treatments-xml/data/03/CF/5A/03CF5A00720DFFBEFF17FC83F1FEF85D.xml

<document id="63E91D6D84DE29F496E4585EC129C0CC" ID-DOI="10.11646/zootaxa.3630.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261763" ID-ZooBank="C7529E9C-73F7-4E52-92E7-FCACA8F99A7D" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629846447618" checkinUser="felipe" docAuthor="Liu, Ye, Shi, Hongliang &amp; Liang, Hongbin" docDate="2013" docId="03CF5A00720DFFBEFF17FC83F1FEF85D" docLanguage="en" docName="zt03630p518.pdf" docOrigin="Zootaxa 3630 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Chlaenioctenus) bodhidharma Liu &amp; Shi &amp; Liang 2013, sp. n." docType="treatment" docVersion="1" lastPageNumber="508" masterDocId="FFF62278720EFFBDFF80FFC0F270FF81" masterDocTitle="Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus" masterLastPageNumber="518" masterPageNumber="505" pageNumber="508" updateTime="1730973406347" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="9D571337E0115D9B2BBA849E0E7FDB1F">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:name id="A89B8AEE68D7A0AD55B9C6E33378F40E" type="personal">
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<mods:namePart id="64B58B08F5B519001F25D79F616B8A23">Liu, Ye</mods:namePart>
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<mods:namePart id="A4B931F6E3A8C2A7AE6FE68E07580C65">Shi, Hongliang</mods:namePart>
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<mods:namePart id="501D1ACD7E3C389BDA49A088880C18D8">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="A05E257E18054389A1E7F626B50CDDA3" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="494ED20ACCD94312377A23920A53A84E">Zootaxa</mods:title>
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<mods:date id="DDB16C82E1918F05A48532AAE2CAFC95">2013</mods:date>
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<mods:number id="A2F5EFD6900EB450D8F7335EA22510A5">2013-03-21</mods:number>
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<mods:number id="227FF642971D6258803159691491594C">3630</mods:number>
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</mods:mods>
<treatment id="03CF5A00720DFFBEFF17FC83F1FEF85D" LSID="urn:lsid:plazi:treatment:03CF5A00720DFFBEFF17FC83F1FEF85D" httpUri="http://treatment.plazi.org/id/03CF5A00720DFFBEFF17FC83F1FEF85D" lastPageNumber="508" pageId="3" pageNumber="508">
<subSubSection id="C37CB89D720DFFBEFF17FC83F111FCDD" box="[151,865,835,860]" pageId="3" pageNumber="508" type="nomenclature">
<paragraph id="8BD9EB16720DFFBEFF17FC83F111FCDD" blockId="3.[151,865,835,895]" box="[151,865,835,860]" pageId="3" pageNumber="508">
<taxonomicName id="4C669095720DFFBEFF17FC83F16AFCDD" ID-CoL="5Y87J" authority="Liu &amp; Liang" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,794,835,860]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFF17FC83F37DFCDD" bold="true" box="[151,269,835,860]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFE9DFC83F3A1FCDD" bold="true" box="[285,465,835,860]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
) 
<emphasis id="B9123704720DFFBEFE62FC83F16AFCDD" bold="true" box="[482,794,835,860]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFE62FC83F00DFCDD" bold="true" box="[482,637,835,860]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
Liu &amp; Liang
</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFCA1FC84F111FCDD" bold="true" box="[801,865,836,860]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFCA1FC84F111FCDD" box="[801,865,836,860]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF17FCA7F07FFCFE" box="[151,527,871,895]" pageId="3" pageNumber="508" type="description">
<paragraph id="8BD9EB16720DFFBEFF17FCA7F07FFCFE" blockId="3.[151,865,835,895]" box="[151,527,871,895]" pageId="3" pageNumber="508">
(
<figureCitation id="135DF793720DFFBEFF1FFCA7F37DFCFE" box="[159,269,871,895]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 5–8. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="3" pageNumber="508">Figs. 5−8</figureCitation>
, 
<figureCitation id="135DF793720DFFBEFE98FCA7F346FCFE" box="[280,310,871,895]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">21</figureCitation>
, 
<figureCitation id="135DF793720DFFBEFEC2FCA7F310FCFE" box="[322,352,871,895]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">27</figureCitation>
, 
<figureCitation id="135DF793720DFFBEFEECFCA7F3FAFCFE" box="[364,394,871,895]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">33</figureCitation>
, 
<figureCitation id="135DF793720DFFBEFE16FCA7F3AEFCFE" box="[406,478,871,895]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">40, 41</figureCitation>
, 
<figureCitation id="135DF793720DFFBEFE6AFCA7F077FCFE" box="[490,519,871,895]" captionStart="FIGURES 48–51" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 48–51. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF17FC6FF382FB1E" pageId="3" pageNumber="508" type="materials_examined">
<paragraph id="8BD9EB16720DFFBEFF17FC6FF382FB1E" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF17FC6FF0F4FC46" box="[151,644,943,968]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF17FC6FF3F9FC49" bold="true" box="[151,393,943,968]" pageId="3" pageNumber="508">Specimen examined</emphasis>
(
<emphasis id="B9123704720DFFBEFE1EFC70F004FC49" bold="true" box="[414,628,943,968]" pageId="3" pageNumber="508">
total 
<specimenCount id="9D60209F720DFFBEFE61FC6FF004FC49" box="[481,628,943,968]" pageId="3" pageNumber="508" type="generic">3 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720DFFBEFD10FC6FF3EDFC6D" collectingDate="1982-04-25" collectorName="Ch. J. Rai'" elevation="2900" location="Alghera" pageId="3" pageNumber="508" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DD55B4720DFFBEFD10FC6FF08DFC46" box="[656,765,943,967]" pageId="3" pageNumber="508" type="holotype">Holotype</typeStatus>
: male (NHMB), “Darjeeling D., 
<collectorName id="26938EC0720DFFBEFB05FC6FF77AFC46" box="[1157,1290,943,967]" pageId="3" pageNumber="508">Ch. J. Rai’</collectorName>
’ / “ 
<location id="8EB9BDCD720DFFBEFAB9FC6FF7E8FC46" LSID="urn:lsid:plazi:treatment:03CF5A00720DFFBEFF17FC83F1FEF85D:8EB9BDCD720DFFBEFAB9FC6FF7E8FC46" box="[1337,1432,943,967]" name="Alghera" pageId="3" pageNumber="508">Alghera</location>
, 
<quantity id="4C9E46F3720DFFBEFF17FC14F298FC6D" box="[151,232,980,1004]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">
<elevation id="004B0C25720DFFBEFF17FC14F298FC6D" box="[151,232,980,1004]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">2900m</elevation>
</quantity>
, 
<date id="FFD8CDD6720DFFBEFF72FC14F303FC6D" box="[242,371,980,1004]" pageId="3" pageNumber="508" value="1982-04-25">
<collectingDate id="EF9C343E720DFFBEFF72FC14F303FC6D" box="[242,371,980,1004]" pageId="3" pageNumber="508" value="1982-04-25">25.IV.1982</collectingDate>
</date>
’’ / “
</materialsCitation>
<typeStatus id="54DD55B4720DFFBEFE1DFC14F046FC6D" box="[413,566,980,1004]" pageId="3" pageNumber="508" type="holotype">HOLOTYPE</typeStatus>
, 
<taxonomicName id="4C669095720DFFBEFDC1FC14F667FC6D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[577,1047,980,1004]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFDC1FC14F0C3FC6D" box="[577,691,980,1004]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFD42FC14F100FC6D" box="[706,880,980,1004]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
) 
<emphasis id="B9123704720DFFBEFC00FC14F667FC6D" box="[896,1047,980,1004]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFB9EFC14F62CFC6D" bold="true" box="[1054,1116,980,1004]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFB9EFC14F62CFC6D" box="[1054,1116,980,1004]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [red label]. 
<materialsCitation id="3B0EE14B720DFFBEFEA3FC38F333FBB5" collectingDate="1982-04-25" elevation="2900" pageId="3" pageNumber="508" specimenCount="3" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54DD55B4720DFFBEFEA3FC38F3E0FB91" box="[291,400,1016,1040]" pageId="3" pageNumber="508" type="paratype">Paratypes</typeStatus>
(total 
<specimenCount id="9D60209F720DFFBEFE63FC37F008FB8E" box="[483,632,1015,1039]" pageId="3" pageNumber="508" type="generic">2 specimens</specimenCount>
): 
<specimenCount id="9D60209F720DFFBEFD0FFC37F090FB8E" box="[655,736,1015,1039]" pageId="3" pageNumber="508" type="male">1 male</specimenCount>
(NHMB), “Darjeeling D., Ch. J. Rai’’ / “Alghera, 
<quantity id="4C9E46F3720DFFBEFAC7FC37F7E8FB8E" box="[1351,1432,1015,1039]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">
<elevation id="004B0C25720DFFBEFAC7FC37F7E8FB8E" box="[1351,1432,1015,1039]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">2900m</elevation>
</quantity>
, 
<date id="FFD8CDD6720DFFBEFF17FBDCF368FBB5" box="[151,280,1052,1076]" pageId="3" pageNumber="508" value="1982-04-25">
<collectingDate id="EF9C343E720DFFBEFF17FBDCF368FBB5" box="[151,280,1052,1076]" pageId="3" pageNumber="508" value="1982-04-25">25.IV.1982</collectingDate>
</date>
’’ / “
</materialsCitation>
<typeStatus id="54DD55B4720DFFBEFEC3FBDCF3A4FBB5" box="[323,468,1052,1076]" pageId="3" pageNumber="508" type="paratype">PARATYPE</typeStatus>
, 
<taxonomicName id="4C669095720DFFBEFE5FFBDCF1C6FBB5" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[479,950,1052,1076]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFE5FFBDCF021FBB5" box="[479,593,1052,1076]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFDE1FBDCF17FFBB5" box="[609,783,1052,1076]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
) 
<emphasis id="B9123704720DFFBEFC9FFBDCF1C6FBB5" box="[799,950,1052,1076]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFC3DFBDCF18CFBB5" bold="true" box="[957,1020,1052,1076]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFC3DFBDCF18CFBB5" box="[957,1020,1052,1076]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by 
<collectorName id="26938EC0720DFFBEFBE1FBDCF6F8FBB5" box="[1121,1160,1052,1076]" pageId="3" pageNumber="508">Liu</collectorName>
&amp; 
<collectorName id="26938EC0720DFFBEFB2BFBDCF69FFBB5" box="[1195,1263,1052,1076]" pageId="3" pageNumber="508">Liang</collectorName>
, 2012” [yellow label]; 
<materialsCitation id="3B0EE14B720DFFBEFF67FBFFF7EBFBD6" box="[231,1435,1087,1111]" elevation="1800.0" pageId="3" pageNumber="508">
<specimenCount id="9D60209F720DFFBEFF67FBFFF33AFBD6" box="[231,330,1087,1111]" pageId="3" pageNumber="508" type="female">1 female</specimenCount>
(NHMB), “ 
<collectingCountry id="F371AB86720DFFBEFE53FBFFF06FFBD6" box="[467,543,1087,1111]" name="India" pageId="3" pageNumber="508">Indien</collectingCountry>
, Darjeeling D., B. Bhakta’’ / “Sukhay Simana, 
<quantity id="4C9E46F3720DFFBEFBC0FBFFF6A8FBD6" box="[1088,1240,1087,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="3" pageNumber="508" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">
<elevation id="004B0C25720DFFBEFBC0FBFFF6A8FBD6" box="[1088,1240,1087,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="3" pageNumber="508" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">1600−2000m</elevation>
</quantity>
, 31.IV−11.V.86’’
</materialsCitation>
[date 31.IV is wrong, possibly 30.IV] / “ 
<typeStatus id="54DD55B4720DFFBEFDD9FBA4F09AFBFD" box="[601,746,1124,1148]" pageId="3" pageNumber="508" type="paratype">PARATYPE</typeStatus>
, 
<taxonomicName id="4C669095720DFFBEFD74FBA4F6BBFBFD" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[756,1227,1124,1148]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFD74FBA4F115FBFD" box="[756,869,1124,1148]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFCF6FBA4F654FBFD" box="[886,1060,1124,1148]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
) 
<emphasis id="B9123704720DFFBEFBB4FBA4F6BBFBFD" box="[1076,1227,1124,1148]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFB52FBA4F761FBFD" bold="true" box="[1234,1297,1124,1148]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFB52FBA4F761FBFD" box="[1234,1297,1124,1148]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label].
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FB6CF74AF885" pageId="3" pageNumber="508" type="description">
<paragraph id="8BD9EB16720DFFBEFF49FB6CF008FAAE" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FB6CF30FFB42" box="[201,383,1196,1220]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FB6CF308FB45" bold="true" box="[201,376,1196,1220]" pageId="3" pageNumber="508">Measurements</emphasis>
.
</emphasis>
BL = 
<quantity id="4C9E46F3720DFFBEFE4EFB6CF004FB42" box="[462,628,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="1.69" metricValueMin="1.53" pageId="3" pageNumber="508" unit="mm" value="16.1" valueMax="16.9" valueMin="15.3">15.3−16.9 mm</quantity>
(mean = 
<quantity id="4C9E46F3720DFFBEFD65FB6CF13FFB45" box="[741,847,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="3" pageNumber="508" unit="mm" value="15.9">15.9 mm</quantity>
), HW = 
<quantity id="4C9E46F3720DFFBEFC35FB6CF64FFB42" box="[949,1087,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="3" pageNumber="508" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4−2.9 mm</quantity>
(mean = 
<quantity id="4C9E46F3720DFFBEFB30FB6CF77CFB45" box="[1200,1292,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="508" unit="mm" value="2.7">2.7 mm</quantity>
), EYL = 1.0 mm, PL = 
<quantity id="4C9E46F3720DFFBEFE95FB0FF3EFFB66" box="[277,415,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="3" pageNumber="508" unit="mm" value="3.45" valueMax="3.6" valueMin="3.3">3.3−3.6 mm</quantity>
(mean = 
<quantity id="4C9E46F3720DFFBEFD90FB0FF01CFB66" box="[528,620,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="508" unit="mm" value="3.4">3.4 mm</quantity>
), PW = 
<quantity id="4C9E46F3720DFFBEFD4DFB0FF128FB66" box="[717,856,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="3" pageNumber="508" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.7−4.3 mm</quantity>
(mean = 4.0 mm), EL = 
<quantity id="4C9E46F3720DFFBEFBFDFB0FF778FB66" box="[1149,1288,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="9.149999999999999" metricValueMax="9.6" metricValueMin="8.7" pageId="3" pageNumber="508" unit="mm" value="9.149999999999999" valueMax="9.6" valueMin="8.7">8.7−9.6 mm</quantity>
(mean = 9.0 mm), EW = 
<quantity id="4C9E46F3720DFFBEFEA3FB34F3DBFA8A" box="[291,427,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.1" metricValueMin="5.4" pageId="3" pageNumber="508" unit="mm" value="5.75" valueMax="6.1" valueMin="5.4">5.4−6.1 mm</quantity>
(mean = 
<quantity id="4C9E46F3720DFFBEFD96FB34F000FA8D" box="[534,624,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="508" unit="mm" value="5.6">5.6 mm</quantity>
), ratio PW/PL = 1.12−1.19 (mean = 1.15), EL/EW = 1.56−1.63 (mean = 1.60), EW/PW = 1.38−1.46 (mean = 1.42).
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FAFBF36DFA3E" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FAFBF32AFAD5" bold="true" box="[201,346,1339,1364]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">Description.</emphasis>
Head and pronotum black, pronotum with green metallic shine laterally; elytra black, with a yellow spot in the middle of each elytron; ventral surface, coxae, trochanters, tibiae black, femora yellow in the middle, black at base and apex (less so in female, 
<figureCitation id="135DF793720DFFBEFD42FA44F174FA1D" box="[706,772,1412,1436]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 5–8. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="3" pageNumber="508">Fig. 8</figureCitation>
); antennomeres yellow or reddish brown; mandibles, palpi, tarsi brown.
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FA0CF74AF885" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
Head convex, vertex and occiput finely punctate; genae glabrous, oblique, as long as one-fifth of eyes in dorsal view; lateral margins of mentum nearly straight. Pronotum (
<figureCitation id="135DF793720DFFBEFCC3FA2FF1E5F986" box="[835,917,1519,1543]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">Fig. 21</figureCitation>
) moderately ovoid, widest slightly before the middle; lateral margins distinctly bordered, slightly sinuate before posterior angles; disc finely punctate, glabrous, median line distinct; posterior angles rounded; base punctate; basal foveae deep, coarsely punctate. Elytra moderately convex, widest slightly behind the middle; intervals 2−8 convex, glabrous medially, with a row of punctures laterally (near striae) (
<figureCitation id="135DF793720DFFBEFD91F9BFF018F916" box="[529,616,1663,1687]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">Fig. 27</figureCitation>
), intervals 1 and 9 irregularly pubescent; yellow spot covering half of interval 4 to interval 8. Proepisterna sparsely punctate. Hind femora with basal black areas slightly longer than hind trochanters; claws (
<figureCitation id="135DF793720DFFBEFEF8F908F3BCF961" box="[376,460,1736,1760]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">Fig. 33</figureCitation>
) curved, with 7−9 long teeth. Aedeagus (
<figureCitation id="135DF793720DFFBEFC33F908F630F961" box="[947,1088,1736,1760]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 40, 41</figureCitation>
) robust in lateral view, apical lamella narrow and long. Gonostylus (
<figureCitation id="135DF793720DFFBEFDC5F92CF0E6F885" box="[581,662,1772,1796]" captionStart="FIGURES 48–51" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 48–51. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 49</figureCitation>
) of female ovipositor with five long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49F8D0F6B7F8A9" box="[201,1223,1808,1832]" pageId="3" pageNumber="508" type="etymology">
<paragraph id="8BD9EB16720DFFBEFF49F8D0F6B7F8A9" blockId="3.[151,1436,943,2012]" box="[201,1223,1808,1832]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49F8D0F33FF8A6" box="[201,335,1808,1832]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49F8D0F338F8A9" bold="true" box="[201,328,1808,1832]" pageId="3" pageNumber="508">Etymology</emphasis>
.
</emphasis>
Latinized name 
<taxonomicName id="4C669095720DFFBEFD8AF8D0F0D1F8A9" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[522,673,1808,1832]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFD8AF8D0F0D1F8A9" box="[522,673,1808,1832]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
refers to the great Buddhist monk Bodhidharma.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49F8F4F1FEF85D" pageId="3" pageNumber="508" type="discussion">
<paragraph id="8BD9EB16720DFFBEFF49F8F4F1FEF85D" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49F8F4F34CF8CA" box="[201,316,1844,1868]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49F8F4F346F8CD" bold="true" box="[201,310,1844,1868]" pageId="3" pageNumber="508">Remarks</emphasis>
.
</emphasis>
<taxonomicName id="4C669095720DFFBEFEC6F8F4F028F8CD" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[326,600,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFEC6F8F4F028F8CD" box="[326,600,1844,1868]" italics="true" pageId="3" pageNumber="508">Chlaenius bodhidharma</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C669095720DFFBEFD75F8F4F1C8F8CD" authorityName="Kirschenhofer" authorityYear="2003" box="[757,952,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B9123704720DFFBEFD75F8F4F1C8F8CD" box="[757,952,1844,1868]" italics="true" pageId="3" pageNumber="508">C. schillhammeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C669095720DFFBEFC74F8F4F61CF8CD" authorityName="Bates" authorityYear="1892" box="[1012,1132,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFC74F8F4F61CF8CD" box="[1012,1132,1844,1868]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
in having elytral intervals glabrous in the middle, but differs from the latter in having yellow spot at the middle of each elytron and covering only half of interval 
<quantity id="4C9E46F3720DFFBEFE06F8BCF3C3F815" box="[390,435,1916,1940]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="508" unit="in" value="4.0">4 in</quantity>
contrast to yellow spot placed slightly behind the middle and covering whole width of interval 
<quantity id="4C9E46F3720DFFBEFF74F860F36FF839" box="[244,287,1952,1976]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="508" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C669095720DFFBEFEA6F860F397F839" authorityName="Kirschenhofer" authorityYear="2003" box="[294,487,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B9123704720DFFBEFEA6F860F397F839" box="[294,487,1952,1976]" italics="true" pageId="3" pageNumber="508">C. schillhammeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C669095720DFFBEFD9FF860F0E5F839" authorityName="Bates" authorityYear="1892" box="[543,661,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFD9FF860F0E5F839" box="[543,661,1952,1976]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
. Additionally, aedeagus of 
<taxonomicName id="4C669095720DFFBEFC4AF860F6F2F839" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[970,1154,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFC4AF860F1ADF839" box="[970,989,1952,1976]" italics="true" pageId="3" pageNumber="508">C</emphasis>
. 
<emphasis id="B9123704720DFFBEFC6BF860F6F2F839" box="[1003,1154,1952,1976]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
is more robust (
<figureCitation id="135DF793720DFFBEFABBF860F7FFF839" box="[1339,1423,1952,1976]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Fig. 40</figureCitation>
), and apical lamella (
<figureCitation id="135DF793720DFFBEFEF2F804F3B3F85D" box="[370,451,1988,2012]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Fig. 41</figureCitation>
) longer than in 
<taxonomicName id="4C669095720DFFBEFDF5F804F09AF85D" authorityName="Bates" authorityYear="1892" box="[629,746,1988,2012]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFDF5F804F09AF85D" box="[629,746,1988,2012]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
(
<figureCitation id="135DF793720DFFBEFD79F804F10FF85D" box="[761,895,1988,2012]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 46, 47</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>