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<document id="938697F0F5CEC9D9F13EAD3C4FAC1884" ID-DOI="10.5070/P9401362375" ID-ISSN="0031-0298" ID-ZooBank="EE96562A-AEB9-4E4C-AAAF-036F41D8D3EF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712167387144" checkinUser="felipe" docAuthor="Kiernan, Caitlin R. &amp; Ebersole, Jun A." docDate="2023" docId="03964C6FFF9C672CFC4415608550FF6F" docLanguage="en" docName="PaleoBios.40.13.1-28.pdf" docOrigin="PaleoBios 40 (13)" docSource="http://dx.doi.org/10.5070/p9401362375" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Ectenosaurus Russell 1967" docType="treatment" docVersion="1" lastPageNumber="4" masterDocId="FFAF3417FF99672AFFAC1461872FFFB2" masterDocTitle="Two new plioplatecarpine mosasaurs (Mosasauridae; Plioplatecarpinae) of the genus Ectenosaurus from the Upper Cretaceous of North America" masterLastPageNumber="28" masterPageNumber="1" pageNumber="4" updateTime="1727997956931" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="D8EA95383D01CB061DF32B89196DBB29">Two new plioplatecarpine mosasaurs (Mosasauridae; Plioplatecarpinae) of the genus Ectenosaurus from the Upper Cretaceous of North America</mods:title>
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<mods:namePart id="F3175EDC66567F76A72AC3780358106E">Kiernan, Caitlin R.</mods:namePart>
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<mods:namePart id="6C3E877385CD5ABE6297578DC53F5AE1">Ebersole, Jun A.</mods:namePart>
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<mods:date id="8FE01652924FADEBB6CE7B888FD250C3">2023</mods:date>
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<treatment id="03964C6FFF9C672CFC4415608550FF6F" LSID="urn:lsid:plazi:treatment:03964C6FFF9C672CFC4415608550FF6F" httpUri="http://treatment.plazi.org/id/03964C6FFF9C672CFC4415608550FF6F" lastPageId="6" lastPageNumber="4" pageId="5" pageNumber="4">
<subSubSection id="C325AEF2FF9C672FFC4415608267FEAF" box="[1000,1352,257,285]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8B80FD79FF9C672FFC4415608267FEAF" blockId="5.[1000,1352,257,285]" box="[1000,1352,257,285]" pageId="5" pageNumber="4">
<taxonomicName id="4C3F86FAFF9C672FFC4415608267FEAF" authority="Russell, 1967" authorityName="Russell" authorityYear="1967" box="[1000,1352,257,285]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="B94B216BFF9C672FFC4415608385FEAF" box="[1000,1194,257,285]" italics="true" pageId="5" pageNumber="4">ECTENOSAURUS</emphasis>
<bibRefCitation id="EFAE8088FF9C672FFB1D15648267FEAF" author="Russell, D. A." box="[1201,1352,261,285]" pageId="5" pageNumber="4" pagination="1 - 241" refId="ref16943" refString="Russell, D. A. 1967. Systematics and morphology of American mosasaurs (Reptilia, Sauria). Bulletin of the Peabody Museum of Natural History 23: 1 - 241." type="journal article" year="1967">Russell, 1967</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C325AEF2FF9C672FFCCC155284BDFEC3" pageId="5" pageNumber="4" type="type_taxon">
<paragraph id="8B80FD79FF9C672FFCCC155284BDFEC3" blockId="5.[832,1521,305,1945]" pageId="5" pageNumber="4">
<emphasis id="B94B216BFF9C672FFCCC15528304FEFF" bold="true" box="[864,1067,307,333]" pageId="5" pageNumber="4">Generic Type―</emphasis>
<taxonomicName id="4C3F86FAFF9C672FFB87155084A2FEC3" authority="(Merriam, 1894)" baseAuthorityName="Merriam" baseAuthorityYear="1894" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="4" phylum="Chordata" rank="species" species="clidastoides">
<emphasis id="B94B216BFF9C672FFB8715508247FEFF" box="[1067,1384,305,333]" italics="true" pageId="5" pageNumber="4">Ectenosaurus clidastoides</emphasis>
(
<bibRefCitation id="EFAE8088FF9C672FFAD0155284AEFEC2" author="Merriam, J. C." pageId="5" pageNumber="4" pagination="1 - 39" refId="ref16134" refString="Merriam, J. C. 1894. Ueber die Pythonomorphen der Kansas- Kreide. Paleontographica 41: 1 - 39." type="journal article" year="1894">Merriam, 1894</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C325AEF2FF9C672FFCCC151882A1FC58" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8B80FD79FF9C672FFCCC1518831AFDF6" blockId="5.[832,1521,305,1945]" pageId="5" pageNumber="4">
<emphasis id="B94B216BFF9C672FFCCC15188375FE21" bold="true" box="[864,1114,377,403]" pageId="5" pageNumber="4">
Generic
<materialsCitation id="3B57F724FF9C672FFC671518831AFDF6" country="Germany" county="Niobrara Chalk of Logan County" location="Bayerishe Staatssammlung" municipality="Formerly" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Kansas" typeStatus="holotype">
<typeStatus id="548443DBFF9C672FFC6715188313FE21" box="[971,1084,377,403]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
<collectingMunicipality id="6BE46703FF9C672FFBF6151883E4FE21" box="[1114,1227,377,403]" pageId="5" pageNumber="4">Formerly</collectingMunicipality>
at
<location id="8EE0ABA2FF9C672FFB40151884A0FE04" LSID="urn:lsid:plazi:treatment:03964C6FFF9C672CFC4415608550FF6F:8EE0ABA2FF9C672FFB40151884A0FE04" country="Germany" county="Niobrara Chalk of Logan County" municipality="Formerly" name="Bayerishe Staatssammlung" pageId="5" pageNumber="4" stateProvince="Kansas">Bayerishe Staatssammlung</location>
für
<location id="8EE0ABA2FF9C672FFC6B15FD835EFE04" LSID="urn:lsid:plazi:treatment:03964C6FFF9C672CFC4415608550FF6F:8EE0ABA2FF9C672FFC6B15FD835EFE04" box="[967,1137,412,438]" country="Germany" county="Niobrara Chalk of Logan County" municipality="Formerly" name="Palaontologie" pageId="5" pageNumber="4" stateProvince="Kansas">Paläontologie</location>
(
<location id="8EE0ABA2FF9C672FFB2F15FD83CAFE04" LSID="urn:lsid:plazi:treatment:03964C6FFF9C672CFC4415608550FF6F:8EE0ABA2FF9C672FFB2F15FD83CAFE04" box="[1155,1253,412,438]" country="Germany" county="Niobrara Chalk of Logan County" municipality="Formerly" name="Munich" pageId="5" pageNumber="4" stateProvince="Kansas">Munich</location>
,
<collectingCountry id="F328BDE9FF9C672FFB5815FC8245FE05" box="[1268,1386,413,439]" name="Germany" pageId="5" pageNumber="4">Germany</collectingCountry>
), probably destroyed in
<location id="8EE0ABA2FF9C672FFC4215A183CCFE68" LSID="urn:lsid:plazi:treatment:03964C6FFF9C672CFC4415608550FF6F:8EE0ABA2FF9C672FFC4215A183CCFE68" box="[1006,1251,448,474]" country="Germany" county="Niobrara Chalk of Logan County" municipality="Formerly" name="Second World War" pageId="5" pageNumber="4" stateProvince="Kansas">Second World War</location>
, included &quot;posterior portion of the skull and several vertebrae&quot; collected from the
<collectingCounty id="62E185F5FF9C672FFC1916678266FD92" box="[949,1353,518,545]" pageId="5" pageNumber="4">Niobrara Chalk of Logan County</collectingCounty>
,
<collectingRegion id="49FB339BFF9C672FFAFA1666829FFD93" box="[1366,1456,519,545]" country="United States of America" name="Kansas" pageId="5" pageNumber="4">Kansas</collectingRegion>
, USA (
<bibRefCitation id="EFAE8088FF9C672FFCE71648830BFDF6" author="Russell, D. A." box="[843,1060,553,580]" pageId="5" pageNumber="4" pagination="1 - 241" refId="ref16943" refString="Russell, D. A. 1967. Systematics and morphology of American mosasaurs (Reptilia, Sauria). Bulletin of the Peabody Museum of Natural History 23: 1 - 241." type="journal article" year="1967">Russell 1967:158</bibRefCitation>
).
</materialsCitation>
</emphasis>
</paragraph>
<paragraph id="8B80FD79FF9C672FFCCC162D82A1FC58" blockId="5.[832,1521,305,1945]" pageId="5" pageNumber="4">
<emphasis id="B94B216BFF9C672FFCCC162D834DFDD5" bold="true" box="[864,1122,588,615]" pageId="5" pageNumber="4">
Generic
<typeStatus id="548443DBFF9C672FFC74162C836CFDD5" box="[984,1091,589,615]" pageId="5" pageNumber="4" type="neotype">Neotype</typeStatus>
</emphasis>
<collectionCode id="ED2E65BCFF9C672FFBCF162C8380FDD5" box="[1123,1199,589,615]" country="0" httpUri="http://grbio.org/cool/5bc7-5n0c" name="Fort Hays Sternberg Museum" pageId="5" pageNumber="4">FHSM</collectionCode>
<collectionCode id="ED2E65BCFF9C672FFB17162C83C0FDD5" box="[1211,1263,589,615]" pageId="5" pageNumber="4">VP-</collectionCode>
401, designated the
<typeStatus id="548443DBFF9C672FFCEC1611848AFD38" box="[832,933,624,650]" pageId="5" pageNumber="4" type="neotype">neotype</typeStatus>
by
<bibRefCitation id="EFAE8088FF9C672FFC61161183FFFD38" author="Willman, A. J. &amp; T. Konishi &amp; M. W. Caldwell" box="[973,1232,624,650]" pageId="5" pageNumber="4" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. (2021)</bibRefCitation>
, includes an essentially complete, articulated skull with both mandibles, articulated vertebral column to the 29th prepygal, ribs (with fragments of calcified costal cartilage), sternum, both scapulo-coracoids, and the majority of both forelimbs. It should be stressed that portions of the
<typeStatus id="548443DBFF9C672FFAA21741825DFC88" box="[1294,1394,800,826]" pageId="5" pageNumber="4" type="neotype">neotype</typeStatus>
have been heavily restored with plaster. Specimen was collected from the Smoky Hill Member of the Niobrara Chalk (upper Santonian-lower Campanian) at &quot;Garret Ranch, seven or eight miles northwest of Wakeeney, Trego County,
<collectingRegion id="49FB339BFF9C672FFCEC17B184B4FC58" box="[832,923,976,1002]" country="United States of America" name="Kansas" pageId="5" pageNumber="4">Kansas</collectingRegion>
&quot; (
<bibRefCitation id="EFAE8088FF9C672FFC1A17B183A0FC58" author="Russell, D. A." box="[950,1167,975,1002]" pageId="5" pageNumber="4" pagination="1 - 241" refId="ref16943" refString="Russell, D. A. 1967. Systematics and morphology of American mosasaurs (Reptilia, Sauria). Bulletin of the Peabody Museum of Natural History 23: 1 - 241." type="journal article" year="1967">Russell 1967:158</bibRefCitation>
,
<bibRefCitation id="EFAE8088FF9C672FFB3017B18252FC58" author="Schumacher, B." box="[1180,1405,976,1002]" pageId="5" pageNumber="4" refId="ref17191" refString="Schumacher, B. 1993. Biostratigraphy of Mosasauridae (Squamata, Varanoidea) from the Smoky Hill Member, Niobrara Chalk (Upper Cretaceous) of western Kansas. Master's Thesis. Fort Hays State University, Hays." type="book" year="1993">Schumacher 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C325AEF2FF9C672CFCCC17928550FF6F" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8B80FD79FF9C672CFCCC17928550FF6F" blockId="5.[832,1521,305,1945]" lastBlockId="6.[112,639,193,221]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<emphasis id="B94B216BFF9C672FFCCC179284FBFB82" bold="true" pageId="5" pageNumber="4">Generic Diagnosis (modified from Willman at al. 2021)―</emphasis>
Plioplatecarpine possessing the following apomorphies: premaxilla-maxilla suture extending posteriorly beyond posterior end of third maxillary tooth; maxillary and dentary tooth count each exceeding 12; prefrontal excluded from posterolateral margin of external nares by wing of maxilla; frontal emarginate above orbits; prefrontal and postorbitofrontal not in contact; parietal table wider than long; posteroventral process on jugal present; anterior border of jugal vertical ramus convex near jugal corner; 911 teeth in pterygoid; pterygoid tooth row anterior to ectopterygoid process straight; pterygoid teeth subequal in size; suprastapedial process medially excavated; posterolateral infrastapedial process of quadrate overlapping posteriorly onto distal terminus of surprastapedial process; stapedial pit narrowly rectangular; medial parapet of dentary higher than lateral wall; coronoid dorsal process extending posteriorly over free dorsal border of surangular, terminating acutely in profile; post-glenoid notch on dorsal rim of articular; at least 29 presacral, five pygal, and 28 intermediate caudal vertebrae; hypapophysis laterally compressed; anteroventral ridge extending from hypapophysis distinct, almost reaching cotyle; articular surface of pygal square in outline, not triangular; up to seven well ossified carpals. See
<bibRefCitation id="EFAE8088FF9C672FFCD9133C8384F8C5" author="Russell, D. A." box="[885,1195,1885,1912]" pageId="5" pageNumber="4" pagination="1 - 241" refId="ref16943" refString="Russell, D. A. 1967. Systematics and morphology of American mosasaurs (Reptilia, Sauria). Bulletin of the Peabody Museum of Natural History 23: 1 - 241." type="journal article" year="1967">Russell (1967:157158)</bibRefCitation>
for additional diagnostic postcranial characters, based on the
<typeStatus id="548443DBFF9C672FFAA0131E825DF82B" box="[1292,1394,1919,1945]" pageId="5" pageNumber="4" type="neotype">neotype</typeStatus>
specimen (
<collectionCode id="ED2E65BCFF9F672CFFD414A287E9FF6F" box="[120,198,195,221]" country="0" httpUri="http://grbio.org/cool/5bc7-5n0c" name="Fort Hays Sternberg Museum" pageId="6" pageNumber="5">FHSM</collectionCode>
<collectionCode id="ED2E65BCFF9F672CFF6014A287D2FF6F" box="[204,253,195,221]" pageId="6" pageNumber="5">VP-</collectionCode>
401,
<taxonomicName id="4C3F86FAFF9F672CFE9814A08541FF6F" baseAuthorityName="Merriam" baseAuthorityYear="1894" box="[308,622,193,221]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="5" phylum="Chordata" rank="species" species="clidastoides">
<emphasis id="B94B216BFF9F672CFE9814A08541FF6F" box="[308,622,193,221]" italics="true" pageId="6" pageNumber="5">Ectenosaurus clidastoides</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>