Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae) Author Câmara, J. T. Author Rafael, J. A. text Zootaxa 2014 3846 4 502 546 journal article 10.11646/zootaxa.3846.4.2 ea136b1c-dc96-4c41-b078-01ca553453a0 1175-5326 252519 71A80CC3-CDC1-4291-863B-FA7501D60AF5 Opeatocerata megalophallus sp. nov. ( Figs 114–122 ) Diagnosis . Epandrium lobe acuminate at apex ( Figs 116, 117 ); phallus narrow, much longer than the hypandrium ( Fig. 121 ); subepandrial sclerite as wide as long ( Fig. 120 ); bacilliform sclerite shorter than subepandrial sclerite ( Fig. 120 ). Description. Male ( Fig. 114 ). Head : Face slightly divergent toward the proboscis, dark brown, with gray pruinescence visible in ventral view, about 4X longer than lower width. Ocellar tubercle protrubeant, dark brown with gray pruinescence; ocelli brown. Antenna brown to black bristles; postpedicel about 1.5X longer than pedicel; stylus about 1.5X longer than postpedicel. Proboscis yellow shorter than head height. Thorax yellow, shiny ( Fig. 114 ). Legs yellow, except for hind femur and tibia with apical black ring and all tarsomeres 3–5 brown; hind tibia with anterodorsal and posterodorsal rows of longer bristles. Wing ( Fig. 115 ) hyaline, brown pterostigma, conspicuous, about 2.3X longer than wide. Halter yellow. Abdomen yellow, shiny ( Fig. 114 ); tergite 8 divided in two subtriangular plates; sternite 8 divided in two trapezoidal plates. Terminalia : Anterior cercus with proclinate dorsal projection in lateral view ( Figs 116, 117 ), with posterior margin slightly concave in dorsal view ( Fig. 118 ); descendant plate subrectangular in posterior view ( Fig. 119 ). Posterior cercus with acuminate apex in lateral view ( Figs 116, 117 ); with dorsal projections, a subbasal pointed and a submedian subtriangular ( Fig. 118 ). Hypoproct bacilliform lateral view and covering inner surface of the posterior cercus in posterior view ( Fig. 117 ). Epandrial ventral lobe with acuminate apex ( Figs 116, 117 ), with spiniform bristles mesially and longer bristles apically ( Fig. 117 ). Hypandrium longer than wide, with concave apex ( Fig. 122 ), without long bristles. Phallus narrow, much longer than the hypandrium ( Fig. 121 ). Subepandrial sclerite as wide as long, U-shaped ( Fig. 120 ). Bacilliform sclerite without folds and shorter than subepandrial sclerite ( Fig. 120 ). Specimen length : 2.9 mm ; wing length : 2.9 mm . Female . Unknown. FIGURES 114–122 . Opeatocerata megalophallus sp. nov. , holotype ♂. 114 . habitus , lateral view; 115 . wing; 116 . abdomen from tergite 6 to terminalia, lateral view; 117 . terminalia, lateral view; 118 . lobes of cerci, dorsal view; 119 . descendant plate of anterior cercus and hypoproct, posterior view; 120 . subepandrial sclerite and bacilliform sclerite, dorsal view; 121 . ejaculatory apodeme, phallus and hypandrium, lateral view; 122 . hypandrium, ventral view. Geographical distribution . Costa Rica (San Mateo, Guanacaste) ( Fig. 211 ). Type material . HOLOTYPE ♂, labelled: Higuito, San Mateo C[osta] R[ica] / Pablo Schild Coll. ( USNM ). Paratype : Costa Rica , Guanacaste . Bagaces. P.N. Palo Verde. Ext. E. Campo de Aterrizaje, 0–50 m , 16.vii–17.viii.1999 , L. Jiménez, Malaise. L_N_ 260952 _385020 #53259 ( 1 ♂ , INBio ). Holotype condition . Left legs lost; right wing mounted on microslide; abdomen in microtube with glycerin. Etymology . From the Greek mega (long, great) and phallos (phallus), referring to the length of the phallus. Remarks . Opeatocerata megalophallus sp. nov. differs from other species due to the narrow phallus, much longer than the hypandrium ( Fig. 121 ) (in other species the phallus is either subcylindrical, with a narrow base widening towards the apex or some other shape) and subepandrial sclerite as wide as long ( Fig. 120 ) (in other species it is wider than long or longer than wide).