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<document id="48077A6285C31207CDB7E3EA655E3310" ID-DOI="10.11646/phytotaxa.395.4.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13716734" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725650394261" checkinUser="felipe" docAuthor="Martínez-Sagarra, Gloria &amp; Devesa, Juan A." docDate="2019" docId="2052AF45FFEFFFF5C5965138C071EF3F" docLanguage="en" docName="phytotaxa.395.4.1.pdf" docOrigin="Phytotaxa 395 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.395.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Festuca greuteri Martinez-Sagarra &amp; Devesa 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="DC6BD73DFFEDFFF0C51E522CC125EC7E" masterDocTitle="Two new taxa of Festuca (Poaceae) from the southeastern Iberian Peninsula" masterLastPageNumber="264" masterPageNumber="251" pageNumber="253" updateTime="1725657136954" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0230D5F54EB1BE66F3698A643F2FBC2C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B79C0D2B2BD2B14B1AFA646012810B55">Two new taxa of Festuca (Poaceae) from the southeastern Iberian Peninsula</mods:title>
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<mods:name id="B6A640A80504173B130C27B54ACC1857" type="personal">
<mods:role id="6808B5C27FA20C3B24858D23EFC35F41">
<mods:roleTerm id="C3A6D8C2B0F4740A85D3389E19449039">Author</mods:roleTerm>
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<mods:namePart id="1E73EA5F66AF13E479911A8B6C135604">Martínez-Sagarra, Gloria</mods:namePart>
</mods:name>
<mods:name id="45A7A5C4901139A56F5E223745F5A4ED" type="personal">
<mods:role id="9D7FD95DD7C4BFC184E6C6360F5502C5">
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<mods:namePart id="394D314CEB16049D8CA274CF9973DEE2">Devesa, Juan A.</mods:namePart>
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<mods:typeOfResource id="C6E2997BED75318FB95D74BF9B6B4166">text</mods:typeOfResource>
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<mods:title id="239BED58C2A178253BC06508CAC64175">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0E75BE1BF349D17D204C19AA3D8EF1B1">
<mods:date id="50EA4A73317FA9BF804965943F8BE9D3">2019</mods:date>
<mods:detail id="A93728D3E106CE22F9CAE6FC7F848834" type="pubDate">
<mods:number id="ED5D0EBD4597115C247391A5F9F36090">2019-03-12</mods:number>
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<mods:detail id="EC4270D4F184F711FBA8585F49F52AA5" type="volume">
<mods:number id="A3ED6369CC6ECC9D7BF71DADA48D49D3">395</mods:number>
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<mods:number id="F761FEA48606E3F5389CD162B4DB699A">4</mods:number>
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<mods:start id="49301F55B762534DC5034728D5AC89C5">251</mods:start>
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<mods:url id="4CFEC0E66D49D1C8E3781CCEC7BB9AA3">http://dx.doi.org/10.11646/phytotaxa.395.4.1</mods:url>
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<mods:classification id="4A6437685DFDAE34327CED20670B4A8D">journal article</mods:classification>
<mods:identifier id="AE223E1482880D549D7BA003C95B4F8A" type="DOI">10.11646/phytotaxa.395.4.1</mods:identifier>
<mods:identifier id="8BFAD6E4C1BF135475E700032D5EF2CD" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="DE1EF03D96F8480852AD8C4B837027B7" type="Zenodo-Dep">13716734</mods:identifier>
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<treatment id="2052AF45FFEFFFF5C5965138C071EF3F" LSID="urn:lsid:plazi:treatment:2052AF45FFEFFFF5C5965138C071EF3F" httpUri="http://treatment.plazi.org/id/2052AF45FFEFFFF5C5965138C071EF3F" lastPageId="5" lastPageNumber="256" pageId="2" pageNumber="253">
<subSubSection id="E0E14DD8FFEFFFF2C5965138C243EF51" box="[136,870,788,815]" pageId="2" pageNumber="253" type="nomenclature">
<paragraph id="A8441E53FFEFFFF2C5965138C243EF51" blockId="2.[136,870,788,815]" box="[136,870,788,815]" pageId="2" pageNumber="253">
<heading id="F30CA93FFFEFFFF2C5965138C243EF51" box="[136,870,788,815]" fontSize="11" level="2" pageId="2" pageNumber="253" reason="3">
<taxonomicName id="6FFB65D0FFEFFFF2C5965138C3A2EF51" ID-CoL="8Y3XC" authority="Martinez-Sagarra &amp; Devesa" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[136,647,788,815]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="2" pageNumber="253" phylum="Tracheophyta" rank="species" species="greuteri" status="sp. nov.">
<emphasis id="9A8FC241FFEFFFF2C5965138C062EF50" bold="true" box="[136,327,788,814]" italics="true" pageId="2" pageNumber="253">Festuca greuteri</emphasis>
Martínez-Sagarra &amp; Devesa
</taxonomicName>
<emphasis id="9A8FC241FFEFFFF2C791513AC3C1EF51" box="[655,740,790,815]" italics="true" pageId="2" pageNumber="253">
<taxonomicNameLabel id="81BC7F3AFFEFFFF2C791513AC3C1EF51" box="[655,740,790,815]" pageId="2" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="30C002D6FFEFFFF2C7EC513AC278EF51" box="[754,861,790,815]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[136,229,2029,2050]" captionStartId-1="4.[136,229,1871,1892]" captionTargetBox-0="[249,1338,190,2004]" captionTargetBox-1="[226,1376,194,1833]" captionTargetId-0="figure-14@3.[249,1338,190,2004]" captionTargetId-1="figure-16@4.[197,1391,190,1849]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Holotype of Festuca greuteri (Granada, Sierra de Guillimona, puerto de la Losa, COFC 65825)." captionText-1="FIGURE 3. Festuca greuteri subsp. greuteri (A–C and E–L) and F. greuteri subsp. camarolensis (D). A) Habit; B) Detail of the apex of the sheath and ligule in adaxial view; C−D) Spikelet; E) Lower glume in abaxial view; F) Upper glume in abaxial view; G) Lemma in abaxial view; H) Palea in abaxial view; I) Lodicule; J) Sexual whorls; K) Caryopsis in adaxial view; L) Caryopsis in abaxial view. Vouchers: Spain, Puerto de la Losa, Sierra de Guillimona, Granada, 4 June 2014, E. López &amp; G. Martínez, COFC 61365 (F. greuteri subsp. greuteri); and Spain, Sierra de Camarolos, Málaga, 10 June 2015, B. Cabezudo, J. A. Devesa, G. Martínez &amp; F. Soriguer, COFC 62140 (F. greuteri subsp. camarolensis)." figureDoi-0="http://doi.org/10.5281/zenodo.13716738" figureDoi-1="http://doi.org/10.5281/zenodo.13716740" httpUri-0="https://zenodo.org/record/13716738/files/figure.png" httpUri-1="https://zenodo.org/record/13716740/files/figure.png" pageId="2" pageNumber="253">
Figs. 2
<emphasis id="9A8FC241FFEFFFF2C65E5134C275EF50" box="[832,848,792,814]" italics="true" pageId="2" pageNumber="253">−</emphasis>
3
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0E14DD8FFEFFFF2C5965110C41FEF0B" pageId="2" pageNumber="253" type="materials_examined">
<paragraph id="A8441E53FFEFFFF2C5965110C41FEF0B" blockId="2.[136,1452,828,885]" pageId="2" pageNumber="253">
<emphasis id="9A8FC241FFEFFFF2C5965110C1E7EF2F" bold="true" box="[136,194,828,849]" pageId="2" pageNumber="253">Type:</emphasis>
—
<materialsCitation id="1893140EFFEFFFF2C5C55110C413EF0B" collectingDate="2014-06-04" collectorName="de Guillimona &amp; de la Losa &amp; E. Lopez &amp; G. Martinez-Sagarra" country="Spain" elevation="1780" latitude="38.003613" location="Granada" longLatPrecision="19" longitude="-2.59" municipality="Sierra de Guillimona" pageId="2" pageNumber="253" specimenCount="2" typeStatus="holotype">
<collectingCountry id="D0EC5EC3FFEFFFF2C5C55110C004EF2F" box="[219,289,828,849]" name="Spain" pageId="2" pageNumber="253">SPAIN</collectingCountry>
.
<location id="AD244888FFEFFFF2C4355110C0A5EF2F" LSID="urn:lsid:plazi:treatment:2052AF45FFEFFFF5C5965138C071EF3F:AD244888FFEFFFF2C4355110C0A5EF2F" box="[299,384,828,849]" country="Spain" latitude="38.003613" longLatPrecision="19" longitude="-2.59" municipality="Sierra de Guillimona" name="Granada" pageId="2" pageNumber="253">Granada</location>
:
<collectingMunicipality id="48208429FFEFFFF2C4955110C37EEF2F" box="[395,603,828,849]" pageId="2" pageNumber="253">
Sierra
<collectorName id="050E7B85FFEFFFF2C4D25110C37EEF2F" box="[460,603,828,849]" pageId="2" pageNumber="253">de Guillimona</collectorName>
</collectingMunicipality>
, puerto
<collectorName id="050E7B85FFEFFFF2C7B55110C231EF2F" box="[683,788,828,849]" pageId="2" pageNumber="253">de la Losa</collectorName>
,
<geoCoordinate id="CDCF7894FFEFFFF2C63E5110C2A2EF2F" box="[800,903,828,849]" degrees="38" direction="north" minutes="0" orientation="latitude" pageId="2" pageNumber="253" precision="15" seconds="13" value="38.003613">38º0’13”N</geoCoordinate>
<geoCoordinate id="CDCF7894FFEFFFF2C6915110C2D9EF2F" box="[911,1020,828,849]" degrees="2" direction="west" minutes="35" orientation="longitude" pageId="2" pageNumber="253" precision="15" seconds="24" value="-2.59">2º35’24”W</geoCoordinate>
,
<quantity id="6F03B3B6FFEFFFF2C1165110C577EF2F" box="[1032,1106,828,849]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="2" pageNumber="253" unit="m" value="1780.0">
<elevation id="23D6F960FFEFFFF2C1165110C577EF2F" box="[1032,1106,828,849]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="2" pageNumber="253" unit="m" value="1780.0">1780 m</elevation>
</quantity>
elevation,
<date id="DC453893FFEFFFF2C1DE5110C41FEF2F" box="[1216,1338,828,849]" pageId="2" pageNumber="253" value="2014-06-04">
<collectingDate id="CC01C17BFFEFFFF2C1DE5110C41FEF2F" box="[1216,1338,828,849]" pageId="2" pageNumber="253" value="2014-06-04">4 June 2014</collectingDate>
</date>
, calizas, en márgenes de carretera,
<emphasis id="9A8FC241FFEFFFF2C485514CC3FDEF0B" box="[411,728,864,885]" italics="true" pageId="2" pageNumber="253">
<collectorName id="050E7B85FFEFFFF2C485514CC0CAEF0B" box="[411,495,864,885]" pageId="2" pageNumber="253">E. López</collectorName>
&amp;
<collectorName id="050E7B85FFEFFFF2C710514CC3FDEF0B" box="[526,728,864,885]" pageId="2" pageNumber="253">G. Martínez-Sagarra</collectorName>
</emphasis>
,
<emphasis id="9A8FC241FFEFFFF2C7FB514CC264EF0B" box="[741,833,864,885]" italics="true" pageId="2" pageNumber="253">GM22/14</emphasis>
(
<typeStatus id="7740A0F1FFEFFFF2C653514CC283EF0B" box="[845,934,864,885]" pageId="2" pageNumber="253" type="holotype">holotype</typeStatus>
: COFC 65825!,
<figureCitation id="30C002D6FFEFFFF2C15A514CC55BEF0B" box="[1092,1150,864,885]" captionStart="FIGURE 2" captionStartId="3.[136,229,2029,2050]" captionTargetBox="[249,1338,190,2004]" captionTargetId="figure-14@3.[249,1338,190,2004]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Festuca greuteri (Granada, Sierra de Guillimona, puerto de la Losa, COFC 65825)." figureDoi="http://doi.org/10.5281/zenodo.13716738" httpUri="https://zenodo.org/record/13716738/files/figure.png" pageId="2" pageNumber="253">Fig. 2</figureCitation>
;
<typeStatus id="7740A0F1FFEFFFF2C197514CC5FEEF0B" box="[1161,1243,864,885]" pageId="2" pageNumber="253" type="isotype">isotypes</typeStatus>
: COFC!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E0E14DD8FFEFFFF2C596518AC2D7EB3D" pageId="2" pageNumber="253" type="description">
<paragraph id="A8441E53FFEFFFF2C596518AC2EDEA85" blockId="2.[136,1452,934,2003]" pageId="2" pageNumber="253">
<emphasis id="9A8FC241FFEFFFF2C596518AC039EFC1" bold="true" box="[136,284,934,959]" pageId="2" pageNumber="253">Description:</emphasis>
—Perennial herb, tufted, culm up to 55(65) cm ×
<quantity id="6F03B3B6FFEFFFF2C65F518AC2ECEFC1" box="[833,969,934,959]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="2" pageNumber="253" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
, erect, cylindrical, finely striated, glabrous, smooth or more rarely sparsely antrorse-scabrid under the inflorescence, with (0–)1(–2) visible nodes towards the lower part, covered at the base by imbricate leaf sheaths. Leaf sheath of non-flowering shoots with margins free for about 2/3–3/4 of their length, greenish-glaucous and herbaceous at the beginning, papyraceous and straw-coloured at maturity, pruinose or not, glabrous, smooth; auricles 0.1–0.5(–0.8) mm, sometimes unequal, obtuse, briefly ciliolate at the apex; ligule
<quantity id="6F03B3B6FFEFFFF2C4455676C0C3E80D" box="[347,486,1114,1139]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="2" pageNumber="253" unit="mm" value="0.3" valueMax="0.5" valueMin="0.1">0.1–0.5 mm</quantity>
, membranous, truncate, briefly ciliolate; leaf blade (1.5–)2–15(–20) cm ×
<quantity id="6F03B3B6FFEFFFF2C02C5676C482E80D" box="[1330,1447,1114,1139]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="2" pageNumber="253" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.6–1 mm</quantity>
, setaceous, rarely almost junciform, erect-curved, sometimes ± sigmoid at the end, ± rigid, not pungent, conduplicate, from elliptical to obovate (rarely V-shaped) outline in cross-section, continuous abaxial subepidermal sclerenchyma, sometimes somewhat interrupted, absent on the ribs, 7(–9) vascular bundles, 4(–6) grooves, and 3(–5) ribs in cross-section (
<figureCitation id="30C002D6FFEFFFF2C5F656C6C00BE97D" box="[232,302,1258,1283]" captionStart="FIGURE 4" captionStartId="5.[136,229,1927,1948]" captionTargetBox="[371,1212,1135,1849]" captionTargetId="figure-413@5.[357,1231,1130,1850]" captionTargetPageId="5" captionText="FIGURE 4. Leaf blades in cross-section of Festuca greuteri subsp. greuteri (A–F), and F.greuteri subsp.camarolensis (G–K).Provenance of the material: Sierra de la Sagra (A, F), Sierra de Baza (B), Sierra de Tejeda (E), Sierra de Guillimona (D), and Sierra de Camarolos (C, G–K)." figureDoi="http://doi.org/10.5281/zenodo.13716744" httpUri="https://zenodo.org/record/13716744/files/figure.png" pageId="2" pageNumber="253">Fig. 4</figureCitation>
), green or usually greenish-glaucous, glabrous, smooth or antrorse-scabridulous, apex from obtuse to acute. Cauline leaves 1(–2), similar to those of non-flowering shoots, with auricles
<quantity id="6F03B3B6FFEFFFF2C1235722C5EDE959" box="[1085,1224,1294,1319]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="2" pageNumber="253" unit="mm" value="0.5" valueMax="0.8" valueMin="0.2">0.2–0.8 mm</quantity>
, ligule
<quantity id="6F03B3B6FFEFFFF2C0035722C482E959" box="[1309,1447,1294,1319]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="253" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
, and blade
<quantity id="6F03B3B6FFEFFFF2C5E1571EC04CE935" box="[255,361,1330,1355]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="6.0" metricValueMin="1.3" pageId="2" pageNumber="253" unit="cm" value="3.65" valueMax="6.0" valueMin="1.3">1.3–6 cm</quantity>
×
<quantity id="6F03B3B6FFEFFFF2C495571EC333E935" box="[395,534,1330,1355]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="2" pageNumber="253" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
. Panicle 2–8.5(–9) cm, dense, oblong-ellipsoidal, with 13–29(–32) spikelets and an axis provided with 9–11 nodes
<emphasis id="9A8FC241FFEFFFF2C70A577BC301E913" box="[532,548,1367,1389]" italics="true" pageId="2" pageNumber="253">−</emphasis>
the lowermost internode (0.5–)1.5–2.5(–3) cm
<emphasis id="9A8FC241FFEFFFF2C12D577BC566E913" box="[1075,1091,1367,1389]" italics="true" pageId="2" pageNumber="253">−</emphasis>
, antrorse-scabrid, with 1–3(–5) branches
<emphasis id="9A8FC241FFEFFFF2C5EB5757C020E9EF" box="[245,261,1403,1425]" italics="true" pageId="2" pageNumber="253">−</emphasis>
first simple branch,
<quantity id="6F03B3B6FFEFFFF2C4F25756C367E9ED" box="[492,578,1402,1427]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="2" pageNumber="253" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 cm</quantity>
and with 2–6 spikelets
<emphasis id="9A8FC241FFEFFFF2C6505757C27BE9EF" box="[846,862,1403,1425]" italics="true" pageId="2" pageNumber="253">−</emphasis>
, solitary, erect, ± appressed, usually erect-patent at anthesis. Spikelets (5.5)6–9.5(–11.5) mm, laterally compressed, elliptic, green, greenish-glaucous or greenish-purple, with a pedicel of
<quantity id="6F03B3B6FFEFFFF2C45657EEC0F5E9A5" box="[328,464,1474,1499]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.5" metricValueMin="0.5" pageId="2" pageNumber="253" unit="mm" value="2.5" valueMax="4.5" valueMin="0.5">0.5–4.5 mm</quantity>
, slightly thickened at the apex, (3–)4–7(–8) fertile florets and 1 apical and ± rudimentary sterile floret. Two glumes, unequal, narrowly scarious on margins, smooth or with scattered aculei at the edges of the upper half, acute or acuminate, the lower (1.6–)2–4(–4.3) mm, linear-lanceolate, with 1 nerve; the upper (3–)3.2–5(– 6.3) mm, ovate-lanceolate, with 3 nerves. Lemma (3.6–)
<quantity id="6F03B3B6FFEFFFF2C6165402C2B0EA39" box="[776,917,1582,1607]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="6.7" metricValueMin="4.2" pageId="2" pageNumber="253" unit="mm" value="5.45" valueMax="6.7" valueMin="4.2">4.2–6.7 mm</quantity>
, ovate-lanceolate, acute, awned, with narrowly scarious margins, with sparse aculei in the upper half, greenish or greenish-purple at the margins and towards the apex, with 5 nerves, that of the basal floret with an awn of
<quantity id="6F03B3B6FFEFFFF2C603545AC283EAF1" box="[797,934,1654,1679]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="14.0" metricValueMin="1.0" pageId="2" pageNumber="253" unit="mm" value="0.75" valueMax="1.4" valueMin="0.1">0.1–1.4 mm</quantity>
, that of the second basal floret with an awn of (0.1–)0.2–2.3(–2.5) mm, and awns up to 2.5(–3) mm in the rest. Palea (3.5–)4–5.7(–6) mm, subequal or somewhat shorter than lemma, bidentate, with 2 scabrid keels. Anthers (1.7–)1.9–3.4(–3.6) mm, linear, yellow. Ovary glabrous. Lodicules
<quantity id="6F03B3B6FFEFFFF2C5E354CEC0A2EA85" box="[253,391,1762,1787]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="253" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
, bilobed. Caryopsis
<quantity id="6F03B3B6FFEFFFF2C77554CEC3D1EA85" box="[619,756,1762,1787]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.4" metricValueMin="2.6" pageId="2" pageNumber="253" unit="mm" value="3.0" valueMax="3.4" valueMin="2.6">2.6–3.4 mm</quantity>
, adhering to palea.
</paragraph>
<paragraph id="A8441E53FFEFFFF2C5A3552AC4ABEB61" blockId="2.[136,1452,934,2003]" box="[189,1422,1798,1823]" pageId="2" pageNumber="253">
<emphasis id="9A8FC241FFEFFFF2C5A3552AC01FEB61" bold="true" box="[189,314,1798,1823]" pageId="2" pageNumber="253">Eponymy:</emphasis>
—The specific epithet honours Prof. emer. Dr. Werner Greuter (Università degli Studi di Palermo).
</paragraph>
<paragraph id="A8441E53FFEFFFF2C5A35506C2D7EB3D" blockId="2.[136,1452,934,2003]" box="[189,1010,1834,1859]" pageId="2" pageNumber="253">
<emphasis id="9A8FC241FFEFFFF2C5A35506C066EB3D" bold="true" box="[189,323,1834,1859]" pageId="2" pageNumber="253">Phenology:</emphasis>
—Flowering time, May to July; fruiting time, July to August.
</paragraph>
</subSubSection>
<subSubSection id="E0E14DD8FFEFFFF5C5A35562C071EF3F" lastPageId="5" lastPageNumber="256" pageId="2" pageNumber="253" type="distribution">
<paragraph id="A8441E53FFEFFFF2C5A35562C074EBAD" blockId="2.[136,1452,934,2003]" pageId="2" pageNumber="253">
<emphasis id="9A8FC241FFEFFFF2C5A35562C0CFEB19" bold="true" box="[189,490,1870,1895]" pageId="2" pageNumber="253">Distribution and ecology:</emphasis>
—Endemic to the southeast of the Iberian Peninsula (
<collectingCountry id="D0EC5EC3FFEFFFF2C12D5562C55CEB19" box="[1075,1145,1870,1895]" name="Spain" pageId="2" pageNumber="253">Spain</collectingCountry>
). The species is restricted to Baetic summit areas (Almería, Granada, Jaén, and Málaga provinces). It grows on open grasslands, scree of mountain meadows, and open scrubland, on basic substrates (limestones, dolomites, and marble dolomites);
<quantity id="6F03B3B6FFEFFFF2C01255BAC489EBD1" box="[1292,1452,1942,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.2" metricValueMin="1.3" pageId="2" pageNumber="253" unit="m" value="1750.0" valueMax="2200.0" valueMin="1300.0">1300–2200 m</quantity>
elevation (
<figureCitation id="30C002D6FFEFFFF2C5E05596C01AEBAD" box="[254,319,1978,2003]" captionStart="FIGURE 1" captionStartId="1.[136,229,1921,1942]" captionTargetBox="[370,1218,947,1893]" captionTargetId="figure-416@1.[370,1219,947,1893]" captionTargetPageId="1" captionText="FIGURE 1. A) Distributionmap of Festucagreuteri subsp.greuteri, F.greuteri subsp.camarolensis, F.longiauriculata andF.segimonensis. Main populations studied: 1, Sierra de Guillimona (type population of F. greuteri) and Sierra de la Sagra; 2, Sierra de Alcaraz; 3, Sierra de Cazorla and Sierra de Castril; 4, Sierra de María; 5, Sierra de los Filabres; 6, Sierra de Baza; 7, Sierra Nevada; 8, Sierra de Gádor; 9, Sierra de Mágina; 10, Sierra de Tejeda; 11, Sierra de Camarolos and Sierra de El Jobo. B) Habitat of F. greuteri subsp. greuteri (puerto de la Losa, Sierra de Guillimona)." figureDoi="http://doi.org/10.5281/zenodo.13716736" httpUri="https://zenodo.org/record/13716736/files/figure.png" pageId="2" pageNumber="253">Fig. 1</figureCitation>
).
</paragraph>
<caption id="FC844EDBFFEEFFF3C59655C1C5B8E47C" ID-DOI="http://doi.org/10.5281/zenodo.13716738" ID-Zenodo-Dep="13716738" box="[136,1181,2029,2050]" httpUri="https://zenodo.org/record/13716738/files/figure.png" pageId="3" pageNumber="254" startId="3.[136,229,2029,2050]" targetBox="[249,1338,190,2004]" targetPageId="3" targetType="figure">
<paragraph id="A8441E53FFEEFFF3C59655C1C5B8E47C" blockId="3.[136,1181,2029,2050]" box="[136,1181,2029,2050]" pageId="3" pageNumber="254">
<emphasis id="9A8FC241FFEEFFF3C59655C1C1DBE47C" bold="true" box="[136,254,2029,2050]" pageId="3" pageNumber="254">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="6FFB65D0FFEEFFF3C46355C1C33EE47C" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[381,539,2029,2050]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="3" pageNumber="254" phylum="Tracheophyta" rank="species" species="greuteri">
<emphasis id="9A8FC241FFEEFFF3C46355C1C33EE47C" box="[381,539,2029,2050]" italics="true" pageId="3" pageNumber="254">Festuca greuteri</emphasis>
</taxonomicName>
(Granada, Sierra de Guillimona, puerto de la Losa, COFC 65825).
</paragraph>
</caption>
<caption id="FC844EDBFFE9FFF4C5965563C08FE466" ID-DOI="http://doi.org/10.5281/zenodo.13716740" ID-Zenodo-Dep="13716740" httpUri="https://zenodo.org/record/13716740/files/figure.png" pageId="4" pageNumber="255" startId="4.[136,229,1871,1892]" targetBox="[226,1376,194,1833]" targetPageId="4" targetType="figure">
<paragraph id="A8441E53FFE9FFF4C5965563C08FE466" blockId="4.[136,1455,1871,2072]" pageId="4" pageNumber="255">
<emphasis id="9A8FC241FFE9FFF4C5965563C1DAEB1A" bold="true" box="[136,255,1871,1892]" pageId="4" pageNumber="255">FIGURE 3.</emphasis>
<taxonomicName id="6FFB65D0FFE9FFF4C4195563C31BEB1A" box="[263,574,1871,1892]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="255" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="greuteri">
<emphasis id="9A8FC241FFE9FFF4C4195563C083EB1A" box="[263,422,1871,1892]" italics="true" pageId="4" pageNumber="255">Festuca greuteri</emphasis>
subsp.
<emphasis id="9A8FC241FFE9FFF4C4EC5563C31BEB1A" box="[498,574,1871,1892]" italics="true" pageId="4" pageNumber="255">greuteri</emphasis>
</taxonomicName>
(A–C and E–L) and
<taxonomicName id="6FFB65D0FFE9FFF4C60E5563C564EB1A" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[784,1089,1871,1892]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="255" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="camarolensis">
<emphasis id="9A8FC241FFE9FFF4C60E5563C253EB1A" box="[784,886,1871,1892]" italics="true" pageId="4" pageNumber="255">F. greuteri</emphasis>
subsp.
<emphasis id="9A8FC241FFE9FFF4C6DC5563C564EB1A" box="[962,1089,1871,1892]" italics="true" pageId="4" pageNumber="255">camarolensis</emphasis>
</taxonomicName>
(D).
<emphasis id="9A8FC241FFE9FFF4C1695563C5B4EB1A" bold="true" box="[1143,1169,1871,1892]" pageId="4" pageNumber="255">A)</emphasis>
Habit;
<emphasis id="9A8FC241FFE9FFF4C1C25563C5D1EB1A" bold="true" box="[1244,1268,1871,1892]" pageId="4" pageNumber="255">B)</emphasis>
Detail of the apex of the sheath and ligule in adaxial view;
<emphasis id="9A8FC241FFE9FFF4C70B555FC368EBF6" bold="true" box="[533,589,1907,1928]" pageId="4" pageNumber="255">C −D)</emphasis>
Spikelet;
<emphasis id="9A8FC241FFE9FFF4C7AF555FC3ECEBF6" bold="true" box="[689,713,1907,1928]" pageId="4" pageNumber="255">E)</emphasis>
Lower glume in abaxial view;
<emphasis id="9A8FC241FFE9FFF4C6E5555FC534EBF6" bold="true" box="[1019,1041,1907,1928]" pageId="4" pageNumber="255">F)</emphasis>
Upper glume in abaxial view;
<emphasis id="9A8FC241FFE9FFF4C05E555FC47EEBF6" bold="true" box="[1344,1371,1907,1928]" pageId="4" pageNumber="255">G)</emphasis>
Lemma in abaxial view;
<emphasis id="9A8FC241FFE9FFF4C43255BBC062EBD2" bold="true" box="[300,327,1943,1964]" pageId="4" pageNumber="255">H)</emphasis>
Palea in abaxial view;
<emphasis id="9A8FC241FFE9FFF4C73355BBC31BEBD2" bold="true" box="[557,574,1943,1964]" pageId="4" pageNumber="255">I)</emphasis>
Lodicule;
<emphasis id="9A8FC241FFE9FFF4C7B755BBC398EBD2" bold="true" box="[681,701,1943,1964]" pageId="4" pageNumber="255">J)</emphasis>
Sexual whorls;
<emphasis id="9A8FC241FFE9FFF4C64355BBC25DEBD2" bold="true" box="[861,888,1943,1964]" pageId="4" pageNumber="255">K)</emphasis>
Caryopsis in adaxial view;
<emphasis id="9A8FC241FFE9FFF4C19455BBC587EBD2" bold="true" box="[1162,1186,1943,1964]" pageId="4" pageNumber="255">L)</emphasis>
Caryopsis in abaxial view. Vouchers: Spain, Puerto de la Losa, Sierra de Guillimona, Granada, 4 June 2014,
<emphasis id="9A8FC241FFE9FFF4C6955597C54AEBAE" box="[907,1135,1979,2000]" italics="true" pageId="4" pageNumber="255">E. López &amp; G. Martínez</emphasis>
, COFC 61365 (
<taxonomicName id="6FFB65D0FFE9FFF4C0185597C1F1EB8A" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="255" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="greuteri">
<emphasis id="9A8FC241FFE9FFF4C0185597C44CEBAE" box="[1286,1385,1979,2000]" italics="true" pageId="4" pageNumber="255">F. greuteri</emphasis>
subsp.
<emphasis id="9A8FC241FFE9FFF4C59655F3C1F1EB8A" box="[136,212,2015,2036]" italics="true" pageId="4" pageNumber="255">greuteri</emphasis>
</taxonomicName>
); and Spain, Sierra de Camarolos, Málaga, 10 June 2015,
<emphasis id="9A8FC241FFE9FFF4C7E155F3C422EB8A" box="[767,1287,2015,2036]" italics="true" pageId="4" pageNumber="255">B. Cabezudo, J. A. Devesa, G. Martínez &amp; F. Soriguer,</emphasis>
COFC 62140 (
<taxonomicName id="6FFB65D0FFE9FFF4C08455F3C0B9E466" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="255" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="camarolensis">
<emphasis id="9A8FC241FFE9FFF4C08455F3C1F1E466" italics="true" pageId="4" pageNumber="255">F. greuteri</emphasis>
subsp.
<emphasis id="9A8FC241FFE9FFF4C4035A2FC0B9E466" box="[285,412,2051,2072]" italics="true" pageId="4" pageNumber="255">camarolensis</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="A8441E53FFE8FFF5C5A3528CC50CEE3B" blockId="5.[136,1452,160,833]" pageId="5" pageNumber="256">
<emphasis id="9A8FC241FFE8FFF5C5A3528CC091ECC7" bold="true" box="[189,436,160,185]" pageId="5" pageNumber="256">Taxonomic remarks:</emphasis>
—
<taxonomicName id="6FFB65D0FFE8FFF5C4CE528CC3A2ECC7" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[464,647,160,185]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="species" species="greuteri">
<emphasis id="9A8FC241FFE8FFF5C4CE528CC3A2ECC7" box="[464,647,160,185]" italics="true" pageId="5" pageNumber="256">Festuca greuteri</emphasis>
</taxonomicName>
shows some variability in the size of the floral parts (the lowest values of spikelets and lemmas were observed in some specimens from Sierra de Baza and Sierra de Tejeda), and in the diameter, stiffness and anatomical pattern (in cross-section) of the leaf blade. Usually, plants of the easternmost populations show rigid leaf blades, elliptical and with a continuous ring of sclerenchyma in cross-section (e.g., in Sierra de María, Sierra de Guillimona, and Sierra de Castril). The diameter of the leaf blades is particularly small in individuals from Sierra de Mágina (leaf blades in cross-sections not shown), but it is quite variable depending on the plant habitat (see the variation of the leaf cross-sections among populations in
<figureCitation id="30C002D6FFE8FFF5C65E5354C2A0EDEF" box="[832,901,376,401]" captionStart="FIGURE 4" captionStartId="5.[136,229,1927,1948]" captionTargetBox="[371,1212,1135,1849]" captionTargetId="figure-413@5.[357,1231,1130,1850]" captionTargetPageId="5" captionText="FIGURE 4. Leaf blades in cross-section of Festuca greuteri subsp. greuteri (A–F), and F.greuteri subsp.camarolensis (G–K).Provenance of the material: Sierra de la Sagra (A, F), Sierra de Baza (B), Sierra de Tejeda (E), Sierra de Guillimona (D), and Sierra de Camarolos (C, G–K)." figureDoi="http://doi.org/10.5281/zenodo.13716744" httpUri="https://zenodo.org/record/13716744/files/figure.png" pageId="5" pageNumber="256">Fig. 4</figureCitation>
). Plants of the western populations usually have more flaccid leaf blades, with a broken ring of sclerenchyma
<emphasis id="9A8FC241FFE8FFF5C62253B2C26FEDCA" box="[828,842,414,436]" italics="true" pageId="5" pageNumber="256">‒</emphasis>
sometimes on very decurrent strands
<emphasis id="9A8FC241FFE8FFF5C1F853B2C5D1EDCA" box="[1254,1268,414,436]" italics="true" pageId="5" pageNumber="256">‒</emphasis>
in cross-section (e.g., in Sierra de Camarolos,
<figureCitation id="30C002D6FFE8FFF5C4C053ECC375EDA7" box="[478,592,448,473]" captionStart="FIGURE 4" captionStartId="5.[136,229,1927,1948]" captionTargetBox="[371,1212,1135,1849]" captionTargetId="figure-413@5.[357,1231,1130,1850]" captionTargetPageId="5" captionText="FIGURE 4. Leaf blades in cross-section of Festuca greuteri subsp. greuteri (A–F), and F.greuteri subsp.camarolensis (G–K).Provenance of the material: Sierra de la Sagra (A, F), Sierra de Baza (B), Sierra de Tejeda (E), Sierra de Guillimona (D), and Sierra de Camarolos (C, G–K)." figureDoi="http://doi.org/10.5281/zenodo.13716744" httpUri="https://zenodo.org/record/13716744/files/figure.png" pageId="5" pageNumber="256">
Fig. 4G
<emphasis id="9A8FC241FFE8FFF5C72B53EEC366EDA6" box="[565,579,450,472]" italics="true" pageId="5" pageNumber="256">‒</emphasis>
J
</figureCitation>
, although this character combination also appears in some individuals from other mountains when the soils are humid), and obovate or V-shaped outline (
<figureCitation id="30C002D6FFE8FFF5C6C053C8C507ED83" box="[990,1058,484,509]" captionStart="FIGURE 4" captionStartId="5.[136,229,1927,1948]" captionTargetBox="[371,1212,1135,1849]" captionTargetId="figure-413@5.[357,1231,1130,1850]" captionTargetPageId="5" captionText="FIGURE 4. Leaf blades in cross-section of Festuca greuteri subsp. greuteri (A–F), and F.greuteri subsp.camarolensis (G–K).Provenance of the material: Sierra de la Sagra (A, F), Sierra de Baza (B), Sierra de Tejeda (E), Sierra de Guillimona (D), and Sierra de Camarolos (C, G–K)." figureDoi="http://doi.org/10.5281/zenodo.13716744" httpUri="https://zenodo.org/record/13716744/files/figure.png" pageId="5" pageNumber="256">Fig. 4</figureCitation>
). We observed that members of this species are usually glaucous and somewhat pruinose, but it is also a variable character (e.g., plants glaucous-pruinose are frequently found in Sierra de Guillimona, Sierra de Baza and Sierra de Tejeda).
</paragraph>
<paragraph id="A8441E53FFE8FFF5C5A3507CC071EF3F" blockId="5.[136,1452,160,833]" pageId="5" pageNumber="256">
The awn length, however, provides two variation patterns that are recognised here to define two subspecies. Plants with the second basal lemma bearing a short awn, up to 1.3(–1.5) mm, are included in
<taxonomicName id="6FFB65D0FFE8FFF5C1BA5058C46EEEF3" box="[1188,1355,628,653]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="greuteri">
subsp.
<emphasis id="9A8FC241FFE8FFF5C1EC5058C46EEEF3" box="[1266,1355,628,653]" italics="true" pageId="5" pageNumber="256">greuteri</emphasis>
</taxonomicName>
, and are distributed throughout the area of the species; on the contrary, plants with long-awned lemmas (usually more than
<quantity id="6F03B3B6FFE8FFF5C09650B4C191EEAB" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="256" unit="mm" value="1.5">1.5 mm</quantity>
in that of the second floret) are grouped in
<taxonomicName id="6FFB65D0FFE8FFF5C7845090C259EEAB" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[666,892,700,725]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="camarolensis">
subsp.
<emphasis id="9A8FC241FFE8FFF5C7F65090C259EEAB" box="[744,892,700,725]" italics="true" pageId="5" pageNumber="256">camarolensis</emphasis>
</taxonomicName>
, and are restricted to the Sierra de Camarolos and Sierra del Jobo (Málaga province), where it coexists (growing in the same area) with plants of the typical subspecies. Members of
<taxonomicName id="6FFB65D0FFE8FFF5C40B5128C0D0EF63" authorityName="Martinez-Sagarra &amp; Devesa" authorityYear="2019" box="[277,501,772,797]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="camarolensis">
subsp.
<emphasis id="9A8FC241FFE8FFF5C47F5128C0D0EF63" box="[353,501,772,797]" italics="true" pageId="5" pageNumber="256">camarolensis</emphasis>
</taxonomicName>
can have slightly larger floral parts (glumes and lemmas), although overlapping those of
<taxonomicName id="6FFB65D0FFE8FFF5C5B85104C068EF3F" box="[166,333,808,833]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="subSpecies" species="greuteri" subSpecies="greuteri">
subsp.
<emphasis id="9A8FC241FFE8FFF5C5EA5104C068EF3F" box="[244,333,808,833]" italics="true" pageId="5" pageNumber="256">greuteri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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