle slightly pigmented, legs elongated</td>
<td id="491D9C44FFDDF0EE95C2FFAFFA6931D5" box="[1222,1453,1682,1705]" gridcol="2" gridrow="52" pageId="4" pageNumber="103">Chiapas, Mexico</td>
</tr>
</table>
</paragraph>
<paragraph id="B443F77AFFDD0F149247FE0CFD6130D0" blockId="4.[813,1457,1778,2028]" lastBlockId="5.[130,776,1554,2028]" lastPageId="5" lastPageNumber="104" pageId="4" pageNumber="103">
The character “Chelicera with homogenous dentition”, another synapomorphy of 
<taxonomicName id="73FC8CF9FFDD0F159554FE6CFB793017" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[1104,1213,1873,1899]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae</taxonomicName>
, was recovered as a reversion to the plesiomorphic state, heterogeneous dentition, in 
<taxonomicName id="73FC8CF9FFDD0F15924EFEAFFC6330D7" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[842,935,1938,1963]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="86882B68FFDD0F15924EFEAFFC6330D7" box="[842,935,1938,1963]" italics="true" pageId="4" pageNumber="103">M. inops</emphasis>
</taxonomicName>
in Cruz-López &amp; Francke (2015). In a broader phylogeny of 
<taxonomicName id="73FC8CF9FFDD0F1592C7FE8CFB9030B7" authorityName="SOrensen" authorityYear="1932" box="[963,1108,1969,1995]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="103" phylum="Arthropoda" rank="family">Stygnopsidae</taxonomicName>
, cheliceral dentition is a homoplastic character through the family, causing conflict to a subfamilial allocation of 
<taxonomicName id="73FC8CF9FFDC0F149092FF2FFE323150" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[406,502,1554,1580]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDC0F149092FF2FFE323150" box="[406,502,1554,1580]" italics="true" pageId="5" pageNumber="104">Mictlana</emphasis>
</taxonomicName>
and 
<taxonomicName id="73FC8CF9FFDC0F14932BFF2FFD063150" authorityName="Silhavy" authorityYear="1977" box="[559,706,1554,1580]" class="Arachnida" family="Stygnopsidae" genus="Mexotroglinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDC0F14932BFF2FFD063150" box="[559,706,1554,1580]" italics="true" pageId="5" pageNumber="104">Mexotroglinus</emphasis>
</taxonomicName>
based only on this character. According to Cruz-López &amp; Francke (2017), the character describing the shape of the mesotergal sulci has two states evenly distributed between 
<taxonomicName id="73FC8CF9FFDC0F14937DFF4FFCC331F0" box="[633,775,1650,1676]" pageId="5" pageNumber="106" rank="subFamily" subFamily="Stygnopsinae">Stygnopsinae</taxonomicName>
and 
<taxonomicName id="73FC8CF9FFDC0F1491B1FFAFFEE731D0" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[181,291,1682,1708]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae</taxonomicName>
. For that, in the present work, the character “mesotergal sulci sinuous” is considered a putative synapomorphy of 
<taxonomicName id="73FC8CF9FFDC0F149006FFEFFEB43190" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[258,368,1746,1772]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae</taxonomicName>
, character clearly illustrated by SEM photographs in Cruz-López &amp; Francke (2017, 2019a) and observed on the 
<typeStatus id="6B4749D8FFDC0F14903EFE2FFE5D3050" box="[314,409,1810,1836]" pageId="5" pageNumber="104" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="73FC8CF9FFDC0F1490BAFE2FFDE23050" authorityName="Goodnight &amp; Goodnight" authorityYear="1973" box="[446,550,1810,1836]" class="Arachnida" family="Stygnopsidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="86882B68FFDC0F1490BAFE2FFDE23050" box="[446,550,1810,1836]" italics="true" pageId="5" pageNumber="104">H. planus</emphasis>
</taxonomicName>
. Additionally, 
<bibRefCitation id="D06D8A8BFFDC0F1493CCFE2FFE5C3030" author="Cruz- Lopez &amp; Francke" firstAuthor="Cruz- Lopez" journalOrPublisher="Journal of Natural History" pageId="5" pageNumber="104" pagination="1451 - 1464" part="53" refId="ref6797" refString="CRUZ- LOPEZ J. A. &amp; FRANCKE O. F. 2019 a. - New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae), from Northern Oaxaca, Mexico, with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus. Journal of Natural History 53: 1451 - 1464. https: // doi. org / 10.1080 / 00222933.2019.1657194" title="New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae), from Northern Oaxaca, Mexico, with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus" type="journal article" year="2019" yearSuffix="a">Cruz-López &amp; Francke (2019a)</bibRefCitation>
detected the presence of cheliceral comb in several 
<taxonomicName id="73FC8CF9FFDC0F14902FFE6CFE513017" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2017" box="[299,405,1873,1899]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Karosinae">Karosinae</taxonomicName>
representatives, structure observed on the 
<typeStatus id="6B4749D8FFDC0F1491CFFE4CFEED30F7" box="[203,297,1905,1931]" pageId="5" pageNumber="104" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="73FC8CF9FFDC0F14904EFE4FFE6C30F7" authorityName="Cruz-López" authorityYear="2021" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1973" box="[330,424,1905,1931]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="plana" status="comb. nov.">
<emphasis id="86882B68FFDC0F14904EFE4FFE6C30F7" box="[330,424,1905,1931]" italics="true" pageId="5" pageNumber="104">M. plana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DBB9613FFDC0F1490ABFE4FFDD430F7" box="[431,528,1905,1932]" pageId="5" pageNumber="104" rank="species">n. comb.</taxonomicNameLabel>
, but unfortunately that could not be checked in any specimen of 
<taxonomicName id="73FC8CF9FFDC0F149346FEAFFD5930D7" baseAuthorityName="Goodnight &amp; Goodnight" baseAuthorityYear="1971" box="[578,669,1938,1963]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="86882B68FFDC0F149346FEAFFD5930D7" box="[578,669,1938,1963]" italics="true" pageId="5" pageNumber="104">M. inops</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="E083A7F2FFDC0F14901CFCEDFAD1329E" ID-DOI="http://doi.org/10.5281/zenodo.4569988" ID-Zenodo-Dep="4569988" box="[280,1301,1488,1506]" httpUri="https://zenodo.org/record/4569988/files/figure.png" pageId="5" pageNumber="104" startId="5.[280,291,1488,1506]" targetBox="[132,1454,215,1446]" targetPageId="5">
<paragraph id="B443F77AFFDC0F14901CFCEDFAD1329E" blockId="5.[280,1301,1488,1506]" box="[280,1301,1488,1506]" pageId="5" pageNumber="104">
FIG. 1. — Habitus of the holotype (CNAN-T01342) of 
<taxonomicName id="73FC8CF9FFDC0F1493C2FCEDFC93329E" authority="Cruz-López, 2021" authorityName="Cruz-López" authorityYear="2021" box="[710,855,1488,1506]" class="Arachnida" family="Stygnopsidae" genus="Brujita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="chapulapa" status="n. gen., n. sp.">
<emphasis id="86882B68FFDC0F1493C2FCEDFC93329E" box="[710,855,1488,1506]" italics="true" pageId="5" pageNumber="104">Brujita chapulapa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DBB9613FFDC0F149258FCEDFC14329E" box="[860,976,1488,1506]" pageId="5" pageNumber="104" rank="species">n. gen., n. sp.</taxonomicNameLabel>
<emphasis id="86882B68FFDC0F1492D7FCEDFC24329E" bold="true" box="[979,992,1488,1506]" pageId="5" pageNumber="104">A</emphasis>
, dorsal; 
<emphasis id="86882B68FFDC0F14952CFCEDFBF1329E" bold="true" box="[1064,1077,1488,1506]" pageId="5" pageNumber="104">B</emphasis>
, lateral. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="B443F77AFFDC0F14919FFE8CFC323190" blockId="5.[130,776,1554,2028]" lastBlockId="5.[813,1456,1554,1772]" pageId="5" pageNumber="104">
Additionally, in the phylogeny of 
<taxonomicName id="73FC8CF9FFDC0F149308FE8CFD5F30B7" authorityName="SOrensen" authorityYear="1932" box="[524,667,1969,1995]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="family">Stygnopsidae</taxonomicName>
proposed by Cruz-López &amp; Francke (2017), 
<taxonomicName id="73FC8CF9FFDC0F14930CFEECFDAC3097" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[520,616,2001,2027]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDC0F14930CFEECFDAC3097" box="[520,616,2001,2027]" italics="true" pageId="5" pageNumber="104">Mictlana</emphasis>
</taxonomicName>
was recovered as monophyletic with high support (posterior probability 0.91), including a putative female and uncertain immatures of 
<taxonomicName id="73FC8CF9FFDC0F149283FF6EFC343110" authorityName="Goodnight &amp; Goodnight" authorityYear="1973" box="[903,1008,1618,1644]" class="Arachnida" family="Stygnopsidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="86882B68FFDC0F149283FF6EFC343110" box="[903,1008,1618,1644]" italics="true" pageId="5" pageNumber="104">H. planus</emphasis>
</taxonomicName>
. Now with the inclusion of 
<taxonomicName id="73FC8CF9FFDC0F14942DFF6EFA553110" authorityName="Goodnight &amp; Goodnight" authorityYear="1973" box="[1321,1425,1618,1644]" class="Arachnida" family="Stygnopsidae" genus="Hoplobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="86882B68FFDC0F14942DFF6EFA553110" box="[1321,1425,1618,1644]" italics="true" pageId="5" pageNumber="104">H. planus</emphasis>
</taxonomicName>
in 
<taxonomicName id="73FC8CF9FFDC0F149229FF4FFC4931F0" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[813,909,1650,1676]" class="Arachnida" family="Stygnopsidae" genus="Mictlana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="86882B68FFDC0F149229FF4FFC4931F0" box="[813,909,1650,1676]" italics="true" pageId="5" pageNumber="104">Mictlana</emphasis>
</taxonomicName>
, the presence of a large rounded anterior lobe on prosoma could be considered a putative synapomorphy of this genus, regardless whether or not it is the ocularium or the anterior bulge.
</paragraph>
</subSubSection>
</treatment>
</document>