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<document id="AC099248255217E355029779C05586F5" ID-CLB-Dataset="306597" ID-DOI="10.1093/zoolinnean/zlad177" ID-GBIF-Dataset="dcaf5aa0-c7c6-4dc1-871a-8815b1aabb88" ID-ISSN="0024-4082" ID-Zenodo-Dep="14510261" ID-ZooBank="80D1924-E984-4900-88E5-85FE2C5688D8" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1734027918295" checkinUser="plazi" docAuthor="Hinckley, Arlo, Camacho-Sanchez, Miguel, Chua, Marcus A. H., Ruedi, Manuel, Lunde, Darrin, Maldonado, Jesús E., Omar, Hasmahzaiti, Leonard, Jennifer A. &amp; Hawkins, Melissa T. R." docDate="2024" docId="038C8797FF88FFDFFF706694E08DFA90" docLanguage="en" docName="zlad177.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlad177" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Hylomys maxi Sody 1933" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="FFB5FFEFFF85FFD1FF89661BE21AFFDF" masterDocTitle="An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="14" updateTime="1734460109358" updateUser="ExternalLinkService">
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<mods:title id="BFB430204BDDD2A41EAA03141F56E88D">An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia</mods:title>
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<mods:namePart id="0C87931C1894754C6410CDE658CF8610">Hinckley, Arlo</mods:namePart>
<mods:affiliation id="5A489DF931EE2FD73AC86CA3D2AA7826">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
<mods:nameIdentifier id="6344E6B62C8C62431C5807B42982AC93" type="email">hinckleya@si.edu</mods:nameIdentifier>
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<mods:namePart id="E47844FE7FACBC95B4AC7713E453732B">Camacho-Sanchez, Miguel</mods:namePart>
<mods:affiliation id="92BC42D40EE0249FA522AECA6493A5F5">Conservation &amp; Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<mods:namePart id="0BB00719A23DB271D08C6B0DBD62C5B4">Chua, Marcus A. H.</mods:namePart>
<mods:affiliation id="7FE91D5FC1E1082F7B6521C9B2643A07">Smithsonian National Zoo and Conservation Biology Institute, Center for Conservation Genomics, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="DAD2BC5308A955A79E4B5C9881015287">Ruedi, Manuel</mods:namePart>
<mods:affiliation id="B130EFE56AC8B4E8A9E658739F681899">Natural History Museum of Geneva, Geneva, Switzerland</mods:affiliation>
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<mods:namePart id="B7C4D37F6BE3E88B8B1F8D81662942CB">Lunde, Darrin</mods:namePart>
<mods:affiliation id="186D86F05622B163B37D9934D2FAD2C8">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="7A39ED4D7C378C7EDE6380D8541DEE55">Maldonado, Jesús E.</mods:namePart>
<mods:affiliation id="1A7F5A18067B272C478B373713041A84">Smithsonian National Zoo and Conservation Biology Institute, Center for Conservation Genomics, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="C1D127B7D60C47E88C6EA1A5521A9DC7">Omar, Hasmahzaiti</mods:namePart>
<mods:affiliation id="1628A15BDF2C95834A9BA412AAC59201">Museum of Zoology, Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:namePart id="8A157695088BD0DE7CA6A394646DED3A">Leonard, Jennifer A.</mods:namePart>
<mods:affiliation id="F0199AAB2F25B2620BA5363A1C3455F1">Conservation &amp; Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<mods:namePart id="2F1E2558670FAD24983537D9036F5B62">Hawkins, Melissa T. R.</mods:namePart>
<mods:affiliation id="A619D9FED5C892DF7E8DA0979DC19ACA">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:title id="B1EAA529F46CB1B81702EB98131F5E5E">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="CFB12D64DA3A95353CCF7AE4A85AEA7B">
<mods:date id="74C903A99D18D1F461EFA119082B5BE5">2024</mods:date>
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<mods:number id="3EC595D413B77005DD8A65B5C401CD53">202</mods:number>
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<mods:url id="42F531696A74FFEE28E6CAD7A3B3FC90">https://doi.org/10.1093/zoolinnean/zlad177</mods:url>
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<mods:classification id="2E954C514B0D54514FB4B28B7DBD2ED1">journal article</mods:classification>
<mods:identifier id="9FE4A3D2DBF982C3E38D58504545AC2C" type="CLB-Dataset">306597</mods:identifier>
<mods:identifier id="4F2F95F9798243B6762FE15D593B15C3" type="DOI">10.1093/zoolinnean/zlad177</mods:identifier>
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<treatment id="038C8797FF88FFDFFF706694E08DFA90" ID-DOI="http://doi.org/10.5281/zenodo.14510271" ID-Zenodo-Dep="14510271" LSID="urn:lsid:plazi:treatment:038C8797FF88FFDFFF706694E08DFA90" httpUri="http://treatment.plazi.org/id/038C8797FF88FFDFFF706694E08DFA90" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C33F650AFF88FFDCFF706694E068FF75" box="[249,626,143,170]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B9A3681FF88FFDCFF706694E068FF75" blockId="13.[113,713,143,269]" box="[249,626,143,170]" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFF706694E068FF75" bold="true" box="[249,626,143,170]" pageId="13" pageNumber="14">
<taxonomicName id="4C254D02FF88FFDCFF706694E3C9FF75" authority="Sody" authorityName="Sody" authorityYear="1933" box="[249,467,143,170]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF88FFDCFF706694E383FF75" bold="true" box="[249,409,143,170]" italics="true" pageId="13" pageNumber="14">Hylomys maxi</emphasis>
Sody
</taxonomicName>
, 19&amp;&amp; stat. nov.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF88FFDCFFF866ADE04FFED2" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8B9A3681FF88FFDCFFF866ADE0D3FF11" blockId="13.[113,713,143,269]" box="[113,713,182,206]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB3511AFFF88FFDCFFF866ADE0D3FF11" box="[113,713,182,206]" pageId="13" pageNumber="14">
<treatmentCitation id="0A841090FF88FFDCFFF866ADE3F1FF11" author="Sodv HJV" box="[113,491,182,206]" page="438" pageId="13" pageNumber="14" year="1933">
<taxonomicName id="4C254D02FF88FFDCFFF866ADE3F1FF11" ID-CoL="5J2D5" authority="Sodv, 1933: 438" authorityName="Sodv" authorityPageNumber="438" authorityYear="1933" box="[113,491,182,206]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="maxi">
<emphasis id="B951EA93FF88FFDCFFF866ADE325FF11" box="[113,319,182,206]" italics="true" pageId="13" pageNumber="14">Hylomys suillus maxi</emphasis>
<bibRefCitation id="EFB44B70FF88FFDCFECD66ADE3F1FF11" author="Sodv HJV" box="[324,491,182,206]" pageId="13" pageNumber="14" pagination="430 - 42" refId="ref35700" refString="Sodv HJV. XLI - Tbn nbw mammals foom thb Dutch East Indibs. Annals and Magazine of Natural History 1933; 12: 430 - 42. hups: // doi. oog / 10.1080 / 00222933308673706" type="journal article" year="1933">Sody, 1933: 438</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original description.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A3681FF88FFDCFFF866CDE3E5FF31" blockId="13.[113,713,143,269]" box="[113,511,214,238]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB3511AFFF88FFDCFFF866CDE3E5FF31" box="[113,511,214,238]" pageId="13" pageNumber="14">
<treatmentCitation id="0A841090FF88FFDCFFF866CDE3E1FF31" author="Chasbn FN" box="[113,507,214,238]" page="12" pageId="13" pageNumber="14" year="1940">
<taxonomicName id="4C254D02FF88FFDCFFF866CDE3E1FF31" ID-CoL="7K7J7" authority="Chasbn, 1940: 12" authorityName="Chasbn" authorityPageNumber="12" authorityYear="1940" box="[113,507,214,238]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="tionis">
<emphasis id="B951EA93FF88FFDCFFF866CDE35BFF32" box="[113,321,214,238]" italics="true" pageId="13" pageNumber="14">Hylomys suillus tionis</emphasis>
<bibRefCitation id="EFB44B70FF88FFDCFECE66CDE3E1FF31" author="Chasbn FN" box="[327,507,214,238]" pageId="13" pageNumber="14" pagination="1 - 209" refId="ref31447" refString="Chasbn FN. A handlist of Malavsian mammals (a svstbmatic list of thb mammals of thb Malav Pbninsula, Sumatoa, Boonbo and Java, including thb adjacbnt small islands). Bulletin of the Raffles Museum 1940; 15: 1 - 209." type="journal article" year="1940">Chasen, 1940: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A3681FF88FFDCFFF866EEE04FFED2" blockId="13.[113,713,143,269]" box="[113,597,245,269]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB3511AFFF88FFDCFFF866EEE04FFED2" box="[113,597,245,269]" pageId="13" pageNumber="14">
<treatmentCitation id="0A841090FF88FFDCFFF866EEE010FED2" author="Coobbt GB" box="[113,522,245,269]" page="122" pageId="13" pageNumber="14" year="1988">
<taxonomicName id="4C254D02FF88FFDCFFF866EEE010FED2" ID-CoL="5J2D9" authority="Coobbt, 1988: 122" authorityName="Coobbt" authorityPageNumber="122" authorityYear="1988" box="[113,522,245,269]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="suillus">
<emphasis id="B951EA93FF88FFDCFFF866EEE353FED2" box="[113,329,245,269]" italics="true" pageId="13" pageNumber="14">Hylomys suillus suillus</emphasis>
<bibRefCitation id="EFB44B70FF88FFDCFEC766EEE010FED2" author="Coobbt GB" box="[334,522,245,269]" pageId="13" pageNumber="14" pagination="117 - 72" refId="ref31585" refString="Coobbt GB. Nb familv Eoinacbidab: a svnthbsis of its taxonomv, phvlogbnv, bcologv and zoogbogoaphv. Mammal Reoiew 1988; 18: 117 - 72. hups: // doi. oog / 10.1111 / j. 1365 - 2907.1988. tb 00082. x" type="journal article" year="1988">Corbet, 1988: 122</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF88FFDCFFF86735E2E6FDA4" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8B9A3681FF88FFDCFFF86735E2E5FEB9" blockId="13.[113,762,302,358]" pageId="13" pageNumber="14">
<materialsCitation id="3B4D3CDCFF88FFDCFFF86735E2E5FEB9" collectingDate="1930-01-04" collectionCode="RMNH" collectorName="Sody" country="Indonesia" location="Giesting" municipality="Lampung" pageId="13" pageNumber="14" specimenCode="RMNH 23895" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
<emphasis id="B951EA93FF88FFDCFFF86735E2CBFE99" box="[113,209,302,326]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="549E8823FF88FFDCFFF86735E2D6FE99" box="[113,204,302,326]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DB839EFAFF88FFDCFF576734E362FE99" box="[222,376,302,326]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH 23895</specimenCode>
, an
<specimenCount id="9D23FD08FF88FFDCFE296735E011FE99" box="[416,523,302,326]" pageId="13" pageNumber="14" type="adult">adult male</specimenCount>
collected by
<collectorName id="26D05357FF88FFDCFD1B6735E0DCFE99" box="[658,710,302,326]" pageId="13" pageNumber="14">Sody</collectorName>
on
<date id="FF9B1041FF88FFDCFD646735E2E1FEB9" pageId="13" pageNumber="14" value="1930-01-04">
<collectingDate id="EFDFE9A9FF88FFDCFD646735E2E1FEB9" pageId="13" pageNumber="14" value="1930-01-04">4 January 1930</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9A3681FF88FFDCFFF86795E0F8FE1B" blockId="13.[113,763,397,453]" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFFF86795E2E9FE7A" box="[113,243,397,421]" italics="true" pageId="13" pageNumber="14">Type locality:</emphasis>
‘Giesting, Lampoengs, S. Sumatra’ (=
<location id="8EFA605AFF88FFDCFDE66796E0D7FE7A" LSID="urn:lsid:plazi:treatment:038C8797FF88FFDFFF706694E08DFA90:8EFA605AFF88FFDCFDE66796E0D7FE7A" box="[623,717,397,421]" country="Indonesia" municipality="Lampung" name="Giesting" pageId="13" pageNumber="14" stateProvince="Sumatra">Giesting</location>
,
<location id="8EFA605AFF88FFDCFD5F6796E2F2FE1A" LSID="urn:lsid:plazi:treatment:038C8797FF88FFDFFF706694E08DFA90:8EFA605AFF88FFDCFD5F6796E2F2FE1A" country="Indonesia" municipality="Lampung" name="Mt. Tanggamus" pageId="13" pageNumber="14" stateProvince="Sumatra">Mt. Tanggamus</location>
,
<collectingMunicipality id="6BFEACFBFF88FFDCFF7867B6E34DFE1A" box="[241,343,429,453]" pageId="13" pageNumber="14">Lampung</collectingMunicipality>
,
<collectingRegion id="49E1F863FF88FFDCFEE967B7E3ADFE1B" box="[352,439,428,452]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="14">Sumatra</collectingRegion>
,
<collectingCountry id="F3327611FF88FFDCFE4967B7E032FE1B" box="[448,552,428,452]" name="Indonesia" pageId="13" pageNumber="14">Indonesia</collectingCountry>
;
<emphasis id="B951EA93FF88FFDCFDB867B6E05AFE1B" box="[561,576,429,452]" italics="true" pageId="13" pageNumber="14">c.</emphasis>
–5.44, 104.71).
</paragraph>
<paragraph id="8B9A3681FF88FFDCFFF867FDE2E6FDA4" blockId="13.[113,762,486,635]" pageId="13" pageNumber="14">
<materialsCitation id="3B4D3CDCFF88FFDCFFF867FDE2E3FDA4" collectionCode="RMNH" country="Indonesia" location="Giesting" municipality="Lampung" pageId="13" pageNumber="14" specimenCode="RMNH 23896-98" specimenCount="5" stateProvince="Sumatra" typeStatus="paratype">
<emphasis id="B951EA93FF88FFDCFFF867FDE319FE22" box="[113,259,486,509]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="549E8823FF88FFDCFFF867FDE2CAFE22" box="[113,208,486,509]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D23FD08FF88FFDCFF6B67FDE2E8FE22" box="[226,242,486,509]" pageId="13" pageNumber="14">5</specimenCount>
):
</emphasis>
Five according to
<bibRefCitation id="EFB44B70FF88FFDCFE4667FDE04FFE21" author="Sodv HJV" box="[463,597,486,510]" pageId="13" pageNumber="14" pagination="430 - 42" refId="ref35700" refString="Sodv HJV. XLI - Tbn nbw mammals foom thb Dutch East Indibs. Annals and Magazine of Natural History 1933; 12: 430 - 42. hups: // doi. oog / 10.1080 / 00222933308673706" type="journal article" year="1933">Sody (1933)</bibRefCitation>
, although Sody did not examine them: ‘For examining the Atchinese specimens I have to thank Mr. F.J. Naing-Golan’. Naturalis Biodiversity Centre holds three of these (
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), the other two might be lost
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF88FFDCFFF86487E17FFEFA" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B9A3681FF88FFDCFFF86487E2E1FA3C" blockId="13.[113,763,668,1507]" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFFF86487E320FD6B" box="[113,314,668,692]" italics="true" pageId="13" pageNumber="14">Emended diagnosis:</emphasis>
A large-sized
<taxonomicName id="4C254D02FF88FFDCFE5B6487E030FD6B" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[466,554,668,692]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B951EA93FF88FFDCFE5B6487E030FD6B" box="[466,554,668,692]" italics="true" pageId="13" pageNumber="14">Hylomys</emphasis>
</taxonomicName>
(average
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=
<quantity id="4CDD9B64FF88FFDCFD5B6487E287FD0C" metricMagnitude="-1" metricUnit="m" metricValue="1.39" pageId="13" pageNumber="14" unit="mm" value="139.0">139 mm</quantity>
, W =
<quantity id="4CDD9B64FF88FFDCFF6864A7E30BFD0B" box="[225,273,700,724]" metricMagnitude="-2" metricUnit="kg" metricValue="6.4" pageId="13" pageNumber="14" unit="g" value="64.0">64 g</quantity>
,
<collectionCode id="ED34AE44FF88FFDCFEAB64A7E349FD0B" box="[290,339,700,724]" pageId="13" pageNumber="14">GLS</collectionCode>
=
<quantity id="4CDD9B64FF88FFDCFEF864A7E3C8FD0C" box="[369,466,700,724]" metricMagnitude="-2" metricUnit="m" metricValue="3.69" pageId="13" pageNumber="14" unit="mm" value="36.9">36.9 mm</quantity>
) characterized by a brown suffusion in throat (from late April to mid-November but absent from late February to late April), bicoloured (except specimen
<collectionCode id="ED34AE44FF88FFDCFF6D6501E32DFCEE" box="[228,311,794,817]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH</collectionCode>
23896) and short tail (average
<collectionCode id="ED34AE44FF88FFDCFD266501E0DBFCEE" box="[687,705,794,817]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="14">T</collectionCode>
=
<quantity id="4CDD9B64FF88FFDCFD696501E287FC8F" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="16.0">16 mm</quantity>
,
<collectionCode id="ED34AE44FF88FFDCFF206522E2A1FC8E" box="[169,187,825,849]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="14">T</collectionCode>
/
<collectionCode id="ED34AE44FF88FFDCFF4C6522E2F7FC8E" box="[197,237,825,849]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="13" pageNumber="14" type="Herbarium">HB</collectionCode>
= 11.7%), long hindfoot (average
<collectionCode id="ED34AE44FF88FFDCFDEC6522E093FC8F" box="[613,649,825,848]" country="Brazil" lsid="urn:lsid:biocol.org:col:14339" name="Universidade Federal do Pará" pageId="13" pageNumber="14" type="Herbarium">HF</collectionCode>
=
<quantity id="4CDD9B64FF88FFDCFD216522E0EFFC8F" box="[680,757,825,849]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="mm" value="25.0">25 mm</quantity>
, HF + nail =
<quantity id="4CDD9B64FF88FFDCFF736543E350FCAF" box="[250,330,856,880]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="14" unit="mm" value="26.0">26 mm</quantity>
), uniformly brown-coloured and relatively short ears (average E =
<quantity id="4CDD9B64FF88FFDCFE4C6563E002FC50" box="[453,536,888,912]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="16.0">16 mm</quantity>
, E/
<collectionCode id="ED34AE44FF88FFDCFDC86563E06FFC4F" box="[577,629,888,912]" pageId="13" pageNumber="14">GLS</collectionCode>
= 45%). Its skull can be distinguished from all relatives by its relatively larger molars (Supporting Information, File S1,
<figureCitation id="131E2A04FF88FFDCFD1765ADE0F5FC11" box="[670,751,950,974]" captionStart="Figure 5" captionStartId="9.[113,178,1006,1030]" captionTargetBox="[113,1457,145,977]" captionTargetId="figure-518@9.[112,1459,144,979]" captionTargetPageId="9" captionText="Figure 5. Morphometric variation in Hylomys. A, first and second principal components from an analysis of 16 cranial dimensions, computed with Dataset 4 (with the exclusion of external measurements); B, second and third principal components of the same analysis.Asterisks denote undamaged holotype or neotype specimens that could be included in this analysis." figureDoi="http://doi.org/10.5281/zenodo.14510273" httpUri="https://zenodo.org/record/14510273/files/figure.png" pageId="13" pageNumber="14">Fig. S5</figureCitation>
) and from all relatives but allopatric
<taxonomicName id="4C254D02FF88FFDCFE7A65CDE049FC31" authorityName="Mullbo" authorityYear="1840" box="[499,595,982,1006]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suillus">
<emphasis id="B951EA93FF88FFDCFE7A65CDE049FC31" box="[499,595,982,1006]" italics="true" pageId="13" pageNumber="14">H. suillus</emphasis>
</taxonomicName>
by its long and relatively broad rostrum (
<figureCitation id="131E2A04FF88FFDCFE0E65EEE3D9FBD2" box="[391,451,1013,1037]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." figureDoi="http://doi.org/10.5281/zenodo.14510275" httpUri="https://zenodo.org/record/14510275/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
). Its skull can also be distinguished from all other relatives by the combination of the following characters: obtuse-angled notch between anterior side of premaxilla; posterior end of nasals does not extend to the level of the antorbital rim; dorsal region of maxillary and frontal have a rugged appearance with capillary grooves and pores; prominent antorbital ridge/flange; anterior side of the antorbital rim is posterior to second cusp of M1; robust supraorbital processes and dentition; short alisphenoid canal (average
<collectionCode id="ED34AE44FF88FFDCFE986314E359FAF8" box="[273,323,1295,1319]" country="USA" lsid="urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="13" pageNumber="14" type="Herbarium">ASC</collectionCode>
=
<quantity id="4CDD9B64FF88FFDCFEE86314E3ABFAF9" box="[353,433,1295,1319]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="0.2">0.2 mm</quantity>
), prominent posterior cuspule in P
<quantity id="4CDD9B64FF88FFDCFF296335E2D1FA99" box="[160,203,1326,1350]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="14" unit="in" value="2.0">2 in</quantity>
Sumatran specimens with unworn dentition; P3 is much larger than P1; P4 has a thick labial cingulum; posterior nasal spine present; prominent occipital crest that extends more ventrally than in most other species; single root/two well fused roots in p3; anterior margin of p1 crown extends to root of c1.
</paragraph>
<paragraph id="8B9A3681FF88FFDCFFF8601EE16AFF18" blockId="13.[113,762,1540,1972]" lastBlockId="13.[810,1459,144,293]" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFFF8601EE2E1F9C3" box="[113,251,1541,1564]" italics="true" pageId="13" pageNumber="14">Comparisons:</emphasis>
Distinguished from parapatric
<taxonomicName id="4C254D02FF88FFDCFDDA601EE0A2F9C3" baseAuthorityName="Robinson and Kloss" baseAuthorityYear="1916" box="[595,696,1541,1564]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFDDA601EE0A2F9C3" box="[595,696,1541,1564]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
by its larger size (
<collectionCode id="ED34AE44FF88FFDCFF64603FE302F9E3" box="[237,280,1572,1596]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="13" pageNumber="14" type="Herbarium">HB</collectionCode>
:
<quantity id="4CDD9B64FF88FFDCFEAD6038E3A9F9E4" box="[292,435,1571,1595]" metricMagnitude="-1" metricUnit="m" metricValue="1.395" metricValueMax="1.56" metricValueMin="1.23" pageId="13" pageNumber="14" unit="mm" value="139.5" valueMax="156.0" valueMin="123.0">123–156 mm</quantity>
vs.
<quantity id="4CDD9B64FF88FFDCFE696038E06BF9E4" box="[480,625,1571,1595]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="13" pageNumber="14" unit="mm" value="107.5" valueMax="115.0" valueMin="100.0">100–115 mm</quantity>
,
<collectionCode id="ED34AE44FF88FFDCFDF7603FE0A9F9E3" box="[638,691,1572,1596]" pageId="13" pageNumber="14">GLS</collectionCode>
:
<quantity id="4CDD9B64FF88FFDCFD366038E2D7F985" metricMagnitude="-2" metricUnit="m" metricValue="3.695" metricValueMax="3.84" metricValueMin="3.55" pageId="13" pageNumber="14" unit="mm" value="36.95" valueMax="38.4" valueMin="35.5">35.5– 38.4 mm</quantity>
vs.
<quantity id="4CDD9B64FF88FFDCFF7A6058E395F984" box="[243,399,1603,1627]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" metricValueMax="3.34" metricValueMin="2.92" pageId="13" pageNumber="14" unit="mm" value="31.299999999999997" valueMax="33.4" valueMin="29.2">29.2–33.4 mm</quantity>
), relatively shorter tail (average: 16 vs.
<quantity id="4CDD9B64FF88FFDCFF1A6079E2C5F9A5" box="[147,223,1634,1658]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="13" pageNumber="14" unit="mm" value="23.0">23 mm</quantity>
) and harsher dorsal fur. Its skull has a more angular and robust appearance than
<taxonomicName id="4C254D02FF88FFDCFE136099E3E6F946" baseAuthorityName="Robinson and Kloss" baseAuthorityYear="1916" box="[410,508,1666,1689]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFE136099E3E6F946" box="[410,508,1666,1689]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
, its supraorbital process is well developed vs. absent in
<taxonomicName id="4C254D02FF88FFDCFE3760B9E038F966" baseAuthorityName="Robinson and Kloss" baseAuthorityYear="1916" box="[446,546,1698,1721]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFE3760B9E038F966" box="[446,546,1698,1721]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
, and its dentition is robust (particularly the canines and adjacent teeth) vs. gracile in
<taxonomicName id="4C254D02FF88FFDCFF0260FBE3ACF927" authority="(Rubdi et al. 1994)" baseAuthorityName="Rubdi" baseAuthorityYear="1994" box="[139,438,1759,1784]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFF0260FBE2F6F928" box="[139,236,1760,1783]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
(
<bibRefCitation id="EFB44B70FF88FFDCFF7460C4E3B0F927" author="Rubdi M &amp; Chapuisat M &amp; Iskandao D" box="[253,426,1759,1784]" pageId="13" pageNumber="14" pagination="965 - 78" refId="ref35216" refString="Rubdi M, Chapuisat M, Iskandao D. Taxonomic status of Hylomys parous and Hylomys suillus (Insbctivooa: Eoinacbidab): biochbmical and moophological analvsbs. Journal of Mammalogy 1994; 75: 965 - 78. hups: // doi. oog / 10.2307 / 1382478" type="journal article" year="1994">
Ruedi
<emphasis id="B951EA93FF88FFDCFEC860FBE36AF928" box="[321,368,1759,1783]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
1994
</bibRefCitation>
)
</taxonomicName>
. The notch between premaxilla tips is obtuse vs. acute-angled in
<taxonomicName id="4C254D02FF88FFDCFE5C611BE023F8C8" box="[469,569,1792,1815]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFE5C611BE023F8C8" box="[469,569,1792,1815]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
. The frontal bone and dorsal region of maxillary is more rugged, with greater capillary grooves and pores, than in
<taxonomicName id="4C254D02FF88FFDCFE7A6125E04DF88A" box="[499,599,1854,1877]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFE7A6125E04DF88A" box="[499,599,1854,1877]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
. It has a prominent antorbital ridge and occipital crest, and thick P4 labial cingulum, while these are absent or vestigial/inconspicuous in
<taxonomicName id="4C254D02FF88FFDCFFF86187E2CFF86C" box="[113,213,1948,1971]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFFF86187E2CFF86C" box="[113,213,1948,1971]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
</taxonomicName>
. It has a single root/two well fused roots in P2 and p3 vs. two non-fused roots in
<taxonomicName id="4C254D02FF88FFDCFBD3668AE171FF18" authority="(Jbnkins and Robinson 2002)" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF88FFDCFBD3668AE6A6FF77" box="[1114,1212,145,168]" italics="true" pageId="13" pageNumber="14">H. parvus</emphasis>
(
<bibRefCitation id="EFB44B70FF88FFDCFB46668BE145FF18" author="Jbnkins PD &amp; Robinson MF" pageId="13" pageNumber="14" pagination="1 - 11" refId="ref32930" refString="Jbnkins PD, Robinson MF. Anothbo vaoiation on thb gvmnuob thbmb: dbscoiption of a nbw spbcibs of Hylomys (Lipotvphla, Eoinacbidab, Galboicinab). Bulletin of Ne Natural History Museum, Zoology Series 2002; 68: 1 - 11." type="journal article" year="2002">Jenkins and Robinson 2002</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A3681FF88FFDCFCCC66D5E17FFEFA" blockId="13.[810,1459,144,293]" pageId="13" pageNumber="14">Comparisons with additional species have been included in the following accounts, after such species have been formally defined.</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF88FFDCFCA3675DE747FA96" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B9A3681FF88FFDCFCA3675DE747FA96" blockId="13.[810,1461,326,1823]" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFCA3675DE6F0FE81" box="[810,1258,326,350]" italics="true" pageId="13" pageNumber="14">Distribution, habitat, and natural history:</emphasis>
Currently known from forests between
<emphasis id="B951EA93FF88FFDCFB99677CE605FEA1" box="[1040,1055,359,382]" italics="true" pageId="13" pageNumber="14">c.</emphasis>
100 and
<quantity id="4CDD9B64FF88FFDCFB08677DE6CBFEA2" box="[1153,1233,358,382]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="2000.0">2000 m</quantity>
a.s.l. on Sumatra and 600 and
<quantity id="4CDD9B64FF88FFDCFC04679EE1FBFE42" box="[909,993,389,413]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="m" value="1700.0">1700 m</quantity>
a.s.l. on the Malay peninsula, possibly extending north up to the Kangar Pattani Line (K-P) vegetation transition, which cuts across the Thailand–Malaysia border (see comments on the following species account).
<taxonomicName id="4C254D02FF88FFDCFAD267F8E147FDC5" authorityName="Sody" authorityYear="1933" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF88FFDCFAD267F8E147FDC5" italics="true" pageId="13" pageNumber="14">Hylomys maxi</emphasis>
</taxonomicName>
seems more abundant in hill and montane forests and was not recorded in dipterocarp forest in surveys on Northern Sumatra and Peninsular Malaysia (
<bibRefCitation id="EFB44B70FF88FFDCFB1B645AE734FD86" author="Langham N" box="[1170,1326,577,601]" pageId="13" pageNumber="14" pagination="199 - 206" refId="ref33394" refString="Langham N. Distoibution and bcologv of small mammals in thobb oain foobst localitibs of pbninsula Malavsia with paoticulao obfbobncbs to Kbdah Pbak. Biotropica 1983; 15: 199 - 206. hups: // doi. oog / 10.2307 / 2387829" type="journal article" year="1983">Langham 1983</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFAB0645AE145FDA6" author="Boubli JP &amp; Gobllb CEV &amp; van Schaik CP" pageId="13" pageNumber="14" pagination="149 - 54" refId="ref31019" refString="Boubli JP, Gobllb CEV, van Schaik CP. Small mammal spbcibs divbositv and composition in two bcologicallv distinct oainfooobst sitbs in noothbon Sumatoa, Indonbsia. Ecotropica 2004; 10: 149 - 54." type="journal article" year="2004">
Boubli
<emphasis id="B951EA93FF88FFDCFA0C6459E7AEFD86" box="[1413,1460,577,601]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2004
</bibRefCitation>
). This species has been collected in ‘jungle filled hollow surrounded by grasslands’ (Fred Ulmer field notes,
<collectionCode id="ED34AE44FF88FFDCFAF8649BE7A9FD47" box="[1393,1459,640,664]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="13" pageNumber="14">ANSP</collectionCode>
archives) and in secondary vegetation on the edge of mountain forests, in highly anthropized areas (gardens, golf courses, etc.; authors’ unpublished data).
<bibRefCitation id="EFB44B70FF88FFDCFB0364C5E7A9FD29" author="Robinson HC &amp; Kloss CB" box="[1162,1459,734,758]" pageId="13" pageNumber="14" pagination="1 - 80" refId="ref35040" refString="Robinson HC, Kloss CB. Rbsults of an bxpbdition to Kooinchi Pbak, Sumatoa I Mammals. Journal of the Federated Malay States Museum 1918; 8: 1 - 80." type="journal article" year="1918">Robinson and Kloss (1918)</bibRefCitation>
stated that the species was ‘quite numerous at dusk, running about in a patch of sweet potato (
<taxonomicName id="4C254D02FF88FFDCFB066506E72CFCEB" box="[1167,1334,796,820]" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="batatas">
<emphasis id="B951EA93FF88FFDCFB066506E72CFCEB" box="[1167,1334,796,820]" italics="true" pageId="13" pageNumber="14">Ipomoea batatas</emphasis>
</taxonomicName>
) planted in a flat place among rocks at the edge of a torrent’. In Peninsular Malaysia it shares habitat with
<taxonomicName id="4C254D02FF88FFDCFBFA6540E75BFCAC" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[1139,1345,859,883]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fuliginosa">
<emphasis id="B951EA93FF88FFDCFBFA6540E75BFCAC" box="[1139,1345,859,883]" italics="true" pageId="13" pageNumber="14">Crocidura fuliginosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFAC76540E18DFC4D" authorityName="Jentink" authorityYear="1888" class="Mammalia" family="Soricidae" genus="Crocidura" isUncertain="true" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="neglecta">
<emphasis id="B951EA93FF88FFDCFAC76540E7A9FCAC" box="[1358,1459,859,883]" italics="true" pageId="13" pageNumber="14">Crocidura</emphasis>
cf.
<emphasis id="B951EA93FF88FFDCFCCE6561E18DFC4D" box="[839,919,890,914]" italics="true" pageId="13" pageNumber="14">neglecta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFC286561E674FC4D" authorityName="Robinson &amp; Kloss" authorityYear="1911" box="[929,1134,890,914]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="malayana">
<emphasis id="B951EA93FF88FFDCFC286561E674FC4D" box="[929,1134,890,914]" italics="true" pageId="13" pageNumber="14">Crocidura malayana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFBF06560E735FC4D" baseAuthorityName="Kloss" baseAuthorityYear="1917" box="[1145,1327,890,914]" class="Mammalia" family="Soricidae" genus="Suncus" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B951EA93FF88FFDCFBF06560E735FC4D" box="[1145,1327,890,914]" italics="true" pageId="13" pageNumber="14">Suncus malayanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFAB36560E18CFC6D" class="Mammalia" family="Talpidae" genus="Euroscaptor" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B951EA93FF88FFDCFAB36560E18CFC6D" italics="true" pageId="13" pageNumber="14">Euroscaptor malayanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFC2B6581E615FC6D" baseAuthorityName="Diard" baseAuthorityYear="1820" box="[930,1039,922,946]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="glis">
<emphasis id="B951EA93FF88FFDCFC2B6581E615FC6D" box="[930,1039,922,946]" italics="true" pageId="13" pageNumber="14">Tupaia glis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB926581E6DDFC6D" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[1051,1223,922,946]" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="bowersi">
<emphasis id="B951EA93FF88FFDCFB926581E6DDFC6D" box="[1051,1223,922,946]" italics="true" pageId="13" pageNumber="14">Berylmys bowersi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB5A6581E7B7FC6D" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1235,1453,922,946]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="sabanus">
<emphasis id="B951EA93FF88FFDCFB5A6581E7B7FC6D" box="[1235,1453,922,946]" italics="true" pageId="13" pageNumber="14">Leopoldamys sabanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA365A2E616FC0E" baseAuthorityName="Bonhote" baseAuthorityYear="1900" box="[810,1036,953,977]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="B951EA93FF88FFDCFCA365A2E616FC0E" box="[810,1036,953,977]" italics="true" pageId="13" pageNumber="14">Leopoldamys ciliatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFBA065A1E6F8FC0E" box="[1065,1250,953,977]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="surifer">
<emphasis id="B951EA93FF88FFDCFBA065A1E6F8FC0E" box="[1065,1250,953,977]" italics="true" pageId="13" pageNumber="14">Maxomys surifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB7765A1E7B6FC0E" baseAuthorityName="Thomas" baseAuthorityYear="1894" box="[1278,1452,953,977]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rajah">
<emphasis id="B951EA93FF88FFDCFB7765A1E7B6FC0E" box="[1278,1452,953,977]" italics="true" pageId="13" pageNumber="14">Maxomys rajah</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA365C2E618FC2F" baseAuthorityName="Thomas" baseAuthorityYear="1894" box="[810,1026,984,1008]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="whiteheadi">
<emphasis id="B951EA93FF88FFDCFCA365C2E618FC2F" box="[810,1026,984,1008]" italics="true" pageId="13" pageNumber="14">Maxomys whiteheadi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB9865C2E706FC2F" authority=", Nioioenter" authorityName="Nioioenter" box="[1041,1308,985,1008]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inas">
<emphasis id="B951EA93FF88FFDCFB9865C2E6BFFC2F" box="[1041,1189,985,1008]" italics="true" pageId="13" pageNumber="14">Maxomys inas</emphasis>
,
<emphasis id="B951EA93FF88FFDCFB3D65C2E7B6FC2F" box="[1204,1452,985,1008]" italics="true" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFB3D65C2E706FC2F" box="[1204,1308,985,1008]" italics="true" pageId="13" pageNumber="14">Niviventer</emphasis>
<emphasis id="B951EA93FF88FFDCFAAD65C2E7B6FC2F" box="[1316,1452,985,1008]" italics="true" pageId="13" pageNumber="14">cremoriventer</emphasis>
</emphasis>
</taxonomicName>
,
<emphasis id="B951EA93FF88FFDCFCA365E3E1EEFBD0" box="[810,1012,1016,1039]" italics="true" pageId="13" pageNumber="14">Niviventer cameroni</emphasis>
,
<taxonomicName id="4C254D02FF88FFDCFB8B65E3E6DBFBCF" baseAuthorityName="Jentink" baseAuthorityYear="1879" box="[1026,1217,1016,1040]" class="Mammalia" family="Muridae" genus="Sundamys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="B951EA93FF88FFDCFB8B65E3E6DBFBCF" box="[1026,1217,1016,1040]" italics="true" pageId="13" pageNumber="14">Sundamys muelleri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB4765E3E7B7FBCF" baseAuthorityName="Bonhote" baseAuthorityYear="1903" box="[1230,1453,1016,1040]" class="Mammalia" family="Muridae" genus="Sundamys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="B951EA93FF88FFDCFB4765E3E7B7FBCF" box="[1230,1453,1016,1040]" italics="true" pageId="13" pageNumber="14">Sundamys annandalei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA36203E1D0FBF0" baseAuthorityName="Peale" baseAuthorityYear="1848" box="[810,970,1047,1071]" genus="Ratus" pageId="13" pageNumber="14" rank="species" species="exulans">
<emphasis id="B951EA93FF88FFDCFCA36203E1D0FBF0" box="[810,970,1047,1071]" italics="true" pageId="13" pageNumber="14">Rattus exulans</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFC6B620CE6D8FBF0" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[994,1218,1047,1071]" class="Mammalia" family="Sciuridae" genus="Callosciurus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="caniceps">
<emphasis id="B951EA93FF88FFDCFC6B620CE6D8FBF0" box="[994,1218,1047,1071]" italics="true" pageId="13" pageNumber="14">Callosciurus caniceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB526203E7B7FBF0" baseAuthorityName="Blanford" baseAuthorityYear="1878" box="[1243,1453,1047,1071]" class="Mammalia" family="Sciuridae" genus="Dremomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufigenis">
<emphasis id="B951EA93FF88FFDCFB526203E7B7FBF0" box="[1243,1453,1047,1071]" italics="true" pageId="13" pageNumber="14">Dremomys rufigenis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA3622CE1CFFB91" baseAuthorityName="F. Cuvier" baseAuthorityYear="1821" box="[810,981,1079,1102]" class="Mammalia" family="Sciuridae" genus="Lariscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="insignis">
<emphasis id="B951EA93FF88FFDCFCA3622CE1CFFB91" box="[810,981,1079,1102]" italics="true" pageId="13" pageNumber="14">Lariscus insignis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D02FF88FFDCFB94622CE6F0FB90" baseAuthorityName="Bonhote" baseAuthorityYear="1908" box="[1053,1258,1079,1103]" class="Mammalia" family="Sciuridae" genus="Sundasciurus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tahan">
<emphasis id="B951EA93FF88FFDCFB94622CE6F0FB90" box="[1053,1258,1079,1103]" italics="true" pageId="13" pageNumber="14">Sundasciurus tahan</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB44B70FF88FFDCFA8A622CE7B7FB90" author="Robinson HC" box="[1283,1453,1079,1103]" pageId="13" pageNumber="14" pagination="235 - 41" refId="ref34956" refString="Robinson HC. A list of small collbction of mammals and biods foom thb mountains of Ulu Langat, Sblangoo. Journal of the Federated Malay States Museum 1911; 4: 235 - 41." type="journal article" year="1911">Robinson 1911</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFCA3624DE622FBB1" author="Haooison JL &amp; Toaub R" box="[810,1080,1110,1134]" pageId="13" pageNumber="14" pagination="337 - 46" refId="ref32314" refString="Haooison JL, Toaub R. Rodbnts and Insbctivoobs foom Sblangoo, Malava. Journal of Mammalogy 1950; 31: 337 - 46. hups: // doi. oog / 10.2307 / 1375305" type="journal article" year="1950">Harrison and Traub 1950</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFBCC624DE6A1FBB1" author="Rudd RL" box="[1093,1211,1110,1134]" pageId="13" pageNumber="14" pagination="588 - 94" refId="ref35114" refString="Rudd RL. Wbight and goowth in Malavsian oain foobst mammals. Journal of Mammalogy 1965; 46: 588 - 94." type="journal article" year="1965">Rudd 1965</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFB41624DE77CFBB1" author="Langham N" box="[1224,1382,1110,1134]" pageId="13" pageNumber="14" pagination="199 - 206" refId="ref33394" refString="Langham N. Distoibution and bcologv of small mammals in thobb oain foobst localitibs of pbninsula Malavsia with paoticulao obfbobncbs to Kbdah Pbak. Biotropica 1983; 15: 199 - 206. hups: // doi. oog / 10.2307 / 2387829" type="journal article" year="1983">Langham 1983</bibRefCitation>
, Ruedi 1995,
<bibRefCitation id="EFB44B70FF88FFDCFCF9626EE634FB52" author="Omao H &amp; Adamson EAS &amp; Bhassu S" box="[880,1070,1141,1165]" pageId="13" pageNumber="14" pagination="237 - 43" refId="ref34520" refString="Omao H, Adamson EAS, Bhassu S et al. Phvlogbnbtic oblationships of Malavan and Malagasv Pvmgv Shobws of thb gbnus Suncus (Sooicomoopha: Sooicidab) infboobd foom mitochondoial cytochrome b gbnb sbqubncbs. Raffles Bulletin of Zoology 2011; 59: 237 - 43." type="journal article" year="2011">
Omar
<emphasis id="B951EA93FF88FFDCFC30626DE1F5FB52" box="[953,1007,1141,1165]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2011
</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFBB6626EE66EFB52" author="Omao H &amp; Hashim R &amp; Bhassu S" box="[1087,1140,1141,1165]" pageId="13" pageNumber="14" pagination="446 - 54" refId="ref34568" refString="Omao H, Hashim R, Bhassu S et al. Moophological and gbnbtic oblationships of thb Crocidura monticola spbcibs complbx (Sooicidab: Coociduoinab) in Sundaland. Mammalian Biology 2013; 78: 446 - 54. hups: // doi. oog / 10.1016 / j. mambio. 2013.04.004" type="journal article" year="2013">2013</bibRefCitation>
, authors’ unpublished data). In Sumatra,
<taxonomicName id="4C254D02FF88FFDCFC24628EE61BFB73" authorityName="Sody" authorityYear="1933" box="[941,1025,1173,1196]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF88FFDCFC24628EE61BFB73" box="[941,1025,1173,1196]" italics="true" pageId="13" pageNumber="14">H. maxi</emphasis>
</taxonomicName>
shares its habitat with
<taxonomicName id="4C254D02FF88FFDCFB73628EE7B7FB72" authorityName="Dobson" authorityYear="1887" box="[1274,1453,1173,1197]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="beccari">
<emphasis id="B951EA93FF88FFDCFB73628EE7B7FB72" box="[1274,1453,1173,1197]" italics="true" pageId="13" pageNumber="14">Crocidura beccari</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA362AFE1EBFB13" authorityName="Lyon" authorityYear="1908" box="[810,1009,1204,1228]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lepidura">
<emphasis id="B951EA93FF88FFDCFCA362AFE1EBFB13" box="[810,1009,1204,1228]" italics="true" pageId="13" pageNumber="14">Crocidura lepidura</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB8F62AFE6F7FB13" authorityName="Robinson &amp; Kloss" authorityYear="1918" box="[1030,1261,1204,1228]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="aequicauda">
<emphasis id="B951EA93FF88FFDCFB8F62AFE6F7FB13" box="[1030,1261,1204,1228]" italics="true" pageId="13" pageNumber="14">Crocidura aequicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFA8B62AEE7B7FB13" authorityName="Horsfield" authorityYear="1822" box="[1282,1453,1205,1228]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="jaoanica">
<emphasis id="B951EA93FF88FFDCFA8B62AEE7B7FB13" box="[1282,1453,1205,1228]" italics="true" pageId="13" pageNumber="14">Tupaia javanica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCA362CFE1F9FB34" authorityName="Raffles" authorityYear="1821" box="[810,995,1235,1259]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ferruginea">
<emphasis id="B951EA93FF88FFDCFCA362CFE1F9FB34" box="[810,995,1235,1259]" italics="true" pageId="13" pageNumber="14">Tupaia ferruginea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFC7C62CFE66FFB34" authorityName="Raffles" authorityYear="1821" box="[1013,1141,1236,1259]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tana">
<emphasis id="B951EA93FF88FFDCFC7C62CFE66FFB34" box="[1013,1141,1236,1259]" italics="true" pageId="13" pageNumber="14">Tupaia tana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB0E62CFE75AFB34" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1916" box="[1159,1344,1235,1259]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inflatus">
<emphasis id="B951EA93FF88FFDCFB0E62CFE75AFB34" box="[1159,1344,1235,1259]" italics="true" pageId="13" pageNumber="14">Maxomys inflatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFAD862CFE145FAD4" baseAuthorityName="Thomas" baseAuthorityYear="1894" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rajah">
<emphasis id="B951EA93FF88FFDCFAD862CFE145FAD4" italics="true" pageId="13" pageNumber="14">Maxomys rajah</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFCE262E8E608FAD4" box="[875,1042,1267,1291]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="surifer">
<emphasis id="B951EA93FF88FFDCFCE262E8E608FAD4" box="[875,1042,1267,1291]" italics="true" pageId="13" pageNumber="14">Maxomys surifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFB9562E8E6EDFAD5" authority=", Nioioenter" authorityName="Nioioenter" box="[1052,1271,1267,1291]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="caroli">
<emphasis id="B951EA93FF88FFDCFB9562E8E69EFAD4" box="[1052,1156,1267,1291]" italics="true" pageId="13" pageNumber="14">Mus caroli</emphasis>
,
<emphasis id="B951EA93FF88FFDCFB0662E8E74FFAD5" box="[1167,1365,1267,1290]" italics="true" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFB0662E8E6EDFAD5" box="[1167,1271,1267,1290]" italics="true" pageId="13" pageNumber="14">Niviventer</emphasis>
<emphasis id="B951EA93FF88FFDCFB7562E8E74FFAD5" box="[1276,1365,1267,1290]" italics="true" pageId="13" pageNumber="14">fraternus</emphasis>
</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF88FFDCFAD662E8E17AFAF5" baseAuthorityName="Thomas" baseAuthorityYear="1898" class="Mammalia" family="Sciuridae" genus="Lariscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="niobe">
<emphasis id="B951EA93FF88FFDCFAD662E8E17AFAF5" italics="true" pageId="13" pageNumber="14">Lariscus niobe</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C254D02FF88FFDCFC106309E691FAF5" baseAuthorityName="Millbo" baseAuthorityYear="1942" box="[921,1163,1298,1322]" class="Mammalia" family="Sciuridae" genus="Sundasciurus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="altitudinis">
<emphasis id="B951EA93FF88FFDCFC106309E691FAF5" box="[921,1163,1298,1322]" italics="true" pageId="13" pageNumber="14">Sundasciurus altitudinis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB44B70FF88FFDCFB146309E7B7FAF5" author="Robinson HC &amp; Kloss CB" box="[1181,1453,1298,1322]" pageId="13" pageNumber="14" pagination="1 - 80" refId="ref35040" refString="Robinson HC, Kloss CB. Rbsults of an bxpbdition to Kooinchi Pbak, Sumatoa I Mammals. Journal of the Federated Malay States Museum 1918; 8: 1 - 80." type="journal article" year="1918">Robinson and Kloss 1918</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFCA3632AE1BCFA96" author="Millbo GS" box="[810,934,1329,1353]" pageId="13" pageNumber="14" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller 1942</bibRefCitation>
, Ruedi 1995, authors’ unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF88FFDFFCCF634AE3C5FD24" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8B9A3681FF88FFDCFCCF634AE628F8C0" blockId="13.[810,1461,326,1823]" pageId="13" pageNumber="14">
<taxonomicName id="4C254D02FF88FFDCFCCF634AE1C8FAB7" authorityName="Sody" authorityYear="1933" box="[838,978,1361,1385]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF88FFDCFCCF634AE1C8FAB7" box="[838,978,1361,1385]" italics="true" pageId="13" pageNumber="14">Hylomys maxi</emphasis>
</taxonomicName>
is omnivorous. In Peninsular Malaysia stomach contents contained tapioca bait and several crickets (
<emphasis id="B951EA93FF88FFDCFAE6636AE798FA57" box="[1391,1410,1393,1416]" italics="true" pageId="13" pageNumber="14">N</emphasis>
= 1;
<bibRefCitation id="EFB44B70FF88FFDCFCA56394E1D2FA78" author="Haooison JL" box="[812,968,1423,1447]" pageId="13" pageNumber="14" pagination="5 - 18" refId="ref32289" refString="Haooison JL. Nb natuoal food of somb Malavan mammals. Bulletin of the Singapore National Museum 1961; 30: 5 - 18." type="journal article" year="1961">Harrison 1961</bibRefCitation>
). In Sumatra, stomachs contained insects of various orders, principally small beetles (
<emphasis id="B951EA93FF88FFDCFB4663B4E6F8FA19" box="[1231,1250,1455,1478]" italics="true" pageId="13" pageNumber="14">N</emphasis>
= 2–3). In captivity they feed on fish, shrimp, insects (cockroaches, mealworms, grasshoppers), raw meat, and soft fruit, showing less interest in the latter (
<bibRefCitation id="EFB44B70FF88FFDCFC326016E629F9FB" author="Davis D" box="[955,1075,1548,1573]" pageId="13" pageNumber="14" pagination="147 - 8" refId="ref31652" refString="Davis D. Nb Lbssbo Gvmnuob, Hylomys suillus. Ne Malayan Nature Journal 1965; 19: 147 - 8." type="journal article" year="1965">Davis 1965</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFBCB6016E6CDF9FA" author="Mbdwav L" box="[1090,1239,1549,1573]" pageId="13" pageNumber="14" refId="ref33924" refString="Mbdwav L. Nb wild mammals of Malava and offshoob islands including Singapoob. In: The Wild Mammals of Malaya and Offshore Islands Including Singapore. London: Oxfood Univbositv Pobss, 1969." type="book" year="1969">Medway 1969</bibRefCitation>
,
<bibRefCitation id="EFB44B70FF88FFDCFB6E6016E144F99B" author="Gbnoud M &amp; Rubdi M" pageId="13" pageNumber="14" pagination="309 - 16" refId="ref32145" refString="Gbnoud M, Rubdi M. Ratb of mbtabolism, tbmpboatuob obgulations, and bvapooativb watbo loss in thb lbssbo gvmnuob Hylomys suillus (Insbctivooa, Mammalia). Journal of Zoology 1996; 240: 309 - 16. hups: // doi. oog / 10.1111 / j. 1469 - 7998.1996. tb 05287. x" type="journal article" year="1996">Genoud and Ruedi 1996</bibRefCitation>
). In Selangor, a pregnant individual with two embryos was recorded in March and females with enlarged mammae were recorded from May to November (
<bibRefCitation id="EFB44B70FF88FFDCFB416070E740F95C" author="Mbdwav L" box="[1224,1370,1643,1667]" pageId="13" pageNumber="14" refId="ref33924" refString="Mbdwav L. Nb wild mammals of Malava and offshoob islands including Singapoob. In: The Wild Mammals of Malaya and Offshore Islands Including Singapore. London: Oxfood Univbositv Pobss, 1969." type="book" year="1969">Medway 1969</bibRefCitation>
; USNM specimens).
<taxonomicName id="4C254D02FF88FFDCFC256091E620F97D" authorityName="Sody" authorityYear="1933" box="[940,1082,1674,1698]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF88FFDCFC256091E620F97D" box="[940,1082,1674,1698]" italics="true" pageId="13" pageNumber="14">Hylomys maxi</emphasis>
</taxonomicName>
is cathemeral and found alone or in small groups (2–3 individuals; Fred Ulmer field notes,
<bibRefCitation id="EFB44B70FF88FFDCFAD560B2E144F93F" author="Mbdwav L" pageId="13" pageNumber="14" refId="ref33924" refString="Mbdwav L. Nb wild mammals of Malava and offshoob islands including Singapoob. In: The Wild Mammals of Malaya and Offshore Islands Including Singapore. London: Oxfood Univbositv Pobss, 1969." type="book" year="1969">Medway 1969</bibRefCitation>
). In West Malaysia, average densities in favourable habitats are estimated at 3–4 individuals/ha. Home ranges are
<emphasis id="B951EA93FF88FFDCFACC60F2E754F8DF" box="[1349,1358,1769,1792]" italics="true" pageId="13" pageNumber="14">c</emphasis>
.
<quantity id="4CDD9B64FF88FFDCFAD160F3E7A9F920" box="[1368,1459,1768,1792]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">30–40 m</quantity>
in diameter (
<bibRefCitation id="EFB44B70FF88FFDCFC27611CE63BF8C0" author="Rudd RL" box="[942,1057,1799,1823]" pageId="13" pageNumber="14" pagination="1 - 112" refId="ref35137" refString="Rudd RL. Population dbnsitv and movbmbnts of thb lbssbo gvmnuob, Hylomys suillus. Malayan Nature Journal 1980; 34: 1 - 112." type="journal article" year="1980">Rudd 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A3681FF88FFDFFCA3615AE3C5FD24" blockId="13.[810,1460,1857,1975]" lastBlockId="14.[129,778,144,763]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B951EA93FF88FFDCFCA3615AE1A2F887" box="[810,952,1857,1880]" italics="true" pageId="13" pageNumber="14">Conservation:</emphasis>
Recorded in Taman Negara, Kerinci Seblat and Leuser national parks, Pulau Tioman Wildlife Reserve, and Ulu Gombak, Ulu Langat, and Temengor forest reserves. Temengor has been selectively logged since 2001 and captures of
<taxonomicName id="4C254D02FF8BFFDFFF14668AE2EBFF77" authorityName="Sody" authorityYear="1933" box="[157,241,145,168]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF8BFFDFFF14668AE2EBFF77" box="[157,241,145,168]" italics="true" pageId="14" pageNumber="15">H. maxi</emphasis>
</taxonomicName>
have not changed between sites with and without logging residues (
<bibRefCitation id="EFB44B70FF8BFFDFFECB66B4E017FF18" author="Yamada T &amp; Yoshida S &amp; Hosaka T" box="[322,525,175,199]" pageId="14" pageNumber="15" pagination="100 - 4" refId="ref36399" refString="Yamada T, Yoshida S, Hosaka T et al. Logging obsidubs consbovb small mammalian divbositv in a Malavsian pooduction foobst. Biological Conseroation 2016; 194: 100 - 4. hups: // doi. oog / 10.1016 / j. biocon. 2015.12. 004" type="journal article" year="2016">
Yamada
<emphasis id="B951EA93FF8BFFDFFE1566ABE3CAFF18" box="[412,464,175,199]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2016
</bibRefCitation>
). It is unclear how ongoing deforestation in Kerinci Seblat and Leuser National Parks (see
<taxonomicName id="4C254D02FF8BFFDFFF7066F4E34FFED9" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[249,341,239,262]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="oorax">
<emphasis id="B951EA93FF8BFFDFFF7066F4E34FFED9" box="[249,341,239,262]" italics="true" pageId="14" pageNumber="15">H. vorax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D02FF8BFFDFFE0666F4E3EDFED9" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[399,503,239,262]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="parous">
<emphasis id="B951EA93FF8BFFDFFE0666F4E3EDFED9" box="[399,503,239,262]" italics="true" pageId="14" pageNumber="15">H. parvus</emphasis>
</taxonomicName>
accounts) is affecting
<taxonomicName id="4C254D02FF8BFFDFFD7966F4E2AEFEFA" authorityName="Sody" authorityYear="1933" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF8BFFDFFD7966F4E2AEFEFA" italics="true" pageId="14" pageNumber="15">H. maxi</emphasis>
</taxonomicName>
, nor the degree of deforestation in Malaysian protected areas.
<taxonomicName id="4C254D02FF8BFFDFFF4D6736E34DFE9B" authorityName="Sody" authorityYear="1933" box="[196,343,301,325]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF8BFFDFFF4D6736E34DFE9B" box="[196,343,301,325]" italics="true" pageId="14" pageNumber="15">Hylomys maxi</emphasis>
</taxonomicName>
has been recorded in anthropized habitats such as gardens and sweet potato patches, but not in palm oil plantations. To assess the conservation status of
<taxonomicName id="4C254D02FF8BFFDFFD266777E119FE5C" authorityName="Sody" authorityYear="1933" box="[687,771,364,387]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF8BFFDFFD266777E119FE5C" box="[687,771,364,387]" italics="true" pageId="14" pageNumber="15">H. maxi</emphasis>
</taxonomicName>
, further research is required to determine population trends within protected areas and to evaluate whether this species can adapt to palm oil plantations, considering much of the regional habitat is undergoing land-use change. Likewise, additional field campaigns combined with habitat suitability modelling seem pivotal to evaluate the species area of occupancy. Conservation planning must also acknowledge that
<taxonomicName id="4C254D02FF8BFFDFFD79645CE2AEFDA1" authorityName="Sody" authorityYear="1933" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF8BFFDFFD79645CE2AEFDA1" italics="true" pageId="14" pageNumber="15">H. maxi</emphasis>
</taxonomicName>
contains two divergent lineages that, pending additional research, may deserve species-level recognition (see below), and an altered conservation status. In the meantime, we suggest these two lineages are treated independently by regional government agencies and NGOs.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF8BFFDFFF086538E08DFA90" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B9A3681FF8BFFDFFF086538E08DFA90" blockId="14.[129,778,802,1359]" pageId="14" pageNumber="15">
<emphasis id="B951EA93FF8BFFDFFF086538E2E8FCE5" box="[129,242,803,826]" italics="true" pageId="14" pageNumber="15">Comments:</emphasis>
Sumatran populations seem to be smaller on average than Peninsular Malaysia populations and are somewhat differentiated in the craniodental PCA but do overlap, although this could be due to ontogenetical sampling biases. Sumatran populations also differ from Peninsular Malaysia populations with cytochrome
<emphasis id="B951EA93FF8BFFDFFEBE65A4E34CFC08" box="[311,342,959,983]" italics="true" pageId="14" pageNumber="15">b p</emphasis>
-distances of
<emphasis id="B951EA93FF8BFFDFFE5565DBE3F6FC08" box="[476,492,960,983]" italics="true" pageId="14" pageNumber="15">c.</emphasis>
6% and have a private allele for the
<emphasis id="B951EA93FF8BFFDFFF4365C4E314FC29" box="[202,270,991,1014]" italics="true" pageId="14" pageNumber="15">BDNF</emphasis>
fragment, but not for
<emphasis id="B951EA93FF8BFFDFFE7965C4E03CFC29" box="[496,550,991,1014]" italics="true" pageId="14" pageNumber="15">GHR</emphasis>
. Further geographical and nuclear loci sampling, particularly across Sumatra (which is poorly represented in museums), will be required to assess whether Sumatran and Peninsular Malaysia’s populations represent two different species.
<bibRefCitation id="EFB44B70FF8BFFDFFE236247E0B4FBAB" author="Abd Wahab MF &amp; Motokawa M &amp; Anwaoali Khan FA" box="[426,686,1116,1140]" pageId="14" pageNumber="15" pagination="3125 - 41" refId="ref30291" refString="Abd Wahab MF, Motokawa M, Anwaoali Khan FA et al. Phvlogbnbtic oblationships and moophological vaoiation of Gvmnuobs (Galboicidab: Hylomys) foomGbntingHighlands, Pahang, Malavsia. SainsMalaysiana 2022; 51: 3125 - 41. hups: // doi. oog / 10.17576 / jsm- 2022 - 5110 - 01" type="journal article" year="2022">
Abd Wahab
<emphasis id="B951EA93FF8BFFDFFDA26247E04FFBAB" box="[555,597,1116,1140]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2022)
</bibRefCitation>
retained Tioman Island populations as a subspecies of
<taxonomicName id="4C254D02FF8BFFDFFDE06267E0DDFB4C" authorityName="Mullbo" authorityYear="1840" box="[617,711,1147,1171]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="suillus">
<emphasis id="B951EA93FF8BFFDFFDE06267E0DDFB4C" box="[617,711,1147,1171]" italics="true" pageId="14" pageNumber="15">H. suillus</emphasis>
</taxonomicName>
(
<emphasis id="B951EA93FF8BFFDFFD526267E2ACFB6D" italics="true" pageId="14" pageNumber="15">H. s. tionis</emphasis>
), but our results suggest it is found within the diversity of
<emphasis id="B951EA93FF8BFFDFFF0862A1E2CDFB0E" box="[129,215,1210,1233]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C254D02FF8BFFDFFF0862A1E2C9FB0E" authorityName="Sody" authorityYear="1933" box="[129,211,1210,1233]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maxi">H. maxi</taxonomicName>
.
</emphasis>
We do not believe this population deserves subspecieslevel recognition given that this taxon was described based on pelage coloration, which we now know, after looking at a much larger series than
<bibRefCitation id="EFB44B70FF8BFFDFFECD6303E3F2FAEF" author="Chasbn FN" box="[324,488,1304,1328]" pageId="14" pageNumber="15" pagination="1 - 209" refId="ref31447" refString="Chasbn FN. A handlist of Malavsian mammals (a svstbmatic list of thb mammals of thb Malav Pbninsula, Sumatoa, Boonbo and Java, including thb adjacbnt small islands). Bulletin of the Raffles Museum 1940; 15: 1 - 209." type="journal article" year="1940">Chasen (1940)</bibRefCitation>
, that it is highly prone to ecophenotypic, ontogenetic, and seasonal variation.
</paragraph>
</subSubSection>
</treatment>
</document>
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