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<document id="0ABE02757B29DCB2B64B8433CA84F5DE" ID-DOI="10.3897/zookeys.1223.139346" ID-ZooBank="451749E9-009E-43E9-A6F1-892035BDF1B0" ID-publisher-id="139346" URI-arpha="6EA2BAFE-82AB-5570-95B8-EE1F5FA60D46" URI-zoobank="http://zoobank.org/451749E9-009E-43E9-A6F1-892035BDF1B0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736349736103" checkinUser="pensoft" docAuthor="Wesener, Thomas, Akkari, Nesrine &amp; Golovatch, Sergei I." docDate="2025" docId="272301F93D9B5E349317C4940B710DBF" docLanguage="en" docName="ZooKeys 1223: 185-220" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/139346/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dalodesmus odontopezus" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="220" masterDocId="6EA2BAFE82AB557095B8EE1F5FA60D46" masterDocTitle="Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)" masterLastPageNumber="220" masterPageNumber="185" pageNumber="185" updateTime="1736355520980" updateUser="admin">
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<mods:title id="0C88CDA9F07CDDFC63B0AC6C650A5080">Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="F41A7E09704B0A6F4D9F81492F979D11">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="9A59A3E9D9BBC4CBEC4C1927DFBB7A81">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:namePart id="029F93C82513CB9B762BEE16BB0710D8">Akkari, Nesrine</mods:namePart>
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<mods:affiliation id="B04AF7A83EF5567FC65421AD4A854624">Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:namePart id="F577F03CC33184AF5CAC13BC436CD950">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="76AAD4D3D07FE30B3E2116E175A79D55">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
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<treatment id="272301F93D9B5E349317C4940B710DBF" ID-arpha="272301F9-3D9B-5E34-9317-C4940B710DBF" LSID="urn:lsid:plazi:treatment:272301F93D9B5E349317C4940B710DBF" httpUri="http://treatment.plazi.org/id/272301F93D9B5E349317C4940B710DBF" scope_class="Diplopoda" scope_family="Dalodesmidae" scope_order="Polydesmida">
<subSubSection id="3927C4022F1065053C726F33C993E78F" type="nomenclature">
<paragraph id="213DE656A4DD4C97228688DE9705989D">
<taxonomicName id="982F84045676777F51D10E19608E7990" ID-CoL="342DZ" authority="(Attems, 1898)" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezus">
<emphasis id="DF75F48458E6601E9FC08409B79020AC" italics="true">Dalodesmus odontopezus</emphasis>
(
<bibRefCitation id="758A6BC14BC7503F0F021A7307F8BB3B" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9673EA0B5FAB73B162E5B0F509F96A45" type="description">
<paragraph id="97C1343AE1BCA856980FA233274CE8DE">
<figureCitation id="D2622C3CCC04F6388BB99A0FF9AC7FB8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dalodesmus odontopezus (Attems, 1898), ♂ holotype A right paratergum 14, dorsal view (after Attems 1898) B slide of holotype C both gonopods, ventral view (after Attems 1940) D both gonopods, ventral view (slide NHMW MY 4429, photograph by O. Macek) E midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm." figureDoi="10.3897/zookeys.1223.139346.figure9" httpUri="https://binary.pensoft.net/fig/1215833">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F223A6F412EC6E8D6517906A57A8DC7F" type="reference_group">
<paragraph id="DAB4D037784912E3EFB69FB1AC353416">
<treatmentCitationGroup id="C86512F1B36C09E0B72D577510B86FBE">
<treatmentCitation id="282944D03CCDC9F7440080195ABBC8FD" author="Attems C" page="360" year="1898">
<taxonomicName id="4BF187565688B8A73CE43AAD2F89F715" ID-CoL="59J9Y" authority="Attems, 1898: 360" authorityName="Attems" authorityPageNumber="360" authorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum">
<emphasis id="FA6353533662BA1E2881BFD36FCFBE86" italics="true">Tubercularium odontopezum</emphasis>
<comment id="753DDFD5CE7B4AC408298287C2F984F8">
<bibRefCitation id="C36F42E344760634F5BB3844C635AF01" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898: 360</bibRefCitation>
, plate 7, figs 158161 (D).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="4755BC08D05074AF2A2B186FAF12BCF9">
<treatmentCitationGroup id="E59D7CDFA14726D6165480A453F982F4">
<taxonomicName id="06B965C8997E433A5831466066BCB48B" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum" subGenus="Tubercularium">
<emphasis id="67F64B977F051DCDA8DEC92BB35EB4A5" italics="true">Polydesmus</emphasis>
(
<emphasis id="90275F9F51E8A7C97E4DE2461B14D7F4" italics="true">Tubercularium</emphasis>
)
<emphasis id="2A3C275CA2E05B5B1CDEFDC638009B60" italics="true">odontopezum</emphasis>
</taxonomicName>
<comment id="C611103E9702386B6F23249F0227FDB1">
(sic!)
<treatmentCitation id="4E413A147654CF0206FEAB5EF89A20C6" author="de Saussure H &amp; Zehntner L" page="89" year="1902">
<bibRefCitation id="50F366A855B6265FE89213A21EB609A6" author="de Saussure H &amp; Zehntner L" editor="Grandidier A" firstAuthor="de Saussure" journalOrPublisher="Imprimérie nationale, Paris" page="89" pagination="1-356" refId="B9" refString="de Saussure H, Zehntner L (1902) Myriapodes de Madagascar. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Imprimérie nationale, Paris, 1356." title="Myriapodes de Madagascar." volumeTitle="Histoire physique, naturelle et politique de Madagascar." year="1902">de Saussure and Zehntner 1902: 89</bibRefCitation>
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(D).
</comment>
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</paragraph>
<paragraph id="B580A39D93E83FE97A56494CB46B0DC6">
<treatmentCitationGroup id="450B6F996F4E0E6977C71E041960797F">
<taxonomicName id="29EF2AD36AE5399C297D15000D8DEC57" ID-CoL="59J9Y" authorityName="Attems" authorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum">
<emphasis id="1E54100127B8CF78E34815C11D9316FD" italics="true">Tubercularium odontopezum</emphasis>
</taxonomicName>
<comment id="DBDD83F3D2F508FEA04F729E9F0FE27B">
<treatmentCitation id="F44562F67D87B0F18AF07CCBDDA3A176" author="Attems C" page="435" year="1940">
<bibRefCitation id="0ED6D8279332C16BB7F93C8BF01CCCDD" DOI="10.1515/9783111609645.1" author="Attems C" firstAuthor="Attems" page="435" refId="B2" refString="Attems C (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich 70, i xvii + 1577. https://doi.org/10.1515/9783111609645.1" year="1940">Attems 1940: 435</bibRefCitation>
</treatmentCitation>
, fig. 618 (D, K).
</comment>
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</paragraph>
<paragraph id="2499C4A71A97D55CB31CA5C6B0845C2A">
<treatmentCitationGroup id="59D2C841DDA04FF9C36DC4F94FF274BB">
<taxonomicName id="3709AB99E3C1FB2D6ED60514D3323016" ID-CoL="59J9Y" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum">
<emphasis id="654CDA89E4FBA8A40EAFFEFE2051B51E" italics="true">Dalodesmus odontopezum</emphasis>
</taxonomicName>
<comment id="D2EBDC42025808E137AF94A9084E4E08">
(sic!)
<treatmentCitation id="49891EA45AC59DDED09E8CBB5127061C" author="Hoffman RL" page="230" year="1974">
<bibRefCitation id="2793C4CE18743AA54D4E46D7561A72C4" author="Hoffman RL" firstAuthor="Hoffman" issue="2" journalOrPublisher="Wasmann Journal of Biology" page="230" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman 1974: 230</bibRefCitation>
</treatmentCitation>
(L).
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="271E43B6A4B74A05A47E79C876D1BDD2">
<treatmentCitationGroup id="F16008D6E9269ABEB9D04EDDAAEED9F4">
<taxonomicName id="64D4E1EA171864F33C65DD26DCE54997" ID-CoL="342DZ" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezus">
<emphasis id="E87D770986E2E8AD8E3C873404E51144" italics="true">Dalodesmus odontopezus</emphasis>
</taxonomicName>
<comment id="25FC9F3FB807BF499C9C694B912B1879">
<treatmentCitation id="41A64D01686911A714B135BDCE255B3A" author="Jeekel CAW" page="238" year="1965">
<bibRefCitation id="5C292EA4334DC19E0BD45E0A08939B38" author="Jeekel CAW" firstAuthor="Jeekel" journalOrPublisher="Entomologische Berichten" page="238" pagination="236-239" refId="B17" refString="Jeekel CAW (1965) The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida). Entomologische Berichten 25: 236239." title="The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida)." volume="25" year="1965">Jeekel 1965: 238</bibRefCitation>
</treatmentCitation>
(L);
<treatmentCitation id="8040B47B55BC2766C431624BAE37E9AE" author="Golovatch SI &amp; Hoffman RL" page="161" year="1989">
<bibRefCitation id="36C7BB0FE4AB05CE58A3882BD2421270" DOI="10.1080/03946975.1989.10539436" author="Golovatch SI &amp; Hoffman RL" firstAuthor="Golovatch" journalOrPublisher="Tropical Zoology" page="161" pagination="159-164" refId="B12" refString="Golovatch SI, Hoffman RL (1989) Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae). Tropical Zoology 2: 159164. https://doi.org/10.1080/03946975.1989.10539436" title="Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae)." volume="2" year="1989">Golovatch and Hoffman 1989: 161</bibRefCitation>
</treatmentCitation>
(L);
<treatmentCitation id="AC1D6E7BD7720C2AA3D043A2DB7C38C7" author="Enghoff H" page="623" year="2003">
<bibRefCitation id="27BF66438A0AE963BDDB32769F666447" author="Enghoff H" editor="Goodman SM &amp; Benstead JP" firstAuthor="Enghoff" journalOrPublisher="University of Chicago Press, Chicago and London" page="623" pagination="617-627" refId="B10" refString="Enghoff H (2003) Diplopoda, millipedes. In: Goodman SM, Benstead JP (Eds) The Natural History of Madagascar. University of Chicago Press, Chicago and London, 617627." title="Diplopoda, millipedes." volumeTitle="The Natural History of Madagascar." year="2003">Enghoff 2003: 623</bibRefCitation>
</treatmentCitation>
(L);
<treatmentCitation id="98A9380C9AD3FD7EA3CA5FBB02E22564" author="Wesener T &amp; Enghoff H" page="926" year="2022">
<bibRefCitation id="6BC60837EBF0F833BCE1DAC175AE38C7" DOI="10.2307/j.ctv2ks6tbb.118" author="Wesener T &amp; Enghoff H" editor="Fisher BL &amp; Goodman SM" firstAuthor="Wesener" journalOrPublisher="Princeton University Press USA" page="926" pagination="918-933" refId="B22" refString="Wesener T, Enghoff H (2022) Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy. In: Fisher BL, Goodman SM (Eds) The New Natural History of Madagascar. Princeton University Press USA, 918933. https://doi.org/10.2307/j.ctv2ks6tbb.118" title="Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy." volumeTitle="The New Natural History of Madagascar." year="2022">Wesener and Enghoff 2022: 926</bibRefCitation>
</treatmentCitation>
(L).
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0ESEBG" type="notes">
<paragraph id="405FB4FF2C9F90F89E8DC4CA10EE2B35">
<heading id="07F43515B8500330C3288688B9F65C4B" reason="title">Note.</heading>
</paragraph>
<paragraph id="518FC712FDF62CF73AEB3E11203DEFE1">
This species was described from a single
<specimenCount id="451F93CDCEF274165EC5A0C2AD5724EE" count="1" type="male"></specimenCount>
<typeStatus id="5C9AC81832BAB0E89FA619ED12D7FE0B" type="holotype">holotype</typeStatus>
, from Nosy Be Islet (
<bibRefCitation id="C8950FD654CE53D943EB2078C8E16C4E" author="Attems" firstAuthor="Attems" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems 1898</bibRefCitation>
), slide
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<collectionCode id="25060FC44B64F4DD029E213E718A9DCE" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
MY 4429 containing its two legs and gonopods (Fig.
<figureCitation id="C58D50EFBCA93F27C777EE98FC305408" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dalodesmus odontopezus (Attems, 1898), ♂ holotype A right paratergum 14, dorsal view (after Attems 1898) B slide of holotype C both gonopods, ventral view (after Attems 1940) D both gonopods, ventral view (slide NHMW MY 4429, photograph by O. Macek) E midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm." figureDoi="10.3897/zookeys.1223.139346.figure9" httpUri="https://binary.pensoft.net/fig/1215833">9 C</figureCitation>
), labelled
<taxonomicName id="272C256B90A0CB9E4208C0FBBD9CDB8D" authority="(Attems, 1898)" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum">
<emphasis id="C350273B0496C530C2DFEFB593CB3A4E" italics="true">Tubercularium odontopezum</emphasis>
(
<bibRefCitation id="DDB0C248D0195B96151B39A922D32F16" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898</bibRefCitation>
)
</taxonomicName>
, revised, in the
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<collectionCode id="E8D3D0DE696ED200A0553FB39640F21E" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
collection. The torso seems to be misplaced.
</paragraph>
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<paragraph id="A32DF41C9191CDFF7BB3378EA5DE08F5">
<label id="7F573F981C7DA32434DBEBED7D78B034">Figure 9.</label>
</paragraph>
<paragraph id="EC29AB7D07C5C6A38E2666BD3A995573">
<taxonomicName id="02397CFC5CA60828B22E533413AA3FA1" authority="(Attems, 1898)" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezus">
<emphasis id="8E4DDEC00B7227E80EE53A736674EC0F" italics="true">Dalodesmus odontopezus</emphasis>
(
<bibRefCitation id="F6BF0A40DA8C58306CC642F82EEE3F50" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898</bibRefCitation>
)
</taxonomicName>
, ♂ holotype
<emphasis id="70ADB794231AF59B9C613253B026F490" bold="true">A</emphasis>
right paratergum 14, dorsal view (after
<bibRefCitation id="F681DD8B8EE76434FAE920A1C2F247C8" author="Attems" firstAuthor="Attems" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems 1898</bibRefCitation>
)
<emphasis id="9EE2478651E338BF746DC470A646D6C2" bold="true">B</emphasis>
slide of holotype
<emphasis id="AAC5E692746EB99DE9AC96FCBE256D78" bold="true">C</emphasis>
both gonopods, ventral view (after
<bibRefCitation id="DD9DACB5A0F25CEDB69C12005D9692B6" DOI="10.1515/9783111609645.1" author="Attems" firstAuthor="Attems" refId="B2" refString="Attems C (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich 70, i xvii + 1577. https://doi.org/10.1515/9783111609645.1" year="1940">Attems 1940</bibRefCitation>
)
<emphasis id="A79CB36CF5C12FA9A6CAF6CF7053F6F3" bold="true">D</emphasis>
both gonopods, ventral view (slide
<named-content id="7C6BD3405F7302A18D799DEEBBEB5F2B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Austria">NHMW</named-content>
MY 4429, photograph by O. Macek)
<emphasis id="B49A5F69AB5A9F1E6634A7B35464C3D0" bold="true">E</emphasis>
midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEHBG" type="description">
<paragraph id="0478C62C9AD91C6E17CF82FD6D7A043E">
<heading id="2885D6E21746352B23C35B72B5A1A6C5" reason="title">Brief description.</heading>
</paragraph>
<paragraph id="E20283D1316C3197E5A2C86D2C6F1A4E">
(After
<bibRefCitation id="2863C48A7FE0F4C2DFE98EC08CF4F4C0" author="Attems" firstAuthor="Attems" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems 1898</bibRefCitation>
,
<bibRefCitation id="A8B6DC12F16124472221A6783262B533" DOI="10.1515/9783111609645.1" author="Attems" firstAuthor="Attems" refId="B2" refString="Attems C (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich 70, i xvii + 1577. https://doi.org/10.1515/9783111609645.1" year="1940">1940</bibRefCitation>
.)
<specimenCount id="A036066EB89D8A4FC210A53AD1D722DC" count="1" type="male"></specimenCount>
<typeStatus id="6ED11ED9FA77BD927D1363C3548B53B1" type="holotype">holotype</typeStatus>
<quantity id="69A7BA740C4C09B946E09BAE3E6FC190" metricMagnitude="-2" metricUnit="m" metricValue="2.8" unit="mm" value="28.0">28 mm</quantity>
long and 2.0 and
<quantity id="7D606D724D032D753182A3C11DC0B715" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
wide on pro- and metazona, respectively. Colouration of metaterga more intense brown in posterior halves, lighter in anterior ones; sides of paraterga yellowish, venter light yellowish brown, antennae and legs dirty yellow. Paraterga horizontal, not upturned above dorsum, sharpened caudally; dorsal tuberculations between paraterga mostly round (Fig.
<figureCitation id="AB129D70DE420D57BF7756567F2E3FCA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dalodesmus odontopezus (Attems, 1898), ♂ holotype A right paratergum 14, dorsal view (after Attems 1898) B slide of holotype C both gonopods, ventral view (after Attems 1940) D both gonopods, ventral view (slide NHMW MY 4429, photograph by O. Macek) E midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm." figureDoi="10.3897/zookeys.1223.139346.figure9" httpUri="https://binary.pensoft.net/fig/1215833">9 A</figureCitation>
).
</paragraph>
<paragraph id="06538CA568B51554183FB484E0E159EC">
Gonopods (Fig.
<figureCitation id="C155CC8F80D8D803D6B38670A5819933" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dalodesmus odontopezus (Attems, 1898), ♂ holotype A right paratergum 14, dorsal view (after Attems 1898) B slide of holotype C both gonopods, ventral view (after Attems 1940) D both gonopods, ventral view (slide NHMW MY 4429, photograph by O. Macek) E midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm." figureDoi="10.3897/zookeys.1223.139346.figure9" httpUri="https://binary.pensoft.net/fig/1215833">9 B, C</figureCitation>
) showing setose femorites (fe), each with a small roundish, pulvillus-like field (p) of particularly dense setae ventrally near apex, coupled with complex acropodites: a slightly sigmoid and apically bifid solenomere (sl) lying between the highest, erect and subspiniform mesal branch (mb) and a much shorter, but prominent, roughly laciniate and fimbriate lateral branch (lb) directed ventrad.
</paragraph>
</subSubSection>
</treatment>
</document>