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<document id="C316ADB00A539C781B3C9DE704EB6A40" ID-DOI="10.11646/zootaxa.3630.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261763" ID-ZooBank="C7529E9C-73F7-4E52-92E7-FCACA8F99A7D" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629846447618" checkinUser="felipe" docAuthor="Liu, Ye, Shi, Hongliang &amp; Liang, Hongbin" docDate="2013" docId="03CF5A00720FFFBEFF17FA83F6C5FD6E" docLanguage="en" docName="zt03630p518.pdf" docOrigin="Zootaxa 3630 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Chlaenioctenus) cheni Liu &amp; Shi &amp; Liang 2013, sp. n." docType="treatment" docVersion="1" lastPageNumber="508" masterDocId="FFF62278720EFFBDFF80FFC0F270FF81" masterDocTitle="Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus" masterLastPageNumber="518" masterPageNumber="505" pageNumber="506" updateTime="1730973406347" updateUser="diego" zenodo-license-document="CLOSED">
<mods:mods id="84AA7D1D3B2AF446FAA3FA293444A5EE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="274D5CEC94F0BA9CF8FBF5B78B5788E9">
<mods:title id="A9D8EAA55F7BB0BB33247996B8F33C89">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
</mods:titleInfo>
<mods:name id="96104E25D1691681642A5DF78F3D4740" type="personal">
<mods:role id="CEC9AC76FDF0A05E31CA066CA66D00BD">
<mods:roleTerm id="60F1EF954820C63EF238ACF389882FDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0CE30528BBE0F015694208F4C3EC6D0">Liu, Ye</mods:namePart>
</mods:name>
<mods:name id="82ECD88A5BED077AAFA8D9785D7030D5" type="personal">
<mods:role id="0E894569B052C715D4D1CA01A5FBCC82">
<mods:roleTerm id="7D674A8419703F8E0F4240CF87D4FC7C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB6877F21D5D1869CDE885FF1D88B006">Shi, Hongliang</mods:namePart>
</mods:name>
<mods:name id="BE5A6EA2AE1A821550B29A472EAC63DF" type="personal">
<mods:role id="3CC0DFADC3213FD5EC03A43DA9E2633C">
<mods:roleTerm id="209F0C62000122F7FF81255219697FF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6185BE2EFA0C54DC3D9E0527736A900F">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="575CB6BF1F7A18E26B77DF3FDB6FBE6D" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="85B389F97A89223E52F9C44966BC2314">text</mods:typeOfResource>
<mods:relatedItem id="559EAC948B6D353B3B5FCD7BB9812763" type="host">
<mods:titleInfo id="F250437E3C248E88DE7375B75F00F15A">
<mods:title id="030BE960817B767F7B33A973133D7089">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0B1630DB94E4798B58BEE39CD0C3AE68">
<mods:date id="CEC22356D2DAFA1CC6A57AFE5995B331">2013</mods:date>
<mods:detail id="3AC35D7E31F633D06A0678CC1E5AA774" type="pubDate">
<mods:number id="F158B3C6F70052BC7115EE66DB52D9FE">2013-03-21</mods:number>
</mods:detail>
<mods:detail id="CF0A7A93C290F4FD18EFC577FEBCF8D1" type="volume">
<mods:number id="6EFFA6D6550C4803CBE072FDF0931B45">3630</mods:number>
</mods:detail>
<mods:detail id="F529FD4A6E1FF630007BCAE79FEE328A" type="issue">
<mods:number id="2ACC6F92FCF244C6EC206F484E571527">3</mods:number>
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<mods:start id="9C26FD4D4974F8AE90C217C28B0A605C">505</mods:start>
<mods:end id="A1A36AD768D6262A871B1AE6A70AAFF6">518</mods:end>
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<mods:classification id="F7502F5A23D2F2550A0EB52D1D10AEB5">journal article</mods:classification>
<mods:identifier id="1E708A22593956F75BAF6E80270118C3" type="DOI">10.11646/zootaxa.3630.3.6</mods:identifier>
<mods:identifier id="DA23E417E24936D64384677B5C5FCA4E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="87A4B54E32404C31FFF4E6041149D7C9" type="Zenodo-Dep">5261763</mods:identifier>
<mods:identifier id="CDE62049FA4C1BD1512FB24950110A7A" type="ZooBank">C7529E9C-73F7-4E52-92E7-FCACA8F99A7D</mods:identifier>
</mods:mods>
<treatment id="03CF5A00720FFFBEFF17FA83F6C5FD6E" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E" httpUri="http://treatment.plazi.org/id/03CF5A00720FFFBEFF17FA83F6C5FD6E" lastPageId="3" lastPageNumber="508" pageId="1" pageNumber="506">
<subSubSection id="C37CB89D720FFFBCFF17FA83F174FADA" box="[151,772,1347,1372]" pageId="1" pageNumber="506" type="nomenclature">
<paragraph id="8BD9EB16720FFFBCFF17FA83F174FADA" blockId="1.[150,772,1347,1407]" box="[151,772,1347,1372]" pageId="1" pageNumber="506">
<taxonomicName id="4C669095720FFFBCFF17FA83F0CEFADD" ID-CoL="69VVJ" authority="Liu &amp; Liang" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,702,1347,1372]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFF17FA83F37DFADD" bold="true" box="[151,269,1347,1372]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFE9DFA83F3A1FADD" bold="true" box="[285,465,1347,1372]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFE62FA83F0CEFADD" bold="true" box="[482,702,1347,1372]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFE62FA83F051FADD" bold="true" box="[482,545,1347,1372]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
Liu &amp; Liang
</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFD45FA84F174FADA" bold="true" box="[709,772,1348,1371]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFD45FA84F174FADA" box="[709,772,1348,1371]" pageId="1" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBCFF16FAA7F07FFAFE" box="[150,527,1383,1407]" pageId="1" pageNumber="506" type="description">
<paragraph id="8BD9EB16720FFFBCFF16FAA7F07FFAFE" blockId="1.[150,772,1347,1407]" box="[150,527,1383,1407]" pageId="1" pageNumber="506">
(
<figureCitation id="135DF793720FFFBCFF1FFAA7F37CFAFE" box="[159,268,1383,1407]" captionStart="FIGURES 1–4" captionStartId="2.[151,263,1909,1932]" captionTargetBox="[224,1309,194,1888]" captionTargetId="figure-51@2.[151,1436,193,1888]" captionTargetPageId="2" captionText="FIGURES 1–4. Chlaenius cheni sp. n. 1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="1" pageNumber="506">Figs. 1−4</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE98FAA7F347FAFE" box="[280,311,1383,1407]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">20</figureCitation>
,
<figureCitation id="135DF793720FFFBCFEC2FAA7F310FAFE" box="[322,352,1383,1407]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="1" pageNumber="506">26</figureCitation>
,
<figureCitation id="135DF793720FFFBCFEECFAA7F3FAFAFE" box="[364,394,1383,1407]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="1" pageNumber="506">32</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE16FAA7F3AEFAFE" box="[406,478,1383,1407]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="1" pageNumber="506">38, 39</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE6AFAA7F077FAFE" box="[490,519,1383,1407]" captionStart="FIGURES 48–51" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 48–51. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="1" pageNumber="506">48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBCFF17FA6FF1BFF91E" pageId="1" pageNumber="506" type="materials_examined">
<paragraph id="8BD9EB16720FFFBCFF17FA6FF1BFF91E" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF17FA6FF005FA46" box="[151,629,1455,1480]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">
<emphasis id="B9123704720FFFBCFF17FA6FF3F5FA49" bold="true" box="[151,389,1455,1480]" pageId="1" pageNumber="506">Specimen examined</emphasis>
(
<emphasis id="B9123704720FFFBCFE17FA70F015FA49" bold="true" box="[407,613,1455,1480]" pageId="1" pageNumber="506">
total
<specimenCount id="9D60209F720FFFBCFE56FA6FF015FA49" box="[470,613,1455,1480]" pageId="1" pageNumber="506" type="generic">3 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720FFFBCFDFEFA6FF3DFF98E" collectingDate="1989-03-21" collectionCode="SCAU" collectorName="Chen Shoujian" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Guangdong Prov." typeStatus="holotype">
<typeStatus id="54DD55B4720FFFBCFDFEFA6FF09BFA46" box="[638,747,1455,1479]" pageId="1" pageNumber="506" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60209F720FFFBCFD76FA6FF15CFA46" box="[758,812,1455,1479]" pageId="1" pageNumber="506" type="male">male</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFCBDFA6FF1FDFA46" box="[829,909,1455,1479]" pageId="1" pageNumber="506">SCAU</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFC2CFA6FF184FA46" box="[940,1012,1455,1479]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFB80FA6FF6B8FA46" box="[1024,1224,1455,1479]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFB54FA6FF285FA6D" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFB54FA6FF285FA6D" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
,
<date id="FFD8CDD6720FFFBCFF79FA14F31BFA6D" box="[249,363,1492,1516]" pageId="1" pageNumber="506" value="1989-03-21">
<collectingDate id="EF9C343E720FFFBCFF79FA14F31BFA6D" box="[249,363,1492,1516]" pageId="1" pageNumber="506" value="1989-03-21">1989.3.21</collectingDate>
</date>
,
<collectorName id="26938EC0720FFFBCFEF7FA14F06EFA6D" box="[375,542,1492,1516]" pageId="1" pageNumber="506">Chen Shoujian</collectorName>
leg.’’ / “HOLOTYPE,
<taxonomicName id="4C669095720FFFBCFCA7FA14F6D6FA6D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[807,1190,1492,1516]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFCA7FA14F1E8FA6D" box="[807,920,1492,1516]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFC2BFA14F629FA6D" box="[939,1113,1492,1516]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFBEAFA14F6D6FA6D" box="[1130,1190,1492,1516]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFB2FFA14F69FFA6D" bold="true" box="[1199,1263,1492,1516]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFB2FFA14F69FFA6D" box="[1199,1263,1492,1516]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [red label]
</materialsCitation>
.
<materialsCitation id="3B0EE14B720FFFBCFE3DFA38F62DF9D6" collectingDate="1989-03-21" collectionCode="SCAU" collectorName="Chen Shoujian" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong Prov." typeStatus="paratype">
<typeStatus id="54DD55B4720FFFBCFE3DFA38F05AF991" box="[445,554,1528,1552]" pageId="1" pageNumber="506" type="paratype">Paratypes</typeStatus>
(total 2 specimens):
<specimenCount id="9D60209F720FFFBCFC9DFA37F1F0F98E" box="[797,896,1527,1551]" pageId="1" pageNumber="506" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFC11FA37F191F98E" box="[913,993,1527,1551]" pageId="1" pageNumber="506">SCAU</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFB80FA37F638F98E" box="[1024,1096,1527,1551]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFBD5FA37F76EF98E" box="[1109,1310,1527,1551]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFAABFA37F339F9B2" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFAABFA37F339F9B2" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
,
<date id="FFD8CDD6720FFFBCFED6F9DCF3B9F9B5" box="[342,457,1564,1588]" pageId="1" pageNumber="506" value="1989-03-21">
<collectingDate id="EF9C343E720FFFBCFED6F9DCF3B9F9B5" box="[342,457,1564,1588]" pageId="1" pageNumber="506" value="1989-03-21">1989.3.21</collectingDate>
</date>
,
<collectorName id="26938EC0720FFFBCFE56F9DCF00DF9B5" box="[470,637,1564,1588]" pageId="1" pageNumber="506">Chen Shoujian</collectorName>
leg.’’ / “
<taxonomicName id="4C669095720FFFBCFD66F9DCF1BCF9B5" authorityName="Motschulsky" authorityYear="1854" box="[742,972,1564,1588]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="posticalis">
<emphasis id="B9123704720FFFBCFD66F9DCF1BCF9B5" box="[742,972,1564,1588]" italics="true" pageId="1" pageNumber="506">Chlaenius posticalis</emphasis>
</taxonomicName>
M. det. Chen Shoujian” / “PARATYPE,
<taxonomicName id="4C669095720FFFBCFF17F980F067F9D9" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,535,1599,1624]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFF17F980F378F9D9" box="[151,264,1600,1624]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFE9BF980F3B9F9D9" box="[283,457,1600,1624]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFE5BF980F067F9D9" box="[475,535,1600,1624]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFDA2F980F011F9D9" bold="true" box="[546,609,1600,1624]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFDA2F980F011F9D9" box="[546,609,1600,1624]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label];
</materialsCitation>
<materialsCitation id="3B0EE14B720FFFBCFBE6F9FFF1BFF91E" collectionCode="IZCAS" collectorName="An Shuwen" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong Prov." typeStatus="paratype">
<specimenCount id="9D60209F720FFFBCFBE6F9FFF6B9F9D6" box="[1126,1225,1599,1623]" pageId="1" pageNumber="506" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFB5AF9FFF740F9D6" box="[1242,1328,1599,1623]" pageId="1" pageNumber="506">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFAD0F9FFF7E8F9D6" box="[1360,1432,1599,1623]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFF17F9A4F31EF9FA" box="[151,366,1636,1660]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFE09F9A4F0A3F9FA" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFE09F9A4F0A3F9FA" box="[393,723,1636,1660]" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
, 2003.7,
<collectorName id="26938EC0720FFFBCFCDAF9A4F19DF9FD" box="[858,1005,1636,1660]" pageId="1" pageNumber="506">An Shuwen</collectorName>
leg.’’ / “PARATYPE,
<taxonomicName id="4C669095720FFFBCFAAAF9A4F3EBF921" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFAAAF9A4F7EBF9FD" box="[1322,1435,1636,1660]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFF20F948F33EF921" box="[160,334,1672,1696]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFEDFF948F3EBF921" box="[351,411,1672,1696]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFE22F948F391F921" bold="true" box="[418,481,1672,1696]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFE22F948F391F921" box="[418,481,1672,1696]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBEFF49F96CF1E5FE4E" lastPageId="3" lastPageNumber="508" pageId="1" pageNumber="506" type="description">
<paragraph id="8BD9EB16720FFFBCFF49F96CF040F8AE" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF49F96CF30FF942" box="[201,383,1708,1732]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">
<emphasis id="B9123704720FFFBCFF49F96CF308F945" bold="true" box="[201,376,1708,1732]" pageId="1" pageNumber="506">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F3720FFFBCFE4EF96CF004F942" box="[462,628,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6199999999999999" metricValueMin="1.48" pageId="1" pageNumber="506" unit="mm" value="15.5" valueMax="16.2" valueMin="14.8">14.8−16.2 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFD65F96CF13FF945" box="[741,847,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="506" unit="mm" value="15.4">15.4 mm</quantity>
), HW =
<quantity id="4C9E46F3720FFFBCFC35F96CF64FF942" box="[949,1087,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="1" pageNumber="506" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.6−2.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFB30F96CF77CF945" box="[1200,1292,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="506" unit="mm" value="2.6">2.6 mm</quantity>
), EYL = 1.0 mm, PL =
<quantity id="4C9E46F3720FFFBCFE8EF90FF3E5F966" box="[270,405,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.4" metricValueMin="3.3" pageId="1" pageNumber="506" unit="mm" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.3−3.4 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFD80F90FF02AF966" box="[512,602,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="506" unit="mm" value="3.4">3.4 mm</quantity>
), PW =
<quantity id="4C9E46F3720FFFBCFD34F90FF14BF966" box="[692,827,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="1" pageNumber="506" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5−3.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFC26F90FF189F966" box="[934,1017,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="506" unit="mm" value="3.6">3.6mm</quantity>
), EL =
<quantity id="4C9E46F3720FFFBCFBCBF90FF6A3F966" box="[1099,1235,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.5" metricValueMin="8.7" pageId="1" pageNumber="506" unit="mm" value="9.1" valueMax="9.5" valueMin="8.7">8.7−9.5 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFABDF90FF7E0F966" box="[1341,1424,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="506" unit="mm" value="9.1">9.1mm</quantity>
), EW =
<quantity id="4C9E46F3720FFFBCFF5FF934F317F88A" box="[223,359,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="5.7" metricValueMin="5.2" pageId="1" pageNumber="506" unit="mm" value="5.45" valueMax="5.7" valueMin="5.2">5.2−5.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFE52F934F05BF88D" box="[466,555,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="506" unit="mm" value="5.5">5.5 mm</quantity>
), ratio PW/PL = 1.06−1.09 (mean = 1.07), EL/EW = 1.58−1.70 (mean = 1.65), EW/PW = 1.49−1.58 (mean = 1.53).
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF49F8FBF6E2F81A" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF49F8FBF32AF8D5" bold="true" box="[201,346,1851,1876]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">Description.</emphasis>
Head and pronotum black, with green metallic shine; elytra black, with a round yellow spot at posterior third of each elytron; ventral surface, coxae, trochanters, and tibiae black; antennae brown; palpi, tarsi reddish brown or dark brown; femora yellow in the middle portion, black at base and apex.
</paragraph>
<caption id="DF19BB9E720CFFBFFF17F8B5F14DF82B" ID-DOI="http://doi.org/10.5281/zenodo.5261767" ID-Zenodo-Dep="5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="2" pageNumber="507" startId="2.[151,263,1909,1932]" targetBox="[224,1309,194,1888]" targetPageId="2">
<paragraph id="8BD9EB16720CFFBFFF17F8B5F14DF82B" blockId="2.[151,1436,1909,1962]" pageId="2" pageNumber="507">
<emphasis id="B9123704720CFFBFFF17F8B5F34BF80D" bold="true" box="[151,315,1909,1932]" pageId="2" pageNumber="507">FIGURES 1–4.</emphasis>
<taxonomicName id="4C669095720CFFBFFEC3F8B6F399F80D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[323,489,1910,1932]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov.">
<emphasis id="B9123704720CFFBFFEC3F8B6F399F80D" box="[323,489,1910,1932]" italics="true" pageId="2" pageNumber="507">Chlaenius cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720CFFBFFE70F8B6F05BF80A" bold="true" box="[496,555,1910,1931]" pageId="2" pageNumber="507">
<taxonomicNameLabel id="A2218A7F720CFFBFFE70F8B6F05BF80A" box="[496,555,1910,1931]" pageId="2" pageNumber="507" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm.
</paragraph>
</caption>
<paragraph id="8BD9EB16720DFFBEFF49FF57F1E5FE4E" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
Head with vertex convex, finely punctate; occiput coarsely punctate and sparsely pubescent; genae glabrous, oblique, as long as one-fifth of eyes in dorsal view; lateral margins of mentum rounded. Pronotum (
<figureCitation id="135DF793720DFFBEFB74FF7CF736FF55" box="[1268,1350,188,212]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">Fig. 20</figureCitation>
) ovoid, widest in the middle; lateral margins distinctly bordered, sinuate before posterior angles; disc finely punctate, pubescent, median line distinct, shallow; posterior angles obtuse (&gt;90 degrees), rounded at apex; base rugosepunctate, pubescent; basal foveae deep, coarsely punctate. Elytra moderately convex, widest near posterior third; intervals (
<figureCitation id="135DF793720DFFBEFE8CFE8CF313FEE5" box="[268,355,332,356]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">Fig. 26</figureCitation>
) convex, densely punctate and pubescent; yellow spot covering intervals 4−8. Proepisterna sparsely punctate. Hind femora with basal black areas slightly shorter than hind trochanters, claws (
<figureCitation id="135DF793720DFFBEFB69FEAFF74AFE06" box="[1257,1338,367,391]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">Fig. 32</figureCitation>
) curved, with 6−7 short teeth. Aedeagus (
<figureCitation id="135DF793720DFFBEFD90FE54F0EBFE2D" box="[528,667,404,428]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 38, 39</figureCitation>
) thin in lateral view, apical lamella rounded and short. Gonostylus (
<figureCitation id="135DF793720DFFBEFF1FFE77F281FE4E" box="[159,241,439,463]" captionStart="FIGURES 48–51" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 48–51. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 48</figureCitation>
) of female ovipositor with five long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FE1CF7FAFE75" box="[201,1418,476,500]" pageId="3" pageNumber="508" type="etymology">
<paragraph id="8BD9EB16720DFFBEFF49FE1CF7FAFE75" blockId="3.[151,1436,151,752]" box="[201,1418,476,500]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FE1CF33FFE72" box="[201,335,476,500]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FE1CF338FE75" bold="true" box="[201,328,476,500]" pageId="3" pageNumber="508">Etymology</emphasis>
.
</emphasis>
Latinized name
<taxonomicName id="4C669095720DFFBEFD8AFE1CF037FE75" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[522,583,476,500]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFD8AFE1CF037FE75" box="[522,583,476,500]" italics="true" pageId="3" pageNumber="508">cheni</emphasis>
</taxonomicName>
refers to Professor Chen Shoujian, a respected carabid specialist in
<collectingCountry id="F371AB86720DFFBEFAC0FE1CF7F6FE75" box="[1344,1414,476,500]" name="China" pageId="3" pageNumber="508">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FDC0F6C5FD6E" pageId="3" pageNumber="508" type="discussion">
<paragraph id="8BD9EB16720DFFBEFF49FDC0F35CFD02" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FDC0F34CFD96" box="[201,316,512,536]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FDC0F346FD99" bold="true" box="[201,310,512,536]" pageId="3" pageNumber="508">Remarks</emphasis>
.
</emphasis>
<taxonomicName id="4C669095720DFFBEFEC7FDC0F070FD99" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[327,512,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFEC7FDC0F070FD99" box="[327,512,512,536]" italics="true" pageId="3" pageNumber="508">Chlaenius cheni</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C669095720DFFBEFD1FFDC0F164FD99" authorityName="Kirschenhofer" authorityYear="2008" box="[671,788,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B9123704720DFFBEFD1FFDC0F164FD99" box="[671,788,512,536]" italics="true" pageId="3" pageNumber="508">C. sehnali</emphasis>
</taxonomicName>
in having yellow spot on posterior third of elytron and femora yellow in the middle portion, but differs from the latter in having pronotal disc finely punctate and elytral intervals regularly pubescent in contrast to pronotal disc coarsely punctate and elytral intervals glabrous medially in
<taxonomicName id="4C669095720DFFBEFF34FDACF355FD05" authorityName="Kirschenhofer" authorityYear="2008" box="[180,293,620,644]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B9123704720DFFBEFF34FDACF355FD05" box="[180,293,620,644]" italics="true" pageId="3" pageNumber="508">C. sehnali</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FD50F6C5FD6E" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
<taxonomicName id="4C669095720DFFBEFF49FD50F36EFD29" authorityName="Jedlicka" authorityYear="1960" box="[201,286,656,680]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFF49FD50F36EFD29" box="[201,286,656,680]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
is the nearest species with
<taxonomicName id="4C669095720DFFBEFDDEFD50F0CFFD29" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[606,703,656,680]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFDDEFD50F0CFFD29" box="[606,703,656,680]" italics="true" pageId="3" pageNumber="508">C. cheni</emphasis>
</taxonomicName>
in distribution (
<figureCitation id="135DF793720DFFBEFCFAFD4FF1A0FD26" box="[890,976,655,679]" captionStart="FIGURE 52" captionStartId="12.[151,250,1844,1867]" captionTargetBox="[151,1436,935,1822]" captionTargetId="figure-69@12.[151,1436,935,1822]" captionTargetPageId="12" captionText="FIGURE 52. Geographical distribution of Chlaenius (Chlaenioctenus) species." figureDoi="http://doi.org/10.5281/zenodo.5261797" httpUri="https://zenodo.org/record/5261797/files/figure.png" pageId="3" pageNumber="508">Fig. 52</figureCitation>
), and their position of yellow spots and interval pubescence on elytra are also similar. However, the new species differs from
<taxonomicName id="4C669095720DFFBEFBE3FD74F6C7FD4D" authorityName="Jedlicka" authorityYear="1960" box="[1123,1207,692,716]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFBE3FD74F606FD4D" box="[1123,1142,692,716]" italics="true" pageId="3" pageNumber="508">C</emphasis>
.
<emphasis id="B9123704720DFFBEFB05FD74F6C7FD4D" box="[1157,1207,692,716]" italics="true" pageId="3" pageNumber="508">freyi</emphasis>
</taxonomicName>
in having bicolored femora (wholly black in
<taxonomicName id="4C669095720DFFBEFE2CFD18F38EFD71" authorityName="Jedlicka" authorityYear="1960" box="[428,510,728,752]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFE2CFD18F38EFD71" box="[428,510,728,752]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
), and long gonostylus spines (
<figureCitation id="135DF793720DFFBEFCD4FD17F1D5FD6E" box="[852,933,727,751]" captionStart="FIGURES 48–51" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 48–51. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 48</figureCitation>
) (very short in
<taxonomicName id="4C669095720DFFBEFBD2FD18F6D5FD71" authorityName="Jedlicka" authorityYear="1960" box="[1106,1189,728,752]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFBD2FD18F6D5FD71" box="[1106,1189,728,752]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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