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<document id="F1D8DAC3A4C7C6E72D77DAD577E32EBD" ID-CLB-Dataset="303278" ID-DOI="10.11646/zootaxa.5506.2.1" ID-GBIF-Dataset="376491fe-cadb-454b-82bb-16edc4df6ef8" ID-ISSN="1175-5326" ID-Zenodo-Dep="13746744" ID-ZooBank="F51D2CFF-49FF-4ED9-8A44-1FD0F1B10379" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1726053386088" checkinUser="plazi" docAuthor="Aidek, Ahmad E., Saad, Adib, Jablonski, Daniel, Esterbauer, Hans &amp; Fritz, Uwe" docDate="2024" docId="03B9F52AC72AFF9404B62C93F598F874" docLanguage="en" docName="zootaxa.5506.2.1.pdf" docOrigin="Zootaxa 5506 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mauremys rivulata" docType="treatment" docVersion="4" lastPageNumber="167" masterDocId="FF808D52C725FF840421296BF408FFB5" masterDocTitle="Turtles and tortoises of Syria: Diversity, distribution, and conservation" masterLastPageNumber="193" masterPageNumber="151" pageNumber="166" updateTime="1726487354855" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1770CA6A9A847D6F3C0E6F1F687C0786" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CBB057A434050145DD1F87364456B975">
<mods:title id="059F5FD14FB13DA5BA68FE652D1D6413">Turtles and tortoises of Syria: Diversity, distribution, and conservation</mods:title>
</mods:titleInfo>
<mods:name id="76960E8552E98840BB431C31C38E20DA" type="personal">
<mods:role id="A36F5FEE347B892D1F23D692BE20795A">
<mods:roleTerm id="96F57871146D4FC40ECF858BC7445799">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="236AF46D14F5042DECCA194D7AC3CBBB">Aidek, Ahmad E.</mods:namePart>
<mods:affiliation id="D81DF17909C70DBC909326470EEEAE07">General Commission for Scientific Agricultural Research, Deir ez-Zor, Syria</mods:affiliation>
</mods:name>
<mods:name id="A767B8875C06B66EA3A264BE539818D9" type="personal">
<mods:role id="B8949ECA32764C128A4D93D13613F21C">
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</mods:role>
<mods:namePart id="4B1A3D6052244E04019EA00011A6D2FF">Saad, Adib</mods:namePart>
<mods:affiliation id="F6846EE96434052A034C0B3E599EDC15">Al-Manara University, Latakia, Syria</mods:affiliation>
</mods:name>
<mods:name id="81604A60C96621AED3FA3305697C710B" type="personal">
<mods:role id="A30F5077DE268C3055EA815B7D6D92A7">
<mods:roleTerm id="D57636174E4176E3E888B6353ADD435F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E87626948B4837862534C2D0AFC6A3F">Jablonski, Daniel</mods:namePart>
<mods:affiliation id="66EB929E21921444DCF810C2F11ABBEE">Department of Zoology, Comenius University in Bratislava, Ilkovičova 6, Mlynská dolina, 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name id="4ACC418274D7AB7F93162C300351A7C1" type="personal">
<mods:role id="E09F6F1A9E3C039CD24C774C41405316">
<mods:roleTerm id="C94FEAC5036D1C22EF83A80581C10165">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC4F50AE3F37CE813E968F6395E3E3B7">Esterbauer, Hans</mods:namePart>
<mods:affiliation id="09548190B58A9EC649C65E3B8C9E2E78">Johann-Puch-Strasse 27 / III / 5, 4400 Steyr, Austria</mods:affiliation>
</mods:name>
<mods:name id="B8CF7A0279696439E8A8A7DF37F380D5" type="personal">
<mods:role id="FB6614DFB71E1639616DA482B17D1D18">
<mods:roleTerm id="97FB727B065681E272BA0728A1AA2DB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF0674B020F0ED5EFF6C95407A0C56F7">Fritz, Uwe</mods:namePart>
<mods:affiliation id="795AFE0B7CA9B5DE1904905AA8902185">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9F8768872EF0C5A0508FEC87067E5E27">text</mods:typeOfResource>
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<mods:title id="E5F8B893ADCB0C1CF860439F3220ACB9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF2BB7373318ECA7EE8008F92A04334D">
<mods:date id="AA4211C289E8B203DA9AA64EA982AC0D">2024</mods:date>
<mods:detail id="ECB4C5869A3A616F4480DEEC31FB2670" type="pubDate">
<mods:number id="B10BE6A033DDCB40E43ADEA82D142A1B">2024-09-06</mods:number>
</mods:detail>
<mods:detail id="C81173D632694E959A1932A76B80C2E5" type="volume">
<mods:number id="6876FBE615D1C03436CDB3CB06B5215B">5506</mods:number>
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<mods:url id="3E42F7336BA49B9BFBED9ACBBF688214">http://dx.doi.org/10.11646/zootaxa.5506.2.1</mods:url>
</mods:location>
<mods:classification id="3DEAA431D28758732F1C144EFFE66147">journal article</mods:classification>
<mods:identifier id="D58D74EB13AA3DE3D80C52055C4586A3" type="CLB-Dataset">303278</mods:identifier>
<mods:identifier id="ED0E266128B268FCB85A4BCA1762A3EE" type="DOI">10.11646/zootaxa.5506.2.1</mods:identifier>
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<treatment id="03B9F52AC72AFF9404B62C93F598F874" ID-DOI="http://doi.org/10.5281/zenodo.13767995" ID-Zenodo-Dep="13767995" LSID="urn:lsid:plazi:treatment:03B9F52AC72AFF9404B62C93F598F874" httpUri="http://treatment.plazi.org/id/03B9F52AC72AFF9404B62C93F598F874" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">
<subSubSection id="C30A17B7C72AFF8B04B62C93F6C1F980" pageId="15" pageNumber="166" type="nomenclature">
<paragraph id="8BAF443CC72AFF8B04B62C93F7C8F9A4" blockId="15.[151,960,1527,1590]" box="[151,960,1527,1554]" pageId="15" pageNumber="166">
<heading id="D0E7F350C72AFF8B04B62C93F7C8F9A4" bold="true" box="[151,960,1527,1554]" fontSize="11" level="1" pageId="15" pageNumber="166" reason="1">
<emphasis id="B964982EC72AFF8B04B62C93F7C8F9A4" bold="true" box="[151,960,1527,1554]" pageId="15" pageNumber="166">
<taxonomicName id="4C103FBFC72AFF8B04B62C93F576F9A7" ID-CoL="3YFWM" authority="(Valenciennes in Bory de Saint-Vincent, 1833)" box="[151,382,1527,1554]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="15" pageNumber="166" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC72AFF8B04B62C93F576F9A7" bold="true" box="[151,382,1527,1554]" italics="true" pageId="15" pageNumber="166">Mauremys rivulata</emphasis>
</taxonomicName>
(Valenciennes
<emphasis id="B964982EC72AFF8B06192C9CF658F9A4" bold="true" box="[568,592,1527,1553]" italics="true" pageId="15" pageNumber="166">in</emphasis>
Bory de
<collectingCountry id="F30704ACC72AFF8B06E32C9CF778F9A4" box="[706,880,1527,1553]" name="Saint Vincent and the Grenadines" pageId="15" pageNumber="166">Saint-Vincent</collectingCountry>
, 1833)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF443CC72AFF8B04B62F77F6C1F980" blockId="15.[151,960,1527,1590]" box="[151,713,1563,1590]" pageId="15" pageNumber="166">
<materialsCitation id="3B784E61C72AFF8B04B62F77F6C1F980" box="[151,713,1563,1590]" location="Western Caspian Turtle" pageId="15" pageNumber="166" specimenCount="1" stateProvince="Balkan Welayaty">
<location id="8ECF12E7C72AFF8B04B62F77F5A8F980" LSID="urn:lsid:plazi:treatment:03B9F52AC72AFF9404B62C93F598F874:8ECF12E7C72AFF8B04B62F77F5A8F980" box="[151,416,1563,1590]" name="Western Caspian Turtle" pageId="15" pageNumber="166" stateProvince="Balkan Welayaty">Western Caspian Turtle</location>
,
<collectingRegion id="49D48ADEC72AFF8B058A2F70F5F2F980" box="[427,506,1563,1589]" country="Turkmenistan" name="Balkan Welayaty" pageId="15" pageNumber="166">Balkan</collectingRegion>
Terrapin (
<figureCitation id="132B58B9C72AFF8B064E2F70F6C8F980" box="[623,704,1563,1589]" captionStart="FIGURE 14" captionStartId="16.[151,250,1120,1144]" captionTargetBox="[151,1436,237,1097]" captionTargetId="figure-54@16.[151,1436,237,1097]" captionTargetPageId="16" captionText="FIGURE 14. Balkan terrapins, Mauremys rivulata. (A) Al-Alani, upper Orontes (photo: H. Baniasi); (B) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (C) SaddAl-Kafat, southeast Hamah (photo: Ehab Jbr); (D) Bhalu, Nahr Al-Qash, east of Latakia (photo: Basel Barakat)." figureDoi="http://doi.org/10.5281/zenodo.13746778" httpUri="https://zenodo.org/record/13746778/files/figure.png" pageId="15" pageNumber="166">Fig. 14</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72AFF8B04B62F08F6A3F9C8" box="[151,683,1635,1662]" pageId="15" pageNumber="166" type="description">
<paragraph id="8BAF443CC72AFF8B04B62F08F6A3F9C8" blockId="15.[151,1437,1635,2022]" box="[151,683,1635,1662]" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04B62F08F5C9F9C8" bold="true" box="[151,449,1635,1661]" pageId="15" pageNumber="166">IUCN Red List Category</emphasis>
: Not evaluated (NE)
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C72AFF9404E62FECF549F9E8" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166" type="materials_examined">
<paragraph id="8BAF443CC72AFF8B04E62FECF551F95C" blockId="15.[151,1437,1635,2022]" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04E62FECF56DF914" bold="true" box="[199,357,1671,1697]" pageId="15" pageNumber="166">
<typeStatus id="54ABFA9EC72AFF8B04E62FECF50AF914" box="[199,258,1671,1697]" pageId="15" pageNumber="166">Type</typeStatus>
locality
</emphasis>
: Restricted to Modon, Morée [
<collectingRegion id="49D48ADEC72AFF8B06FB2FECF765F914" box="[730,877,1671,1697]" country="Greece" name="Peloponnisos" pageId="15" pageNumber="166">Peloponnese</collectingRegion>
,
<collectingCountry id="F30704ACC72AFF8B075F2FECF7D8F914" box="[894,976,1671,1697]" name="Greece" pageId="15" pageNumber="166">Greece</collectingCountry>
] by
<bibRefCitation id="EF8139CDC72AFF8B002A2FE3F16FF914" author="Bory de Saint-Vincent, J. B." box="[1035,1383,1671,1698]" pageId="15" pageNumber="166" refId="ref13644" refString="Bory de Saint-Vincent, J. B. (1836) Expedition Scientifique de Moree. Section des Sciences Physiques. Tome Premier. Relation. F. G. Levrault, Paris, 472 pp." type="book" year="1836">
Bory de
<collectingCountry id="F30704ACC72AFF8B00572FECF118F914" box="[1142,1296,1671,1697]" name="Saint Vincent and the Grenadines" pageId="15" pageNumber="166">Saint-Vincent</collectingCountry>
(1836)
</bibRefCitation>
and emended to “Umgebung von Modon,
<collectingRegion id="49D48ADEC72AFF8B06172FC7F677F973" box="[566,639,1708,1734]" country="Greece" name="Peloponnisos" pageId="15" pageNumber="166">Morea</collectingRegion>
[environs of Modon,
<collectingRegion id="49D48ADEC72AFF8B074C2FC0F008F970" box="[877,1024,1707,1733]" country="Greece" name="Peloponnisos" pageId="15" pageNumber="166">Peloponnese</collectingRegion>
],
<collectingCountry id="F30704ACC72AFF8B00312FC0F0A2F970" box="[1040,1194,1707,1733]" name="Greece" pageId="15" pageNumber="166">Griechenland</collectingCountry>
” [
<collectingCountry id="F30704ACC72AFF8B009E2FC0F11CF970" box="[1215,1300,1707,1733]" name="Greece" pageId="15" pageNumber="166">Greece</collectingCountry>
] by
<bibRefCitation id="EF8139CDC72AFF8B01612FC7F55EF95C" author="Mertens, R. &amp; Muller, L." pageId="15" pageNumber="166" pagination="1 - 62" refId="ref16694" refString="Mertens, R. &amp; Muller, L. (1928) Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 41, 1 - 62." type="journal article" year="1928">Mertens &amp; Müller (1928)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAF443CC72AFF8B04E62F98F039F884" blockId="15.[151,1437,1635,2022]" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04E62F98F589F8B8" bold="true" box="[199,385,1779,1805]" pageId="15" pageNumber="166">Type specimens</emphasis>
:
<materialsCitation id="3B784E61C72AFF8B05AF2F98F025F884" collectionCode="MNHN" pageId="15" pageNumber="166" specimenCode="MNHN 1930, 1930" specimenCount="6" typeStatus="holotype">
<specimenCount id="9D168FB5C72AFF8B05AF2F98F614F8BB" box="[398,540,1779,1806]" count="6" pageId="15" pageNumber="166" type="generic" typeStatus="syntype">Six syntypes</specimenCount>
in the Muséum National d’Historie Naturelle, Paris,
<specimenCode id="DBB6EC47C72AFF8B00452F9FF0F2F8B8" box="[1124,1274,1779,1806]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="166">MNHN 1930</specimenCode>
,
<specimenCode id="DBB6EC47C72AFF8B01252F98F136F8B8" box="[1284,1342,1779,1805]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="166">1930</specimenCode>
A, 4094, 4095, 9491, and 9492.
<specimenCode id="DBB6EC47C72AFF8B05B82E73F625F884" box="[409,557,1815,1842]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="166">MNHN 1930</specimenCode>
listed as
<typeStatus id="54ABFA9EC72AFF8B06B72E7CF6F0F884" box="[662,760,1815,1841]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="EF8139CDC72AFF8B07032E7CF025F884" author="Fritz, U. &amp; Wischuf, T." box="[802,1069,1815,1841]" pageId="15" pageNumber="166" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf (1997)</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF443CC72AFF8B04E62E50F655F8E0" blockId="15.[151,1437,1635,2022]" box="[199,605,1851,1878]" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04E62E50F54EF8E0" bold="true" box="[199,326,1851,1877]" pageId="15" pageNumber="166">Chorotype</emphasis>
: Eastern-Mediterranean.
</paragraph>
<paragraph id="8BAF443CC72AFF8B04E62E34F0BFF8CC" blockId="15.[151,1437,1635,2022]" box="[199,1207,1887,1914]" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04E62E34F5C0F8CC" bold="true" box="[199,456,1887,1913]" pageId="15" pageNumber="166">
First record for
<collectingCountry id="F30704ACC72AFF8B05A92E34F5C0F8CC" box="[392,456,1887,1913]" name="Syria" pageId="15" pageNumber="166">Syria</collectingCountry>
</emphasis>
:
<collectingRegion id="49D48ADEC72AFF8B05F42E34F622F8CC" box="[469,554,1887,1913]" country="Syria" name="Halab" pageId="15" pageNumber="166">Aleppo</collectingRegion>
, Nahr Al-Dhahab, Sabkhat Al-Jabboul (
<bibRefCitation id="EF8139CDC72AFF8B07C62E34F0A2F8CC" author="Siebenrock, F." box="[999,1194,1887,1913]" pageId="15" pageNumber="166" pagination="171 - 225" refId="ref19218" refString="Siebenrock, F. (1913) Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Schildkroten aus Syrien und Mesopotamien. Annalen des Naturhistorischen Museums Wien, 27, 171 - 225." type="journal article" year="1913">Siebenrock 1913</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF443CC72AFF9404E62EE8F0DCFF60" blockId="15.[151,1437,1635,2022]" lastBlockId="16.[151,1436,151,213]" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">
<emphasis id="B964982EC72AFF8B04E62EE8F577F828" bold="true" box="[199,383,1923,1949]" pageId="15" pageNumber="166">Literature data</emphasis>
:
<collectingRegion id="49D48ADEC72AFF8B05AA2EE8F5E8F828" box="[395,480,1923,1949]" country="Syria" name="Halab" pageId="15" pageNumber="166">Aleppo</collectingRegion>
, Nahr Al-Dhahab, Sabkhat Al-Jabboul (
<bibRefCitation id="EF8139CDC72AFF8B07BA2EE8F055F828" author="Siebenrock, F." box="[923,1117,1923,1949]" pageId="15" pageNumber="166" pagination="171 - 225" refId="ref19218" refString="Siebenrock, F. (1913) Wissenschaftliche Ergebnisse der Expedition nach Mesopotamien, 1910. Schildkroten aus Syrien und Mesopotamien. Annalen des Naturhistorischen Museums Wien, 27, 171 - 225." type="journal article" year="1913">Siebenrock 1913</bibRefCitation>
), Utaibeh (
<bibRefCitation id="EF8139CDC72AFF8B00F62EE8F187F828" author="Boulenger, G. A." box="[1239,1423,1923,1949]" pageId="15" pageNumber="166" pagination="41 - 55" refId="ref13716" refString="Boulenger, G. A. (1923) Etude sur les Batraciens et les Reptiles rapportes par M. Henri Gadeau de Kerville de son voyage zoologique en Syrie (avril-juin 1908). In: Pellegrin, J. (Ed.), Voyage zoologique d'Henri Gadeau de Kerville en Syrie (avril-juin 1908). Tome Quatrieme et Dernier. J. - B. Bailliere, Paris, pp. 41 - 55, pl. vi." type="book chapter" year="1923">Boulenger 1923</bibRefCitation>
), Masil Al-Fawwar, Al-Ka’b, Mas’ada, N Al-Qunaitra, SW Jubata Al-Khashab, Arnah (
<bibRefCitation id="EF8139CDC72AFF8B00572ECCF149F877" author="Esterbauer, H." box="[1142,1345,1959,1986]" pageId="15" pageNumber="166" pagination="23 - 34" refId="ref14303" refString="Esterbauer, H. (1985 a) Zur Herpetofauna Sudwestsyriens. Herpetofauna, 7 (38), 23 - 34." type="journal article" year="1985">Esterbauer 1985a</bibRefCitation>
,
<bibRefCitation id="EF8139CDC72AFF8B016E2ECCF190F877" author="Esterbauer, H." box="[1359,1432,1959,1986]" pageId="15" pageNumber="166" pagination="389 - 392" refId="ref14326" refString="Esterbauer, H. (1985 b) Okologische und verhaltensbiologische Beobachtungen an der Maurischen Landschildkrote, Testudo graeca terrestris Forskal, 1775 in Sudwestsyrien. Aquarien-Terrarien, 32, 389 - 392." type="journal article" year="1985">1985b</bibRefCitation>
,
<bibRefCitation id="EF8139CDC72AFF8B04B62EA0F4DAF850" author="Esterbauer, H." box="[151,210,1995,2021]" pageId="15" pageNumber="166" pagination="21 - 54" refId="ref14360" refString="Esterbauer, H. (1992) Die Herpetofauna des ostlichen Golan und Hermongebietes. Funde und Bemerkungen zur Systematik und Okologie. Zoology in the Middle East, 7, 21 - 54. https: // doi. org / 10.1080 / 09397140.1992.10637623" type="journal article" year="1992">1992</bibRefCitation>
,
<bibRefCitation id="EF8139CDC72AFF8B04FC2EA0F511F850" author="Esterbauer, H." box="[221,281,1995,2021]" pageId="15" pageNumber="166" pagination="20 - 27" refId="ref14439" refString="Esterbauer, H. (2022) Die Froschlurche des syrischen Bachsystems Masil al Fawwar. Aqua-Terra Austria, 17, 20 - 27." type="journal article" year="2022">2022</bibRefCitation>
), Golan (
<bibRefCitation id="EF8139CDC72AFF8B055F2EA0F67EF850" author="Sivan, N. &amp; Werner, Y. L." box="[382,630,1995,2022]" pageId="15" pageNumber="166" pagination="193 - 211" refId="ref19439" refString="Sivan, N. &amp; Werner, Y. L. (1992) Survey of the reptiles of the Golan plateau and Mt. Hermon. Israel Journal of Ecology and Evolution, 37, 193 - 211. https: // doi. org / 10.1080 / 00212210.1992.10688655" type="journal article" year="1992">Sivan &amp; Werner 1992</bibRefCitation>
),
<collectingRegion id="49D48ADEC72AFF8B06A92EA0F6D5F850" box="[648,733,1995,2021]" country="Syria" name="Halab" pageId="15" pageNumber="166">Aleppo</collectingRegion>
, Nahr Qweiq, Nahr Al-Dhahab, Oasis of
<collectingRegion id="49D48ADEC72AFF8B00892EA7F116F853" box="[1192,1310,1996,2022]" country="Syria" name="Dimashq" pageId="15" pageNumber="166">Damascus</collectingRegion>
, Housh Al-Admal (= Hosch-Odmol), Utaibeh Lake, Yarmouk River, Mzeirib Lake (
<bibRefCitation id="EF8139CDC735FF9407E329FCF0B4FF04" author="Fritz, U. &amp; Wischuf, T." box="[962,1212,151,177]" pageId="16" pageNumber="167" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf 1997</bibRefCitation>
), Ein Taqa, Mazraet Mardido, Orontes River near Jourine,
<quantity id="4CE8E9D9C735FF94066329D0F673FF60" box="[578,635,187,213]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="167" unit="km" value="4.0">4 km</quantity>
NE Masyaf (= Mesiaf;
<bibRefCitation id="EF8139CDC735FF9407A429D0F089FF60" author="Vamberger, M. &amp; Stuckas, H. &amp; Ayaz, D. &amp; Lymberakis, P. &amp; Siroky, P. &amp; Fritz, U." box="[901,1153,187,213]" pageId="16" pageNumber="167" pagination="313 - 322" refId="ref20818" refString="Vamberger, M., Stuckas, H., Ayaz, D., Lymberakis, P., Siroky, P. &amp; Fritz, U. (2014) Massive transoceanic gene flow in a freshwater turtle (Testudines: Geoemydidae: Mauremys rivulata). Zoologica Scripta, 43, 313 - 322. https: // doi. org / 10.1111 / zsc. 12055" type="journal article" year="2014">
Vamberger
<emphasis id="B964982EC735FF94002629D0F037FF60" box="[1031,1087,187,213]" italics="true" pageId="16" pageNumber="167">et al.</emphasis>
2014
</bibRefCitation>
,
<bibRefCitation id="EF8139CDC735FF9400AD29D0F0C1FF60" author="Vamberger, M. &amp; Stuckas, H. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Ayaz, D. &amp; Aloufi, A. A. &amp; Lymberakis, P. &amp; Siroky, P. &amp; Fritz, U." box="[1164,1225,187,213]" pageId="16" pageNumber="167" pagination="401 - 413" refId="ref20888" refString="Vamberger, M., Stuckas, H., Vargas-Ramirez, M., Kehlmaier, C., Ayaz, D., Aloufi, A. A., Lymberakis, P., Siroky, P. &amp; Fritz, U. (2017) Unexpected hybridization patterns in Near Eastern terrapins (Mauremys caspica, M. rivulata) indicate ancient gene flow across the Fertile Crescent. Zoologica Scripta, 46, 401 - 413. https: // doi. org / 10.1111 / zsc. 12227" type="journal article" year="2017">2017</bibRefCitation>
).
</paragraph>
<caption id="DF6F14B4C735FF9404B62D0BF617FB75" ID-DOI="http://doi.org/10.5281/zenodo.13746778" ID-Zenodo-Dep="13746778" httpUri="https://zenodo.org/record/13746778/files/figure.png" pageId="16" pageNumber="167" startId="16.[151,250,1120,1144]" targetBox="[151,1436,237,1097]" targetPageId="16" targetType="figure">
<paragraph id="8BAF443CC735FF9404B62D0BF617FB75" blockId="16.[151,1437,1120,1216]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404B62D0BF52AFBCD" bold="true" box="[151,290,1120,1144]" pageId="16" pageNumber="167">FIGURE 14.</emphasis>
Balkan terrapins,
<taxonomicName id="4C103FBFC735FF9405C32D0AF6ADFBCD" box="[482,677,1120,1144]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC735FF9405C32D0AF6ADFBCD" box="[482,677,1120,1144]" italics="true" pageId="16" pageNumber="167">Mauremys rivulata</emphasis>
</taxonomicName>
. (
<emphasis id="B964982EC735FF94069D2D0BF6C6FBCD" bold="true" box="[700,718,1120,1144]" pageId="16" pageNumber="167">A</emphasis>
) Al-Alani, upper Orontes (photo: H. Baniasi); (
<emphasis id="B964982EC735FF9400E02D0BF0DAFBCD" bold="true" box="[1217,1234,1120,1144]" pageId="16" pageNumber="167">B</emphasis>
) Masil Al-Fawwar, southwestern Syria (photo: Hans Esterbauer); (
<emphasis id="B964982EC735FF94064E2DEFF689FB29" bold="true" box="[623,641,1156,1180]" pageId="16" pageNumber="167">C</emphasis>
) Sadd Al-Kafat, southeast Hamah (photo: Ehab Jbr); (
<emphasis id="B964982EC735FF9400802DEFF0BBFB29" bold="true" box="[1185,1203,1156,1180]" pageId="16" pageNumber="167">D</emphasis>
) Bhalu, Nahr Al-Qash, east of Latakia (photo: Basel Barakat).
</paragraph>
</caption>
<paragraph id="8BAF443CC735FF9404E62C6FF693FA18" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404E62C6FF5BAFAAB" bold="true" box="[199,434,1284,1310]" pageId="16" pageNumber="167">Museum specimens</emphasis>
:
<collectingRegion id="49D48ADEC735FF9405E22C6EF60EFAAA" box="[451,518,1285,1311]" country="Syria" name="Hims" pageId="16" pageNumber="167">Homs</collectingRegion>
(
<collectionCode id="ED01DCF9C735FF9406362C6FF68BFAAB" box="[535,643,1284,1310]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="167" type="Museum">NHMUK</collectionCode>
1971.1861),
<collectingRegion id="49D48ADEC735FF94073A2C6FF765FAAB" box="[795,877,1284,1310]" country="Syria" name="Halab" pageId="16" pageNumber="167">Aleppo</collectingRegion>
(
<collectionCode id="ED01DCF9C735FF94075F2C6FF7B7FAAB" box="[894,959,1284,1310]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="16" pageNumber="167">NMB</collectionCode>
8297,
<collectionCode id="ED01DCF9C735FF9400302C6EF050FAAA" box="[1041,1112,1285,1311]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="167">NMW</collectionCode>
2201:1, 2212:1, 29656:1–2,
<collectionCode id="ED01DCF9C735FF9404B62C43F4E5FAF7" box="[151,237,1320,1346]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="167">MNHN</collectionCode>
5834A), Nahr Al-Dhahab (some of them putative hybrids:
<collectionCode id="ED01DCF9C735FF9407B82C43F7E8FAF7" box="[921,992,1320,1346]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="167">NMW</collectionCode>
29659:1–6, 29660:1–10), Nahr Qweiq (
<collectionCode id="ED01DCF9C735FF9404BE2C20F4E0FAD0" box="[159,232,1355,1381]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="167">NMW</collectionCode>
1741:1, 1746:1, 15082:1–2, 15084:1–3, 15098:1–3, 29655:1–5, 29657:1–3, 29658:1–4), Naseeb (
<collectionCode id="ED01DCF9C735FF9401722C20F194FAD0" box="[1363,1436,1355,1381]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="167">NMW</collectionCode>
18881:1–2), Housh Al-Admal (
<collectionCode id="ED01DCF9C735FF9405D72C04F638FA3C" box="[502,560,1391,1417]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="167">SMF</collectionCode>
75481), Mzeirib Lake (
<collectionCode id="ED01DCF9C735FF94071D2C04F77EFA3C" box="[828,886,1391,1417]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="167">SMF</collectionCode>
75487, 75489), Utaibeh (
<collectionCode id="ED01DCF9C735FF9400B72C04F10AFA3C" box="[1174,1282,1391,1417]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="167" type="Museum">NHMUK</collectionCode>
1909.4.20.1– 6), Yarmouk River (
<collectionCode id="ED01DCF9C735FF9405572CF9F5EAFA19" box="[374,482,1426,1452]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="167" type="Museum">NHMUK</collectionCode>
1864.8.23.156).
</paragraph>
<paragraph id="8BAF443CC735FF9404E62CDEF64FFA46" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404E62CDEF5DEFA65" bold="true" box="[199,470,1461,1488]" pageId="16" pageNumber="167">Citizen science records</emphasis>
: Al-Qatriyeh, Az-Zubar,
<collectingRegion id="49D48ADEC735FF9406CD2CDDF74BFA65" box="[748,835,1462,1488]" country="Syria" name="Al Ladhiqiyah" pageId="16" pageNumber="167">Latakia</collectingRegion>
, Psnada, Qismeen,
<quantity id="4CE8E9D9C735FF9400392CDDF058FA65" box="[1048,1104,1462,1488]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="167" unit="km" value="4.0">4 km</quantity>
N
<collectingRegion id="49D48ADEC735FF94004E2CDDF0CEFA65" box="[1135,1222,1462,1488]" country="Syria" name="Tartus" pageId="16" pageNumber="167">Tartous</collectingRegion>
, (iNaturalist 2023),
<collectingRegion id="49D48ADEC735FF9404B62CB1F531FA46" box="[151,313,1497,1524]" isEnumeration="true" pageId="16" pageNumber="167">Homs, Hamah</collectingRegion>
(observation.org 2023).
</paragraph>
<paragraph id="8BAF443CC735FF9404E62C96F549F9E8" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404E62C96F577F9A2" bold="true" box="[199,383,1533,1559]" pageId="16" pageNumber="167">Recent records</emphasis>
: Al-Alani, Al-Bassah, Al-Nawras Beach, Bhalu, Daraa,
<collectingRegion id="49D48ADEC735FF9400342C96F061F9A2" box="[1045,1129,1533,1559]" country="Syria" name="Hamah" pageId="16" pageNumber="167">Hamah</collectingRegion>
, Helfaya,
<collectingRegion id="49D48ADEC735FF9400C52C96F122F9A2" box="[1252,1322,1533,1559]" country="Syria" name="Hims" pageId="16" pageNumber="167">Homs</collectingRegion>
, Jisr ech-Choghur, Jourin,
<collectingRegion id="49D48ADEC735FF9405792F4BF5A7F98F" box="[344,431,1568,1594]" country="Syria" name="Al Ladhiqiyah" pageId="16" pageNumber="167">Latakia</collectingRegion>
, Nahr Al-Kabir Al-Shemali, Nahr Qweiq (
<collectingRegion id="49D48ADEC735FF9407A32F4BF7D2F98F" box="[898,986,1568,1594]" country="Syria" name="Halab" pageId="16" pageNumber="167">Aleppo</collectingRegion>
), Sadd Al-Kafat,
<collectingRegion id="49D48ADEC735FF9400B92F4AF0E7F98E" box="[1176,1263,1569,1595]" country="Syria" name="Tartus" pageId="16" pageNumber="167">Tartous</collectingRegion>
, Um et-Toyour, Ya’four, Zakia.
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C735FF9404E62F0CF7C3F8BA" pageId="16" pageNumber="167" type="distribution">
<paragraph id="8BAF443CC735FF9404E62F0CF7C3F8BA" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404E62F0CF5B5F934" bold="true" box="[199,445,1639,1665]" pageId="16" pageNumber="167">
Distribution in
<collectingCountry id="F30704ACC735FF94055C2F0CF5B5F934" box="[381,445,1639,1665]" name="Syria" pageId="16" pageNumber="167">Syria</collectingCountry>
</emphasis>
: Widespread and abundant. Its distribution in Syria extends from
<collectingRegion id="49D48ADEC735FF9400B52F0CF0EEF934" box="[1172,1254,1639,1665]" country="Syria" name="Halab" pageId="16" pageNumber="167">Aleppo</collectingRegion>
and the Orontes Basin over the coastal region to the surroundings of
<collectingRegion id="49D48ADEC735FF9406CE2FE0F76AF910" box="[751,866,1675,1701]" country="Syria" name="Dimashq" pageId="16" pageNumber="167">Damascus</collectingRegion>
and the southern region in Daraa and Golan (
<figureCitation id="132B58B9C735FF94014E2FE1F4BEF97C" captionStart="FIGURE 15" captionStartId="17.[151,250,1389,1413]" captionTargetBox="[161,1429,191,1356]" captionTargetId="figure-17@17.[151,1436,181,1365]" captionTargetPageId="17" captionText="FIGURE 15. Records of Mauremys rivulata in Syria. Black circles: previous records, blue circles: recently confirmed previous records, red circles: new records." figureDoi="http://doi.org/10.5281/zenodo.13746780" httpUri="https://zenodo.org/record/13746780/files/figure.png" pageId="16" pageNumber="167">Fig. 15</figureCitation>
). In the Sabkhat Al-Jabboul,
<taxonomicName id="4C103FBFC735FF9406252FC4F68CF97D" box="[516,644,1710,1736]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC735FF9406252FC4F68CF97D" box="[516,644,1710,1736]" italics="true" pageId="16" pageNumber="167">M. rivulata</emphasis>
</taxonomicName>
became extinct more than fifty years ago after the drying up of Nahr Al-Dhahab. According to morphology, some
<taxonomicName id="4C103FBFC735FF9406AD2FB9F709F95E" box="[652,769,1746,1771]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="genus">
<emphasis id="B964982EC735FF9406AD2FB9F709F95E" box="[652,769,1746,1771]" italics="true" pageId="16" pageNumber="167">Mauremys</emphasis>
</taxonomicName>
in the Nahr Al-Dahab showed traits of
<taxonomicName id="4C103FBFC735FF9400922FB9F126F95E" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[1203,1326,1746,1771]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC735FF9400922FB9F126F95E" box="[1203,1326,1746,1771]" italics="true" pageId="16" pageNumber="167">M. caspica</emphasis>
</taxonomicName>
, although they resembled
<taxonomicName id="4C103FBFC735FF9405692F9DF5CFF8BA" box="[328,455,1781,1807]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC735FF9405692F9DF5CFF8BA" box="[328,455,1781,1807]" italics="true" pageId="16" pageNumber="167">M. rivulata</emphasis>
</taxonomicName>
in gross morphology (
<bibRefCitation id="EF8139CDC735FF9406E52F9EF7B6F8BA" author="Fritz, U. &amp; Wischuf, T." box="[708,958,1781,1807]" pageId="16" pageNumber="167" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A17B7C735FF9404E62E73F598F874" pageId="16" pageNumber="167" type="discussion">
<paragraph id="8BAF443CC735FF9404E62E73F102F8CC" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
<emphasis id="B964982EC735FF9404E62E73F53CF887" bold="true" box="[199,308,1816,1842]" pageId="16" pageNumber="167">Remarks</emphasis>
: Based on morphology,
<bibRefCitation id="EF8139CDC735FF94066E2E73F755F887" author="Fritz, U. &amp; Wischuf, T." box="[591,861,1816,1842]" pageId="16" pageNumber="167" pagination="223 - 260" refId="ref15001" refString="Fritz, U. &amp; Wischuf, T. (1997) Zur Systematik westasiatisch - sudosteuropaischer Bachschildkroten (Gattung Mauremys) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen, 49, 223 - 260." type="journal article" year="1997">Fritz &amp; Wischuf (1997)</bibRefCitation>
reported a few
<taxonomicName id="4C103FBFC735FF9400372E72F083F887" box="[1046,1163,1817,1842]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="genus">
<emphasis id="B964982EC735FF9400372E72F083F887" box="[1046,1163,1817,1842]" italics="true" pageId="16" pageNumber="167">Mauremys</emphasis>
</taxonomicName>
with intermediate traits between
<taxonomicName id="4C103FBFC735FF9404DD2E56F570F8E3" baseAuthorityName="Gmelin" baseAuthorityYear="1774" box="[252,376,1853,1878]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="caspica">
<emphasis id="B964982EC735FF9404DD2E56F570F8E3" box="[252,376,1853,1878]" italics="true" pageId="16" pageNumber="167">M. caspica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C103FBFC735FF9405912E56F627F8E3" box="[432,559,1852,1878]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="rivulata">
<emphasis id="B964982EC735FF9405912E56F627F8E3" box="[432,559,1852,1878]" italics="true" pageId="16" pageNumber="167">M. rivulata</emphasis>
</taxonomicName>
from the Nahr Al-Dhahab (“Nahr ed Deheb”) in
<collectingCountry id="F30704ACC735FF94007D2E57F09FF8E3" box="[1116,1175,1852,1878]" name="Syria" pageId="16" pageNumber="167">Syria</collectingCountry>
(voucher specimens in the Naturhistorisches Museum Wien, NMW 29659:1–6, 29660:1–10). This population is now extinct.
</paragraph>
<paragraph id="8BAF443CC735FF9404E62EE8F598F874" blockId="16.[151,1437,1284,1985]" pageId="16" pageNumber="167">
GBIF contains 349 putative records for
<collectingCountry id="F30704ACC735FF9406B32EE8F6D9F828" box="[658,721,1923,1949]" name="Syria" pageId="16" pageNumber="167">Syria</collectingCountry>
. However, only 10 of these are located within
<collectingCountry id="F30704ACC735FF9400CA2EE8F122F828" box="[1259,1322,1923,1949]" name="Syria" pageId="16" pageNumber="167">Syria</collectingCountry>
; the other records lie in
<collectingCountry id="F30704ACC735FF9405112ECCF584F874" box="[304,396,1959,1985]" name="Turkey" pageId="16" pageNumber="167">Türkiye</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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