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<document id="FD5178B9CA3B20BAAD08FF354DA6B104" ID-DOI="10.11646/zootaxa.5523.5.1" ID-GBIF-Dataset="f466b472-1eeb-4caa-848d-6b7dbaef811e" ID-ISSN="1175-5326" ID-Zenodo-Dep="13949809" ID-ZooBank="710BF1DD-0C2C-426A-B6BE-A0BB54BFEB06" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729236455931" checkinUser="plazi" docAuthor="Reid, Chris A. M." docDate="2024" docId="0392420FA05EF838FF770D363EE3D4A1" docLanguage="en" docName="Zootaxa.5523.5.1.pdf" docOrigin="Zootaxa 5523 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleoptera" docType="treatment" docVersion="1" lastPageNumber="516" masterDocId="FFAB3A77A050F837FFE008113B7ED154" masterDocTitle="The chrysomeline leaf beetles of Timor, Wallacea (Coleoptera: Chrysomelidae: Chrysomelinae)" masterLastPageNumber="518" masterPageNumber="501" pageNumber="515" updateTime="1732042923365" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="51F8149013C9367BFB3077A18B11CC3B">The chrysomeline leaf beetles of Timor, Wallacea (Coleoptera: Chrysomelidae: Chrysomelinae)</mods:title>
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<treatment id="0392420FA05EF838FF770D363EE3D4A1" ID-DOI="http://doi.org/10.5281/zenodo.13974122" ID-GBIF-Taxon="244134646" ID-Zenodo-Dep="13974122" LSID="urn:lsid:plazi:treatment:0392420FA05EF838FF770D363EE3D4A1" httpUri="http://treatment.plazi.org/id/0392420FA05EF838FF770D363EE3D4A1" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515">
<subSubSection id="C321A092A05EF839FF770D363962D415" box="[151,540,1319,1345]" pageId="14" pageNumber="515" type="nomenclature">
<paragraph id="8B84F319A05EF839FF770D363962D415" blockId="14.[151,540,1319,1381]" box="[151,540,1319,1345]" pageId="14" pageNumber="515">
<heading id="D0CC4475A05EF839FF770D363962D415" bold="true" box="[151,540,1319,1345]" fontSize="11" level="1" pageId="14" pageNumber="515" reason="1">
<emphasis id="B94F2F0BA05EF839FF770D363962D415" bold="true" box="[151,540,1319,1345]" pageId="14" pageNumber="515">
Other
<taxonomicName id="4C3B889AA05EF839FF090D363AD8D415" box="[233,422,1319,1345]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Chrysomelinae</taxonomicName>
in Timor
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C321A092A05EF839FF770D5A3E2FD609" pageId="14" pageNumber="515" type="description">
<paragraph id="8B84F319A05EF839FF770D5A3B84D431" blockId="14.[151,540,1319,1381]" box="[151,250,1355,1381]" pageId="14" pageNumber="515">
(
<figureCitation id="1300EF9CA05EF839FF7F0D5A3B8FD431" box="[159,241,1355,1381]" captionStart="FIGURE 31" captionStartId="15.[151,250,725,749]" captionTargetBox="[633,954,196,689]" captionTargetId="figure-15@15.[622,966,181,701]" captionTargetPageId="15" captionText="FIGURE 31. Phola octodecimguttata (Fabricius, 1775), collected in Timor Leste." figureDoi="http://doi.org/10.5281/zenodo.13949839" httpUri="https://zenodo.org/record/13949839/files/figure.png" pageId="14" pageNumber="515">Fig. 31</figureCitation>
)
</paragraph>
<paragraph id="8B84F319A05EF839FF770D823E77D4A1" blockId="14.[151,1437,1427,1886]" pageId="14" pageNumber="515">
<taxonomicName id="4C3B889AA05EF839FF770D823957D4F9" authority="(Jacoby, 1884)" baseAuthorityName="Jacoby" baseAuthorityYear="1884" box="[151,553,1427,1453]" class="Insecta" family="Chrysomelidae" genus="Platymela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B94F2F0BA05EF839FF770D823A0AD4F9" box="[151,372,1427,1453]" italics="true" pageId="14" pageNumber="515">Platymela unicolor</emphasis>
(Jacoby, 1884)
</taxonomicName>
is endemic to Timor, but it was not found during the Australian Museum expedition. The host of two similar northern Australian species of
<taxonomicName id="4C3B889AA05EF839FC950DA63899D485" authorityName="Baly" authorityYear="1856" box="[885,999,1463,1489]" class="Insecta" family="Chrysomelidae" genus="Platymela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B94F2F0BA05EF839FC950DA63899D485" box="[885,999,1463,1489]" italics="true" pageId="14" pageNumber="515">Platymela</emphasis>
</taxonomicName>
is the small tree
<taxonomicName id="4C3B889AA05EF839FB400DA63F89D485" ID-CoL="34TL" box="[1184,1271,1463,1489]" class="Magnoliopsida" family="Sapindaceae" genus="Atalaya" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05EF839FB400DA63F89D485" box="[1184,1271,1463,1489]" italics="true" pageId="14" pageNumber="515">Atalaya</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3B889AA05EF839FAE50DA63EEBD485" ID-CoL="FY3" box="[1285,1429,1463,1489]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
) (
<bibRefCitation id="EFAA8EE8A05EF839FF7F0DCA3A66D4A1" author="Reid, C. A. M." box="[159,280,1499,1525]" pageId="14" pageNumber="515" pagination="1 - 119" refId="ref10459" refString="Reid, C. A. M. (2006) A taxonomic revision of the Australian Chrysomelinae, with a key to the genera (Coleoptera: Chrysomelidae). Zootaxa, 1292 (1), 1 - 119. https: // doi. org / 10.11646 / zootaxa. 1292.1.1" type="journal article" year="2006">Reid 2006</bibRefCitation>
), which occurs in Timor (
<bibRefCitation id="EFAA8EE8A05EF839FDD70DCA39BBD4A1" author="Cowie, I." box="[567,709,1499,1525]" pageId="14" pageNumber="515" refId="ref9514" refString="Cowie, I. (2006) A survey of flora and vegetation of the proposed Jaco-Tutuala-Lore National Park, Timor-Leste (East Timor). Report to Birdlife International from Northern Teritory Herbarium. Version 1.0. Palmerston, Northern Territory, 63 pp." type="book" year="2006">Cowie 2006</bibRefCitation>
), but was not sampled during the AMS expeditions.
</paragraph>
<paragraph id="8B84F319A05EF839FF270DEE39CAD7D1" blockId="14.[151,1437,1427,1886]" pageId="14" pageNumber="515">
<taxonomicName id="4C3B889AA05EF839FF270DEE39E9D74D" authority="(Linnaeus, 1763)" baseAuthorityName="Linnaeus" baseAuthorityYear="1763" box="[199,663,1535,1561]" class="Magnoliopsida" family="Campanulaceae" genus="Phyllocharis" kingdom="Plantae" order="Asterales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis id="B94F2F0BA05EF839FF270DEE3AB9D74D" box="[199,455,1535,1561]" italics="true" pageId="14" pageNumber="515">Phyllocharis undulata</emphasis>
(Linnaeus, 1763)
</taxonomicName>
was recorded from Timor by
<bibRefCitation id="EFAA8EE8A05EF839FBE50DEE3F80D74D" author="Mohamedsaid, M. S." box="[1029,1278,1535,1561]" pageId="14" pageNumber="515" refId="ref10011" refString="Mohamedsaid, M. S. (2009) Chrysomelidae of the Lesser Sunda Islands: Wallace's Line and the crossing of worlds. In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Vol. 2. Brill, Leiden, pp. 57 - 104. https: // doi. org / 10.1163 / ej. 9789004169470.1 - 299.21" type="book" year="2009">Mohamedsaid (2009)</bibRefCitation>
. This species was not collected on the Australian Museum expedition, probably because its host was not sampled. The host of
<taxonomicName id="4C3B889AA05EF839FF770E563AEAD735" baseAuthorityName="Linnaeus" baseAuthorityYear="1763" box="[151,404,1607,1633]" class="Magnoliopsida" family="Campanulaceae" genus="Phyllocharis" kingdom="Plantae" order="Asterales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis id="B94F2F0BA05EF839FF770E563AEAD735" box="[151,404,1607,1633]" italics="true" pageId="14" pageNumber="515">Phyllocharis undulata</emphasis>
</taxonomicName>
in
<collectingRegion id="49FF3DFBA05EF839FE5D0E563964D735" box="[445,538,1607,1633]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="515">Sumatra</collectingRegion>
is the coastal shrub
<taxonomicName id="4C3B889AA05EF839FCED0E563F79D735" ID-CoL="W3V5" box="[781,1031,1607,1633]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="species" species="inerme">
<emphasis id="B94F2F0BA05EF839FCED0E563F79D735" box="[781,1031,1607,1633]" italics="true" pageId="14" pageNumber="515">Clerodendrum inerme</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3B889AA05EF839FBF80E563FD7D735" ID-CoL="HST" box="[1048,1193,1607,1633]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
) (C. Reid,
<emphasis id="B94F2F0BA05EF839FACD0E593EEBD735" box="[1325,1429,1607,1633]" italics="true" pageId="14" pageNumber="515">pers. obs</emphasis>
. 2010), which is present on Timor (
<bibRefCitation id="EFAA8EE8A05EF839FDFA0E7A39D6D7D1" author="Cowie, I." box="[538,680,1643,1669]" pageId="14" pageNumber="515" refId="ref9514" refString="Cowie, I. (2006) A survey of flora and vegetation of the proposed Jaco-Tutuala-Lore National Park, Timor-Leste (East Timor). Report to Birdlife International from Northern Teritory Herbarium. Version 1.0. Palmerston, Northern Territory, 63 pp." type="book" year="2006">Cowie 2006</bibRefCitation>
).
</paragraph>
<paragraph id="8B84F319A05EF839FF270E9E3E2FD609" blockId="14.[151,1437,1427,1886]" pageId="14" pageNumber="515">
<taxonomicName id="4C3B889AA05EF839FF270E9E39D4D7FE" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[199,682,1679,1706]" class="Insecta" family="Chrysomelidae" genus="Phola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="octodecimguttata">
<emphasis id="B94F2F0BA05EF839FF270E9E3AA9D7FD" box="[199,471,1679,1705]" italics="true" pageId="14" pageNumber="515">Phola octodecimguttata</emphasis>
(Fabricius, 1775)
</taxonomicName>
, not previously known from Timor (
<bibRefCitation id="EFAA8EE8A05EF839FBBB0E9E3E3FD7FD" author="Mohamedsaid, M. S." box="[1115,1345,1679,1705]" pageId="14" pageNumber="515" refId="ref10011" refString="Mohamedsaid, M. S. (2009) Chrysomelidae of the Lesser Sunda Islands: Wallace's Line and the crossing of worlds. In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Vol. 2. Brill, Leiden, pp. 57 - 104. https: // doi. org / 10.1163 / ej. 9789004169470.1 - 299.21" type="book" year="2009">Mohamedsaid 2009</bibRefCitation>
;
<bibRefCitation id="EFAA8EE8A05EF839FAB10E813A5AD799" author="Lee, C. - F. &amp; Geiser, M. F." pageId="14" pageNumber="515" pagination="197 - 218" refId="ref9853" refString="Lee, C. - F. &amp; Geiser, M. F. (2023) Taxonomic review of the genus Phola Weise, 1890 (Coleoptera: Chrysomelidae: Chrysomelinae). Journal of Taiwan Agricultural Science, 72 (3), 197 - 218." type="journal article" year="2023">Lee &amp; Geiser 2023</bibRefCitation>
), was collected by the Australian Museum expedition on a coastal shrub,
<taxonomicName id="4C3B889AA05EF839FBBE0EA53F90D799" box="[1118,1262,1715,1741]" pageId="14" pageNumber="515">
<emphasis id="B94F2F0BA05EF839FBBE0EA53FEBD799" box="[1118,1173,1716,1741]" italics="true" pageId="14" pageNumber="515">Vitex</emphasis>
species
</taxonomicName>
(
<taxonomicName id="4C3B889AA05EF839FB1D0EA23EEFD799" ID-CoL="HST" box="[1277,1425,1715,1741]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
), probably
<taxonomicName id="4C3B889AA05EF839FEE10EC93A14D7A5" ID-CoL="5BLDY" box="[257,362,1751,1777]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="species" species="trifolia">
<emphasis id="B94F2F0BA05EF839FEE10EC93A14D7A5" box="[257,362,1751,1777]" italics="true" pageId="14" pageNumber="515">V. trifolia</emphasis>
</taxonomicName>
based on its distribution and habitat (
<bibRefCitation id="EFAA8EE8A05EF839FCE90EC638E1D7A5" author="de Kok, R. P. J." box="[777,927,1751,1777]" pageId="14" pageNumber="515" pagination="587 - 603" refId="ref9560" refString="de Kok, R. P. J. (2007) The genus Vitex L. (Lamiaceae) in New Guinea and the South Pacific Islands. Kew Bulletin, 62, 587 - 603." type="journal article" year="2007">de Kok 2007</bibRefCitation>
).
<taxonomicName id="4C3B889AA05EF839FC500EC63FC2D7A5" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[944,1212,1751,1777]" class="Insecta" family="Chrysomelidae" genus="Phola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="octodecimguttata">
<emphasis id="B94F2F0BA05EF839FC500EC63FC2D7A5" box="[944,1212,1751,1777]" italics="true" pageId="14" pageNumber="515">Phola octodecimguttata</emphasis>
</taxonomicName>
is known to feed on coastal
<taxonomicName id="4C3B889AA05EF839FF0C0EED3A5DD641" ID-CoL="8W4MP" box="[236,291,1788,1813]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05EF839FF0C0EED3A5DD641" box="[236,291,1788,1813]" italics="true" pageId="14" pageNumber="515">Vitex</emphasis>
</taxonomicName>
and is widespread from eastern
<collectingCountry id="F32CB389A05EF839FD700EEA3983D641" box="[656,765,1787,1813]" name="Indonesia" pageId="14" pageNumber="515">Indonesia</collectingCountry>
to
<collectingCountry id="F32CB389A05EF839FCC20EED38ACD641" box="[802,978,1787,1814]" name="New Caledonia" pageId="14" pageNumber="515">New Caledonia</collectingCountry>
, including coastal
<collectingCountry id="F32CB389A05EF839FB430EEA3E74D641" box="[1187,1290,1787,1813]" name="Australia" pageId="14" pageNumber="515">Australia</collectingCountry>
(
<bibRefCitation id="EFAA8EE8A05EF839FAF80EEA3EE9D642" author="Chen, S. H." box="[1304,1431,1787,1814]" pageId="14" pageNumber="515" pagination="465 - 475" refId="ref9382" refString="Chen, S. H. (1985) Phylogeny and classification of the Chrysomeloidea. Entomography, 3, 465 - 475." type="journal article" year="1985">Chen 1985</bibRefCitation>
; Reid &amp; Smith 2009;
<bibRefCitation id="EFAA8EE8A05EF839FE670F313919D66D" author="Lee, C. - F. &amp; Geiser, M. F." box="[391,615,1823,1850]" pageId="14" pageNumber="515" pagination="197 - 218" refId="ref9853" refString="Lee, C. - F. &amp; Geiser, M. F. (2023) Taxonomic review of the genus Phola Weise, 1890 (Coleoptera: Chrysomelidae: Chrysomelinae). Journal of Taiwan Agricultural Science, 72 (3), 197 - 218." type="journal article" year="2023">Lee &amp; Geiser 2023</bibRefCitation>
). Material examined (
<figureCitation id="1300EF9CA05EF839FC800F0E38C9D66D" box="[864,951,1823,1849]" captionStart="FIGURE 31" captionStartId="15.[151,250,725,749]" captionTargetBox="[633,954,196,689]" captionTargetId="figure-15@15.[622,966,181,701]" captionTargetPageId="15" captionText="FIGURE 31. Phola octodecimguttata (Fabricius, 1775), collected in Timor Leste." figureDoi="http://doi.org/10.5281/zenodo.13949839" httpUri="https://zenodo.org/record/13949839/files/figure.png" pageId="14" pageNumber="515">Fig. 31</figureCitation>
):
<specimenCount id="9D3D3890A05EF839FC2C0F0E3894D66D" box="[972,1002,1823,1849]" count="1" pageId="14" pageNumber="515" type="female">1♀</specimenCount>
/ Timor Leste, Lautem/Baucau border
<geoCoordinate id="EE0F95DEA05EF839FF770F523A73D60A" box="[151,269,1859,1886]" degrees="8" direction="south" minutes="25.596" orientation="latitude" pageId="14" pageNumber="515" precision="1" value="-8.4266">8°25.596S</geoCoordinate>
<geoCoordinate id="EE0F95DEA05EF839FEF40F523AD6D609" box="[276,424,1859,1885]" degrees="126" direction="east" minutes="41.617" orientation="longitude" pageId="14" pageNumber="515" precision="1" value="126.69362">126°41.617E</geoCoordinate>
acacia/herbs shingle beach
<taxonomicName id="4C3B889AA05EF839FD000F553869D609" ID-CoL="8W4MP" box="[736,791,1860,1885]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="515" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05EF839FD000F553869D609" box="[736,791,1860,1885]" italics="true" pageId="14" pageNumber="515">Vitex</emphasis>
</taxonomicName>
<quantity id="4CC35EFCA05EF839FCFE0F523820D609" box="[798,862,1859,1885]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="14" pageNumber="515" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2m</quantity>
<date id="FF85D5D9A05EF839FC850F5238A2D609" box="[869,988,1859,1885]" pageId="14" pageNumber="515" value="2011-11-10">10.xi.2011</date>
C Reid site 8/ K414869/ (AMS).
</paragraph>
</subSubSection>
<caption id="DF44A391A05FF838FF770AC438A7D3B9" ID-DOI="http://doi.org/10.5281/zenodo.13949839" ID-Zenodo-Dep="13949839" box="[151,985,725,749]" httpUri="https://zenodo.org/record/13949839/files/figure.png" pageId="15" pageNumber="516" startId="15.[151,250,725,749]" targetBox="[633,954,196,689]" targetPageId="15" targetType="figure">
<paragraph id="8B84F319A05FF838FF770AC438A7D3B9" blockId="15.[151,985,725,749]" box="[151,985,725,749]" pageId="15" pageNumber="516">
<emphasis id="B94F2F0BA05FF838FF770AC43A5ED3B9" bold="true" box="[151,288,725,749]" pageId="15" pageNumber="516">FIGURE 31.</emphasis>
<taxonomicName id="4C3B889AA05FF838FEC60AC439AED3B9" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[294,720,725,749]" class="Insecta" family="Chrysomelidae" genus="Phola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="octodecimguttata">
<emphasis id="B94F2F0BA05FF838FEC60AC43967D3B9" box="[294,537,725,749]" italics="true" pageId="15" pageNumber="516">Phola octodecimguttata</emphasis>
(Fabricius, 1775)
</taxonomicName>
, collected in Timor Leste.
</paragraph>
</caption>
<subSubSection id="C321A092A05FF838FF770B233EE3D4A1" pageId="15" pageNumber="516" type="key">
<paragraph id="8B84F319A05FF838FF770B2339FFD218" blockId="15.[151,641,818,844]" box="[151,641,818,844]" pageId="15" pageNumber="516">
<emphasis id="B94F2F0BA05FF838FF770B2339FFD218" bold="true" box="[151,641,818,844]" pageId="15" pageNumber="516">
Key to
<taxonomicName id="4C3B889AA05FF838FF110B233AD0D218" box="[241,430,818,844]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda">Chrysomelinae</taxonomicName>
species of Timor
</emphasis>
</paragraph>
<keyStep id="30CFEBBCA05FF838FF770B663EE2D2B0" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770B663EE2D2FE" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770B663EE2D2FE" pageId="15" pageNumber="516">
1. Procoxal cavities externally closed; elytra strongly patterned with permanent colours, with pale spots or blotches against a dark background (at least anterior half of epipleuron visible laterally; feeding on
<taxonomicName id="4C3B889AA05FF838FC5A0B853F4BD2FE" ID-CoL="HST" box="[954,1077,916,938]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
)............................ 2
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770BA33EE2D2B0" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770BA33EE2D2B0" pageId="15" pageNumber="516">
- Procoxal cavities externally wide open; dead insect without strongly patterned elytra (feeding on
<taxonomicName id="4C3B889AA05FF838FB680BA33F8DD293" ID-CoL="4F2P" box="[1160,1267,946,967]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05FF838FB680BA33F8DD293" box="[1160,1267,946,967]" italics="true" pageId="15" pageNumber="516">Eucalyptus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C3B889AA05FF838FAF20BA33EF2D29C" ID-CoL="FY3" box="[1298,1420,946,968]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
) ................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CFEBBCA05FF838FF770BFC3EE2D50D" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770BFC3EE3D54B" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770BFC3EE3D54B" pageId="15" pageNumber="516">
2(1) Elytra dark brown with creamy-yellow spots (about 8 on each); appendages brown; apical half of epipleura concealed laterally; on
<taxonomicName id="4C3B889AA05FF838FEE00C1B3A4ED54B" ID-CoL="8W4MP" box="[256,304,1034,1055]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05FF838FEE00C1B3A4ED54B" box="[256,304,1034,1055]" italics="true" pageId="15" pageNumber="516">Vitex</emphasis>
</taxonomicName>
...........................................................................
<taxonomicName id="4C3B889AA05FF838FB560C1B3EE3D54B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1206,1437,1034,1055]" class="Insecta" family="Chrysomelidae" genus="Phola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="octodecimguttata">
<emphasis id="B94F2F0BA05FF838FB560C1B3EE3D54B" box="[1206,1437,1034,1055]" italics="true" pageId="15" pageNumber="516">Phola octodecimguttata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770C393EE2D50D" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770C393EE2D50D" pageId="15" pageNumber="516">
- Elytra dark metallic blue with transverse orange blotches; appendages metallic bluish-black; epipleura entirely visible laterally; on
<taxonomicName id="4C3B889AA05FF838FEE00C553AF5D50D" ID-CoL="8VWBC" box="[256,395,1092,1113]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05FF838FEE00C553AF5D50D" box="[256,395,1092,1113]" italics="true" pageId="15" pageNumber="516">Clerodendrum</emphasis>
</taxonomicName>
....................................................................
<taxonomicName id="4C3B889AA05FF838FB260C553EE2D50D" baseAuthorityName="Linnaeus" baseAuthorityYear="1763" box="[1222,1436,1092,1113]" class="Magnoliopsida" family="Campanulaceae" genus="Phyllocharis" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis id="B94F2F0BA05FF838FB260C553EE2D50D" box="[1222,1436,1092,1113]" italics="true" pageId="15" pageNumber="516">Phyllocharis undulata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CFEBBCA05FF838FF770C733EE2D59B" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770C733EE2D5C0" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770C733EE2D5C0" pageId="15" pageNumber="516">
3(1) Epipleura wholly hidden in lateral view; body shape ovate, strongly convex in lateral view; clypeus not laterally ridged (host
<taxonomicName id="4C3B889AA05FF838FF020C6E3A33D5C0" ID-CoL="4F2P" box="[226,333,1151,1172]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05FF838FF020C6E3A33D5C0" box="[226,333,1151,1172]" italics="true" pageId="15" pageNumber="516">Eucalyptus</emphasis>
</taxonomicName>
).......................................................................................... 4
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770C8C3EE2D59B" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770C8C3EE2D59B" pageId="15" pageNumber="516">
- Epipleura wholly visible in lateral view; body partly parallel-sided, almost flat in lateral view; clypeus laterally ridged (host probably
<taxonomicName id="4C3B889AA05FF838FEDD0CAB3AF6D59B" ID-CoL="34TL" box="[317,392,1210,1231]" class="Magnoliopsida" family="Sapindaceae" genus="Atalaya" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F2F0BA05FF838FEDD0CAB3AF6D59B" box="[317,392,1210,1231]" italics="true" pageId="15" pageNumber="516">Atalaya</emphasis>
</taxonomicName>
).......................................................................
<taxonomicName id="4C3B889AA05FF838FB040CAB3EE2D59B" baseAuthorityName="Jacoby" baseAuthorityYear="1884" box="[1252,1436,1210,1231]" class="Insecta" family="Chrysomelidae" genus="Platymela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B94F2F0BA05FF838FB040CAB3EE2D59B" box="[1252,1436,1210,1231]" italics="true" pageId="15" pageNumber="516">Platymela unicolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CFEBBCA05FF838FF770CC63EE2D45E" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770CC63EE2D5B9" blockId="15.[151,1437,887,1525]" box="[151,1436,1239,1261]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770CC63EE2D5B9" box="[151,1436,1239,1261]" pageId="15" pageNumber="516">
4(3) Larger, length&gt;
<quantity id="4CC35EFCA05FF838FE9D0CC63AC8D5B9" box="[381,438,1239,1261]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="516" unit="mm" value="8.0">8 mm</quantity>
; elytra partly dark brown, not striate.................................................... 5
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770CE43EE2D45E" blockId="15.[151,1437,887,1525]" box="[151,1436,1268,1290]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770CE43EE2D45E" box="[151,1436,1268,1290]" pageId="15" pageNumber="516">
- Smaller, length &lt;
<quantity id="4CC35EFCA05FF838FE670CE43ABFD45E" box="[391,449,1269,1290]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="516" unit="mm" value="7.0">7 mm</quantity>
; elytra pale brown, striate............................................................ 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CFEBBCA05FF838FF770D033EE2D4D4" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770D033EE2D411" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770D033EE2D411" pageId="15" pageNumber="516">
5(4) Larger, length about
<quantity id="4CC35EFCA05FF838FE490D033A91D473" box="[425,495,1298,1319]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="516" unit="mm" value="12.0">12 mm</quantity>
; body strongly rounded at sides and convex in profile, height at least half length; elytral punctures irregularly linked by small grooves (acervate); prosternum with pair of lateral flanges at base of process....
<taxonomicName id="4C3B889AA05FF838FB1C0D213EE2D411" box="[1276,1436,1327,1349]" pageId="15" pageNumber="516">
<emphasis id="B94F2F0BA05FF838FB1C0D213E2ED411" box="[1276,1360,1328,1349]" italics="true" pageId="15" pageNumber="516">Paropsis</emphasis>
species
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770D5C3EE2D4D4" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770D5C3EE2D4D4" pageId="15" pageNumber="516">
- Smaller, length about
<quantity id="4CC35EFCA05FF838FE530D5C3A87D436" box="[435,505,1357,1378]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="516" unit="mm" value="10.0">10 mm</quantity>
; basal half of elytra almost parallel-sided, profile flatter, height much less than half length; elytral interspaces smooth; prosternum without flanges................................
<taxonomicName id="4C3B889AA05FF838FBCE0D7B3E5DD42B" authorityName="Reid" authorityYear="2024" box="[1070,1315,1386,1407]" class="Insecta" family="Chrysomelidae" genus="Paropsisterna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="tatamailau" status="sp. nov.">
<emphasis id="B94F2F0BA05FF838FBCE0D7B3E5DD42B" box="[1070,1315,1386,1407]" italics="true" pageId="15" pageNumber="516">Paropsisterna tatamailau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C9270A05FF838FAC90D7B3EE2D4D4" box="[1321,1436,1386,1408]" pageId="15" pageNumber="516" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CFEBBCA05FF838FF770D963EE3D4A1" pageId="15" pageNumber="516">
<paragraph id="8B84F319A05FF838FF770D963EE2D4EE" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770D963EE2D4EE" pageId="15" pageNumber="516">
6(4) Elytral striae strongly punctured, 1 &amp; 2 irregular, not entirely linear; base of stria 5 obliquely grooved; mesoventrite process narrow..............................................................................
<taxonomicName id="4C3B889AA05FF838FB2A0DB43EE2D4EE" box="[1226,1436,1444,1466]" pageId="15" pageNumber="516">
<emphasis id="B94F2F0BA05FF838FB2A0DB43E2FD4EE" box="[1226,1361,1445,1466]" italics="true" pageId="15" pageNumber="516">Paropsisterna</emphasis>
species
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B84F319A05FF838FF770DD33EE3D4A1" blockId="15.[151,1437,887,1525]" pageId="15" pageNumber="516">
<keyLead id="30CA502CA05FF838FF770DD33EE3D4A1" pageId="15" pageNumber="516">
- Elytral striae weakly punctured, 1 &amp; 2 regular, linear; base of stria 5 not grooved; mesoventrite process broad..................................................................................................
<taxonomicName id="4C3B889AA05FF838FB0C0DF13EE3D4A1" authorityName="Reid" authorityYear="2024" box="[1260,1437,1504,1525]" class="Insecta" family="Chrysomelidae" genus="Paropsisterna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="tais">
<emphasis id="B94F2F0BA05FF838FB0C0DF13EE3D4A1" box="[1260,1437,1504,1525]" italics="true" pageId="15" pageNumber="516">Paropsisterna tais</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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