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<document id="36A9594F1C508AC19E1B8983F3672B6C" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFA1FFC67EFCFA50F7D3DE7B" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis parishi Banks, New combination" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="25" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="3941EF8FAC91D31545CBDFF35150C3C3">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<paragraph id="8BE52311FFA1FFC67EFCFA50F7A0DCED" blockId="24.[151,683,1490,1516]" box="[151,683,1490,1516]" pageId="24" pageNumber="25">
<heading id="D0AD947DFFA1FFC67EFCFA50F7A0DCED" bold="true" box="[151,683,1490,1516]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B92EFF03FFA1FFC67EFCFA50F7A0DCED" bold="true" box="[151,683,1490,1516]" pageId="24" pageNumber="25">
<taxonomicName id="4C5A5892FFA1FFC67EFCFA50F4C2DCED" ID-CoL="57H2F" authority="Banks" authorityName="Banks" box="[151,457,1490,1516]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="parishi" status="comb. nov.">
<emphasis id="B92EFF03FFA1FFC67EFCFA50F46ADCED" bold="true" box="[151,353,1490,1516]" italics="true" pageId="24" pageNumber="25">Tortopsis parishi</emphasis>
(Banks)
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<taxonomicName id="4C5A5892FFA1FFC67EFCF999F718DF33" authority="Banks 1918: 10" authorityName="Banks" authorityPageNumber="10" authorityYear="1918" box="[151,531,1563,1586]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="parishi">
<emphasis id="B92EFF03FFA1FFC67EFCF999F469DF33" box="[151,354,1563,1586]" italics="true" pageId="24" pageNumber="25">Campsurus parishi</emphasis>
<treatmentCitation id="0AFB0500FFA1FFC67F02F999F718DF33" author="Banks" box="[361,531,1563,1586]" page="10" pageId="24" pageNumber="25" year="1918">
<bibRefCitation id="EFCB5EE0FFA1FFC67F02F999F4E0DF33" author="Banks" box="[361,491,1563,1586]" pageId="24" pageNumber="34" refString="Banks, N. (1918) New neuropteroid insects. Bulletin of the Museum of Comparative Zoology, 62 (1), 1 - 22." type="journal article" year="1918">Banks 1918</bibRefCitation>
: 10
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</taxonomicName>
.
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<emphasis id="B92EFF03FFA1FFC67EFCF9B9F44CDF53" box="[151,327,1595,1618]" italics="true" pageId="24" pageNumber="25">Tortopus parishi</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AFB0500FFA1FFC67F3EF9B9F706DF53" author="Ulmer" box="[341,525,1595,1618]" page="110" pageId="24" pageNumber="25" year="1942">
<bibRefCitation id="EFCB5EE0FFA1FFC67F3EF9B9F4D2DF53" author="Ulmer" box="[341,473,1595,1618]" pageId="24" pageNumber="35" refString="Ulmer, G. (1942) Alte und neue Eintagsfliegen (Ephemeropteren) aus Sud- und Mittelamerika. Stettiner Entomologische Zeitung, 103, 98 - 128." type="journal article" year="1942">Ulmer 1942</bibRefCitation>
: 110
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.
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</paragraph>
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<paragraph id="8BE52311FFA1FFC67EFCF902F11ADFBE" blockId="24.[151,1437,1664,1914]" pageId="24" pageNumber="25">
Material: the only specimen known (male
<typeStatus id="54E19DB3FFA1FFC67CEEF902F7E6DF9B" box="[645,749,1664,1690]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
) is deposited at Harvard. In spite of the attempts to borrow the material, neither the specimen nor photographs were available for study.
</paragraph>
<paragraph id="8BE52311FFA1FFC67EADF948F7D3DE7B" blockId="24.[151,1437,1664,1914]" pageId="24" pageNumber="25">
Discussion and diagnosis. This species is known from one male imago from
<collectingCountry id="F34D6381FFA1FFC67AF8F948F1FEDFE5" box="[1171,1269,1738,1764]" name="Ecuador" pageId="24" pageNumber="25">Ecuador</collectingCountry>
(Durar). The original description and figures are inadequate to species characterization but clearly indicate that this species belongs to
<taxonomicName id="4C5A5892FFA1FFC67F70F894F483DE2E" box="[283,392,1814,1839]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA1FFC67F70F894F483DE2E" box="[283,392,1814,1839]" italics="true" pageId="24" pageNumber="25">Tortopsis</emphasis>
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, as is recovered in the phylogenetic tree (
<figureCitation id="13613F94FFA1FFC67D16F897F6D1DE2E" box="[893,986,1813,1839]" captionStart="FIGURE 1. A" captionStartId="4.[151,256,1731,1755]" captionTargetBox="[207,1379,197,1701]" captionTargetId="figure@4.[207,1380,197,1701]" captionTargetPageId="4" captionText="FIGURE 1. A, strict consensus of 234 shortest trees obtained with the complete data set (58 steps, Ci = 81, Ri = 93); B, strict consensus of 3 shortest trees obtained with the reduced data set (58 steps, Ci = 81, Ri = 92). Values near nodes indicate group support (frequency difference / absolute Bremer Support / relative BS). Black circles indicate unique apomorphies, white circles indicate parallelism or reversal, the small numbers above and below these marks are character and state numbers respectively." httpUri="https://zenodo.org/record/195432/files/figure.png" pageId="24" pageNumber="25">Fig. 1A</figureCitation>
). The main feature used to assign the species to
<taxonomicName id="4C5A5892FFA1FFC67F7EF8B9F48BDE55" box="[277,384,1851,1876]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA1FFC67F7EF8B9F48BDE55" box="[277,384,1851,1876]" italics="true" pageId="24" pageNumber="25">Tortopsis</emphasis>
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is the form of the penes, widely separated from each other from their bases. Fore wing length is reported to be
<quantity id="4CA28EF4FFA1FFC67FC7F8E2F720DE7B" box="[428,555,1888,1914]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="24" pageNumber="25" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
(
<bibRefCitation id="EFCB5EE0FFA1FFC67C51F8E2F7C0DE7B" author="Banks" box="[570,715,1888,1914]" pageId="24" pageNumber="34" refString="Banks, N. (1918) New neuropteroid insects. Bulletin of the Museum of Comparative Zoology, 62 (1), 1 - 22." type="journal article" year="1918">Banks 1918</bibRefCitation>
).
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