treatments-xml/data/1D/03/E8/1D03E8542C4FCE2CFC7BC2B39BEDFEA3.xml

<document id="DA73084188AAD88407B1A3C9489648CC" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C4FCE2CFC7BC2B39BEDFEA3" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Lophoplax bicristata Tesch 1918" docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="338" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1DDC1C336237E11D878A2D59793CA624">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="05B95EBD16A73A44B4F1216086398BEB">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="BBA2D4DA2E5724B4F6114F4AC31A380D">Ahyong,</mods:namePart>
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<mods:namePart id="6CB1006C3E2448599EC210DBE8CCD910">Shane T.</mods:namePart>
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<paragraph id="951559422C4FCE29FC7BC2B39B79FD23" blockId="3.[945,1318,701,757]" box="[945,1318,701,725]" pageId="3" pageNumber="338">
<heading id="CE5DEE2E2C4FCE29FC7BC2B39B79FD23" bold="true" box="[945,1318,701,725]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="338" reason="2">
<taxonomicName id="52AA22C12C4FCE29FC7BC2B39B79FD23" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[945,1318,701,725]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4FCE29FC7BC2B39B79FD23" bold="true" box="[945,1318,701,725]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FC7BC2B39AC8FD23" bold="true" box="[945,1175,701,725]" italics="true" pageId="3" pageNumber="338">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4FCE29FB55C2B39B79FD23" author="Tesch JJ" box="[1183,1318,701,725]" pageId="3" pageNumber="338" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
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<paragraph id="951559422C4FCE29FBE5C2D39AF6FD03" blockId="3.[945,1318,701,757]" box="[1071,1193,733,757]" pageId="3" pageNumber="338">
(
<figureCitation id="0D9145C72C4FCE29FBFDC2D39ADAFD03" box="[1079,1157,733,757]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2</figureCitation>
, 
<figureCitation id="0D9145C72C4FCE29FB58C2D39AC0FD03" box="[1170,1183,733,757]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FCE7C3139BF5FBE3" pageId="3" pageNumber="338" type="materials_examined">
<paragraph id="951559422C4FCE29FCE7C3139BF5FBE3" blockId="3.[813,1452,797,1045]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C3139A52FCC3" bold="true" box="[813,1037,797,821]" pageId="3" pageNumber="338">Material examined.</emphasis>
<materialsCitation id="25C2531F2C4FCE29FBDFC3109A8CFC43" collectingDate="1899-12-16" collectingDateMax="1899-12-18" collectingDateMin="1899-12-16" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="90" latitude="-5.6083336" location="Nuhu Jaan" longLatPrecision="130" longitude="132.92" municipality="Kei Islands" pageId="3" pageNumber="338" specimenCode="D103002" specimenCount="1" typeStatus="lectotype">
<typeStatus id="4A11E7E02C4FCE29FBDFC3109ADFFCC0" box="[1045,1152,798,822]" pageId="3" pageNumber="338" type="lectotype">Lectotype</typeStatus>
(here designated): 
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<specimenCode id="C50CF1392C4FCE29FCA5C3339D89FCA3" box="[879,982,829,853]" collectionCode="RMNH-ZMA" pageId="3" pageNumber="338">D103002</specimenCode>
, male (5.8 × 
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), 
<quantity id="5252F4A72C4FCE29FB04C3339B6EFCA3" box="[1230,1329,829,853]" metricMagnitude="3" metricUnit="m" metricValue="3.7014912" pageId="3" pageNumber="338" unit="mi" value="2.3">2.3 miles</quantity>
from north point of 
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, northwest of 
<collectingMunicipality id="7571C3382C4FCE29FB66C3539B72FC83" box="[1196,1325,861,885]" pageId="3" pageNumber="338">Kei Islands</collectingMunicipality>
, 
<collectingCountry id="EDBD19D22C4FCE29FAF6C3539BF7FC83" box="[1340,1448,861,885]" name="Indonesia" pageId="3" pageNumber="338">Indonesia</collectingCountry>
, 
<geoCoordinate id="F09E3F852C4FCE29FCE7C3729DD7FC63" box="[813,904,892,917]" degrees="5" direction="south" minutes="36.5" orientation="latitude" pageId="3" pageNumber="338" precision="92" value="-5.6083336">5°36.5′S</geoCoordinate>
<geoCoordinate id="F09E3F852C4FCE29FC5AC3729A55FC63" box="[912,1034,892,917]" degrees="132" direction="east" minutes="55.2" orientation="longitude" pageId="3" pageNumber="338" precision="92" value="132.92">132°55.2′E</geoCoordinate>
, 
<quantity id="5252F4A72C4FCE29FBDDC3739A0EFC63" box="[1047,1105,893,917]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="338" unit="m" value="90.0">
<elevation id="1E87BE712C4FCE29FBDDC3739A0EFC63" box="[1047,1105,893,917]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="338" unit="m" value="90.0">90 m</elevation>
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, 
<collectingCounty id="7C7421CE2C4FCE29FB94C3739B2BFC63" box="[1118,1396,893,917]" pageId="3" pageNumber="338">Siboga Expedition station</collectingCounty>
260, coll. 
<collectorName id="385F3C942C4FCE29FCA8C3909D8EFC43" box="[866,977,925,949]" pageId="3" pageNumber="338">M. Weber</collectorName>
, 
<date id="E1147F822C4FCE29FC16C3939A8CFC43" box="[988,1235,925,949]" pageId="3" pageNumber="338" value="1899-12-16" valueMax="1899-12-18" valueMin="1899-12-16">
<collectingDate id="F150866A2C4FCE29FC16C3939A8CFC43" box="[988,1235,925,949]" pageId="3" pageNumber="338" value="1899-12-16" valueMax="1899-12-18" valueMin="1899-12-16">16–18 December 1899</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="25C2531F2C4FCE29FB17C3939BF9FBE3" collectingDate="1899-06-10" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="59" location="Borneo Bank" municipality="Strait of Makassar" pageId="3" pageNumber="338" specimenCode="D103001" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="4A11E7E02C4FCE29FB17C3939B31FC43" box="[1245,1390,925,949]" pageId="3" pageNumber="338" type="paralectotype">Paralectotype</typeStatus>
(here designated): 
<collectionCode id="F3BBC1872C4FCE29FC7DC3B09A14FC20" box="[951,1099,958,982]" pageId="3" pageNumber="338">RMNH-ZMA</collectionCode>
<specimenCode id="C50CF1392C4FCE29FB99C3B39AE6FC23" box="[1107,1209,957,981]" collectionCode="RMNH-ZMA" pageId="3" pageNumber="338">D103001</specimenCode>
, male (4.5 × 
<quantity id="5252F4A72C4FCE29FA83C3B39BC0FC23" box="[1353,1439,957,981]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="338" unit="mm" value="3.7">3.7 mm</quantity>
), 
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, 
<collectingMunicipality id="7571C3382C4FCE29FC06C3D39AC9FC03" box="[972,1174,989,1013]" pageId="3" pageNumber="338">Strait of Makassar</collectingMunicipality>
, 
<collectingCountry id="EDBD19D22C4FCE29FB69C3D39B52FC03" box="[1187,1293,989,1013]" name="Indonesia" pageId="3" pageNumber="338">Indonesia</collectingCountry>
, 
<quantity id="5252F4A72C4FCE29FAD0C3D39B0BFC03" box="[1306,1364,989,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="338" unit="m" value="59.0">
<elevation id="1E87BE712C4FCE29FAD0C3D39B0BFC03" box="[1306,1364,989,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="338" unit="m" value="59.0">59 m</elevation>
</quantity>
, 
<collectingCounty id="7C7421CE2C4FCE29FAABC3D39DAEFBE3" pageId="3" pageNumber="338">Siboga Expedition station</collectingCounty>
77, coll. coll. 
<collectorName id="385F3C942C4FCE29FB59C3F09B5BFBE3" box="[1171,1284,1021,1045]" pageId="3" pageNumber="338">M. Weber</collectorName>
, 
<date id="E1147F822C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">
<collectingDate id="F150866A2C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">10 June 1899</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FCE7C4339B6BF9E3" pageId="3" pageNumber="338" type="diagnosis">
<paragraph id="951559422C4FCE29FCE7C4339B6BF9E3" blockId="3.[811,1452,1085,1557]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C4339DC1FBA3" bold="true" box="[813,926,1085,1109]" pageId="3" pageNumber="338">Diagnosis.</emphasis>
Carapace frontal margin almost straight, granulate, not distinctly produced anteriorly, lateral lobe very low but just visible (
<figureCitation id="0D9145C72C4FCE29FC25C4739A2EFB63" box="[1007,1137,1149,1173]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); epigastric areole low but visible; protogastric areolet distinct, smooth (
<figureCitation id="0D9145C72C4FCE29FAE9C4939BC1FB43" box="[1315,1438,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); metabranchial region with prominent raised granules (
<figureCitation id="0D9145C72C4FCE29FA4AC4B39D0EFB03" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
); median lobe on posterior margin of epistome obtusely triangular (
<figureCitation id="0D9145C72C4FCE29FC65C4F39A54FAE3" box="[943,1035,1277,1301]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3B</figureCitation>
). Distal margin of ischium of third maxilliped smooth (
<figureCitation id="0D9145C72C4FCE29FBCBC5139A09FAC3" box="[1025,1110,1309,1333]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3A</figureCitation>
). Cheliped carpus dorsal surface with subquadrate areole with crenulate ridged margins (
<figureCitation id="0D9145C72C4FCE29FA4AC5339D2DFA83" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A–C</figureCitation>
); proximal basal part of male chela almost smooth or with very low tubercles (
<figureCitation id="0D9145C72C4FCE29FB96C5739AEBFA63" box="[1116,1204,1405,1429]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2D</figureCitation>
). P2–4 merus extensor margin distinctly crenulated, with low lobes. P5 merus unarmed (
<figureCitation id="0D9145C72C4FCE29FC6AC5B39A7FFA23" box="[928,1056,1469,1493]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3C, D</figureCitation>
). Male thoracic sternum anteriorly relatively wide (
<figureCitation id="0D9145C72C4FCE29FC16C5D39A6FFA03" box="[988,1072,1501,1525]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2E</figureCitation>
). G1 gently sinuous, distal strongly recurved (
<figureCitation id="0D9145C72C4FCE29FC51C5F39A5BF9E3" box="[923,1028,1533,1557]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3F–I</figureCitation>
). G2 about ¼ length of G1.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE2CFCE7C6339BEDFEA3" lastPageId="6" lastPageNumber="341" pageId="3" pageNumber="338" type="description">
<paragraph id="951559422C4FCE2FFCE7C6339A93F7E3" blockId="3.[813,1452,1597,2101]" lastBlockId="5.[813,1453,1725,2069]" lastPageId="5" lastPageNumber="340" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C6339DE8F9A3" bold="true" box="[813,951,1597,1621]" pageId="3" pageNumber="338">Description.</emphasis>
Carapace subquadrate; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like, not concealing margins; regions well demarcated with few areoles distinct: protogastric areole transversely ovate, distinct; epigastric region low, visible when denuded, not areolate; hepatic region with 1 tubercle, no areole visible; cardiac region raised, covered with granules, setose, not areolate, lateral parts with raised patches of granules, but no visible cardio-branchial areole; gastro-cardiac region depressed, with grooves visible but not deep (
<figureCitation id="0D9145C72C4FCE29FAEFC7539BC1F883" box="[1317,1438,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); intestinal region convex, forming rounded areole, adjacent surfaces gently raised, with small granules, setose, separated from cristate posterior carapace margin by wide space, no obvious channel (
<figureCitation id="0D9145C72C4FCE29FC3DC7D39A29F803" box="[1015,1142,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); meso- and metabranchial regions each with 1 gently raised area covered with granules; metabranchial region with tubercle just posterior to raised patch of granules (
<figureCitation id="0D9145C72C49CE2FFE84C6B39FF4F923" box="[334,427,1725,1749]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Subhepatic and suborbital regions slightly rugose to almost smooth; pterygostomial region with small granules. Frontal margin weakly bilobed, almost straight, with shallow median concavity, margins granulate; lateral lobe, small, low, just visible in dorsal view (
<figureCitation id="0D9145C72C49CE2FFFB3C7539E8FF883" box="[121,208,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Supraorbital margin short, sinuous, with distinct submedian lobe, without fissure or cleft (
<figureCitation id="0D9145C72C49CE2FFD8FC7739CC0F863" box="[581,671,1917,1941]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Orbit dorsal, supraorbital margin, eyes clearly visible in dorsal view; eyes freely movable, ocular peduncle relatively long, filling orbit (
<figureCitation id="0D9145C72C49CE2FFECAC7D39F06F803" box="[256,345,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Anterolateral margin with 4 small, sharp teeth (including external orbital tooth), external orbital tooth broader, next 3 teeth acutely triangular, tip slightly enlarged (
<figureCitation id="0D9145C72C49CE2FFC52C6D39A53F903" box="[920,1036,1757,1781]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="5" pageNumber="340">Fig. 2A, B</figureCitation>
). Posterolateral margin uneven, with 2 tubercles, each with smaller basal granules; margins gently converging towards posterior carapace margin (
<figureCitation id="0D9145C72C49CE2FFA8EC7139BC1F8C3" box="[1348,1438,1821,1845]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally. Basal antennal article subquadrate; flagellum entering orbital hiatus. Epistome with median lobe of posterior margin obtusely triangular, no obvious fissure in frontal view; lateral parts sinuous with small cleft (
<figureCitation id="0D9145C72C49CE2FFA81C7B39BC1F823" box="[1355,1438,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340">Fig. 3B</figureCitation>
). Endostome distinct, sloping gradually posteriorly into buccal cavern, endostomial ridges low, short.
</paragraph>
<caption id="C1D509CA2C48CE2EFF4DC7FD9B55F7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682765" ID-Zenodo-Dep="14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="4" pageNumber="339" startId="4.[135,170,2035,2057]" targetBox="[162,1447,189,2020]" targetPageId="4" targetType="figure">
<paragraph id="951559422C48CE2EFF4DC7FD9B55F7B5" blockId="4.[135,1474,2035,2115]" pageId="4" pageNumber="339">
Fig. 2. 
<taxonomicName id="52AA22C12C48CE2EFF04C7FD9C49F7FE" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[206,534,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C48CE2EFF04C7FD9FC8F7FE" box="[206,407,2035,2056]" italics="true" pageId="4" pageNumber="339">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C48CE2EFE57C7FD9C49F7FE" author="Tesch JJ" box="[413,534,2035,2057]" pageId="4" pageNumber="339" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon.
</paragraph>
</caption>
<caption id="C1D509CA2C49CE2FFFBBC6179C11F973" ID-DOI="http://doi.org/10.5281/zenodo.14682767" ID-Zenodo-Dep="14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340" startId="5.[113,148,1561,1583]" targetBox="[115,1452,189,1546]" targetPageId="5" targetType="figure">
<paragraph id="951559422C49CE2FFFBBC6179C11F973" blockId="5.[113,1452,1561,1670]" pageId="5" pageNumber="340">
Fig. 3. 
<taxonomicName id="52AA22C12C49CE2FFF76C6179C52F9D8" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[188,525,1561,1583]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C49CE2FFF76C6179FD6F9D8" box="[188,393,1561,1582]" italics="true" pageId="5" pageNumber="340">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C49CE2FFE58C6179C52F9D8" author="Tesch JJ" box="[402,525,1561,1583]" pageId="5" pageNumber="340" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm.
</paragraph>
</caption>
<paragraph id="951559422C4ACE2CFF42C0B39FC7FEA3" blockId="6.[136,776,189,341]" pageId="6" pageNumber="341">
Third maxilliped with ischium relatively short, subquadrate, length 1.2× width, with shallow median oblique sulcus; merus subquadrate, anterolateral angle gently auriculiform. Exopod relatively stout, tip reaching to just before distal edge of merus (
<figureCitation id="0D9145C72C4ACE2CFEFEC1339FD3FEA3" box="[308,396,317,341]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C1739F15FD63" blockId="6.[136,776,381,661]" pageId="6" pageNumber="341">
Male thoracic sternum surface relatively smooth, slightly rugose; sternites 1 and 2 completely fused, wide, lateral margins gently concave; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 almost straight (
<figureCitation id="0D9145C72C4ACE2CFD6CC1F39CA4FDE3" box="[678,763,509,533]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Fig. 2E</figureCitation>
). Sternite 8 clearly visible as subrectangular plate adjacent to pleonal somite 2 when pleon closed (
<figureCitation id="0D9145C72C4ACE2CFDF4C2339CCCFDA3" box="[574,659,573,597]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior third of thoracic somite 5.
</paragraph>
<paragraph id="951559422C4ACE2CFF42C2B39FBFFB63" blockId="6.[136,776,701,1174]" pageId="6" pageNumber="341">
Adult male chelipeds almost symmetrical, stout (
<figureCitation id="0D9145C72C4ACE2CFD60C2B39EC3FD03" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1A, B</figureCitation>
). Basis-ischium fused but suture visible; extensor margins uneven but unarmed (
<figureCitation id="0D9145C72C4ACE2CFEBAC2F39F9CFCE3" box="[368,451,765,789]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with tubercles and granules; inner distal angle dentiform with subdistal angle tuberculate (
<figureCitation id="0D9145C72C4ACE2CFEC2C3539F07FC83" box="[264,344,861,885]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Carpus surface granulate; dorsal surface with subquadrate areole with crenulate ridged margins; inner distal angle with low sharp tooth (
<figureCitation id="0D9145C72C4ACE2CFDC9C3939C24FC43" box="[515,635,925,949]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1A–C</figureCitation>
). Chela with dorsal surface covered with granules; inner surface almost smooth; median and ventral surfaces almost smooth or with very low tubercles; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, slightly shorter than palm, pigmented brown along distal half; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C4ACE2CFE90C4739F8CFB63" box="[346,467,1149,1173]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1–t4 = anterolateral teeth 1–4." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C4B39CCAF9C3" blockId="6.[136,776,1213,1589]" pageId="6" pageNumber="341">
Ambulatory legs long, P4 longest, about 1.8× carapace width; P4 merus length about 5× height, distinctly longer than half carapace width; outer surface slightly rugose to almost smooth; coxa rounded; merus elongate, P2–4 extensor margin not cristate with low, broad, well-spaced setose prominences, P5 merus extensor margin smooth, fully setose; surface of P2–5 gently convex, without median swelling or ridge; P2–5 carpus relatively smooth, with very low submarginal ridge, lowest in P5; propodus laterally flattened, unarmed, relatively long in P2–4, shortest in P5; P2–4 dactylus falciform, unarmed, P5 dactylus shorter, slightly upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C4ACE2CFE1FC6139C69F9C3" box="[469,566,1565,1589]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, D–F, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Figs. 2A</figureCitation>
, 
<figureCitation id="0D9145C72C4ACE2CFD89C6139CD7F9C3" box="[579,648,1565,1589]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">3C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C6539FD1F8A3" blockId="6.[136,776,1629,1877]" pageId="6" pageNumber="341">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely much narrower than somite 1, trapezoidal; somite 3 broadly subtrapezoidal, widest, somites 4–6 trapezoidal, gradually decreasing in width with somite 6 almost rectangular; telson slightly wider than long, rounded, slightly narrowly distally, lateral margins convex, longer than somite 6 (
<figureCitation id="0D9145C72C4ACE2CFEE7C7339FDDF8A3" box="[301,386,1853,1877]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C7739F80F803" blockId="6.[136,775,1917,2037]" pageId="6" pageNumber="341">
G1 relatively stout, gently sinuous, distally tapering to narrowly rounded tip, strongly hooked inwards; subdistal surface with 2 long stout setae (
<figureCitation id="0D9145C72C4ACE2CFE32C7B39C38F823" box="[504,615,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3F–I</figureCitation>
). G2 sigmoid, about ¼ length of G1 (
<figureCitation id="0D9145C72C4ACE2CFE4EC7D39F8CF803" box="[388,467,2013,2037]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; C–E = 1.0 mm; F, J = 0.25 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3J</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C8139BEDFEA3" blockId="6.[136,356,2077,2101]" lastBlockId="6.[835,1475,189,341]" pageId="6" pageNumber="341">
Females not known. 
<emphasis id="A7DE85502C4ACE2CFC89C0B39DEBFF23" bold="true" box="[835,948,189,213]" pageId="6" pageNumber="341">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4ACE2CFC74C0B39AFBFF23" authorityName="Tesch" authorityYear="1918" box="[958,1188,189,213]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4ACE2CFC74C0B39AFBFF23" box="[958,1188,189,213]" italics="true" pageId="6" pageNumber="341">Lophoplax bicristata</emphasis>
</taxonomicName>
is only known from 
<specimenCount id="83AC92CB2C4ACE2CFA53C0B09DDDFF03" count="2" pageId="6" pageNumber="341" type="male">two males</specimenCount>
<materialsCitation id="25C2531F2C4ACE2CFC43C0D39BEDFEA3" country="Indonesia" county="They" elevation="75" location="Kei Islands" municipality="Both" pageId="6" pageNumber="341" specimenCount="1">
collected from relatively near each other (
<location id="90750F992C4ACE2CFA82C0D39B9CFF03" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFA82C0D39B9CFF03" box="[1352,1475,221,245]" country="Indonesia" county="They" municipality="Both" name="Kei Islands" pageId="6" pageNumber="341">Kei Islands</location>
and 
<location id="90750F992C4ACE2CFCBFC0F09A54FEE3" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFCBFC0F09A54FEE3" box="[885,1035,253,277]" country="Indonesia" county="They" municipality="Both" name="Borneo Bank" pageId="6" pageNumber="341">Borneo Bank</location>
, 
<collectingCountry id="EDBD19D22C4ACE2CFBD3C0F39AD7FEE3" box="[1049,1160,253,277]" name="Indonesia" pageId="6" pageNumber="341">Indonesia</collectingCountry>
). 
<collectingMunicipality id="7571C3382C4ACE2CFB57C0F39A8DFEE3" box="[1181,1234,253,277]" pageId="6" pageNumber="341">Both</collectingMunicipality>
are mature and were covered with silt and mud, suggesting they live on soft substrates. 
<collectingCounty id="7C7421CE2C4ACE2CFC76C1339DACFEA3" box="[956,1011,317,341]" pageId="6" pageNumber="341">They</collectingCounty>
were collected from depths of 
<quantity id="5252F4A72C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">
<elevation id="1E87BE712C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">59–90 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>