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<document id="D9E957DDA73FB50CC6EAB35B2FF14B82" ID-DOI="10.11646/zootaxa.5543.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386776" ID-ZooBank="E34AF3EF-4D03-4C08-8E11-C9514D42021B" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1733919186538" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2024" docId="03C83A1CFF8CC353FF772A2A577B4063" docLanguage="en" docName="zootaxa.5543.4.1.pdf" docOrigin="Zootaxa 5543 (4)" docSource="https://doi.org/10.11646/zootaxa.5543.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Circeaster pullus Mah 2006" docType="treatment" docVersion="2" lastPageNumber="469" masterDocId="FFF14264FF9DC341FFE02E495341417D" masterDocTitle="New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific" masterLastPageNumber="500" masterPageNumber="451" pageNumber="468" updateTime="1734380713031" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="511778B7FD79AA2A4D04C4672250EA56">New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific</mods:title>
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<mods:namePart id="D06E8A8F32B94210DBAAABD8C59217B1">Mah, Christopher L.</mods:namePart>
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<mods:date id="801C39DF4B5E350C4F77B7A704143420">2024</mods:date>
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<treatment id="03C83A1CFF8CC353FF772A2A577B4063" ID-DOI="http://doi.org/10.5281/zenodo.14503272" ID-Zenodo-Dep="14503272" LSID="urn:lsid:plazi:treatment:03C83A1CFF8CC353FF772A2A577B4063" httpUri="http://treatment.plazi.org/id/03C83A1CFF8CC353FF772A2A577B4063" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<subSubSection id="C37BD881FF8CC350FF772A2A52B04501" box="[151,497,1122,1149]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8BDE8B0AFF8CC350FF772A2A52B04501" blockId="17.[151,497,1122,1185]" box="[151,497,1122,1149]" pageId="17" pageNumber="468">
<heading id="D0963C66FF8CC350FF772A2A52B04501" bold="true" box="[151,497,1122,1149]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<taxonomicName id="4C61F089FF8CC350FF772A2A52B04501" ID-CoL="5Z78M" authority="Mah, 2006" authorityName="Mah" authorityYear="2006" box="[151,497,1122,1149]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B9155718FF8CC350FF772A2A52B04501" bold="true" box="[151,497,1122,1149]" pageId="17" pageNumber="468">
<emphasis id="B9155718FF8CC350FF772A2A52244500" bold="true" box="[151,357,1122,1149]" italics="true" pageId="17" pageNumber="468">Circeaster pullus</emphasis>
<bibRefCitation id="EFF0F6FBFF8CC350FE8D2A2A52B04501" author="Mah, C. L." box="[365,497,1122,1149]" pageId="17" pageNumber="468" pagination="917 - 954" refId="ref22084" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37BD881FF8CC350FF772ACE5115459E" pageId="17" pageNumber="468" type="description">
<paragraph id="8BDE8B0AFF8CC350FF772ACE52EE45DC" blockId="17.[151,497,1122,1185]" box="[151,431,1158,1185]" pageId="17" pageNumber="468">
<figureCitation id="135A978FFF8CC350FF772ACE521045DC" box="[151,337,1158,1185]" captionStart="FIGURE 15" captionStartId="36.[151,250,1909,1933]" captionTargetBox="[189,1399,181,1886]" captionTargetId="figure-24@36.[189,1399,181,1886]" captionTargetPageId="36" captionText="FIGURE 15. Circeaster pullus Mah 2006 feeding observations. A–B. Feeding on Narella (Chrysogorgiidae). Santa Rosa North 467.3 m, Lasers on B are 10 cm apart, others lacking scale. C. On bamboo coral, Lone Cone, 1543.8 m. D. On bamboo coral, Unamed Seamount, 1605.8 m E. Johnson Atoll site 1, 520 m. F. feeding on bamboo coral, Johnston Atoll region, 1582 m." figureDoi="http://doi.org/10.5281/zenodo.14386830" httpUri="https://zenodo.org/record/14386830/files/figure.png" pageId="17" pageNumber="468">FIGURE 15A–F</figureCitation>
,
<figureCitation id="135A978FFF8CC350FEBD2ACE52EE45DC" box="[349,431,1159,1185]" captionStart="FIGURE 16" captionStartId="37.[151,250,1781,1805]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-15@37.[151,1436,181,1757]" captionTargetPageId="37" captionText="FIGURE 16. Circeaster pullus Mah 2006 feeding observations. A.Feeding on Isididae, Johnston Atoll, 492 m. B–C. Feeding on Isididae, Santa Rosa North site, 467.3 m. Lasers are 10 cm apart. D. Feeding? on stoloniferous soft corals, Sibelius Seamount, 2489 m. Lasers=10cm apart." figureDoi="http://doi.org/10.5281/zenodo.14386834" httpUri="https://zenodo.org/record/14386834/files/figure.png" pageId="17" pageNumber="468">16A–D</figureCitation>
</paragraph>
<paragraph id="8BDE8B0AFF8CC350FF772A855115459E" blockId="17.[151,596,1227,1251]" box="[151,596,1227,1251]" pageId="17" pageNumber="468">
<taxonomicName id="4C61F089FF8CC350FF772A855207459E" authorityName="Mah" authorityYear="2006" box="[151,326,1227,1251]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B9155718FF8CC350FF772A855207459E" box="[151,326,1227,1251]" italics="true" pageId="17" pageNumber="468">Circeaster pullus</emphasis>
</taxonomicName>
, 2006: 940,
<figureCitation id="135A978FFF8CC350FE252A825179459E" box="[453,568,1227,1251]" captionStart="FIGURE 6" captionStartId="20.[151,250,1801,1825]" captionTargetBox="[151,1436,181,1777]" captionTargetId="figure-24@20.[151,1436,181,1777]" captionTargetPageId="20" captionText="FIGURE 6. Cladaster kiko n. sp. MCZ 162797. Holotype. A. Abactinal B. In situ observation. C. Abactinal-lateral view. D. Abactinal-lateral view showing abactinal and superomarginal surface. E. Actinal F. Adambulacral plates showing large furrow spines. Scale bar=10.0 mm, A, E; Scale bar=5.0 mm: C, D; Scale bar=1.0 mm: F." figureDoi="http://doi.org/10.5281/zenodo.14386794" httpUri="https://zenodo.org/record/14386794/files/figure.png" pageId="17" pageNumber="468">Figs. 6D–F</figureCitation>
,
<figureCitation id="135A978FFF8CC350FDA22A825110459E" box="[578,593,1227,1251]" captionStart="FIGURE 8" captionStartId="26.[151,250,1490,1514]" captionTargetBox="[151,1436,181,1466]" captionTargetId="figure-24@26.[151,1436,181,1466]" captionTargetPageId="26" captionText="FIGURE 8. Pillsburiaster hyugaensis Mah &amp; Fujita n. sp. NSMT E-11392. Holotype, A. Abactinal B. Abactinal-lateral view. C. Abactinal Closeup showing coarse granules. D.Abactinal-lateral view showing surface superomarginal plates. E. Actinal. F. Adambulacral plates showing furrow and subambulacral spination. Scale bar=20 mm, A, E; Scale bar=3.0: B, D; Scale bar=2.0 mm: C, F." figureDoi="http://doi.org/10.5281/zenodo.14386806" httpUri="https://zenodo.org/record/14386806/files/figure.png" pageId="17" pageNumber="468">8</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37BD881FF8CC350FF772B58523F44A2" pageId="17" pageNumber="468" type="diagnosis">
<paragraph id="8BDE8B0AFF8CC350FF772B5852514456" blockId="17.[151,272,1297,1323]" box="[151,272,1297,1323]" pageId="17" pageNumber="468">
<emphasis id="B9155718FF8CC350FF772B5852514456" bold="true" box="[151,272,1297,1323]" pageId="17" pageNumber="468">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BDE8B0AFF8CC350FF772B10523F44A2" blockId="17.[151,1436,1369,1503]" pageId="17" pageNumber="468">R/r = 2.0–3.0. Abactinal arm plates bare, enlarged, at base of arm. Round, hemispherical granules on disk, marginal plate surface. Superomarginal plates abutted over arm midline. Enlarged subambulacral spine abradial to adambulacral pedicellaria, adjacent to actinal intermediate region. Bivalve pedicellariae present on actinal surface, adambulacral plates.</paragraph>
</subSubSection>
<subSubSection id="C37BD881FF8CC350FF772878508646F2" pageId="17" pageNumber="468" type="discussion">
<paragraph id="8BDE8B0AFF8CC350FF772878525F4736" blockId="17.[151,286,1585,1611]" box="[151,286,1585,1611]" pageId="17" pageNumber="468">
<heading id="D0963C66FF8CC350FF772878525F4736" bold="true" box="[151,286,1585,1611]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="B9155718FF8CC350FF772878525F4736" bold="true" box="[151,286,1585,1611]" pageId="17" pageNumber="468">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE8B0AFF8CC350FF77283350A447A1" blockId="17.[151,1436,1657,1936]" pageId="17" pageNumber="468">
<emphasis id="B9155718FF8CC350FF77283353A347EE" box="[151,226,1658,1683]" italics="true" pageId="17" pageNumber="468">In situ</emphasis>
observations of what was identified as
<taxonomicName id="4C61F089FF8CC350FD5B2833506547EE" authorityName="Mah" authorityYear="2006" box="[699,804,1657,1684]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B9155718FF8CC350FD5B2833506547EE" box="[699,804,1657,1684]" italics="true" pageId="17" pageNumber="468">C. pullus</emphasis>
</taxonomicName>
were made based on the identification of the abutted superomarginal plates along the arm. None of the video observations showed the furrow spines which was another of the primary characters which distinguished this species from
<taxonomicName id="4C61F089FF8CC350FC83288B509F47A6" authorityName="Mah" authorityYear="2006" box="[867,990,1729,1756]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="sandrae">
<emphasis id="B9155718FF8CC350FC83288B509F47A6" box="[867,990,1729,1756]" italics="true" pageId="17" pageNumber="468">C. sandrae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE8B0AFF8CC350FF2728AF508646F2" blockId="17.[151,1436,1657,1936]" pageId="17" pageNumber="468">
Two color variants were observed that were consistent with the morphological definition of
<taxonomicName id="4C61F089FF8CC350FB3428AF56D44782" authorityName="Mah" authorityYear="2006" box="[1236,1429,1765,1792]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B9155718FF8CC350FB3428AF56D44782" box="[1236,1429,1765,1792]" italics="true" pageId="17" pageNumber="468">Circeaster pullus</emphasis>
</taxonomicName>
, one a straw color with white highlights, with shorter, broader arms that was observed at relatively shallow depth (
<quantity id="4C9926EFFF8CC350FF7F29645260463A" box="[159,289,1837,1864]" metricMagnitude="2" metricUnit="m" metricValue="4.72" metricValueMax="5.2" metricValueMin="4.24" pageId="17" pageNumber="468" unit="m" value="472.0" valueMax="520.0" valueMin="424.0">424–520 m</quantity>
) (
<figureCitation id="135A978FFF8CC350FED82964529E4635" box="[312,479,1837,1864]" captionStart="FIGURE 15" captionStartId="36.[151,250,1909,1933]" captionTargetBox="[189,1399,181,1886]" captionTargetId="figure-24@36.[189,1399,181,1886]" captionTargetPageId="36" captionText="FIGURE 15. Circeaster pullus Mah 2006 feeding observations. A–B. Feeding on Narella (Chrysogorgiidae). Santa Rosa North 467.3 m, Lasers on B are 10 cm apart, others lacking scale. C. On bamboo coral, Lone Cone, 1543.8 m. D. On bamboo coral, Unamed Seamount, 1605.8 m E. Johnson Atoll site 1, 520 m. F. feeding on bamboo coral, Johnston Atoll region, 1582 m." figureDoi="http://doi.org/10.5281/zenodo.14386830" httpUri="https://zenodo.org/record/14386830/files/figure.png" pageId="17" pageNumber="468">Fig. 15A, B, E</figureCitation>
;
<figureCitation id="135A978FFF8CC350FE0B2964517F463A" box="[491,574,1837,1863]" captionStart="FIGURE 16" captionStartId="37.[151,250,1781,1805]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-15@37.[151,1436,181,1757]" captionTargetPageId="37" captionText="FIGURE 16. Circeaster pullus Mah 2006 feeding observations. A.Feeding on Isididae, Johnston Atoll, 492 m. B–C. Feeding on Isididae, Santa Rosa North site, 467.3 m. Lasers are 10 cm apart. D. Feeding? on stoloniferous soft corals, Sibelius Seamount, 2489 m. Lasers=10cm apart." figureDoi="http://doi.org/10.5281/zenodo.14386834" httpUri="https://zenodo.org/record/14386834/files/figure.png" pageId="17" pageNumber="468">16A–C</figureCitation>
) and a dark orange variant with more elongate arms present in deeper waters (
<quantity id="4C9926EFFF8CC350FF7F2918527D4616" box="[159,316,1873,1900]" metricMagnitude="3" metricUnit="m" metricValue="2.0469999999999997" metricValueMax="2.489" metricValueMin="1.605" pageId="17" pageNumber="468" unit="m" value="2047.0" valueMax="2489.0" valueMin="1605.0">1605–2489 m</quantity>
) (
<figureCitation id="135A978FFF8CC350FEB3291852B94611" box="[339,504,1873,1900]" captionStart="FIGURE 15" captionStartId="36.[151,250,1909,1933]" captionTargetBox="[189,1399,181,1886]" captionTargetId="figure-24@36.[189,1399,181,1886]" captionTargetPageId="36" captionText="FIGURE 15. Circeaster pullus Mah 2006 feeding observations. A–B. Feeding on Narella (Chrysogorgiidae). Santa Rosa North 467.3 m, Lasers on B are 10 cm apart, others lacking scale. C. On bamboo coral, Lone Cone, 1543.8 m. D. On bamboo coral, Unamed Seamount, 1605.8 m E. Johnson Atoll site 1, 520 m. F. feeding on bamboo coral, Johnston Atoll region, 1582 m." figureDoi="http://doi.org/10.5281/zenodo.14386830" httpUri="https://zenodo.org/record/14386830/files/figure.png" pageId="17" pageNumber="468">Fig. 15C, D, F</figureCitation>
;
<figureCitation id="135A978FFF8CC350FDE2291851764616" box="[514,567,1873,1899]" captionStart="FIGURE 16" captionStartId="37.[151,250,1781,1805]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-15@37.[151,1436,181,1757]" captionTargetPageId="37" captionText="FIGURE 16. Circeaster pullus Mah 2006 feeding observations. A.Feeding on Isididae, Johnston Atoll, 492 m. B–C. Feeding on Isididae, Santa Rosa North site, 467.3 m. Lasers are 10 cm apart. D. Feeding? on stoloniferous soft corals, Sibelius Seamount, 2489 m. Lasers=10cm apart." figureDoi="http://doi.org/10.5281/zenodo.14386834" httpUri="https://zenodo.org/record/14386834/files/figure.png" pageId="17" pageNumber="468">16D</figureCitation>
). The
<typeStatus id="54DA35A8FF8CC350FD9B291B51EA4611" box="[635,683,1874,1900]" pageId="17" pageNumber="468">type</typeStatus>
series for
<taxonomicName id="4C61F089FF8CC350FCFF291B50C24616" authorityName="Mah" authorityYear="2006" box="[799,899,1873,1900]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="468" phylum="Echinodermata" rank="species" species="pullus">
<emphasis id="B9155718FF8CC350FCFF291B50C24616" box="[799,899,1873,1900]" italics="true" pageId="17" pageNumber="468">C. pullus</emphasis>
</taxonomicName>
was collected from depths more consistent with the shallow, yellow variant. Full data on videos is listed in the Appendix.
</paragraph>
</subSubSection>
<subSubSection id="C37BD881FF8FC353FF772EDE577B4063" pageId="18" pageNumber="469" type="distribution">
<paragraph id="8BDE8B0AFF8FC353FF772EDE526841CC" blockId="18.[151,297,151,177]" box="[151,297,151,177]" pageId="18" pageNumber="469">
<heading id="D0963C66FF8FC353FF772EDE526841CC" bold="true" box="[151,297,151,177]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="B9155718FF8FC353FF772EDE526841CC" bold="true" box="[151,297,151,177]" pageId="18" pageNumber="469">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE8B0AFF8FC353FF772E96577B4063" blockId="18.[151,1082,223,286]" pageId="18" pageNumber="469">
<collectingCountry id="F376CB9AFF8FC353FF772E96527D4184" box="[151,316,223,249]" name="United States Pacific Island Wildlife Refuges" pageId="18" pageNumber="469">Johnston Atoll</collectingCountry>
, North Pacific,
<quantity id="4C9926EFFF8FC353FE0C2E96513D4187" box="[492,636,223,250]" metricMagnitude="3" metricUnit="m" metricValue="1.478" metricValueMax="2.489" metricValueMin="0.46699999999999997" pageId="18" pageNumber="469" unit="m" value="1478.0" valueMax="2489.0" valueMin="467.0">467–2489 m</quantity>
. South Pacific Ocean,
<collectingCountry id="F376CB9AFF8FC353FE592F4D51404063" box="[441,513,260,286]" name="Tonga" pageId="18" pageNumber="469">Tonga</collectingCountry>
,
<collectingCountry id="F376CB9AFF8FC353FDED2F4D51FC4060" box="[525,701,259,286]" name="New Caledonia" pageId="18" pageNumber="469">New Caledonia</collectingCountry>
, Marquesas Islands.
<quantity id="4C9926EFFF8FC353FC462F4A57774063" box="[934,1078,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.525" metricValueMax="10.0" metricValueMin="7.05" pageId="18" pageNumber="469" unit="m" value="852.5" valueMax="1000.0" valueMin="705.0">705–1000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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