Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida) Author Wesener, Thomas 0000-0002-2028-3541 Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany Author Akkari, Nesrine 0000-0001-5019-4833 Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Golovatch, Sergei I. Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia text ZooKeys 2025 2025-01-08 1223 185 220 journal article 10.3897/zookeys.1223.139346 451749E9-009E-43E9-A6F1-892035BDF1B0 Dalodesmus odontopezus ( Attems, 1898 ) Fig. 9 Tubercularium odontopezum Attems, 1898: 360 , plate 7, figs 158–161 (D). Polydesmus ( Tubercularium ) odontopezum (sic!) – de Saussure and Zehntner 1902: 89 (D). Tubercularium odontopezum Attems 1940: 435 , fig. 618 (D, K). Dalodesmus odontopezum (sic!) – Hoffman 1974: 230 (L). Dalodesmus odontopezus Jeekel 1965: 238 (L); Golovatch and Hoffman 1989: 161 (L); Enghoff 2003: 623 (L); Wesener and Enghoff 2022: 926 (L). Note. This species was described from a single holotype , from Nosy Be Islet ( Attems 1898 ), slide NHMW MY 4429 containing its two legs and gonopods (Fig. 9 C ), labelled Tubercularium odontopezum ( Attems, 1898 ) , revised, in the NHMW collection. The torso seems to be misplaced. Dalodesmus odontopezus ( Attems, 1898 ) , ♂ holotype A right paratergum 14, dorsal view (after Attems 1898 ) B slide of holotype C both gonopods, ventral view (after Attems 1940 ) D both gonopods, ventral view (slide NHMW MY 4429, photograph by O. Macek) E midbody leg. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = prefemorite; sl = solenomere branch. Scale bars: 500 µm. Brief description. (After Attems 1898 , 1940 .) holotype 28 mm long and 2.0 and 3.5 mm wide on pro- and metazona, respectively. Colouration of metaterga more intense brown in posterior halves, lighter in anterior ones; sides of paraterga yellowish, venter light yellowish brown, antennae and legs dirty yellow. Paraterga horizontal, not upturned above dorsum, sharpened caudally; dorsal tuberculations between paraterga mostly round (Fig. 9 A ). Gonopods (Fig. 9 B, C ) showing setose femorites (fe), each with a small roundish, pulvillus-like field (p) of particularly dense setae ventrally near apex, coupled with complex acropodites: a slightly sigmoid and apically bifid solenomere (sl) lying between the highest, erect and subspiniform mesal branch (mb) and a much shorter, but prominent, roughly laciniate and fimbriate lateral branch (lb) directed ventrad.