196 lines
19 KiB
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196 lines
19 KiB
XML
<document id="E9FB2715A190E26B33F2C0FF7B3518B1" ID-DOI="10.11646/zootaxa.5477.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="12733301" ID-ZooBank="5F7FB3F5-BC72-40C3-BB2A-63C4C6F119CA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720790623788" checkinUser="plazi" docAuthor="Amorim, Danielle C. De A., Dos Santos Junior, José N. A., Carvalho- Filho, Fernando Da S., Silveira, Orlando T. & Carpenter, James M." docDate="2024" docId="038C1B72FFA4FFDFFF6E5C406A52FABC" docLanguage="en" docName="zootaxa.5477.5.2.pdf" docOrigin="Zootaxa 5477 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5477.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polybia scrobalis subsp. pronotalis Richards 1978" docType="treatment" docVersion="2" lastPageNumber="557" masterDocId="FFB5630AFFB0FFCBFFF95F5E6942FFD5" masterDocTitle="Taxonomic study of wasp species in the Polybia (Myrapetra) occidentalis group morphologically related to P. scrobalis and P. flavifrons, with description of a new species (Hymenoptera, Vespidae, Polistinae)" masterLastPageNumber="562" masterPageNumber="537" pageNumber="557" updateTime="1720791207115" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="822D2C02D999E7501C28B493EB875A76">Taxonomic study of wasp species in the Polybia (Myrapetra) occidentalis group morphologically related to P. scrobalis and P. flavifrons, with description of a new species (Hymenoptera, Vespidae, Polistinae)</mods:title>
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<mods:namePart id="F527099A8BE0B6F203058F0C5CFC898B">Amorim, Danielle C. De A.</mods:namePart>
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<mods:affiliation id="CEB2ED529889447C43F39C6C5C584FA5">Museu Paraense Emílio Goeldi-MPEG, Coordenação de Zoologia, Laboratório de Entomologia, Avenida Perimetral, 1901, Terra Firme, CEP: 66077 - 830, Belém, Pará, Brasil.</mods:affiliation>
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<mods:roleTerm id="BA24BEFC901EF087D2C19004490F3F89">Author</mods:roleTerm>
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<mods:namePart id="DF6B8B550004600B8DC87E8877A75C57">Dos Santos Junior, José N. A.</mods:namePart>
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<mods:affiliation id="4F6BE41760B0E26ADB055D3EC5154209">Universidade Federal do Sul e Sudeste do Pará-Folha 31, Quadra 07, Lote Especial, s / n. º-Nova Marabá, Marabá-PA, 68507 - 590, São Félix do Xingú, Pará, Brasil.</mods:affiliation>
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<mods:roleTerm id="A6BB715F0EC4A84103E14ADAD1472564">Author</mods:roleTerm>
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<mods:namePart id="1ABBC3BC30703880967498C89CEE80FE">Carvalho- Filho, Fernando Da S.</mods:namePart>
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<mods:affiliation id="C2C5DCAC468DD4BE3729D6E04996C5C7">Museu Paraense Emílio Goeldi-MPEG, Coordenação de Zoologia, Laboratório de Entomologia, Avenida Perimetral, 1901, Terra Firme, CEP: 66077 - 830, Belém, Pará, Brasil.</mods:affiliation>
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<mods:nameIdentifier id="563E9D8996C1B45E5C8AAB8085C83BE0" type="email">fernandofilho@museu-goeldi.br</mods:nameIdentifier>
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<mods:name id="7D81A21BDF0D9C180C5BB756C06E7F4C" type="personal">
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<mods:roleTerm id="3E30850F24FC3C22005A24A6844FBCB8">Author</mods:roleTerm>
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<mods:namePart id="BCFD5BBBC25DBED25061B8AE4BDEA5BC">Silveira, Orlando T.</mods:namePart>
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<mods:nameIdentifier id="4988C833214D3180781AAFD503883149" type="ORCID">0000-0002-5899-199X</mods:nameIdentifier>
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<mods:affiliation id="9BA1FA85063E4AA2585265524E88BAFA">Museu Paraense Emílio Goeldi-MPEG, Coordenação de Zoologia, Laboratório de Entomologia, Avenida Perimetral, 1901, Terra Firme, CEP: 66077 - 830, Belém, Pará, Brasil.</mods:affiliation>
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<mods:nameIdentifier id="AF41A9E9D20FA91EFCCB117839A9285C" type="email">orlando@museu-goeldi.br</mods:nameIdentifier>
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<mods:namePart id="5142B77C7A4F9FC853D8F7EF1B4FFA9B">Carpenter, James M.</mods:namePart>
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<mods:affiliation id="9B387A0CA03CF47613267B980817326D">American Museum of Natural History- 200 Central Park W, New York, NY 10024, USA.</mods:affiliation>
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<mods:typeOfResource id="6E866532F50221AFDEA1E4324EE259EC">text</mods:typeOfResource>
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<mods:title id="8B05192D0F0E71EC3D9D79530C928B06">Zootaxa</mods:title>
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<mods:part id="6E35C98B0A857E01BD25C3F1F5BE5913">
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<mods:date id="6BBDC6A1C9C599FA98C28C91E384BF49">2024</mods:date>
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<mods:number id="D343EF5FCF51BAE8602AC6D48B47316C">2024-07-11</mods:number>
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<mods:number id="AAFF337211AC9A9B93E1602B2FE4578B">5477</mods:number>
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<mods:number id="2EF564C6114E61290F37CDB2BFB8021D">5</mods:number>
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<mods:start id="4E170AE4AAE461B5F807C1A76DA74C02">537</mods:start>
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<mods:url id="6FD44A28CFD4A24EDD16AACFDE706EA2">http://dx.doi.org/10.11646/zootaxa.5477.5.2</mods:url>
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<treatment id="038C1B72FFA4FFDFFF6E5C406A52FABC" ID-DOI="http://doi.org/10.5281/zenodo.12733363" ID-Zenodo-Dep="12733363" LSID="urn:lsid:plazi:treatment:038C1B72FFA4FFDFFF6E5C406A52FABC" httpUri="http://treatment.plazi.org/id/038C1B72FFA4FFDFFF6E5C406A52FABC" lastPageNumber="557" pageId="20" pageNumber="557">
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<subSubSection id="C33FF9EFFFA4FFDFFF6E5C406BE8FCED" box="[151,682,798,825]" pageId="20" pageNumber="557" type="nomenclature">
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<paragraph id="8B9AAA64FFA4FFDFFF6E5C406BE8FCED" blockId="20.[151,682,798,825]" box="[151,682,798,825]" pageId="20" pageNumber="557">
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<heading id="D0D21D08FFA4FFDFFF6E5C406BE8FCED" bold="true" box="[151,682,798,825]" fontSize="11" level="1" pageId="20" pageNumber="557" reason="1">
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<taxonomicName id="4C25D1E7FFA4FFDFFF6E5C406BE8FCED" ID-CoL="5KKGJ" authority="Richards, 1978" authorityName="Richards" authorityYear="1978" box="[151,682,798,825]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="557" phylum="Arthropoda" rank="subSpecies" species="scrobalis" subSpecies="pronotalis">
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<emphasis id="B9517676FFA4FFDFFF6E5C406BE8FCED" bold="true" box="[151,682,798,825]" pageId="20" pageNumber="557">
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<emphasis id="B9517676FFA4FFDFFF6E5C4068A5FCEC" bold="true" box="[151,487,798,825]" italics="true" pageId="20" pageNumber="557">Polybia scrobalis pronotalis</emphasis>
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<bibRefCitation id="EFB4D795FFA4FFDFFE175C406BE8FCED" author="Richards, O. W." box="[494,682,798,824]" pageId="20" pageNumber="557" refId="ref26791" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FF9EFFFA4FFDFFF6E5C006B2CFC40" pageId="20" pageNumber="557" type="reference_group">
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<paragraph id="8B9AAA64FFA4FFDFFF6E5C006B2CFC40" blockId="20.[151,1436,862,917]" pageId="20" pageNumber="557">
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<treatmentCitationGroup id="AB358D4AFFA4FFDFFF6E5C006B2CFC40" pageId="20" pageNumber="557">
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<treatmentCitation id="0A848C75FFA4FFDFFF6E5C006BC0FCA3" author="Richards, O. W." box="[151,642,862,886]" page="40" pageId="20" pageNumber="557" year="1978">
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<taxonomicName id="4C25D1E7FFA4FFDFFF6E5C006BC0FCA3" ID-CoL="5KKGJ" authority="Richards, 1978: 40" authorityName="Richards" authorityPageNumber="40" authorityYear="1978" box="[151,642,862,886]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="557" phylum="Arthropoda" rank="subSpecies" species="scrobalis" subSpecies="pronotalis">
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<emphasis id="B9517676FFA4FFDFFF6E5C0068F5FCA3" box="[151,439,862,886]" italics="true" pageId="20" pageNumber="557">Polybia scrobalis pronotalis</emphasis>
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<materialsCitation id="3B4DA039FFA4FFDFFE475C006BC0FCA3" box="[446,642,862,886]" collectionCode="NHM" collectorName="M. Cooper & Rasmussen & Ansejo" country="Colombia" elevation="900" location="Valle" municipality="Cali" pageId="20" pageNumber="557" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<bibRefCitation id="EFB4D795FFA4FFDFFE475C006BC0FCA3" author="Richards, O. W." box="[446,642,862,886]" pageId="20" pageNumber="557" refId="ref26791" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978: 40</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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, 88;
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<typeStatus id="549E14C6FFA4FFDFFD4D5C006A50FCA3" box="[692,786,862,886]" pageId="20" pageNumber="557" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D2361EDFFA4FFDFFCE35C006A6FFCA0" box="[794,813,862,885]" count="1" pageId="20" pageNumber="557" type="female">♀</specimenCount>
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,
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<collectingCountry id="F332EAF4FFA4FFDFFCC35C006AE3FCA3" box="[826,929,862,886]" name="Colombia" pageId="20" pageNumber="557">Colombia</collectingCountry>
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,
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<location id="8EFAFCBFFFA4FFDFFC525C006AA0FCA3" LSID="urn:lsid:plazi:treatment:038C1B72FFA4FFDFFF6E5C406A52FABC:8EFAFCBFFFA4FFDFFC525C006AA0FCA3" box="[939,994,862,886]" country="Colombia" municipality="Cali" name="Valle" pageId="20" pageNumber="557">Valle</location>
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,
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<collectingMunicipality id="6BFE301EFFA4FFDFFC155C006D5BFCA3" box="[1004,1049,862,886]" pageId="20" pageNumber="557">Cali</collectingMunicipality>
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,
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<quantity id="4CDD0781FFA4FFDFFBDD5C006D23FCA3" box="[1060,1121,862,886]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="557" unit="m" value="900.0">
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<elevation id="00084D57FFA4FFDFFBDD5C006D23FCA3" box="[1060,1121,862,886]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="557" unit="m" value="900.0">900m</elevation>
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</quantity>
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, 20.iv.72 (
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<collectorName id="26D0CFB2FFA4FFDFFB325C006C7EFCA3" box="[1227,1340,862,886]" pageId="20" pageNumber="557">M. Cooper</collectorName>
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) (NHM), examined.
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<collectorName id="26D0CFB2FFA4FFDFFECF5C2368EAFC41" box="[310,424,893,916]" pageId="20" pageNumber="557">Rasmussen</collectorName>
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&
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<collectorName id="26D0CFB2FFA4FFDFFE3E5C226B4DFC41" box="[455,527,892,916]" pageId="20" pageNumber="557">Ansejo</collectorName>
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2009: 48
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FF9EFFFA4FFDFFF6E5C9C6D39FBB8" pageId="20" pageNumber="557" type="diagnosis">
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<paragraph id="8B9AAA64FFA4FFDFFF6E5C9C6D72FBF0" blockId="20.[151,1436,962,1385]" pageId="20" pageNumber="557">
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<emphasis id="B9517676FFA4FFDFFF6E5C9C6850FC09" bold="true" box="[151,274,962,988]" pageId="20" pageNumber="557">Diagnosis.</emphasis>
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FEMALE. Very similar to the nominotypical form; occipital-genal margin distinct on the sides; pronotal carina absent; prominence in front of the fovea moderately elevated; more extensive yellow marks, as on the anterior margin of the pronotum and on the second metasomal segment. MALE. Unknown
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</paragraph>
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<paragraph id="8B9AAA64FFA4FFDFFF3E5B706D39FBB8" blockId="20.[151,1436,962,1385]" pageId="20" pageNumber="557">
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<emphasis id="B9517676FFA4FFDFFF3E5B7068F5FB9D" bold="true" box="[199,439,1070,1096]" pageId="20" pageNumber="557">Examined material.</emphasis>
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<materialsCitation id="3B4DA039FFA4FFDFFE3B5B716CDAFB9C" box="[450,1432,1071,1097]" collectionCode="NHM" collectorName="Valle & Cali & M. Cooper" country="Colombia" location="COLOMBIA" pageId="20" pageNumber="557" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="549E14C6FFA4FFDFFE3B5B716B68FB9C" box="[450,554,1071,1097]" pageId="20" pageNumber="557" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D2361EDFFA4FFDFFDCC5B716B08FB9C" box="[565,586,1071,1097]" count="1" pageId="20" pageNumber="557" type="female">♀</specimenCount>
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,
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<collectingCountry id="F332EAF4FFA4FFDFFDA55B716BB4FB9C" box="[604,758,1071,1097]" name="Colombia" pageId="20" pageNumber="557">COLOMBIA</collectingCountry>
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:
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<collectorName id="26D0CFB2FFA4FFDFFCFA5B716A7DFB9C" box="[771,831,1071,1097]" pageId="20" pageNumber="557">Valle</collectorName>
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,
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<collectorName id="26D0CFB2FFA4FFDFFCB75B716A3DFB9C" box="[846,895,1071,1097]" pageId="20" pageNumber="557">Cali</collectorName>
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, 20. iv.72 (
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<collectorName id="26D0CFB2FFA4FFDFFBFE5B716DC4FB9C" box="[1031,1158,1071,1097]" pageId="20" pageNumber="557">M. Cooper</collectorName>
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) (
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<collectionCode id="ED3432A1FFA4FFDFFB595B716DA7FB9C" box="[1184,1253,1071,1097]" pageId="20" pageNumber="557">NHM</collectionCode>
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) (photographs)
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</materialsCitation>
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;
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<materialsCitation id="3B4DA039FFA4FFDFFF6E5B0D6D35FBB8" box="[151,1143,1107,1133]" collectionCode="MNHN" collectorName="R. Benoist" country="Ecuador" location="Tandapi" pageId="20" pageNumber="557" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
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<typeStatus id="549E14C6FFA4FFDFFF6E5B0D69BEFBB8" box="[151,252,1107,1133]" pageId="20" pageNumber="557" type="paratype">Paratype</typeStatus>
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,
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<collectingCountry id="F332EAF4FFA4FFDFFEFE5B0D68D6FBB8" box="[263,404,1107,1133]" name="Ecuador" pageId="20" pageNumber="557">EQUADOR</collectingCountry>
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:
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<collectingRegion id="49E16486FFA4FFDFFE665B0D689CFBB8" box="[415,478,1107,1133]" country="Ecuador" name="Napo" pageId="20" pageNumber="557">Napo</collectingRegion>
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,
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<location id="8EFAFCBFFFA4FFDFFE105B0D6B04FBB8" LSID="urn:lsid:plazi:treatment:038C1B72FFA4FFDFFF6E5C406A52FABC:8EFAFCBFFFA4FFDFFE105B0D6B04FBB8" box="[489,582,1107,1133]" country="Ecuador" name="Tandapi" pageId="20" pageNumber="557" stateProvince="Napo">Tandapi</location>
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,
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<specimenCount id="9D2361EDFFA4FFDFFDA85B0D6B36FBB8" box="[593,628,1107,1133]" count="1" pageId="20" pageNumber="557" type="female">1♀</specimenCount>
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2.vii.31 (
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<collectorName id="26D0CFB2FFA4FFDFFD195B0D6A1AFBB8" box="[736,856,1107,1133]" pageId="20" pageNumber="557">R. Benoist</collectorName>
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) (
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<collectionCode id="ED3432A1FFA4FFDFFC965B0D6A8AFBB8" box="[879,968,1107,1133]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="557">MNHN</collectionCode>
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) (photographs)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FF9EFFFA4FFDFFF3E5B286B0BFB44" box="[199,585,1142,1169]" pageId="20" pageNumber="557" type="distribution">
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<paragraph id="8B9AAA64FFA4FFDFFF3E5B286B0BFB44" blockId="20.[151,1436,962,1385]" box="[199,585,1142,1169]" pageId="20" pageNumber="557">
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|
<emphasis id="B9517676FFA4FFDFFF3E5B286823FB45" bold="true" box="[199,353,1142,1168]" pageId="20" pageNumber="557">Distribution.</emphasis>
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|
<collectingCountry id="F332EAF4FFA4FFDFFE9E5B29689AFB44" box="[359,472,1143,1169]" name="Colombia" pageId="20" pageNumber="557">Colômbia</collectingCountry>
|
|
,
|
|
<collectingCountry id="F332EAF4FFA4FFDFFE1D5B296B07FB44" box="[484,581,1143,1169]" name="Ecuador" pageId="20" pageNumber="557">Equador</collectingCountry>
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|
.
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</paragraph>
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|
</subSubSection>
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|
<subSubSection id="C33FF9EFFFA4FFDFFF3E5BC56A52FABC" pageId="20" pageNumber="557" type="discussion">
|
|
<paragraph id="8B9AAA64FFA4FFDFFF3E5BC56A52FABC" blockId="20.[151,1436,962,1385]" pageId="20" pageNumber="557">
|
|
<emphasis id="B9517676FFA4FFDFFF3E5BC56879FB60" bold="true" box="[199,315,1179,1205]" pageId="20" pageNumber="557">Remarks.</emphasis>
|
|
<bibRefCitation id="EFB4D795FFA4FFDFFEBD5BC568B9FB60" author="Richards, O. W." box="[324,507,1179,1205]" pageId="20" pageNumber="557" refId="ref26791" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards (1978)</bibRefCitation>
|
|
separated this form from typical
|
|
<taxonomicName id="4C25D1E7FFA4FFDFFC855BC56AA0FB60" authorityName="Richards" authorityYear="1970" box="[892,994,1179,1205]" class="Insecta" family="Eumenidae" genus="Polybia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="557" phylum="Arthropoda" rank="species" species="scrobalis">
|
|
<emphasis id="B9517676FFA4FFDFFC855BC56AA0FB60" box="[892,994,1179,1205]" italics="true" pageId="20" pageNumber="557">scrobalis</emphasis>
|
|
</taxonomicName>
|
|
mainly based on size (with the ranges overlapping), and thickness of genal margin and pronotal prominence. Such differences could not be observed in the examined type material in the NHM, except for those on color, since the
|
|
<typeStatus id="549E14C6FFA4FFDFFC065BBD6D23FB28" box="[1023,1121,1251,1277]" pageId="20" pageNumber="557" type="holotype">holotype</typeStatus>
|
|
of
|
|
<emphasis id="B9517676FFA4FFDFFB705BBD6C73FB28" box="[1161,1329,1251,1277]" italics="true" pageId="20" pageNumber="557">P. s. pronotalis</emphasis>
|
|
is in fact more extensively marked with yellow spots. Importantly, a
|
|
<typeStatus id="549E14C6FFA4FFDFFCDE5A596AC5FAF4" box="[807,903,1287,1313]" pageId="20" pageNumber="557" type="paratype">paratype</typeStatus>
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|
at the Paris Museum is much darker, with fewer yellow marks than the
|
|
<typeStatus id="549E14C6FFA4FFDFFE6A5A7568BBFA90" box="[403,505,1323,1349]" pageId="20" pageNumber="557" type="holotype">holotype</typeStatus>
|
|
. The distributions of the two subspecies seem to be disjunct, with all known records of
|
|
<emphasis id="B9517676FFA4FFDFFF4F5A11681EFABC" box="[182,348,1359,1385]" italics="true" pageId="20" pageNumber="557">P. s. pronotalis</emphasis>
|
|
coming from western Andean regions.
|
|
</paragraph>
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|
</subSubSection>
|
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</treatment>
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</document> |