A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae) Author Molineri, Carlos text Zootaxa 2010 2481 1 36 journal article 10.5281/zenodo.195431 2df8bd5a-e125-42aa-aa6f-ae3e78b83929 1175-5326 195431 Tortopsis parishi (Banks) New combination Campsurus parishi Banks 1918 : 10 . Tortopus parishi ; Ulmer 1942 : 110 . Material: the only specimen known (male holotype ) is deposited at Harvard. In spite of the attempts to borrow the material, neither the specimen nor photographs were available for study. Discussion and diagnosis. This species is known from one male imago from Ecuador (Durar). The original description and figures are inadequate to species characterization but clearly indicate that this species belongs to Tortopsis , as is recovered in the phylogenetic tree ( Fig. 1A ). The main feature used to assign the species to Tortopsis is the form of the penes, widely separated from each other from their bases. Fore wing length is reported to be 10–12 mm ( Banks 1918 ).