268 lines
25 KiB
XML
268 lines
25 KiB
XML
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<mods:title id="B46A4C02A7126CD7F246D97841B707F6">Figs. 4 - 10 in Descriptions of Two New Species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) in Korea</mods:title>
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<mods:namePart id="D605663473B329632190E1BFBAFC3ADA">Arias-Buriticá, Jorge Armando</mods:namePart>
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<mods:namePart id="AA270569B971480DD86C71B90E885FED">Bach, Andressa</mods:namePart>
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<mods:namePart id="9761DB833B257149FAB6622CEB86A6C9">Peres, Carlos A.</mods:namePart>
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<mods:namePart id="FBC9E475FE72BEA60DA289D4ED99633F">Haugaasen, Torbjørn</mods:namePart>
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<mods:namePart id="ED93C3D34EE7CFB027EF220E68A14E5E">Hawes, Joseph E.</mods:namePart>
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<mods:namePart id="50B053629E30F7D787226C91E1887072">Azevedo, Renato A.</mods:namePart>
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<mods:namePart id="C163E2ABC13C054949C51A1BB40B340B">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="D3DEA7900A6920DACE99F226658521F6">The Coleopterists Bulletin</mods:title>
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<mods:date id="00052240F870A9C5F7FC5BA7F2E361C7">2023</mods:date>
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<mods:number id="28DCDFD8C0AC46E438D93B9724B9C64A">2023-12-21</mods:number>
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<mods:url id="3F0E830A94E47B39A6C56813156357D9">http://dx.doi.org/10.1649/0010-065x-77.4.629</mods:url>
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<mods:identifier id="CA2AA779A8BF08E5527FE84B8FFFF2BD" type="DOI">10.1649/0010-065X-77.4.629</mods:identifier>
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<taxonomicName id="4C4E4D58FFC5FFDDFE51E6B0A473FF6F" ID-CoL="55F6" authority="PEREIRA AND MARTINEZ, 1960" authorityName="PEREIRA AND MARTINEZ" authorityYear="1960" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
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<smallCapsWord id="8D17A007FFC5FFDDFE51E6B0A460FF4E" baselines="211,212" box="[461,562,194,217]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Isocopris" pageId="6" pageNumber="634">
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<emphasis id="B93AEAC9FFC5FFDDFE51E6B0A460FF4E" bold="true" box="[461,562,194,217]" italics="true" pageId="6" pageNumber="634">ISOCOPRIS</emphasis>
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<smallCapsWord id="8D17A007FFC5FFDDFED1E695A727FF6E" baselines="244" box="[333,373,231,248]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="6" pageNumber="634">AND</smallCapsWord>
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, 1960
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<paragraph id="8BF136DBFFC5FFDDFF2CE77CA75AFE3F" blockId="6.[152,638,270,425]" pageId="6" pageNumber="634">
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This key is an adaptation of Rossini and Vaz-de-Mello (2017), updated to include the new species. The sentences between quotation marks (“”) have been transcribed
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<emphasis id="B93AEAC9FFC5FFDDFEDCE72CA7E6FEE5" box="[320,436,350,371]" italics="true" pageId="6" pageNumber="634">ipsis litteris</emphasis>
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from this paper and the figures of the others species are available in the same work.
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<keyLead id="30BF95EEFFC5FFDDFF04E7CCA42FFDE5" pageId="6" pageNumber="634">
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1. “Pronotum abruptly vertical above the head, superiorly with a pair of transverse humps. Male with a cephalic horn always flanked by two low and transverse carinae; female with a cephalic carina tuberculate at middle. Large-sized species, body length>
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” ..................................................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="8BF136DBFFC5FFDDFF04E405A42FFCB8" blockId="6.[152,638,446,1722]" pageId="6" pageNumber="634">
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<keyLead id="30BF95EEFFC5FFDDFF04E405A42FFCB8" pageId="6" pageNumber="634">
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1′. “Pronotum simply convex, never with distinct humps on the superior side. Male either with a simple and conical horn or a transverse and strong carina; female with either a cephalic horn or carina, but never tuberculate at middle. Medium-sized species, body length <
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<quantity id="4CB69B3EFFC5FFDDFF7DE56BA774FCB8" box="[225,294,793,814]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="634" unit="mm" value="22.0">22 mm</quantity>
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” .................................................... 3
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</keyLead>
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</paragraph>
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<paragraph id="8BF136DBFFC5FFDDFF04E546A42CFBC2" blockId="6.[152,638,446,1722]" pageId="6" pageNumber="634">
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<keyLead id="30BF95EEFFC5FFDDFF04E546A42CFBC2" pageId="6" pageNumber="634">
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2. “Clypeus wide and evenly curved, slightly sinuated at middle, male with cephalic horn cylindrical to conical (from above), frontal side of the horn always finely wrinkled; female with cephalic carina on the fronto-clypeal region, anteromedial region of pronotum feebly excavated, pronotal protuberance very weak (figs. 1a, b). Anterior pronotal bead simple, metasternal surface completely covered by long and straight orange hairs.
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||
<collectingCountry id="F359764BFFC5FFDDFE53E256A458FBAC" box="[463,522,1060,1082]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
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||
,
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||
<collectingCountry id="F359764BFFC5FFDDFD8AE256A424FBAC" box="[534,630,1060,1082]" name="Argentina" pageId="6" pageNumber="634">Argentina</collectingCountry>
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” ..............
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<taxonomicName id="4C4E4D58FFC5FFDDFEB5E24CA42CFBC2" authority="(Germar, 1824)" baseAuthorityName="Germar" baseAuthorityYear="1824" box="[297,638,1086,1108]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="inhiatus">
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||
<emphasis id="B93AEAC9FFC5FFDDFEB5E24CA42CFBC2" bold="true" box="[297,638,1086,1108]" pageId="6" pageNumber="634">
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||
<emphasis id="B93AEAC9FFC5FFDDFEB5E24CA789FBC2" bold="true" box="[297,475,1086,1108]" italics="true" pageId="6" pageNumber="634">Isocopris inhiatus</emphasis>
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(Germar, 1824)
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</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph id="8BF136DBFFC5FFDDFF04E22AA42CFA39" blockId="6.[152,638,446,1722]" pageId="6" pageNumber="634">
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<keyLead id="30BF95EEFFC5FFDDFF04E22AA42CFA39" pageId="6" pageNumber="634">
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2′. “Clypeus elongated forward and clearly narrower at middle, lateral margins straight, male with cephalic horn rectangular and transversally elongated, frontal side of the horn smooth to finely punctuated; female with cephalic carina between eyes, anteromedial region of the pronotum distinctly excavated, pronotal humps distinct (figs. 2a, b). Anterior pronotal bead slightly concave at the middle of the posterior margin, metasternum with long hairs along the inner side of mesocoxae, metasternal disc bare.
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||
<collectingCountry id="F359764BFFC5FFDDFE38E30DA78CFA03" box="[420,478,1407,1429]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
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||
”......................... ...............
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||
<taxonomicName id="4C4E4D58FFC5FFDDFEACE3EBA42CFA39" authority="(Lucas, 1857)" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[304,638,1433,1455]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="hypocrita">
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||
<emphasis id="B93AEAC9FFC5FFDDFEACE3EBA42CFA39" bold="true" box="[304,638,1433,1455]" pageId="6" pageNumber="634">
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||
<emphasis id="B93AEAC9FFC5FFDDFEACE3EBA7BCFA39" bold="true" box="[304,494,1433,1455]" italics="true" pageId="6" pageNumber="634">Isocopris hypocrita</emphasis>
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(Lucas, 1857)
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</emphasis>
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30BA2E7EFFC5FFDDFF04E3C6A2C1FEBE" pageId="6" pageNumber="634">
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<paragraph id="8BF136DBFFC5FFDDFF04E3C6A42FF912" blockId="6.[152,638,446,1722]" pageId="6" pageNumber="634">
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||
<keyLead id="30BF95EEFFC5FFDDFF04E3C6A42FF912" pageId="6" pageNumber="634">3. “Both sexes with a strong fronto-clypeal carina flanked by two hump-like weak tubercles, carina acuminated on both sides in male, straight in female, frons deeply depressed (figs. 5a, b). Margin of the head distinctly notched at the clypeo-genal junction, body completely black, head and pronotum shining, elytra more opaque” ....................................................... 4</keyLead>
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||
</paragraph>
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||
<paragraph id="8BF136DBFFC5FFDDFF04E0FAA2C1FEBE" blockId="6.[152,638,446,1722]" lastBlockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
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<keyLead id="30BF95EEFFC5FFDDFF04E0FAA2C1FEBE" pageId="6" pageNumber="634">3′. “Both sexes with a conical cephalic horn, margin of the head either notched or angulated (genal margin wider than clypeus) at the clypeo-genal junction, frons either depressed or normal, clypeus weakly to distinctly reflexed at middle” ......................................... 5</keyLead>
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</paragraph>
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||
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<paragraph id="8BF136DBFFC5FFDDFD31E75CA2C1FD96" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
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<keyLead id="30BF95EEFFC5FFDDFD31E75CA2C1FD96" pageId="6" pageNumber="634">
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4. Both sexes with ventral clypeal process large and trapezoidal in lateral view (figs. 1e, f). Metaventral anterior lobe glabrous (figs. 1b, c). Females with elytral striae from the first to the fourth widened in the central area (fig. 1c). Southwestern Amazonia,
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||
<collectingCountry id="F359764BFFC5FFDDFB97E7C6A214FE5C" box="[1035,1094,436,458]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
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||
(fig. 3) .................
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<emphasis id="B93AEAC9FFC5FFDDFCD6E7BDA2C1FD96" bold="true" pageId="6" pageNumber="634">
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFCD6E7BDA503FE69" authority="Arias-Buritica, Bach" authorityName="Arias-Buritica, Bach" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="rossinii">
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||
<emphasis id="B93AEAC9FFC5FFDDFCD6E7BDA5A6FE72" bold="true" box="[842,1012,463,484]" italics="true" pageId="6" pageNumber="634">Isocopris rossinii</emphasis>
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||
Arias-Buriticá, Bach
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</taxonomicName>
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, and Vaz-de-Mello, new species
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||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E471A2C1FD37" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E471A2C1FD37" pageId="6" pageNumber="634">
|
||
4′. Both sexes with ventral clypeal process small and tooth-shaped in lateral view. Metaventral anterior lobe with setae. Females without widened elytral striae. Cerrado habitat.
|
||
<collectingCountry id="F359764BFFC5FFDDFBC5E427A2C1FDFD" box="[1113,1171,597,619]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
|
||
.............
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFCA5E402A2C1FD37" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B93AEAC9FFC5FFDDFCA5E402A2C1FD37" bold="true" pageId="6" pageNumber="634">
|
||
<emphasis id="B93AEAC9FFC5FFDDFCA5E402A5AEFD10" bold="true" box="[825,1020,624,646]" italics="true" pageId="6" pageNumber="634">Isocopris foveolatus</emphasis>
|
||
(Luederwaldt, 1931)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E7EFFC5FFDDFD31E4D4A2C1FC5F" pageId="6" pageNumber="634">
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E4D4A2C1FCD4" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E4D4A2C1FCD4" pageId="6" pageNumber="634">
|
||
5. “Fore tibia with an inner-apical projection, tooth-like and directed inward, last protarsal segment strongly enlarged at the apex, tarsal claws very long and strongly folded (figs. 6a, b, e).
|
||
<collectingCountry id="F359764BFFC5FFDDFC85E560A504FCBE" box="[793,854,786,808]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
|
||
”......................
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFC76E560A2C1FCD4" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="B93AEAC9FFC5FFDDFC76E560A2C1FCD4" bold="true" pageId="6" pageNumber="634">
|
||
<emphasis id="B93AEAC9FFC5FFDDFC76E560A2C1FCB1" bold="true" box="[1002,1171,785,807]" italics="true" pageId="6" pageNumber="634">Isocopris tarsalis</emphasis>
|
||
(Luederwaldt, 1931)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E534A2C1FC5F" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E534A2C1FC5F" pageId="6" pageNumber="634">5′. “Fore tibia either normal or with an inner-apical projection directed forward, last protarsal segment normal and never strongly enlarged, tarsal claws short and weakly curved” ..................................................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E7EFFC5FFDDFD31E5BCA2C1FA75" pageId="6" pageNumber="634">
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E5BCA2C0FB66" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E5BCA2C0FB66" pageId="6" pageNumber="634">
|
||
6. “Clypeus acuminate and sides feebly curved, weakly reflexed and distinctly sinuated at middle, with two obtuse teeth, head of both sexes with three conical tubercles, central tubercle stronger, lateral ones slightly backward, frons not depressed. Apical spur of the fore tibia with superior margin normal. Body black, elytra opaque to feebly shining and completely sericeous.
|
||
<collectingCountry id="F359764BFFC5FFDDFCCCE2D7A5DFFB2D" box="[848,909,1189,1211]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
|
||
(only known of north of
|
||
<collectingRegion id="498AF839FFC5FFDDFD7FE2B2A531FB40" box="[739,867,1216,1238]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="634">Minas Gerais</collectingRegion>
|
||
), dry forest” ................
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFBA6E2B2A2C0FB66" authority="Rossini" authorityName="Rossini" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="xacriaba">
|
||
<emphasis id="B93AEAC9FFC5FFDDFBA6E2B2A2C0FB66" bold="true" pageId="6" pageNumber="634">
|
||
<emphasis id="B93AEAC9FFC5FFDDFBA6E2B2A506FB67" bold="true" italics="true" pageId="6" pageNumber="634">Isocopris xacriaba</emphasis>
|
||
Rossini and Vaz-de-Mello, 2017
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E287A2C1FA75" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E287A2C1FA75" pageId="6" pageNumber="634">6′. “Clypeus with sides sinuated, strongly reflexed at middle, either transversally truncated or weakly sinuated, both sexes with a central conical horn flanked by two weak hump-like tubercles, frons slightly to distinctly depressed. Apical spur of the fore tibia with superior margin bifid (fig. 3h). Body completely shining, black to brownish. Known of Amazon and dry forests”............ 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E7EFFC5FFDDFD31E39BA2C1F92C" pageId="6" pageNumber="634">
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E39BA2C1F9D9" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E39BA2C1F9D9" pageId="6" pageNumber="634">
|
||
7. “Pronotum and elytra with a dense and shallow punctuation evenly distributed (figs. 3b, c). Size of
|
||
<quantity id="4CB69B3EFFC5FFDDFCD7E06CA5E7F9A5" box="[843,949,1566,1588]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="6" pageNumber="634" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15–22 mm</quantity>
|
||
.
|
||
<collectingCountry id="F359764BFFC5FFDDFC22E06CA5A9F9A2" box="[958,1019,1566,1588]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
|
||
”........
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFBA4E06CA2C1F9D9" authority="(Felsche, 1901)" baseAuthorityName="Felsche" baseAuthorityYear="1901" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B93AEAC9FFC5FFDDFBA4E06CA2C1F9D9" bold="true" pageId="6" pageNumber="634">
|
||
<emphasis id="B93AEAC9FFC5FFDDFBA4E06CA5A7F9D8" bold="true" italics="true" pageId="6" pageNumber="634">Isocopris imitator</emphasis>
|
||
(Felsche, 1901)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136DBFFC5FFDDFD31E021A2C1F92C" blockId="6.[685,1177,194,1722]" pageId="6" pageNumber="634">
|
||
<keyLead id="30BF95EEFFC5FFDDFD31E021A2C1F92C" pageId="6" pageNumber="634">
|
||
7′. “Pronotum and elytra with a very fine and inconspicuous punctuation (figs. 4b, c). Size of
|
||
<quantity id="4CB69B3EFFC5FFDDFD7FE0F8A51FF909" box="[739,845,1674,1695]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="6" pageNumber="634" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
|
||
.
|
||
<collectingCountry id="F359764BFFC5FFDDFCCBE0F8A5BAF936" box="[855,1000,1674,1696]" name="French Guiana" pageId="6" pageNumber="634">French Guiana</collectingCountry>
|
||
,
|
||
<collectingCountry id="F359764BFFC5FFDDFC6EE0F8A27FF936" box="[1010,1069,1674,1696]" name="Brazil" pageId="6" pageNumber="634">Brazil</collectingCountry>
|
||
and
|
||
<collectingCountry id="F359764BFFC5FFDDFBC0E0F8A2D8F909" box="[1116,1162,1674,1695]" name="Peru" pageId="6" pageNumber="634">Peru</collectingCountry>
|
||
” ........
|
||
<taxonomicName id="4C4E4D58FFC5FFDDFC84E0D6A2C1F92C" authority="(Luederwaldt, 1922)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[792,1171,1700,1722]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="B93AEAC9FFC5FFDDFC84E0D6A2C1F92C" bold="true" box="[792,1171,1700,1722]" pageId="6" pageNumber="634">
|
||
<emphasis id="B93AEAC9FFC5FFDDFC84E0D6A5EFF92C" bold="true" box="[792,957,1700,1722]" italics="true" pageId="6" pageNumber="634">Isocopris nitidus</emphasis>
|
||
(Luederwaldt, 1922)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |