Add updates up until 2024-10-11 19:33:09

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Body slender, pubescent. Vertex inclined. Cheeks narrow. Ocelli arranged linearly. Antennal scape as wide as frontal rostrum. Pronotum with anterior field narrower than posterior field. Anterior tibiae only equipped with internal tympanum. Forelegs and midlegs almost equal in length. Tegmen ornamented with dark patches, patchy near the basal field, striped near the apical field. Forewings extending beyond the abdominal tip. Hindwings longer than the forewings.
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<emphasis id="B976EAA6FFA8790CBCF17C0EFAF361E3" bold="true" box="[151,937,366,393]" pageId="1" pageNumber="586">
Key to the species of
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<taxonomicName id="4C024D37FFA8790CBDFB7C0EFBEE61E3" ID-CoL="8NX99" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[413,692,366,393]" class="Insecta" family="Gryllidae" genus="Pentacentrus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAA6FFA8790CBDFB7C0EFB6761E2" bold="true" box="[413,573,367,393]" italics="true" pageId="1" pageNumber="586">Pentacentrus</emphasis>
Saussure
</taxonomicName>
@ -75,7 +76,7 @@ known from
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</paragraph>
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<subSubSection id="C318653FFFA8790CBCF17CD5F84064D2" pageId="1" pageNumber="586" type="key">
<keyStep id="30F62E11FFA8790CBCF17CD5FCC6618D" pageId="1" pageNumber="586">
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<keyLead id="30F39581FFA8790CBCF17CD5FCC661A2" box="[151,1436,436,458]" pageId="1" pageNumber="586">
@ -236,6 +237,74 @@ Li, Xu, &amp; Liu, 2019
</keyLead>
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</keyStep>
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<emphasis id="B976EAA6FFA8790CBCF179FEF84064D2" bold="true" box="[151,282,1183,1209]" pageId="1" pageNumber="586">Taxonomy</emphasis>
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<heading id="D0F581D8FFA8790CBCF17987FB09656B" bold="true" box="[151,595,1254,1280]" fontSize="11" level="1" pageId="1" pageNumber="586" reason="1">
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Family
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<heading id="D0F581D8FFA8790CBCF1784FFBC76523" bold="true" box="[151,669,1326,1352]" fontSize="11" level="1" pageId="1" pageNumber="586" reason="1">
<emphasis id="B976EAA6FFA8790CBCF1784FFBC76523" bold="true" box="[151,669,1326,1352]" pageId="1" pageNumber="586">
Subfamily
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Genus
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<emphasis id="B976EAA6FFA8790CBC887816F8D465FA" bold="true" box="[238,398,1399,1425]" italics="true" pageId="1" pageNumber="586">Pentacentrus</emphasis>
Saussure, 1878
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<taxonomicName id="4C024D37FFA8790CBCF178D6F8AA65A5" ID-CoL="8NX99" authority="Saussure, 1878: 539" authorityName="Saussure" authorityPageNumber="539" authorityYear="1878" box="[151,496,1462,1486]" class="Insecta" family="Gryllidae" genus="Pentacentrus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAA6FFA8790CBCF178D6F84765A5" box="[151,285,1463,1486]" italics="true" pageId="1" pageNumber="586">Pentacentrus</emphasis>
Saussure, 1878: 539
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<emphasis id="B976EAA6FFA8790CBCF178B5F8D96587" bold="true" box="[151,387,1492,1517]" pageId="1" pageNumber="586">
Chinese name:
<emphasis id="B976EAA6FFA8790CBD5178B4F8D96587" bold="true" box="[311,387,1493,1516]" italics="true" pageId="1" pageNumber="586">K̊ĸ</emphasis>
</emphasis>
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<emphasis id="B976EAA6FFA8790CBCF17892F87C6660" bold="true" box="[151,294,1523,1547]" pageId="1" pageNumber="586">
<typeStatus id="54B98816FFA8790CBCF17892F9966660" box="[151,204,1523,1547]" pageId="1" pageNumber="586">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C024D37FFA8790CBD4A7895FB9A6660" ID-CoL="76MPN" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[300,704,1523,1547]" class="Insecta" family="Gryllidae" genus="Pentacentrus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B976EAA6FFA8790CBD4A7895FB7A6660" box="[300,544,1523,1547]" italics="true" pageId="1" pageNumber="586">Pentacentrus pulchellus</emphasis>
Saussure, 1878
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<emphasis id="B976EAA6FFA8790CBCF17B58F8486638" bold="true" box="[151,274,1593,1619]" pageId="1" pageNumber="586">Diagnosis.</emphasis>
Body slender, pubescent. Vertex inclined. Cheeks narrow. Ocelli arranged linearly. Antennal scape as wide as frontal rostrum. Pronotum with anterior field narrower than posterior field. Anterior tibiae only equipped with internal tympanum. Forelegs and midlegs almost equal in length. Tegmen ornamented with dark patches, patchy near the basal field, striped near the apical field. Forewings extending beyond the abdominal tip. Hindwings longer than the forewings.
</paragraph>
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</treatment>
</document>

View file

@ -1,555 +0,0 @@
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<mods:title id="5001B8E800A4A76D87330BA9A2C5996A">Devadatta adii sp. nov. (Odonata: Zygoptera: Devadattidae) from Upper Siang District, Arunachal Pradesh, India</mods:title>
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<mods:namePart id="0B8A962CD233240E3DAC24D9A8C768FC">Kunte, Krushnamegh</mods:namePart>
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<mods:affiliation id="24FE75887D1F2F02220AF9F0A0DABAEB">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, Karnataka, India 560065</mods:affiliation>
<mods:nameIdentifier id="FBF56008BCFABB962F1496DAE9B9E959" type="email">krushnamegh@ncbs.res.in</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo id="07125EE80776CB973D693A305C947CAD">
<mods:title id="DB0C7A78F16D71F7D93D317CF066344C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="17333E76C3F24988CDD110CF131F61FB">
<mods:date id="358B9A00AF35320C469A0EC307BC5239">2024</mods:date>
<mods:detail id="C4CD99C36940A25568267B8785A5CF2C" type="pubDate">
<mods:number id="F49E20D23F26FA21E3DAEC26215D14BB">2024-10-10</mods:number>
</mods:detail>
<mods:detail id="F5C00CF35A452D2F86F12165E02FC881" type="volume">
<mods:number id="084432F905820995DEEB24604FEB0E39">5519</mods:number>
</mods:detail>
<mods:detail id="743BF4D38AC3D2A053F97C2DCC157C93" type="issue">
<mods:number id="1372F04E550A08C5E828B12EB1F98999">4</mods:number>
</mods:detail>
<mods:extent id="85286ACC5FE5FE7D20488BE225960117" unit="page">
<mods:start id="9ED5AB5CA13048F679D59E5AB1603A9E">571</mods:start>
<mods:end id="2F7769F4064A330E3760B09A7B2B101F">589</mods:end>
</mods:extent>
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<mods:location id="B4EDD7D1D24017F53B42999B00DC093D">
<mods:url id="35588A22A696F2671E8DE7A94FA4CAC3">http://zoobank.org/f46dc8a0-ebf2-431a-b486-75595cea886a</mods:url>
</mods:location>
<mods:classification id="0897BD89449B7C6CB0907C4F70E49CE7">journal article</mods:classification>
<mods:identifier id="004AFE27218DF2F7915DE9205804C648" type="DOI">10.11646/zootaxa.5519.4.6</mods:identifier>
<mods:identifier id="06F33D283BE8EFEBA81FEBFCF4CD8CC8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="011EC2123CA6D449CC3EE2A12E24E8B8" type="Zenodo-Dep">13916293</mods:identifier>
<mods:identifier id="75B33348D352EBA216966EB4712B59F5" type="ZooBank">F46DC8A0-EBF2-431A-B486-75595CEA886A</mods:identifier>
</mods:mods>
<treatment id="03D387630C328E4472B0FC7BFB95572D" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D" httpUri="http://treatment.plazi.org/id/03D387630C328E4472B0FC7BFB95572D" lastPageId="10" lastPageNumber="581" pageId="1" pageNumber="572">
<subSubSection id="C36065FE0C328E4F72B0FC7BFD8E55A5" box="[151,624,942,969]" pageId="1" pageNumber="572" type="nomenclature">
<paragraph id="8BC536750C328E4F72B0FC7BFD8E55A5" blockId="1.[151,624,942,1005]" box="[151,624,942,969]" pageId="1" pageNumber="572">
<heading id="D08D81190C328E4F72B0FC7BFD8E55A5" bold="true" box="[151,624,942,969]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
<taxonomicName id="4C7A4DF60C328E4F72B0FC7BFD8E55A5" authority="Joshi, Sawant &amp; Kunte" authorityName="Joshi, Sawant &amp; Kunte" box="[151,624,942,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="adii">
<emphasis id="B90EEA670C328E4F72B0FC7BFD8E55A5" bold="true" box="[151,624,942,969]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0FC7BFEB955A5" bold="true" box="[151,327,942,969]" italics="true" pageId="1" pageNumber="572">Devadatta adii</emphasis>
Joshi, Sawant &amp; Kunte
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4F72B0FC06FEEC5581" box="[151,274,979,1005]" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC536750C328E4F72B0FC06FEEC5581" blockId="1.[151,624,942,1005]" box="[151,274,979,1005]" pageId="1" pageNumber="572">
(
<figureCitation id="13412AF00C328E4F72B8FC06FEF75581" box="[159,265,979,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1849,1873]" captionStartId-4="7.[151,250,1943,1967]" captionStartId-5="8.[151,250,1650,1674]" captionStartId-6="9.[151,250,1943,1967]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[214,1373,181,1919]" captionTargetBox-2="[214,1373,181,1919]" captionTargetBox-3="[232,1356,151,1837]" captionTargetBox-4="[214,1373,181,1919]" captionTargetBox-5="[151,1436,181,1627]" captionTargetBox-6="[214,1373,181,1919]" captionTargetId-0="figure-18@2.[214,1373,181,1919]" captionTargetId-1="figure-17@3.[214,1373,181,1919]" captionTargetId-2="figure-17@5.[214,1373,181,1919]" captionTargetId-3="figure-18@6.[232,1356,151,1837]" captionTargetId-4="figure-17@7.[214,1373,181,1919]" captionTargetId-5="figure-18@8.[151,1436,181,1627]" captionTargetId-6="figure-17@9.[214,1373,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." captionText-1="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (ce) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (fh) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." captionText-2="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-3="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (ef) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (kl) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (qr) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." captionText-4="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." captionText-5="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (bd) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." captionText-6="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916295" figureDoi-1="http://doi.org/10.5281/zenodo.13916297" figureDoi-2="http://doi.org/10.5281/zenodo.13916299" figureDoi-3="http://doi.org/10.5281/zenodo.13916301" figureDoi-4="http://doi.org/10.5281/zenodo.13916303" figureDoi-5="http://doi.org/10.5281/zenodo.13916305" figureDoi-6="http://doi.org/10.5281/zenodo.13916307" httpUri-0="https://zenodo.org/record/13916295/files/figure.png" httpUri-1="https://zenodo.org/record/13916297/files/figure.png" httpUri-2="https://zenodo.org/record/13916299/files/figure.png" httpUri-3="https://zenodo.org/record/13916301/files/figure.png" httpUri-4="https://zenodo.org/record/13916303/files/figure.png" httpUri-5="https://zenodo.org/record/13916305/files/figure.png" httpUri-6="https://zenodo.org/record/13916307/files/figure.png" pageId="1" pageNumber="572">Figs. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4F72B0FBCEFA9C5339" pageId="1" pageNumber="572" type="materials_examined">
<paragraph id="8BC536750C328E4F72B0FBCEFBDF5235" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<materialsCitation id="3B123C280C328E4F72B0FBCEFBDF5235" collectingDate="2022-05-23" collectorName="Fahim Khan &amp; Krushnamefh Kunte &amp; Viraj Nawge" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="B90EEA670C328E4F72B0FBCEFEC65254" bold="true" box="[151,312,1051,1080]" pageId="1" pageNumber="572">
<typeStatus id="54C188D70C328E4F72B0FBCEFEF85259" box="[151,262,1051,1077]" pageId="1" pageNumber="572" type="holotype">Holotype</typeStatus>
.
<specimenCount id="9D7CFDFC0C328E4F7332FBC9FEC65254" box="[277,312,1052,1080]" count="1" pageId="1" pageNumber="572" type="male">1♂</specimenCount>
</emphasis>
(IBC-BR299),
<location id="8EA560AE0C328E4F73CAFBCEFDAD5259" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D:8EA560AE0C328E4F73CAFBCEFDAD5259" box="[493,595,1051,1077]" country="India" latitude="28.6337" municipality="Upper Siang District" name="Ramsing" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Ramsing</location>
,
<collectingMunicipality id="6BA1AC0F0C328E4F7045FBCEFCAC5259" box="[610,850,1051,1077]" pageId="1" pageNumber="572">Upper Siang District</collectingMunicipality>
,
<collectingRegion id="49BEF8970C328E4F7178FBCEFBC45259" box="[863,1082,1051,1077]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F7660FBCEFB7F5259" box="[1095,1153,1051,1077]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F76B5FBCEFAEE5259" box="[1170,1296,1051,1077]" degrees="28.6337" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.6337">28.6337 N</geoCoordinate>
, 94.957 0 E;
<elevation id="0057D1460C328E4F72B0FBEAFED65235" box="[151,296,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="572" unit="m" value="1090.0">
<quantity id="4C829B900C328E4F72B0FBEAFF125235" box="[151,236,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="572" unit="m" value="1090.0">1090 m</quantity>
a.s.l.
</elevation>
),
<date id="FFC410B50C328E4F7319FBEAFE515235" box="[318,431,1087,1113]" pageId="1" pageNumber="572" value="2022-05-23">
<collectingDate id="EF80E95D0C328E4F7319FBEAFE515235" box="[318,431,1087,1113]" pageId="1" pageNumber="572" value="2022-05-23">23.v.2022</collectingDate>
</date>
,
<collectorName id="268F53A30C328E4F739CFBEAFDB65235" box="[443,584,1087,1113]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
,
<collectorName id="268F53A30C328E4F7073FBEAFCC45235" box="[596,826,1087,1113]" pageId="1" pageNumber="572">Krushnamefh Kunte</collectorName>
, &amp;
<collectorName id="268F53A30C328E4F7145FBEAFC0E5235" box="[866,1008,1087,1113]" pageId="1" pageNumber="572">Viraj Nawge</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BC536750C328E4F72E0FBB6FA9C5339" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FBB6FE7352EC" bold="true" box="[199,397,1122,1152]" pageId="1" pageNumber="572">
<typeStatus id="54C188D70C328E4F72E0FBB6FEBF5211" box="[199,321,1123,1149]" pageId="1" pageNumber="572" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D7CFDFC0C328E4F736AFBB7FE7352EC" box="[333,397,1122,1152]" count="2" pageId="1" pageNumber="572" type="male">2 ♂♂</specimenCount>
</emphasis>
(IBC-BR297, IBC-BR300), location, date of collection and collector same as
<typeStatus id="54C188D70C328E4F7727FBB6FA985211" box="[1280,1382,1123,1149]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
;
<materialsCitation id="3B123C280C328E4F7755FBB7FC7852CD" collectingDate="2022-05-22" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F7755FBB7FA6252EC" bold="true" box="[1394,1436,1122,1152]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F7755FBB7FA6252EC" box="[1394,1436,1122,1152]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(IBC-BR383) same location and collectors as
<typeStatus id="54C188D70C328E4F7084FB52FCF652CD" box="[675,776,1159,1185]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
,
<date id="FFC410B50C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">
<collectingDate id="EF80E95D0C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">22.v.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B123C280C328E4F71B4FB53FE0E52A9" collectingDate="2022-05-29" collectorName="Shantanu Joshi" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F71B4FB53FC4052C8" bold="true" box="[915,958,1158,1188]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F71B4FB53FC4052C8" box="[915,958,1158,1188]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(IBC-BR451), same location as holotype,
<date id="FFC410B50C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">
<collectingDate id="EF80E95D0C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">29.v.2022</collectingDate>
</date>
<collectorName id="268F53A30C328E4F7328FB7EFE4552A9" box="[271,443,1195,1221]" pageId="1" pageNumber="572">Shantanu Joshi</collectorName>
leg.
</materialsCitation>
;
<emphasis id="B90EEA670C328E4F73D8FB7FFDD552A4" bold="true" box="[511,555,1194,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F73D8FB7FFDD552A4" box="[511,555,1194,1224]" count="1" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
</emphasis>
(IBC-BR314) all details are the same as
<typeStatus id="54C188D70C328E4F762AFB7EFB8D52A9" box="[1037,1139,1195,1221]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
.
<materialsCitation id="3B123C280C328E4F76A7FB79FA925361" collectingDate="2024-05-23" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" elevation="460" latitude="28.6642" location="Sirring Nala" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCount="3" specimenCount-female="3" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F76A7FB79FB4952A4" bold="true" box="[1152,1207,1196,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F76A7FB79FB4952A4" box="[1152,1207,1196,1224]" count="3" pageId="1" pageNumber="572" type="female">3♀♀</specimenCount>
</emphasis>
(IBC-BX307, IBC-BX309, IBC-BX313)
<location id="8EA560AE0C328E4F73B2FB1AFDDA5285" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D:8EA560AE0C328E4F73B2FB1AFDDA5285" box="[405,548,1231,1257]" country="India" latitude="28.6642" municipality="Upper Siang District" name="Sirring Nala" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Sirring Nala</location>
,
<collectingMunicipality id="6BA1AC0F0C328E4F7016FB1AFCDF5285" box="[561,801,1231,1257]" pageId="1" pageNumber="572">Upper Siang District</collectingMunicipality>
,
<collectingRegion id="49BEF8970C328E4F7109FB1AFBF65285" box="[814,1032,1231,1257]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F7632FB1AFBB15285" box="[1045,1103,1231,1257]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F7647FB1AFB235285" box="[1120,1245,1231,1257]" degrees="28.6642" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.6642">28.6642 N</geoCoordinate>
, 94.994 3 E;
<elevation id="0057D1460C328E4F7755FB1AFF145361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">
<quantity id="4C829B900C328E4F7755FB1AFF535361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">460 m</quantity>
a.s.l.
</elevation>
),
<date id="FFC410B50C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">
<collectingDate id="EF80E95D0C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">23.v.2024</collectingDate>
</date>
,
<collectorName id="268F53A30C328E4F7358FB26FD925361" box="[383,620,1267,1293]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
,
<collectorName id="268F53A30C328E4F705EFB26FCF95361" box="[633,775,1267,1293]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
,
<collectorName id="268F53A30C328E4F7133FB26FC1F5361" box="[788,993,1267,1293]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
,
<collectorName id="268F53A30C328E4F71CBFB26FB4B5361" box="[1004,1205,1267,1293]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
,
<collectorName id="268F53A30C328E4F76E6FB26FAC95361" box="[1217,1335,1267,1293]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B123C280C328E4F775EFB21FA9C5339" collectingDate="2024-06-02" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" county="Upper Siang District" elevation="360" latitude="28.7639" location="Yinkiong" municipality="Likor Road" pageId="1" pageNumber="572" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F775EFB21FA62537C" bold="true" box="[1401,1436,1268,1296]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F775EFB21FA62537C" box="[1401,1436,1268,1296]" count="1" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
</emphasis>
(IBC-BX390)
<location id="8EA560AE0C328E4F731FFAC2FE5E535D" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D:8EA560AE0C328E4F731FFAC2FE5E535D" box="[312,416,1303,1329]" country="India" county="Upper Siang District" latitude="28.7639" municipality="Likor Road" name="Yinkiong" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Yinkiong</location>
to
<collectingMunicipality id="6BA1AC0F0C328E4F73E4FAC2FDB8535D" box="[451,582,1303,1329]" pageId="1" pageNumber="572">Likor Road</collectingMunicipality>
,
<collectingCounty id="62A44EF90C328E4F7076FAC2FCC4535D" box="[593,826,1303,1329]" pageId="1" pageNumber="572">Upper Siang District</collectingCounty>
,
<collectingRegion id="49BEF8970C328E4F7164FAC2FBE5535D" box="[835,1051,1303,1329]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F7602FAC2FBA1535D" box="[1061,1119,1303,1329]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F764BFAC2FB19535D" box="[1132,1255,1303,1329]" degrees="28.7639" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.7639">28.7639 N</geoCoordinate>
, 94.899 9 E;
<elevation id="0057D1460C328E4F7755FAC2FF175339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">
<quantity id="4C829B900C328E4F7755FAC2FF535339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">360 m</quantity>
a.s.l.
</elevation>
),
<date id="FFC410B50C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">
<collectingDate id="EF80E95D0C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">02.vi.2024</collectingDate>
</date>
,
<collectorName id="268F53A30C328E4F73A2FAEEFD8E5339" box="[389,624,1339,1365]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
,
<collectorName id="268F53A30C328E4F705BFAEEFCF75339" box="[636,777,1339,1365]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
,
<collectorName id="268F53A30C328E4F7132FAEEFC1E5339" box="[789,992,1339,1365]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
,
<collectorName id="268F53A30C328E4F71CDFAEEFB4C5339" box="[1002,1202,1339,1365]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
,
<collectorName id="268F53A30C328E4F769BFAEEFACF5339" box="[1212,1329,1339,1365]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4F72E0FA8AFE5B5389" pageId="1" pageNumber="572" type="etymology">
<paragraph id="8BC536750C328E4F72E0FA8AFDB553F1" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA8AFEB05315" bold="true" box="[199,334,1375,1401]" pageId="1" pageNumber="572">Etymology.</emphasis>
The specific epithet
<emphasis id="B90EEA670C328E4F7018FA8AFD8A5315" box="[575,628,1375,1401]" italics="true" pageId="1" pageNumber="572">adii</emphasis>
honours the indigenous Adi tribe of the Upper Siang District,
<collectingRegion id="49BEF8970C328E4F770FFA8AFF0D53F1" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F72D9FA56FEC653F1" box="[254,312,1411,1437]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
and neighbouring areas.
</paragraph>
<paragraph id="8BC536750C328E4F72E0FA72FE5B5389" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA72FE0C53AD" bold="true" box="[199,498,1446,1473]" pageId="1" pageNumber="572">Proposed English Name.</emphasis>
Golden Grissette: The common English name is given due to the golden yellow pterothorax of the male.
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4472B0F9C7FB95572D" lastPageId="10" lastPageNumber="581" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC536750C328E4F72B0F9C7FCFD5041" blockId="1.[151,1437,1554,2013]" box="[151,771,1554,1581]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9C7FD7F5040" bold="true" box="[151,641,1554,1581]" pageId="1" pageNumber="572">
Description of
<typeStatus id="54C188D70C328E4F7360F9C6FE505041" box="[327,430,1555,1581]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
male IBC-BR299
</emphasis>
(
<figureCitation id="13412AF00C328E4F70B7F9C6FD045041" box="[656,762,1555,1581]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[214,1373,181,1919]" captionTargetId-0="figure-18@2.[214,1373,181,1919]" captionTargetId-1="figure-17@3.[214,1373,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." captionText-1="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (ce) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (fh) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916295" figureDoi-1="http://doi.org/10.5281/zenodo.13916297" httpUri-0="https://zenodo.org/record/13916295/files/figure.png" httpUri-1="https://zenodo.org/record/13916297/files/figure.png" pageId="1" pageNumber="572">Figs. 12</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C328E4F72B0F9E2FACE50D1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9E2FF2A503D" box="[151,212,1591,1617]" italics="true" pageId="1" pageNumber="572">Head</emphasis>
(
<figureCitation id="13412AF00C328E4F72CFF9E2FEC2503D" box="[232,316,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1a</figureCitation>
). Head black, marked as follows: labrum yellowish brown; lower half of mandibles pale blue; labium bluish-yellow except for a round dark area at the center of the upper margin in life; anteclypeus pale blue; postclypeus blackish-brown; frons black. Median ocellus creamy yellow, rest shiny brown; antennae black, upper margin of pedicel pale bluish yellow. Eyes upper half brown, lower half grey in life, darker post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F912FED05145" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F912FE69508D" box="[199,407,1735,1761]" italics="true" pageId="1" pageNumber="572">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7381F912FDEA508D" box="[422,532,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1b, d</figureCitation>
). Anterior and posterior lobes black, middle lobes brown; propleuron dark brown with downward projection at posterior end: posterior margin pruinosed bluish-white in life, dark brown to black post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F8E6FDE851B1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F8E6FEB35121" box="[199,333,1843,1869]" italics="true" pageId="1" pageNumber="572">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7347F8E6FE155121" box="[352,491,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1c, ef</figureCitation>
). Mesostigmal plates and mesepisternum matt black with greenish metallic sheen; yellow stripe on mesepisternum above mesopleural suture; pterothorax laterally golden yellow in life, pale brown post-mortem; upper part of mesepimeron pale brown, faint brown markings on the interpleural suture and anterior part of metepisternum; mesinfraepisternum brown, metinfraepisternum yellow. Pterothorax ventrally pale yellow, small black spot at posterior edge.
</paragraph>
<caption id="DF0566FD0C318E4C72B0F842FDE3519B" ID-DOI="http://doi.org/10.5281/zenodo.13916295" ID-Zenodo-Dep="13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BC536750C318E4C72B0F842FDE3519B" blockId="2.[151,1437,1943,2039]" pageId="2" pageNumber="573">
<emphasis id="B90EEA670C318E4C72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="573">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4DF60C318E4C733FF842FE5151C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[280,431,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C318E4C733FF842FE5151C3" box="[280,431,1943,1967]" italics="true" pageId="2" pageNumber="573">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C318E4C7392F842FDFB51C3" bold="true" box="[437,517,1943,1967]" pageId="2" pageNumber="573">
<taxonomicNameLabel id="A23D571C0C318E4C7392F842FDFB51C3" box="[437,517,1943,1967]" pageId="2" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c).
</paragraph>
</caption>
<caption id="DF0566FD0C308E4D72B0F842FDB2519B" ID-DOI="http://doi.org/10.5281/zenodo.13916297" ID-Zenodo-Dep="13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="8BC536750C308E4D72B0F842FDB2519B" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="574">
<emphasis id="B90EEA670C308E4D72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="3" pageNumber="574">FIGURE 2.</emphasis>
<taxonomicName id="4C7A4DF60C308E4D733DF842FE4C51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[282,434,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C308E4D733DF842FE4C51C3" box="[282,434,1943,1967]" italics="true" pageId="3" pageNumber="574">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C308E4D739EF842FDF451C3" bold="true" box="[441,522,1943,1967]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A23D571C0C308E4D739EF842FDF451C3" box="[441,522,1943,1967]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (ce) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (fh) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C378E4A72E0FF42FD7D56B9" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF42FF0256DC" box="[199,252,151,176]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae pale brown with pruinosed blue in life, pale brown post-mortem; femur, tibiae brown. Tibiofemoral joint, tarsus and spines dark brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FF0AFF355709" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF0AFEF55694" box="[199,267,223,248]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A7339FF0AFE725695" box="[286,396,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1cg</figureCitation>
) very faintly tinted with brown; venation black; Pt trapezoid, upper margin is ½ of lower margin, blackish brown, nervures black, covers almost 3 ½ cells below; a slight bulge in the upper wing margin just proximal to pt on anterior side. Arculus at level of Ax 45. Ax
<quantity id="4C829B900C378E4A7147FEF2FC72572D" box="[864,908,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
right FW,
<quantity id="4C829B900C378E4A7620FEF2FBCD572D" box="[1031,1075,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="4" pageNumber="575" unit="in" value="8.0">8 in</quantity>
rest. Px
<quantity id="4C829B900C378E4A76B0FEF3FB05572D" box="[1175,1275,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">3435 in</quantity>
FW,
<quantity id="4C829B900C378E4A771EFEF3FA62572D" box="[1337,1436,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">2526 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FEBAFB1457BD" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FEBAFECF57E5" box="[199,305,367,393]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7366FEBAFE6D57E5" box="[321,403,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1c</figureCitation>
). Pale brown, marked with S1 pale brown, minute small black spots on both ends of lower margin of tergite; S2 with a darker brown lateral patch; S36 brownish black lateral stripe, fainter on S45, S710 black, S610 bluish pruinescence, S910 almost completely covered in bluish pruinescence in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FE0EFB8A540D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FE0EFECF5799" box="[199,305,475,501]" italics="true" pageId="4" pageNumber="575">Genitalia</emphasis>
(
<figureCitation id="13412AF00C378E4A7318FE0EFE525799" box="[319,428,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (ce) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (fh) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Fig. 2ae</figureCitation>
). First and second segments ventrally and dorsally dark brown. Third segment pale yellow. Apical segment translucent medially raised on dorsal side ending with a hood, lamina curved anteriorly and extended laterally, segmented in two parts on both sides, upper part forming a shoulder and ending in a thin filamentous arm slightly thicker at apex. Lower part canine-shaped, rounded, curved outward &amp; laterally.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FDBEFE84549D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FDBEFE5854E9" box="[199,422,619,645]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A7392FDBEFDD254E9" box="[437,556,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (ce) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (fh) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Figs. 2fh</figureCitation>
). Cerci black, bent inwards at midpoint making ~90° angle with proximal part; cerci apex rounded and expanded; inner margin of proximal part has bulging and numerous small black spines; cerci length more than S10; paraproct dark brown to black, 12 of the cerci length, broad at base, apex sharply pointed and curved inwards.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FD2EFB7B5579" blockId="4.[151,1437,151,789]" box="[199,1157,762,789]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 3536; HW 2930; total length 4546.</paragraph>
<paragraph id="8BC536750C378E4A72B0FC97FE015531" blockId="4.[151,1436,834,1293]" box="[151,511,834,861]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FC97FE8B5531" bold="true" box="[151,373,834,861]" pageId="4" pageNumber="575">Variation in Males</emphasis>
(
<figureCitation id="13412AF00C378E4A73A3FC96FE115531" box="[388,495,835,861]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (ef) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (kl) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (qr) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 34</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C378E4A72B0FCB2FEFB5235" blockId="4.[151,1436,834,1293]" pageId="4" pageNumber="575">
Though in life all males have similar golden yellow lateral pterothorax, post-mortem it changes to dull brown in some individuals (
<figureCitation id="13412AF00C378E4A7340FC5EFE3D55C9" box="[359,451,907,933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Figs. 1c</figureCitation>
,
<figureCitation id="13412AF00C378E4A73F6FC5EFE0E55C9" box="[465,496,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">3b</figureCitation>
). IBC-BR297, IBC-BR
<specimenCount id="9D7CFDFC0C378E4A70DDFC5EFC9255C9" box="[762,876,907,933]" count="383" pageId="4" pageNumber="575" type="male">383 males</specimenCount>
have brown upper part of mesepimeron, whereas IBC-BR300 and IBC-BR451 have clear golden yellow thorax without any post-mortem browning (
<figureCitation id="13412AF00C378E4A7738FC7AFA6E55A5" box="[1311,1424,943,969]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (ef) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (kl) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (qr) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 34</figureCitation>
). IBC-BR
<specimenCount id="9D7CFDFC0C378E4A72D2FC06FEA55581" box="[245,347,979,1005]" count="297" pageId="4" pageNumber="575" type="male">297 male</specimenCount>
has 910 Ax in FW and HW and 34 Px in FW, 27 Px in HW. IBC-BR
<specimenCount id="9D7CFDFC0C378E4A7641FC06FB325581" box="[1126,1228,979,1005]" count="300" pageId="4" pageNumber="575" type="male">300 male</specimenCount>
has 78 Ax in FW and HW and 3132 Px in FW and 2627 Px in HW (
<figureCitation id="13412AF00C378E4A70CCFC22FCCE527D" box="[747,816,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Similarly, there are 37 Px in FW and 28 Px in HW in IBC-BR
<specimenCount id="9D7CFDFC0C378E4A72D1FBCEFEA25259" box="[246,348,1051,1077]" count="383" pageId="4" pageNumber="575" type="male">383 male</specimenCount>
and 36 Px in FW and 27 Px in HW in IBC-BR
<specimenCount id="9D7CFDFC0C378E4A7154FBCFFC275259" box="[883,985,1050,1077]" count="451" pageId="4" pageNumber="575" type="male">451 male</specimenCount>
(
<figureCitation id="13412AF00C378E4A71CEFBCEFBD05259" box="[1001,1070,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Both these males have 8 Ax in all wings.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FBB6FE8C5211" blockId="4.[151,1436,834,1293]" box="[199,370,1123,1149]" pageId="4" pageNumber="575">Measurements.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB52FB9752CD" blockId="4.[151,1436,834,1293]" box="[199,1129,1159,1185]" pageId="4" pageNumber="575">IBC-BR297: Abdomen without appendages 3637; HW 3233; total length 4647.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB7EFBC152A9" blockId="4.[151,1436,834,1293]" box="[199,1087,1195,1221]" pageId="4" pageNumber="575">IBC-BR300: Abdomen without appendages 3132; HW 28; total length 4142.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB1AFB975285" blockId="4.[151,1436,834,1293]" box="[199,1129,1231,1257]" pageId="4" pageNumber="575">IBC-BR383: Abdomen without appendages 3132; HW 2930; total length 4344.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB27FB975361" blockId="4.[151,1436,834,1293]" box="[199,1129,1266,1293]" pageId="4" pageNumber="575">IBC-BR451: Abdomen without appendages 3233; HW 2829; total length 4142.</paragraph>
<paragraph id="8BC536750C378E4A72B0FAEFFCE35339" blockId="4.[151,1437,1338,1941]" box="[151,797,1338,1365]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FAEFFD625338" bold="true" box="[151,668,1338,1365]" pageId="4" pageNumber="575">
Description of
<typeStatus id="54C188D70C378E4A7360FAEEFE4C5339" box="[327,434,1339,1365]" pageId="4" pageNumber="575" type="paratype">paratype</typeStatus>
female IBC-BX390
</emphasis>
(
<figureCitation id="13412AF00C378E4A708CFAEEFCEB5339" box="[683,789,1338,1365]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1650,1674]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[151,1436,181,1627]" captionTargetId-0="figure-17@7.[214,1373,181,1919]" captionTargetId-1="figure-18@8.[151,1436,181,1627]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." captionText-1="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (bd) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13916303" figureDoi-1="http://doi.org/10.5281/zenodo.13916305" httpUri-0="https://zenodo.org/record/13916303/files/figure.png" httpUri-1="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Figs. 56</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C378E4A72B0FA8AFAA253AD" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FA8AFF2A5315" box="[151,212,1375,1401]" italics="true" pageId="4" pageNumber="575">Head</emphasis>
(
<figureCitation id="13412AF00C378E4A72C2FA8AFEA85315" box="[229,342,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5ab</figureCitation>
). Labrum, labium and mandible pale blue in life, brown post-mortem, labrum dark brown post-mortem; anteclypeus, postclypeus and frons black with blue shine. Rest of head black. Ocelli brown, antennae black. Eyes upper half brown, lower half grey in life, darker post-mortem. Postocular area and occiput black.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA1EFBFC5389" blockId="4.[151,1437,1338,1941]" box="[199,1026,1482,1509]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA1EFE665389" box="[199,408,1483,1509]" italics="true" pageId="4" pageNumber="575">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7380FA1EFDEA5389" box="[423,532,1482,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5ce</figureCitation>
). Similar to male. Slightly pruinosed in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA3AFA98503D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA3AFEB35065" box="[199,333,1519,1545]" italics="true" pageId="4" pageNumber="575">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7347FA3AFE305065" box="[352,462,1518,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5fg</figureCitation>
). Similar to male, except browner laterally. Mesepisternum black with bronze tinge; brownish yellow stripe on mesepisternum just above the mesopleural suture which is greyish blue in life; upper part of mesepimeron brown; faint brown marking on interpleural suture; rest of the lateral pterothorax pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F98EFD5150F5" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F98EFF025018" box="[199,252,1627,1652]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae brown. Femur, tibiae brown with bluish pruinescence in life, dark brown post-mortem. Tibiofemoral joint, tarsus and spines blackish brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F976FA8950D1" blockId="4.[151,1437,1338,1941]" box="[199,1399,1698,1725]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F976FEF550D0" box="[199,267,1699,1724]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A733CF976FE9350D1" box="[283,365,1699,1725]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (bd) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6a</figureCitation>
). Hyaline, similar to male; Pt pale brown. Ax
<quantity id="4C829B900C378E4A7157F976FC6550D1" box="[880,923,1699,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
all wings. Px
<quantity id="4C829B900C378E4A761DF977FB6350D1" box="[1082,1181,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">3435 in</quantity>
FW,
<quantity id="4C829B900C378E4A76FEF977FAC250D1" box="[1241,1340,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">2526 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F912FE0A5145" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F912FECF508D" box="[199,305,1735,1761]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7364F912FE69508D" box="[323,407,1734,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5c</figureCitation>
). S12 dorsally yellowish brown, laterally brown, S37 brown with faint, pale yellowish basal rings; dorsum of S6 with a large yellow, rounded, dorsal marking towards posterior margin; S910 black; abdomen ventrally pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8E6FD94511D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F8E6FE575121" box="[199,425,1843,1869]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A739DF8E6FDD25121" box="[442,556,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (bd) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6bd</figureCitation>
). Cerci short, conical, pointed, black, less than length of S10. Paraproct very small, globular, brown; ovipositor brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8AEFB7B51F9" blockId="4.[151,1437,1338,1941]" box="[199,1157,1915,1941]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 3031; HW 2930; total length 3940.</paragraph>
<caption id="DF0566FD0C368E4B72B0F842FC5451BF" ID-DOI="http://doi.org/10.5281/zenodo.13916299" ID-Zenodo-Dep="13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="5" targetType="figure">
<paragraph id="8BC536750C368E4B72B0F842FC5451BF" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="576">
<emphasis id="B90EEA670C368E4B72B0F842FEEA51C3" bold="true" box="[151,276,1943,1967]" pageId="5" pageNumber="576">FIGURE 3.</emphasis>
<taxonomicName id="4C7A4DF60C368E4B733CF842FE4D51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[283,435,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C368E4B733CF842FE4D51C3" box="[283,435,1943,1967]" italics="true" pageId="5" pageNumber="576">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C368E4B739CF842FDF251C3" bold="true" box="[443,524,1943,1967]" pageId="5" pageNumber="576">
<taxonomicNameLabel id="A23D571C0C368E4B739CF842FDF251C3" box="[443,524,1943,1967]" pageId="5" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<caption id="DF0566FD0C358E4872B0F8ECFD72519D" ID-DOI="http://doi.org/10.5281/zenodo.13916301" ID-Zenodo-Dep="13916301" httpUri="https://zenodo.org/record/13916301/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,1849,1873]" targetBox="[232,1356,151,1837]" targetPageId="6" targetType="figure">
<paragraph id="8BC536750C358E4872B0F8ECFD72519D" blockId="6.[151,1436,1849,2034]" pageId="6" pageNumber="577">
<emphasis id="B90EEA670C358E4872B0F8ECFEED513D" bold="true" box="[151,275,1849,1873]" pageId="6" pageNumber="577">FIGURE 4.</emphasis>
<taxonomicName id="4C7A4DF60C358E48733EF8EFFE4E513E" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1850,1874]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C358E48733EF8EFFE4E513E" box="[281,432,1850,1874]" italics="true" pageId="6" pageNumber="577">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C358E487391F8EFFDF9513E" bold="true" box="[438,519,1850,1874]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A23D571C0C358E487391F8EFFDF9513E" box="[438,519,1850,1874]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (ef) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (kl) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (qr) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
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<paragraph id="8BC536750C348E4972B0F842FE2D519B" blockId="7.[151,1437,1943,2039]" pageId="7" pageNumber="578">
<emphasis id="B90EEA670C348E4972B0F842FEEC51C3" bold="true" box="[151,274,1943,1967]" pageId="7" pageNumber="578">FIGURE 5.</emphasis>
<taxonomicName id="4C7A4DF60C348E497330F842FE5351C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[279,429,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C348E497330F842FE5351C3" box="[279,429,1943,1967]" italics="true" pageId="7" pageNumber="578">Devadatta adii</emphasis>
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<emphasis id="B90EEA670C348E497395F842FDFC51C3" bold="true" box="[434,514,1943,1967]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A23D571C0C348E497395F842FDFC51C3" box="[434,514,1943,1967]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c).
</paragraph>
</caption>
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<paragraph id="8BC536750C3B8E4672B0F9A7FCCD50C3" blockId="8.[151,1437,1650,1711]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F9A7FEED50E7" bold="true" box="[151,275,1650,1675]" pageId="8" pageNumber="579">FIGURE 6.</emphasis>
<taxonomicName id="4C7A4DF60C3B8E46733EF9A6FE4E50E7" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1651,1675]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3B8E46733EF9A6FE4E50E7" box="[281,432,1651,1675]" italics="true" pageId="8" pageNumber="579">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3B8E467391F9A6FDF950E7" bold="true" box="[438,519,1651,1675]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A23D571C0C3B8E467391F9A6FDF950E7" box="[438,519,1651,1675]" pageId="8" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC- BX390): (a) left forewing and hindwing; (bd) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view.
</paragraph>
</caption>
<paragraph id="8BC536750C3B8E4672B0F908FE0D509B" blockId="8.[151,1436,1757,1999]" box="[151,499,1757,1783]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F908FE71509B" bold="true" box="[151,399,1757,1783]" pageId="8" pageNumber="579">Variation in Females</emphasis>
(
<figureCitation id="13412AF00C3B8E4673B9F908FE1C509B" box="[414,482,1757,1783]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C3B8E4672B0F8D4FF4251A3" blockId="8.[151,1436,1757,1999]" pageId="8" pageNumber="579">
Stripe on mesepisternum is usually greyish blue in life, it has variable post-mortem colours (
<figureCitation id="13412AF00C3B8E467690F8D4FB035177" box="[1207,1277,1793,1819]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BX307 and IBC-BX
<specimenCount id="9D7CFDFC0C3B8E467300F8F0FE4E5153" box="[295,432,1829,1855]" count="309" pageId="8" pageNumber="579" type="female">309 females</specimenCount>
have pale bluish yellow stripe on mesepisternum, whereas IBC-BR314 and IBC-BX313 have brownish yellow stripe on mesepisternum (
<figureCitation id="13412AF00C3B8E467091F89CFD04510F" box="[694,762,1865,1891]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BR
<specimenCount id="9D7CFDFC0C3B8E46714AF89CFC17510F" box="[877,1001,1865,1891]" count="314" pageId="8" pageNumber="579" type="female">314 female</specimenCount>
has 78 Ax in all the wings and 3133 Px in FW and 2327 Px in HW. IBC-BX
<specimenCount id="9D7CFDFC0C3B8E46707AF8B8FD2751EB" box="[605,729,1901,1927]" count="307" pageId="8" pageNumber="579" type="female">307 female</specimenCount>
has clearer wings and 78 Ax and 3336, whereas IBC-BX
<specimenCount id="9D7CFDFC0C3B8E467754F8B8FF1D51C7" count="309" pageId="8" pageNumber="579" type="female">309 female</specimenCount>
has faintly brown enfumed wings and 710 Ax and 3436 Px. IBC-BX
<specimenCount id="9D7CFDFC0C3B8E46762AF844FB7451C7" box="[1037,1162,1937,1963]" count="313" pageId="8" pageNumber="579" type="female">313 female</specimenCount>
has 810 Ax and 3437 Px.
</paragraph>
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<paragraph id="8BC536750C3A8E4772B0F842FBF551BF" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="580">
<emphasis id="B90EEA670C3A8E4772B0F842FEEB51C3" bold="true" box="[151,277,1943,1967]" pageId="9" pageNumber="580">FIGURE 7.</emphasis>
<taxonomicName id="4C7A4DF60C3A8E477339F842FE4951C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[286,439,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3A8E477339F842FE4951C3" box="[286,439,1943,1967]" italics="true" pageId="9" pageNumber="580">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3A8E4773E7F842FDED51C3" bold="true" box="[448,531,1943,1967]" pageId="9" pageNumber="580">
<taxonomicNameLabel id="A23D571C0C3A8E4773E7F842FDED51C3" box="[448,531,1943,1967]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C398E4472E0FF42FE8C56DD" blockId="10.[199,1131,151,321]" box="[199,370,151,177]" pageId="10" pageNumber="581">Measurements.</paragraph>
<paragraph id="8BC536750C398E4472E0FF6EFB9756B9" blockId="10.[199,1131,151,321]" box="[199,1129,186,213]" pageId="10" pageNumber="581">IBC-BR314: Abdomen without appendages 2829; HW 2425; total length 3435.</paragraph>
<paragraph id="8BC536750C398E4472E0FF0AFB955695" blockId="10.[199,1131,151,321]" box="[199,1131,223,249]" pageId="10" pageNumber="581">IBC-BX307: Abdomen without appendages 3132; HW 3132; total length 4041.</paragraph>
<paragraph id="8BC536750C398E4472E0FED6FB955771" blockId="10.[199,1131,151,321]" box="[199,1131,259,285]" pageId="10" pageNumber="581">IBC-BX309: Abdomen without appendages 3232; HW 3031; total length 4041.</paragraph>
<paragraph id="8BC536750C398E4472E0FEF2FB95572D" blockId="10.[199,1131,151,321]" box="[199,1131,295,321]" pageId="10" pageNumber="581">IBC-BX313: Abdomen without appendages 3132; HW 3122; total length 4142.</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="CED7EAAA7F3BB829D2DFF8BD4480F8DE" bold="true" box="[151,951,1821,1848]" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D2DFF8BD458AF8DE" ID-CoL="5WNR5" authority="Friese, 1903" authorityName="Friese" authorityYear="1903" box="[151,701,1821,1848]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" sensu="lato" species="weisi" subGenus="Thoracobombus">
<emphasis id="CED7EAAA7F3BB829D2DFF8BD4529F8DE" bold="true" box="[151,542,1821,1848]" italics="true" pageId="10" pageNumber="311">Bombus (Thoracobombus) weisi</emphasis>
<bibRefCitation id="98324B497F3BB829D06EF8BD458AF8DE" author="Friese, H." box="[550,701,1821,1847]" pageId="10" pageNumber="311" pagination="253 - 255" refId="ref11221" refString="Friese, H. (1903) Neue Bombus - Arten aus der Neotropischen Region. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 3, 253 - 255." type="journal article" year="1903">Friese, 1903</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="D5E457D17F3BB829D082F8BD4472F8DE" box="[714,837,1821,1847]" pageId="10" pageNumber="311" sensu="lato">sensu lato</taxonomicNameLabel>
stat. rev.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB828D2DFF8C24441FF24" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311" type="reference_group">
<paragraph id="FC1C36B87F3BB829D2DFF8C24476F871" blockId="10.[151,1436,1890,2037]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFF8C24476F871" pageId="10" pageNumber="311">
<treatmentCitation id="7D0210A97F3BB829D2DFF8C2452EF893" author="Smith, F." box="[151,537,1890,1914]" page="153" pageId="10" pageNumber="311" year="1861">
<taxonomicName id="3BA34D3B7F3BB829D2DFF8C2452EF893" ID-CoL="MFMJ" authority="Smith, 1861: 153" authorityName="Smith" authorityPageNumber="153" authorityYear="1861" box="[151,537,1890,1914]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="laboriosus">
<emphasis id="CED7EAAA7F3BB829D2DFF8C24668F893" box="[151,351,1890,1914]" italics="true" pageId="10" pageNumber="311">Bombus laboriosus</emphasis>
<bibRefCitation id="98324B497F3BB829D321F8C2452EF893" author="Smith, F." box="[361,537,1890,1914]" pageId="10" pageNumber="311" pagination="146 - 155" refId="ref13586" refString="Smith, F. (1861) Descriptions of new genera and species of exotic Hymenoptera. Journal of Entomology, 1, 146 - 155." type="journal article" year="1861">Smith, 1861:153</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, not of
<treatmentCitation id="7D0210A97F3BB829D03BF8C2440BF893" author="Fabricius, J. C." box="[627,828,1890,1914]" page="352" pageId="10" pageNumber="311" year="1804">
<bibRefCitation id="98324B497F3BB829D03BF8C2440BF893" author="Fabricius, J. C." box="[627,828,1890,1914]" pageId="10" pageNumber="311" refId="ref11008" refString="Fabricius, J. C. (1804) Systema Piezatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Carolum Reichard, Brunsvigae, 439 + 30 pp. https: // doi. org / 10.5962 / bhl. title. 10490" type="book" year="1804">Fabricius 1804:352</bibRefCitation>
</treatmentCitation>
(=
<taxonomicName id="3BA34D3B7F3BB829D120F8C243DEF893" authority="(Fabricius))" authorityName="Smith" authorityYear="1861" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[872,1257,1890,1914]" class="Insecta" family="Apidae" genus="Emphoropsis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="laboriosus">
<emphasis id="CED7EAAA7F3BB829D120F8C24354F893" box="[872,1123,1890,1914]" italics="true" pageId="10" pageNumber="311">Emphoropsis laboriosus</emphasis>
(Fabricius))
</taxonomicName>
.
<typeStatus id="2318881A7F3BB829D6BFF8C24262F893" box="[1271,1365,1890,1914]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
queen
<collectingRegion id="3E67F85A7F3BB829D28FF8204623F871" box="[199,276,1920,1944]" country="Mexico" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingRegion>
NHMUK (images at NHMUK014025497), examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3BB829D2DFF83F42BAF85E" blockId="10.[151,1436,1890,2037]" box="[151,1421,1951,1976]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFF83F42BAF85E" box="[151,1421,1951,1976]" pageId="10" pageNumber="311">
<treatmentCitation id="7D0210A97F3BB829D2DFF83F46E0F85E" author="Friese, H." box="[151,471,1951,1975]" page="253" pageId="10" pageNumber="311" year="1903">
<taxonomicName id="3BA34D3B7F3BB829D2DFF83F46E0F85E" ID-CoL="5WNR5" authority="Friese, 1903: 253" authorityName="Friese" authorityPageNumber="253" authorityYear="1903" box="[151,471,1951,1975]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3BB829D2DFF83F4614F85E" box="[151,291,1951,1975]" italics="true" pageId="10" pageNumber="311">Bombus weisi</emphasis>
<bibRefCitation id="98324B497F3BB829D362F83F46E0F85E" author="Friese, H." box="[298,471,1951,1975]" pageId="10" pageNumber="311" pagination="253 - 255" refId="ref11221" refString="Friese, H. (1903) Neue Bombus - Arten aus der Neotropischen Region. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 3, 253 - 255." type="journal article" year="1903">Friese, 1903:253</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="2318881A7F3BB829D3A9F8004571F851" box="[481,582,1952,1976]" pageId="10" pageNumber="311" type="lectotype">Lectotype</typeStatus>
queen by designation of
<treatmentCitation id="7D0210A97F3BB829D10DF83F4333F85E" author="Milliron, H. E." box="[837,1028,1951,1975]" page="98" pageId="10" pageNumber="311" year="1960">
<bibRefCitation id="98324B497F3BB829D10DF83F4333F85E" author="Milliron, H. E." box="[837,1028,1951,1975]" pageId="10" pageNumber="311" pagination="87 - 99" refId="ref12572" refString="Milliron, H. E. (1960) Recognition of bumblebee type specimens, with notes on some dubious names (Hymenoptera: Apidae). Bulletin of the Brooklyn Entomological Society, 55, 87 - 99." type="journal article" year="1960">Milliron (1960:98)</bibRefCitation>
</treatmentCitation>
<collectingCountry id="84B476287F3BB829D642F83F434FF85E" box="[1034,1144,1951,1975]" name="Costa Rica" pageId="10" pageNumber="311">Costa Rica</collectingCountry>
ZMHB, images examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3BB829D2DFF81E4227F83F" blockId="10.[151,1436,1890,2037]" box="[151,1296,1982,2006]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFF81E4227F83F" box="[151,1296,1982,2006]" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D2DFF81E4652F83F" ID-CoL="MFHL" authorityName="Say" authorityYear="1837" box="[151,357,1982,2006]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="CED7EAAA7F3BB829D2DFF81E4652F83F" box="[151,357,1982,2006]" italics="true" pageId="10" pageNumber="311">Bombus ephippiatus</emphasis>
</taxonomicName>
[subsp.]
<treatmentCitation id="7D0210A97F3BB829D38BF81F4425F83F" author="Cockerell, T. D. A." box="[451,786,1982,2006]" page="344" pageId="10" pageNumber="311" year="1908">
<taxonomicName id="3BA34D3B7F3BB829D38BF81F4425F83F" ID-CoL="5FYSH" authority="Cockerell, 1908: 344" authorityName="Cockerell" authorityPageNumber="344" authorityYear="1908" box="[451,786,1982,2006]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="montezumae">
<emphasis id="CED7EAAA7F3BB829D38BF81F450CF83F" box="[451,571,1983,2006]" italics="true" pageId="10" pageNumber="311">montezumae</emphasis>
<bibRefCitation id="98324B497F3BB829D009F81E4425F83F" author="Cockerell, T. D. A." box="[577,786,1982,2006]" pageId="10" pageNumber="311" pagination="259 - 267" refId="ref10579" refString="Cockerell, T. D. A. (1908) Descriptions and records of bees. Annals and Magazine of Natural History, Series 8, 1, 259 - 267. [inc 344]" type="journal article" year="1908">Cockerell, 1908:344</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, replacement name for
<taxonomicName id="3BA34D3B7F3BB829D648F81F423BF83F" authority="Smith, 1861" authorityName="Smith" authorityYear="1861" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[1024,1292,1982,2006]" class="Insecta" family="Apidae" genus="Emphoropsis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="laboriosus">
<emphasis id="CED7EAAA7F3BB829D648F81F43B0F83F" box="[1024,1159,1982,2006]" italics="true" pageId="10" pageNumber="311">B. laboriosus</emphasis>
<bibRefCitation id="98324B497F3BB829D6C5F81E423BF83F" author="Smith, F." box="[1165,1292,1982,2006]" pageId="10" pageNumber="311" pagination="146 - 155" refId="ref13586" refString="Smith, F. (1861) Descriptions of new genera and species of exotic Hymenoptera. Journal of Entomology, 1, 146 - 155." type="journal article" year="1861">Smith, 1861</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3BB829D2DFF87D42A3F81D" blockId="10.[151,1436,1890,2037]" box="[151,1428,2012,2037]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFF87D42A3F81D" box="[151,1428,2012,2037]" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D2DFF87D4666F81D" ID-CoL="MFP5" authorityName="Cockerell" authorityYear="1951" box="[151,337,2013,2036]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="mateonis">
<emphasis id="CED7EAAA7F3BB829D2DFF87D4666F81D" box="[151,337,2013,2036]" italics="true" pageId="10" pageNumber="311">BOMBUS MATEONIS</emphasis>
</taxonomicName>
[sic]
<treatmentCitation id="7D0210A97F3BB829D3C3F87C456AF81C" author="Cockerell, T. D. A." box="[395,605,2012,2037]" page="487" pageId="10" pageNumber="311" year="1949">
<bibRefCitation id="98324B497F3BB829D3C3F87C456AF81C" author="Cockerell, T. D. A." box="[395,605,2012,2037]" pageId="10" pageNumber="311" pagination="429 - 490" refId="ref10616" refString="Cockerell, T. D. A. (1949) Bees from Central America, principally Honduras. Proceedings of the US National Museum, 98, 429 - 490. https: // doi. org / 10.5479 / si. 00963801.98 - 3233.429" type="journal article" year="1949">Cockerell, 1949:487</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="2318881A7F3BB829D02EF87C45F3F81D" box="[614,708,2012,2036]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
male
<collectingCountry id="84B476287F3BB829D14AF87C4447F81D" box="[770,880,2012,2036]" name="Guatemala" pageId="10" pageNumber="311">Guatemala</collectingCountry>
<collectionCode id="9AB2AE7D7F3BB829D13EF87D4488F81C" box="[886,959,2013,2037]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="311" type="Museum">USNM</collectionCode>
(images at USNMENT00534366), examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3AB828D2DFFF374441FF46" blockId="11.[151,886,151,205]" box="[151,886,151,175]" pageId="11" pageNumber="312">
<treatmentCitationGroup id="DCB311967F3AB828D2DFFF374441FF46" box="[151,886,151,175]" pageId="11" pageNumber="312">
[
<taxonomicName id="3BA34D3B7F3AB828D2E8FF374663FF47" ID-CoL="5WNR5" authority="Brasero et al., 2019" box="[160,340,151,175]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="subSpecies" species="ombus" subSpecies="weisi">
<emphasis id="CED7EAAA7F3AB828D2E8FF374663FF47" box="[160,340,151,175]" italics="true" pageId="11" pageNumber="312">B.&lt;ombus&gt; weisi</emphasis>
</taxonomicName>
(part B) [
<collectingCountry id="84B476287F3AB828D3FEFF37453FFF46" box="[438,520,151,175]" name="Mexico" pageId="11" pageNumber="312">Mexico</collectingCountry>
,
<collectingRegion id="3E67F85A7F3AB828D05BFF374550FF46" box="[531,615,151,175]" country="Mexico" name="Chiapas" pageId="11" pageNumber="312">Chiapas</collectingRegion>
]
<treatmentCitation id="7D0210A97F3AB828D03BFF37445BFF46" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[627,876,151,175]" page="190" pageId="11" pageNumber="312" year="2019">
<bibRefCitation id="98324B497F3AB828D03BFF37445BFF46" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[627,876,151,175]" pageId="11" pageNumber="312" pagination="183 - 194" refId="ref10378" refString="Brasero, N., Vandame, R., Sagot, P., Martinet, B., Valterova, I. &amp; Rasmont, P. (2019) Thoracobombus from Mexico: a description of the male species-specific cephalic labial gland secretons. Apidologie, 50, 183 - 194. https: // doi. org / 10.1007 / s 13592 - 018 - 0629 - 4" type="journal article" year="2019">
Brasero
<emphasis id="CED7EAAA7F3AB828D080FF3745CBFF46" box="[712,764,151,175]" italics="true" pageId="11" pageNumber="312">et al.</emphasis>
, 2019:190
</bibRefCitation>
</treatmentCitation>
.]
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3AB828D2DFFF164441FF24" blockId="11.[151,886,151,205]" box="[151,886,181,205]" pageId="11" pageNumber="312">
<treatmentCitationGroup id="DCB311967F3AB828D2DFFF164441FF24" box="[151,886,181,205]" pageId="11" pageNumber="312">
[
<taxonomicName id="3BA34D3B7F3AB828D2E8FF154663FF24" ID-CoL="5WNR5" authority="Brasero et al., 2019" box="[160,340,181,205]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="subSpecies" species="ombus" subSpecies="weisi">
<emphasis id="CED7EAAA7F3AB828D2E8FF154663FF24" box="[160,340,181,205]" italics="true" pageId="11" pageNumber="312">B.&lt;ombus&gt; weisi</emphasis>
</taxonomicName>
(part C) [
<collectingCountry id="84B476287F3AB828D3FEFF15453FFF24" box="[438,520,181,205]" name="Mexico" pageId="11" pageNumber="312">Mexico</collectingCountry>
,
<collectingRegion id="3E67F85A7F3AB828D05BFF154550FF24" box="[531,615,181,205]" country="Mexico" name="Chiapas" pageId="11" pageNumber="312">Chiapas</collectingRegion>
]
<treatmentCitation id="7D0210A97F3AB828D03BFF16445BFF24" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[627,876,181,205]" page="190" pageId="11" pageNumber="312" year="2019">
<bibRefCitation id="98324B497F3AB828D03BFF16445BFF24" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[627,876,181,205]" pageId="11" pageNumber="312" pagination="183 - 194" refId="ref10378" refString="Brasero, N., Vandame, R., Sagot, P., Martinet, B., Valterova, I. &amp; Rasmont, P. (2019) Thoracobombus from Mexico: a description of the male species-specific cephalic labial gland secretons. Apidologie, 50, 183 - 194. https: // doi. org / 10.1007 / s 13592 - 018 - 0629 - 4" type="journal article" year="2019">
Brasero
<emphasis id="CED7EAAA7F3AB828D080FF1645CBFF24" box="[712,764,181,205]" italics="true" pageId="11" pageNumber="312">et al.</emphasis>
, 2019:190
</bibRefCitation>
</treatmentCitation>
.]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3AB828D2DFFF5B47ECFDB0" pageId="11" pageNumber="312" type="diagnosis">
<paragraph id="FC1C36B87F3AB828D2DFFF5B4612FE4C" blockId="11.[151,1437,251,1394]" pageId="11" pageNumber="312">
<emphasis id="CED7EAAA7F3AB828D2DFFF5B463DFEFC" bold="true" box="[151,266,251,277]" pageId="11" pageNumber="312">Diagnosis</emphasis>
.
<emphasis id="CED7EAAA7F3AB828D350FF5B465BFEFC" box="[280,364,251,277]" italics="true" pageId="11" pageNumber="312">Female</emphasis>
. (cf.
<emphasis id="CED7EAAA7F3AB828D3EDFF5C45ADFEFC" box="[421,666,251,277]" italics="true" pageId="11" pageNumber="312">
<taxonomicName id="3BA34D3B7F3AB828D3EDFF5C456DFEFC" authorityName="Franklin. Our" authorityYear="1907" box="[421,602,251,277]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="nigrodorsalis">B. nigrodorsalis</taxonomicName>
<taxonomicNameLabel id="D5E457D17F3AB828D028FF5C45ADFEFC" box="[608,666,252,277]" pageId="11" pageNumber="312" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
) Metasomal T
<quantity id="3B5B9B5D7F3AB828D175FF5C4450FEFC" box="[829,871,251,278]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="312" unit="in" value="2.0">2 in</quantity>
the middle between the large pits (with hair bases) with the surface weakly sculptured, smooth and shining. Short hair between the antennal base and the ocelli black; hair of the thoracic dorsum usually anteriorly black, at most with a minority of yellow hairs intermixed in a weak transverse band; metasomal T3 usually yellow at least medially but black laterally; hind basitarsus posterior long fringe black.
</paragraph>
<paragraph id="FC1C36B87F3AB828D28FFE0F47ECFDB0" blockId="11.[151,1437,251,1394]" pageId="11" pageNumber="312">
<emphasis id="CED7EAAA7F3AB828D28FFE0F4636FE20" box="[199,257,431,457]" italics="true" pageId="11" pageNumber="312">Male</emphasis>
. (cf.
<emphasis id="CED7EAAA7F3AB828D377FE10450DFE20" box="[319,570,431,457]" italics="true" pageId="11" pageNumber="312">
<taxonomicName id="3BA34D3B7F3AB828D377FE1046C2FE20" authorityName="Franklin. Our" authorityYear="1907" box="[319,501,431,457]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="nigrodorsalis">B. nigrodorsalis</taxonomicName>
<taxonomicNameLabel id="D5E457D17F3AB828D3B6FE10450DFE20" box="[510,570,432,457]" pageId="11" pageNumber="312" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
) Gonostylus as long as broad, triangular, with the inwardly projecting inner basal process strongly sclerotised for a length as much as the breadth of the volsella (
<figureCitation id="64982A3D7F3AB828D6CEFE7343F9FE07" box="[1158,1230,467,494]" captionStart="FIGURES 45" captionStartId="9.[151,275,910,936]" captionTargetBox="[167,1397,190,901]" captionTargetPageId="9" captionText="FIGURES 45. Male genitalia of (4) B. nigrodorsalis and (5) B. weisi from the right lateral aspect, with posterior extremities at the top of the images and dorsal parts to the right of the images." pageId="11" pageNumber="312">Fig. 5</figureCitation>
); volsella distally (posteriorly) remaining broad and nearly parallel-sided throughout its length beyond the gonostylus; penis-valve head distally straight with the tip rounded. Hair of metasomal T3 with at least a distinct small lateral patch of black.
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3AB828D28FFDC344C5FBE3" pageId="11" pageNumber="312" type="description">
<paragraph id="FC1C36B87F3AB828D28FFDC344B7FCD8" blockId="11.[151,1437,251,1394]" pageId="11" pageNumber="312">
<emphasis id="CED7EAAA7F3AB828D28FFDC34666FD94" bold="true" box="[199,337,611,637]" pageId="11" pageNumber="312">Description</emphasis>
.
<emphasis id="CED7EAAA7F3AB828D32CFDC3468FFD94" box="[356,440,611,637]" italics="true" pageId="11" pageNumber="312">Female</emphasis>
. Queen body length
<quantity id="3B5B9B5D7F3AB828D0FAFDC34405FD97" box="[690,818,611,638]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="11" pageNumber="312" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">1517 mm</quantity>
, worker
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. Head with the oculo-malar area length / breadth ratio 1.3. Thorax with the mid basitarsus distal posterior corner acutely and spinosely pointed. Hair long, uneven, and black; pronotum and anterior scutum either sometimes with yellow intermixed, forming a weak transverse band connected with the yellow hypoepimeral area, or more usually this band reduced or replaced with black; scutellum posteriorly with a weak yellow fringe, which may be much reduced with black; lateral propodeum yellow; legs including bases, femora, and corbicular fringes black.
</paragraph>
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<emphasis id="CED7EAAA7F3AB828D28FFC9B4636FCBC" box="[199,257,827,853]" italics="true" pageId="11" pageNumber="312">Male</emphasis>
. Body length
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. Hair long, uneven, and black; mandibular ventral fringe with long curved hair brown; vertex and occiput sometimes mixed with yellow; pronotum and anterior scutum either yellow, forming a transverse band connected with the yellow hypoepimeral area, or often replaced with black; mesepisternum, hypoepimeral area and metepisternum with yellow; lateral propodeum yellow; leg bases and femora sometimes partly yellow; metasomal T12 yellow; T3 yellow, but always with patches of black at least at the sides; T6 at least with a posterior orange fringe; T7 entirely orange. Genitalia shown in
<figureCitation id="64982A3D7F3AB828D1E2FC4F44DAFBE3" box="[938,1005,1007,1034]" captionStart="FIGURES 45" captionStartId="9.[151,275,910,936]" captionTargetBox="[167,1397,190,901]" captionTargetPageId="9" captionText="FIGURES 45. Male genitalia of (4) B. nigrodorsalis and (5) B. weisi from the right lateral aspect, with posterior extremities at the top of the images and dorsal parts to the right of the images." pageId="11" pageNumber="312">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3AB828D28FFBB34219FBB8" pageId="11" pageNumber="312" type="materials_examined">
<paragraph id="FC1C36B87F3AB828D28FFBB34219FBB8" blockId="11.[151,1437,251,1394]" pageId="11" pageNumber="312">
<emphasis id="CED7EAAA7F3AB828D28FFBB3469EFBC4" bold="true" box="[199,425,1043,1069]" pageId="11" pageNumber="312">Material examined</emphasis>
.
<materialsCitation id="4CCB3CE57F3AB828D3FFFBB44219FBB8" pageId="11" pageNumber="312">
<collectionCode id="9AB2AE7D7F3AB828D3FFFBB4451DFBC7" box="[439,554,1044,1070]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="11" pageNumber="312" type="Museum">ECOSUR</collectionCode>
: 1195 specimens. Data and barcodes for specimens from
<figureCitation id="64982A3D7F3AB828D6EFFBB3423FFBC7" box="[1191,1288,1043,1070]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,260,1590,1616]" captionStartId-1="8.[151,260,1760,1786]" captionTargetBox-0="[151,1435,180,1565]" captionTargetBox-1="[151,1434,181,1733]" captionTargetId-0="figure-30@6.[151,1435,179,1566]" captionTargetId-1="figure-30@8.[151,1435,179,1736]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 2. Support for apparent species gene coalescents from unique haplotypes for COI barcode-like sequences including numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MRBAYES and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intraspecies branching PTP models (species coalescent nodes). Sequence labels include: sequence length in number of bases; taxon name; country in capitals (if not shown then MEXICO) and province; sequence identification code (E# = ECOSUR#). Sequence labels are followed by the percentage of the bases A and T at the third codon position, where low scores (≤97.6%) are likely to indicate numts. Labels for sequences interpreted as likely lower-divergence numts are shown in orange and for likely higher-divergence numts are shown in red (see Fig. 1). The scale bar represents branch length on the tree in substitutions per nucleotide." captionText-1="FIGURE 3. Support for species gene coalescents from unique haplotypes for COI barcode (orthologous) sequences excluding likely numts (orange and red bars and labels in Figs 12), obtained from an evolutionary tree estimated with MRBAYES and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at nodes, treebranch colours, sequence labels, and scale bar as in Fig. 2." figureDoi-0="http://doi.org/10.5281/zenodo.13914596" figureDoi-1="http://doi.org/10.5281/zenodo.13914598" httpUri-0="https://zenodo.org/record/13914596/files/figure.png" httpUri-1="https://zenodo.org/record/13914598/files/figure.png" pageId="11" pageNumber="312">Figs 23</figureCitation>
are available from https://www.ecosur.mx/beesofmesoamerica/ and barcodes are available from a Supplementary File.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3AB828D28FFBFB46ACFA9B" pageId="11" pageNumber="312" type="distribution">
<paragraph id="FC1C36B87F3AB828D28FFBFB46ACFA9B" blockId="11.[151,1437,251,1394]" pageId="11" pageNumber="312">
<emphasis id="CED7EAAA7F3AB828D28FFBFB466EFB9C" bold="true" box="[199,345,1115,1141]" pageId="11" pageNumber="312">Distribution</emphasis>
.
<taxonomicName id="3BA34D3B7F3AB828D323FBFB453DFB9C" authorityName="Friese" authorityYear="1903" box="[363,522,1115,1141]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3AB828D323FBFB453DFB9C" box="[363,522,1115,1141]" italics="true" pageId="11" pageNumber="312">Bombus weisi</emphasis>
</taxonomicName>
is distributed from
<collectingRegion id="3E67F85A7F3AB828D0BCFBFC446EFB9F" box="[756,857,1116,1142]" country="Mexico" name="Durango" pageId="11" pageNumber="312">Durango</collectingRegion>
,
<collectingCountry id="84B476287F3AB828D12FFBFB448AFB9C" box="[871,957,1115,1141]" name="Mexico" pageId="11" pageNumber="312">Mexico</collectingCountry>
(northernmost point
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) south to
<collectingRegion id="3E67F85A7F3AB828D2DFFB2047CBFB73" box="[151,252,1152,1178]" country="Costa Rica" name="San Jose" pageId="11" pageNumber="312">San José</collectingRegion>
,
<collectingCountry id="84B476287F3AB828D340FB2046B4FB70" box="[264,387,1151,1178]" name="Costa Rica" pageId="11" pageNumber="312">Costa Rica</collectingCountry>
(southernmost point
<geoCoordinate id="9997507F7F3AB828D03CFBDF45EAFB73" box="[628,733,1151,1178]" degrees="9.369" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="55" value="9.369">9.369° N</geoCoordinate>
), occurring on both sides of the Tehuantepec Isthmus (
<figureCitation id="64982A3D7F3AB828D701FBDF42B9FB70" box="[1353,1422,1151,1177]" captionStart="FIGURE 6" captionStartId="9.[151,260,1632,1658]" captionTargetBox="[357,1229,1030,1608]" captionTargetId="figure-72@9.[357,1229,1030,1608]" captionTargetPageId="9" captionText="FIGURE 6. Map showing the distributions of samples examined for this study of the weisi-complex: (white spots) taxon nigrodorsalis s. str. (=B. nigrodorsalis stat. rev.); (grey spots) taxon montezumae (=B. weisi stat. rev.); and (black spots) taxon weisi s. str. (=B. weisi stat. rev.). Larger symbols with spots in their centres show locations from which specimens were sequenced for each taxon. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI." figureDoi="http://doi.org/10.5281/zenodo.13914600" httpUri="https://zenodo.org/record/13914600/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
). Its elevational range is
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asl through a variety of habitats, from pine or mixed forest to cloud forest, including cultivated land in semi-open areas. 95% of
<taxonomicName id="3BA34D3B7F3AB828D0ABFB6844A0FB08" authorityName="Franklin. Our" authorityYear="1907" box="[739,919,1223,1249]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3AB828D0ABFB6844A0FB08" box="[739,919,1223,1249]" italics="true" pageId="11" pageNumber="312">B. nigrodorsalis</emphasis>
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records are from above
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, while
<taxonomicName id="3BA34D3B7F3AB828D70DFB6842AAFB08" authorityName="Friese" authorityYear="1903" box="[1349,1437,1224,1249]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3AB828D70DFB6842AAFB08" box="[1349,1437,1224,1249]" italics="true" pageId="11" pageNumber="312">B. weisi</emphasis>
</taxonomicName>
is more abundant below this elevation.
<taxonomicName id="3BA34D3B7F3AB828D01EFB4B45C5FAEC" authorityName="Friese" authorityYear="1903" box="[598,754,1259,1285]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3AB828D01EFB4B45C5FAEC" box="[598,754,1259,1285]" italics="true" pageId="11" pageNumber="312">Bombus weisi</emphasis>
</taxonomicName>
has been collected all year round, except in May, with 95% of the records in July to December.
<taxonomicName id="3BA34D3B7F3AB828D07AFAAF45F8FAC0" authorityName="Friese" authorityYear="1903" box="[562,719,1295,1321]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3AB828D07AFAAF45F8FAC0" box="[562,719,1295,1321]" italics="true" pageId="11" pageNumber="312">Bombus weisi</emphasis>
</taxonomicName>
and
<taxonomicName id="3BA34D3B7F3AB828D140FAB04488FAC0" authorityName="Franklin. Our" authorityYear="1907" box="[776,959,1295,1321]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3AB828D140FAB04488FAC0" box="[776,959,1295,1321]" italics="true" pageId="11" pageNumber="312">B. nigrodorsalis</emphasis>
</taxonomicName>
are broadly sympatric in some regions of
<collectingCountry id="84B476287F3AB828D2DFFA9347D8FAA4" box="[151,239,1331,1357]" name="Mexico" pageId="11" pageNumber="312">Mexico</collectingCountry>
, from
<collectingRegion id="3E67F85A7F3AB828D372FA9446DCFAA7" box="[314,491,1331,1358]" country="Mexico" name="San Luis Potosi" pageId="11" pageNumber="312">San Luis Potosí</collectingRegion>
to
<collectingRegion id="3E67F85A7F3AB828D058FA944553FAA7" box="[528,612,1332,1358]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="312">Oaxaca</collectingRegion>
(
<figureCitation id="64982A3D7F3AB828D03CFA93458EFAA4" box="[628,697,1331,1357]" captionStart="FIGURE 6" captionStartId="9.[151,260,1632,1658]" captionTargetBox="[357,1229,1030,1608]" captionTargetId="figure-72@9.[357,1229,1030,1608]" captionTargetPageId="9" captionText="FIGURE 6. Map showing the distributions of samples examined for this study of the weisi-complex: (white spots) taxon nigrodorsalis s. str. (=B. nigrodorsalis stat. rev.); (grey spots) taxon montezumae (=B. weisi stat. rev.); and (black spots) taxon weisi s. str. (=B. weisi stat. rev.). Larger symbols with spots in their centres show locations from which specimens were sequenced for each taxon. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI." figureDoi="http://doi.org/10.5281/zenodo.13914600" httpUri="https://zenodo.org/record/13914600/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
), but
<taxonomicName id="3BA34D3B7F3AB828D0B3FA944463FAA4" authorityName="Friese" authorityYear="1903" box="[763,852,1332,1357]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3AB828D0B3FA944463FAA4" box="[763,852,1332,1357]" italics="true" pageId="11" pageNumber="312">B. weisi</emphasis>
</taxonomicName>
extends beyond
<collectingCountry id="84B476287F3AB828D65BFA93435EFAA4" box="[1043,1129,1331,1357]" name="Mexico" pageId="11" pageNumber="312">Mexico</collectingCountry>
southwards as far as
<collectingCountry id="84B476287F3AB828D714FA9447FDFA98" name="Costa Rica" pageId="11" pageNumber="312">Costa Rica</collectingCountry>
(
<bibRefCitation id="98324B497F3AB828D293FAF746B8FA9B" author="Labougle, J. M." box="[219,399,1367,1394]" pageId="11" pageNumber="312" pagination="35 - 73" refId="ref11948" refString="Labougle, J. M. (1990) Bombus of Mexico and Central America (Hymenoptera, Apidae). Kansas University Science Bulletin, 54, 35 - 73." type="journal article" year="1990">Labougle, 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,259 @@
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<mods:url id="85F4677D41EABFC9407CF379C1F32877">http://dx.doi.org/10.11646/zootaxa.5514.4.1</mods:url>
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<mods:classification id="06C76193CE6D9D57BD6EFB85368F4AA4">journal article</mods:classification>
<mods:identifier id="A13452756123915D98465DCD445250E6" type="DOI">10.11646/zootaxa.5514.4.1</mods:identifier>
<mods:identifier id="B5A15E60F2D7928F33158827579BB0FD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C0C02A2C38FB512700708DA5C5714708" type="Zenodo-Dep">13914592</mods:identifier>
<mods:identifier id="B2F76C86A9135128548E6BB37FADFC2E" type="ZooBank">436EC045-C3AE-4BB9-9709-202B8E2E7CEB</mods:identifier>
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<treatment id="740A87AE7F3BB829D2DFFE8647EDF922" LSID="urn:lsid:plazi:treatment:740A87AE7F3BB829D2DFFE8647EDF922" httpUri="http://treatment.plazi.org/id/740A87AE7F3BB829D2DFFE8647EDF922" lastPageNumber="311" pageId="10" pageNumber="311">
<subSubSection id="B4B965337F3BB829D2DFFE86436AFEA8" box="[151,1117,294,321]" pageId="10" pageNumber="311" type="nomenclature">
<paragraph id="FC1C36B87F3BB829D2DFFE86436AFEA8" blockId="10.[151,1117,294,321]" box="[151,1117,294,321]" pageId="10" pageNumber="311">
<heading id="A75481D47F3BB829D2DFFE86436AFEA8" bold="true" box="[151,1117,294,321]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
<emphasis id="CED7EAAA7F3BB829D2DFFE86436AFEA8" bold="true" box="[151,1117,294,321]" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D2DFFE864474FEA9" ID-CoL="MFR7" authority="Franklin, 1907" authorityName="Franklin. Our" authorityYear="1907" box="[151,835,294,321]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" sensu="stricto" species="nigrodorsalis" subGenus="Thoracobombus">
<emphasis id="CED7EAAA7F3BB829D2DFFE8645B5FEA8" bold="true" box="[151,642,294,321]" italics="true" pageId="10" pageNumber="311">Bombus (Thoracobombus) nigrodorsalis</emphasis>
<bibRefCitation id="98324B497F3BB829D0C1FE864474FEA9" author="Franklin, H. J." box="[649,835,294,320]" pageId="10" pageNumber="311" pagination="90 - 93" refId="ref11108" refString="Franklin, H. J. (1907) Notes on Bombinae, with descriptions of new species. Entomological News, 18, 90 - 93." type="journal article" year="1907">Franklin, 1907</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="D5E457D17F3BB829D118FE8744DBFEA9" box="[848,1004,294,321]" pageId="10" pageNumber="311" sensu="stricto">sensu stricto</taxonomicNameLabel>
stat. rev.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB829D2DFFECB449AFE36" pageId="10" pageNumber="311" type="reference_group">
<paragraph id="FC1C36B87F3BB829D2DFFECB426FFE6A" blockId="10.[151,1436,363,479]" box="[151,1368,363,387]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFFECB426FFE6A" box="[151,1368,363,387]" pageId="10" pageNumber="311">
<treatmentCitation id="7D0210A97F3BB829D2DFFECB4501FE6A" author="Franklin, H. J." box="[151,566,363,387]" page="90" pageId="10" pageNumber="311" year="1907">
<taxonomicName id="3BA34D3B7F3BB829D2DFFECB4501FE6A" ID-CoL="MFR7" authority="Franklin, 1907: 90" authorityName="Franklin. Our" authorityPageNumber="90" authorityYear="1907" box="[151,566,363,387]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3BB829D2DFFECB4640FE6A" box="[151,375,363,387]" italics="true" pageId="10" pageNumber="311">Bombus nigrodorsalis</emphasis>
<bibRefCitation id="98324B497F3BB829D335FECB4501FE6A" author="Franklin, H. J." box="[381,566,363,387]" pageId="10" pageNumber="311" pagination="90 - 93" refId="ref11108" refString="Franklin, H. J. (1907) Notes on Bombinae, with descriptions of new species. Entomological News, 18, 90 - 93." type="journal article" year="1907">Franklin, 1907:90</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="2318881A7F3BB829D077FECB45AAFE6A" box="[575,669,363,387]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
queen
<collectingRegion id="3E67F85A7F3BB829D0AFFECB4403FE6A" box="[743,820,363,387]" country="Mexico" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingRegion>
USNM (images at USNMENT01044337), examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3BB829D2DFFE2A4606FE29" blockId="10.[151,1436,363,479]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFFE2A4606FE29" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D2DFFE2A46BDFE4B" box="[151,394,394,418]" italics="true" pageId="10" pageNumber="311">
&lt;
<taxonomicName id="3BA34D3B7F3BB829D2EDFE2A46BDFE4B" ID-CoL="MFR7" authorityName="Franklin. Our" authorityYear="1907" box="[165,394,394,418]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">Bombus nigrodorsalis</taxonomicName>
</emphasis>
<treatmentCitation id="7D0210A97F3BB829D3C1FE2A45C2FE4B" author="Franklin, H. J." box="[393,757,394,418]" page="91" pageId="10" pageNumber="311" year="1907">
<emphasis id="CED7EAAA7F3BB829D3C1FE2A46A0FE4B" box="[393,407,394,418]" italics="true" pageId="10" pageNumber="311">&gt;</emphasis>
subsp.
<emphasis id="CED7EAAA7F3BB829D3A9FE2A450DFE4B" box="[481,570,394,418]" italics="true" pageId="10" pageNumber="311">laticollis</emphasis>
<bibRefCitation id="98324B497F3BB829D076FE2A45C2FE4B" author="Franklin, H. J." box="[574,757,394,418]" pageId="10" pageNumber="311" pagination="90 - 93" refId="ref11108" refString="Franklin, H. J. (1907) Notes on Bombinae, with descriptions of new species. Entomological News, 18, 90 - 93." type="journal article" year="1907">Franklin, 1907:91</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="2318881A7F3BB829D0B5FE2A446CFE4B" box="[765,859,394,418]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
queen
<collectingRegion id="3E67F85A7F3BB829D1EAFE2A44D8FE4B" box="[930,1007,394,418]" country="Mexico" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingRegion>
USNM (images at USNMENT00534364), examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC1C36B87F3BB829D2DFFE68449AFE36" blockId="10.[151,1436,363,479]" box="[151,941,455,479]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="DCB311967F3BB829D2DFFE68449AFE36" box="[151,941,455,479]" pageId="10" pageNumber="311">
[
<taxonomicName id="3BA34D3B7F3BB829D2E8FE674663FE36" ID-CoL="5WNR5" authority="Brasero et al., 2019:" box="[160,340,455,479]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="subSpecies" species="ombus" subSpecies="weisi">
<emphasis id="CED7EAAA7F3BB829D2E8FE674663FE36" box="[160,340,455,479]" italics="true" pageId="10" pageNumber="311">B.&lt;ombus&gt; weisi</emphasis>
</taxonomicName>
(part
<taxonomicName id="3BA34D3B7F3BB829D3C7FE674500FE36" ID-CoL="MFR7" authorityName="Franklin. Our" authorityYear="1907" box="[399,567,455,479]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">
A:
<emphasis id="CED7EAAA7F3BB829D3E7FE674500FE36" box="[431,567,455,479]" italics="true" pageId="10" pageNumber="311">nigrodorsalis</emphasis>
</taxonomicName>
) [
<collectingCountry id="84B476287F3BB829D003FE6745A9FE36" box="[587,670,455,479]" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingCountry>
]
<treatmentCitation id="7D0210A97F3BB829D0E2FE684494FE36" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[682,931,455,479]" page="190" pageId="10" pageNumber="311" year="2019">
<bibRefCitation id="98324B497F3BB829D0E2FE684494FE36" author="Brasero, N. &amp; Vandame, R. &amp; Sagot, P. &amp; Martinet, B. &amp; Valterova, I. &amp; Rasmont, P." box="[682,931,455,479]" pageId="10" pageNumber="311" pagination="183 - 194" refId="ref10378" refString="Brasero, N., Vandame, R., Sagot, P., Martinet, B., Valterova, I. &amp; Rasmont, P. (2019) Thoracobombus from Mexico: a description of the male species-specific cephalic labial gland secretons. Apidologie, 50, 183 - 194. https: // doi. org / 10.1007 / s 13592 - 018 - 0629 - 4" type="journal article" year="2019">
Brasero
<emphasis id="CED7EAAA7F3BB829D0B7FE684405FE36" box="[767,818,455,479]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
, 2019:190
</bibRefCitation>
</treatmentCitation>
.]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB829D2DFFDAD4407FCAE" pageId="10" pageNumber="311" type="diagnosis">
<paragraph id="FC1C36B87F3BB829D2DFFDAD4402FD7D" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D2DFFDAD463DFDCE" bold="true" box="[151,266,525,551]" pageId="10" pageNumber="311">Diagnosis</emphasis>
.
<emphasis id="CED7EAAA7F3BB829D35FFDAD465CFDCE" box="[279,363,525,551]" italics="true" pageId="10" pageNumber="311">Female</emphasis>
. (cf.
<emphasis id="CED7EAAA7F3BB829D3ECFDAE451FFDCE" box="[420,552,525,551]" italics="true" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D3ECFDAE46CBFDCE" authorityName="Williams &amp; Sagot &amp; Martínez-López &amp; Barajas &amp; Mérida-Rivas &amp; Vandame" authorityYear="2024" box="[420,508,526,551]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="weisi">B. weisi</taxonomicName>
<taxonomicNameLabel id="D5E457D17F3BB829D049FDAE451FFDCE" box="[513,552,525,551]" pageId="10" pageNumber="311" sensu="lato">s. l.</taxonomicNameLabel>
</emphasis>
) Metasomal T
<quantity id="3B5B9B5D7F3BB829D081FDAD45C4FDCE" box="[713,755,525,551]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="311" unit="in" value="2.0">2 in</quantity>
the middle between the pits (with hair bases) with the surface strongly sculptured, irregular and dull. Short hair between the antennal base and the ocelli predominantly yellow; hair of the thoracic dorsum usually anteriorly with a transverse yellow band; metasomal T3 entirely yellow; hind basitarsus posterior long fringe with at least the tips orange.
</paragraph>
<paragraph id="FC1C36B87F3BB829D28FFD3D4407FCAE" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D28FFD3D4636FD5E" box="[199,257,669,695]" italics="true" pageId="10" pageNumber="311">Male</emphasis>
. (cf.
<emphasis id="CED7EAAA7F3BB829D371FD3E468CFD5E" box="[313,443,669,695]" italics="true" pageId="10" pageNumber="311">
<taxonomicName id="3BA34D3B7F3BB829D371FD3E46A7FD5E" authorityName="Williams &amp; Sagot &amp; Martínez-López &amp; Barajas &amp; Mérida-Rivas &amp; Vandame" authorityYear="2024" box="[313,400,670,695]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="weisi">B. weisi</taxonomicName>
<taxonomicNameLabel id="D5E457D17F3BB829D3DDFD3E468CFD5E" box="[405,443,669,695]" pageId="10" pageNumber="311" sensu="lato">s. l.</taxonomicNameLabel>
</emphasis>
) Genitalia with the gonostylus shorter than long, rectangular, with the inwardly projecting inner basal process strongly sclerotised for a length less than half of the breadth of the volsella (
<figureCitation id="64982A3D7F3BB829D6AFFD614219FD32" box="[1255,1326,705,731]" captionStart="FIGURES 45" captionStartId="9.[151,275,910,936]" captionTargetBox="[167,1397,190,901]" captionTargetPageId="9" captionText="FIGURES 45. Male genitalia of (4) B. nigrodorsalis and (5) B. weisi from the right lateral aspect, with posterior extremities at the top of the images and dorsal parts to the right of the images." pageId="10" pageNumber="311">Fig. 4</figureCitation>
); volsella distally (posteriorly) beyond the gonostylus strongly narrowed or constricted proximally to the inner distal process; penis-valve head distally (posteriorly) slightly pointed and slightly incurved towards the midline. Hair of metasomal T3 laterally yellow without a distinct lateral patch of black.
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB829D28FFCF14410FA6E" pageId="10" pageNumber="311" type="description">
<paragraph id="FC1C36B87F3BB829D28FFCF14626FB65" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D28FFCF14666FC82" bold="true" box="[199,337,849,875]" pageId="10" pageNumber="311">Description</emphasis>
.
<emphasis id="CED7EAAA7F3BB829D328FCF14683FC82" box="[352,436,849,875]" italics="true" pageId="10" pageNumber="311">Female</emphasis>
. Queen body length
<quantity id="3B5B9B5D7F3BB829D0E8FCF1442BFC85" box="[672,796,849,876]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="10" pageNumber="311" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">1517 mm</quantity>
, worker
<quantity id="3B5B9B5D7F3BB829D137FCF144E6FC85" box="[895,977,849,876]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="311" unit="mm" value="13.0">13 mm</quantity>
. Head with the oculo-malar area length / breadth ratio 1.6. Thorax with the mid basitarsus distal posterior corner acutely and spinosely pointed. Hair long, uneven, and black; on the head above the antennal socket the shorter hair yellow; pronotum and anterior scutum either often yellow, forming a transverse band connected with the yellow hypoepimeral area, or this band reduced with black in some queens and most workers; scutellum posteriorly with a yellow fringe, which may be much reduced with black; lateral propodeum yellow; hind tibia with the ends of the corbicular fringes and on the hind basitarsus orange-brown; all leg bases and hind femur with yellow ventrally; metasomal T13 entirely yellow; T4 black, with a narrow medial posterior fringe of rufous orange; T5 black anteriorly and in the posterior half orange; T6 orange.
</paragraph>
<paragraph id="FC1C36B87F3BB829D28FFB354410FA6E" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D28FFB354636FB46" box="[199,257,1173,1199]" italics="true" pageId="10" pageNumber="311">Male</emphasis>
. Body length
<quantity id="3B5B9B5D7F3BB829D3EAFB354529FB59" box="[418,542,1173,1200]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="10" pageNumber="311" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
. Hair long, uneven, and black; mandibular ventral fringe with long curved hair black; face with many mixed short hair yellow; pronotum and anterior scutum either yellow, forming a transverse band connected with the yellow hypoepimeral area, or often this band reduced with black; mesepisternum, hypoepimeral area and metepisternum with yellow; lateral propodeum yellow; hind tibia with dorsal fringe longer than the breadth of the tibia; hind basitarsus with keirotrichia entirely orange; metasomal T13 yellow; T45 posterior fringe often mixed with yellow; T
<quantity id="3B5B9B5D7F3BB829D3CFFAE94686FA8A" box="[391,433,1353,1379]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="311" unit="in" value="6.0">6 in</quantity>
the posterior half with orange; T7 entirely orange; sterna with long pale hairs on posterior fringes but S6 short and orange. Genitalia shown in
<figureCitation id="64982A3D7F3BB829D096FACD4414FA6E" box="[734,803,1389,1415]" captionStart="FIGURES 45" captionStartId="9.[151,275,910,936]" captionTargetBox="[167,1397,190,901]" captionTargetPageId="9" captionText="FIGURES 45. Male genitalia of (4) B. nigrodorsalis and (5) B. weisi from the right lateral aspect, with posterior extremities at the top of the images and dorsal parts to the right of the images." pageId="10" pageNumber="311">Fig. 4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB829D28FFA314219FA26" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="FC1C36B87F3BB829D28FFA314219FA26" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D28FFA31469DFA42" bold="true" box="[199,426,1425,1451]" pageId="10" pageNumber="311">Material examined</emphasis>
.
<materialsCitation id="4CCB3CE57F3BB829D3F0FA31421DFA26" collectionCode="ECOSUR" pageId="10" pageNumber="311" specimenCount="596">
<collectionCode id="9AB2AE7D7F3BB829D3F0FA31451CFA42" box="[440,555,1425,1451]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="10" pageNumber="311" type="Museum">ECOSUR</collectionCode>
:
<specimenCount id="EAA5FD317F3BB829D07EFA3145D7FA42" box="[566,736,1425,1452]" count="596" pageId="10" pageNumber="311" type="generic">596 specimens</specimenCount>
. Data and barcodes for specimens from
<figureCitation id="64982A3D7F3BB829D6ECFA314231FA42" box="[1188,1286,1425,1451]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,260,1590,1616]" captionStartId-1="8.[151,260,1760,1786]" captionTargetBox-0="[151,1435,180,1565]" captionTargetBox-1="[151,1434,181,1733]" captionTargetId-0="figure-30@6.[151,1435,179,1566]" captionTargetId-1="figure-30@8.[151,1435,179,1736]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 2. Support for apparent species gene coalescents from unique haplotypes for COI barcode-like sequences including numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MRBAYES and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intraspecies branching PTP models (species coalescent nodes). Sequence labels include: sequence length in number of bases; taxon name; country in capitals (if not shown then MEXICO) and province; sequence identification code (E# = ECOSUR#). Sequence labels are followed by the percentage of the bases A and T at the third codon position, where low scores (≤97.6%) are likely to indicate numts. Labels for sequences interpreted as likely lower-divergence numts are shown in orange and for likely higher-divergence numts are shown in red (see Fig. 1). The scale bar represents branch length on the tree in substitutions per nucleotide." captionText-1="FIGURE 3. Support for species gene coalescents from unique haplotypes for COI barcode (orthologous) sequences excluding likely numts (orange and red bars and labels in Figs 12), obtained from an evolutionary tree estimated with MRBAYES and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at nodes, treebranch colours, sequence labels, and scale bar as in Fig. 2." figureDoi-0="http://doi.org/10.5281/zenodo.13914596" figureDoi-1="http://doi.org/10.5281/zenodo.13914598" httpUri-0="https://zenodo.org/record/13914596/files/figure.png" httpUri-1="https://zenodo.org/record/13914598/files/figure.png" pageId="10" pageNumber="311">Figs 23</figureCitation>
are available from https://www.ecosur.mx/beesofmesoamerica/ and barcodes are available from a Supplementary File
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4B965337F3BB829D28FFA7947EDF922" pageId="10" pageNumber="311" type="distribution">
<paragraph id="FC1C36B87F3BB829D28FFA7947EDF922" blockId="10.[151,1437,525,1739]" pageId="10" pageNumber="311">
<emphasis id="CED7EAAA7F3BB829D28FFA79466EFA1A" bold="true" box="[199,345,1497,1523]" pageId="10" pageNumber="311">Distribution</emphasis>
.
<taxonomicName id="3BA34D3B7F3BB829D322FA794554FA1A" authorityName="Franklin. Our" authorityYear="1907" box="[362,611,1497,1523]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3BB829D322FA794554FA1A" box="[362,611,1497,1523]" italics="true" pageId="10" pageNumber="311">Bombus nigrodorsalis</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="84B476287F3BB829D147FA794452FA1A" box="[783,869,1497,1523]" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingCountry>
(
<figureCitation id="64982A3D7F3BB829D13EFA79448BFA1A" box="[886,956,1497,1523]" captionStart="FIGURE 6" captionStartId="9.[151,260,1632,1658]" captionTargetBox="[357,1229,1030,1608]" captionTargetId="figure-72@9.[357,1229,1030,1608]" captionTargetPageId="9" captionText="FIGURE 6. Map showing the distributions of samples examined for this study of the weisi-complex: (white spots) taxon nigrodorsalis s. str. (=B. nigrodorsalis stat. rev.); (grey spots) taxon montezumae (=B. weisi stat. rev.); and (black spots) taxon weisi s. str. (=B. weisi stat. rev.). Larger symbols with spots in their centres show locations from which specimens were sequenced for each taxon. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI." figureDoi="http://doi.org/10.5281/zenodo.13914600" httpUri="https://zenodo.org/record/13914600/files/figure.png" pageId="10" pageNumber="311">Fig. 6</figureCitation>
), distributed from
<collectingRegion id="3E67F85A7F3BB829D6DCFA79423AFA1A" box="[1172,1293,1497,1523]" country="Mexico" name="Chihuahua" pageId="10" pageNumber="311">Chihuahua</collectingRegion>
in the north (
<geoCoordinate id="9997507F7F3BB829D2D7FA5D462FF9FE" box="[159,280,1533,1559]" degrees="29.189" direction="north" orientation="latitude" pageId="10" pageNumber="311" precision="55" value="29.189">29.189° N</geoCoordinate>
) to
<collectingRegion id="3E67F85A7F3BB829D30FFA5D46ACF9FE" box="[327,411,1533,1559]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="311">Oaxaca</collectingRegion>
in the south (
<geoCoordinate id="9997507F7F3BB829D073FA5D4582F9FE" box="[571,693,1533,1559]" degrees="16.149" direction="north" orientation="latitude" pageId="10" pageNumber="311" precision="55" value="16.149">16.149° N</geoCoordinate>
). It is absent south of the Tehuantepec Isthmus and from beyond
<collectingCountry id="84B476287F3BB829D2DFF98147C7F9D2" box="[151,240,1569,1595]" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingCountry>
. The altitudinal range is
<quantity id="3B5B9B5D7F3BB829D042F9814591F9D5" box="[522,678,1569,1596]" metricMagnitude="3" metricUnit="m" metricValue="2.8169999999999997" metricValueMax="3.6339999999999995" metricValueMin="2.0" pageId="10" pageNumber="311" unit="m" value="2817.0" valueMax="3634.0" valueMin="2000.0">20003634 m</quantity>
asl, in areas with pine forest. This species has been collected from July to January, with only two queens collected in April and June. Males were collected from late September to mid-January.
<taxonomicName id="3BA34D3B7F3BB829D367F9C94511F96A" authorityName="Franklin. Our" authorityYear="1907" box="[303,550,1641,1667]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3BB829D367F9C94511F96A" box="[303,550,1641,1667]" italics="true" pageId="10" pageNumber="311">Bombus nigrodorsalis</emphasis>
</taxonomicName>
is broadly sympatric with
<taxonomicName id="3BA34D3B7F3BB829D118F9CA449EF96A" authorityName="Williams &amp; Sagot &amp; Martínez-López &amp; Barajas &amp; Mérida-Rivas &amp; Vandame" authorityYear="2024" box="[848,937,1642,1667]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="weisi">
<emphasis id="CED7EAAA7F3BB829D118F9CA449EF96A" box="[848,937,1642,1667]" italics="true" pageId="10" pageNumber="311">B. weisi</emphasis>
</taxonomicName>
in some regions of
<collectingCountry id="84B476287F3BB829D6CDF9C943EAF96A" box="[1157,1245,1641,1667]" name="Mexico" pageId="10" pageNumber="311">Mexico</collectingCountry>
, but within those regions
<taxonomicName id="3BA34D3B7F3BB829D2BBF92E469CF94E" authorityName="Franklin. Our" authorityYear="1907" box="[243,427,1677,1703]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nigrodorsalis">
<emphasis id="CED7EAAA7F3BB829D2BBF92E469CF94E" box="[243,427,1677,1703]" italics="true" pageId="10" pageNumber="311">B. nigrodorsalis</emphasis>
</taxonomicName>
is present at higher elevations and has a geographical range that extends further to the north.
</paragraph>
</subSubSection>
</treatment>
</document>