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<document id="F79205B80E271B4D0F06B18E9A436259" ID-CLB-Dataset="306597" ID-DOI="10.1093/zoolinnean/zlad177" ID-GBIF-Dataset="dcaf5aa0-c7c6-4dc1-871a-8815b1aabb88" ID-ISSN="0024-4082" ID-Zenodo-Dep="14510261" ID-ZooBank="80D1924-E984-4900-88E5-85FE2C5688D8" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1734027918295" checkinUser="plazi" docAuthor="Hinckley, Arlo, Camacho-Sanchez, Miguel, Chua, Marcus A. H., Ruedi, Manuel, Lunde, Darrin, Maldonado, Jesús E., Omar, Hasmahzaiti, Leonard, Jennifer A. &amp; Hawkins, Melissa T. R." docDate="2024" docId="038C8797FF96FFC4FF16631EE76CFD80" docLanguage="en" docName="zlad177.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlad177" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Hylomys vorax Hinckley, Lunde &amp; Hawkins 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFB5FFEFFF85FFD1FF89661BE21AFFDF" masterDocTitle="An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="20" updateTime="1734460109358" updateUser="ExternalLinkService">
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<mods:title id="2E96D024B6F537FACA61F2B3F23ED45D">An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia</mods:title>
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<mods:namePart id="4961BD9F675EA2FA36FB0C7C21E09666">Hinckley, Arlo</mods:namePart>
<mods:affiliation id="24883ABC8CEC246C1D9F9D5CE74B119B">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="DAD7E4D18A82A28A90A94B3BD5A74753">Camacho-Sanchez, Miguel</mods:namePart>
<mods:affiliation id="27E166DD592C5ADDCC6C927EAC86E379">Conservation &amp; Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<mods:namePart id="7C6D65781F266F95928789AC4349A29A">Chua, Marcus A. H.</mods:namePart>
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<mods:namePart id="9674F11EB8C3836D87FFB7B3C910E4EA">Ruedi, Manuel</mods:namePart>
<mods:affiliation id="431A8C3CB8010972D7A3C857B7F633B8">Natural History Museum of Geneva, Geneva, Switzerland</mods:affiliation>
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<mods:namePart id="BA8E3800DFA79C1A1FF0066862C67220">Lunde, Darrin</mods:namePart>
<mods:affiliation id="F35E9E42F357B29B432D9B3B75289426">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="4E5F7E2222DFAB000D8B4211C066683E">Maldonado, Jesús E.</mods:namePart>
<mods:affiliation id="AEF69076AC9AC19CB7B555A6F894C8AD">Smithsonian National Zoo and Conservation Biology Institute, Center for Conservation Genomics, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="BFB4F35E667AE2FA4A6E1086A594E373">Omar, Hasmahzaiti</mods:namePart>
<mods:affiliation id="3350CF648F23C24689AA8FE7FD72D607">Museum of Zoology, Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:namePart id="40CC69308B28300272B49CC6EE8AEE0E">Leonard, Jennifer A.</mods:namePart>
<mods:affiliation id="0E62A6E4821DBDAE740BD111FCD80920">Conservation &amp; Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<mods:namePart id="8497E358753A722A2578BAE183F20157">Hawkins, Melissa T. R.</mods:namePart>
<mods:affiliation id="0D0FBF98B2D35345DAD9F4A76EB9F057">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:date id="2D9FA798DF8B2DD8F9E325E17537692F">2024</mods:date>
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<treatment id="038C8797FF96FFC4FF16631EE76CFD80" ID-DOI="http://doi.org/10.5281/zenodo.14510289" ID-Zenodo-Dep="14510289" LSID="urn:lsid:plazi:treatment:038C8797FF96FFC4FF16631EE76CFD80" httpUri="http://treatment.plazi.org/id/038C8797FF96FFC4FF16631EE76CFD80" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C33F650AFF96FFC2FF16631EE0D6FAFF" box="[159,716,1285,1312]" pageId="19" pageNumber="20" type="nomenclature">
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<emphasis id="B951EA93FF96FFC2FF16631EE0D6FAFF" bold="true" box="[159,716,1285,1312]" pageId="19" pageNumber="20">
<taxonomicName id="4C254D02FF96FFC2FF16631EE06CFAC0" authority="Hinckley, Lunde &amp; Hawkins" authorityName="Hinckley, Lunde &amp; Hawkins" authorityYear="2024" box="[159,630,1285,1312]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="vorax" status="sp. nov.">
<emphasis id="B951EA93FF96FFC2FF16631EE35DFAFF" bold="true" box="[159,327,1285,1312]" italics="true" pageId="19" pageNumber="20">Hylomys vorax</emphasis>
Hinckley, Lunde &amp; Hawkins
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,
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</emphasis>
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<paragraph id="8B9A3681FF96FFC2FFF86337E02AFA9B" blockId="19.[113,762,1285,1442]" box="[113,560,1324,1348]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB3511AFFF96FFC2FFF86337E02AFA9B" box="[113,560,1324,1348]" pageId="19" pageNumber="20">
<treatmentCitation id="0A841090FF96FFC2FFF86337E036FA9B" author="Millbo GS" box="[113,556,1324,1348]" page="111" pageId="19" pageNumber="20" year="1942">
<taxonomicName id="4C254D02FF96FFC2FFF86337E036FA9B" ID-CoL="5J2D5" authority="Millbo, 1942: 109 - 111" authorityName="Millbo" authorityPageNumber="109 - 111" authorityYear="1942" box="[113,556,1324,1348]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="maxi">
<emphasis id="B951EA93FF96FFC2FFF86337E325FA9B" box="[113,319,1324,1348]" italics="true" pageId="19" pageNumber="20">Hylomys suillus maxi</emphasis>
<bibRefCitation id="EFB44B70FF96FFC2FECD6337E036FA9B" author="Millbo GS" box="[324,556,1324,1348]" pageId="19" pageNumber="20" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller, 1942: 109–111</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8B9A3681FF96FFC2FFF86350E3F0FABC" blockId="19.[113,762,1285,1442]" box="[113,490,1355,1379]" pageId="19" pageNumber="20">
<taxonomicName id="4C254D02FF96FFC2FFF86350E3FFFABC" ID-CoL="5J2D5" authority="Bbst, 2018: 328" authorityName="Bbst" authorityPageNumber="328" authorityYear="2018" box="[113,485,1355,1379]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="maxi">
<emphasis id="B951EA93FF96FFC2FFF86350E325FABC" box="[113,319,1355,1379]" italics="true" pageId="19" pageNumber="20">Hylomys suillus maxi</emphasis>
Best, 2018: 328
</taxonomicName>
.
</paragraph>
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<subSubSection id="C33F650AFF96FFC2FFF86370E3F3FA7D" pageId="19" pageNumber="20" type="description">
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<emphasis id="B951EA93FF96FFC2FFF86370E343FA5D" box="[113,345,1387,1411]" italics="true" pageId="19" pageNumber="20">ZooBank registration:</emphasis>
urn:lsid:zoobank.org:act:
<uuid id="FF830C54FF96FFC2FDF86370E3FCFA7D" pageId="19" pageNumber="20">BA2B0BCE-AF92-4583-8C74-1A3EAA60AE7B</uuid>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF96FFC2FFF863D8E6B7FADF" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B9A3681FF96FFC2FFF863D8E7B9FF18" blockId="19.[113,763,1475,1969]" lastBlockId="19.[810,1458,144,199]" pageId="19" pageNumber="20">
<materialsCitation id="3B4D3CDCFF96FFC2FFF863D8E393FA24" accessionNumber="OR138102, OR554454" collectingDate="1939-05-01" collectionCode="USNM" collectorName="Frederick A. Ulmer, Jr" country="Indonesia" elevation="2408" latitude="3.85" location="Mt. Leuser" longLatPrecision="784" longitude="97.12" municipality="Aceh" pageId="19" pageNumber="20" specimenCode="USNM 271034" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
<emphasis id="B951EA93FF96FFC2FFF863D8E2CBFA04" box="[113,209,1475,1499]" italics="true" pageId="19" pageNumber="20">
<typeStatus id="549E8823FF96FFC2FFF863D8E2D6FA04" box="[113,204,1475,1499]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DB839EFAFF96FFC2FF5163D8E360FA04" box="[216,378,1475,1499]" pageId="19" pageNumber="20">USNM 271034</specimenCode>
, an
<specimenCount id="9D23FD08FF96FFC2FE2D63D8E00AFA04" box="[420,528,1475,1499]" pageId="19" pageNumber="20" type="adult">adult male</specimenCount>
collected by
<collectorName id="26D05357FF96FFC2FD1063D8E2F0FA25" pageId="19" pageNumber="20">Frederick A. Ulmer, Jr</collectorName>
on
<date id="FF9B1041FF96FFC2FE8663F8E393FA24" box="[271,393,1507,1531]" pageId="19" pageNumber="20" value="1939-05-01">
<collectingDate id="EFDFE9A9FF96FFC2FE8663F8E393FA24" box="[271,393,1507,1531]" pageId="19" pageNumber="20" value="1939-05-01">1 May 1939</collectingDate>
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. The specimen consists of a museum study skin with accompanying cranium and mandible; skin well prepared, with a small hairless patch on the two thighs and the most external toe of the left hindfoot missing (clipped for DNA extraction), otherwise in good condition; skull well prepared (with some remaining dried tissue attached, allowing for subsequent historical DNA extraction), but has damaged pterygoid processes, left P2 crown, and ectotympanic bone of the right bulla. Total length
<quantity id="4CDD9B64FF96FFC2FEBD60C6E397F92A" box="[308,397,1757,1781]" metricMagnitude="-1" metricUnit="m" metricValue="1.48" pageId="19" pageNumber="20" unit="mm" value="148.0">148 mm</quantity>
, tail
<quantity id="4CDD9B64FF96FFC2FE3760C6E012F92A" box="[446,520,1757,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="mm" value="22.0">22 mm</quantity>
(17.5 % HB), hindfoot
<quantity id="4CDD9B64FF96FFC2FFF860E7E2D5F8CB" box="[113,207,1788,1812]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="19" pageNumber="20" unit="mm" value="27.5">27.5 mm</quantity>
, ear length
<quantity id="4CDD9B64FF96FFC2FECD60E6E395F8CB" box="[324,399,1789,1813]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="mm" value="19.0">19 mm</quantity>
, weight
<quantity id="4CDD9B64FF96FFC2FE6A60E7E039F8CA" box="[483,547,1788,1813]" metricMagnitude="-2" metricUnit="kg" metricValue="4.25" pageId="19" pageNumber="20" unit="g" value="42.5">42.5 g</quantity>
. Skull measurements in Supporting Information, Table S2. 3D surface scans of this holotype skull are available at: https://doi.org/10.17602/ M2/M560018 (cranium); https://doi.org/10.17602/M2/ M560022 (mandible). Pictures of this holotype skull and skin are shown in
<figureCitation id="131E2A04FF96FFC2FF706182E344F86E" box="[249,350,1945,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="19" pageNumber="20">Figure 9A</figureCitation>
and the Supporting Information,
<figureCitation id="131E2A04FF96FFC2FD316182E15FFF77" box="[696,837,144,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="19" pageNumber="20">Figure S9</figureCitation>
, respectively. DNA sequences associated with this holotype have been deposited in GenBank:
<accessionNumber id="9476AB62FF96FFC2FB0C66B4E6E2FF18" box="[1157,1272,175,199]" pageId="19" pageNumber="20">OR138102</accessionNumber>
and
<accessionNumber id="9476AB62FF96FFC2FAA066B4E785FF18" box="[1321,1439,175,199]" pageId="19" pageNumber="20">OR554454</accessionNumber>
.
</paragraph>
<paragraph id="8B9A3681FF96FFC2FCA366F2E6FDFCAC" blockId="19.[810,1460,232,883]" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FCA366F2E1A5FEDF" box="[810,959,232,256]" italics="true" pageId="19" pageNumber="20">Type locality: ‘</emphasis>
Bivouac No. 5, Loser Trail, Atjeh’ (now known as
<location id="8EFA605AFF96FFC2FCA36713E18DFEFF" LSID="urn:lsid:plazi:treatment:038C8797FF96FFC4FF16631EE76CFD80:8EFA605AFF96FFC2FCA36713E18DFEFF" box="[810,919,264,288]" country="Indonesia" latitude="3.85" longLatPrecision="784" longitude="97.12" municipality="Aceh" name="Mt. Leuser" pageId="19" pageNumber="20" stateProvince="Sumatra">Mt. Leuser</location>
trail,
<collectingMunicipality id="6BFEACFBFF96FFC2FC456713E618FEFF" box="[972,1026,264,288]" pageId="19" pageNumber="20">Aceh</collectingMunicipality>
,
<collectingRegion id="49E1F863FF96FFC2FB836713E67BFEFF" box="[1034,1121,264,288]" country="Indonesia" name="Sumatera Utara" pageId="19" pageNumber="20">Sumatra</collectingRegion>
,
<collectingCountry id="F3327611FF96FFC2FBE16713E6CBFEFF" box="[1128,1233,264,288]" name="Indonesia" pageId="19" pageNumber="20">Indonesia</collectingCountry>
). This locality is shown in the map of de
<bibRefCitation id="EFB44B70FF96FFC2FC47673CE71BFEE0" author="Schaubnsbb RM &amp; Riplbv SD" box="[974,1281,295,319]" pageId="19" pageNumber="20" pagination="311 - 68" refId="ref35395" refString="db Schaubnsbb RM, Riplbv SD. Zoological obsults of thb Gboogb Vandbobilt Sumatoan bxpbdition, 1936 - 1939 paot I: biods foom Atjbh. Proceedings of the Academy of Natural Sciences of Philadelphia 1939; 91: 311 - 68." type="journal article" year="1939">Schauensee and Ripley (1939)</bibRefCitation>
, and we have estimated it at
<emphasis id="B951EA93FF96FFC2FC1A675CE1B8FE81" box="[915,930,327,350]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
<geoCoordinate id="EE115046FF96FFC2FC2C675DE1EBFE81" box="[933,1009,326,350]" degrees="3.85" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="3.85">3.85º N</geoCoordinate>
,
<geoCoordinate id="EE115046FF96FFC2FC70675CE655FE81" box="[1017,1103,327,351]" degrees="97.12" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="97.12">97.12º E</geoCoordinate>
,
<quantity id="4CDD9B64FF96FFC2FBDF675CE6B8FE81" box="[1110,1186,327,351]" metricMagnitude="3" metricUnit="m" metricValue="2.408" pageId="19" pageNumber="20" unit="m" value="2408.0">
<elevation id="0008D1B2FF96FFC2FBDF675CE6B8FE81" box="[1110,1186,327,351]" metricMagnitude="3" metricUnit="m" metricValue="2.408" pageId="19" pageNumber="20" unit="m" value="2408.0">2408 m</elevation>
</quantity>
a.s.l.The type locality could be georeferenced after we located two flanking sampling sites of the George Vanderbilt Sumatran Expedition (Ulmer’s field notes,
<bibRefCitation id="EFB44B70FF96FFC2FCA367BFE654FE62" author="Schaubnsbb RM &amp; Riplbv SD" box="[810,1102,420,445]" pageId="19" pageNumber="20" pagination="311 - 68" refId="ref35395" refString="db Schaubnsbb RM, Riplbv SD. Zoological obsults of thb Gboogb Vandbobilt Sumatoan bxpbdition, 1936 - 1939 paot I: biods foom Atjbh. Proceedings of the Academy of Natural Sciences of Philadelphia 1939; 91: 311 - 68." type="journal article" year="1939">Schauensee and Ripley 1939</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF96FFC2FBD067BFE6CAFE62" author="Millbo GS" box="[1113,1232,420,445]" pageId="19" pageNumber="20" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller 1942</bibRefCitation>
): Mt. Leuser peak no. 1 (
<geoCoordinate id="EE115046FF96FFC2FCC167DFE1C9FE04" box="[840,979,452,476]" degrees="3.791072" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="3.791072">3.791072º N</geoCoordinate>
,
<geoCoordinate id="EE115046FF96FFC2FC6867DFE669FE03" box="[993,1139,452,476]" degrees="97.211351" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="97.21135">97.211351º E</geoCoordinate>
) and Blangbeke (
<emphasis id="B951EA93FF96FFC2FAB867DEE720FE03" box="[1329,1338,453,476]" italics="true" pageId="19" pageNumber="20">c</emphasis>
.
<geoCoordinate id="EE115046FF96FFC2FAC367DFE124FE24" degrees="3.891223" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="3.891223">3.891223º N</geoCoordinate>
,
<geoCoordinate id="EE115046FF96FFC2FCC267F8E1C6FE24" box="[843,988,483,507]" degrees="97.152307" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="97.152306">97.152307º E</geoCoordinate>
). Blangbeke could be easily georeferenced by the authors of this manuscript since it is the only ‘blang’ (open area) with a river on a ‘flat plateau of about 7000 ft’ found to the south of ‘Base camp (3600 ft), Blangnanga’, which is
<quantity id="4CDD9B64FF96FFC2FAB2645AE766FD86" box="[1339,1404,577,601]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="km" value="20.0">20 km</quantity>
West of the town of Blankejeren according to Fred Ulmer’s notes. The river is called ‘Waih Blang Bebeke’, which resembles the locality’s name. The type locality
<emphasis id="B951EA93FF96FFC2FBA96484E63CFD69" box="[1056,1062,671,694]" italics="true" pageId="19" pageNumber="20">‘</emphasis>
Bivouac No. 5’ is shown on the map of de
<bibRefCitation id="EFB44B70FF96FFC2FCC364A5E69EFD09" author="Schaubnsbb RM &amp; Riplbv SD" box="[842,1156,702,726]" pageId="19" pageNumber="20" pagination="311 - 68" refId="ref35395" refString="db Schaubnsbb RM, Riplbv SD. Zoological obsults of thb Gboogb Vandbobilt Sumatoan bxpbdition, 1936 - 1939 paot I: biods foom Atjbh. Proceedings of the Academy of Natural Sciences of Philadelphia 1939; 91: 311 - 68." type="journal article" year="1939">Schauensee and Ripley (1939)</bibRefCitation>
, just south of the Waih Blang Bebeke headwater. The approximate bivouac location was estimated in this manuscript through the combination of this map and information retrieved with Google Earth topographic layers and street view, and Fred Ulmer’s references and pictures housed at the
<collectionCode id="ED34AE44FF96FFC2FCEE6540E1BDFCAC" box="[871,935,859,883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">ANSP</collectionCode>
Mammal Department archives.
</paragraph>
<paragraph id="8B9A3681FF96FFC2FCA3658EE75BFBF5" blockId="19.[810,1460,917,1066]" pageId="19" pageNumber="20">
<materialsCitation id="3B4D3CDCFF96FFC2FCA3658EE7AAFC72" box="[810,1456,917,941]" collectionCode="ANSP" pageId="19" pageNumber="20" specimenCode="ANSP 20379, 20380, 20381" specimenCount="9" typeStatus="paratype">
<emphasis id="B951EA93FF96FFC2FCA3658EE1ACFC72" box="[810,950,917,941]" italics="true" pageId="19" pageNumber="20">
<typeStatus id="549E8823FF96FFC2FCA3658EE192FC73" box="[810,904,917,940]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D23FD08FF96FFC2FC1C658EE1BFFC72" box="[917,933,917,941]" pageId="19" pageNumber="20">9</specimenCount>
):
</emphasis>
<specimenCode id="DB839EFAFF96FFC2FC34658EE65EFC72" box="[957,1092,917,941]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">ANSP 20379</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FBC2658EE697FC72" box="[1099,1165,917,941]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20380</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FB1C658EE6C0FC72" box="[1173,1242,917,941]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20381</specimenCode>
: same as type locality
</materialsCitation>
;
<materialsCitation id="3B4D3CDCFF96FFC2FCA365AFE710FC13" box="[810,1290,948,972]" collectionCode="ANSP" elevation="2104" location="Blang Beke" municipality="Aceh" pageId="19" pageNumber="20" specimenCode="ANSP 20374, 20375, 20376, 20377, 20378" specimenCount="1" stateProvince="Sumatra" typeStatus="paratype">
<specimenCode id="DB839EFAFF96FFC2FCA365AFE1ABFC13" box="[810,945,948,972]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">ANSP 20374</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FC3365AFE1E6FC13" box="[954,1020,948,972]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20375</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FB8D65AFE65CFC13" box="[1028,1094,948,972]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20376</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FBC665AFE68BFC13" box="[1103,1169,948,972]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20377</specimenCode>
,
<specimenCode id="DB839EFAFF96FFC2FB1065AFE6C1FC13" box="[1177,1243,948,972]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">20378</specimenCode>
, and
</materialsCitation>
<materialsCitation id="3B4D3CDCFF96FFC2FA8665AFE1ADFBD4" collectionCode="USNM" elevation="2104" location="Blang Beke" municipality="Aceh" pageId="19" pageNumber="20" specimenCode="USNM 271033" specimenCount="1" stateProvince="Sumatra" typeStatus="paratype">
<specimenCode id="DB839EFAFF96FFC2FA8665AFE7AAFC13" box="[1295,1456,948,972]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">USNM 271033</specimenCode>
: ‘Blangbeke, Atjeh’ (=
<location id="8EFA605AFF96FFC2FC7E65C8E677FC34" LSID="urn:lsid:plazi:treatment:038C8797FF96FFC4FF16631EE76CFD80:8EFA605AFF96FFC2FC7E65C8E677FC34" box="[1015,1133,979,1003]" municipality="Aceh" name="Blang Beke" pageId="19" pageNumber="20" stateProvince="Sumatra">Blang Beke</location>
,
<collectingMunicipality id="6BFEACFBFF96FFC2FBFD65C8E6B0FC34" box="[1140,1194,979,1003]" pageId="19" pageNumber="20">Aceh</collectingMunicipality>
,
<collectingRegion id="49E1F863FF96FFC2FB3B65C8E710FC34" box="[1202,1290,979,1003]" pageId="19" pageNumber="20">Sumatra</collectingRegion>
; elevation
<elevation id="0008D1B2FF96FFC2FAFB65C8E1B3FBD4" metricMagnitude="3" metricUnit="m" metricValue="2.1035" metricValueMax="2.134" metricValueMin="2.073" pageId="19" pageNumber="20" unit="m" value="2103.5" valueMax="2134.0" valueMin="2073.0">
<quantity id="4CDD9B64FF96FFC2FAFB65C8E16DFBD5" metricMagnitude="3" metricUnit="m" metricValue="2.1035" metricValueMax="2.134" metricValueMin="2.073" pageId="19" pageNumber="20" unit="m" value="2103.5" valueMax="2134.0" valueMin="2073.0">2073– 2134 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
<materialsCitation id="3B4D3CDCFF96FFC2FC3265E8E75BFBF5" collectionCode="USNM" elevation="2104" location="Blang Beke" municipality="Aceh" pageId="19" pageNumber="20" specimenCode="USNM 271033" specimenCount="1" stateProvince="Sumatra" typeStatus="paratype">
<typeStatus id="549E8823FF96FFC2FC3265E8E60EFBD4" box="[955,1044,1011,1035]" pageId="19" pageNumber="20">Paratype</typeStatus>
<specimenCode id="DB839EFAFF96FFC2FB9165E8E684FBD4" box="[1048,1182,1011,1035]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">ANSP 20377</specimenCode>
was illustrated by
<collectorName id="26D05357FF96FFC2FADA65E8E192FBF5" pageId="19" pageNumber="20">Frederick A. Ulmer</collectorName>
(Supporting Information, File S1,
<figureCitation id="131E2A04FF96FFC2FB6D6209E722FBF5" box="[1252,1336,1042,1066]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." figureDoi="http://doi.org/10.5281/zenodo.14510291" httpUri="https://zenodo.org/record/14510291/files/figure.png" pageId="19" pageNumber="20">Fig. S11</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="8B9A3681FF96FFC2FCA36257E6B7FADF" blockId="19.[810,1460,1099,1280]" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FCA36257E666FBBC" box="[810,1148,1100,1123]" italics="true" pageId="19" pageNumber="20">Representative DNA sequences:</emphasis>
Deposited in GenBank with the following accession numbers: OR138101 and OR138102 (mitochondrial genomes, two representatives), OR554452-OR554454 (
<emphasis id="B951EA93FF96FFC2FBBA62B1E673FB1E" box="[1075,1129,1194,1217]" italics="true" pageId="19" pageNumber="20">
<collectionCode id="ED34AE44FF96FFC2FBBA62B1E673FB1E" box="[1075,1129,1194,1217]" pageId="19" pageNumber="20">GHR</collectionCode>
</emphasis>
, three representatives). DNA sequences associated with the following voucher specimens:
<collectionCode id="ED34AE44FF96FFC2FCA362F3E16CFADF" box="[810,886,1256,1280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">USNM</collectionCode>
271034,
<collectionCode id="ED34AE44FF96FFC2FC5D62F3E60FFADF" box="[980,1045,1256,1280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="20">ANSP</collectionCode>
20377, 20380.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF96FFC2FCA3633AE634F98C" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8B9A3681FF96FFC2FCA3633AE634F98C" blockId="19.[810,1459,1313,1619]" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FCA3633AE183FAE6" box="[810,921,1313,1337]" italics="true" pageId="19" pageNumber="20">Etymology:</emphasis>
The specific name acknowledges the voracious behaviour that Frederick Ulmer, the collector of the type series, described in his field notes: ‘They were voracious beasts often devouring the whole bait before springing the trap. Ham rind, coconut, meat, and walnuts were eaten. One shrew partially devoured the chicken head bait of a steel trap before getting caught in a nearby Schuyler trap baited with ham rind’ (
<collectionCode id="ED34AE44FF96FFC2FAED63C6E7A9FA2A" box="[1380,1459,1501,1525]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">USNM</collectionCode>
271033). We recommend the common names Leuser Gymnure, Salak Ba’a Leuser, and Gimnuro de Leuser, in English, Malay, and Spanish, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF96FFC5FCA36065E66BFAB1" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8B9A3681FF96FFC5FCA36065E063FD24" blockId="19.[810,1460,1661,1967]" lastBlockId="20.[129,778,144,764]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FCA36065E18AF94A" box="[810,912,1662,1685]" italics="true" pageId="19" pageNumber="20">Diagnosis:</emphasis>
A medium-sized
<taxonomicName id="4C254D02FF96FFC2FBDE6066E6B6F94A" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1111,1196,1661,1685]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B951EA93FF96FFC2FBDE6066E6B6F94A" box="[1111,1196,1661,1685]" italics="true" pageId="19" pageNumber="20">Hylomys</emphasis>
</taxonomicName>
(average
<collectionCode id="ED34AE44FF96FFC2FA9A6065E72DF94A" box="[1299,1335,1662,1685]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="19" pageNumber="20" type="Herbarium">HB</collectionCode>
=
<quantity id="4CDD9B64FF96FFC2FADD6066E7B5F94A" box="[1364,1455,1661,1685]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="19" pageNumber="20" unit="mm" value="123.0">123 mm</quantity>
, W =
<quantity id="4CDD9B64FF96FFC2FCD66086E185F96A" box="[863,927,1693,1717]" metricMagnitude="-2" metricUnit="kg" metricValue="4.4399999999999995" pageId="19" pageNumber="20" unit="g" value="44.4">44.4 g</quantity>
) characterized by a thick and soft, dark brown fur, a long and monocoloured black tail (average
<collectionCode id="ED34AE44FF96FFC2FA9560A7E734F90C" box="[1308,1326,1724,1747]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="20">T</collectionCode>
=
<quantity id="4CDD9B64FF96FFC2FAC360A7E7B4F90C" box="[1354,1454,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="19" pageNumber="20" unit="mm" value="21.4">21.4 mm</quantity>
;
<collectionCode id="ED34AE44FF96FFC2FCA360C0E126F92C" box="[810,828,1755,1779]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="20">T</collectionCode>
/
<collectionCode id="ED34AE44FF96FFC2FCCC60C0E177F92C" box="[837,877,1755,1779]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="19" pageNumber="20" type="Herbarium">HB</collectionCode>
= 17.5%), and long ears (average E =
<quantity id="4CDD9B64FF96FFC2FA9660C0E79FF92C" box="[1311,1413,1755,1779]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="19" pageNumber="20" unit="mm" value="19.1">19.1 mm</quantity>
, E/
<collectionCode id="ED34AE44FF96FFC2FCA360E0E140F8CC" box="[810,858,1787,1811]" pageId="19" pageNumber="20">GLS</collectionCode>
= 56.3%). Ventral coloration is paler than the dorsum, with buff-tipped, dark-grey hairs. Brown-yellowish hue in the throat area is generally inconspicuous and just present in females (but these could be due to phenological sampling biases: specimen series collected in April). All specimens have an ochraceous hue around the cloaca/inguinal area. Fore/hindfeet and tail have a lighter and darker coloration than dorsum, respectively. The skull of this species is distinguished from all other congeneric species by a unique combination of the following characters: acute-angled notch between premaxilla tips; long nasals; narrow rostrum; poorly developed antorbital ridge/flange; broad interorbital constriction; small but prominent supraorbital process in males, which is generally inconspicuous in females; relatively smooth frontal bone and dorsal region of maxillary; anterior palatine foramina (generally) anterior or (rarely) at level of maxilla/palatine suture; generally thick and robust postpalatine torus; lack of posterior nasal spine; elongated pterygoid process; robust and globular ectotympanic process that tappers dorsoventrally towards proximal end; gracile mandible, with relatively short vertical height at coronoid and length of angular process; procumbent incisors; single root in P2; P2 and P3 lack posterior cuspules, even in subadults and young specimens; absent labial cingulum in P4; anterior margin of p1 crown does not extend to root of c1; p3 is much larger than p2 and larger than p1, p1 is generally larger than p2; posterior cuspule of p2 absent, even in subadults and young specimens; single root in p3.
</paragraph>
<paragraph id="8B9A3681FF91FFC5FF086538E0D2FACE" blockId="20.[128,779,802,1986]" pageId="20" pageNumber="21">
<emphasis id="B951EA93FF91FFC5FF086538E312FCE5" box="[129,264,803,826]" italics="true" pageId="20" pageNumber="21">Comparisons:</emphasis>
Compared with its closest geographical and genetic relative,
<taxonomicName id="4C254D02FF91FFC5FEBB6559E3DFFC85" authorityName="Sody" authorityYear="1933" box="[306,453,834,858]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FEBB6559E3DFFC85" box="[306,453,834,858]" italics="true" pageId="20" pageNumber="21">Hylomys maxi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF91FFC5FE5F6558E028FC85" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[470,562,835,858]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FE5F6558E028FC85" box="[470,562,835,858]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
is distinguished by its smaller (average
<collectionCode id="ED34AE44FF91FFC5FED16579E366FCA6" box="[344,380,866,889]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FE10657AE3EEFCA6" box="[409,500,865,889]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="21" unit="mm" value="123.0">123 mm</quantity>
; W =
<quantity id="4CDD9B64FF91FFC5FDB3657AE064FCA5" box="[570,638,865,890]" metricMagnitude="-2" metricUnit="kg" metricValue="4.4399999999999995" pageId="20" pageNumber="21" unit="g" value="44.4">44.4 g</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FD19657AE0DAFCA6" box="[656,704,865,889]" pageId="20" pageNumber="21">GLS</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FD54657AE2B7FC47" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="20" pageNumber="21" unit="mm" value="34.5">34.5 mm</quantity>
; LMA =
<quantity id="4CDD9B64FF91FFC5FE86659AE344FC47" box="[271,350,897,921]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="21" unit="mm" value="24.0">24 mm</quantity>
;
<figureCitation id="131E2A04FF91FFC5FEE2659AE3AFFC46" box="[363,437,897,921]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
) vs. larger size (
<collectionCode id="ED34AE44FF91FFC5FDEC659AE096FC46" box="[613,652,897,921]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FD20659AE118FC47" box="[681,770,897,921]" metricMagnitude="-1" metricUnit="m" metricValue="1.39" pageId="20" pageNumber="21" unit="mm" value="139.0">139 mm</quantity>
; W =
<quantity id="4CDD9B64FF91FFC5FF3E65BBE2E0FC67" box="[183,250,928,952]" metricMagnitude="-2" metricUnit="kg" metricValue="6.42" pageId="20" pageNumber="21" unit="g" value="64.2">64.2 g</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FE8E65BBE32DFC67" box="[263,311,928,952]" pageId="20" pageNumber="21">GLS</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FEDA65BBE3AEFC68" box="[339,436,928,952]" metricMagnitude="-2" metricUnit="m" metricValue="3.69" pageId="20" pageNumber="21" unit="mm" value="36.9">36.9 mm</quantity>
; LMA =
<quantity id="4CDD9B64FF91FFC5FD9D65BBE06EFC68" box="[532,628,928,952]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="20" pageNumber="21" unit="mm" value="26.3">26.3 mm</quantity>
;
<figureCitation id="131E2A04FF91FFC5FD0965BBE0D2FC67" box="[640,712,928,952]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="20" pageNumber="21">Fig. 9B</figureCitation>
), uniformly black and relatively longer (average
<collectionCode id="ED34AE44FF91FFC5FDC765DBE07AFC08" box="[590,608,960,983]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FDF565A4E0C4FC08" box="[636,734,959,983]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="20" pageNumber="21" unit="mm" value="21.4">21.4 mm</quantity>
,
<collectionCode id="ED34AE44FF91FFC5FD6265A4E0E5FC08" box="[747,767,959,983]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
/ HB = 17.1–18.8%) vs. bicoloured (except specimen
<collectionCode id="ED34AE44FF91FFC5FD3165C4E113FC29" box="[696,777,991,1014]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">RMNH</collectionCode>
23896) and relatively shorter tail (average
<collectionCode id="ED34AE44FF91FFC5FDC465E5E045FBCA" box="[589,607,1022,1045]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FDF265E5E0C7FBCA" box="[635,733,1022,1046]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="20" pageNumber="21" unit="mm" value="16.2">16.2 mm</quantity>
,
<collectionCode id="ED34AE44FF91FFC5FD6265E5E0E5FBC9" box="[747,767,1022,1046]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
/ HB = 5.11–16.9%) (Supporting Information, File S1,
<figureCitation id="131E2A04FF91FFC5FD6A6206E2A2FB8A" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10A</figureCitation>
), and a relatively longer (E/
<collectionCode id="ED34AE44FF91FFC5FD806226E027FB8A" box="[521,573,1085,1109]" pageId="20" pageNumber="21">GLS</collectionCode>
= 54.8–55.9%) vs. shorter ear (E/
<collectionCode id="ED34AE44FF91FFC5FE966247E349FBAB" box="[287,339,1116,1140]" pageId="20" pageNumber="21">GLS</collectionCode>
= 39.3–48.2%) (Supporting Information, File S1,
<figureCitation id="131E2A04FF91FFC5FF526260E35AFB4C" box="[219,320,1147,1171]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10B</figureCitation>
). Its fur is softer, denser, and longer (
<quantity id="4CDD9B64FF91FFC5FD566260E2B3FB6D" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="20" pageNumber="21" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
at axial region) than
<taxonomicName id="4C254D02FF91FFC5FE0E6280E3C3FB6D" authorityName="Sody" authorityYear="1933" box="[391,473,1179,1202]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FE0E6280E3C3FB6D" box="[391,473,1179,1202]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(
<quantity id="4CDD9B64FF91FFC5FE616281E058FB6D" box="[488,578,1178,1202]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="20" pageNumber="21" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
at axial region), although higher elevation (&gt;
<quantity id="4CDD9B64FF91FFC5FE0662A1E3FBFB0E" box="[399,481,1210,1234]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="m" value="1100.0">1100 m</quantity>
a.s.l.) Sumatran
<taxonomicName id="4C254D02FF91FFC5FD0362A0E0C0FB0D" authorityName="Sody" authorityYear="1933" box="[650,730,1211,1234]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FD0362A0E0C0FB0D" box="[650,730,1211,1234]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
specimen furs were not measured, and this trait is subject to great ecophenotypic variation in this genus (
<bibRefCitation id="EFB44B70FF91FFC5FD8762E2E0A2FACE" author="Rubdi M &amp; Chapuisat M &amp; Iskandao D" box="[526,696,1273,1297]" pageId="20" pageNumber="21" pagination="965 - 78" refId="ref35216" refString="Rubdi M, Chapuisat M, Iskandao D. Taxonomic status of Hylomys parous and Hylomys suillus (Insbctivooa: Eoinacbidab): biochbmical and moophological analvsbs. Journal of Mammalogy 1994; 75: 965 - 78. hups: // doi. oog / 10.2307 / 1382478" type="journal article" year="1994">
Ruedi
<emphasis id="B951EA93FF91FFC5FDD862E2E09AFACE" box="[593,640,1273,1297]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
1994
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A3681FF91FFC5FF156303E786FE9A" blockId="20.[128,779,802,1986]" lastBlockId="20.[825,1476,144,1390]" pageId="20" pageNumber="21">
The skull of
<taxonomicName id="4C254D02FF91FFC5FEAD6302E367FAEF" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[292,381,1305,1328]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FEAD6302E367FAEF" box="[292,381,1305,1328]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
has a narrower and more gracile appearance than
<taxonomicName id="4C254D02FF91FFC5FE906323E371FA90" authorityName="Sody" authorityYear="1933" box="[281,363,1336,1359]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FE906323E371FA90" box="[281,363,1336,1359]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. The notch between the premaxilla tips is acute vs. obtuse in
<taxonomicName id="4C254D02FF91FFC5FEE8634CE3AEFAB1" authorityName="Sody" authorityYear="1933" box="[353,436,1367,1390]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FEE8634CE3AEFAB1" box="[353,436,1367,1390]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A04FF91FFC5FE4E634CE014FAB0" box="[455,526,1367,1391]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
). Its rostrum breadth is narrower (4.5–5.0 mm) vs. broader (
<quantity id="4CDD9B64FF91FFC5FD88636DE09FFA51" box="[513,645,1398,1422]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.4" metricValueMin="5.2" pageId="20" pageNumber="21" unit="mm" value="5.8" valueMax="6.4" valueMin="5.2">5.2–6.4 mm</quantity>
) in
<taxonomicName id="4C254D02FF91FFC5FD38636CE11EFA51" authorityName="Sody" authorityYear="1933" box="[689,772,1399,1422]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FD38636CE11EFA51" box="[689,772,1399,1422]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Its nasals are relatively longer (average
<collectionCode id="ED34AE44FF91FFC5FD96638EE05EFA72" box="[543,580,1429,1453]" pageId="20" pageNumber="21">NL</collectionCode>
/
<collectionCode id="ED34AE44FF91FFC5FDC5638EE064FA72" box="[588,638,1429,1453]" pageId="20" pageNumber="21">GLS</collectionCode>
= 36.1%) vs. shorter in
<taxonomicName id="4C254D02FF91FFC5FF7363AEE349FA13" authorityName="Sody" authorityYear="1933" box="[250,339,1461,1484]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FF7363AEE349FA13" box="[250,339,1461,1484]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(average
<collectionCode id="ED34AE44FF91FFC5FE4C63AEE3F0FA12" box="[453,490,1461,1485]" pageId="20" pageNumber="21">NL</collectionCode>
/
<collectionCode id="ED34AE44FF91FFC5FE7B63AEE03EFA12" box="[498,548,1461,1485]" pageId="20" pageNumber="21">GLS</collectionCode>
= 32.4%; Supporting Information, File S1,
<figureCitation id="131E2A04FF91FFC5FED463CFE3D9FA33" box="[349,451,1492,1516]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10C</figureCitation>
). Antorbital ridge/flange is undeveloped vs. well developed in
<taxonomicName id="4C254D02FF91FFC5FE5C63EFE032F9D4" authorityName="Sody" authorityYear="1933" box="[469,552,1524,1547]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FE5C63EFE032F9D4" box="[469,552,1524,1547]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
, particularly between the lacrimal flange and supraorbital process. Its frontal bone and dorsal region of maxillary has a relatively smooth vs. rugged appearance in
<emphasis id="B951EA93FF91FFC5FF776049E34CF9B6" box="[254,342,1618,1641]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C254D02FF91FFC5FF776049E348F9B6" authorityName="Sody" authorityYear="1933" box="[254,338,1618,1641]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">H. maxi</taxonomicName>
.
</emphasis>
Its interorbital constriction is proportionately broader than
<taxonomicName id="4C254D02FF91FFC5FEC3606AE384F957" authorityName="Sody" authorityYear="1933" box="[330,414,1649,1672]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FEC3606AE384F957" box="[330,414,1649,1672]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(Supporting Information, File S1,
<figureCitation id="131E2A04FF91FFC5FF08608BE2FCF977" box="[129,230,1680,1704]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10D</figureCitation>
). Zygomata and supraorbital processes are less robust than in
<taxonomicName id="4C254D02FF91FFC5FF5C60ABE333F918" authorityName="Sody" authorityYear="1933" box="[213,297,1712,1735]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FF5C60ABE333F918" box="[213,297,1712,1735]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Anterior palatine foramina end anteriorly or (rarely) at maxilla/palatine suture vs. posterior to maxilla/palatine suture in
<taxonomicName id="4C254D02FF91FFC5FE8260F4E340F8D9" authorityName="Sody" authorityYear="1933" box="[267,346,1775,1798]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FE8260F4E340F8D9" box="[267,346,1775,1798]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Postpalatine torus is generally thicker and more robust than in
<taxonomicName id="4C254D02FF91FFC5FEDC6115E3BDF8FA" authorityName="Sody" authorityYear="1933" box="[341,423,1806,1829]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FEDC6115E3BDF8FA" box="[341,423,1806,1829]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Posterior nasal spine is absent vs. present in
<taxonomicName id="4C254D02FF91FFC5FF606136E323F89B" authorityName="Sody" authorityYear="1933" box="[233,313,1837,1860]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FF606136E323F89B" box="[233,313,1837,1860]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Postpalatal length is shorter (
<quantity id="4CDD9B64FF91FFC5FDE16136E118F89A" box="[616,770,1837,1861]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.23" metricValueMin="1.1099999999999999" pageId="20" pageNumber="21" unit="mm" value="11.7" valueMax="12.3" valueMin="11.1">11.1–12.3 mm</quantity>
) vs. longer (
<quantity id="4CDD9B64FF91FFC5FF676157E393F8BB" box="[238,393,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.42" metricValueMin="1.23" pageId="20" pageNumber="21" unit="mm" value="13.25" valueMax="14.2" valueMin="12.3">12.3–14.2 mm</quantity>
) in
<taxonomicName id="4C254D02FF91FFC5FE396156E01AF8BB" authorityName="Sody" authorityYear="1933" box="[432,512,1869,1892]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FE396156E01AF8BB" box="[432,512,1869,1892]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
, while braincase is similar in breadth (average
<collectionCode id="ED34AE44FF91FFC5FEDC6177E36EF85C" box="[341,372,1900,1923]" country="0" name="Buffalo Bill Museum" pageId="20" pageNumber="21">BB</collectionCode>
= 14.8 vs.
<quantity id="4CDD9B64FF91FFC5FE636170E020F85C" box="[490,570,1899,1923]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="mm" value="15.0">15 mm</quantity>
). Epipterygoid processes are lateroventrally vs. lateral oriented in
<taxonomicName id="4C254D02FF91FFC5FDD86190E0BAF87D" authorityName="Sody" authorityYear="1933" box="[593,672,1931,1954]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FDD86190E0BAF87D" box="[593,672,1931,1954]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Its robust ectotympanic process thickens dorsoventrally towards the tip, while it is more gracile and dorsoventrally flattened in
<taxonomicName id="4C254D02FF91FFC5FAF9668AE7D8FF77" authorityName="Sody" authorityYear="1933" box="[1392,1474,145,168]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FAF9668AE7D8FF77" box="[1392,1474,145,168]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A04FF91FFC5FCCD66ABE18DFF18" box="[836,919,175,199]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">
<emphasis id="B951EA93FF91FFC5FCCD66ABE178FF18" box="[836,866,176,199]" italics="true" pageId="20" pageNumber="21">Fig</emphasis>
.
<emphasis id="B951EA93FF91FFC5FCE766B4E18DFF18" box="[878,919,175,199]" italics="true" pageId="20" pageNumber="21">10B</emphasis>
</figureCitation>
). Occipital crest absent or inconspicuous and shorter vs. prominent and longer in
<emphasis id="B951EA93FF91FFC5FBE966D4E6A2FF39" box="[1120,1208,207,230]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C254D02FF91FFC5FBE966D4E6AEFF39" authorityName="Sody" authorityYear="1933" box="[1120,1204,207,230]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">H. maxi</taxonomicName>
.
</emphasis>
Its mandible is more gracile, with a shorter vertical height at coronoid and shorter length of angular process (
<collectionCode id="ED34AE44FF91FFC5FB8F6716E65EFEFA" box="[1030,1092,269,293]" pageId="20" pageNumber="21">HCO</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FBE86716E6C4FEFA" box="[1121,1246,269,293]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.6" metricValueMin="7.4" pageId="20" pageNumber="21" unit="mm" value="8.0" valueMax="8.6" valueMin="7.4">7.4–8.6 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FB676715E704FEF9" box="[1262,1310,270,294]" pageId="20" pageNumber="21">LAP</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FAB36716E7ADFEFA" box="[1338,1463,269,293]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.1" metricValueMin="7.9" pageId="20" pageNumber="21" unit="mm" value="8.5" valueMax="9.1" valueMin="7.9">7.9–9.1 mm</quantity>
) than
<taxonomicName id="4C254D02FF91FFC5FCE56736E1A7FE9B" authorityName="Sody" authorityYear="1933" box="[876,957,301,324]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FCE56736E1A7FE9B" box="[876,957,301,324]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
(
<collectionCode id="ED34AE44FF91FFC5FC436736E612FE9A" box="[970,1032,301,325]" pageId="20" pageNumber="21">HCO</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FBAD6736E6B6FE9B" box="[1060,1196,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="9.649999999999999" metricValueMax="10.7" metricValueMin="8.6" pageId="20" pageNumber="21" unit="mm" value="9.649999999999999" valueMax="10.7" valueMin="8.6">8.6–10.7 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FB306736E6F3FE9A" box="[1209,1257,301,325]" pageId="20" pageNumber="21">LAP</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FA8C6736E796FE9B" box="[1285,1420,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.8" metricValueMin="8.8" pageId="20" pageNumber="21" unit="mm" value="9.8" valueMax="10.8" valueMin="8.8">8.8–10.8 mm</quantity>
).
</paragraph>
<paragraph id="8B9A3681FF91FFC5FCDC6757E71EFCC5" blockId="20.[825,1476,144,1390]" pageId="20" pageNumber="21">
Its dentition appearance is less robust than that of
<taxonomicName id="4C254D02FF91FFC5FAE06756E7A6FEBB" authorityName="Sody" authorityYear="1933" box="[1385,1468,333,356]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FAE06756E7A6FEBB" box="[1385,1468,333,356]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
, with smaller teeth and more space between incisors, canine, and premolars than
<taxonomicName id="4C254D02FF91FFC5FC546790E635FE7D" authorityName="Sody" authorityYear="1933" box="[989,1071,395,418]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FC546790E635FE7D" box="[989,1071,395,418]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. I1 is more procumbent than
<taxonomicName id="4C254D02FF91FFC5FAE36790E7A6FE7D" authorityName="Sody" authorityYear="1933" box="[1386,1468,395,418]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FAE36790E7A6FE7D" box="[1386,1468,395,418]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. P1 is usually similar in size to P3 vs. generally much larger in
<taxonomicName id="4C254D02FF91FFC5FCB067D1E194FE3E" authorityName="Sody" authorityYear="1933" box="[825,910,458,481]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FCB067D1E194FE3E" box="[825,910,458,481]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. P4 labial cingulum absent or inconspicuous vs. well developed in
<taxonomicName id="4C254D02FF91FFC5FC4067F2E607FDDF" authorityName="Sody" authorityYear="1933" box="[969,1053,489,512]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FC4067F2E607FDDF" box="[969,1053,489,512]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Molars are smaller (M1M1 =
<quantity id="4CDD9B64FF91FFC5FAE167F2E17FFDC0" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.08" metricValueMin="0.98" pageId="20" pageNumber="21" unit="mm" value="10.3" valueMax="10.8" valueMin="9.8">9.8–10.8 mm</quantity>
; P4M3 =
<quantity id="4CDD9B64FF91FFC5FC456413E65FFDC0" box="[972,1093,520,544]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="8.2" metricValueMin="7.5" pageId="20" pageNumber="21" unit="mm" value="7.85" valueMax="8.2" valueMin="7.5">7.5–8.2 mm</quantity>
) vs. larger (M1M1 =
<quantity id="4CDD9B64FF91FFC5FAAF6413E7A1FDC0" box="[1318,1467,520,544]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.25" metricValueMin="1.09" pageId="20" pageNumber="21" unit="mm" value="11.7" valueMax="12.5" valueMin="10.9">10.9–12.5 mm</quantity>
; P4M3 =
<quantity id="4CDD9B64FF91FFC5FC11643CE60BFDE0" box="[920,1041,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.9" metricValueMin="8.3" pageId="20" pageNumber="21" unit="mm" value="9.1" valueMax="9.9" valueMin="8.3">8.3–9.9 mm</quantity>
) in
<taxonomicName id="4C254D02FF91FFC5FBB66433E68BFDE0" authorityName="Sody" authorityYear="1933" box="[1087,1169,552,575]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FBB66433E68BFDE0" box="[1087,1169,552,575]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. Its lower toothrow is shorter (
<quantity id="4CDD9B64FF91FFC5FCC8645CE1C6FD80" box="[833,988,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.77" metricValueMin="1.66" pageId="20" pageNumber="21" unit="mm" value="17.15" valueMax="17.7" valueMin="16.6">16.6–17.7 mm</quantity>
) vs. longer (
<quantity id="4CDD9B64FF91FFC5FBD3645CE6ECFD80" box="[1114,1270,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.825" metricValueMax="1.89" metricValueMin="1.76" pageId="20" pageNumber="21" unit="mm" value="18.25" valueMax="18.9" valueMin="17.6">17.6–18.9 mm</quantity>
) in
<emphasis id="B951EA93FF91FFC5FA97645CE76FFD81" box="[1310,1397,583,606]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C254D02FF91FFC5FA97645CE76BFD81" authorityName="Sody" authorityYear="1933" box="[1310,1393,583,606]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">H. maxi</taxonomicName>
.
</emphasis>
Crown height of c1 is generally greater in
<taxonomicName id="4C254D02FF91FFC5FB21647CE71BFDA1" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1192,1281,615,638]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FB21647CE71BFDA1" box="[1192,1281,615,638]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
than
<taxonomicName id="4C254D02FF91FFC5FAC9647CE789FDA1" authorityName="Sody" authorityYear="1933" box="[1344,1427,615,638]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FAC9647CE789FDA1" box="[1344,1427,615,638]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
, the latter species c1 is more expanded anteriorly, giving it a less prominent appearance. Anterior margin of p1 crown does not extend anteriorly to the root of c1, but it does in
<taxonomicName id="4C254D02FF91FFC5FABA64DEE79FFD03" authorityName="Sody" authorityYear="1933" box="[1331,1413,709,732]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FABA64DEE79FFD03" box="[1331,1413,709,732]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
. p3 is usually much larger than p
<quantity id="4CDD9B64FF91FFC5FBCA64F8E673FD24" box="[1091,1129,739,763]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C254D02FF91FFC5FBE464FFE6D8FD24" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1133,1218,740,763]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FBE464FFE6D8FD24" box="[1133,1218,740,763]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
, while it has a similar size or p3 is just slightly larger than p
<quantity id="4CDD9B64FF91FFC5FB0A6518E6B3FCC4" box="[1155,1193,771,795]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C254D02FF91FFC5FB276518E6E4FCC5" authorityName="Sody" authorityYear="1933" box="[1198,1278,771,794]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FB276518E6E4FCC5" box="[1198,1278,771,794]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A3681FF91FFC5FCDC6539E66BFAB1" blockId="20.[825,1476,144,1390]" pageId="20" pageNumber="21">
Distinguished from
<taxonomicName id="4C254D02FF91FFC5FBAF6538E693FCE5" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1062,1161,803,826]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FBAF6538E693FCE5" box="[1062,1161,803,826]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
by its larger size (
<collectionCode id="ED34AE44FF91FFC5FAC36539E76BFCE5" box="[1354,1393,802,826]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FA046539E187FC86" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.28" metricValueMin="1.17" pageId="20" pageNumber="21" unit="mm" value="122.5" valueMax="128.0" valueMin="117.0">117– 128 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FC38655AE1FBFC86" box="[945,993,833,857]" pageId="20" pageNumber="21">GLS</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FC74655AE684FC86" box="[1021,1182,833,857]" metricMagnitude="-2" metricUnit="m" metricValue="3.4849999999999994" metricValueMax="3.63" metricValueMin="3.34" pageId="20" pageNumber="21" unit="mm" value="34.849999999999994" valueMax="36.3" valueMin="33.4">33.4–36.3 mm</quantity>
; LMA =
<quantity id="4CDD9B64FF91FFC5FA80655AE78DFC86" box="[1289,1431,833,857]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" metricValueMax="2.56" metricValueMin="2.3" pageId="20" pageNumber="21" unit="mm" value="24.3" valueMax="25.6" valueMin="23.0">23–25.6 mm</quantity>
vs. HB =
<quantity id="4CDD9B64FF91FFC5FCF5657BE611FCA7" box="[892,1035,864,888]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="20" pageNumber="21" unit="mm" value="107.5" valueMax="115.0" valueMin="100.0">100–115 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FB9F657AE65CFCA6" box="[1046,1094,865,889]" pageId="20" pageNumber="21">GLS</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FBEA657AE6E7FCA7" box="[1123,1277,865,889]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" metricValueMax="3.34" metricValueMin="2.92" pageId="20" pageNumber="21" unit="mm" value="31.299999999999997" valueMax="33.4" valueMin="29.2">29.2–33.4 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FA80657AE75BFCA6" box="[1289,1345,865,889]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13714" name="Institute for Agricultural Research of Mozambique" pageId="20" pageNumber="21" type="Herbarium">LMA</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FAD4657AE17CFC47" metricMagnitude="-2" metricUnit="m" metricValue="2.1149999999999998" metricValueMax="2.2199999999999998" metricValueMin="2.0100000000000002" pageId="20" pageNumber="21" unit="mm" value="21.15" valueMax="22.2" valueMin="20.1">20.1–22.2 mm</quantity>
) and black monocoloured and relatively shorter (average
<collectionCode id="ED34AE44FF91FFC5FA2F659BE7A0FC47" box="[1446,1466,896,920]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
/
<collectionCode id="ED34AE44FF91FFC5FCB065BBE147FC68" box="[825,861,928,951]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
= 17.5%) vs. bicoloured and relatively longer tail (average
<collectionCode id="ED34AE44FF91FFC5FCB065A4E151FC08" box="[825,843,959,983]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
/
<collectionCode id="ED34AE44FF91FFC5FCDD65A4E166FC08" box="[852,892,959,983]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
= 22.4%). Rostrum is longer (
<collectionCode id="ED34AE44FF91FFC5FB5565A4E709FC08" box="[1244,1299,959,983]" pageId="20" pageNumber="21">ROL</collectionCode>
= 15.2–17.3 vs. 13.6–15.0 mm) but similar in width (
<collectionCode id="ED34AE44FF91FFC5FB5665C5E70DFC29" box="[1247,1303,990,1014]" pageId="20" pageNumber="21">ROB</collectionCode>
= 4.52–5.05 vs.
<quantity id="4CDD9B64FF91FFC5FCB065E6E1A8FBCA" box="[825,946,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="4.715" metricValueMax="5.0" metricValueMin="4.43" pageId="20" pageNumber="21" unit="mm" value="4.715" valueMax="5.0" valueMin="4.43">4.43–5 mm</quantity>
) to
<taxonomicName id="4C254D02FF91FFC5FC5265E5E627FBCA" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[987,1085,1022,1045]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FC5265E5E627FBCA" box="[987,1085,1022,1045]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
, giving
<taxonomicName id="4C254D02FF91FFC5FB0465E5E6FEFBCA" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1165,1252,1022,1045]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FB0465E5E6FEFBCA" box="[1165,1252,1022,1045]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
a more elongated appearance. Toothrow length is longer (
<collectionCode id="ED34AE44FF91FFC5FB376206E6FAFBEA" box="[1214,1248,1053,1077]" country="India" name="Indian Museum" pageId="20" pageNumber="21">IM</collectionCode>
3Sa =
<quantity id="4CDD9B64FF91FFC5FAAF6207E7A1FBEB" box="[1318,1467,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" metricValueMax="1.85" metricValueMin="1.72" pageId="20" pageNumber="21" unit="mm" value="17.85" valueMax="18.5" valueMin="17.2">17.2–18.5 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FCB26227E147FB8B" box="[827,861,1084,1108]" country="India" name="Indian Museum" pageId="20" pageNumber="21">IM</collectionCode>
3I =
<quantity id="4CDD9B64FF91FFC5FC056227E639FB8C" box="[908,1059,1084,1108]" metricMagnitude="-2" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.77" metricValueMin="1.66" pageId="20" pageNumber="21" unit="mm" value="17.15" valueMax="17.7" valueMin="16.6">16.6–17.7 mm</quantity>
) than in
<taxonomicName id="4C254D02FF91FFC5FB0E6226E6F3FB8B" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1159,1257,1085,1108]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FB0E6226E6F3FB8B" box="[1159,1257,1085,1108]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
(
<collectionCode id="ED34AE44FF91FFC5FB716227E700FB8B" box="[1272,1306,1084,1108]" country="India" name="Indian Museum" pageId="20" pageNumber="21">IM</collectionCode>
3Sa =
<quantity id="4CDD9B64FF91FFC5FAD66227E7A5FB8C" box="[1375,1471,1084,1108]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="20" pageNumber="21" unit="mm" value="16.1">16.1 mm</quantity>
;
<collectionCode id="ED34AE44FF91FFC5FCB26240E147FBAC" box="[827,861,1115,1139]" country="India" name="Indian Museum" pageId="20" pageNumber="21">IM</collectionCode>
3I =
<quantity id="4CDD9B64FF91FFC5FC056240E60BFBAC" box="[908,1041,1115,1139]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="20" pageNumber="21" unit="mm" value="15.25" valueMax="16.0" valueMin="14.5">14.5–16 mm</quantity>
). Braincase and palate breadth are wider in
<taxonomicName id="4C254D02FF91FFC5FCD16260E1A8FB4D" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[856,946,1147,1170]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FCD16260E1A8FB4D" box="[856,946,1147,1170]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
(
<collectionCode id="ED34AE44FF91FFC5FC4D6260E1FDFB4C" box="[964,999,1147,1171]" country="0" name="Buffalo Bill Museum" pageId="20" pageNumber="21">BB</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FB8A6261E687FB4D" box="[1027,1181,1146,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" metricValueMax="1.51" metricValueMin="1.45" pageId="20" pageNumber="21" unit="mm" value="14.8" valueMax="15.1" valueMin="14.5">14.5–15.1 mm</quantity>
; M3B =
<quantity id="4CDD9B64FF91FFC5FA896260E765FB4D" box="[1280,1407,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="20" pageNumber="21" unit="mm" value="6.25" valueMax="6.4" valueMin="6.1">6.1–6.4 mm</quantity>
) than
<taxonomicName id="4C254D02FF91FFC5FCB06280E184FB6D" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[825,926,1179,1202]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FCB06280E184FB6D" box="[825,926,1179,1202]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
(
<collectionCode id="ED34AE44FF91FFC5FC266281E1C8FB6D" box="[943,978,1178,1202]" country="0" name="Buffalo Bill Museum" pageId="20" pageNumber="21">BB</collectionCode>
=
<quantity id="4CDD9B64FF91FFC5FC676281E66FFB6E" box="[1006,1141,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="1.355" metricValueMax="1.4" metricValueMin="1.31" pageId="20" pageNumber="21" unit="mm" value="13.55" valueMax="14.0" valueMin="13.1">13.1–14 mm</quantity>
; M3B =
<quantity id="4CDD9B64FF91FFC5FB5E6281E759FB6E" box="[1239,1347,1178,1202]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="6.0" metricValueMin="4.7" pageId="20" pageNumber="21" unit="mm" value="5.35" valueMax="6.0" valueMin="4.7">4.7–6 mm</quantity>
). Height of the coronoid process is proportionately shorter than
<taxonomicName id="4C254D02FF91FFC5FAE962A1E7D8FB0E" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1376,1474,1210,1233]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FAE962A1E7D8FB0E" box="[1376,1474,1210,1233]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
(Supporting Information, File S1,
<figureCitation id="131E2A04FF91FFC5FB1A62C2E6E8FB2E" box="[1171,1266,1241,1265]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10E</figureCitation>
). First upper incisor is more procumbent than
<taxonomicName id="4C254D02FF91FFC5FBC962E2E6BAFACF" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1088,1184,1273,1296]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FBC962E2E6BAFACF" box="[1088,1184,1273,1296]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
</taxonomicName>
. Males have small but prominent supraorbital processes, which are absent in
<emphasis id="B951EA93FF91FFC5FAAA6303E792FAF0" box="[1315,1416,1304,1327]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C254D02FF91FFC5FAAA6303E79FFAF0" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1315,1413,1304,1327]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">H. parvus</taxonomicName>
.
</emphasis>
It has a single root in P2 and p3 vs. two non-fused roots in
<taxonomicName id="4C254D02FF91FFC5FAE9632CE677FAB1" authority="(Jbnkins and Robinson 2002)" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF91FFC5FAE9632CE7D8FA91" box="[1376,1474,1335,1358]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
(
<bibRefCitation id="EFB44B70FF91FFC5FCCE634DE67BFAB1" author="Jbnkins PD &amp; Robinson MF" box="[839,1121,1366,1390]" pageId="20" pageNumber="21" pagination="1 - 11" refId="ref32930" refString="Jbnkins PD, Robinson MF. Anothbo vaoiation on thb gvmnuob thbmb: dbscoiption of a nbw spbcibs of Hylomys (Lipotvphla, Eoinacbidab, Galboicinab). Bulletin of Ne Natural History Museum, Zoology Series 2002; 68: 1 - 11." type="journal article" year="2002">Jenkins and Robinson 2002</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF91FFC4FCB06394E072FCA6" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8B9A3681FF91FFC4FCB06394E072FCA6" blockId="20.[825,1475,1423,1980]" lastBlockId="21.[113,764,144,889]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B951EA93FF91FFC5FCB06394E6D4FA78" box="[825,1230,1423,1447]" italics="true" pageId="20" pageNumber="21">Distribution, habitat, and natural history:</emphasis>
Currently known from montane forest from
<quantity id="4CDD9B64FF91FFC5FB8663B4E6AAFA19" box="[1039,1200,1454,1479]" metricMagnitude="3" metricUnit="m" metricValue="2.4539999999999997" metricValueMax="2.835" metricValueMin="2.073" pageId="20" pageNumber="21" unit="m" value="2454.0" valueMax="2835.0" valueMin="2073.0">2073 to 2835 m</quantity>
a.s.l. on Mount Leuser, but it seems to be most abundant at elevations ranging from
<quantity id="4CDD9B64FF91FFC5FA0763D5E1BFF9DA" metricMagnitude="3" metricUnit="m" metricValue="2.2405" metricValueMax="2.408" metricValueMin="2.073" pageId="20" pageNumber="21" unit="m" value="2240.5" valueMax="2408.0" valueMin="2073.0">2073 to 2408 m</quantity>
a.s.l. (
<figureCitation id="131E2A04FF91FFC5FC6163F6E635F9DA" box="[1000,1071,1517,1541]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." figureDoi="http://doi.org/10.5281/zenodo.14510291" httpUri="https://zenodo.org/record/14510291/files/figure.png" pageId="20" pageNumber="21">Fig. 11</figureCitation>
). This habitat is characterized by ‘trees averaging only 15 to 40 feet in height, very hard, knotted and twisted. Everywhere the ground and the branches of the trees were covered with a deep carpet of moss and ferns’ (Ulmer’s field notes,
<bibRefCitation id="EFB44B70FF91FFC5FC266070E632F95C" author="Millbo GS" box="[943,1064,1643,1667]" pageId="20" pageNumber="21" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller 1942</bibRefCitation>
). However, one specimen was recorded at
<quantity id="4CDD9B64FF91FFC5FCDF6091E1B2F97E" box="[854,936,1674,1698]" metricMagnitude="3" metricUnit="m" metricValue="2.835" pageId="20" pageNumber="21" unit="m" value="2835.0">2835 m</quantity>
a.s.l., described as ‘flat heathy plateau mostly bare or covered with grass interspersed with patches of bushes and low trees’ such as ‘rhododendrons,
<taxonomicName id="4C254D02FF91FFC5FB4B60D2E73DF93F" box="[1218,1319,1737,1760]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B951EA93FF91FFC5FB4B60D2E73DF93F" box="[1218,1319,1737,1760]" italics="true" pageId="20" pageNumber="21">Vaccinium</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D02FF91FFC5FAE460D2E7A6F93F" box="[1389,1468,1737,1760]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B951EA93FF91FFC5FAE460D2E7A6F93F" box="[1389,1468,1737,1760]" italics="true" pageId="20" pageNumber="21">Quercus</emphasis>
</taxonomicName>
’ (Ulmer’s field notes, de
<bibRefCitation id="EFB44B70FF91FFC5FBA760F3E749F8DF" author="Schaubnsbb RM &amp; Riplbv SD" box="[1070,1363,1768,1792]" pageId="20" pageNumber="21" pagination="311 - 68" refId="ref35395" refString="db Schaubnsbb RM, Riplbv SD. Zoological obsults of thb Gboogb Vandbobilt Sumatoan bxpbdition, 1936 - 1939 paot I: biods foom Atjbh. Proceedings of the Academy of Natural Sciences of Philadelphia 1939; 91: 311 - 68." type="journal article" year="1939">Schauensee and Ripley 1939</bibRefCitation>
). Its lower elevational limit is unknown, but possibly around
<quantity id="4CDD9B64FF91FFC5FAB0611CE792F8C0" box="[1337,1416,1799,1823]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="m" value="2000.0">2000 m</quantity>
since there is a noticeable change from moss forest to foothill jungle according to
<bibRefCitation id="EFB44B70FF91FFC5FC49615DE64AF881" author="Millbo GS" box="[960,1104,1862,1886]" pageId="20" pageNumber="21" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller (1942)</bibRefCitation>
. Furthermore, the lower elevational limits of
<taxonomicName id="4C254D02FF91FFC5FC1F617DE1F4F8A2" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[918,1006,1894,1917]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF91FFC5FC1F617DE1F4F8A2" box="[918,1006,1894,1917]" italics="true" pageId="20" pageNumber="21">H. vorax</emphasis>
</taxonomicName>
might be constrained by
<taxonomicName id="4C254D02FF91FFC5FB73617DE757F8A2" authorityName="Sody" authorityYear="1933" box="[1274,1357,1894,1917]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FB73617DE757F8A2" box="[1274,1357,1894,1917]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
, which has been recorded between less than 100 and
<quantity id="4CDD9B64FF91FFC5FB6E619EE72CF843" box="[1255,1334,1925,1949]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="m" value="1100.0">1100 m</quantity>
a.s.l. in Aceh, and up to
<quantity id="4CDD9B64FF91FFC5FC2B61BFE1EBF864" box="[930,1009,1956,1980]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="21" unit="m" value="2200.0">2200 m</quantity>
a.s.l. on Mt. Kerinci. We suggest
<taxonomicName id="4C254D02FF91FFC5FACC61BEE78CF863" authorityName="Sody" authorityYear="1933" box="[1349,1430,1957,1980]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF91FFC5FACC61BEE78CF863" box="[1349,1430,1957,1980]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF90FFC4FFF8668AE2D3FF77" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[113,201,145,168]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FFF8668AE2D3FF77" box="[113,201,145,168]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
have a parapatric distribution in the Gayo Highlands mirroring the elevational segregation of
<taxonomicName id="4C254D02FF90FFC4FD9866ABE079FF18" authorityName="Sody" authorityYear="1933" box="[529,611,176,199]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FD9866ABE079FF18" box="[529,611,176,199]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF90FFC4FD1166ABE0E0FF18" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[664,762,176,199]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF90FFC4FD1166ABE0E0FF18" box="[664,762,176,199]" italics="true" pageId="21" pageNumber="22">H. parvus</emphasis>
</taxonomicName>
on Mt. Kerinci (
<bibRefCitation id="EFB44B70FF90FFC4FE9E66D4E3DBFF38" author="Rubdi M &amp; Chapuisat M &amp; Iskandao D" box="[279,449,207,231]" pageId="21" pageNumber="22" pagination="965 - 78" refId="ref35216" refString="Rubdi M, Chapuisat M, Iskandao D. Taxonomic status of Hylomys parous and Hylomys suillus (Insbctivooa: Eoinacbidab): biochbmical and moophological analvsbs. Journal of Mammalogy 1994; 75: 965 - 78. hups: // doi. oog / 10.2307 / 1382478" type="journal article" year="1994">
Ruedi
<emphasis id="B951EA93FF90FFC4FED366D4E393FF38" box="[346,393,207,231]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
1994
</bibRefCitation>
). Additional sampling at Gayo Highlands between 1100 and
<quantity id="4CDD9B64FF90FFC4FE2566F5E3E7FEDA" box="[428,509,238,262]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="m" value="2000.0">2000 m</quantity>
a.s.l. will be required to confirm this hypothesis.
<taxonomicName id="4C254D02FF90FFC4FEFF6716E011FEFA" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[374,523,269,293]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FEFF6716E011FEFA" box="[374,523,269,293]" italics="true" pageId="21" pageNumber="22">Hylomys vorax</emphasis>
</taxonomicName>
seems to show preference for moist microhabitats with high cover, since six over nine specimens with associated habitat field notes were caught in a mossy area next to a river. These specimens were caught either under mossy logs (
<emphasis id="B951EA93FF90FFC4FEC96790E349FE7D" box="[320,339,395,418]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 4), under tree roots in thick scrubby jungle (
<emphasis id="B951EA93FF90FFC4FF4967B0E2C9FE1D" box="[192,211,427,450]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 2), in thick growth near blang (=meadow) (
<emphasis id="B951EA93FF90FFC4FD2467B0E0DAFE1D" box="[685,704,427,450]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 1), mossy hole at base of tree (
<emphasis id="B951EA93FF90FFC4FE0567D1E385FE3E" box="[396,415,458,481]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 1), or in grass in an open glade near forest (
<emphasis id="B951EA93FF90FFC4FF6667F2E318FDDF" box="[239,258,489,512]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 1). Stomachs contained the chewed remains of earthworms (
<emphasis id="B951EA93FF90FFC4FF756412E315FDFF" box="[252,271,521,544]" italics="true" pageId="21" pageNumber="22">N</emphasis>
= 2), and specimens were caught with ham rind or coconut bait (Ulmer’s field notes). A subadult male and two females were caught under the same log. The most likely scenario is that a mature female with teats (
<collectionCode id="ED34AE44FF90FFC4FD88647DE04AFDA1" box="[513,592,614,638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
271033) and its offspring (subadults
<collectionCode id="ED34AE44FF90FFC4FEC8649EE398FD42" box="[321,386,645,669]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="21" pageNumber="22">ANSP</collectionCode>
20375 and 20378) were collected together, perhaps suggesting parental care and/or territory overlap with offspring at least until adulthood. Three adult specimens had a very strong porcine odour, but this was not mentioned for any of the subadults in Fred Ulmer’s field notes. It shares habitat with
<taxonomicName id="4C254D02FF90FFC4FF2E6538E0B7FCE5" authority="(Robinson and Kloss 1916)" baseAuthorityName="Robinson and Kloss" baseAuthorityYear="1916" box="[167,685,802,826]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="hylomyoides">
<emphasis id="B951EA93FF90FFC4FF2E6538E39EFCE5" box="[167,388,802,826]" italics="true" pageId="21" pageNumber="22">Maxomys hylomyoides</emphasis>
(
<bibRefCitation id="EFB44B70FF90FFC4FE1F6539E0B8FCE5" author="Robinson HC &amp; Kloss CB" box="[406,674,802,826]" pageId="21" pageNumber="22" pagination="269 - 78" refId="ref34992" refString="Robinson HC, Kloss CB. Pobliminaov diagnosbs of somb nbw spbcibs and subspbcibs of mammals and biods obtainbd in Kooinchi, Wbst Sumatoa, Fbb - Junb 1914. Journal of the Straits Branch of the Royal Asiatic Society 1916; 73: 269 - 78." type="journal article" year="1916">Robinson and Kloss 1916</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C254D02FF90FFC4FD336538E386FC86" authority="(Chasbn 1939)" baseAuthorityName="Chasbn" baseAuthorityYear="1939" genus="Ratus" pageId="21" pageNumber="22" rank="species" species="hoogerwerfi">
<emphasis id="B951EA93FF90FFC4FD336538E2F8FC86" italics="true" pageId="21" pageNumber="22">Rattus hoogerwerfi</emphasis>
(Chasen 1939)
</taxonomicName>
,
<taxonomicName id="4C254D02FF90FFC4FE38655AE2A8FCA6" authority="(Nomas 1888)" baseAuthorityName="Nomas" baseAuthorityYear="1888" class="Mammalia" family="Muridae" genus="Sundamys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="inssaluteus">
<emphasis id="B951EA93FF90FFC4FE38655AE096FC86" box="[433,652,833,857]" italics="true" pageId="21" pageNumber="22">Sundamys infraluteus</emphasis>
(Thomas 1888)
</taxonomicName>
, and
<taxonomicName id="4C254D02FF90FFC4FF6E657AE07EFCA6" authority="(Millbo 1942)" baseAuthorityName="Millbo" baseAuthorityYear="1942" box="[231,612,865,889]" class="Mammalia" family="Sciuridae" genus="Sundasciurus" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="altitudinis">
<emphasis id="B951EA93FF90FFC4FF6E657AE3D5FCA6" box="[231,463,865,889]" italics="true" pageId="21" pageNumber="22">Sundasciurus altitudinis</emphasis>
(
<bibRefCitation id="EFB44B70FF90FFC4FE56657AE042FCA6" author="Millbo GS" box="[479,600,865,889]" pageId="21" pageNumber="22" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller 1942</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF90FFC4FFF86581E037F8E1" pageId="21" pageNumber="22" type="conservation">
<paragraph id="8B9A3681FF90FFC4FFF86581E037F8E1" blockId="21.[113,763,922,1854]" pageId="21" pageNumber="22">
<emphasis id="B951EA93FF90FFC4FFF86581E2E3FC6E" box="[113,249,922,945]" italics="true" pageId="21" pageNumber="22">Conservation:</emphasis>
This species is found in Gunung Leuser National Park (
<collectionCode id="ED34AE44FF90FFC4FF2565A2E2E3FC0E" box="[172,249,953,977]" country="USA" httpUri="http://grbio.org/cool/seqy-90es" name="Glacier National Park" pageId="21" pageNumber="22">GLNP</collectionCode>
). The Leuser ecosystem is designated as a ‘national strategic area’ (
<collectionCode id="ED34AE44FF90FFC4FE9E65C2E34AFC2E" box="[279,336,985,1009]" pageId="21" pageNumber="22">NSA</collectionCode>
) in Indonesia, to safeguard ecosystem services and limit infrastructure and agricultural expansion for ecological preservation. However, recent actions by the Aceh provincial government, which omitted Leuser’s
<collectionCode id="ED34AE44FF90FFC4FDD2622CE097FB90" box="[603,653,1079,1103]" pageId="21" pageNumber="22">NSA</collectionCode>
status and proposed conflicting development projects, have created uncertainty about its conservation status and the effectiveness of decentralized forest governance in Indonesia (
<bibRefCitation id="EFB44B70FF90FFC4FDB5628FE0F3FB72" author="Sloan S &amp; Campbbll MJ &amp; Alamgio M" box="[572,745,1172,1197]" pageId="21" pageNumber="22" pagination="298 - 309" refId="ref35611" refString="Sloan S, Campbbll MJ, Alamgio M et al. Infoastouctuob dbvblopmbnt and contbstbd foobst govbonancb thobatbn thb Lbusbo Ecosvstbm, Indonbsia. Land Use Policy 2018; 77: 298 - 309. hups: // doi. oog / 10.1016 / j. landusbpol. 2018.05.043" type="journal article" year="2018">
Sloan
<emphasis id="B951EA93FF90FFC4FDF4628EE0B4FB73" box="[637,686,1172,1196]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2018
</bibRefCitation>
).
<collectionCode id="ED34AE44FF90FFC4FFF862AFE2ACFB13" box="[113,182,1204,1228]" country="USA" httpUri="http://grbio.org/cool/seqy-90es" name="Glacier National Park" pageId="21" pageNumber="22">GLNP</collectionCode>
has lost
<emphasis id="B951EA93FF90FFC4FE8362AEE303FB13" box="[266,281,1205,1228]" italics="true" pageId="21" pageNumber="22">c.</emphasis>
8000 ha of its primary forest from 2012 to 2017, possibly due to a combination of industrial oil palm plantations, dryland agriculture, and forest fires (
<bibRefCitation id="EFB44B70FF90FFC4FE6A62E9E093FAD4" author="Lubis MI &amp; Puspaoini W &amp; Poabowo SA &amp; Maothv W &amp; Taomizi &amp; Andavani N &amp; Linkib M" box="[483,649,1266,1291]" pageId="21" pageNumber="22" pagination="741 - 9" refId="ref33633" refString="Lubis MI, Puspaoini W, Poabowo SA, Maothv W, Taomizi, Andavani N, Linkib M. Unoavbling thb complbxitv of human - tigbo conflicts in thb Lbusbo Ecosvstbm, Sumatoa. Animal Conseroation 2020; 23: 741 - 9." type="journal article" year="2020">
Lubis
<emphasis id="B951EA93FF90FFC4FDAB62E8E04BFAD5" box="[546,593,1266,1290]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2020
</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF90FFC4FD1D62E9E2D9FAF5" author="Condoo AA &amp; Poasbtvo LB &amp; Rushavati SB" pageId="21" pageNumber="22" refId="ref31536" refString="Condoo AA, Poasbtvo LB, Rushavati SB et al. Mbasuoing mbtoics of climatb changb and its implication on thb bndangbobd mammal consbovation in thb Lbusbo Ecosvstbm. Frontiers in Enoironmental Science 2021; 9. hups: // doi. oog / 10.3389 / fbnvs. 2021.713837" type="journal volume" year="2021">
Condro
<emphasis id="B951EA93FF90FFC4FD6062E8E290FAF5" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2021
</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF90FFC4FF476309E3AFFAF5" author="Dwivahobni A &amp; Fuad HA &amp; Sunaovo S" box="[206,437,1298,1322]" pageId="21" pageNumber="22" pagination="1235 - 1242" refId="ref31806" refString="Dwivahobni A, Fuad HA, Sunaovo S et al. Foobst covbo changbs in Indonbsia's tboobstoial national paoks bbtwbbn 2012 and 2017. Biodioersitas Journal of Biological Dioersity 2021; 22: 1235 - 1242. hups: // doi. oog / 10.13057 / biodiv / d 220320" type="journal article" year="2021">
Dwiyahreni
<emphasis id="B951EA93FF90FFC4FEC46308E366FAF5" box="[333,380,1298,1322]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2021
</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF90FFC4FE486309E06BFAF5" author="Noha AS &amp; Saoaswita N &amp; Sulistivono N" box="[449,625,1298,1322]" pageId="21" pageNumber="22" pagination="1420 - 6" refId="ref35840" refString="Noha AS, Saoaswita N, Sulistivono N et al. Analvsis of land covbo changbs dub to foobst fiobs in Gunung Lbusbo National Paok, Nooth Sumatoa Poovincb, Indonbsia. Biodioersitas Journal of Biological Dioersity 2022; 23: 1420 - 6." type="journal article" year="2022">
Thoha
<emphasis id="B951EA93FF90FFC4FD816308E022FAF5" box="[520,568,1298,1322]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2022
</bibRefCitation>
). The Leuser Ecosystem is also threatened by habitat fragmentation due to unofficial road development and conversion of ‘sustainable managed’ buffer forests to palm oil plantations (
<bibRefCitation id="EFB44B70FF90FFC4FDB5636BE0F3FA57" author="Sloan S &amp; Campbbll MJ &amp; Alamgio M" box="[572,745,1392,1416]" pageId="21" pageNumber="22" pagination="298 - 309" refId="ref35611" refString="Sloan S, Campbbll MJ, Alamgio M et al. Infoastouctuob dbvblopmbnt and contbstbd foobst govbonancb thobatbn thb Lbusbo Ecosvstbm, Indonbsia. Land Use Policy 2018; 77: 298 - 309. hups: // doi. oog / 10.1016 / j. landusbpol. 2018.05.043" type="journal article" year="2018">
Sloan
<emphasis id="B951EA93FF90FFC4FDF4636AE0B5FA57" box="[637,687,1392,1416]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2018
</bibRefCitation>
). While most deforestation is taking place in lowland areas, it is suggested that the magnitude of climate change effects will be higher on the montane areas of the
<collectionCode id="ED34AE44FF90FFC4FE5163D5E007FA39" box="[472,541,1486,1510]" country="USA" httpUri="http://grbio.org/cool/seqy-90es" name="Glacier National Park" pageId="21" pageNumber="22">GLNP</collectionCode>
(
<bibRefCitation id="EFB44B70FF90FFC4FDA463D5E0F3FA39" author="Condoo AA &amp; Poasbtvo LB &amp; Rushavati SB" box="[557,745,1486,1510]" pageId="21" pageNumber="22" refId="ref31536" refString="Condoo AA, Poasbtvo LB, Rushavati SB et al. Mbasuoing mbtoics of climatb changb and its implication on thb bndangbobd mammal consbovation in thb Lbusbo Ecosvstbm. Frontiers in Enoironmental Science 2021; 9. hups: // doi. oog / 10.3389 / fbnvs. 2021.713837" type="journal volume" year="2021">
Condro
<emphasis id="B951EA93FF90FFC4FD0A63D5E0ABFA39" box="[643,689,1486,1510]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2021
</bibRefCitation>
). Finally, it is anticipated that the Trans-Sumatra Highway’s completion, if constructed according to the current plan, would encourage increased anthropogenic pressure on the outskirts of the
<collectionCode id="ED34AE44FF90FFC4FF146050E2FBF9BC" box="[157,225,1611,1635]" country="USA" httpUri="http://grbio.org/cool/seqy-90es" name="Glacier National Park" pageId="21" pageNumber="22">GLNP</collectionCode>
, potentially impacting its fragile ecosystem (
<bibRefCitation id="EFB44B70FF90FFC4FD486050E2FBF95D" author="Sloan S &amp; Alamgio M &amp; Campbbll MJ" pageId="21" pageNumber="22" refId="ref35660" refString="Sloan S, Alamgio M, Campbbll MJ et al. Dbvblopmbnt coooidoos and obmnant-foobst consbovation in Sumatoa, Indonbsia. Tropical Conseroation Science 2019; 12: 194008291988950. hups: // doi. oog / 10.1177 / 1940082919889509" type="journal volume" year="2019">
Sloan
<emphasis id="B951EA93FF90FFC4FFF86070E2BEF95D" box="[113,164,1642,1666]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2019
</bibRefCitation>
). Species co-distributed with
<taxonomicName id="4C254D02FF90FFC4FDAA6070E067F95D" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[547,637,1643,1666]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FDAA6070E067F95D" box="[547,637,1643,1666]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
, like
<taxonomicName id="4C254D02FF90FFC4FD336070E2F8F97D" baseAuthorityName="Chasbn" baseAuthorityYear="1939" genus="Ratus" pageId="21" pageNumber="22" rank="species" species="hoogerwerfi">
<emphasis id="B951EA93FF90FFC4FD336070E2F8F97D" italics="true" pageId="21" pageNumber="22">Rattus hoogerwerfi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF90FFC4FE906091E3DFF97D" baseAuthorityName="Collbu" baseAuthorityYear="1893" box="[281,453,1674,1698]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="thomasi">
<emphasis id="B951EA93FF90FFC4FE906091E3DFF97D" box="[281,453,1674,1698]" italics="true" pageId="21" pageNumber="22">Presbytis thomasi</emphasis>
</taxonomicName>
, have similar or larger occurrence ranges and are listed as vulnerable in the
<collectionCode id="ED34AE44FF90FFC4FDD060B2E081F91E" box="[601,667,1705,1729]" pageId="21" pageNumber="22">IUCN</collectionCode>
Red List of Threatened Species. Further research is required to assess
<taxonomicName id="4C254D02FF90FFC4FD6860D2E2B2F920" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FD6860D2E2B2F920" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
population trends, explore its distribution beyond Mt. Leuser, and investigate whether
<taxonomicName id="4C254D02FF90FFC4FE486113E00EF8C0" authorityName="Sody" authorityYear="1933" box="[449,532,1800,1823]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FE486113E00EF8C0" box="[449,532,1800,1823]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
is displacing
<taxonomicName id="4C254D02FF90FFC4FD2B6113E0E0F8C0" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[674,762,1800,1823]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FD2B6113E0E0F8C0" box="[674,762,1800,1823]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
due to climate-induced elevational changes.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF90FFC4FFF8617BE76CFD80" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8B9A3681FF90FFC4FFF8617BE76CFD80" blockId="21.[113,763,1887,1974]" lastBlockId="21.[810,1460,143,607]" pageId="21" pageNumber="22">
<emphasis id="B951EA93FF90FFC4FFF8617BE2FAF8A8" box="[113,224,1888,1911]" italics="true" pageId="21" pageNumber="22">Comments:</emphasis>
Five specimens (
<collectionCode id="ED34AE44FF90FFC4FE21617BE3F9F8A7" box="[424,483,1888,1912]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="21" pageNumber="22" type="Museum">MZB</collectionCode>
3172–5,
<collectionCode id="ED34AE44FF90FFC4FDC5617BE086F8A8" box="[588,668,1888,1911]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
5137-8) with damaged skulls collected in Pajatoeng Kalang (south of Isaq) at
<quantity id="4CDD9B64FF90FFC4FF4B6185E30AF869" box="[194,272,1950,1974]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="m" value="2000.0">2000 m</quantity>
were clustered in the morphospace of
<taxonomicName id="4C254D02FF90FFC4FD076184E0FBF869" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[654,737,1951,1974]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FD076184E0FBF869" box="[654,737,1951,1974]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
in some preliminary uni/bivariate analyses (
<collectionCode id="ED34AE44FF90FFC4FB59668BE711FF77" box="[1232,1291,144,168]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="21" pageNumber="22" type="Museum">MZB</collectionCode>
3172–5: HB/E,
<collectionCode id="ED34AE44FF90FFC4FCA366B4E146FF18" box="[810,860,175,199]" pageId="21" pageNumber="22">GLS</collectionCode>
/
<collectionCode id="ED34AE44FF90FFC4FCEC66B4E19CFF18" box="[869,902,175,199]" country="India" name="Indian Museum" pageId="21" pageNumber="22">IM</collectionCode>
3Sb, M1M1,
<collectionCode id="ED34AE44FF90FFC4FB8066B4E637FF18" box="[1033,1069,175,199]" country="India" name="Indian Museum" pageId="21" pageNumber="22">IM</collectionCode>
3I;
<collectionCode id="ED34AE44FF90FFC4FBC566B4E686FF19" box="[1100,1180,175,198]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
5137-8: HB/E, P4M3) but in between
<taxonomicName id="4C254D02FF90FFC4FC2A66D4E1EFFF39" authorityName="Sody" authorityYear="1933" box="[931,1013,207,230]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FC2A66D4E1EFFF39" box="[931,1013,207,230]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF90FFC4FBA066D4E665FF39" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1065,1151,207,230]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FBA066D4E665FF39" box="[1065,1151,207,230]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
(
<collectionCode id="ED34AE44FF90FFC4FB0766D5E6D3FF39" box="[1166,1225,206,230]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="21" pageNumber="22" type="Museum">MZB</collectionCode>
3172–5: HB/
<collectionCode id="ED34AE44FF90FFC4FAD266D5E776FF39" box="[1371,1388,206,230]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="22">T</collectionCode>
,
<collectionCode id="ED34AE44FF90FFC4FAFE66D5E7B0FF39" box="[1399,1450,206,230]" pageId="21" pageNumber="22">GLS</collectionCode>
/
<collectionCode id="ED34AE44FF90FFC4FCA366F5E157FEDA" box="[810,845,238,261]" country="0" name="Buffalo Bill Museum" pageId="21" pageNumber="22">BB</collectionCode>
;
<collectionCode id="ED34AE44FF90FFC4FCDF66F5E1BCFEDA" box="[854,934,238,261]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
5137-8: M1M1,
<collectionCode id="ED34AE44FF90FFC4FBDA66F5E66DFED9" box="[1107,1143,238,262]" country="India" name="Indian Museum" pageId="21" pageNumber="22">IM</collectionCode>
3I) or in the range of
<taxonomicName id="4C254D02FF90FFC4FAC366F5E780FEDA" authorityName="Sody" authorityYear="1933" box="[1354,1434,238,261]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FAC366F5E780FEDA" box="[1354,1434,238,261]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
in others (
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5137:
<collectionCode id="ED34AE44FF90FFC4FB996716E63AFEFB" box="[1040,1056,269,292]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="22">T</collectionCode>
,
<collectionCode id="ED34AE44FF90FFC4FBA06716E641FEFA" box="[1065,1115,269,293]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12552" name="Agricultural Development and Advisory Service" pageId="21" pageNumber="22" type="Herbarium">PPL</collectionCode>
). Specimens
<collectionCode id="ED34AE44FF90FFC4FB546716E737FEFB" box="[1245,1325,269,292]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
5137-8 skins and skulls were examined through pictures and have a bicoloured tail (
<taxonomicName id="4C254D02FF90FFC4FCD36757E1B3FEBC" authorityName="Sody" authorityYear="1933" box="[858,937,332,355]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FCD36757E1B3FEBC" box="[858,937,332,355]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
feature), but most cranial or mandible features were consistent with
<taxonomicName id="4C254D02FF90FFC4FC426777E63AFE5C" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[971,1056,364,387]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FC426777E63AFE5C" box="[971,1056,364,387]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
. These were excluded from
<tableCitation id="C6A7033AFF90FFC4FAB56770E79DFE5C" box="[1340,1415,363,387]" captionStart="Table 1" captionStartId="11.[113,168,143,167]" captionTargetPageId="11" captionText="Table 1. Craniodental and selected external measurements (mm), weight (g) of adult Hylomys, including new species from Mount Leuser and the Da Lat–Dak Lak Plateaus. Mean, SD (w), ranges, and sample size (N) are specified for each measurement." httpUri="http://table.plazi.org/id/DF5A6609FF8EFFDAFFF86694E662FF1C" pageId="21" pageNumber="22" tableUuid="DF5A6609FF8EFFDAFFF86694E662FF1C">Table 1</tableCitation>
and all bivariate plot figures since they were collected in a habitat transition and potentially parapatric (and perhaps even hybridization) area of
<taxonomicName id="4C254D02FF90FFC4FC4467D1E605FE3E" authorityName="Sody" authorityYear="1933" box="[973,1055,458,481]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF90FFC4FC4467D1E605FE3E" box="[973,1055,458,481]" italics="true" pageId="21" pageNumber="22">H. maxi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF90FFC4FBDD67D1E6B1FE3E" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[1108,1195,458,481]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF90FFC4FBDD67D1E6B1FE3E" box="[1108,1195,458,481]" italics="true" pageId="21" pageNumber="22">H. vorax</emphasis>
</taxonomicName>
, and were not sequenced nor included in the craniodental PCA due to their damaged state. Lastly, diagnostic measurements of
<collectionCode id="ED34AE44FF90FFC4FB076413E6C4FDC0" box="[1166,1246,520,543]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
5137-8 were not collected by the authors, but by Pepijn Kamminga (senior collection manager at
<collectionCode id="ED34AE44FF90FFC4FC14645CE1EBFD80" box="[925,1009,583,607]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
) and could be subject to observer bias.
</paragraph>
</subSubSection>
</treatment>
</document>
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