treatments-xml/data/03/D3/87/03D387630C328E5E72B0FC7BFBE057E6.xml

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<mods:title id="563C3276A123B499C7291ADD7460DB31">Devadatta adii sp. nov. (Odonata: Zygoptera: Devadattidae) from Upper Siang District, Arunachal Pradesh, India</mods:title>
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<mods:namePart id="4CF2982F4CBC76087EECAA9D4BD52ADC">Sawant, Dattaprasad</mods:namePart>
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<materialsCitation id="3B123C280C328E4F72E0FBB6FA985211" box="[199,1382,1122,1152]" collectingDate="2022-05-23" collectorName="Fahim Khan &amp; Krushnamefh Kunte &amp; Viraj Nawge" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCode="IBC-BR297, IBC-BR300" specimenCount="2" specimenCount-male="2" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F72E0FBB6FE7352EC" bold="true" box="[199,397,1122,1152]" pageId="1" pageNumber="572">
<typeStatus id="54C188D70C328E4F72E0FBB6FEBF5211" box="[199,321,1123,1149]" pageId="1" pageNumber="572" type="paratype">Paratypes</typeStatus>
. 
<specimenCount id="9D7CFDFC0C328E4F736AFBB7FE7352EC" box="[333,397,1122,1152]" count="2" pageId="1" pageNumber="572" type="male">2 ♂♂</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F73BBFBB6FDD65211" box="[412,552,1123,1149]" pageId="1" pageNumber="572">IBC-BR297</specimenCode>
, 
<specimenCode id="DBDC9E0E0C328E4F7013FBB6FD3E5211" box="[564,704,1123,1149]" pageId="1" pageNumber="572">IBC-BR300</specimenCode>
), location, date of collection and collector same as holotype
</materialsCitation>
; 
<materialsCitation id="3B123C280C328E4F7755FBB7FC7852CD" collectingDate="2022-05-22" collectorName="Fahim Khan &amp; Krushnamefh Kunte &amp; Viraj Nawge" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCode="IBC-BR383" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F7755FBB7FA6252EC" bold="true" box="[1394,1436,1122,1152]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F7755FBB7FA6252EC" box="[1394,1436,1122,1152]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F72B8FB52FED452CD" box="[159,298,1159,1185]" pageId="1" pageNumber="572">IBC-BR383</specimenCode>
) same location and collectors as holotype, 
<date id="FFC410B50C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">
<collectingDate id="EF80E95D0C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">22.v.2022</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B123C280C328E4F71B4FB53FE0E52A9" collectingDate="2022-05-29" collectorName="Shantanu Joshi" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCode="IBC-BR451" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F71B4FB53FC4052C8" bold="true" box="[915,958,1158,1188]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F71B4FB53FC4052C8" box="[915,958,1158,1188]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F71E9FB53FBA552CD" box="[974,1115,1158,1185]" pageId="1" pageNumber="572">IBC-BR451</specimenCode>
), same location as holotype, 
<date id="FFC410B50C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">
<collectingDate id="EF80E95D0C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">29.v.2022</collectingDate>
</date>
<collectorName id="268F53A30C328E4F7328FB7EFE4552A9" box="[271,443,1195,1221]" pageId="1" pageNumber="572">Shantanu Joshi</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B123C280C328E4F73D8FB7FFB8D52A9" box="[511,1139,1194,1224]" collectingDate="2022-05-23" collectorName="Fahim Khan &amp; Krushnamefh Kunte &amp; Viraj Nawge" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCode="IBC-BR314" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<emphasis id="B90EEA670C328E4F73D8FB7FFDD552A4" bold="true" box="[511,555,1194,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F73D8FB7FFDD552A4" box="[511,555,1194,1224]" count="1" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F701AFB7EFD3652A9" box="[573,712,1195,1221]" pageId="1" pageNumber="572">IBC-BR314</specimenCode>
) all details are the same as holotype
</materialsCitation>
. 
<materialsCitation id="3B123C280C328E4F76A7FB79FA925361" collectingDate="2024-05-23" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" elevation="460" latitude="28.6642" location="Sirring Nala" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCode="IBC-BX307, IBC-BX309, IBC-BX313" specimenCount="3" specimenCount-female="3" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F76A7FB79FB4952A4" bold="true" box="[1152,1207,1196,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F76A7FB79FB4952A4" box="[1152,1207,1196,1224]" count="3" pageId="1" pageNumber="572" type="female">3♀♀</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F76EEFB7EFAA852A9" box="[1225,1366,1195,1221]" pageId="1" pageNumber="572">IBC-BX307</specimenCode>
, 
<specimenCode id="DBDC9E0E0C328E4F7743FB7EFF155285" pageId="1" pageNumber="572">IBC-BX309</specimenCode>
, 
<specimenCode id="DBDC9E0E0C328E4F72DEFB1AFE7D5285" box="[249,387,1231,1257]" pageId="1" pageNumber="572">IBC-BX313</specimenCode>
) 
<location id="8EA560AE0C328E4F73B2FB1AFDDA5285" LSID="urn:lsid:plazi:treatment:03D387630C328E5E72B0FC7BFBE057E6:8EA560AE0C328E4F73B2FB1AFDDA5285" box="[405,548,1231,1257]" country="India" latitude="28.6642" municipality="Upper Siang District" name="Sirring Nala" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Sirring Nala</location>
, 
<collectingMunicipality id="6BA1AC0F0C328E4F7016FB1AFCDF5285" box="[561,801,1231,1257]" pageId="1" pageNumber="572">Upper Siang District</collectingMunicipality>
, 
<collectingRegion id="49BEF8970C328E4F7109FB1AFBF65285" box="[814,1032,1231,1257]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="F36D76E50C328E4F7632FB1AFBB15285" box="[1045,1103,1231,1257]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F7647FB1AFB235285" box="[1120,1245,1231,1257]" degrees="28.6642" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.6642">28.6642 N</geoCoordinate>
, 
<geoCoordinate id="EE4E50B20C328E4F76CCFB1AFA9A5285" box="[1259,1380,1231,1257]" pageId="1" pageNumber="572">94.994 3 E</geoCoordinate>
; 
<elevation id="0057D1460C328E4F7755FB1AFF145361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">
<quantity id="4C829B900C328E4F7755FB1AFF535361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">460 m</quantity>
a.s.l.
</elevation>
), 
<date id="FFC410B50C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">
<collectingDate id="EF80E95D0C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">23.v.2024</collectingDate>
</date>
, 
<collectorName id="268F53A30C328E4F7358FB26FD925361" box="[383,620,1267,1293]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
, 
<collectorName id="268F53A30C328E4F705EFB26FCF95361" box="[633,775,1267,1293]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
, 
<collectorName id="268F53A30C328E4F7133FB26FC1F5361" box="[788,993,1267,1293]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
, 
<collectorName id="268F53A30C328E4F71CBFB26FB4B5361" box="[1004,1205,1267,1293]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
, 
<collectorName id="268F53A30C328E4F76E6FB26FAC95361" box="[1217,1335,1267,1293]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation id="3B123C280C328E4F775EFB21FA9C5339" collectingDate="2024-06-02" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" county="Upper Siang District" elevation="360" latitude="28.7639" location="Yinkiong" municipality="Likor Road" pageId="1" pageNumber="572" specimenCode="IBC-BX390" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F775EFB21FA62537C" bold="true" box="[1401,1436,1268,1296]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F775EFB21FA62537C" box="[1401,1436,1268,1296]" count="1" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
</emphasis>
(
<specimenCode id="DBDC9E0E0C328E4F72B8FAC2FED5535D" box="[159,299,1303,1329]" pageId="1" pageNumber="572">IBC-BX390</specimenCode>
) 
<location id="8EA560AE0C328E4F731FFAC2FE5E535D" LSID="urn:lsid:plazi:treatment:03D387630C328E5E72B0FC7BFBE057E6:8EA560AE0C328E4F731FFAC2FE5E535D" box="[312,416,1303,1329]" country="India" county="Upper Siang District" latitude="28.7639" municipality="Likor Road" name="Yinkiong" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Yinkiong</location>
to 
<collectingMunicipality id="6BA1AC0F0C328E4F73E4FAC2FDB8535D" box="[451,582,1303,1329]" pageId="1" pageNumber="572">Likor Road</collectingMunicipality>
, 
<collectingCounty id="62A44EF90C328E4F7076FAC2FCC4535D" box="[593,826,1303,1329]" pageId="1" pageNumber="572">Upper Siang District</collectingCounty>
, 
<collectingRegion id="49BEF8970C328E4F7164FAC2FBE5535D" box="[835,1051,1303,1329]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="F36D76E50C328E4F7602FAC2FBA1535D" box="[1061,1119,1303,1329]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F764BFAC2FB19535D" box="[1132,1255,1303,1329]" degrees="28.7639" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.7639">28.7639 N</geoCoordinate>
, 
<geoCoordinate id="EE4E50B20C328E4F76D6FAC2FA99535D" box="[1265,1383,1303,1329]" pageId="1" pageNumber="572">94.899 9 E</geoCoordinate>
; 
<elevation id="0057D1460C328E4F7755FAC2FF175339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">
<quantity id="4C829B900C328E4F7755FAC2FF535339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">360 m</quantity>
a.s.l.
</elevation>
), 
<date id="FFC410B50C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">
<collectingDate id="EF80E95D0C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">02.vi.2024</collectingDate>
</date>
, 
<collectorName id="268F53A30C328E4F73A2FAEEFD8E5339" box="[389,624,1339,1365]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
, 
<collectorName id="268F53A30C328E4F705BFAEEFCF75339" box="[636,777,1339,1365]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
, 
<collectorName id="268F53A30C328E4F7132FAEEFC1E5339" box="[789,992,1339,1365]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
, 
<collectorName id="268F53A30C328E4F71CDFAEEFB4C5339" box="[1002,1202,1339,1365]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
, 
<collectorName id="268F53A30C328E4F769BFAEEFACF5339" box="[1212,1329,1339,1365]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4F72E0FA8AFE5B5389" pageId="1" pageNumber="572" type="etymology">
<paragraph id="8BC536750C328E4F72E0FA8AFDB553F1" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA8AFEB05315" bold="true" box="[199,334,1375,1401]" pageId="1" pageNumber="572">Etymology.</emphasis>
The specific epithet ‘ 
<emphasis id="B90EEA670C328E4F7018FA8AFD8A5315" box="[575,628,1375,1401]" italics="true" pageId="1" pageNumber="572">adii’</emphasis>
honours the indigenous Adi tribe of the Upper Siang District, 
<collectingRegion id="49BEF8970C328E4F770FFA8AFF0D53F1" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="F36D76E50C328E4F72D9FA56FEC653F1" box="[254,312,1411,1437]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
and neighbouring areas.
</paragraph>
<paragraph id="8BC536750C328E4F72E0FA72FE5B5389" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA72FE0C53AD" bold="true" box="[199,498,1446,1473]" pageId="1" pageNumber="572">Proposed English Name.</emphasis>
Golden Grissette: The common English name is given due to the golden yellow pterothorax of the male.
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4472B0F9C7FB95572D" lastPageId="10" lastPageNumber="581" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC536750C328E4F72B0F9C7FCFD5041" blockId="1.[151,1437,1554,2013]" box="[151,771,1554,1581]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9C7FD7F5040" bold="true" box="[151,641,1554,1581]" pageId="1" pageNumber="572">
Description of 
<typeStatus id="54C188D70C328E4F7360F9C6FE505041" box="[327,430,1555,1581]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
male IBC-BR299
</emphasis>
(
<figureCitation id="13412AF00C328E4F70B7F9C6FD045041" box="[656,762,1555,1581]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[214,1373,181,1919]" captionTargetId-0="figure-18@2.[214,1373,181,1919]" captionTargetId-1="figure-17@3.[214,1373,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." captionText-1="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916295" figureDoi-1="http://doi.org/10.5281/zenodo.13916297" httpUri-0="https://zenodo.org/record/13916295/files/figure.png" httpUri-1="https://zenodo.org/record/13916297/files/figure.png" pageId="1" pageNumber="572">Figs. 1–2</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C328E4F72B0F9E2FACE50D1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9E2FF2A503D" box="[151,212,1591,1617]" italics="true" pageId="1" pageNumber="572">Head</emphasis>
(
<figureCitation id="13412AF00C328E4F72CFF9E2FEC2503D" box="[232,316,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1a</figureCitation>
). Head black, marked as follows: labrum yellowish brown; lower half of mandibles pale blue; labium bluish-yellow except for a round dark area at the center of the upper margin in life; anteclypeus pale blue; postclypeus blackish-brown; frons black. Median ocellus creamy yellow, rest shiny brown; antennae black, upper margin of pedicel pale bluish yellow. Eyes upper half brown, lower half grey in life, darker post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F912FED05145" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F912FE69508D" box="[199,407,1735,1761]" italics="true" pageId="1" pageNumber="572">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7381F912FDEA508D" box="[422,532,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1b, d</figureCitation>
). Anterior and posterior lobes black, middle lobes brown; propleuron dark brown with downward projection at posterior end: posterior margin pruinosed bluish-white in life, dark brown to black post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F8E6FDE851B1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F8E6FEB35121" box="[199,333,1843,1869]" italics="true" pageId="1" pageNumber="572">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7347F8E6FE155121" box="[352,491,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1c, e–f</figureCitation>
). Mesostigmal plates and mesepisternum matt black with greenish metallic sheen; yellow stripe on mesepisternum above mesopleural suture; pterothorax laterally golden yellow in life, pale brown post-mortem; upper part of mesepimeron pale brown, faint brown markings on the interpleural suture and anterior part of metepisternum; mesinfraepisternum brown, metinfraepisternum yellow. Pterothorax ventrally pale yellow, small black spot at posterior edge.
</paragraph>
<caption id="DF0566FD0C318E4C72B0F842FDE3519B" ID-DOI="http://doi.org/10.5281/zenodo.13916295" ID-Zenodo-Dep="13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BC536750C318E4C72B0F842FDE3519B" blockId="2.[151,1437,1943,2039]" pageId="2" pageNumber="573">
<emphasis id="B90EEA670C318E4C72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="573">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4DF60C318E4C733FF842FE5151C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[280,431,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C318E4C733FF842FE5151C3" box="[280,431,1943,1967]" italics="true" pageId="2" pageNumber="573">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C318E4C7392F842FDFB51C3" bold="true" box="[437,517,1943,1967]" pageId="2" pageNumber="573">
<taxonomicNameLabel id="A23D571C0C318E4C7392F842FDFB51C3" box="[437,517,1943,1967]" pageId="2" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c).
</paragraph>
</caption>
<caption id="DF0566FD0C308E4D72B0F842FDB2519B" ID-DOI="http://doi.org/10.5281/zenodo.13916297" ID-Zenodo-Dep="13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="8BC536750C308E4D72B0F842FDB2519B" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="574">
<emphasis id="B90EEA670C308E4D72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="3" pageNumber="574">FIGURE 2.</emphasis>
<taxonomicName id="4C7A4DF60C308E4D733DF842FE4C51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[282,434,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C308E4D733DF842FE4C51C3" box="[282,434,1943,1967]" italics="true" pageId="3" pageNumber="574">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C308E4D739EF842FDF451C3" bold="true" box="[441,522,1943,1967]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A23D571C0C308E4D739EF842FDF451C3" box="[441,522,1943,1967]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C378E4A72E0FF42FD7D56B9" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF42FF0256DC" box="[199,252,151,176]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae pale brown with pruinosed blue in life, pale brown post-mortem; femur, tibiae brown. Tibiofemoral joint, tarsus and spines dark brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FF0AFF355709" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF0AFEF55694" box="[199,267,223,248]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A7339FF0AFE725695" box="[286,396,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1c–g</figureCitation>
) very faintly tinted with brown; venation black; Pt trapezoid, upper margin is ½ of lower margin, blackish brown, nervures black, covers almost 3 ½ cells below; a slight bulge in the upper wing margin just proximal to pt on anterior side. Arculus at level of Ax 4–5. Ax 
<quantity id="4C829B900C378E4A7147FEF2FC72572D" box="[864,908,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
right FW, 
<quantity id="4C829B900C378E4A7620FEF2FBCD572D" box="[1031,1075,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="4" pageNumber="575" unit="in" value="8.0">8 in</quantity>
rest. Px 
<quantity id="4C829B900C378E4A76B0FEF3FB05572D" box="[1175,1275,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">34–35 in</quantity>
FW, 
<quantity id="4C829B900C378E4A771EFEF3FA62572D" box="[1337,1436,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">25–26 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FEBAFB1457BD" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FEBAFECF57E5" box="[199,305,367,393]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7366FEBAFE6D57E5" box="[321,403,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1c</figureCitation>
). Pale brown, marked with S1 pale brown, minute small black spots on both ends of lower margin of tergite; S2 with a darker brown lateral patch; S3–6 brownish black lateral stripe, fainter on S4–5, S7–10 black, S6–10 bluish pruinescence, S9–10 almost completely covered in bluish pruinescence in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FE0EFB8A540D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FE0EFECF5799" box="[199,305,475,501]" italics="true" pageId="4" pageNumber="575">Genitalia</emphasis>
(
<figureCitation id="13412AF00C378E4A7318FE0EFE525799" box="[319,428,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Fig. 2a–e</figureCitation>
). First and second segments ventrally and dorsally dark brown. Third segment pale yellow. Apical segment translucent medially raised on dorsal side ending with a hood, lamina curved anteriorly and extended laterally, segmented in two parts on both sides, upper part forming a shoulder and ending in a thin filamentous arm slightly thicker at apex. Lower part canine-shaped, rounded, curved outward &amp; laterally.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FDBEFE84549D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FDBEFE5854E9" box="[199,422,619,645]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A7392FDBEFDD254E9" box="[437,556,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Figs. 2f–h</figureCitation>
). Cerci black, bent inwards at midpoint making ~90° angle with proximal part; cerci apex rounded and expanded; inner margin of proximal part has bulging and numerous small black spines; cerci length more than S10; paraproct dark brown to black, 1⁄2 of the cerci length, broad at base, apex sharply pointed and curved inwards.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FD2EFB7B5579" blockId="4.[151,1437,151,789]" box="[199,1157,762,789]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 35–36; HW 29–30; total length 45–46.</paragraph>
<paragraph id="8BC536750C378E4A72B0FC97FE015531" blockId="4.[151,1436,834,1293]" box="[151,511,834,861]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FC97FE8B5531" bold="true" box="[151,373,834,861]" pageId="4" pageNumber="575">Variation in Males</emphasis>
(
<figureCitation id="13412AF00C378E4A73A3FC96FE115531" box="[388,495,835,861]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 3–4</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C378E4A72B0FCB2FEFB5235" blockId="4.[151,1436,834,1293]" pageId="4" pageNumber="575">
Though in life all males have similar golden yellow lateral pterothorax, post-mortem it changes to dull brown in some individuals (
<figureCitation id="13412AF00C378E4A7340FC5EFE3D55C9" box="[359,451,907,933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Figs. 1c</figureCitation>
, 
<figureCitation id="13412AF00C378E4A73F6FC5EFE0E55C9" box="[465,496,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">3b</figureCitation>
). IBC-BR297, IBC-BR 
<specimenCount id="9D7CFDFC0C378E4A70DDFC5EFC9255C9" box="[762,876,907,933]" count="383" pageId="4" pageNumber="575" type="male">383 males</specimenCount>
have brown upper part of mesepimeron, whereas IBC-BR300 and IBC-BR451 have clear golden yellow thorax without any post-mortem browning (
<figureCitation id="13412AF00C378E4A7738FC7AFA6E55A5" box="[1311,1424,943,969]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 3–4</figureCitation>
). IBC-BR 
<specimenCount id="9D7CFDFC0C378E4A72D2FC06FEA55581" box="[245,347,979,1005]" count="297" pageId="4" pageNumber="575" type="male">297 male</specimenCount>
has 9–10 Ax in FW and HW and 34 Px in FW, 27 Px in HW. IBC-BR 
<specimenCount id="9D7CFDFC0C378E4A7641FC06FB325581" box="[1126,1228,979,1005]" count="300" pageId="4" pageNumber="575" type="male">300 male</specimenCount>
has 7–8 Ax in FW and HW and 31–32 Px in FW and 26–27 Px in HW (
<figureCitation id="13412AF00C378E4A70CCFC22FCCE527D" box="[747,816,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Similarly, there are 37 Px in FW and 28 Px in HW in IBC-BR 
<specimenCount id="9D7CFDFC0C378E4A72D1FBCEFEA25259" box="[246,348,1051,1077]" count="383" pageId="4" pageNumber="575" type="male">383 male</specimenCount>
and 36 Px in FW and 27 Px in HW in IBC-BR 
<specimenCount id="9D7CFDFC0C378E4A7154FBCFFC275259" box="[883,985,1050,1077]" count="451" pageId="4" pageNumber="575" type="male">451 male</specimenCount>
(
<figureCitation id="13412AF00C378E4A71CEFBCEFBD05259" box="[1001,1070,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Both these males have 8 Ax in all wings.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FBB6FE8C5211" blockId="4.[151,1436,834,1293]" box="[199,370,1123,1149]" pageId="4" pageNumber="575">Measurements.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB52FB9752CD" blockId="4.[151,1436,834,1293]" box="[199,1129,1159,1185]" pageId="4" pageNumber="575">IBC-BR297: Abdomen without appendages 36–37; HW 32–33; total length 46–47.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB7EFBC152A9" blockId="4.[151,1436,834,1293]" box="[199,1087,1195,1221]" pageId="4" pageNumber="575">IBC-BR300: Abdomen without appendages 31–32; HW 28; total length 41–42.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB1AFB975285" blockId="4.[151,1436,834,1293]" box="[199,1129,1231,1257]" pageId="4" pageNumber="575">IBC-BR383: Abdomen without appendages 31–32; HW 29–30; total length 43–44.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB27FB975361" blockId="4.[151,1436,834,1293]" box="[199,1129,1266,1293]" pageId="4" pageNumber="575">IBC-BR451: Abdomen without appendages 32–33; HW 28–29; total length 41–42.</paragraph>
<paragraph id="8BC536750C378E4A72B0FAEFFCE35339" blockId="4.[151,1437,1338,1941]" box="[151,797,1338,1365]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FAEFFD625338" bold="true" box="[151,668,1338,1365]" pageId="4" pageNumber="575">
Description of 
<typeStatus id="54C188D70C378E4A7360FAEEFE4C5339" box="[327,434,1339,1365]" pageId="4" pageNumber="575" type="paratype">paratype</typeStatus>
female IBC-BX390
</emphasis>
(
<figureCitation id="13412AF00C378E4A708CFAEEFCEB5339" box="[683,789,1338,1365]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1650,1674]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[151,1436,181,1627]" captionTargetId-0="figure-17@7.[214,1373,181,1919]" captionTargetId-1="figure-18@8.[151,1436,181,1627]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." captionText-1="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13916303" figureDoi-1="http://doi.org/10.5281/zenodo.13916305" httpUri-0="https://zenodo.org/record/13916303/files/figure.png" httpUri-1="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Figs. 5–6</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C378E4A72B0FA8AFAA253AD" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FA8AFF2A5315" box="[151,212,1375,1401]" italics="true" pageId="4" pageNumber="575">Head</emphasis>
(
<figureCitation id="13412AF00C378E4A72C2FA8AFEA85315" box="[229,342,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5a–b</figureCitation>
). Labrum, labium and mandible pale blue in life, brown post-mortem, labrum dark brown post-mortem; anteclypeus, postclypeus and frons black with blue shine. Rest of head black. Ocelli brown, antennae black. Eyes upper half brown, lower half grey in life, darker post-mortem. Postocular area and occiput black.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA1EFBFC5389" blockId="4.[151,1437,1338,1941]" box="[199,1026,1482,1509]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA1EFE665389" box="[199,408,1483,1509]" italics="true" pageId="4" pageNumber="575">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7380FA1EFDEA5389" box="[423,532,1482,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5c–e</figureCitation>
). Similar to male. Slightly pruinosed in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA3AFA98503D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA3AFEB35065" box="[199,333,1519,1545]" italics="true" pageId="4" pageNumber="575">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7347FA3AFE305065" box="[352,462,1518,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5f–g</figureCitation>
). Similar to male, except browner laterally. Mesepisternum black with bronze tinge; brownish yellow stripe on mesepisternum just above the mesopleural suture which is greyish blue in life; upper part of mesepimeron brown; faint brown marking on interpleural suture; rest of the lateral pterothorax pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F98EFD5150F5" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F98EFF025018" box="[199,252,1627,1652]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae brown. Femur, tibiae brown with bluish pruinescence in life, dark brown post-mortem. Tibiofemoral joint, tarsus and spines blackish brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F976FA8950D1" blockId="4.[151,1437,1338,1941]" box="[199,1399,1698,1725]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F976FEF550D0" box="[199,267,1699,1724]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A733CF976FE9350D1" box="[283,365,1699,1725]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6a</figureCitation>
). Hyaline, similar to male; Pt pale brown. Ax 
<quantity id="4C829B900C378E4A7157F976FC6550D1" box="[880,923,1699,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
all wings. Px 
<quantity id="4C829B900C378E4A761DF977FB6350D1" box="[1082,1181,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">34–35 in</quantity>
FW, 
<quantity id="4C829B900C378E4A76FEF977FAC250D1" box="[1241,1340,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">25–26 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F912FE0A5145" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F912FECF508D" box="[199,305,1735,1761]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7364F912FE69508D" box="[323,407,1734,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5c</figureCitation>
). S1–2 dorsally yellowish brown, laterally brown, S3–7 brown with faint, pale yellowish basal rings; dorsum of S6 with a large yellow, rounded, dorsal marking towards posterior margin; S9–10 black; abdomen ventrally pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8E6FD94511D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F8E6FE575121" box="[199,425,1843,1869]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A739DF8E6FDD25121" box="[442,556,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6b–d</figureCitation>
). Cerci short, conical, pointed, black, less than length of S10. Paraproct very small, globular, brown; ovipositor brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8AEFB7B51F9" blockId="4.[151,1437,1338,1941]" box="[199,1157,1915,1941]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 30–31; HW 29–30; total length 39–40.</paragraph>
<caption id="DF0566FD0C368E4B72B0F842FC5451BF" ID-DOI="http://doi.org/10.5281/zenodo.13916299" ID-Zenodo-Dep="13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="5" targetType="figure">
<paragraph id="8BC536750C368E4B72B0F842FC5451BF" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="576">
<emphasis id="B90EEA670C368E4B72B0F842FEEA51C3" bold="true" box="[151,276,1943,1967]" pageId="5" pageNumber="576">FIGURE 3.</emphasis>
<taxonomicName id="4C7A4DF60C368E4B733CF842FE4D51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[283,435,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C368E4B733CF842FE4D51C3" box="[283,435,1943,1967]" italics="true" pageId="5" pageNumber="576">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C368E4B739CF842FDF251C3" bold="true" box="[443,524,1943,1967]" pageId="5" pageNumber="576">
<taxonomicNameLabel id="A23D571C0C368E4B739CF842FDF251C3" box="[443,524,1943,1967]" pageId="5" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<caption id="DF0566FD0C358E4872B0F8ECFD72519D" ID-DOI="http://doi.org/10.5281/zenodo.13916301" ID-Zenodo-Dep="13916301" httpUri="https://zenodo.org/record/13916301/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,1849,1873]" targetBox="[232,1356,151,1837]" targetPageId="6" targetType="figure">
<paragraph id="8BC536750C358E4872B0F8ECFD72519D" blockId="6.[151,1436,1849,2034]" pageId="6" pageNumber="577">
<emphasis id="B90EEA670C358E4872B0F8ECFEED513D" bold="true" box="[151,275,1849,1873]" pageId="6" pageNumber="577">FIGURE 4.</emphasis>
<taxonomicName id="4C7A4DF60C358E48733EF8EFFE4E513E" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1850,1874]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C358E48733EF8EFFE4E513E" box="[281,432,1850,1874]" italics="true" pageId="6" pageNumber="577">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C358E487391F8EFFDF9513E" bold="true" box="[438,519,1850,1874]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A23D571C0C358E487391F8EFFDF9513E" box="[438,519,1850,1874]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
<caption id="DF0566FD0C348E4972B0F842FE2D519B" ID-DOI="http://doi.org/10.5281/zenodo.13916303" ID-Zenodo-Dep="13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="7" pageNumber="578" startId="7.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="7" targetType="figure">
<paragraph id="8BC536750C348E4972B0F842FE2D519B" blockId="7.[151,1437,1943,2039]" pageId="7" pageNumber="578">
<emphasis id="B90EEA670C348E4972B0F842FEEC51C3" bold="true" box="[151,274,1943,1967]" pageId="7" pageNumber="578">FIGURE 5.</emphasis>
<taxonomicName id="4C7A4DF60C348E497330F842FE5351C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[279,429,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C348E497330F842FE5351C3" box="[279,429,1943,1967]" italics="true" pageId="7" pageNumber="578">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C348E497395F842FDFC51C3" bold="true" box="[434,514,1943,1967]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A23D571C0C348E497395F842FDFC51C3" box="[434,514,1943,1967]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c).
</paragraph>
</caption>
<caption id="DF0566FD0C3B8E4672B0F9A7FCCD50C3" ID-DOI="http://doi.org/10.5281/zenodo.13916305" ID-Zenodo-Dep="13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="8" pageNumber="579" startId="8.[151,250,1650,1674]" targetBox="[151,1436,181,1627]" targetPageId="8" targetType="figure">
<paragraph id="8BC536750C3B8E4672B0F9A7FCCD50C3" blockId="8.[151,1437,1650,1711]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F9A7FEED50E7" bold="true" box="[151,275,1650,1675]" pageId="8" pageNumber="579">FIGURE 6.</emphasis>
<taxonomicName id="4C7A4DF60C3B8E46733EF9A6FE4E50E7" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1651,1675]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3B8E46733EF9A6FE4E50E7" box="[281,432,1651,1675]" italics="true" pageId="8" pageNumber="579">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3B8E467391F9A6FDF950E7" bold="true" box="[438,519,1651,1675]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A23D571C0C3B8E467391F9A6FDF950E7" box="[438,519,1651,1675]" pageId="8" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view.
</paragraph>
</caption>
<paragraph id="8BC536750C3B8E4672B0F908FE0D509B" blockId="8.[151,1436,1757,1999]" box="[151,499,1757,1783]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F908FE71509B" bold="true" box="[151,399,1757,1783]" pageId="8" pageNumber="579">Variation in Females</emphasis>
(
<figureCitation id="13412AF00C3B8E4673B9F908FE1C509B" box="[414,482,1757,1783]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C3B8E4672B0F8D4FF4251A3" blockId="8.[151,1436,1757,1999]" pageId="8" pageNumber="579">
Stripe on mesepisternum is usually greyish blue in life, it has variable post-mortem colours (
<figureCitation id="13412AF00C3B8E467690F8D4FB035177" box="[1207,1277,1793,1819]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BX307 and IBC-BX 
<specimenCount id="9D7CFDFC0C3B8E467300F8F0FE4E5153" box="[295,432,1829,1855]" count="309" pageId="8" pageNumber="579" type="female">309 females</specimenCount>
have pale bluish yellow stripe on mesepisternum, whereas IBC-BR314 and IBC-BX313 have brownish yellow stripe on mesepisternum (
<figureCitation id="13412AF00C3B8E467091F89CFD04510F" box="[694,762,1865,1891]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BR 
<specimenCount id="9D7CFDFC0C3B8E46714AF89CFC17510F" box="[877,1001,1865,1891]" count="314" pageId="8" pageNumber="579" type="female">314 female</specimenCount>
has 7–8 Ax in all the wings and 31–33 Px in FW and 23–27 Px in HW. IBC-BX 
<specimenCount id="9D7CFDFC0C3B8E46707AF8B8FD2751EB" box="[605,729,1901,1927]" count="307" pageId="8" pageNumber="579" type="female">307 female</specimenCount>
has clearer wings and 7–8 Ax and 33–36, whereas IBC-BX 
<specimenCount id="9D7CFDFC0C3B8E467754F8B8FF1D51C7" count="309" pageId="8" pageNumber="579" type="female">309 female</specimenCount>
has faintly brown enfumed wings and 7–10 Ax and 34–36 Px. IBC-BX 
<specimenCount id="9D7CFDFC0C3B8E46762AF844FB7451C7" box="[1037,1162,1937,1963]" count="313" pageId="8" pageNumber="579" type="female">313 female</specimenCount>
has 8–10 Ax and 34–37 Px.
</paragraph>
<caption id="DF0566FD0C3A8E4772B0F842FBF551BF" ID-DOI="http://doi.org/10.5281/zenodo.13916307" ID-Zenodo-Dep="13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="9" pageNumber="580" startId="9.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="9" targetType="figure">
<paragraph id="8BC536750C3A8E4772B0F842FBF551BF" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="580">
<emphasis id="B90EEA670C3A8E4772B0F842FEEB51C3" bold="true" box="[151,277,1943,1967]" pageId="9" pageNumber="580">FIGURE 7.</emphasis>
<taxonomicName id="4C7A4DF60C3A8E477339F842FE4951C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[286,439,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3A8E477339F842FE4951C3" box="[286,439,1943,1967]" italics="true" pageId="9" pageNumber="580">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3A8E4773E7F842FDED51C3" bold="true" box="[448,531,1943,1967]" pageId="9" pageNumber="580">
<taxonomicNameLabel id="A23D571C0C3A8E4773E7F842FDED51C3" box="[448,531,1943,1967]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C398E4472E0FF42FE8C56DD" blockId="10.[199,1131,151,321]" box="[199,370,151,177]" pageId="10" pageNumber="581">Measurements.</paragraph>
<paragraph id="8BC536750C398E4472E0FF6EFB9756B9" blockId="10.[199,1131,151,321]" box="[199,1129,186,213]" pageId="10" pageNumber="581">IBC-BR314: Abdomen without appendages 28–29; HW 24–25; total length 34–35.</paragraph>
<paragraph id="8BC536750C398E4472E0FF0AFB955695" blockId="10.[199,1131,151,321]" box="[199,1131,223,249]" pageId="10" pageNumber="581">IBC-BX307: Abdomen without appendages 31–32; HW 31–32; total length 40–41.</paragraph>
<paragraph id="8BC536750C398E4472E0FED6FB955771" blockId="10.[199,1131,151,321]" box="[199,1131,259,285]" pageId="10" pageNumber="581">IBC-BX309: Abdomen without appendages 32–32; HW 30–31; total length 40–41.</paragraph>
<paragraph id="8BC536750C398E4472E0FEF2FB95572D" blockId="10.[199,1131,151,321]" box="[199,1131,295,321]" pageId="10" pageNumber="581">IBC-BX313: Abdomen without appendages 31–32; HW 31–22; total length 41–42.</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C398E4472B0FE47FD4D53AD" pageId="10" pageNumber="581" type="diagnosis">
<paragraph id="8BC536750C398E4472B0FE47FE5D57C0" blockId="10.[151,419,402,428]" box="[151,419,402,428]" pageId="10" pageNumber="581">
<heading id="D08D81190C398E4472B0FE47FE5D57C0" bold="true" box="[151,419,402,428]" fontSize="11" level="1" pageId="10" pageNumber="581" reason="1">
<emphasis id="B90EEA670C398E4472B0FE47FE5D57C0" bold="true" box="[151,419,402,428]" pageId="10" pageNumber="581">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC536750C398E4472B0FE0EFC1055ED" blockId="10.[151,1437,475,1473]" pageId="10" pageNumber="581">
<taxonomicName id="4C7A4DF60C398E4472B0FE0EFEBC5799" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[151,322,475,501]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4472B0FE0EFEBC5799" box="[151,322,475,501]" italics="true" pageId="10" pageNumber="581">Devadatta adii</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C7A4DF60C398E447005FE0EFD2B5799" authorityName="Sasamoto" authorityYear="2003" box="[546,725,475,501]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447005FE0EFD2B5799" box="[546,725,475,501]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
in morphological features and caudal appendages. Following characters distinguish it from 
<taxonomicName id="4C7A4DF60C398E4473C5FE2AFD6F5475" authorityName="Sasamoto" authorityYear="2003" box="[482,657,511,537]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E4473C5FE2AFD6F5475" box="[482,657,511,537]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
. In males, (a) cerci curved at ~90° angle near midpoint in 
<taxonomicName id="4C7A4DF60C398E447733FE2AFA9F5475" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1300,1377,511,537]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447733FE2AFA9F5475" box="[1300,1377,511,537]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
(
<figureCitation id="13412AF00C398E447748FE2AFF4A5451" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8a</figureCitation>
), cerci less curved and at an obtuse angle near midpoint in 
<taxonomicName id="4C7A4DF60C398E447165FDF6FC0F5451" authorityName="Sasamoto" authorityYear="2003" box="[834,1009,547,573]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447165FDF6FC0F5451" box="[834,1009,547,573]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
(
<figureCitation id="13412AF00C398E4471D8FDF6FBAC5451" box="[1023,1106,547,573]" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8h</figureCitation>
); (b) inner border of cerci has numerous small spines at anterior 1/3 
<superScript id="7C0F9B3D0C398E44701CFD93FDB75438" attach="left" box="[571,585,582,596]" fontSize="6" pageId="10" pageNumber="581">rd</superScript>
in 
<taxonomicName id="4C7A4DF60C398E44704AFD92FD42540D" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[621,700,583,609]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E44704AFD92FD42540D" box="[621,700,583,609]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
(
<figureCitation id="13412AF00C398E4470ECFD92FCE2540D" box="[715,796,583,609]" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8a</figureCitation>
), inner border of cerci is smooth in 
<taxonomicName id="4C7A4DF60C398E447688FD92FA9E540D" authorityName="Sasamoto" authorityYear="2003" box="[1199,1376,583,609]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447688FD92FA9E540D" box="[1199,1376,583,609]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
(
<figureCitation id="13412AF00C398E447748FD92FF4854E9" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8h</figureCitation>
); (c) segment 3 of genitalia is broad anteriorly and filaments are relatively longer (~ 
<quantity id="4C829B900C398E44767DFDBFFB4254E9" box="[1114,1212,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="581" unit="mm" value="0.75">0.75mm</quantity>
) in 
<taxonomicName id="4C7A4DF60C398E4476C1FDBEFACD54E9" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1254,1331,619,645]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4476C1FDBEFACD54E9" box="[1254,1331,619,645]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
, whereas anterior part of segment 3 of genitalia is narrow and rectangular shape and filaments are shorter in 
<taxonomicName id="4C7A4DF60C398E4476C2FD5AFA6B54C5" authorityName="Sasamoto" authorityYear="2003" box="[1253,1429,655,681]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E4476C2FD5AFA6B54C5" box="[1253,1429,655,681]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
; (d) paraproct is slightly longer than 1⁄2 the length of cerci length with sharply pointed apex in 
<taxonomicName id="4C7A4DF60C398E4476F4FD66FADD54A1" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1235,1315,691,717]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4476F4FD66FADD54A1" box="[1235,1315,691,717]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
, paraproct slightly shorter than 1⁄2 the length of cerci with round apex in 
<taxonomicName id="4C7A4DF60C398E447172FD02FBFB549D" authorityName="Sasamoto" authorityYear="2003" box="[853,1029,727,753]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447172FD02FBFB549D" box="[853,1029,727,753]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
; (e) pterothorax of 
<taxonomicName id="4C7A4DF60C398E4476C6FD02FAD1549D" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1249,1327,727,753]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4476C6FD02FAD1549D" box="[1249,1327,727,753]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
is yellow laterally without any distinct black marking on second lateral suture (
<figureCitation id="13412AF00C398E447194FD2EFBF85579" box="[947,1030,763,789]" captionStart="FIGURE 9" captionStartId="12.[151,250,1835,1859]" captionTargetBox="[226,1362,123,1827]" captionTargetId="figure-18@12.[226,1362,123,1827]" captionTargetPageId="12" captionText="FIGURE 9. Lateral view of pterothorax of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale.(Illustrations of D.aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are drawn from Dow et al. 2015. Illustrations of D. cyanocephala and D. glaucinotata are drawn from the photos provided by Akihiko Sasamoto. Illustrations of D. ducatrix, D. kompieri and D. yokoii are drawn from Phan et al. 2015. Illustration of D. argyoides is drawn from the image photographed by Dennis Ferrel.)" figureDoi="http://doi.org/10.5281/zenodo.13916311" httpUri="https://zenodo.org/record/13916311/files/figure.png" pageId="10" pageNumber="581">Fig. 9a</figureCitation>
), whereas the lateral pterothorax of 
<taxonomicName id="4C7A4DF60C398E4472B0FCCAFEB65555" authorityName="Sasamoto" authorityYear="2003" box="[151,328,799,825]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E4472B0FCCAFEB65555" box="[151,328,799,825]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
is bluish grey with a black stripe above the second lateral suture (
<figureCitation id="13412AF00C398E447608FCCAFB7D5555" box="[1071,1155,799,825]" captionStart="FIGURE 9" captionStartId="12.[151,250,1835,1859]" captionTargetBox="[226,1362,123,1827]" captionTargetId="figure-18@12.[226,1362,123,1827]" captionTargetPageId="12" captionText="FIGURE 9. Lateral view of pterothorax of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale.(Illustrations of D.aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are drawn from Dow et al. 2015. Illustrations of D. cyanocephala and D. glaucinotata are drawn from the photos provided by Akihiko Sasamoto. Illustrations of D. ducatrix, D. kompieri and D. yokoii are drawn from Phan et al. 2015. Illustration of D. argyoides is drawn from the image photographed by Dennis Ferrel.)" figureDoi="http://doi.org/10.5281/zenodo.13916311" httpUri="https://zenodo.org/record/13916311/files/figure.png" pageId="10" pageNumber="581">Fig. 9h</figureCitation>
); (f) prothorax of 
<taxonomicName id="4C7A4DF60C398E447769FCCAFA635555" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1358,1437,799,825]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447769FCCAFA635555" box="[1358,1437,799,825]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
is brownish black, whereas 
<taxonomicName id="4C7A4DF60C398E4473F2FC96FD765531" authorityName="Sasamoto" authorityYear="2003" box="[469,648,835,861]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E4473F2FC96FD765531" box="[469,648,835,861]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
has creamy white spots on median lobe; (g) wing tips of 
<taxonomicName id="4C7A4DF60C398E447706FC96FA8F5531" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1313,1393,835,861]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447706FC96FA8F5531" box="[1313,1393,835,861]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
are transparent, whereas wing tips of 
<taxonomicName id="4C7A4DF60C398E447036FCB2FD3F55ED" authorityName="Sasamoto" authorityYear="2003" box="[529,705,871,897]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447036FCB2FD3F55ED" box="[529,705,871,897]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
have brown pigmentation.
</paragraph>
<paragraph id="8BC536750C398E4472E0FC5EFB2F5235" blockId="10.[151,1437,475,1473]" pageId="10" pageNumber="581">
<taxonomicName id="4C7A4DF60C398E4472E0FC5EFE9355C9" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[199,365,907,933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4472E0FC5EFE9355C9" box="[199,365,907,933]" italics="true" pageId="10" pageNumber="581">Devadatta adii</emphasis>
</taxonomicName>
male is easily distinguished from males of 
<taxonomicName id="4C7A4DF60C398E447161FC5EFC6255C8" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[838,924,907,932]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C398E447161FC5EFC6255C8" box="[838,924,907,932]" italics="true" pageId="10" pageNumber="581">D. aran</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E44718FFC5EFBC855C9" baseAuthorityName="Selys" baseAuthorityYear="1859" box="[936,1078,907,933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="argyoides">
<emphasis id="B90EEA670C398E44718FFC5EFBC855C9" box="[936,1078,907,933]" italics="true" pageId="10" pageNumber="581">D. argyoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447665FC5EFB1355C9" authorityName="Laidlaw" authorityYear="1934" box="[1090,1261,907,933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C398E447665FC5EFB1355C9" box="[1090,1261,907,933]" italics="true" pageId="10" pageNumber="581">D. basilanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4476DDFC5EFA6B55C9" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1274,1429,907,933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C398E4476DDFC5EFA6B55C9" box="[1274,1429,907,933]" italics="true" pageId="10" pageNumber="581">D. clavicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4472B0FC7AFEAC55A5" authorityName="Hamalainen, Sasamota &amp; Karube" authorityYear="2006" box="[151,338,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="cyanocephala">
<emphasis id="B90EEA670C398E4472B0FC7AFEAC55A5" box="[151,338,943,969]" italics="true" pageId="10" pageNumber="581">D. cyanocephala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447378FC7AFE2255A5" authorityName="Lieftinck" authorityYear="1969" box="[351,476,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C398E447378FC7AFE2255A5" box="[351,476,943,969]" italics="true" pageId="10" pageNumber="581">D. ducatrix</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4473CFFC7AFD9255A5" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[488,620,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C398E4473CFFC7AFD9255A5" box="[488,620,943,969]" italics="true" pageId="10" pageNumber="581">D. kompieri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E44705FFC7AFCD455A5" authorityName="Fraser" authorityYear="1933" box="[632,810,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="multinervosa">
<emphasis id="B90EEA670C398E44705FFC7AFCD455A5" box="[632,810,943,969]" italics="true" pageId="10" pageNumber="581">D. multinervosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447111FC7AFC0E55A5" authorityName="Laidlaw" authorityYear="1934" box="[822,1008,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C398E447111FC7AFC0E55A5" box="[822,1008,943,969]" italics="true" pageId="10" pageNumber="581">D. podolestoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4471DBFC7AFB9B55A5" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1020,1125,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C398E4471DBFC7AFB9B55A5" box="[1020,1125,943,969]" italics="true" pageId="10" pageNumber="581">D. somoh</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447656FC7AFB2355A5" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1137,1245,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C398E447656FC7AFB2355A5" box="[1137,1245,943,969]" italics="true" pageId="10" pageNumber="581">D. tanduk</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7A4DF60C398E447730FC7AFA8555A5" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[1303,1403,943,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C398E447730FC7AFA8555A5" box="[1303,1403,943,969]" italics="true" pageId="10" pageNumber="581">D. yokoii</emphasis>
</taxonomicName>
by its characteristic yellow lateral pterothorax (
<figureCitation id="13412AF00C398E44705CFC06FD495581" box="[635,695,979,1005]" captionStart="FIGURE 9" captionStartId="12.[151,250,1835,1859]" captionTargetBox="[226,1362,123,1827]" captionTargetId="figure-18@12.[226,1362,123,1827]" captionTargetPageId="12" captionText="FIGURE 9. Lateral view of pterothorax of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale.(Illustrations of D.aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are drawn from Dow et al. 2015. Illustrations of D. cyanocephala and D. glaucinotata are drawn from the photos provided by Akihiko Sasamoto. Illustrations of D. ducatrix, D. kompieri and D. yokoii are drawn from Phan et al. 2015. Illustration of D. argyoides is drawn from the image photographed by Dennis Ferrel.)" figureDoi="http://doi.org/10.5281/zenodo.13916311" httpUri="https://zenodo.org/record/13916311/files/figure.png" pageId="10" pageNumber="581">Fig 9</figureCitation>
). All the other species have black pterothorax with intermittent pale creamy yellow markings. Additionally, 
<taxonomicName id="4C7A4DF60C398E447077FC22FD59527C" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[592,679,1015,1040]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C398E447077FC22FD59527C" box="[592,679,1015,1040]" italics="true" pageId="10" pageNumber="581">D. aran</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447093FC22FC9E527D" authorityName="Laidlaw" authorityYear="1934" box="[692,864,1015,1041]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C398E447093FC22FC9E527D" box="[692,864,1015,1041]" italics="true" pageId="10" pageNumber="581">D. basilanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E447149FC22FBF5527D" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[878,1035,1015,1041]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C398E447149FC22FBF5527D" box="[878,1035,1015,1041]" italics="true" pageId="10" pageNumber="581">D. clavicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E44763FFC22FB32527D" authorityName="Fraser" authorityYear="1933" box="[1048,1228,1015,1041]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="multinervosa">
<emphasis id="B90EEA670C398E44763FFC22FB32527D" box="[1048,1228,1015,1041]" italics="true" pageId="10" pageNumber="581">D. multinervosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4476FDFC22FA6B527D" authorityName="Laidlaw" authorityYear="1934" box="[1242,1429,1015,1041]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C398E4476FDFC22FA6B527D" box="[1242,1429,1015,1041]" italics="true" pageId="10" pageNumber="581">D. podolestoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E4472B0FBCEFEFC5259" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[151,258,1051,1077]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C398E4472B0FBCEFEFC5259" box="[151,258,1051,1077]" italics="true" pageId="10" pageNumber="581">D. somoh</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7A4DF60C398E447318FBCEFE525259" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[319,428,1051,1077]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C398E447318FBCEFE525259" box="[319,428,1051,1077]" italics="true" pageId="10" pageNumber="581">D. tanduk</emphasis>
</taxonomicName>
have a very short paraproct which is barely visible in dorsal view (
<figureCitation id="13412AF00C398E4476B0FBCEFB255259" box="[1175,1243,1051,1077]" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8</figureCitation>
). Wing tips of all 
<taxonomicName id="4C7A4DF60C398E4472B0FBEAFEBD5235" box="[151,323,1087,1113]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEA670C398E4472B0FBEAFEF25235" box="[151,268,1087,1113]" italics="true" pageId="10" pageNumber="581">Devadatta</emphasis>
spp.
</taxonomicName>
, except 
<taxonomicName id="4C7A4DF60C398E4473B8FBEAFDAD5235" authorityName="Fraser" authorityYear="1933" box="[415,595,1087,1113]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="multinervosa">
<emphasis id="B90EEA670C398E4473B8FBEAFDAD5235" box="[415,595,1087,1113]" italics="true" pageId="10" pageNumber="581">D. multinervosa</emphasis>
</taxonomicName>
are variably pigmented. This feature is absent in 
<taxonomicName id="4C7A4DF60C398E44765BFBEAFB355235" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1148,1227,1087,1113]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E44765BFBEAFB355235" box="[1148,1227,1087,1113]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC536750C398E4472E0FBB6FB4052A9" blockId="10.[151,1437,475,1473]" pageId="10" pageNumber="581">
Additionally, 
<taxonomicName id="4C7A4DF60C398E447346FBB6FE515211" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[353,431,1123,1149]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447346FBB6FE515211" box="[353,431,1123,1149]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E44739BFBB6FDC55211" authorityName="Lieftinck" authorityYear="1969" box="[444,571,1123,1149]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C398E44739BFBB6FDC55211" box="[444,571,1123,1149]" italics="true" pageId="10" pageNumber="581">D. ducatrix</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C398E44706EFBB6FD315211" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[585,719,1123,1149]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C398E44706EFBB6FD315211" box="[585,719,1123,1149]" italics="true" pageId="10" pageNumber="581">D. kompieri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C398E447123FBB6FC945211" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[772,874,1123,1149]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C398E447123FBB6FC945211" box="[772,874,1123,1149]" italics="true" pageId="10" pageNumber="581">D. yokoii</emphasis>
</taxonomicName>
have similar caudal appendages and genital ligula (
<figureCitation id="13412AF00C398E4472B8FB52FF1852CD" box="[159,230,1159,1185]" captionStart="FIGURE 8" captionStartId="11.[151,250,1806,1830]" captionTargetBox="[232,1356,113,1799]" captionTargetId="figure-17@11.[232,1356,113,1799]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal view of caudal appendages of Devadatta spp.: (a) Devadatta adii sp. nov.; (b) D. aran; (c) D. argyoides; (d) D. basilanensis; (e) D. clavicauda; (f) D. cyanocephala; (g) D. ducatrix; (h) D. glaucinotata; (i) D. kompieri; (j) D. podolestoides; (k) D. somoh; (l) D. tanduk; (m) D. yokoii. Not to scale. (Caudal appendages of D. aran, D. basilanensis, D. clavicauda, D. podolestoides, D. somoh and D. tanduk are redrawn from Dow et al. 2015. Caudal appendages of D. argyoides and D. glaucinotata are redrawn from Kitagawa &amp; Katatani 2013. Caudal appendages of D. cyanocephala are redrawn from Hämäläinen et al. 2006. Caudal appendages of D. ducatrix are redrawn from Lieftinck 1969. Caudal appendages of D. kompieri and D. yokoii are redrawn from Phan et al. 2015.)" figureDoi="http://doi.org/10.5281/zenodo.13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="10" pageNumber="581">Fig. 8</figureCitation>
). However, the spinous inner margin of cerci, broader anterior part of segment 3 and longer filaments of penile organ and transparent wing tips help to distinguish the new species from the latter three.
</paragraph>
<paragraph id="8BC536750C398E4472E0FB1AFAFC5339" blockId="10.[151,1437,475,1473]" pageId="10" pageNumber="581">
<taxonomicName id="4C7A4DF60C398E4472E0FB1AFE265285" authorityName="Fraser" authorityYear="1933" box="[199,472,1231,1257]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="multinervosa">
<emphasis id="B90EEA670C398E4472E0FB1AFE265285" box="[199,472,1231,1257]" italics="true" pageId="10" pageNumber="581">Devadatta multinervosa</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="54C188D70C398E4470D8FB1AFC9F5285" box="[767,865,1231,1257]" pageId="10" pageNumber="581" type="holotype">holotype</typeStatus>
male which is currently deposited in the Natural History Museum (NHM), London, 
<collectingCountry id="F36D76E50C398E447001FB26FDB05361" box="[550,590,1267,1293]" name="United Kingdom" pageId="10" pageNumber="581">UK</collectingCountry>
(
<figureCitation id="13412AF00C398E447078FB26FD4D5361" box="[607,691,1267,1293]" captionStart="FIGURE 10" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@13.[214,1373,181,1919]" captionTargetPageId="13" captionText="FIGURE 10. Devadatta multinervosa, holotype male (NHMUK 013383752): (a) dorsal habitus; (b) lateral view of pterothorax; (c) ventral view of segment 2; (d–e) caudal appendages: (d) dorsal view; (e) lateral view.All scale bars 1 mm. (©Natural History Museum 2024)." figureDoi="http://doi.org/10.5281/zenodo.13916313" httpUri="https://zenodo.org/record/13916313/files/figure.png" pageId="10" pageNumber="581">Fig. 10</figureCitation>
). Although the cerci of this specimen are damaged and unsuitable for comparison (
<bibRefCitation id="EFEB4B840C398E447372FAC2FD47535D" author="Natural History Museum" box="[341,697,1303,1329]" pageId="10" pageNumber="581" refId="ref6203" refString="Natural History Museum (2024) Data Portal query on 1 resources created at 12 July 2024. 10: 32: 18.657615 PID. Available from: https: // data. nhm. ac. uk (asccessed 17 September 2024) https: // doi. org / 10.5519 / qd. 6 ny 304 oz" type="url" year="2024">Natural History Museum 2024</bibRefCitation>
), its larger size, black pterothorax, shorter paraproct, and higher nodal index (Px 
<quantity id="4C829B900C398E447377FAEEFE4D5339" box="[336,435,1339,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.1811" metricValueMax="1.2446" metricValueMin="1.1176" pageId="10" pageNumber="581" unit="in" value="46.5" valueMax="49.0" valueMin="44.0">44–49 in</quantity>
FW, 
<quantity id="4C829B900C398E4473C8FAEEFDAC5339" box="[495,594,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" metricValueMax="9.906" metricValueMin="9.398" pageId="10" pageNumber="581" unit="in" value="38.0" valueMax="39.0" valueMin="37.0">37–39 in</quantity>
HW, Ax 
<quantity id="4C829B900C398E44709DFAEEFD0D5339" box="[698,755,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="10" pageNumber="581" unit="in" value="13.0">13 in</quantity>
FW, 
<quantity id="4C829B900C398E447108FAEEFC6F5339" box="[815,913,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="10" pageNumber="581" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
HW) distinguish it from 
<taxonomicName id="4C7A4DF60C398E44768AFAEEFB055339" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1197,1275,1339,1365]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E44768AFAEEFB055339" box="[1197,1275,1339,1365]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC536750C398E4472E0FA8AFD4D53AD" blockId="10.[151,1437,475,1473]" pageId="10" pageNumber="581">
<taxonomicName id="4C7A4DF60C398E4472E0FA8AFE905315" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[199,366,1375,1401]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4472E0FA8AFE905315" box="[199,366,1375,1401]" italics="true" pageId="10" pageNumber="581">Devadatta adii</emphasis>
</taxonomicName>
female is readily distinguished from all described females of 
<taxonomicName id="4C7A4DF60C398E447633FA8AFB435315" box="[1044,1213,1375,1401]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEA670C398E447633FA8AFB775315" box="[1044,1161,1375,1401]" italics="true" pageId="10" pageNumber="581">Devadatta</emphasis>
spp.
</taxonomicName>
by its yellow lateral pterothorax. Moreover, the female of 
<taxonomicName id="4C7A4DF60C398E447066FA56FD0C53F1" authorityName="Sasamoto" authorityYear="2003" box="[577,754,1411,1437]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C398E447066FA56FD0C53F1" box="[577,754,1411,1437]" italics="true" pageId="10" pageNumber="581">D. glaucinotata</emphasis>
</taxonomicName>
can be distinguished from that of 
<taxonomicName id="4C7A4DF60C398E447650FA56FB3853F1" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1143,1222,1411,1437]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447650FA56FB3853F1" box="[1143,1222,1411,1437]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
by the dark brown lateral pterothorax (
<bibRefCitation id="EFEB4B840C398E447353FA72FD5653AD" author="Kitagawa, K. &amp; Katatani, N." box="[372,680,1447,1473]" pageId="10" pageNumber="581" pagination="47 - 56" refId="ref6050" refString="Kitagawa, K. &amp; Katatani, N. (2013) Notes on the odonate fauna of Southeast Asia. Part 3. Amphipterygidae (Genus Devadatta). Aeschna, 49, 47 - 56. [in Japanese, with English title / summary]" type="journal article" year="2013">Kitagawa &amp; Katatani 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C398E4472B0F9C7FB6A511D" pageId="10" pageNumber="581" type="biology_ecology">
<paragraph id="8BC536750C398E4472B0F9C7FE7C5040" blockId="10.[151,386,1554,1581]" box="[151,386,1554,1581]" pageId="10" pageNumber="581">
<heading id="D08D81190C398E4472B0F9C7FE7C5040" bold="true" box="[151,386,1554,1581]" fontSize="11" level="1" pageId="10" pageNumber="581" reason="1">
<emphasis id="B90EEA670C398E4472B0F9C7FE7C5040" bold="true" box="[151,386,1554,1581]" pageId="10" pageNumber="581">Habitat and habits</emphasis>
</heading>
</paragraph>
<paragraph id="8BC536750C398E4472B0F98EFB6A511D" blockId="10.[151,1436,1627,1905]" pageId="10" pageNumber="581">
<taxonomicName id="4C7A4DF60C398E4472B0F98EFEC05019" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[151,318,1627,1653]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E4472B0F98EFEC05019" box="[151,318,1627,1653]" italics="true" pageId="10" pageNumber="581">Devadatta adii</emphasis>
</taxonomicName>
was observed in forested streams and along a waterfall in the Ramsing, Sirring Nala and Yinkiong to Likor Road areas of Upper Siang District. The waterfall was surrounded by tall grasses, where males of 
<taxonomicName id="4C7A4DF60C398E447734F9AAFA9E50F5" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[1299,1376,1663,1689]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E447734F9AAFA9E50F5" box="[1299,1376,1663,1689]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
were observed perching at heights ranging from 
<quantity id="4C829B900C398E447056F976FD1250D1" box="[625,748,1699,1725]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" metricValueMax="3.048" metricValueMin="0.6096" pageId="10" pageNumber="581" unit="ft" value="6.0" valueMax="10.0" valueMin="2.0">2 to 10 feet</quantity>
above the ground (
<figureCitation id="13412AF00C398E4471E7F976FBE050D1" box="[960,1054,1699,1725]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-18@14.[151,1435,181,1894]" captionTargetPageId="14" captionText="FIGURE 11. Devadatta adii sp. nov. in life: (a) paratype male (IBC-BR300); (b) female (not collected)." figureDoi="http://doi.org/10.5281/zenodo.13916315" httpUri="https://zenodo.org/record/13916315/files/figure.png" pageId="10" pageNumber="581">Fig. 11a</figureCitation>
). Females were observed perching on twigs and fern leaves at a different location (
<figureCitation id="13412AF00C398E447097F912FCEF508D" box="[688,785,1735,1761]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-18@14.[151,1435,181,1894]" captionTargetPageId="14" captionText="FIGURE 11. Devadatta adii sp. nov. in life: (a) paratype male (IBC-BR300); (b) female (not collected)." figureDoi="http://doi.org/10.5281/zenodo.13916315" httpUri="https://zenodo.org/record/13916315/files/figure.png" pageId="10" pageNumber="581">Fig. 11b</figureCitation>
). The new species exhibits flight characteristics similar to other members of the 
<taxonomicName id="4C7A4DF60C398E4473AFF93EFDE25169" authorityName="Dijkstra, Kalkman, Dow, Stokvis &amp; van Tol" authorityYear="2014" box="[392,540,1771,1797]" class="Insecta" family="Devadattidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="family">Devadattidae</taxonomicName>
family. Individuals were active in the morning and afternoon, and were observed in both shaded and sunny areas. The specific location of 
<taxonomicName id="4C7A4DF60C398E44710FF8DAFC875145" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[808,889,1807,1833]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C398E44710FF8DAFC875145" box="[808,889,1807,1833]" italics="true" pageId="10" pageNumber="581">D. adii</emphasis>
</taxonomicName>
is accessed via a rugged road bordered by cliff forests on one side and a deep valley on the other (
<figureCitation id="13412AF00C398E4470E1F8E6FCE75121" box="[710,793,1843,1869]" captionStart="FIGURE 12" captionStartId="15.[151,250,1918,1942]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-17@15.[151,1435,181,1894]" captionTargetPageId="15" captionText="FIGURE 12. Type locality of Devadatta adii sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.13916317" httpUri="https://zenodo.org/record/13916317/files/figure.png" pageId="10" pageNumber="581">Fig. 12</figureCitation>
). Other species like 
<taxonomicName id="4C7A4DF60C398E4471DDF8E6FA975121" authority="(Selys, 1863)" baseAuthorityName="Selys" baseAuthorityYear="1863" box="[1018,1385,1843,1869]" class="Insecta" family="Platycnemididae" genus="Calicnemia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="B90EEA670C398E4471DDF8E6FB325120" box="[1018,1228,1843,1869]" italics="true" pageId="10" pageNumber="581">Calicnemia eximia</emphasis>
(Selys, 1863)
</taxonomicName>
, and 
<taxonomicName id="4C7A4DF60C398E4472B0F882FD6E511D" authority="Gurung &amp; Phan, 2023" authorityName="Gurung &amp; Phan" authorityYear="2023" box="[151,656,1879,1905]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="581" phylum="Arthropoda" rank="species" species="samtsensis">
<emphasis id="B90EEA670C398E4472B0F882FE6C511C" box="[151,402,1879,1904]" italics="true" pageId="10" pageNumber="581">Protosticta samtsensis</emphasis>
<bibRefCitation id="EFEB4B840C398E4473BFF882FD6E511D" author="Gurung, M. M. &amp; Phan, Q. T." box="[408,656,1879,1905]" pageId="10" pageNumber="581" pagination="137 - 144" refId="ref5617" refString="Gurung, M. M. &amp; Phan, Q. T. (2023) Description of Protosticta samtsensis sp. nov. (Odonata: Zygoptera: Platystictidae) from Samtse district, Bhutan. Zootaxa, 5352 (1), 137 - 144. https: // doi. org / 10.11646 / zootaxa. 5352.1.5" type="journal article" year="2023">Gurung &amp; Phan, 2023</bibRefCitation>
</taxonomicName>
were commonly observed to co-occur with it.
</paragraph>
</subSubSection>
<caption id="DF0566FD0C388E4572B0F8DBFD70519B" ID-DOI="http://doi.org/10.5281/zenodo.13916309" ID-Zenodo-Dep="13916309" httpUri="https://zenodo.org/record/13916309/files/figure.png" pageId="11" pageNumber="582" startId="11.[151,250,1806,1830]" targetBox="[232,1356,113,1799]" targetPageId="11" targetType="figure">
<paragraph id="8BC536750C388E4572B0F8DBFD70519B" blockId="11.[151,1437,1806,2039]" pageId="11" pageNumber="582">
<emphasis id="B90EEA670C388E4572B0F8DBFEEA514B" bold="true" box="[151,276,1806,1831]" pageId="11" pageNumber="582">FIGURE 8.</emphasis>
Dorsal view of caudal appendages of 
<taxonomicName id="4C7A4DF60C388E4570B8F8DAFCC3514A" box="[671,829,1807,1831]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEA670C388E4570B8F8DAFCF7514B" box="[671,777,1807,1831]" italics="true" pageId="11" pageNumber="582">Devadatta</emphasis>
spp.
</taxonomicName>
: (a) 
<taxonomicName id="4C7A4DF60C388E45714CF8DAFBFD514B" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[875,1027,1807,1831]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C388E45714CF8DAFBFD514B" box="[875,1027,1807,1831]" italics="true" pageId="11" pageNumber="582">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C388E45762CF8DAFBA3514B" bold="true" box="[1035,1117,1807,1831]" pageId="11" pageNumber="582">
<taxonomicNameLabel id="A23D571C0C388E45762CF8DAFBA3514B" box="[1035,1117,1807,1831]" pageId="11" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (b) 
<taxonomicName id="4C7A4DF60C388E4576B7F8DAFB1E514A" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1168,1248,1807,1830]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C388E4576B7F8DAFB1E514A" box="[1168,1248,1807,1830]" italics="true" pageId="11" pageNumber="582">D. aran</emphasis>
</taxonomicName>
; (c) 
<taxonomicName id="4C7A4DF60C388E457735F8DAFA6B514B" baseAuthorityName="Selys" baseAuthorityYear="1859" box="[1298,1429,1807,1831]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="argyoides">
<emphasis id="B90EEA670C388E457735F8DAFA6B514B" box="[1298,1429,1807,1831]" italics="true" pageId="11" pageNumber="582">D. argyoides</emphasis>
</taxonomicName>
; (d) 
<taxonomicName id="4C7A4DF60C388E457298F8E7FE9E5125" authorityName="Laidlaw" authorityYear="1934" box="[191,352,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C388E457298F8E7FE9E5125" box="[191,352,1841,1865]" italics="true" pageId="11" pageNumber="582">D. basilanensis</emphasis>
</taxonomicName>
; (e) 
<taxonomicName id="4C7A4DF60C388E4573BFF8E7FDD45125" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[408,554,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C388E4573BFF8E7FDD45125" box="[408,554,1841,1865]" italics="true" pageId="11" pageNumber="582">D. clavicauda</emphasis>
</taxonomicName>
; (f) 
<taxonomicName id="4C7A4DF60C388E457047F8E7FCEE5125" authorityName="Hamalainen, Sasamota &amp; Karube" authorityYear="2006" box="[608,784,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="cyanocephala">
<emphasis id="B90EEA670C388E457047F8E7FCEE5125" box="[608,784,1841,1865]" italics="true" pageId="11" pageNumber="582">D. cyanocephala</emphasis>
</taxonomicName>
; (g) 
<taxonomicName id="4C7A4DF60C388E45716DF8E7FC3F5125" authorityName="Lieftinck" authorityYear="1969" box="[842,961,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C388E45716DF8E7FC3F5125" box="[842,961,1841,1865]" italics="true" pageId="11" pageNumber="582">D. ducatrix</emphasis>
</taxonomicName>
; (h) 
<taxonomicName id="4C7A4DF60C388E4571DCF8E7FB605125" authorityName="Sasamoto" authorityYear="2003" box="[1019,1182,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C388E4571DCF8E7FB605125" box="[1019,1182,1841,1865]" italics="true" pageId="11" pageNumber="582">D. glaucinotata</emphasis>
</taxonomicName>
; (i) 
<taxonomicName id="4C7A4DF60C388E4576F5F8E7FAB15125" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[1234,1359,1841,1865]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C388E4576F5F8E7FAB15125" box="[1234,1359,1841,1865]" italics="true" pageId="11" pageNumber="582">D. kompieri</emphasis>
</taxonomicName>
; (j) 
<taxonomicName id="4C7A4DF60C388E4577A3F8E7FEDC5100" authorityName="Laidlaw" authorityYear="1934" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C388E4577A3F8E7FEDC5100" italics="true" pageId="11" pageNumber="582">D. podolestoides</emphasis>
</taxonomicName>
; (k) 
<taxonomicName id="4C7A4DF60C388E457370F880FE445100" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[343,442,1876,1900]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C388E457370F880FE445100" box="[343,442,1876,1900]" italics="true" pageId="11" pageNumber="582">D. somoh</emphasis>
</taxonomicName>
; (l) 
<taxonomicName id="4C7A4DF60C388E4573CDF880FDAE5100" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[490,592,1876,1900]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C388E4573CDF880FDAE5100" box="[490,592,1876,1900]" italics="true" pageId="11" pageNumber="582">D. tanduk</emphasis>
</taxonomicName>
; (m) 
<taxonomicName id="4C7A4DF60C388E4570ACF880FD145100" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[651,746,1876,1900]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C388E4570ACF880FD145100" box="[651,746,1876,1900]" italics="true" pageId="11" pageNumber="582">D. yokoii</emphasis>
</taxonomicName>
. Not to scale. (Caudal appendages of 
<taxonomicName id="4C7A4DF60C388E45765FF880FB375100" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1144,1225,1877,1900]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C388E45765FF880FB375100" box="[1144,1225,1877,1900]" italics="true" pageId="11" pageNumber="582">D. aran</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C388E4576F0F880FA8B5100" authorityName="Laidlaw" authorityYear="1934" box="[1239,1397,1876,1900]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C388E4576F0F880FA8B5100" box="[1239,1397,1876,1900]" italics="true" pageId="11" pageNumber="582">D. basilanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C388E4577A3F880FEF851E3" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C388E4577A3F880FEF851E3" italics="true" pageId="11" pageNumber="582">D. clavicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C388E457334F8A2FE4051E3" authorityName="Laidlaw" authorityYear="1934" box="[275,446,1911,1935]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C388E457334F8A2FE4051E3" box="[275,446,1911,1935]" italics="true" pageId="11" pageNumber="582">D. podolestoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C388E4573ECF8A2FDD351E3" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[459,557,1911,1935]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C388E4573ECF8A2FDD351E3" box="[459,557,1911,1935]" italics="true" pageId="11" pageNumber="582">D. somoh</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C388E457078F8A2FD3D51E3" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[607,707,1911,1935]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C388E457078F8A2FD3D51E3" box="[607,707,1911,1935]" italics="true" pageId="11" pageNumber="582">D. tanduk</emphasis>
</taxonomicName>
are redrawn from 
<bibRefCitation id="EFEB4B840C388E4571A6F8A2FBD751E3" author="Dow, R. A. &amp; Hamalainen, M. &amp; Stokvis, F. R." box="[897,1065,1911,1935]" pageId="11" pageNumber="582" pagination="301 - 349" refId="ref5460" refString="Dow, R. A., Hamalainen, M. &amp; Stokvis, F. R. (2015) Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae). Zootaxa, 4033 (3), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4033.3.1" type="journal article" year="2015">
Dow 
<emphasis id="B90EEA670C388E45719EF8A2FC1851E3" box="[953,998,1911,1935]" italics="true" pageId="11" pageNumber="582">et al</emphasis>
. 2015
</bibRefCitation>
. Caudal appendages of 
<taxonomicName id="4C7A4DF60C388E45773DF8A2FA6251E3" baseAuthorityName="Selys" baseAuthorityYear="1859" box="[1306,1436,1911,1935]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="argyoides">
<emphasis id="B90EEA670C388E45773DF8A2FA6251E3" box="[1306,1436,1911,1935]" italics="true" pageId="11" pageNumber="582">D. argyoides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C388E4572E3F84FFE9A51DD" authorityName="Sasamoto" authorityYear="2003" box="[196,356,1945,1969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C388E4572E3F84FFE9A51DD" box="[196,356,1945,1969]" italics="true" pageId="11" pageNumber="582">D. glaucinotata</emphasis>
</taxonomicName>
are redrawn from 
<bibRefCitation id="EFEB4B840C388E457000F84CFCC151DE" author="Kitagawa, K. &amp; Katatani, N." box="[551,831,1945,1970]" pageId="11" pageNumber="582" pagination="47 - 56" refId="ref6050" refString="Kitagawa, K. &amp; Katatani, N. (2013) Notes on the odonate fauna of Southeast Asia. Part 3. Amphipterygidae (Genus Devadatta). Aeschna, 49, 47 - 56. [in Japanese, with English title / summary]" type="journal article" year="2013">Kitagawa &amp; Katatani 2013</bibRefCitation>
. Caudal appendages of 
<taxonomicName id="4C7A4DF60C388E457613F84FFB1C51DD" authorityName="Hamalainen, Sasamota &amp; Karube" authorityYear="2006" box="[1076,1250,1945,1969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="cyanocephala">
<emphasis id="B90EEA670C388E457613F84FFB1C51DD" box="[1076,1250,1945,1969]" italics="true" pageId="11" pageNumber="582">D. cyanocephala</emphasis>
</taxonomicName>
are redrawn from 
<bibRefCitation id="EFEB4B840C388E4572B0F869FE7B51B8" author="Hamalainen, M. &amp; Sasamoto, A. &amp; Karube, H." box="[151,389,1980,2004]" pageId="11" pageNumber="582" pagination="1 - 6" refId="ref5677" refString="Hamalainen, M., Sasamoto, A. &amp; Karube, H. (2006) Description of Devadatta cyanocephala sp. nov. from Vietnam (Zygoptera: Amphipterygidae). Tombo - Acta Odonatologica Japonica, 48, 1 - 6." type="journal article" year="2006">
Hämäläinen 
<emphasis id="B90EEA670C388E457330F868FEBD51B8" box="[279,323,1980,2004]" italics="true" pageId="11" pageNumber="582">et al</emphasis>
. 2006
</bibRefCitation>
. Caudal appendages of 
<taxonomicName id="4C7A4DF60C388E457055F868FD1A51B8" authorityName="Lieftinck" authorityYear="1969" box="[626,740,1980,2004]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C388E457055F868FD1A51B8" box="[626,740,1980,2004]" italics="true" pageId="11" pageNumber="582">D. ducatrix</emphasis>
</taxonomicName>
are redrawn from 
<bibRefCitation id="EFEB4B840C388E4571B9F869FBC951B8" author="Lieftinck, M. A." box="[926,1079,1980,2004]" pageId="11" pageNumber="582" pagination="205 - 215" refId="ref6153" refString="Lieftinck, M. A. (1969) Two new Odonata from Southeast Asia, with comments on previously described species. Deutsche Entomologische Zeitschrift, N. F., 16, 205 - 215. https: // doi. org / 10.1002 / mmnd. 4810160118" type="journal article" year="1969">Lieftinck 1969</bibRefCitation>
. Caudal appendages of 
<taxonomicName id="4C7A4DF60C388E457703F868FA6251B8" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[1316,1436,1980,2004]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C388E457703F868FA6251B8" box="[1316,1436,1980,2004]" italics="true" pageId="11" pageNumber="582">D. kompieri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C388E4572E5F80AFEE1519B" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[194,287,2015,2039]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C388E4572E5F80AFEE1519B" box="[194,287,2015,2039]" italics="true" pageId="11" pageNumber="582">D. yokoii</emphasis>
</taxonomicName>
are redrawn from 
<bibRefCitation id="EFEB4B840C388E4573FCF80AFD7A519B" author="Phan, Q. T. &amp; Sasamoto, A. &amp; Hayashi, F." box="[475,644,2015,2039]" pageId="11" pageNumber="582" pagination="414 - 420" refId="ref6310" refString="Phan, Q. T., Sasamoto, A. &amp; Hayashi, F. (2015) Description of two new species of the genus Devadatta from northern Vietnam and central Laos (Odonata: Devadattidae). Zootaxa, 3941 (3), 414 - 420. https: // doi. org / 10.11646 / zootaxa. 3941.3.7" type="journal article" year="2015">
Phan 
<emphasis id="B90EEA670C388E457033F80AFDBF519B" box="[532,577,2015,2039]" italics="true" pageId="11" pageNumber="582">et al</emphasis>
. 2015
</bibRefCitation>
.)
</paragraph>
</caption>
<caption id="DF0566FD0C3F8E4272B0F8FEFBC6519B" ID-DOI="http://doi.org/10.5281/zenodo.13916311" ID-Zenodo-Dep="13916311" httpUri="https://zenodo.org/record/13916311/files/figure.png" pageId="12" pageNumber="583" startId="12.[151,250,1835,1859]" targetBox="[226,1362,123,1827]" targetPageId="12" targetType="figure">
<paragraph id="8BC536750C3F8E4272B0F8FEFBC6519B" blockId="12.[151,1437,1835,2039]" pageId="12" pageNumber="583">
<emphasis id="B90EEA670C3F8E4272B0F8FEFEEA512F" bold="true" box="[151,276,1835,1859]" pageId="12" pageNumber="583">FIGURE 9.</emphasis>
Lateral view of pterothorax of 
<taxonomicName id="4C7A4DF60C3F8E42707EF8FEFD09512F" box="[601,759,1835,1859]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEA670C3F8E42707EF8FEFD3D512F" box="[601,707,1835,1859]" italics="true" pageId="12" pageNumber="583">Devadatta</emphasis>
spp.
</taxonomicName>
: (a) 
<taxonomicName id="4C7A4DF60C3F8E427101F8FEFC43512F" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[806,957,1835,1859]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3F8E427101F8FEFC43512F" box="[806,957,1835,1859]" italics="true" pageId="12" pageNumber="583">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3F8E4271E2F8F9FBE95128" bold="true" box="[965,1047,1836,1860]" pageId="12" pageNumber="583">
<taxonomicNameLabel id="A23D571C0C3F8E4271E2F8F9FBE95128" box="[965,1047,1836,1860]" pageId="12" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (b) 
<taxonomicName id="4C7A4DF60C3F8E42766CF8F9FB65512F" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1099,1179,1836,1859]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C3F8E42766CF8F9FB65512F" box="[1099,1179,1836,1859]" italics="true" pageId="12" pageNumber="583">D. aran</emphasis>
</taxonomicName>
; (c) 
<taxonomicName id="4C7A4DF60C3F8E4276EAF8F9FAAE512F" baseAuthorityName="Selys" baseAuthorityYear="1859" box="[1229,1360,1835,1859]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="argyoides">
<emphasis id="B90EEA670C3F8E4276EAF8F9FAAE512F" box="[1229,1360,1835,1859]" italics="true" pageId="12" pageNumber="583">D. argyoides</emphasis>
</taxonomicName>
; (d) 
<taxonomicName id="4C7A4DF60C3F8E4277A3F8F9FEEA510B" authorityName="Laidlaw" authorityYear="1934" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C3F8E4277A3F8F9FEEA510B" italics="true" pageId="12" pageNumber="583">D. basilanensis</emphasis>
</taxonomicName>
; (e) 
<taxonomicName id="4C7A4DF60C3F8E427363F885FE2F510B" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[324,465,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C3F8E427363F885FE2F510B" box="[324,465,1871,1895]" italics="true" pageId="12" pageNumber="583">D. clavicauda</emphasis>
</taxonomicName>
; (f) 
<taxonomicName id="4C7A4DF60C3F8E4273D9F885FD54510B" authorityName="Hamalainen, Sasamota &amp; Karube" authorityYear="2006" box="[510,682,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="cyanocephala">
<emphasis id="B90EEA670C3F8E4273D9F885FD54510B" box="[510,682,1871,1895]" italics="true" pageId="12" pageNumber="583">D. cyanocephala</emphasis>
</taxonomicName>
; (g) 
<taxonomicName id="4C7A4DF60C3F8E4270FDF885FCB3510B" authorityName="Lieftinck" authorityYear="1969" box="[730,845,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C3F8E4270FDF885FCB3510B" box="[730,845,1871,1895]" italics="true" pageId="12" pageNumber="583">D. ducatrix</emphasis>
</taxonomicName>
; (h) 
<taxonomicName id="4C7A4DF60C3F8E427159F885FBE2510B" authorityName="Sasamoto" authorityYear="2003" box="[894,1052,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C3F8E427159F885FBE2510B" box="[894,1052,1871,1895]" italics="true" pageId="12" pageNumber="583">D. glaucinotata</emphasis>
</taxonomicName>
; (i) 
<taxonomicName id="4C7A4DF60C3F8E427660F885FB3E510B" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[1095,1216,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C3F8E427660F885FB3E510B" box="[1095,1216,1871,1895]" italics="true" pageId="12" pageNumber="583">D. kompieri</emphasis>
</taxonomicName>
; (j) 
<taxonomicName id="4C7A4DF60C3F8E4276CCF885FA6B510B" authorityName="Laidlaw" authorityYear="1934" box="[1259,1429,1871,1895]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C3F8E4276CCF885FA6B510B" box="[1259,1429,1871,1895]" italics="true" pageId="12" pageNumber="583">D. podolestoides</emphasis>
</taxonomicName>
; (k) 
<taxonomicName id="4C7A4DF60C3F8E42729EF8A1FEE651E7" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[185,280,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C3F8E42729EF8A1FEE651E7" box="[185,280,1907,1931]" italics="true" pageId="12" pageNumber="583">D. somoh</emphasis>
</taxonomicName>
; (l) 
<taxonomicName id="4C7A4DF60C3F8E427367F8A1FE5C51E7" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[320,418,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C3F8E427367F8A1FE5C51E7" box="[320,418,1907,1931]" italics="true" pageId="12" pageNumber="583">D. tanduk</emphasis>
</taxonomicName>
; (m) 
<taxonomicName id="4C7A4DF60C3F8E4273F1F8A1FDCF51E7" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[470,561,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C3F8E4273F1F8A1FDCF51E7" box="[470,561,1907,1931]" italics="true" pageId="12" pageNumber="583">D. yokoii</emphasis>
</taxonomicName>
. Not to scale. (Illustrations of 
<taxonomicName id="4C7A4DF60C3F8E42717DF8A1FC5951E7" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[858,935,1908,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="B90EEA670C3F8E42717DF8A1FC5951E7" box="[858,935,1908,1931]" italics="true" pageId="12" pageNumber="583">D. aran</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C3F8E427195F8A1FBB251E7" authorityName="Laidlaw" authorityYear="1934" box="[946,1100,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="B90EEA670C3F8E427195F8A1FBB251E7" box="[946,1100,1907,1931]" italics="true" pageId="12" pageNumber="583">D. basilanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C3F8E427670F8A1FB1D51E7" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[1111,1251,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="B90EEA670C3F8E427670F8A1FB1D51E7" box="[1111,1251,1907,1931]" italics="true" pageId="12" pageNumber="583">D. clavicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C3F8E4276C9F8A1FA6851E7" authorityName="Laidlaw" authorityYear="1934" box="[1262,1430,1907,1931]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="B90EEA670C3F8E4276C9F8A1FA6851E7" box="[1262,1430,1907,1931]" italics="true" pageId="12" pageNumber="583">D. podolestoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C3F8E4272B0F84DFF0651C3" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[151,248,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="B90EEA670C3F8E4272B0F84DFF0651C3" box="[151,248,1943,1967]" italics="true" pageId="12" pageNumber="583">D. somoh</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C3F8E42730EF84DFE7351C3" authorityName="Dow, Hamalainen &amp; Stokvis" authorityYear="2015" box="[297,397,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="B90EEA670C3F8E42730EF84DFE7351C3" box="[297,397,1943,1967]" italics="true" pageId="12" pageNumber="583">D. tanduk</emphasis>
</taxonomicName>
are drawn from 
<bibRefCitation id="EFEB4B840C3F8E427012F84DFD2551C3" author="Dow, R. A. &amp; Hamalainen, M. &amp; Stokvis, F. R." box="[565,731,1943,1967]" pageId="12" pageNumber="583" pagination="301 - 349" refId="ref5460" refString="Dow, R. A., Hamalainen, M. &amp; Stokvis, F. R. (2015) Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae). Zootaxa, 4033 (3), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4033.3.1" type="journal article" year="2015">
Dow 
<emphasis id="B90EEA670C3F8E42704AF84DFD6751C3" box="[621,665,1943,1967]" italics="true" pageId="12" pageNumber="583">et al</emphasis>
. 2015
</bibRefCitation>
. Illustrations of 
<taxonomicName id="4C7A4DF60C3F8E427158F84DFBD551C3" authorityName="Hamalainen, Sasamota &amp; Karube" authorityYear="2006" box="[895,1067,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="cyanocephala">
<emphasis id="B90EEA670C3F8E427158F84DFBD551C3" box="[895,1067,1943,1967]" italics="true" pageId="12" pageNumber="583">D. cyanocephala</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C3F8E42767CF84DFB0451C3" authorityName="Sasamoto" authorityYear="2003" box="[1115,1274,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="glaucinotata">
<emphasis id="B90EEA670C3F8E42767CF84DFB0451C3" box="[1115,1274,1943,1967]" italics="true" pageId="12" pageNumber="583">D. glaucinotata</emphasis>
</taxonomicName>
are drawn from the photos provided by Akihiko Sasamoto. Illustrations of 
<taxonomicName id="4C7A4DF60C3F8E4270D5F869FC9851BF" authorityName="Lieftinck" authorityYear="1969" box="[754,870,1979,2003]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="ducatrix">
<emphasis id="B90EEA670C3F8E4270D5F869FC9851BF" box="[754,870,1979,2003]" italics="true" pageId="12" pageNumber="583">D. ducatrix</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7A4DF60C3F8E427152F869FC1151BF" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[885,1007,1979,2003]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B90EEA670C3F8E427152F869FC1151BF" box="[885,1007,1979,2003]" italics="true" pageId="12" pageNumber="583">D. kompieri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7A4DF60C3F8E427605F869FB7F51BF" authorityName="Phan, Sasamoto &amp; Hayashi" authorityYear="2015" box="[1058,1153,1979,2003]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="yokoii">
<emphasis id="B90EEA670C3F8E427605F869FB7F51BF" box="[1058,1153,1979,2003]" italics="true" pageId="12" pageNumber="583">D. yokoii</emphasis>
</taxonomicName>
are drawn from 
<bibRefCitation id="EFEB4B840C3F8E427709F86EFF33519B" author="Phan, Q. T. &amp; Sasamoto, A. &amp; Hayashi, F." pageId="12" pageNumber="583" pagination="414 - 420" refId="ref6310" refString="Phan, Q. T., Sasamoto, A. &amp; Hayashi, F. (2015) Description of two new species of the genus Devadatta from northern Vietnam and central Laos (Odonata: Devadattidae). Zootaxa, 3941 (3), 414 - 420. https: // doi. org / 10.11646 / zootaxa. 3941.3.7" type="journal article" year="2015">
Phan 
<emphasis id="B90EEA670C3F8E42774FF869FA6851BF" box="[1384,1430,1979,2003]" italics="true" pageId="12" pageNumber="583">et al</emphasis>
. 2015
</bibRefCitation>
. Illustration of 
<taxonomicName id="4C7A4DF60C3F8E427340F835FE17519B" baseAuthorityName="Selys" baseAuthorityYear="1859" box="[359,489,2015,2039]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="argyoides">
<emphasis id="B90EEA670C3F8E427340F835FE17519B" box="[359,489,2015,2039]" italics="true" pageId="12" pageNumber="583">D. argyoides</emphasis>
</taxonomicName>
is drawn from the image photographed by Dennis Ferrel.)
</paragraph>
</caption>
<caption id="DF0566FD0C3E8E4372B0F842FEC6519B" ID-DOI="http://doi.org/10.5281/zenodo.13916313" ID-Zenodo-Dep="13916313" httpUri="https://zenodo.org/record/13916313/files/figure.png" pageId="13" pageNumber="584" startId="13.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="13" targetType="figure">
<paragraph id="8BC536750C3E8E4372B0F842FEC6519B" blockId="13.[151,1437,1943,2039]" pageId="13" pageNumber="584">
<emphasis id="B90EEA670C3E8E4372B0F842FEE151C3" bold="true" box="[151,287,1943,1967]" pageId="13" pageNumber="584">FIGURE 10.</emphasis>
<taxonomicName id="4C7A4DF60C3E8E437303F842FDE951C3" authorityName="Fraser" authorityYear="1933" box="[292,535,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="multinervosa">
<emphasis id="B90EEA670C3E8E437303F842FDE951C3" box="[292,535,1943,1967]" italics="true" pageId="13" pageNumber="584">Devadatta multinervosa</emphasis>
</taxonomicName>
, holotype male (NHMUK 013383752): (a) dorsal habitus; (b) lateral view of pterothorax; (c) ventral view of segment 2; (d–e) caudal appendages: (d) dorsal view; (e) lateral view. All scale bars 1 mm. (© 
<bibRefCitation id="EFEB4B840C3E8E4376DAF868FED3519B" author="Natural History Museum" pageId="13" pageNumber="584" refId="ref6203" refString="Natural History Museum (2024) Data Portal query on 1 resources created at 12 July 2024. 10: 32: 18.657615 PID. Available from: https: // data. nhm. ac. uk (asccessed 17 September 2024) https: // doi. org / 10.5519 / qd. 6 ny 304 oz" type="url" year="2024">Natural History Museum 2024</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF0566FD0C3D8E4072B0F8ABFB4351FA" ID-DOI="http://doi.org/10.5281/zenodo.13916315" ID-Zenodo-Dep="13916315" box="[151,1213,1918,1942]" httpUri="https://zenodo.org/record/13916315/files/figure.png" pageId="14" pageNumber="585" startId="14.[151,250,1918,1942]" targetBox="[151,1435,181,1894]" targetPageId="14" targetType="figure">
<paragraph id="8BC536750C3D8E4072B0F8ABFB4351FA" blockId="14.[151,1213,1918,1942]" box="[151,1213,1918,1942]" pageId="14" pageNumber="585">
<emphasis id="B90EEA670C3D8E4072B0F8ABFEE051FA" bold="true" box="[151,286,1918,1942]" pageId="14" pageNumber="585">FIGURE 11.</emphasis>
<taxonomicName id="4C7A4DF60C3D8E407302F8ABFE4251FA" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[293,444,1918,1942]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3D8E407302F8ABFE4251FA" box="[293,444,1918,1942]" italics="true" pageId="14" pageNumber="585">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3D8E4073E5F8ABFDED51FA" bold="true" box="[450,531,1918,1942]" pageId="14" pageNumber="585">
<taxonomicNameLabel id="A23D571C0C3D8E4073E5F8ABFDED51FA" box="[450,531,1918,1942]" pageId="14" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life: (a) paratype male (IBC-BR300); (b) female (not collected).
</paragraph>
</caption>
<caption id="DF0566FD0C3C8E4172B0F8ABFD3F51F9" ID-DOI="http://doi.org/10.5281/zenodo.13916317" ID-Zenodo-Dep="13916317" box="[151,705,1918,1942]" httpUri="https://zenodo.org/record/13916317/files/figure.png" pageId="15" pageNumber="586" startId="15.[151,250,1918,1942]" targetBox="[151,1435,181,1894]" targetPageId="15" targetType="figure">
<paragraph id="8BC536750C3C8E4172B0F8ABFD3F51F9" blockId="15.[151,705,1918,1942]" box="[151,705,1918,1942]" pageId="15" pageNumber="586">
<emphasis id="B90EEA670C3C8E4172B0F8ABFEDE51FA" bold="true" box="[151,288,1918,1942]" pageId="15" pageNumber="586">FIGURE 12.</emphasis>
Type locality of 
<taxonomicName id="4C7A4DF60C3C8E4173EBF8ABFD9D51FA" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[460,611,1918,1942]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3C8E4173EBF8ABFD9D51FA" box="[460,611,1918,1942]" italics="true" pageId="15" pageNumber="586">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3C8E41704DF8ABFD4451FA" bold="true" box="[618,698,1918,1942]" pageId="15" pageNumber="586">
<taxonomicNameLabel id="A23D571C0C3C8E41704DF8ABFD4451FA" box="[618,698,1918,1942]" pageId="15" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C36065FE0C238E5E72B0FF42FBE057E6" pageId="16" pageNumber="587" type="distribution">
<paragraph id="8BC536750C238E5E72B0FF42FE6056DD" blockId="16.[151,414,151,177]" box="[151,414,151,177]" pageId="16" pageNumber="587">
<emphasis id="B90EEA670C238E5E72B0FF42FECE56DD" bold="true" box="[151,304,151,177]" pageId="16" pageNumber="587">Distribution</emphasis>
(
<figureCitation id="13412AF00C238E5E7367FF42FE6B56DD" box="[320,405,151,177]" captionStart="FIGURE 13" captionStartId="16.[151,250,1341,1365]" captionTargetBox="[152,1436,417,1315]" captionTargetId="figure-105@16.[151,1436,417,1317]" captionTargetPageId="16" captionText="FIGURE 13. Distribution map of Devadatta spp. in South and South East Asia [Range data polygons (.shp) are downloaded from International Union for Conservation of Nature (IUCN) Red List of Threatened Species (IUCN 2024) Version 2023-1; https://www.iucnredlist.org]." figureDoi="http://doi.org/10.5281/zenodo.13916319" httpUri="https://zenodo.org/record/13916319/files/figure.png" pageId="16" pageNumber="587">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C238E5E72B0FF0AFBE057E6" blockId="16.[151,1436,223,394]" pageId="16" pageNumber="587">
Currently, 
<taxonomicName id="4C7A4DF60C238E5E7334FF35FE9B5695" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[275,357,223,249]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C238E5E7334FF35FE9B5695" box="[275,357,223,249]" italics="true" pageId="16" pageNumber="587">D. adii</emphasis>
</taxonomicName>
has been found only in Ramsing, Sirring Nala and Yinkiong to Likor Road areas of Upper Siang District. However, the species may also inhabit other parts of the district. 
<figureCitation id="13412AF00C238E5E7603FED6FB6F5771" box="[1060,1169,259,285]" captionStart="FIGURE 13" captionStartId="16.[151,250,1341,1365]" captionTargetBox="[152,1436,417,1315]" captionTargetId="figure-105@16.[151,1436,417,1317]" captionTargetPageId="16" captionText="FIGURE 13. Distribution map of Devadatta spp. in South and South East Asia [Range data polygons (.shp) are downloaded from International Union for Conservation of Nature (IUCN) Red List of Threatened Species (IUCN 2024) Version 2023-1; https://www.iucnredlist.org]." figureDoi="http://doi.org/10.5281/zenodo.13916319" httpUri="https://zenodo.org/record/13916319/files/figure.png" pageId="16" pageNumber="587">Figure 13</figureCitation>
depicts the distribution of all 
<taxonomicName id="4C7A4DF60C238E5E72FCFEF2FE79572E" box="[219,391,295,322]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEA670C238E5E72FCFEF2FEAE572D" box="[219,336,295,321]" italics="true" pageId="16" pageNumber="587">Devadatta</emphasis>
spp.
</taxonomicName>
based on distribution data from International Union for Conservation of Nature Red List of Threatened Species (
<bibRefCitation id="EFEB4B840C238E5E73A4FE9EFDF05709" author="IUCN" box="[387,526,331,357]" pageId="16" pageNumber="587" refId="ref5722" refString="IUCN (2024) The IUCN Red List of Threatened Species. Version 2023 - 1. Avaliable from: https: // www. iucnredlist. org (accessed 12 April 2024)" type="url" year="2024">IUCN 2024</bibRefCitation>
). 
<taxonomicName id="4C7A4DF60C238E5E7005FE9EFD355709" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[546,715,331,357]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C238E5E7005FE9EFD355709" box="[546,715,331,357]" italics="true" pageId="16" pageNumber="587">Devadatta adii</emphasis>
</taxonomicName>
is the northernmost and westernmost distributed species in this genus; all other species are distributed across South 
<collectingCountry id="F36D76E50C238E5E70C7FEA5FCB757E5" box="[736,841,367,394]" name="China" pageId="16" pageNumber="587">East Asia</collectingCountry>
(
<bibRefCitation id="EFEB4B840C238E5E717FFEBAFBED57E6" author="Dow, R. A. &amp; Hamalainen, M. &amp; Stokvis, F. R." box="[856,1043,367,394]" pageId="16" pageNumber="587" pagination="301 - 349" refId="ref5460" refString="Dow, R. A., Hamalainen, M. &amp; Stokvis, F. R. (2015) Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae). Zootaxa, 4033 (3), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4033.3.1" type="journal article" year="2015">
Dow 
<emphasis id="B90EEA670C238E5E71B0FEA5FC3157E5" box="[919,975,367,393]" italics="true" pageId="16" pageNumber="587">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>