566 lines
75 KiB
XML
566 lines
75 KiB
XML
<document id="A529306EDF775926FA953E66CAEF348F" ID-DOI="10.26107/RBZ-2024-0035" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682894" ID-ZooBank="227BB34E-7FF6-492E-A3E3-F82B74C714B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171811771" checkinUser="felipe" docAuthor="Kottelat, Maurice, , Tan Heok Hui, Hasan, & Veryl" docDate="2024" docId="038D87D4AD25FFD6E76AFAF6FC02A267" docLanguage="en" docName="RafflesBZool.72.479-487.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Kapuasia falaris Kottelat, , Hasan & Veryl, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="486" masterDocId="FFB4FFACAD24FFD1E660FFDDFFAFAA52" masterDocTitle="Kapuasia falaris, a new species of nemacheilid loach from Borneo (Teleostei: Nemacheilidae)" masterLastPageNumber="487" masterPageNumber="479" pageNumber="480" updateTime="1737175458869" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="6585F57B88D69258AC0FD96D4A6DA3B0">
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<mods:title id="68DD995FD50520C7051E103E9A26CA7E">Kapuasia falaris, a new species of nemacheilid loach from Borneo (Teleostei: Nemacheilidae)</mods:title>
|
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</mods:titleInfo>
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<mods:name id="08AF3EE6A68E79205004E390F3D4272C" type="personal">
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<mods:role id="4F5ED3A85C1F05B5A885325260ECB354">
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<mods:roleTerm id="4B8837BD27227956CE41EEFAA49B57BD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="605E6F5A3742880BBE1C1DEF49639D1F">Kottelat, Maurice</mods:namePart>
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</mods:name>
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<mods:name id="316C51C062A22B8433E5CBBB06DEDE2D" type="personal">
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<mods:role id="B22B3901151B77809712F4AE55BAB263">
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<mods:roleTerm id="E0E0644F82177AC888CB29C60D12053A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5B8974F12DECE98224AC991351A851CA">, Tan Heok Hui</mods:namePart>
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</mods:name>
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<mods:name id="4CF9AC02E4CA560408DE01DDF4C1C6F7" type="personal">
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<mods:role id="275055911E998D61657C07499D8EACA9">
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<mods:roleTerm id="40741BE1A05D4551A6694655BD653EB9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AC40D3C0E508A944C6038E3E5C43568C">Hasan,</mods:namePart>
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||
</mods:name>
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<mods:name id="9F82B14478B87CAEFA589CCE4050A884" type="personal">
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<mods:role id="3E07BA6AFADC78DE39D1CA20D05F0EC7">
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<mods:roleTerm id="F547BB55ECC644FC076F21BF56A756FD">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="6C78F63CCB7FE6EB12128149F6264D08">Veryl</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A2104735350FD43B756E76F1BA7748DC">text</mods:typeOfResource>
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<mods:titleInfo id="3F3A21C6511FEE0929359DD88FDC46EC">
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<mods:title id="D2414BFA6F3890A4A632F3BAD39DFCA9">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="0DD0201A80C2E5685ED25ED1C335B381">
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<mods:date id="79ED834B7AF8BCF1E3DA8803A16E0691">2024</mods:date>
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<mods:detail id="84DB9F84C615D8C3AA853E4BA97A7481" type="pubDate">
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<mods:number id="50D056F16E36674234CFC41E790A45D3">2024-12-02</mods:number>
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</mods:detail>
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<mods:detail id="54636E514D6B414085BC40C68F96A0E4" type="volume">
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<mods:number id="F70B7ED10258C270158DAC4651E1A813">72</mods:number>
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<mods:extent id="722D8E1DD8062E4E4CFFFEC325E2DAC0" unit="page">
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<mods:start id="068C81BBFCAD73508A0DE672C8A29618">479</mods:start>
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<mods:end id="052280D686A4C676D3D1C992F4AD7ED6">487</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="41ADBE1DCFC50185AE7E63F4CD2ED17D">journal article</mods:classification>
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<mods:identifier id="4473115AA0E0E67CF387E20746D812A4" type="DOI">10.26107/RBZ-2024-0035</mods:identifier>
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<mods:identifier id="F8153ED5801D6B212D77F581F5215D64" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="9F6BC8BF3664F7B1BA0E020C378B99FB" type="Zenodo-Dep">14682894</mods:identifier>
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</mods:mods>
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<treatment id="038D87D4AD25FFD6E76AFAF6FC02A267" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267" httpUri="http://treatment.plazi.org/id/038D87D4AD25FFD6E76AFAF6FC02A267" lastPageId="7" lastPageNumber="486" pageId="1" pageNumber="480">
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<subSubSection id="C33E6549AD25FFD0E76AFAF6FDF8AF11" box="[266,599,1323,1347]" pageId="1" pageNumber="480" type="nomenclature">
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<paragraph id="8B9B36C2AD25FFD0E76AFAF6FDF8AF11" blockId="1.[266,599,1323,1379]" box="[266,599,1323,1347]" pageId="1" pageNumber="480">
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<heading id="D0D381AEAD25FFD0E76AFAF6FDF8AF11" bold="true" box="[266,599,1323,1347]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="480" reason="2">
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||
<emphasis id="B950EAD0AD25FFD0E76AFAF6FDF8AF11" bold="true" box="[266,599,1323,1347]" pageId="1" pageNumber="480">
|
||
<taxonomicName id="4C244D41AD25FFD0E76AFAF6FE6DAF11" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[266,450,1323,1347]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris" status="sp. nov.">
|
||
<emphasis id="B950EAD0AD25FFD0E76AFAF6FE6DAF11" bold="true" box="[266,450,1323,1347]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357ABAD25FFD0E7B1FAF6FDF8AF11" box="[465,599,1323,1347]" pageId="1" pageNumber="480" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD25FFD0E715FA96FE43AF31" box="[373,492,1355,1379]" pageId="1" pageNumber="480" type="description">
|
||
<paragraph id="8B9B36C2AD25FFD0E715FA96FE43AF31" blockId="1.[266,599,1323,1379]" box="[373,492,1355,1379]" pageId="1" pageNumber="480">
|
||
(
|
||
<figureCitation id="131F2A47AD25FFD0E71DFA96FE4CAF31" box="[381,483,1355,1379]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Figs. 1–3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD25FFD0E611FA56FE89AD71" pageId="1" pageNumber="480" type="materials_examined">
|
||
<paragraph id="8B9B36C2AD25FFD0E611FA56FDCDAC51" blockId="1.[113,752,1419,1540]" pageId="1" pageNumber="480">
|
||
<materialsCitation id="3B4C3C9FAD25FFD0E611FA56FDCDAC51" collectingDate="2024-01-03" collectionCode="MZB" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="MZB 17245" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="holotype">
|
||
<emphasis id="B950EAD0AD25FFD0E611FA56FF4AAFF1" bold="true" box="[113,229,1419,1443]" pageId="1" pageNumber="480">
|
||
<typeStatus id="549F8860AD25FFD0E611FA56FF4FAFF1" box="[113,224,1419,1443]" pageId="1" pageNumber="480" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DB829EB9AD25FFD0E690FA51FED1AFF1" box="[240,382,1419,1443]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="480" type="Museum">MZB 17245</specimenCode>
|
||
, male,
|
||
<quantity id="4CDC9B27AD25FFD0E7B6FA56FD93AFF1" box="[470,572,1419,1443]" metricMagnitude="-2" metricUnit="m" metricValue="5.1899999999999995" pageId="1" pageNumber="480" unit="mm" value="51.9">51.9 mm</quantity>
|
||
SL;
|
||
<collectingCountry id="F3337652AD25FFD0E41BFA56FD43AFF1" box="[635,748,1419,1443]" name="Indonesia" pageId="1" pageNumber="480">Indonesia</collectingCountry>
|
||
:
|
||
<collectingCounty id="62FA4E4EAD25FFD0E611FA71FF65AF91" box="[113,202,1452,1475]" pageId="1" pageNumber="480">Borneo</collectingCounty>
|
||
:
|
||
<collectingRegion id="49E0F820AD25FFD0E6BAFA76FE69AF91" box="[218,454,1451,1475]" country="Indonesia" name="Kalimantan Selatan" pageId="1" pageNumber="480">Kalimantan Selatan</collectingRegion>
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||
:
|
||
<collectingMunicipality id="6BFFACB8AD25FFD0E7B5FA76FDB2AF91" box="[469,541,1451,1475]" pageId="1" pageNumber="480">Barito</collectingMunicipality>
|
||
basin, kabupaten
|
||
<location id="8EFB6019AD25FFD0E612FA16FF1CAFB1" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E612FA16FF1CAFB1" box="[114,179,1483,1507]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Tapin" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Tapin</location>
|
||
, kecamatan
|
||
<location id="8EFB6019AD25FFD0E756FA16FEDEAFB1" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E756FA16FEDEAFB1" box="[310,369,1483,1507]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Piani" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Piani</location>
|
||
: desa
|
||
<location id="8EFB6019AD25FFD0E7D1FA16FD9DAFB6" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E7D1FA16FD9DAFB6" box="[433,562,1483,1508]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Pipitak Jaya" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Pipitak Jaya</location>
|
||
[about
|
||
<geoCoordinate id="EE105005AD25FFD0E4E3FA17FD5FAFB1" box="[643,752,1482,1507]" degrees="2" direction="south" minutes="57" orientation="latitude" pageId="1" pageNumber="480" precision="15" seconds="13" value="-2.9536111">2°57′13″S</geoCoordinate>
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||
<geoCoordinate id="EE105005AD25FFD0E611FA37FEAFAC51" box="[113,256,1514,1539]" degrees="115" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="480" precision="15" seconds="38" value="115.327225">115°19′38″E</geoCoordinate>
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||
];
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||
<location id="8EFB6019AD25FFD0E774FA31FE08AC51" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E774FA31FE08AC51" box="[276,423,1515,1539]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Noor Aminin" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Noor Aminin</location>
|
||
,
|
||
<date id="FF9A1002AD25FFD0E7D4FA36FDF1AC51" box="[436,606,1515,1540]" pageId="1" pageNumber="480" value="2024-01-03">
|
||
<collectingDate id="EFDEE9EAAD25FFD0E7D4FA36FDF1AC51" box="[436,606,1515,1540]" pageId="1" pageNumber="480" value="2024-01-03">3 January 2024</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD25FFD0E611F9F6FF53ACF1" blockId="1.[113,751,1579,1699]" pageId="1" pageNumber="480">
|
||
<materialsCitation id="3B4C3C9FAD25FFD0E611F9F6FDECAC31" collectingDate="2024-01-03" collectionCode="ZRC" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="ZRC 65844, 8" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="paratype">
|
||
<emphasis id="B950EAD0AD25FFD0E611F9F6FF45AC11" bold="true" box="[113,234,1579,1603]" pageId="1" pageNumber="480">
|
||
<typeStatus id="549F8860AD25FFD0E611F9F6FF4AAC11" box="[113,229,1579,1603]" pageId="1" pageNumber="480" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DB829EB9AD25FFD0E691F9F6FEDFAC11" box="[241,368,1579,1603]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65844</specimenCode>
|
||
,
|
||
<specimenCode id="DB829EB9AD25FFD0E71BF9F6FE25AC11" box="[379,394,1579,1603]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">8</specimenCode>
|
||
, 43.0–66.0 mm SL; CMK 29184, 5,
|
||
<quantity id="4CDC9B27AD25FFD0E6EBF996FE8AAC31" box="[139,293,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="4.99" metricValueMax="5.66" metricValueMin="4.32" pageId="1" pageNumber="480" unit="mm" value="49.9" valueMax="56.6" valueMin="43.2">43.2–56.6 mm</quantity>
|
||
SL; same data as holotype
|
||
</materialsCitation>
|
||
. —
|
||
<materialsCitation id="3B4C3C9FAD25FFD0E40CF996FF56ACF1" collectingDate="2024-03-04" collectionCode="ZRC" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="ZRC 65998, 12" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="paratype">
|
||
<specimenCode id="DB829EB9AD25FFD0E40CF996FD44AC31" box="[620,747,1611,1635]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65998</specimenCode>
|
||
,
|
||
<specimenCode id="DB829EB9AD25FFD0E611F9B6FF21ACD1" box="[113,142,1643,1667]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">12</specimenCode>
|
||
, 37.0–
|
||
<quantity id="4CDC9B27AD25FFD0E6B9F9B6FE93ACD1" box="[217,316,1643,1667]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="1" pageNumber="480" unit="mm" value="59.2">59.2 mm</quantity>
|
||
SL; same location data as
|
||
<typeStatus id="549F8860AD25FFD0E411F9B6FD7CACD1" box="[625,723,1643,1667]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
|
||
;
|
||
<date id="FF9A1002AD25FFD0E481F9B6FF56ACF1" pageId="1" pageNumber="480" value="2024-03-04">
|
||
<collectingDate id="EFDEE9EAAD25FFD0E481F9B6FF56ACF1" pageId="1" pageNumber="480" value="2024-03-04">4 March 2024</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD25FFD0E611F916FE89AD71" blockId="1.[112,751,1739,1827]" pageId="1" pageNumber="480">
|
||
<materialsCitation id="3B4C3C9FAD25FFD0E611F916FE8EAD71" collectingDate="2024-02-14" collectionCode="ZRC" pageId="1" pageNumber="480" specimenCode="ZRC 65915, 46" specimenCount="2">
|
||
<emphasis id="B950EAD0AD25FFD0E611F916FE93ACB1" bold="true" box="[113,316,1739,1763]" pageId="1" pageNumber="480">
|
||
<typeStatus id="549F8860AD25FFD0E611F916FE93ACB1" box="[113,316,1739,1763]" isOtherTypeStatus="true" pageId="1" pageNumber="480">Non-type material</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="DB829EB9AD25FFD0E728F916FE6BACB1" box="[328,452,1739,1763]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65915</specimenCode>
|
||
,
|
||
<specimenCode id="DB829EB9AD25FFD0E7ADF916FE45ACB1" box="[461,490,1739,1763]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">46</specimenCode>
|
||
,
|
||
<quantity id="4CDC9B27AD25FFD0E793F916FD26ACB1" box="[499,649,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="5.005" metricValueMax="5.74" metricValueMin="4.27" pageId="1" pageNumber="480" unit="mm" value="50.05" valueMax="57.4" valueMin="42.7">42.7–57.4 mm</quantity>
|
||
SL; same location data as
|
||
<typeStatus id="549F8860AD25FFD0E740F936FE2EAD51" box="[288,385,1771,1795]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
|
||
;
|
||
<date id="FF9A1002AD25FFD0E7ECF936FDE4AD51" box="[396,587,1771,1795]" pageId="1" pageNumber="480" value="2024-02-14">
|
||
<collectingDate id="EFDEE9EAAD25FFD0E7ECF936FDE4AD51" box="[396,587,1771,1795]" pageId="1" pageNumber="480" value="2024-02-14">14 February 2024</collectingDate>
|
||
</date>
|
||
[specimens all dead on arrival]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD25FFD0E611F896FAB4AD71" pageId="1" pageNumber="480" type="diagnosis">
|
||
<paragraph id="8B9B36C2AD25FFD0E611F896FAB4AD71" blockId="1.[113,753,1867,2083]" lastBlockId="1.[811,1453,1259,1828]" pageId="1" pageNumber="480">
|
||
<emphasis id="B950EAD0AD25FFD0E611F896FF40AD31" bold="true" box="[113,239,1867,1891]" pageId="1" pageNumber="480">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C244D41AD25FFD0E69BF891FE6EAD31" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[251,449,1867,1891]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD25FFD0E69BF891FE6EAD31" box="[251,449,1867,1891]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C244D41AD25FFD0E4B6F891FF42ADD1" authorityName="Roberts" authorityYear="1989" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD25FFD0E4B6F891FF42ADD1" italics="true" pageId="1" pageNumber="480">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: colour pattern made of 3–7 regular black bars, wider than interspaces and extending across back, of constant width and reaching ventral midline (vs. irregular bars or mostly triangular saddles reaching downward about to lateral line, often alternating or connected with blotches in lower half of body); head black, except for white dorsal side of the snout, a bar behind eye and a bar across the nape and along hind margin of opercle (vs. top of head and cheek spotted); in males, presence of a suborbital flap (
|
||
<figureCitation id="131F2A47AD25FFD0E2A9FA96FAA0AF36" box="[1225,1295,1355,1380]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="1" pageNumber="480">Fig. 4</figureCitation>
|
||
; vs. absence); in males, vertical skin flanges on the dorsal surface of the pectoral fin, between the branched rays (
|
||
<figureCitation id="131F2A47AD25FFD0E2B7FA56FAB8AFF1" box="[1239,1303,1419,1443]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="1" pageNumber="480">Fig. 5</figureCitation>
|
||
; vs. absence); no large flat pectoral axillary lobe (presence), but instead a cylindrical lobe, with pointed posterior extremity, usually separate from skin of flank by a deep groove, often with a second similar lobe above the groove (
|
||
<figureCitation id="131F2A47AD25FFD0E2ABF9D6FAA2AC71" box="[1227,1293,1547,1571]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="1" pageNumber="480">Fig. 6</figureCitation>
|
||
). The shape of this lobe is apparently unique among Southeast Asian species of nemacheilids.
|
||
<taxonomicName id="4C244D41AD25FFD0E58CF991FB0CAC31" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[1004,1187,1611,1635]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD25FFD0E58CF991FB0CAC31" box="[1004,1187,1611,1635]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from all nemacheilids from Southeast Asia by the shape of the suborbital flap: globulous, at some distance below the eye, and with a deep vertical groove from the flap to the eye; and the strongly upwards curled pectoral fin, appearing with an almost vertical orientation in life (
|
||
<figureCitation id="131F2A47AD25FFD0E2A6F936FAA7AD56" box="[1222,1288,1771,1796]" captionStart="Fig" captionStartId="4.[135,170,1513,1535]" captionTargetBox="[135,1473,189,1497]" captionTargetId="figure-7@4.[135,1473,189,1497]" captionTargetPageId="4" captionText="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Fig. 3</figureCitation>
|
||
), is apparently unique among Southeast Asian nemacheilids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD25FFD6E54DF896FD5CAC64" lastPageId="7" lastPageNumber="486" pageId="1" pageNumber="480" type="description">
|
||
<paragraph id="8B9B36C2AD25FFD3E54DF896FEE0ACBE" blockId="1.[813,1453,1867,2083]" lastBlockId="2.[135,776,1524,1772]" lastPageId="2" lastPageNumber="481" pageId="1" pageNumber="480">
|
||
<emphasis id="B950EAD0AD25FFD0E54DF896FC18AD31" bold="true" box="[813,951,1867,1891]" pageId="1" pageNumber="480">Description.</emphasis>
|
||
See
|
||
<figureCitation id="131F2A47AD25FFD0E588F896FBE6AD36" box="[1000,1097,1867,1892]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Figs. 1–3</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="C6A60379AD25FFD0E30FF896FC95ADD6" captionStart="Table 1" captionStartId="2.[135,190,189,211]" captionText="Table 1. Morphometric data of type material of Kapuasia falaris (n=10). Ranges and means include holotype data." pageId="1" pageNumber="480">Table 1</tableCitation>
|
||
for morphometric data of
|
||
<typeStatus id="549F8860AD25FFD0E205F8B6FB6DADD1" box="[1125,1218,1899,1923]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D22FD4BAD25FFD0E29DF8B1FAD1ADD6" box="[1277,1406,1900,1924]" count="9" pageId="1" pageNumber="480" type="generic" typeStatus="paratype">9 paratypes</specimenCount>
|
||
. An elongate nemacheilid with body depth gradually increasing up to dorsal-fin origin. Behind dorsal fin, body depth almost uniform until caudal-fin base. Dorsal profile continuous between head and body. Head slightly depressed; body slightly compressed anteriorly, gradually more compressed posteriorly. Interorbital area flat. In lateral view, eye protruding over dorsal profile of head. Cheeks not swollen. Snout pointed but not acute. Depth of caudal peduncle 1.3–1.5 times in its length, depth uniform. No dorsal crest or keel on post-dorsal area. No ventral crest or keel on caudal peduncle. Vertebrae 18–20+13–14=32–34 (modally 19+14=33). Largest recorded size
|
||
<quantity id="4CDC9B27AD26FFD3E794F969FDFDAC9E" box="[500,594,1716,1740]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" pageId="2" pageNumber="481" unit="mm" value="57.5">57.5 mm</quantity>
|
||
SL in male, 66.0 mm SL in female.
|
||
</paragraph>
|
||
<caption id="DF5B664AAD26FFD3E6E7FF60FB5DAA81" box="[135,1266,189,211]" pageId="2" pageNumber="481" startId="2.[135,190,189,211]" targetType="table">
|
||
<paragraph id="8B9B36C2AD26FFD3E6E7FF60FB5DAA81" blockId="2.[135,1266,189,211]" box="[135,1266,189,211]" pageId="2" pageNumber="481">
|
||
Table 1. Morphometric data of type material of
|
||
<taxonomicName id="4C244D41AD26FFD3E401FF63FCABAA80" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[609,772,189,211]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="481" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD26FFD3E401FF63FCABAA80" box="[609,772,189,211]" italics="true" pageId="2" pageNumber="481">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
(n=10). Ranges and means include holotype data.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD26FFD3E564FED8FAE4AFF4" pageId="2" pageNumber="481">
|
||
<table id="F924C462AD26002EE6F6FED8FAF3AFF4" box="[150,1372,261,1446]" gridcols="4" gridrows="13" pageId="2" pageNumber="481">
|
||
<tr id="35143480AD26002EE6F6FED8FAF3AB48" box="[150,1372,261,282]" gridrow="0" pageId="2" pageNumber="481" rowspan-0="1">
|
||
<th id="76C55DFCAD26002EE564FED8FCCEAB48" box="[772,865,261,282]" gridcol="1" gridrow="0" pageId="2" pageNumber="481">
|
||
<emphasis id="B950EAD0AD26FFD3E564FED8FCCEAB48" bold="true" box="[772,865,261,282]" pageId="2" pageNumber="481">Holotype</emphasis>
|
||
</th>
|
||
<th id="76C55DFCAD26002EE59DFED8FBDAAB48" box="[1021,1141,261,282]" gridcol="2" gridrow="0" pageId="2" pageNumber="481">
|
||
<emphasis id="B950EAD0AD26FFD3E278FED8FBF6AB48" bold="true" box="[1048,1113,261,282]" pageId="2" pageNumber="481">Range</emphasis>
|
||
</th>
|
||
<th id="76C55DFCAD26002EE341FED8FAF3AB48" box="[1313,1372,261,282]" gridcol="3" gridrow="0" pageId="2" pageNumber="481">
|
||
<emphasis id="B950EAD0AD26FFD3E341FED8FAF3AB48" bold="true" box="[1313,1372,261,282]" pageId="2" pageNumber="481">Mean</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FEE6FAF3AB03" box="[150,1372,315,337]" gridrow="1" pageId="2" pageNumber="481" rowspan-3="1">
|
||
<th id="76C55DFCAD26002EE6F6FEE6FE43AB03" box="[150,492,315,337]" gridcol="0" gridrow="1" pageId="2" pageNumber="481">Standard length (mm)</th>
|
||
<td id="76C55DFCAD26002EE564FEE6FCCEAB03" box="[772,865,315,337]" gridcol="1" gridrow="1" pageId="2" pageNumber="481">51.9</td>
|
||
<td id="76C55DFCAD26002EE59DFEE6FBDAAB03" box="[1021,1141,315,337]" gridcol="2" gridrow="1" pageId="2" pageNumber="481">48.6–66.0</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FEB7FAF3ABD2" box="[150,1372,362,384]" gridrow="2" pageId="2" pageNumber="481" rowspan-3="1">
|
||
<th id="76C55DFCAD26002EE6F6FEB7FE43ABD2" box="[150,492,362,384]" gridcol="0" gridrow="2" pageId="2" pageNumber="481">Total length (mm)</th>
|
||
<td id="76C55DFCAD26002EE564FEB7FCCEABD2" box="[772,865,362,384]" gridcol="1" gridrow="2" pageId="2" pageNumber="481">64.4</td>
|
||
<td id="76C55DFCAD26002EE59DFEB7FBDAABD2" box="[1021,1141,362,384]" gridcol="2" gridrow="2" pageId="2" pageNumber="481">59.7–83.4</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FE44FAF3A8DF" box="[150,1372,409,653]" gridrow="3" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FE44FE43A8DF" box="[150,492,409,653]" gridcol="0" gridrow="3" pageId="2" pageNumber="481">In percent of standard length Total length Head length (dorsal) Head length (lateral) Predorsal length Prepelvic length Pre-anus length Pre-anal length</th>
|
||
<td id="76C55DFCAD26002EE564FE44FCCEA8DF" box="[772,865,409,653]" gridcol="1" gridrow="3" pageId="2" pageNumber="481">124.0 21.9 24.5 53.1 48.4 67.4 74.7</td>
|
||
<td id="76C55DFCAD26002EE59DFE44FBDAA8DF" box="[1021,1141,409,653]" gridcol="2" gridrow="3" pageId="2" pageNumber="481">120.5–126.5 18.5–21.9 22.4–25.4 51.8–54.8 48.0–51.5 67.4–73.2 73.1–78.1</td>
|
||
<td id="76C55DFCAD26002EE341FE44FAF3A8DF" box="[1313,1372,409,653]" gridcol="3" gridrow="3" pageId="2" pageNumber="481">123.0 20.3 23.9 53.3 50.0 69.2 75.4</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FD4AFAF3A8FF" box="[150,1372,663,685]" gridrow="4" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FD4AFE43A8FF" box="[150,492,663,685]" gridcol="0" gridrow="4" pageId="2" pageNumber="481">Head depth</th>
|
||
<td id="76C55DFCAD26002EE564FD4AFCCEA8FF" box="[772,865,663,685]" gridcol="1" gridrow="4" pageId="2" pageNumber="481">10.7</td>
|
||
<td id="76C55DFCAD26002EE59DFD4AFBDAA8FF" box="[1021,1141,663,685]" gridcol="2" gridrow="4" pageId="2" pageNumber="481">10.0–11.4</td>
|
||
<td id="76C55DFCAD26002EE341FD4AFAF3A8FF" box="[1313,1372,663,685]" gridcol="3" gridrow="4" pageId="2" pageNumber="481">10.7</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FD6AFAF3A97E" box="[150,1372,695,812]" gridrow="5" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FD6AFE43A97E" box="[150,492,695,812]" gridcol="0" gridrow="5" pageId="2" pageNumber="481">Body depth at dorsal-fin origin Depth of caudal peduncle Length of caudal peduncle Head width</th>
|
||
<td id="76C55DFCAD26002EE564FD6AFCCEA97E" box="[772,865,695,812]" gridcol="1" gridrow="5" pageId="2" pageNumber="481">13.8 12.0 17.5 14.3</td>
|
||
<td id="76C55DFCAD26002EE59DFD6AFBDAA97E" box="[1021,1141,695,812]" gridcol="2" gridrow="5" pageId="2" pageNumber="481">12.6–15.5 10.6–12.0 15.3–17.5 13.5–15.4</td>
|
||
<td id="76C55DFCAD26002EE341FD6AFAF3A97E" box="[1313,1372,695,812]" gridcol="3" gridrow="5" pageId="2" pageNumber="481">13.6 11.3 16.2 14.3</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FCEBFAF3A91E" box="[150,1372,822,844]" gridrow="6" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FCEBFE43A91E" box="[150,492,822,844]" gridcol="0" gridrow="6" pageId="2" pageNumber="481">Body width at dorsal-fin origin</th>
|
||
<td id="76C55DFCAD26002EE564FCEBFCCEA91E" box="[772,865,822,844]" gridcol="1" gridrow="6" pageId="2" pageNumber="481">9.8</td>
|
||
<td id="76C55DFCAD26002EE59DFCEBFBDAA91E" box="[1021,1141,822,844]" gridcol="2" gridrow="6" pageId="2" pageNumber="481">8.8–11.9</td>
|
||
<td id="76C55DFCAD26002EE341FCEBFAF3A91E" box="[1313,1372,822,844]" gridcol="3" gridrow="6" pageId="2" pageNumber="481">10.2</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FC8BFAF3AE78" box="[150,1372,854,1066]" gridrow="7" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FC8BFE43AE78" box="[150,492,854,1066]" gridcol="0" gridrow="7" pageId="2" pageNumber="481">Snout length Eye diameter Interorbital width Length of dorsal fin Length of upper caudal-fin lobe Length of median caudal-fin rays Length of lower caudal-fin lobe</th>
|
||
<td id="76C55DFCAD26002EE564FC8BFCCEAE78" box="[772,865,854,1066]" gridcol="1" gridrow="7" pageId="2" pageNumber="481">8.4 2.9 5.8 16.5 23.4 17.3 23.4</td>
|
||
<td id="76C55DFCAD26002EE59DFC8BFBDAAE78" box="[1021,1141,854,1066]" gridcol="2" gridrow="7" pageId="2" pageNumber="481">7.7–9.8 2.5–3.3 5.2–6.2 16.0–19.9 19.5–23.6 12.6–17.9 20.3–25.4</td>
|
||
<td id="76C55DFCAD26002EE341FC8BFAF3AE78" box="[1313,1372,854,1066]" gridcol="3" gridrow="7" pageId="2" pageNumber="481">9.1 3.0 5.9 17.5 22.1 14.9 23.0</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FBE9FAF3AEDB" box="[150,1372,1076,1161]" gridrow="8" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FBE9FE43AEDB" box="[150,492,1076,1161]" gridcol="0" gridrow="8" pageId="2" pageNumber="481">Length of anal fin Length of pelvic fin Length of pectoral fin</th>
|
||
<td id="76C55DFCAD26002EE564FBE9FCCEAEDB" box="[772,865,1076,1161]" gridcol="1" gridrow="8" pageId="2" pageNumber="481">16.4 15.9 19.1</td>
|
||
<td id="76C55DFCAD26002EE59DFBE9FBDAAEDB" box="[1021,1141,1076,1161]" gridcol="2" gridrow="8" pageId="2" pageNumber="481">16.2–18.7 15.7–18.0 16.4–19.1</td>
|
||
<td id="76C55DFCAD26002EE341FBE9FAF3AEDB" box="[1313,1372,1076,1161]" gridcol="3" gridrow="8" pageId="2" pageNumber="481">17.3 16.4 18.3</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FB7FFAF3AF4A" box="[150,1372,1186,1304]" gridrow="9" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FB7FFE43AF4A" box="[150,492,1186,1304]" gridcol="0" gridrow="9" pageId="2" pageNumber="481">In percent of dorsal head length Snout length Eye diameter Interorbital width</th>
|
||
<td id="76C55DFCAD26002EE564FB7FFCCEAF4A" box="[772,865,1186,1304]" gridcol="1" gridrow="9" pageId="2" pageNumber="481">38 13 27</td>
|
||
<td id="76C55DFCAD26002EE59DFB7FFBDAAF4A" box="[1021,1141,1186,1304]" gridcol="2" gridrow="9" pageId="2" pageNumber="481">38–50 13–16 27–33</td>
|
||
<td id="76C55DFCAD26002EE341FB7FFAF3AF4A" box="[1313,1372,1186,1304]" gridcol="3" gridrow="9" pageId="2" pageNumber="481">45 15 29</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FAEDFAF3AF34" box="[150,1372,1328,1382]" gridrow="10" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FAEDFE43AF34" box="[150,492,1328,1382]" gridcol="0" gridrow="10" pageId="2" pageNumber="481">In percent of lateral head length Snout length</th>
|
||
<td id="76C55DFCAD26002EE564FAEDFCCEAF34" box="[772,865,1328,1382]" gridcol="1" gridrow="10" pageId="2" pageNumber="481">34</td>
|
||
<td id="76C55DFCAD26002EE59DFAEDFBDAAF34" box="[1021,1141,1328,1382]" gridcol="2" gridrow="10" pageId="2" pageNumber="481">34–42</td>
|
||
<td id="76C55DFCAD26002EE341FAEDFAF3AF34" box="[1313,1372,1328,1382]" gridcol="3" gridrow="10" pageId="2" pageNumber="481">38</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FAADFAF3AFD4" box="[150,1372,1392,1414]" gridrow="11" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FAADFE43AFD4" box="[150,492,1392,1414]" gridcol="0" gridrow="11" pageId="2" pageNumber="481">Eye diameter</th>
|
||
<td id="76C55DFCAD26002EE564FAADFCCEAFD4" box="[772,865,1392,1414]" gridcol="1" gridrow="11" pageId="2" pageNumber="481">12</td>
|
||
<td id="76C55DFCAD26002EE59DFAADFBDAAFD4" box="[1021,1141,1392,1414]" gridcol="2" gridrow="11" pageId="2" pageNumber="481">11–14</td>
|
||
<td id="76C55DFCAD26002EE341FAADFAF3AFD4" box="[1313,1372,1392,1414]" gridcol="3" gridrow="11" pageId="2" pageNumber="481">13</td>
|
||
</tr>
|
||
<tr id="35143480AD26002EE6F6FA4DFAF3AFF4" box="[150,1372,1424,1446]" gridrow="12" pageId="2" pageNumber="481">
|
||
<th id="76C55DFCAD26002EE6F6FA4DFE43AFF4" box="[150,492,1424,1446]" gridcol="0" gridrow="12" pageId="2" pageNumber="481">Interorbital width</th>
|
||
<td id="76C55DFCAD26002EE564FA4DFCCEAFF4" box="[772,865,1424,1446]" gridcol="1" gridrow="12" pageId="2" pageNumber="481">24</td>
|
||
<td id="76C55DFCAD26002EE59DFA4DFBDAAFF4" box="[1021,1141,1424,1446]" gridcol="2" gridrow="12" pageId="2" pageNumber="481">22–26</td>
|
||
<td id="76C55DFCAD26002EE341FA4DFAF3AFF4" box="[1313,1372,1424,1446]" gridcol="3" gridrow="12" pageId="2" pageNumber="481">25</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD26FFD3E6E8F8C9FB35AD9E" blockId="2.[134,775,1811,2092]" lastBlockId="2.[835,1476,1524,1996]" pageId="2" pageNumber="481">
|
||
Dorsal fin with 4 unbranched and 7½ (1), 8½ (12) or 9½ (1*) branched rays; distal margin slightly concave. Usually, second branched ray longest. Pectoral fin curled and slanted, with 1 unbranched and 9 branched rays (including small last ray, usually unbranched), rounded, reaching ⅓ to ⅜ of distance to pelvic-fin base; rays without filamentous extensions. Axillary pectoral lobe present, cylindrical, posteriorly pointed, usually separated from skin of flank by a deep groove; groove often margined above by a second pointed lobe (
|
||
<figureCitation id="131F2A47AD26FFD3E43AF7C9FD33A27E" box="[602,668,2068,2092]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="2" pageNumber="481">Fig. 6</figureCitation>
|
||
), more or less regularly shaped, more or less distinct. Pelvic fin with 1 unbranched and 6 (2), 7 (11*) or 8 (1) branched rays (including small last ray, usually unbranched); reaching almost to anus (⅔ of distance to anal-fin origin; triangular to rounded; posterior margin straight; origin below base of unbranched dorsal-fin rays 1–3; axillary lobe present, entirely free, longer than fin base, high, rounded. Anus situated 1.5–2.5 eye diameter in front of anal-fin origin, distance between anus and anal-fin origin about ½ of distance between anus and pelvic-fin base. Anal fin with 3 unbranched and 5½ branched rays; distal margin slightly concave, first branched ray longest. Caudal fin with 9+8 (11*) or 8+7 (1) branched rays; forked (upper lobe 1.3–1.6 times longer than median rays), lobes rounded, subequal, lower lobe slightly longer than upper one (1.0–1.1 times).
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD26FFD5E523F829FE8EACF9" blockId="2.[835,1474,2036,2093]" lastBlockId="4.[136,775,1619,1707]" lastPageId="4" lastPageNumber="483" pageId="2" pageNumber="481">
|
||
Especially visible in life, inner pectoral-fin rays curled upwards and slightly forwards, against body, so that fin may appear vertically orientated (
|
||
<figureCitation id="131F2A47AD20FFD5E7DAF98EFE55AC39" box="[442,506,1619,1643]" captionStart="Fig" captionStartId="4.[135,170,1513,1535]" captionTargetBox="[135,1473,189,1497]" captionTargetId="figure-7@4.[135,1473,189,1497]" captionTargetPageId="4" captionText="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="4" pageNumber="483">Fig. 3</figureCitation>
|
||
). In pelvic fin, innermost rays also curled upwards. No difference between sexes in this character.
|
||
</paragraph>
|
||
<caption id="DF5B664AAD27FFD2E611F82EFD9BA210" pageId="3" pageNumber="482" startId="3.[113,148,2035,2057]" targetBox="[175,1390,189,2020]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B9B36C2AD27FFD2E611F82EFD9BA210" blockId="3.[113,1451,2035,2115]" pageId="3" pageNumber="482">
|
||
Fig. 2.
|
||
<taxonomicName id="4C244D41AD27FFD2E6DCF829FECEA25A" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[188,353,2035,2057]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="482" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD27FFD2E6DCF829FECEA25A" box="[188,353,2035,2057]" italics="true" pageId="3" pageNumber="482">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5B664AAD20FFD5E6E7FA34FE75AC49" pageId="4" pageNumber="483" startId="4.[135,170,1513,1535]" targetBox="[135,1473,189,1497]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B9B36C2AD20FFD5E6E7FA34FE75AC49" blockId="4.[135,1474,1513,1563]" pageId="4" pageNumber="483">
|
||
Fig. 3.
|
||
<taxonomicName id="4C244D41AD20FFD5E6B0FA34FEDBAFAC" authorityName="Kottelat & & Hasan & Veryl" authorityYear="2024" box="[208,372,1513,1534]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="483" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD20FFD5E6B0FA34FEDBAFAC" box="[208,372,1513,1534]" italics="true" pageId="4" pageNumber="483">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD20FFD5E6EAF90EFEC3ADF8" blockId="4.[136,777,1747,1963]" pageId="4" pageNumber="483">Body entirely covered by scales, except in a few specimens on belly, from throat to anus. Scales embedded, not externally visible on anterior part of body. Lateral line complete, with approximately 86–93 pores (difficult to count with accuracy). Cephalic lateral line system with 6 supraorbital, 4 + 10–11 infraorbital, 9–10 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
|
||
<paragraph id="8B9B36C2AD20FFD5E6E8F80EFAF3ADB9" blockId="4.[136,776,2003,2059]" lastBlockId="4.[834,1476,1619,2027]" pageId="4" pageNumber="483">
|
||
Anterior nare pierced in front side of a pointed flap-like tube. Posterior nare adjacent to anterior one. Mouth U-shaped, gape about two times wider than long (
|
||
<figureCitation id="131F2A47AD20FFD5E29CF98EFAEEAC39" box="[1276,1345,1619,1643]" captionStart="Fig" captionStartId="6.[836,871,518,540]" captionTargetBox="[836,1474,189,503]" captionTargetId="figure-56@6.[836,1474,185,505]" captionTargetPageId="6" captionText="Fig. 7. Kapuasia falaris, CMK 29184, 51.4 mm SL; mouth." pageId="4" pageNumber="483">Fig. 7</figureCitation>
|
||
). Lips thin. Upper lip usually with median notch, with a few shallow, thin wrinkles on posterior edge. Processus dentiformis present. Lower lip almost continuous, with a median notch but not forming two median lobes; median part with small irregular ridges, radiating from anterior extremity of postlabial groove, lateral parts smooth. Anterior part of lower lip and space between anterior extremities of postlabial grooves with irregular ridges, continuous with the patch on the lip itself. Tip of lower jaw not exposed when mouth is closed. No median concavity in lower jaw. Inner rostral barbel not reaching corner of mouth; outer one reaching corner of mouth. Maxillary barbel reaching vertical of eye.
|
||
</paragraph>
|
||
<caption id="DF5B664AAD21FFD4E54DFDBFFC19A8E3" pageId="5" pageNumber="484" startId="5.[813,848,610,632]" targetBox="[814,1452,189,595]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B9B36C2AD21FFD4E54DFDBFFC19A8E3" blockId="5.[813,1452,610,689]" pageId="5" pageNumber="484">
|
||
Fig. 5.
|
||
<taxonomicName id="4C244D41AD21FFD4E512FDBFFBFFA825" authority=", CMK" authorityName="CMK" box="[882,1104,610,631]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="484" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD21FFD4E512FDBFFBBBA825" box="[882,1044,610,631]" italics="true" pageId="5" pageNumber="484">Kapuasia falaris</emphasis>
|
||
,CMK
|
||
</taxonomicName>
|
||
29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD21FFD4E54DFD35FB7BAE52" blockId="5.[811,1452,744,1024]" pageId="5" pageNumber="484">Branched rays 4 and following: both branches are branched, with narrow membranes between branches. Some individual variation exists in the thickness and branching pattern. In females, pattern roughly similar, but membranes are present between all branches. Branched ray 1: posterior branch may be slightly branched near tip. Branched rays 2 and 3: anterior branch not branched, posterior branch branched near tip. Branched ray 4 and following: both branches are branched, with membranes between all branches.</paragraph>
|
||
<caption id="DF5B664AAD21FFD4E611FB50FE31AE8E" pageId="5" pageNumber="484" startId="5.[113,148,1165,1187]" targetBox="[113,751,189,1150]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B9B36C2AD21FFD4E611FB50FE31AE8E" blockId="5.[113,753,1165,1245]" pageId="5" pageNumber="484">
|
||
Fig. 4.
|
||
<taxonomicName id="4C244D41AD21FFD4E6D8FB53FE3AAEF0" authority=", ZRC" authorityName="ZRC" box="[184,405,1165,1187]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="484" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD21FFD4E6D8FB53FEF6AEF0" box="[184,345,1165,1187]" italics="true" pageId="5" pageNumber="484">Kapuasia falaris</emphasis>
|
||
, ZRC
|
||
</taxonomicName>
|
||
65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD21FFD4E611FAC9FE19AF3E" blockId="5.[113,752,1300,1388]" pageId="5" pageNumber="484">
|
||
Stomach elongated, intestine with a bend some distance behind stomach (
|
||
<figureCitation id="131F2A47AD21FFD4E759FAE9FE2DAF1E" box="[313,386,1332,1356]" captionStart="Fig" captionStartId="6.[836,872,1329,1351]" captionTargetBox="[1030,1280,581,1315]" captionTargetId="figure-58@6.[989,1325,577,1324]" captionTargetPageId="6" captionText="Fig. 8. Kapuasia falaris, ZRC 65915, 55.0 mm SL; schematic. Scale bar: 1 mm." pageId="5" pageNumber="484">Fig. 8</figureCitation>
|
||
). Air bladder without posterior chamber in abdominal cavity.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E613FA49FD26AC1E" blockId="5.[112,753,1428,1612]" pageId="5" pageNumber="484">
|
||
<emphasis id="B950EAD0AD21FFD4E613FA49FEF5AFFE" bold="true" box="[115,346,1428,1452]" pageId="5" pageNumber="484">Sexual dimorphism.</emphasis>
|
||
Males with suborbital flap (modified lateral ethmoid;
|
||
<figureCitation id="131F2A47AD21FFD4E77EFA69FEC6AF9E" box="[286,361,1460,1484]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="5" pageNumber="484">Fig. 4a</figureCitation>
|
||
), in lower position than in most other nemacheilids, globulous, movable, upper outline continued upwards, almost vertically, by a groove reaching orbit. Flap missing in females, but a shallow crease corresponds to position of lower edge of flap in males (
|
||
<figureCitation id="131F2A47AD21FFD4E44BF9E9FDD2AC1E" box="[555,637,1588,1612]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="5" pageNumber="484">Fig. 4b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E611F9A9FE13AD7E" blockId="5.[112,753,1652,1836]" pageId="5" pageNumber="484">
|
||
No tubercles on body and fins, in both sexes. In males, on the dorsal surface of pectoral fin, a conspicuous vertical skin flange, translucent, between branched rays 1–8, most developed between branched rays 1–6 (
|
||
<figureCitation id="131F2A47AD21FFD4E46EF909FDE2ACBE" box="[526,589,1748,1772]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="5" pageNumber="484">Fig. 5</figureCitation>
|
||
). These flanges not present in females. Females deeper bodied and apparently reach a larger size than males.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E611F889FA02AF72" blockId="5.[112,753,1876,2092]" lastBlockId="5.[811,1453,1064,1313]" pageId="5" pageNumber="484">
|
||
Unbranched and 1 or 2 anterior branched pectoral-fin rays thicker than others (
|
||
<figureCitation id="131F2A47AD21FFD4E733F8A9FE0DADDE" box="[339,418,1908,1932]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="5" pageNumber="484">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="131F2A47AD21FFD4E7D2F8A9FE6DADDE" box="[434,450,1908,1932]" captionStart="Fig" captionStartId="7.[114,149,1140,1162]" captionTargetBox="[114,752,189,1112]" captionTargetId="figure-14@7.[114,752,187,1127]" captionTargetPageId="7" captionText="Fig. 9. Kapuasia falaris, pectoral fin in dorsal view; ZRC 65844. a, female, 58.9 mm SL; b, male 52.3 mm SL (reversed)." pageId="5" pageNumber="484">9</figureCitation>
|
||
). In males, branched ray 1 branched only once, anterior branch usually thicker than posterior one, without membrane between branches. Branched ray 2: anterior branch thicker than posterior one, unbranched; posterior branch maybe branched at tip; no membranes. Branched rays 3: as ray 2 but branches may be branched.
|
||
<emphasis id="B950EAD0AD21FFD4E54DFBF5FC13AE12" bold="true" box="[813,956,1064,1088]" pageId="5" pageNumber="484">Colouration</emphasis>
|
||
(
|
||
<figureCitation id="131F2A47AD21FFD4E5AEFBF5FB94AE13" box="[974,1083,1064,1089]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="5" pageNumber="484">Figs. 1–3</figureCitation>
|
||
). After fixation in formalin and 4 months in ethanol. Head and body background colour yellowish white. Unless otherwise stated, markings black. Head black; snout white on top, between nostrils to extremity, black on sides; white mark or bar on cheek, from immediately behind eye downwards. Posterior part of opercle white, extending across back, over occiput and continuous with contralateral. Outer rostral barbel black, other barbels whitish.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E54BFA95FAA3AFB2" blockId="5.[811,1452,1352,1504]" pageId="5" pageNumber="484">Body with 3–7 bars, extending from dorsal midline to ventral midline (faint on belly in front of anal fin), continuous with their contralaterals on back. Bars mostly of regular width and shape, wider than interspaces, sometimes paired or with paler central area, appearing vertically split.</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E54FF9D4FB0FADF2" blockId="5.[812,1453,1544,1952]" pageId="5" pageNumber="484">
|
||
Pattern at caudal-fin base: a conspicuous black blotch on each lobe of caudal fin, on proximal half, along upper and lower edge, irregular shape, extending for about half of length of lobe, on unbranched rays and adjacent 4 branched rays. Blotch on upper lobe, a spot of more intense black at anterior extremity. In most specimens, anterior extremity of upper blotch more forward than lower one. A faint inner mark in front of lower blotch. Between blotches, rays black, membranes hyaline, sometimes almost forming a bar. In most specimens, a smaller blotch behind both main ones, along dorsal and ventral edges. In most specimens, a narrow margin around part of whole caudal fin. In
|
||
<specimenCount id="9D22FD4BAD21FFD4E36FF8B4FA08ADD2" box="[1295,1447,1896,1920]" count="1" pageId="5" pageNumber="484" type="generic">one specimen</specimenCount>
|
||
, caudal fin almost entirely hyaline.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD21FFD4E54DF815FA03A212" blockId="5.[813,1452,1992,2113]" pageId="5" pageNumber="484">Dorsal fin hyaline, with a black blotch at base of unbranched rays and first branched rays (1–2). A stripe made of melanophores at middle of last unbranched ray, and in middle area of branched rays, on rays near branching points.</paragraph>
|
||
<caption id="DF5B664AAD22FFD7E524FDDBFA3DA84E" box="[836,1426,518,540]" pageId="6" pageNumber="485" startId="6.[836,871,518,540]" targetBox="[836,1474,189,503]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B9B36C2AD22FFD7E524FDDBFA3DA84E" blockId="6.[836,1426,518,540]" box="[836,1426,518,540]" pageId="6" pageNumber="485">
|
||
Fig. 7.
|
||
<taxonomicName id="4C244D41AD22FFD7E5EDFDDAFBDCA849" authority=", CMK" authorityName="CMK" box="[909,1139,518,540]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD22FFD7E5EDFDDAFB81A849" box="[909,1070,518,540]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
|
||
, CMK
|
||
</taxonomicName>
|
||
29184, 51.4 mm SL; mouth.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5B664AAD22FFD7E524FAECFC42AF31" pageId="6" pageNumber="485" startId="6.[836,872,1329,1351]" targetBox="[1030,1280,581,1315]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B9B36C2AD22FFD7E524FAECFC42AF31" blockId="6.[836,1474,1329,1380]" pageId="6" pageNumber="485">
|
||
Fig. 8.
|
||
<taxonomicName id="4C244D41AD22FFD7E5EFFAECFBDAAF14" authority=", ZRC" authorityName="ZRC" box="[911,1141,1329,1350]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD22FFD7E5EFFAECFB99AF14" box="[911,1078,1329,1350]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
|
||
, ZRC
|
||
</taxonomicName>
|
||
65915, 55.0 mm SL; schematic. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD22FFD7E523FA46FB42AF81" blockId="6.[835,1474,1435,1491]" pageId="6" pageNumber="485">No melanophores on membranes, except between branches immediately distal to branching points.</paragraph>
|
||
<paragraph id="8B9B36C2AD22FFD7E523FA26FAC9AC01" blockId="6.[835,1474,1531,1619]" pageId="6" pageNumber="485">Anal fin hyaline, with a row of patches of melanophores on rays near branching points. Pelvic fin hyaline or with a few faint melanophores on rays near branching points.</paragraph>
|
||
<paragraph id="8B9B36C2AD22FFD7E523F9A6FC76ACA1" blockId="6.[835,1475,1659,1779]" pageId="6" pageNumber="485">Pectoral fin hyaline. Unbranched and branched rays 1–2 with a patch of melanophores on dorsal surface of distal ½ to ⅓. On other rays, melanophores along edge of rays and branches.</paragraph>
|
||
<paragraph id="8B9B36C2AD22FFD7E523F8C6FA21AD81" blockId="6.[835,1475,1819,2003]" pageId="6" pageNumber="485">In life: Pattern as in preserved specimens. Body cream-white, dorsum slight golden, venter white. Iris of eye flecked with gold specks. Bars on body brown, darker on dorsum. Caudal fin hyaline to light yellow, can be fully red. Upper part of last pale band (between last black bar and blotch at upper caudal base, and top of snout, bright iridescent white.</paragraph>
|
||
<caption id="DF5B664AAD22FFD7E6E7F81DFD59A27E" pageId="6" pageNumber="485" startId="6.[135,171,1984,2006]" targetBox="[135,773,189,1969]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B9B36C2AD22FFD7E6E7F81DFD59A27E" blockId="6.[135,774,1984,2093]" pageId="6" pageNumber="485">
|
||
Fig. 6.
|
||
<taxonomicName id="4C244D41AD22FFD7E6B4F81CFED3AD87" box="[212,380,1984,2006]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD22FFD7E6B4F81CFED3AD87" box="[212,380,1984,2006]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD22FFD7E523F826FA6DA260" blockId="6.[835,1474,2043,2099]" pageId="6" pageNumber="485">
|
||
<emphasis id="B950EAD0AD22FFD7E523F826FBBFA241" bold="true" box="[835,1040,2043,2067]" pageId="6" pageNumber="485">Notes on biology.</emphasis>
|
||
The stomach content of two dissected females (ZRC 65915,
|
||
<quantity id="4CDC9B27AD22FFD7E258F7C6FB36A260" box="[1080,1177,2075,2099]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="6" pageNumber="485" unit="mm" value="59.2">59.2 mm</quantity>
|
||
SL, ZRC 65844,
|
||
<quantity id="4CDC9B27AD22FFD7E301F7C6FA6DA260" box="[1377,1474,2075,2099]" metricMagnitude="-2" metricUnit="m" metricValue="5.89" pageId="6" pageNumber="485" unit="mm" value="58.9">58.9 mm</quantity>
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DFF60FB99ABC4" blockId="7.[811,1452,189,406]" pageId="7" pageNumber="486">is continuous, with a narrow shallow median longitudinal groove extending from the mental area to the anterior edge of the lip. The median part of the lower lip has 8–10 ridges on each side of the mental groove, radiating from the anterior extremity of the postlabial groove, across the whole lip, and also present on median space between the anterior extremities of the postlabial groove.</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DFE63FA1DA847" blockId="7.[813,1458,445,533]" pageId="7" pageNumber="486">
|
||
In
|
||
<taxonomicName id="4C244D41AD23FFD6E52BFE63FC1DAB87" box="[843,946,445,469]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E52BFE63FC1DAB87" box="[843,946,445,469]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
|
||
</taxonomicName>
|
||
, the lips are thinner, the groove in the median part of the lower lip is wider (but not forming conspicuous lobes) and the ridges are less obvious and less well organised.
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DFDE3FAFBA947" blockId="7.[811,1452,573,789]" pageId="7" pageNumber="486">
|
||
<taxonomicName id="4C244D41AD23FFD6E54DFDE3FC3DA807" box="[813,914,574,597]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EAD0AD23FFD6E54DFDE3FC3DA807" box="[813,914,574,597]" italics="true" pageId="7" pageNumber="486">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
was also distinguished from all known nemacheilid loaches by the presence of a large, conspicuous, rounded suprapectoral lobe above the pectoral fin. In
|
||
<taxonomicName id="4C244D41AD23FFD6E375FDA3FAD3A8C7" box="[1301,1404,637,661]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E375FDA3FAD3A8C7" box="[1301,1404,637,661]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
|
||
</taxonomicName>
|
||
, the lobe is cylindrical, posteriorly pointed, usually separated from skin of flank by a deep groove; groove often margined above by a second pointed lobe (
|
||
<figureCitation id="131F2A47AD23FFD6E2EAFD00FB64A8A7" box="[1162,1227,733,757]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="7" pageNumber="486">Fig. 6</figureCitation>
|
||
), often less regularly shaped and more or less distinct than the first one.
|
||
</paragraph>
|
||
<caption id="DF5B664AAD23FFD6E612FBA9FD30AEF5" pageId="7" pageNumber="486" startId="7.[114,149,1140,1162]" targetBox="[114,752,189,1112]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B9B36C2AD23FFD6E612FBA9FD30AEF5" blockId="7.[114,753,1140,1191]" pageId="7" pageNumber="486">
|
||
Fig. 9.
|
||
<taxonomicName id="4C244D41AD23FFD6E6DBFBA8FEF3AEDB" box="[187,348,1140,1162]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E6DBFBA8FEF3AEDB" box="[187,348,1140,1162]" italics="true" pageId="7" pageNumber="486">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
, pectoral fin in dorsal view; ZRC 65844. a, female, 58.9 mm SL; b, male 52.3 mm SL (reversed).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36C2AD23FFD6E611FB03FD5CAC64" blockId="7.[113,755,1246,1590]" pageId="7" pageNumber="486">
|
||
SL) is uninformative because specimens kept in captivity for several days. The ovaries of the
|
||
<quantity id="4CDC9B27AD23FFD6E79BFB23FDF4AF44" box="[507,603,1278,1302]" metricMagnitude="-2" metricUnit="m" metricValue="5.89" pageId="7" pageNumber="486" unit="mm" value="58.9">58.9 mm</quantity>
|
||
SL specimen contained unripe eggs, the largest ones about
|
||
<quantity id="4CDC9B27AD23FFD6E4F7FAC3FD40AF64" box="[663,751,1310,1334]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="486" unit="mm" value="0.5">0.5 mm</quantity>
|
||
diameter. We have no information on habitat. The curled and slanted orientation of the pectoral and pelvic fins suggest an adaptation to strong current. In captivity: in a
|
||
<quantity id="4CDC9B27AD23FFD6E435FAA3FD39AFC4" box="[597,662,1406,1430]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="486" unit="cm" value="60.0">60 cm</quantity>
|
||
(length) tank setup with rocks and stones, individuals remain not visible as they hide amongst the crevices or burrowed into coarse gravel. When food is introduced, the fish will swim out or even forcefully push against the coarse gravel in its foraging activity. They have a preference for a carnivore diet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD23FFD6E611F983FDA1ACE4" pageId="7" pageNumber="486" type="distribution">
|
||
<paragraph id="8B9B36C2AD23FFD6E611F983FDA1ACE4" blockId="7.[112,753,1630,1718]" pageId="7" pageNumber="486">
|
||
<emphasis id="B950EAD0AD23FFD6E611F983FEACAC24" bold="true" box="[113,259,1630,1654]" pageId="7" pageNumber="486">Distribution.</emphasis>
|
||
<taxonomicName id="4C244D41AD23FFD6E768F982FE17AC24" box="[264,440,1630,1654]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E768F982FE17AC24" box="[264,440,1630,1654]" italics="true" pageId="7" pageNumber="486">Kapuasia falaris</emphasis>
|
||
</taxonomicName>
|
||
is presently known only from one locality in the Barito drainage, Borneo. It is logically expected to have a wider distribution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD23FFD6E611F903FD9DAD24" pageId="7" pageNumber="486" type="etymology">
|
||
<paragraph id="8B9B36C2AD23FFD6E611F903FD9DAD24" blockId="7.[113,752,1758,1910]" pageId="7" pageNumber="486">
|
||
<emphasis id="B950EAD0AD23FFD6E611F903FF5CACA4" bold="true" box="[113,243,1758,1782]" pageId="7" pageNumber="486">Etymology.</emphasis>
|
||
From the Greek ‘phalaros’ (having a patch of white, φάλαρος) and ‘rhis’ (nose, snout, ῥίς); with a white spot on the nose. Used as a noun in apposition. Falaris or phalaris (φαλαρίς), is the name of the coot,
|
||
<taxonomicName id="4C244D41AD23FFD6E439F8E3FD7BAD04" authorityName="Linnaeus" authorityYear="1758" box="[601,724,1854,1878]" class="Aves" family="Rallidae" genus="Fulica" kingdom="Animalia" order="Gruiformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="atra">
|
||
<emphasis id="B950EAD0AD23FFD6E439F8E3FD7BAD04" box="[601,724,1854,1878]" italics="true" pageId="7" pageNumber="486">Fulica atra</emphasis>
|
||
</taxonomicName>
|
||
, a black bird with white beak and forehead.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD23FFD6E611F843FB58AD87" pageId="7" pageNumber="486" type="discussion">
|
||
<paragraph id="8B9B36C2AD23FFD6E611F843FB4BAEC7" blockId="7.[113,754,1950,2103]" lastBlockId="7.[812,1454,829,1173]" pageId="7" pageNumber="486">
|
||
<emphasis id="B950EAD0AD23FFD6E611F843FF4EADE4" bold="true" box="[113,225,1950,1974]" pageId="7" pageNumber="486">Remarks.</emphasis>
|
||
<taxonomicName id="4C244D41AD23FFD6E68BF842FEFDADE4" box="[235,338,1951,1974]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EAD0AD23FFD6E68BF842FEFDADE4" box="[235,338,1951,1974]" italics="true" pageId="7" pageNumber="486">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C244D41AD23FFD6E7F6F842FD9DADE4" authorityName="Roberts" authorityYear="1989" box="[406,562,1950,1974]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E7F6F842FD9DADE4" box="[406,562,1950,1974]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
) is distinguished from all other genera of
|
||
<taxonomicName id="4C244D41AD23FFD6E71FF863FDB0AD84" box="[383,543,1982,2006]" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
in the morphology of the mouth. It is strongly arched, U-shaped, the upper lip is swollen and smooth. The lower lip is swollen, with a deep postlabial groove along the lateral parts; its anterior edge
|
||
<specimenCount id="9D22FD4BAD23FFD6E54DFCE0FC68A907" box="[813,967,829,853]" count="1" pageId="7" pageNumber="486" type="generic">One specimen</specimenCount>
|
||
of
|
||
<taxonomicName id="4C244D41AD23FFD6E58FFCE3FB26A907" authorityName="Roberts" authorityYear="1989" box="[1007,1161,829,853]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E58FFCE3FB26A907" box="[1007,1161,829,853]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
had a few small tubercles on the anterior branched pectoral-fin rays, assumed to be a male character. Both sexes are present in the material of
|
||
<taxonomicName id="4C244D41AD23FFD6E54DFC43FC3CA9E7" box="[813,915,925,949]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E54DFC43FC3CA9E7" box="[813,915,925,949]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
|
||
</taxonomicName>
|
||
; no tubercles were present in males. Males of
|
||
<taxonomicName id="4C244D41AD23FFD6E3F5FC43FCDCA987" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E3F5FC43FCDCA987" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
|
||
</taxonomicName>
|
||
are also distinguished by the presence of vertical skin flanges between the branched rays of the pectoral fin (
|
||
<figureCitation id="131F2A47AD23FFD6E31FFC00FC93AE47" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="7" pageNumber="486">Fig. 5</figureCitation>
|
||
); such flanges had not been reported in
|
||
<taxonomicName id="4C244D41AD23FFD6E368FC23FA0AAE47" authorityName="Roberts" authorityYear="1989" box="[1288,1445,1021,1045]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E368FC23FA0AAE47" box="[1288,1445,1021,1045]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
, but a shallow ridge of skin between the rays is visible on the photographs of ZRC 56404, of unknown sex, and ZRC 61464, apparently a male, and appears possibly homologous (
|
||
<bibRefCitation id="EFB54B33AD23FFD6E553FBA0FB8CAEC7" author="Kottelat M & Tan HH" box="[819,1059,1149,1173]" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M & Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat & Tan, 2024</bibRefCitation>
|
||
: figs. 1b, c, 3, 4).
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DFB60FBCEAC47" blockId="7.[813,1452,1213,1557]" pageId="7" pageNumber="486">
|
||
The males
|
||
<taxonomicName id="4C244D41AD23FFD6E5CCFB63FBB7AE87" box="[940,1048,1213,1237]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
|
||
<emphasis id="B950EAD0AD23FFD6E5CCFB63FBB7AE87" box="[940,1048,1213,1237]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
|
||
</taxonomicName>
|
||
have a conspicuous suborbital flap (modified lateral ethmoid;
|
||
<figureCitation id="131F2A47AD23FFD6E232FB00FB3AAEA7" box="[1106,1173,1245,1269]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="7" pageNumber="486">Fig. 4</figureCitation>
|
||
), which is missing in the single specimen of
|
||
<taxonomicName id="4C244D41AD23FFD6E266FB23FB0EAF47" authorityName="Roberts" authorityYear="1989" box="[1030,1185,1277,1301]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E266FB23FB0EAF47" box="[1030,1185,1277,1301]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
tentatively identified as male. The flap is globulous, movable, at some distance below the eye, with a deep groove connecting it to the edge of the orbit. In most nemacheilds of Southeast Asia, when a flap is present, it is immediately below the eye and the groove is not present. A somewhat similar condition is reported only in ‘
|
||
<taxonomicName id="4C244D41AD23FFD6E58EFA63FABCAF87" box="[1006,1299,1469,1493]" class="Actinopterygii" family="Nemacheilidae" genus="Mustura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="shuangjiangensis">
|
||
<emphasis id="B950EAD0AD23FFD6E58EFA63FBE7AF87" box="[1006,1096,1470,1493]" italics="true" pageId="7" pageNumber="486">Mustura</emphasis>
|
||
’
|
||
<emphasis id="B950EAD0AD23FFD6E23AFA60FABCAF87" box="[1114,1299,1469,1493]" italics="true" pageId="7" pageNumber="486">shuangjiangensis</emphasis>
|
||
</taxonomicName>
|
||
, in which the flap is at a distance from the eye, and the groove slanted (see
|
||
<bibRefCitation id="EFB54B33AD23FFD6E500FA20FC50AC47" author="Kottelat M" box="[864,1023,1533,1557]" pageId="7" pageNumber="486" pagination="289 - 314" refId="ref4912" refString="Kottelat M (2018) Mustura celata, a new genus and species of loaches from northern Myanmar, and an overview of Physoschistura and related taxa (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 28 (4): 289 - 314." type="journal article" year="2018">Kottelat, 2018</bibRefCitation>
|
||
: fig. 26).
|
||
</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DF9E0FADEACA7" blockId="7.[811,1452,1597,1781]" pageId="7" pageNumber="486">Another character shared by the two species and distinguishing them from all sundaic nemacheilids is the colour pattern of the caudal fin, made of a pair of black blotches in the proximal area of the fin, of which the anterior extremity of the lower one is slightly pushed backwards, the black rays between them, and a bar or row of spots in subdistal position.</paragraph>
|
||
<paragraph id="8B9B36C2AD23FFD6E54DF8C0FB58AD87" blockId="7.[811,1453,1821,2005]" pageId="7" pageNumber="486">
|
||
The strongly curled pectoral and pevic fins, with innermost rays positioned very close to the body and appearing in life as if the fins were vertical is a feature not observed in any other nemacheilid in Southeast Asia. However, the preserved specimen of
|
||
<taxonomicName id="4C244D41AD23FFD6E5D2F843FBE7ADE7" authorityName="Roberts" authorityYear="1989" box="[946,1096,1949,1973]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E5D2F843FBE7ADE7" box="[946,1096,1949,1973]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
|
||
</taxonomicName>
|
||
figured by
|
||
<bibRefCitation id="EFB54B33AD23FFD6E2A0F840FA07ADE7" author="Kottelat M & Tan HH" box="[1216,1448,1949,1973]" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M & Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat & Tan (2024</bibRefCitation>
|
||
: fig. 2) shows a possibly similar condition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6549AD23FFD6E54DF820FC02A267" pageId="7" pageNumber="486" type="materials_examined">
|
||
<paragraph id="8B9B36C2AD23FFD6E54DF820FC02A267" blockId="7.[813,1452,2045,2101]" pageId="7" pageNumber="486">
|
||
<emphasis id="B950EAD0AD23FFD6E54DF820FBA2A247" bold="true" box="[813,1037,2045,2069]" pageId="7" pageNumber="486">Material examined.</emphasis>
|
||
<taxonomicName id="4C244D41AD23FFD6E276F823FB52A247" box="[1046,1277,2045,2069]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B950EAD0AD23FFD6E276F823FB52A247" box="[1046,1277,2045,2069]" italics="true" pageId="7" pageNumber="486">Kapuasia maculiceps</emphasis>
|
||
</taxonomicName>
|
||
: see
|
||
<bibRefCitation id="EFB54B33AD23FFD6E358F820FC06A267" author="Kottelat M & Tan HH" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M & Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat & Tan (2024)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |