249 lines
38 KiB
XML
249 lines
38 KiB
XML
<document id="EC30294AF2E31E6595CE684932B52D96" ID-DOI="10.11646/zootaxa.3846.4.2" ID-GBIF-Dataset="ea136b1c-dc96-4c41-b078-01ca553453a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="252519" ID-ZooBank="71A80CC3-CDC1-4291-863B-FA7501D60AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467899762" checkinUser="plazi" docAuthor="Câmara, J. T. & Rafael, J. A." docDate="2014" docId="03F6943FE062FF84C7C7FA07AAE0134F" docLanguage="en" docName="zt03846p546.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opeatocerata megalophallus Câmara & Rafael, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="527" masterDocId="FFCFEC47E075FF9DC750FFEAA8211077" masterDocTitle="Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)" masterLastPageNumber="546" masterPageNumber="502" pageNumber="525" updateTime="1698305230595" updateUser="plazi">
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<mods:mods id="D87FB37F15E15F1FE62C16B714BC9AB1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="50092CEFF464609E166F1CE757C6D9CD">
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<mods:title id="880ABEF615DBBBA9026EEBFD3481B380">Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3EE6383DF440372BC0C3EF24BEB5EE0E" type="personal">
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<mods:role id="001E36CD1191AFD129D3C7862AEC03F8">
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<mods:roleTerm id="751D952E7C4770E3A56F6217E6ECF804">Author</mods:roleTerm>
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<mods:namePart id="EDAE354ADA2B3279FD8A9D4168F85B1C">Câmara, J. T.</mods:namePart>
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</mods:name>
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<mods:name id="0C380C38D43F937789B1408251341E6F" type="personal">
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<mods:roleTerm id="A1BE92A9C26326ABBFE2C6A788C1CB05">Author</mods:roleTerm>
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<mods:namePart id="8943B70357651B67BFCB45D0011DDA36">Rafael, J. A.</mods:namePart>
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<mods:typeOfResource id="01D0924DA5CC84756D1835AAB15FC36D">text</mods:typeOfResource>
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<mods:titleInfo id="F29AA35A4FE32F00A8CD39714FE2C269">
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<mods:title id="5794D4F5018DA45C3146D01F628D18AB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FD98D19741ADED9323A1CA0A41C4E217">
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<mods:date id="4A4501AFF8F80A92E28932210E9E13A2">2014</mods:date>
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<mods:detail id="53ADC5D03224315D4234719218F36863" type="volume">
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<mods:number id="023796629ABA6A247FFE1604327593F0">3846</mods:number>
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<mods:detail id="29126C8C026E8325D90E1FA850DFC696" type="issue">
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<mods:number id="C9689AB0D5E748139FFBD09E20BF4741">4</mods:number>
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<mods:start id="A6A89B0FE0BF25D5FE93D1EDF3E0408B">502</mods:start>
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<mods:classification id="90010A0D95B2D5A5C51011B2B5DF6DD3">journal article</mods:classification>
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<mods:identifier id="DAFAE9157AC7E23BE77A76318CDB1A7E" type="DOI">10.11646/zootaxa.3846.4.2</mods:identifier>
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<mods:identifier id="3260AE5257D7B32CE804110E92FF549D" type="GBIF-Dataset">ea136b1c-dc96-4c41-b078-01ca553453a0</mods:identifier>
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<mods:identifier id="4644470E7819E03C1C24D95C7F055CD2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4BA4D66043D8A19168ECBA1B2C0F0602" type="Zenodo-Dep">252519</mods:identifier>
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<treatment id="03F6943FE062FF84C7C7FA07AAE0134F" ID-DOI="http://doi.org/10.5281/zenodo.5285253" ID-GBIF-Taxon="119514714" ID-Zenodo-Dep="5285253" LSID="urn:lsid:plazi:treatment:03F6943FE062FF84C7C7FA07AAE0134F" httpUri="http://treatment.plazi.org/id/03F6943FE062FF84C7C7FA07AAE0134F" lastPageId="25" lastPageNumber="527" pageId="23" pageNumber="525">
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<subSubSection id="C34576A2E062FF8AC7C7FA07A963165E" pageId="23" pageNumber="525" type="nomenclature">
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<paragraph id="8BE02529E062FF8AC7C7FA07AA74167F" blockId="23.[151,597,1517,1577]" box="[151,597,1517,1544]" pageId="23" pageNumber="525">
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<heading id="D0A89245E062FF8AC7C7FA07AA74167F" bold="true" box="[151,597,1517,1544]" fontSize="11" level="1" pageId="23" pageNumber="525" reason="1">
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<emphasis id="B92BF93BE062FF8AC7C7FA07AA74167F" bold="true" box="[151,597,1517,1544]" pageId="23" pageNumber="525">
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<taxonomicName id="4C5F5EAAE062FF8AC7C7FA07A9D11670" ID-CoL="8D3LL" box="[151,496,1517,1543]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="525" phylum="Arthropoda" rank="species" species="megalophallus" status="sp. nov.">
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<emphasis id="B92BF93BE062FF8AC7C7FA07A9D11670" bold="true" box="[151,496,1517,1543]" italics="true" pageId="23" pageNumber="525">Opeatocerata megalophallus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2184440E062FF8AC6A8FA04AA74167F" box="[504,597,1518,1544]" pageId="23" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE02529E062FF8AC7C7F9FAA963165E" blockId="23.[151,597,1517,1577]" box="[151,322,1552,1577]" pageId="23" pageNumber="525">
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(
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<figureCitation id="136439ACE062FF8AC7CFF9FAA91B165E" box="[159,314,1552,1577]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Figs 114–122</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34576A2E062FF8AC7C7F9B2A92F16CE" pageId="23" pageNumber="525" type="diagnosis">
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<paragraph id="8BE02529E062FF8AC7C7F9B2A92F16CE" blockId="23.[151,1437,1624,2009]" pageId="23" pageNumber="525">
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<emphasis id="B92BF93BE062FF8AC7C7F9B2A92B1606" bold="true" box="[151,266,1624,1649]" pageId="23" pageNumber="525">Diagnosis</emphasis>
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. Epandrium lobe acuminate at apex (
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<figureCitation id="136439ACE062FF8AC5E2F9B2AB701606" box="[690,849,1624,1649]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Figs 116, 117</figureCitation>
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); phallus narrow, much longer than the hypandrium (
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<figureCitation id="136439ACE062FF8AC7CFF996A92316E2" box="[159,258,1660,1685]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 121</figureCitation>
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); subepandrial sclerite as wide as long (
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<figureCitation id="136439ACE062FF8AC59CF996AB1116E2" box="[716,816,1660,1685]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 120</figureCitation>
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); bacilliform sclerite shorter than subepandrial sclerite (
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<figureCitation id="136439ACE062FF8AC7CFF94AA92316CE" box="[159,258,1696,1721]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 120</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34576A2E062FF84C797F92EAC051183" lastPageId="25" lastPageNumber="527" pageId="23" pageNumber="525" type="description">
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<paragraph id="8BE02529E062FF84C797F92EA98D11A7" blockId="23.[151,1437,1624,2009]" lastBlockId="25.[151,1436,151,824]" lastPageId="25" lastPageNumber="527" pageId="23" pageNumber="525">
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<emphasis id="B92BF93BE062FF8AC797F92EA98116AA" bold="true" box="[199,416,1732,1757]" pageId="23" pageNumber="525">Description. Male</emphasis>
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(
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<figureCitation id="136439ACE062FF8AC6E3F92EAA3916AA" box="[435,536,1732,1757]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 114</figureCitation>
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).
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<emphasis id="B92BF93BE062FF8AC57FF92EAA4E16AA" bold="true" box="[559,623,1732,1757]" pageId="23" pageNumber="525">Head</emphasis>
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: Face slightly divergent toward the proboscis, dark brown, with gray pruinescence visible in ventral view, about
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<date id="FFE103E9E062FF8AC5D3F903AA871776" box="[643,678,1769,1793]" pageId="23" pageNumber="525">4X</date>
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longer than lower width. Ocellar tubercle protrubeant, dark brown with gray pruinescence; ocelli brown. Antenna brown to black bristles; postpedicel about 1.5X longer than pedicel; stylus about 1.5X longer than postpedicel. Proboscis yellow shorter than head height.
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<emphasis id="B92BF93BE062FF8AC337F8DAACE1173E" bold="true" box="[1127,1216,1840,1865]" pageId="23" pageNumber="525">Thorax</emphasis>
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yellow, shiny (
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<figureCitation id="136439ACE062FF8AC23EF8DAA8E5171A" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 114</figureCitation>
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). Legs yellow, except for hind femur and tibia with apical black ring and all tarsomeres 3–5 brown; hind tibia with anterodorsal and posterodorsal rows of longer bristles.
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<emphasis id="B92BF93BE062FF8AC42AF892AB9A17E6" bold="true" box="[890,955,1912,1937]" pageId="23" pageNumber="525">Wing</emphasis>
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(
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<figureCitation id="136439ACE062FF8AC483F892AC1817E6" box="[979,1081,1912,1937]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 115</figureCitation>
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) hyaline, brown pterostigma, conspicuous, about 2.3X longer than wide. Halter yellow.
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<emphasis id="B92BF93BE062FF8AC47AF876ABBE17C2" bold="true" box="[810,927,1948,1973]" pageId="23" pageNumber="525">Abdomen</emphasis>
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yellow, shiny (
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<figureCitation id="136439ACE062FF8AC31EF876AC9117C2" box="[1102,1200,1948,1973]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="23" pageNumber="525">Fig. 114</figureCitation>
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); tergite 8 divided in two subtriangular plates; sternite 8 divided in two trapezoidal plates.
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<emphasis id="B92BF93BE062FF8AC4CCF82AAC3E17AE" bold="true" box="[924,1055,1984,2009]" pageId="23" pageNumber="525">Terminalia</emphasis>
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: Anterior cercus with proclinate dorsal projection in lateral view (
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<figureCitation id="136439ACE06CFF84C55EFF7DAA8D10C7" box="[526,684,151,176]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Figs 116, 117</figureCitation>
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), with posterior margin slightly concave in dorsal view (
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<figureCitation id="136439ACE06CFF84C27DFF7DADAE10C7" box="[1325,1423,151,176]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 118</figureCitation>
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); descendant plate subrectangular in posterior view (
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<figureCitation id="136439ACE06CFF84C586FF56AB1910A3" box="[726,824,188,213]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 119</figureCitation>
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). Posterior cercus with acuminate apex in lateral view (
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<figureCitation id="136439ACE06CFF84C7CFFF35A91F108F" box="[159,318,223,248]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Figs 116, 117</figureCitation>
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); with dorsal projections, a subbasal pointed and a submedian subtriangular (
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<figureCitation id="136439ACE06CFF84C3FDFF35AD31108F" box="[1197,1296,223,248]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 118</figureCitation>
|
||
). Hypoproct bacilliform lateral view and covering inner surface of the posterior cercus in posterior view (
|
||
<figureCitation id="136439ACE06CFF84C3E8FEEEAD3A116B" box="[1208,1307,260,285]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 117</figureCitation>
|
||
). Epandrial ventral lobe with acuminate apex (
|
||
<figureCitation id="136439ACE06CFF84C549FECDAA951137" box="[537,692,295,320]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Figs 116, 117</figureCitation>
|
||
), with spiniform bristles mesially and longer bristles apically (
|
||
<figureCitation id="136439ACE06CFF84C23FFECDA8E51113" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 117</figureCitation>
|
||
). Hypandrium longer than wide, with concave apex (
|
||
<figureCitation id="136439ACE06CFF84C47CFEA6ABB11113" box="[812,912,332,357]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 122</figureCitation>
|
||
), without long bristles. Phallus narrow, much longer than the hypandrium (
|
||
<figureCitation id="136439ACE06CFF84C6B8FE85AA6D11FF" box="[488,588,367,392]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 121</figureCitation>
|
||
). Subepandrial sclerite as wide as long, U-shaped (
|
||
<figureCitation id="136439ACE06CFF84C3F0FE85AD2411FF" box="[1184,1285,367,392]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 120</figureCitation>
|
||
). Bacilliform sclerite without folds and shorter than subepandrial sclerite (
|
||
<figureCitation id="136439ACE06CFF84C467FE7EABB911DB" box="[823,920,404,429]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 120</figureCitation>
|
||
).
|
||
<emphasis id="B92BF93BE06CFF84C4FBFE7EAC4F11DA" bold="true" box="[939,1134,404,429]" pageId="25" pageNumber="527">Specimen length</emphasis>
|
||
:
|
||
<quantity id="4CA788CCE06CFF84C32DFE7EACF411DB" box="[1149,1237,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="25" pageNumber="527" unit="mm" value="2.9">2.9 mm</quantity>
|
||
;
|
||
<emphasis id="B92BF93BE06CFF84C3B0FE7EAD4A11DA" bold="true" box="[1248,1387,404,429]" pageId="25" pageNumber="527">wing length</emphasis>
|
||
:
|
||
<quantity id="4CA788CCE06CFF84C229FE7EA8E711B8" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="25" pageNumber="527" unit="mm" value="2.9">2.9 mm</quantity>
|
||
.
|
||
<emphasis id="B92BF93BE06CFF84C780FE5DA90611A7" bold="true" box="[208,295,439,464]" pageId="25" pageNumber="527">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<caption id="DF2075A1E06DFF85C7C7F8E6ABE01708" httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="24" pageNumber="526" targetBox="[169,1423,200,1776]" targetPageId="24">
|
||
<paragraph id="8BE02529E06DFF85C7C7F8E6ABE01708" blockId="24.[151,1436,1804,1919]" pageId="24" pageNumber="526">
|
||
<emphasis id="B92BF93BE06DFF85C7C7F8E6A9471755" bold="true" box="[151,358,1804,1827]" pageId="24" pageNumber="526">FIGURES 114–122</emphasis>
|
||
.
|
||
<taxonomicName id="4C5F5EAAE06DFF85C624F8E4AABD1754" box="[372,668,1805,1827]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="526" phylum="Arthropoda" rank="species" species="megalophallus" status="sp. nov.">
|
||
<emphasis id="B92BF93BE06DFF85C624F8E4AABD1754" box="[372,668,1805,1827]" italics="true" pageId="24" pageNumber="526">Opeatocerata megalophallus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2184440E06DFF85C5F4F8E7AAD71755" box="[676,758,1805,1826]" pageId="24" pageNumber="526" rank="species">
|
||
<emphasis id="B92BF93BE06DFF85C5F4F8E7AAD71755" bold="true" box="[676,758,1805,1826]" pageId="24" pageNumber="526">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype ♂.
|
||
<emphasis id="B92BF93BE06DFF85C4D6F8E7AB881755" bold="true" box="[902,937,1805,1826]" pageId="24" pageNumber="526">114</emphasis>
|
||
.
|
||
<emphasis id="B92BF93BE06DFF85C4E9F8E7AC251754" box="[953,1028,1805,1827]" italics="true" pageId="24" pageNumber="526">habitus</emphasis>
|
||
, lateral view;
|
||
<emphasis id="B92BF93BE06DFF85C3C8F8E7AC9A1755" bold="true" box="[1176,1211,1805,1826]" pageId="24" pageNumber="526">115</emphasis>
|
||
. wing;
|
||
<emphasis id="B92BF93BE06DFF85C25CF8E7AD0E1755" bold="true" box="[1292,1327,1805,1826]" pageId="24" pageNumber="526">116</emphasis>
|
||
. abdomen from tergite 6 to terminalia, lateral view;
|
||
<emphasis id="B92BF93BE06DFF85C568F8C6AA7A1736" bold="true" box="[568,603,1836,1857]" pageId="24" pageNumber="526">117</emphasis>
|
||
. terminalia, lateral view;
|
||
<emphasis id="B92BF93BE06DFF85C430F8C6ABA21736" bold="true" box="[864,899,1836,1857]" pageId="24" pageNumber="526">118</emphasis>
|
||
. lobes of cerci, dorsal view;
|
||
<emphasis id="B92BF93BE06DFF85C3F8F8C6ACED1736" bold="true" box="[1192,1228,1836,1857]" pageId="24" pageNumber="526">119</emphasis>
|
||
. descendant plate of anterior cercus and hypoproct, posterior view;
|
||
<emphasis id="B92BF93BE06DFF85C522F8A0AAB91728" bold="true" box="[626,664,1866,1887]" pageId="24" pageNumber="526">120</emphasis>
|
||
. subepandrial sclerite and bacilliform sclerite, dorsal view;
|
||
<emphasis id="B92BF93BE06DFF85C3A8F8A0AD3F1728" bold="true" box="[1272,1310,1866,1887]" pageId="24" pageNumber="526">121</emphasis>
|
||
. ejaculatory apodeme, phallus and hypandrium, lateral view;
|
||
<emphasis id="B92BF93BE06DFF85C5D4F883AA8B1709" bold="true" box="[644,682,1897,1918]" pageId="24" pageNumber="526">122</emphasis>
|
||
. hypandrium, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE02529E06CFF84C797FE36AC051183" blockId="25.[151,1436,151,824]" box="[199,1060,476,501]" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C797FE36A9DC1182" bold="true" box="[199,509,476,501]" pageId="25" pageNumber="527">Geographical distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F34865B9E06CFF84C55BFE36AAA41182" box="[523,645,476,501]" name="Costa Rica" pageId="25" pageNumber="527">Costa Rica</collectingCountry>
|
||
(San Mateo, Guanacaste) (
|
||
<figureCitation id="136439ACE06CFF84C4E6FE36AC361183" box="[950,1047,476,501]" captionStart="FIGURE 211" captionStartId="42.[151,250,1955,1977]" captionTargetBox="[235,1368,193,1934]" captionTargetId="figure@42.[216,1371,193,1934]" captionTargetPageId="42" captionText="FIGURE 211. Geographic distribution of species of Opeatocerata." httpUri="https://zenodo.org/record/252546/files/figure.png" pageId="25" pageNumber="527">Fig. 211</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34576A2E06CFF84C797FE15ADA412F2" pageId="25" pageNumber="527" type="materials_examined">
|
||
<paragraph id="8BE02529E06CFF84C797FE15AC421217" blockId="25.[151,1436,151,824]" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C797FE15A94F126F" bold="true" box="[199,366,511,536]" pageId="25" pageNumber="527">
|
||
<typeStatus id="54E49B8BE06CFF84C797FE15A923126F" box="[199,258,511,536]" pageId="25" pageNumber="527">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54E49B8BE06CFF84C62DFE15AA3D126F" box="[381,540,511,536]" pageId="25" pageNumber="527" type="holotype">
|
||
<emphasis id="B92BF93BE06CFF84C62DFE15AA3D126F" bold="true" box="[381,540,511,536]" pageId="25" pageNumber="527">HOLOTYPE</emphasis>
|
||
</typeStatus>
|
||
♂, labelled: Higuito,
|
||
<emphasis id="B92BF93BE06CFF84C444FE15AC65126F" bold="true" box="[788,1092,511,536]" pageId="25" pageNumber="527">San Mateo C[osta] R[ica]</emphasis>
|
||
/ Pablo Schild Coll. (
|
||
<collectionCode id="ED4EBDECE06CFF84C269FDEAADB1126F" LSID="urn:lsid:biocol.org:col:34871" box="[1337,1424,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="527">USNM</collectionCode>
|
||
).
|
||
<typeStatus id="54E49B8BE06CFF84C7C7FDCEA925124A" box="[151,260,548,573]" pageId="25" pageNumber="527" type="paratype">
|
||
<emphasis id="B92BF93BE06CFF84C7C7FDCEA925124A" bold="true" box="[151,260,548,573]" pageId="25" pageNumber="527">Paratype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectingCountry id="F34865B9E06CFF84C64CFDCEA988124A" box="[284,425,548,573]" name="Costa Rica" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C64CFDCEA988124A" bold="true" box="[284,425,548,573]" pageId="25" pageNumber="527">Costa Rica</emphasis>
|
||
</collectingCountry>
|
||
,
|
||
<emphasis id="B92BF93BE06CFF84C691FDCEAA6C124A" bold="true" box="[449,589,548,573]" pageId="25" pageNumber="527">Guanacaste</emphasis>
|
||
. Bagaces. P.N. Palo Verde. Ext. E. Campo de Aterrizaje,
|
||
<quantity id="4CA788CCE06CFF84C266FDCEADB6124B" box="[1334,1431,548,573]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="25" pageNumber="527" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50 m</quantity>
|
||
,
|
||
<date id="FFE103E9E06CFF84C7C7FDADA94E1217" box="[151,367,583,608]" pageId="25" pageNumber="527" value="1999-07-16" valueMax="1999-08-17" valueMin="1999-07-16">16.vii–17.viii.1999</date>
|
||
, L. Jiménez, Malaise. L_N_
|
||
<date id="FFE103E9E06CFF84C5E3FDADAB261217" box="[691,775,583,608]" pageId="25" pageNumber="527" value="52-09-26">260952</date>
|
||
_385020 #53259 (
|
||
<specimenCount id="9D59EEA0E06CFF84C486FDA2AC211228" box="[982,1024,584,608]" pageId="25" pageNumber="527" type="generic">1 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED4EBDECE06CFF84C35CFDADAC761217" box="[1036,1111,583,608]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="25" pageNumber="527">INBio</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE02529E06CFF84C797FD86ADA412F2" blockId="25.[151,1436,151,824]" box="[199,1413,620,645]" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C797FD86A98A12F2" bold="true" box="[199,427,620,645]" pageId="25" pageNumber="527">
|
||
<typeStatus id="54E49B8BE06CFF84C797FD86A91512F2" box="[199,308,620,645]" pageId="25" pageNumber="527" type="holotype">Holotype</typeStatus>
|
||
condition
|
||
</emphasis>
|
||
. Left legs lost; right wing mounted on microslide; abdomen in microtube with glycerin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34576A2E06CFF84C797FD65AD5912DF" box="[199,1400,655,680]" pageId="25" pageNumber="527" type="etymology">
|
||
<paragraph id="8BE02529E06CFF84C797FD65AD5912DF" blockId="25.[151,1436,151,824]" box="[199,1400,655,680]" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C797FD65A96712DF" bold="true" box="[199,326,655,680]" pageId="25" pageNumber="527">Etymology</emphasis>
|
||
. From the Greek
|
||
<emphasis id="B92BF93BE06CFF84C55CFD7BAA6812DF" box="[524,585,657,680]" italics="true" pageId="25" pageNumber="527">mega</emphasis>
|
||
(long, great) and
|
||
<emphasis id="B92BF93BE06CFF84C441FD65AB4512DF" box="[785,868,655,680]" italics="true" pageId="25" pageNumber="527">phallos</emphasis>
|
||
(phallus), referring to the length of the phallus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34576A2E06CFF84C797FD5EAAE0134F" pageId="25" pageNumber="527" type="discussion">
|
||
<paragraph id="8BE02529E06CFF84C797FD5EAAE0134F" blockId="25.[151,1436,151,824]" pageId="25" pageNumber="527">
|
||
<emphasis id="B92BF93BE06CFF84C797FD5EA91512BA" bold="true" box="[199,308,692,717]" pageId="25" pageNumber="527">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C5F5EAAE06CFF84C614FD5FAAAD12BA" box="[324,652,692,717]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="527" phylum="Arthropoda" rank="species" species="megalophallus" status="sp. nov.">
|
||
<emphasis id="B92BF93BE06CFF84C614FD5FAAAD12BA" box="[324,652,692,717]" italics="true" pageId="25" pageNumber="527">Opeatocerata megalophallus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2184440E06CFF84C5C5FD5EAAD012BA" box="[661,753,692,717]" pageId="25" pageNumber="527" rank="species">
|
||
<emphasis id="B92BF93BE06CFF84C5C5FD5EAAD012BA" bold="true" box="[661,753,692,717]" pageId="25" pageNumber="527">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from other species due to the narrow phallus, much longer than the hypandrium (
|
||
<figureCitation id="136439ACE06CFF84C6B8FD3DAA6B1287" box="[488,586,727,752]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 121</figureCitation>
|
||
) (in other species the phallus is either subcylindrical, with a narrow base widening towards the apex or some other shape) and subepandrial sclerite as wide as long (
|
||
<figureCitation id="136439ACE06CFF84C394FD16AD061363" box="[1220,1319,764,789]" captionStart="FIGURES 114 – 122" captionStartId="24.[151,264,1804,1827]" captionTargetBox="[169,1423,200,1776]" captionTargetId="figure@24.[163,1424,193,1783]" captionTargetPageId="24" captionText="FIGURES 114 – 122. Opeatocerata megalophallus sp. nov., holotype ♂. 114. habitus, lateral view; 115. wing; 116. abdomen from tergite 6 to terminalia, lateral view; 117. terminalia, lateral view; 118. lobes of cerci, dorsal view; 119. descendant plate of anterior cercus and hypoproct, posterior view; 120. subepandrial sclerite and bacilliform sclerite, dorsal view; 121. ejaculatory apodeme, phallus and hypandrium, lateral view; 122. hypandrium, ventral view." httpUri="https://zenodo.org/record/252534/files/figure.png" pageId="25" pageNumber="527">Fig. 120</figureCitation>
|
||
) (in other species it is wider than long or longer than wide).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |