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<document id="9A4ECCCB1A7B37ADBA3BC193892EB58D" ID-DOI="10.5281/zenodo.157462" ID-GBIF-Dataset="a9ccfc65-1459-414b-8a99-c99649f83d51" ID-ISSN="1175­5326" ID-Zenodo-Dep="157462" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461019330970" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2004" docId="160487E5FA6AFFBCFE95FD04FE87FC11" docLanguage="en" docName="zt00538.pdf" docOrigin="Zootaxa 538" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pneuminion impressum Perkins, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="EA3DFF9DFA66FFB1FF9DF83AFFBAFFCF" masterDocTitle="A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="13" updateTime="1698569824382" updateUser="plazi">
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<mods:title id="6F3A4A3276F03000BA1FE54529F9B91C">A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="2381CD16C0ABEA1F3725A6E22C0C161B" type="personal">
<mods:role id="8B51B8F66F446F845BE218CDBB319123">
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<mods:namePart id="520C1F6A0129E41287FDBA30477D8F59">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="DE2880E8B8DC0C1A6218C38D0EC99518">text</mods:typeOfResource>
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<mods:title id="0C54D7B01689C005A83AA92930910BAD">Zootaxa</mods:title>
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<mods:part id="FF1E4FB364756FE06B6720935AAFFF28">
<mods:date id="028B14B7C6B97518F30A6581D08C2D15">2004</mods:date>
<mods:detail id="A625E4F568F1FE75FEA303D5863CE3A2" type="volume">
<mods:number id="D480B7292098407C931E7390B8909D08">538</mods:number>
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<mods:classification id="BEFC4D22095DE0FBF3A3DB781AFF63D2">journal article</mods:classification>
<mods:identifier id="AADB2033964B40F8E93BDBAAF1560B70" type="DOI">10.5281/zenodo.157462</mods:identifier>
<mods:identifier id="5D9E6534E12CA22A7D3763B5218F006E" type="GBIF-Dataset">a9ccfc65-1459-414b-8a99-c99649f83d51</mods:identifier>
<mods:identifier id="453D34566FC48A08341000D3B2C20990" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="479D3236BBD86BE01D87E1E5C9A54EFB" type="Zenodo-Dep">157462</mods:identifier>
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<treatment id="160487E5FA6AFFBCFE95FD04FE87FC11" ID-DOI="http://doi.org/10.5281/zenodo.6271135" ID-GBIF-Taxon="119602187" ID-Zenodo-Dep="6271135" LSID="urn:lsid:plazi:treatment:160487E5FA6AFFBCFE95FD04FE87FC11" httpUri="http://treatment.plazi.org/id/160487E5FA6AFFBCFE95FD04FE87FC11" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="D6B76578FA6AFFBDFE95FD04FE54FAB4" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="9E1236F3FA6AFFBDFE95FD04FD7AFA97" blockId="12.[264,704,1342,1403]" box="[264,704,1342,1368]" pageId="12" pageNumber="13">
<heading id="C55A819FFA6AFFBDFE95FD04FD7AFA97" bold="true" box="[264,704,1342,1368]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="ACD9EAE1FA6AFFBDFE95FD04FD7AFA97" bold="true" box="[264,704,1342,1368]" pageId="12" pageNumber="13">
<taxonomicName id="59AD4D70FA6AFFBDFE95FD04FD92FA97" ID-CoL="4KKV9" box="[264,552,1342,1368]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="impressum" status="sp. nov.">
<emphasis id="ACD9EAE1FA6AFFBDFE95FD04FD92FA97" bold="true" box="[264,552,1342,1368]" italics="true" pageId="12" pageNumber="13">Pneuminion impressum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7EA579AFA6AFFBDFDACFD04FD7AFA97" box="[561,704,1342,1368]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E1236F3FA6AFFBDFE95FD5BFE54FAB4" blockId="12.[264,704,1342,1403]" box="[264,494,1377,1403]" pageId="12" pageNumber="13">
(
<figureCitation id="06962A76FA6AFFBDFE8DFD5BFE35FAB4" box="[272,399,1377,1403]" captionStart="FIGURE 11" captionStartId="24.[264,368,1520,1544]" captionTargetBox="[294,1292,310,1512]" captionTargetId="figure@24.[294,1292,308,1513]" captionTargetPageId="24" captionText="FIGURE 11. Pneuminion impressum holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169057/files/figure.png" pageId="12" pageNumber="13">Figures 11</figureCitation>
,
<figureCitation id="06962A76FA6AFFBDFE01FD5BFE06FAB4" box="[412,444,1377,1403]" captionStart="FIGURE 20" captionStartId="33.[264,368,1323,1347]" captionTargetBox="[289,1280,330,1276]" captionTargetId="figure@33.[264,1323,326,1299]" captionTargetPageId="33" captionText="FIGURE 20. Pneuminion impressum. Aedeagus of holotype." httpUri="https://zenodo.org/record/157478/files/figure.png" pageId="12" pageNumber="13">20</figureCitation>
,
<figureCitation id="06962A76FA6AFFBDFE55FD5BFE5CFAB4" box="[456,486,1377,1403]" captionStart="FIGURES 26 28" captionStartId="36.[264,383,1835,1859]" captionTargetBox="[275,1306,310,1827]" captionTargetId="figure@36.[275,1307,308,1830]" captionTargetPageId="36" captionText="FIGURES 26 28. Pneuminion distribution maps. — 26. P. balfourbrownei. — 27. P. impressum. — 28. P. nanum." httpUri="https://zenodo.org/record/169060/files/figure.png" pageId="12" pageNumber="13">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBDFE95FDA4FD4DF9C6" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="9E1236F3FA6AFFBDFE95FDA4FD4DF9C6" blockId="12.[264,1324,1438,1905]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFE95FDA4FE05FA77" bold="true" box="[264,447,1438,1464]" pageId="12" pageNumber="13">
<typeStatus id="41168851FA6AFFBDFE95FDA4FEF9FA77" box="[264,323,1438,1464]" pageId="12" pageNumber="13">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="41168851FA6AFFBDFE4FFDA5FD85FA76" box="[466,575,1439,1465]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="E6BA7663FA6AFFBDFD31FDA5FCF2FA76" box="[684,840,1439,1465]" name="South Africa" pageId="12" pageNumber="13">South Africa</collectingCountry>
: W. Cape, Dwarsberge,
<geoCoordinate id="FB995034FA6AFFBDFBFEFDA5FB0CFA76" box="[1123,1206,1439,1465]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="555" value="-34.02">34.02S</geoCoordinate>
­
<geoCoordinate id="FB995034FA6AFFBDFB4DFDA5FA9CFA76" box="[1232,1318,1439,1465]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="555" value="19.01">19.01E</geoCoordinate>
, sifted moss,
<date id="EA131033FA6AFFBDFE3BFDFDFD95FA2E" box="[422,559,1479,1505]" pageId="12" pageNumber="13" value="1973-11-15">15.XI.1973</date>
, Endrödy­Younga (#254); deposited in the
<collectionCode id="F8BCAE36FA6AFFBDFBD4FDFDFB24FA2E" box="[1097,1182,1479,1505]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="12" pageNumber="13">TMSA</collectionCode>
.
<typeStatus id="41168851FA6AFFBDFB21FDFDFA9DFA2E" box="[1212,1319,1479,1505]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
: Same data as
<typeStatus id="41168851FA6AFFBDFE35FDD5FDA8F9C6" box="[424,530,1519,1545]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, (
<specimenCount id="88ABFD7AFA6AFFBDFDB4FDD5FD37F9C6" box="[553,653,1519,1545]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
<collectionCode id="F8BCAE36FA6AFFBDFD09FDD5FD51F9C6" box="[660,747,1519,1545]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="12" pageNumber="13">TMSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBDFEA5FE2CFDF5F966" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="9E1236F3FA6AFFBDFEA5FE2CFDF5F966" blockId="12.[264,1324,1438,1905]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFEA5FE2CFE0DF9FF" bold="true" box="[312,439,1558,1584]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
A very distinct species, recognized by the deeply impressed and wide pronotal foveae, the rugulose head and pronotum, and the highly modified elytra with a saddle and each elytron with five unilinear rows of setae (
<figureCitation id="06962A76FA6AFFBDFC54FE5DFB9BF94E" box="[969,1057,1639,1665]" captionStart="FIGURE 11" captionStartId="24.[264,368,1520,1544]" captionTargetBox="[294,1292,310,1512]" captionTargetId="figure@24.[294,1292,308,1513]" captionTargetPageId="24" captionText="FIGURE 11. Pneuminion impressum holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169057/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
). The basic aedeagal form is unique in the genus.
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBCFEA5FE8CFECCFC41" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="9E1236F3FA6AFFBDFEA5FE8CFCBDF936" blockId="12.[264,1324,1438,1905]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFEA5FE8CFE6AF91F" bold="true" box="[312,464,1718,1744]" pageId="12" pageNumber="13">Description.</emphasis>
Size (mm x 100; length/width): body (length to elytral apices) 155/70, head 30/44, pronotum 41/59, elytra 100/70.
</paragraph>
<paragraph id="9E1236F3FA6AFFBDFEA5FF3DFB03F8BE" blockId="12.[264,1324,1438,1905]" pageId="12" pageNumber="13">
Dorsum dark brown to brown, legs and maxillary palpi brown to testaceous; dorsal setae very conspicuous on head and pronotum, whitish. Dorsum of head and pronotum rugulose, dull. Frons with fovea in front of ocellus deep, ca.
<date id="EA131033FA6AFFBDFC50FF6DFC51F8BE" box="[973,1003,1879,1905]" pageId="12" pageNumber="13">5x</date>
ocellus diameter.
</paragraph>
<paragraph id="9E1236F3FA6BFFBCFEA5F926FE50FE61" blockId="13.[264,1323,284,990]" pageId="13" pageNumber="14">Pronotal postocular emarginations deeper and less symmetrical in females than males. Pronotum dull, impressions very deep and large, midlongitudinal impression as wide as eye, reliefs quite convex; setae short, denser and more slender on reliefs, becoming sparser and wider laterally.</paragraph>
<paragraph id="9E1236F3FA6BFFBCFEA5F986FE81FCD9" blockId="13.[264,1323,284,990]" pageId="13" pageNumber="14">Elytra with shallow transverse depression (“saddle”) at basal 0.3. Each elytron with five unilinear rows of overlapping, whitish setae, rows present on sutural interval and intervals 2, 4, 6, and 7, rows on intervals 2 and 4 interrupted over a short distance at saddle; interval 1 with short row of setae between base and saddle, and interval 5 with short row at base, otherwise odd­numbered intervals smooth, lacking setae; serial punctures moderately large, larger than smooth intervals separating them, each with a short indistinct seta, setae not overlapping and not forming lines; granules at bases of setae very small, if perceptible; sutural margin raised between saddle and apices; explanate margin very narrow.</paragraph>
<paragraph id="9E1236F3FA6BFFBCFEA5FB1EFECCFC41" blockId="13.[264,1323,284,990]" pageId="13" pageNumber="14">
Aedeagus: main­piece length 36, total length 45; parameres comparatively short, setae sparse, grouped at apex; duct joined with main­piece over basal 2/3, apical 2/3 thicker (
<figureCitation id="06962A76FA6BFFBCFE8CFB4EFED2FC41" box="[273,360,884,910]" captionStart="FIGURE 20" captionStartId="33.[264,368,1323,1347]" captionTargetBox="[289,1280,330,1276]" captionTargetId="figure@33.[264,1323,326,1299]" captionTargetPageId="33" captionText="FIGURE 20. Pneuminion impressum. Aedeagus of holotype." httpUri="https://zenodo.org/record/157478/files/figure.png" pageId="13" pageNumber="14">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6BFFBCFEA5FBA6FE87FC11" pageId="13" pageNumber="14" type="etymology">
<paragraph id="9E1236F3FA6BFFBCFEA5FBA6FE87FC11" blockId="13.[264,1323,284,990]" pageId="13" pageNumber="14">
<emphasis id="ACD9EAE1FA6BFFBCFEA5FBA6FE7EFC79" bold="true" box="[312,452,924,950]" pageId="13" pageNumber="14">Etymology.</emphasis>
Named in reference to the deeply impressed pronotal and elytral sculpture.
</paragraph>
</subSubSection>
</treatment>
</document>