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<document id="7513A4671D5C1840601B432070847C94" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657304FFFA7FF8EFD0BBFBD0B61" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Lebia (Metalebia) laterolucida Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="255" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0D8D6428DE6C2DA269F548278DD2123D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="699ADE61A2559429B59B7AFF3E282910">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:name id="426732699AA29FDA3663B4EE41EEE5CB" type="personal">
<mods:role id="AF03FC1EEA32F889AF2E7DABE0C0E40F">
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<mods:namePart id="17C4AE54CCB6C69A165C89660B536031">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="BC15B0A075BE06BFAB25A8C8A74B2014">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="076A122BF3CEE4D8C6FFDFFA49B73CB6">Rainio, Johanna</mods:namePart>
<mods:affiliation id="A564DD2EB5A0A09798D234480A07A6DB">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<mods:title id="88B98D8E4535FFCE2A9F898A2F04F2BD">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="22EE43897D6A15DD448C075E8B177F03">
<mods:date id="5B9304FC553554785704A007B9A26352">2016</mods:date>
<mods:detail id="157F170002E67F694AE5DE7B41CB9BB9" type="pubDate">
<mods:number id="A192072AA1134F19BA35DB0A68DBBC26">2016-04-29</mods:number>
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<mods:detail id="08A125DA468FB511FD8910B770F15873" type="volume">
<mods:number id="241736DEC6E533B239E5647948B45E48">63</mods:number>
</mods:detail>
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<mods:number id="7E0C3B65881DD89A61B40350FAE09E62">7</mods:number>
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<mods:classification id="0AA3C956BDB7C62899BF451D9BDBAA5D">journal article</mods:classification>
<mods:identifier id="0402856E2808771BE3BE8656ACB03725" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="1234B0957634F1B26F1B3403EF771440" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="DCC3446544015FBB0F4E06E38F316109" type="Zenodo-Dep">13799439</mods:identifier>
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<treatment id="0A3E2657304FFFA7FF8EFD0BBFBD0B61" ID-DOI="http://doi.org/10.5281/zenodo.13799529" ID-Zenodo-Dep="13799529" LSID="urn:lsid:plazi:treatment:0A3E2657304FFFA7FF8EFD0BBFBD0B61" httpUri="http://treatment.plazi.org/id/0A3E2657304FFFA7FF8EFD0BBFBD0B61" lastPageId="55" lastPageNumber="256" pageId="54" pageNumber="255">
<subSubSection id="CA8DC4CA304FFFA6FF8EFD0BBE5C0FAA" box="[123,831,665,689]" pageId="54" pageNumber="255" type="nomenclature">
<paragraph id="82289741304FFFA6FF8EFD0BBE5C0FAA" blockId="54.[123,831,665,720]" box="[123,831,665,689]" pageId="54" pageNumber="255">
<heading id="D960202D304FFFA6FF8EFD0BBE5C0FAA" box="[123,831,665,689]" fontSize="10" level="2" pageId="54" pageNumber="255" reason="4">
<emphasis id="B0E34B53304FFFA6FF8EFD0BBE5C0FAA" bold="true" box="[123,831,665,689]" pageId="54" pageNumber="255">
<taxonomicName id="4597ECC2304FFFA6FF8EFD0BBFBF0FAA" ID-CoL="3SQ9Y" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,732,665,689]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B53304FFFA6FF8EFD0BBDDA0FAA" bold="true" box="[123,185,665,689]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
<emphasis id="B0E34B53304FFFA6FF3CFD0BBC540FAA" bold="true" box="[201,311,665,689]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
)
<emphasis id="B0E34B53304FFFA6FEB2FD0BBCA80FAA" bold="true" box="[327,459,665,689]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628304FFFA6FD1CFD0BBE5C0FAA" box="[745,831,665,689]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA304FFFA6FF8EFD28BDBB0FD4" box="[123,216,698,720]" pageId="54" pageNumber="255" type="description">
<paragraph id="82289741304FFFA6FF8EFD28BDBB0FD4" blockId="54.[123,831,665,720]" box="[123,216,698,720]" pageId="54" pageNumber="255">
<figureCitation id="1AAC8BC4304FFFA6FF8EFD28BDBB0FD4" box="[123,216,698,720]" captionStart="FIGURE" captionStartId="55.[155,167,1281,1300]" captionTargetBox="[120,1176,233,1259]" captionTargetId="figure-694@55.[120,1176,216,1278]" captionTargetPageId="55" captionText="FIGURE 33. Digital images of holotype female of Lebia (Metalebia) laterolucida Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799515" httpUri="https://zenodo.org/record/13799515/files/figure.png" pageId="54" pageNumber="255">Figure 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA304FFFA7FF5EFD66BFBD0B61" lastPageId="55" lastPageNumber="256" pageId="54" pageNumber="255" type="materials_examined">
<paragraph id="82289741304FFFA6FF5EFD66B9620E76" blockId="54.[123,1174,756,1725]" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFD66BC3B0E11" bold="true" box="[171,344,756,780]" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFD66BD870E10" baselines="774,774" box="[171,228,756,780]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="54" pageNumber="255">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FF1EFD65BC3B0E11" baselines="773" box="[235,344,759,778]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="54" pageNumber="255">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C304FFFA6FE8AFD66B9620E76" collectingDate="1999-12-21" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="1000" latitude="-21.258333" location="Nacional Ranomafana Bell Vue Trail" longLatPrecision="126" longitude="47.426666" municipality="Parc Nacional Ranomafana Bell Vue" pageId="54" pageNumber="255" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3304FFFA6FE8AFD66BC810E17" box="[383,482,756,780]" pageId="54" pageNumber="255" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4304FFFA6FE04FD66BF1A0E10" box="[497,633,756,780]" captionStart="FIGURE" captionStartId="55.[155,167,1281,1300]" captionTargetBox="[120,1176,233,1259]" captionTargetId="figure-694@55.[120,1176,216,1278]" captionTargetPageId="55" captionText="FIGURE 33. Digital images of holotype female of Lebia (Metalebia) laterolucida Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799515" httpUri="https://zenodo.org/record/13799515/files/figure.png" pageId="54" pageNumber="255">Figs. 33A–B</figureCitation>
), a female, in CAS, labeled: “CASENT 1006164”/
<collectingCountry id="FA80D7D1304FFFA6FF8EFC87BC5A0E36" box="[123,313,789,813]" name="Madagascar" pageId="54" pageNumber="255">MADAGASCAR</collectingCountry>
<collectingRegion id="405359A3304FFFA6FEB6FC86BF6E0E37" box="[323,525,788,812]" country="Madagascar" name="Fianarantsoa" pageId="54" pageNumber="255">Fianarantsoa Prov.</collectingRegion>
<collectingMunicipality id="624C0D3B304FFFA6FDE2FC87BECC0E37" box="[535,943,788,812]" pageId="54" pageNumber="255">
Parc
<location id="8748C19A304FFFA6FDA4FC86BECC0E37" LSID="urn:lsid:plazi:treatment:0A3E2657304FFFA7FF8EFD0BBFBD0B61:8748C19A304FFFA6FDA4FC86BECC0E37" box="[593,943,788,812]" country="Madagascar" latitude="-21.258333" longLatPrecision="126" longitude="47.426666" municipality="Parc Nacional Ranomafana Bell Vue" name="Nacional Ranomafana Bell Vue Trail" pageId="54" pageNumber="255" stateProvince="Fianarantsoa">Nacional Ranomafana Bell Vue</location>
</collectingMunicipality>
Trail, tropical forest
<geoCoordinate id="E7A3F186304FFFA6FF8EFCA7BD840E56" box="[123,231,821,845]" degrees="21" direction="south" minutes="15. 5" orientation="latitude" pageId="54" pageNumber="255" precision="92" value="-21.258333">21°15.5’S</geoCoordinate>
<geoCoordinate id="E7A3F186304FFFA6FF18FCA7BC390E56" box="[237,346,821,845]" degrees="47" direction="east" minutes="25. 6" orientation="longitude" pageId="54" pageNumber="255" precision="92" value="47.426666">47°25.6’E</geoCoordinate>
<quantity id="456F3AA4304FFFA6FE95FCA7BCD20E56" box="[352,433,821,845]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="255" unit="m" value="1000.0">
<elevation id="09BA7072304FFFA6FE95FCA7BCD20E56" box="[352,433,821,845]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="255" unit="m" value="1000.0">1000 m</elevation>
</quantity>
<collectorName id="2F62F297304FFFA6FE42FCA4BF440E56" box="[439,551,821,845]" pageId="54" pageNumber="255">M.E.Irwin</collectorName>
and
<collectorName id="2F62F297304FFFA6FDAFFCA4BF8E0E56" box="[602,749,821,845]" pageId="54" pageNumber="255">E.I. Schlinger</collectorName>
MEI 99-MA-7
<date id="F629B181304FFFA6FC63FCA7B97E0E56" box="[918,1053,821,845]" pageId="54" pageNumber="255" value="1999-12-21">
<collectingDate id="E66D4869304FFFA6FC63FCA7B97E0E56" box="[918,1053,821,845]" pageId="54" pageNumber="255" value="1999-12-21">21-XII-1999</collectingDate>
</date>
”/ “
<typeStatus id="5D2C29E3304FFFA6FBB7FCA7BDDD0E76" pageId="54" pageNumber="255" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2304FFFA6FF30FCC4BF990E75" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[197,762,854,878]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B53304FFFA6FF30FCC4BC610E75" box="[197,258,854,878]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
<emphasis id="B0E34B53304FFFA6FEE7FCC4BC1D0E75" box="[274,382,854,878]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
)
<emphasis id="B0E34B53304FFFA6FE7BFCC4BF6C0E75" box="[398,527,854,878]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FCF7FCC4BE5F0E76" box="[770,828,854,877]" pageId="54" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="82289741304FFFA6FF5EFCE5BDDD090B" blockId="54.[123,1174,756,1725]" pageId="54" pageNumber="255">
<materialsCitation id="32FF9D1C304FFFA6FF5EFCE5BDD4090B" collectingDate="1998-04-13" collectionCode="CAS" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="850" latitude="-21.233334" location="Talatakely area" longLatPrecision="1264" longitude="47.366665" municipality="Ranomafana National Park" pageId="54" pageNumber="255" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3304FFFA6FF5EFCE5BC700E94" box="[171,275,887,911]" pageId="54" pageNumber="255" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F84304FFFA6FDF6FCE4BF540E95" box="[515,567,886,910]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="255" type="Museum">CAS</collectionCode>
) labelled “CASENT 1049030”/ “
<collectingCountry id="FA80D7D1304FFFA6FC6AFCE4B93E0E95" box="[927,1117,886,910]" name="Madagascar" pageId="54" pageNumber="255">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3304FFFA6FB9FFCE4BC250EB4" country="Madagascar" name="Fianarantsoa" pageId="54" pageNumber="255">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B304FFFA6FEA6FC05BF100EB4" box="[339,627,919,943]" pageId="54" pageNumber="255">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A304FFFA6FD75FC05BE420EB5" LSID="urn:lsid:plazi:treatment:0A3E2657304FFFA7FF8EFD0BBFBD0B61:8748C19A304FFFA6FD75FC05BE420EB5" box="[640,801,919,943]" country="Madagascar" latitude="-21.233334" longLatPrecision="1264" longitude="47.366665" municipality="Ranomafana National Park" name="Talatakely area" pageId="54" pageNumber="255" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4304FFFA6FCDBFC05BE120EB5" box="[814,881,919,943]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="54" pageNumber="255" unit="m" value="850.0">
<elevation id="09BA7072304FFFA6FCDBFC05BE120EB5" box="[814,881,919,943]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="54" pageNumber="255" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="F629B181304FFFA6FC8BFC05B9720EB4" box="[894,1041,919,943]" pageId="54" pageNumber="255" value="1998-04-13">
<collectingDate id="E66D4869304FFFA6FC8BFC05B9720EB4" box="[894,1041,919,943]" pageId="54" pageNumber="255" value="1998-04-13">13 April 1998</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186304FFFA6FBCBFC05B9F50EB4" box="[1086,1174,919,943]" degrees="21" direction="south" minutes="14" orientation="latitude" pageId="54" pageNumber="255" precision="925" value="-21.233334">21°14’S</geoCoordinate>
<geoCoordinate id="E7A3F186304FFFA6FF8EFC2ABDB70ED4" box="[123,212,952,976]" degrees="47" direction="east" minutes="22" orientation="longitude" pageId="54" pageNumber="255" precision="925" value="47.366665">47°22’E</geoCoordinate>
<collectingMethod id="5BD6EF56304FFFA6FF28FC25BC000ED4" box="[221,355,951,975]" pageId="54" pageNumber="255">Malaise trap</collectingMethod>
in mixed tropical forest,
<collectorName id="2F62F297304FFFA6FD8EFC2ABF880ED4" box="[635,747,951,975]" pageId="54" pageNumber="255">M.E.Irwin</collectorName>
and
<collectorName id="2F62F297304FFFA6FCD6FC2ABEDB0ED4" box="[803,952,951,975]" pageId="54" pageNumber="255">E.I. Schlinger</collectorName>
collectors, Stop 98- MAD-1”/ “
<typeStatus id="5D2C29E3304FFFA6FF00FC4ABC1A0EEB" box="[245,377,984,1008]" pageId="54" pageNumber="255" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2304FFFA6FE75FC4ABED70EEB" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[384,948,984,1008]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B53304FFFA6FE75FC4ABCDE0EEB" box="[384,445,984,1008]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
<emphasis id="B0E34B53304FFFA6FE38FC4ABF5A0EEB" box="[461,569,984,1008]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
)
<emphasis id="B0E34B53304FFFA6FDBCFC4ABFA90EEB" box="[585,714,984,1008]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FC4EFC4BBE960EEB" box="[955,1013,985,1008]" pageId="54" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741304FFFA6FF5EFB8BB950092A" blockId="54.[123,1174,756,1725]" box="[171,1075,1049,1073]" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFB8BBC34092B" bold="true" box="[171,343,1049,1073]" pageId="54" pageNumber="255">
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<smallCapsWord id="84CE019D304FFFA6FF19FB8FBC34092B" baselines="1067" box="[236,343,1053,1072]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="54" pageNumber="255">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1304FFFA6FE8BFB8BBC9C092A" box="[382,511,1049,1073]" name="Madagascar" pageId="54" pageNumber="255">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3304FFFA6FDF8FB8BBF9F092A" box="[525,764,1049,1073]" country="Madagascar" name="Fianarantsoa" pageId="54" pageNumber="255">Fianarantsoa Province</collectingRegion>
, Ranomafana National Park.
</paragraph>
<paragraph id="82289741304FFFA6FF5EFBA8BF5109A8" blockId="54.[123,1174,756,1725]" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFBA8BC9A094B" bold="true" box="[171,505,1082,1106]" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFBA8BC4D094B" baselines="1100,1099" box="[171,302,1082,1106]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="54" pageNumber="255">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FECDFBAFBC37094B" baselines="1099" box="[312,340,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="54" pageNumber="255">OF</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FEABFBAFBCD2094B" baselines="1099" box="[350,433,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="54" pageNumber="255">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FE49FBAFBC9A094B" baselines="1099" box="[444,505,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="54" pageNumber="255">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2304FFFA6FCF2FBA8BEEB0949" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[775,904,1082,1106]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FCF2FBA8BEEB0949" box="[775,904,1082,1106]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B53304FFFA6FE9FFBC9BCCD0968" box="[362,430,1115,1139]" italics="true" pageId="54" pageNumber="255">lateris</emphasis>
, meaning side, and
<emphasis id="B0E34B53304FFFA6FD71FBC9BFB20968" box="[644,721,1115,1139]" italics="true" pageId="54" pageNumber="255">lucidus</emphasis>
, meaning bright or full of light. The name refers to the pale, translucent lateral pronotal margins, which contrast markedly with the dark pronotal disc in members of this species.
</paragraph>
<paragraph id="82289741304FFFA7FF5EFB2EBC090B0E" blockId="54.[123,1174,756,1725]" lastBlockId="55.[123,1174,1366,1659]" lastPageId="55" lastPageNumber="256" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFB2EBC2109C8" baselines="1230,1230" box="[171,322,1212,1236]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFB2EBC2109C8" bold="true" box="[171,322,1212,1236]" pageId="54" pageNumber="255">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderate for genus, SBL of females = 4.5–5.0 mm. Members of this species (
<figureCitation id="1AAC8BC4304FFFA6FF2CFB4FBC5B09EE" box="[217,312,1245,1269]" captionStart="FIGURE" captionStartId="55.[155,167,1281,1300]" captionTargetBox="[120,1176,233,1259]" captionTargetId="figure-694@55.[120,1176,216,1278]" captionTargetPageId="55" captionText="FIGURE 33. Digital images of holotype female of Lebia (Metalebia) laterolucida Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799515" httpUri="https://zenodo.org/record/13799515/files/figure.png" pageId="54" pageNumber="255">Fig. 33A</figureCitation>
) key to subgenus
<taxonomicName id="4597ECC2304FFFA6FDF5FB4FBE7009EE" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[512,787,1245,1269]" pageId="54" pageNumber="255" rank="subGenus" subGenus="Metalebia">
<emphasis id="B0E34B53304FFFA6FDF5FB4FBF0F09EE" box="[512,620,1245,1269]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
Jeannel (1949)
</taxonomicName>
and the
<taxonomicName id="4597ECC2304FFFA6FC87FB4FB95009EE" authorityName="Basilewsky" authorityYear="1982" box="[882,1075,1245,1269]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304FFFA6FC87FB4FB95009EE" box="[882,1075,1245,1269]" italics="true" pageId="54" pageNumber="255">madagascariensis</emphasis>
</taxonomicName>
group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody (head plus pronotum) has no metallic reflection and the elytra are uniform in color; and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the
<taxonomicName id="4597ECC2304FFFA6FBD9FACDBDB80883" authorityName="Basilewsky" authorityYear="1982" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304FFFA6FBD9FACDBDB80883" italics="true" pageId="54" pageNumber="255">madagascariensis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4597ECC2304FFFA6FEC7FA13BCB10883" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[306,466,1408,1432]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FEC7FA13BCB10883" box="[306,466,1408,1432]" italics="true" pageId="54" pageNumber="255">L. laterolucida</emphasis>
</taxonomicName>
members are distinguished from those of
<taxonomicName id="4597ECC2304FFFA6FC51FA13BDA108A2" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B53304FFFA6FC51FA13B95F0883" box="[932,1084,1408,1432]" italics="true" pageId="54" pageNumber="255">L. alluaudana</emphasis>
Jeannel (1949)
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FF20FA33BCEE08A2" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[213,397,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov.">
<emphasis id="B0E34B53304FFFA6FF20FA33BCEE08A2" box="[213,397,1441,1465]" italics="true" pageId="54" pageNumber="255">L. apicoviolacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FE6CFA33BC9208A3" box="[409,497,1441,1464]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4597ECC2304FFFA6FDF1FA33BE5708A2" authority="Alluaud (1936 a)" authorityName="Alluaud" authorityYear="1936" box="[516,820,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B53304FFFA6FDF1FA33BF1008A3" box="[516,627,1441,1464]" italics="true" pageId="54" pageNumber="255">L. mirana</emphasis>
<bibRefCitation id="E606EAB0304FFFA6FD8BFA33BE5708A2" author="ALLUAUD, C." box="[638,820,1441,1465]" pageId="54" pageNumber="255" pagination="1 - 13" refId="ref28980" refString="ALLUAUD, C. 1936 a. Carabidae recueillis a Madagascar par MM. Syrig, G. Olsoufieff, Vadon, R. Catala, etc. Afra, Cahiers d'Entomologie 11: 1 - 13." type="journal article" year="1936">Alluaud (1936a)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FCB2FA33BE9408A2" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[839,1015,1441,1465]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53304FFFA6FCB2FA33BE9408A2" box="[839,1015,1441,1465]" italics="true" pageId="54" pageNumber="255">L. ranomafanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FBF1FA33B93F08A3" box="[1028,1116,1441,1464]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4597ECC2304FFFA6FF8EFA50BCE708C2" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[123,388,1473,1497]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B53304FFFA6FF8EFA50BDBF08C2" box="[123,220,1473,1497]" italics="true" pageId="54" pageNumber="255">L. tanala</emphasis>
Jeannel (1949)
</taxonomicName>
by the absence of metallic reflection on the elytra and punctation of the anterior dorsum of the head (both features present in members of the other five species). They differ from members of
<taxonomicName id="4597ECC2304FFFA6FE9FF991BFB60B00" authority="(Fairmaire) (1889)" authorityName="" authorityYear="1889" baseAuthorityName="Fairmaire" baseAuthorityYear="1889" box="[362,725,1538,1563]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="B0E34B53304FFFA6FE9FF991BF620B01" box="[362,513,1538,1562]" italics="true" pageId="54" pageNumber="255">L. sulcipennis</emphasis>
(Fairmaire) (1889)
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FD10F991BE800B00" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[741,995,1538,1563]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="B0E34B53304FFFA6FD10F991BE5A0B01" box="[741,825,1539,1562]" italics="true" pageId="54" pageNumber="255">L. nana</emphasis>
Jeannel (1949)
</taxonomicName>
, and
<taxonomicName id="4597ECC2304FFFA6FBD6F991BC7A0B20" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="B0E34B53304FFFA6FBD6F991B9F60B01" box="[1059,1173,1539,1562]" italics="true" pageId="54" pageNumber="255">L. perrieri</emphasis>
Jeannel (1949)
</taxonomicName>
in having the pronotum broad, ratio PW/PL = at least 1.5, and with lateral margins distinctly rounded at least in the anterior half pronotum (narrower and less rounded anteriorly in members of the three other species). From adults of
<taxonomicName id="4597ECC2304FFFA6FD43F9F7BECB0B67" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[694,936,1636,1660]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B0E34B53304FFFA6FD43F9F7BE630B67" box="[694,768,1636,1660]" italics="true" pageId="54" pageNumber="255">L. rufa</emphasis>
Jeannel (1949)
</taxonomicName>
, they differ in having the pronotal basal lobe distinctly margined (absent from the former).
<taxonomicName id="4597ECC2304FFFA6FC94F917B9450B86" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[865,1062,1669,1693]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FC94F917B9450B86" box="[865,1062,1669,1693]" italics="true" pageId="54" pageNumber="255">Lebia laterolucida</emphasis>
</taxonomicName>
adults are most similar to those of the two remaining species in Jeannel’s key,
<taxonomicName id="4597ECC2304FFFA7FC77F934BDA10875" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" lastPageId="55" lastPageNumber="256" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="brunneipennis">
<emphasis id="B0E34B53304FFFA6FC77F934B9580BA6" box="[898,1083,1701,1725]" italics="true" pageId="54" pageNumber="255">L. brunneipennis</emphasis>
Jeannel (1949)
</taxonomicName>
and
<taxonomicName id="4597ECC2304EFFA7FF01FAC4BFE40875" authority="Chaudoir (1850)" authorityName="Chaudoir" authorityYear="1850" box="[244,647,1366,1390]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304EFFA7FF01FAC4BCB20875" box="[244,465,1366,1390]" italics="true" pageId="55" pageNumber="256">L. madagascariensis</emphasis>
<bibRefCitation id="E606EAB0304EFFA7FE22FAC4BFE40875" author="CHAUDOIR, M." box="[471,647,1366,1390]" pageId="55" pageNumber="256" pagination="3 - 85" refId="ref29725" refString="CHAUDOIR, M. 1850. Memoire sur la famille des Carabiques. 2 e partie. Bulletin de la Societe Imperiale des Naturalistes de Moscou 23: 3 - 85." type="journal article" year="1850">Chaudoir (1850)</bibRefCitation>
</taxonomicName>
, but differ from them in having the dorsum of the head, pronotum and elytra dark piceous, with markedly contrasting pale yellow lateral pronotal margins (dorsum uniformly reddish brown and the lateral pronotal margins not or less markedly contrasting in color in the together two species). In addition, they are distinctly smaller than
<taxonomicName id="4597ECC2304EFFA7FB75FA29BC5F08EF" authorityName="Chaudoir" authorityYear="1850" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304EFFA7FB75FA29BC5F08EF" italics="true" pageId="55" pageNumber="256">L. madagascariensis</emphasis>
</taxonomicName>
adults and have distinct elytral microsculpture (absent or at least very faint in
<taxonomicName id="4597ECC2304EFFA7FB75FA4FBC770B0D" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="brunneipennis">
<emphasis id="B0E34B53304EFFA7FB75FA4FBC770B0D" italics="true" pageId="55" pageNumber="256">L. brunneipennis</emphasis>
</taxonomicName>
adults).
</paragraph>
<caption id="D6E8C7C9304EFFA7FF6EFA93BF4A0835" ID-DOI="http://doi.org/10.5281/zenodo.13799515" ID-Zenodo-Dep="13799515" httpUri="https://zenodo.org/record/13799515/files/figure.png" pageId="55" pageNumber="256" startId="55.[155,167,1281,1300]" targetBox="[120,1176,233,1259]" targetPageId="55" targetType="figure">
<paragraph id="82289741304EFFA7FF6EFA93BF4A0835" blockId="55.[123,1174,1281,1326]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF6EFA93BDBB0809" baselines="1295,1294" box="[155,216,1281,1300]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="55" pageNumber="256">FIGURE</smallCapsWord>
33. Digital images of holotype female of
<taxonomicName id="4597ECC2304EFFA7FDC8FA93B963080F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[573,1024,1281,1300]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="laterolucida" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B53304EFFA7FDC8FA93BF0D080F" box="[573,622,1281,1300]" italics="true" pageId="55" pageNumber="256">Lebia</emphasis>
(
<emphasis id="B0E34B53304EFFA7FD8FFA93BFB3080F" box="[634,720,1281,1300]" italics="true" pageId="55" pageNumber="256">Metalebia</emphasis>
)
<emphasis id="B0E34B53304EFFA7FD28FA93BE27080F" box="[733,836,1281,1300]" italics="true" pageId="55" pageNumber="256">laterolucida</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304EFFA7FBF0FA93B925080F" box="[1029,1094,1281,1300]" pageId="55" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741304EFFA7FF5EF98DB9690B2C" blockId="55.[123,1174,1366,1659]" box="[171,1034,1567,1592]" pageId="55" pageNumber="256">
<emphasis id="B0E34B53304EFFA7FF5EF98DBC920B2D" bold="true" box="[171,497,1567,1591]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF5EF98DBC350B2D" baselines="1585,1585" box="[171,342,1567,1591]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="55" pageNumber="256">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D304EFFA7FEA8F9B1BC920B2D" baselines="1585" box="[349,497,1571,1590]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="55" pageNumber="256">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3304EFFA7FC8AF9B2BECE0B23" box="[895,941,1568,1592]" pageId="55" pageNumber="256">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741304EFFA7FF5EF9D3BFBD0B61" blockId="55.[123,1174,1366,1659]" pageId="55" pageNumber="256">
<emphasis id="B0E34B53304EFFA7FF5EF9D3BCC70B4C" bold="true" box="[171,420,1601,1625]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF5EF9D3BC6A0B4C" baselines="1619,1618" box="[171,265,1601,1625]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="55" pageNumber="256">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D304EFFA7FEE5F9D6BCC70B4C" baselines="1618" box="[272,420,1604,1623]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="55" pageNumber="256">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C304EFFA7FE39F9D3BFB40B61" location="Talatakely area" pageId="55" pageNumber="256" specimenCount="1">
Both specimens of the
<typeStatus id="5D2C29E3304EFFA7FD37F9D3BF930B42" box="[706,752,1601,1625]" pageId="55" pageNumber="256">type</typeStatus>
series were collected in mixed secondary tropical forest in the
<location id="8748C19A304EFFA7FE71F9F0BF450B61" LSID="urn:lsid:plazi:treatment:0A3E2657304FFFA7FF8EFD0BBFBD0B61:8748C19A304EFFA7FE71F9F0BF450B61" box="[388,550,1634,1658]" name="Talatakely area" pageId="55" pageNumber="256">Talatakely area</location>
in malaise traps
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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