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<document id="6B8310D49CB992EFA8376255597235D9" ID-DOI="10.11646/zootaxa.4032.1.1" ID-GBIF-Dataset="aeef3f9e-8a5d-4bb0-b0d9-908830c0f6a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="289744" ID-ZooBank="CB9720EB-7BB7-4199-A835-A3266B0DDA6B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461288662073" checkinUser="plazi" docAuthor="Soto-Adames, Felipe N." docDate="2015" docId="038AC52EFFDCA464FF3DFD8EFC127FAF" docLanguage="en" docName="zt04032p041.pdf" docOrigin="Zootaxa 4032 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trogolaphysa paracarpenteri Soto-Adames, 2015, sp. nov." docType="treatment" docVersion="5" lastPageNumber="32" masterDocId="FFB3BD56FFC0A47BFFAAFF8EFF887F57" masterDocTitle="The dorsal chaetotaxy of first instar Trogolaphysa jataca, with description of twelve new species of Neotropical Trogolaphysa (Hexapoda: Collembola: Paronellidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="29" updateTime="1698615735623" updateUser="plazi">
<mods:mods id="35561BC72924E5ACF8BE774045C792FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CD4684773C947F3C6175B69E293553F6">The dorsal chaetotaxy of first instar Trogolaphysa jataca, with description of twelve new species of Neotropical Trogolaphysa (Hexapoda: Collembola: Paronellidae)</mods:title>
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<mods:namePart id="699A621A25438CCE65366659C6F9AC9C">Soto-Adames, Felipe N.</mods:namePart>
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<mods:title id="3D6B08318C2D3CFE3D59778FEE2A5D53">Zootaxa</mods:title>
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<mods:part id="FCB7CA91CD5864F653CB59EB6F9FB9E5">
<mods:date id="AD28C86B583BDB4AA49093DF6FAE9516">2015</mods:date>
<mods:detail id="DB35D827FF0F676574666DAD721F5053" type="volume">
<mods:number id="629A56D2DF1EF9B1EE613FEF82633C9A">4032</mods:number>
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<mods:identifier id="B3A3C75389923DFC0D4B61801BB8F53B" type="DOI">10.11646/zootaxa.4032.1.1</mods:identifier>
<mods:identifier id="33F48C3359D1861FCFEEC5B04FD3374E" type="GBIF-Dataset">aeef3f9e-8a5d-4bb0-b0d9-908830c0f6a5</mods:identifier>
<mods:identifier id="DA26A88A7DF45941E008CC37D605BAA1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038AC52EFFDCA464FF3DFD8EFC127FAF" ID-DOI="http://doi.org/10.5281/zenodo.6096452" ID-GBIF-Taxon="127693708" ID-Zenodo-Dep="6096452" LSID="urn:lsid:plazi:treatment:038AC52EFFDCA464FF3DFD8EFC127FAF" httpUri="http://treatment.plazi.org/id/038AC52EFFDCA464FF3DFD8EFC127FAF" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<subSubSection id="C33927B3FFDCA467FF3DFD8EFE267D6C" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B9C7438FFDCA467FF3DFD8EFDD37D4D" blockId="28.[151,603,512,571]" box="[151,603,512,538]" pageId="28" pageNumber="29">
<heading id="D0D4C354FFDCA467FF3DFD8EFDD37D4D" bold="true" box="[151,603,512,538]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B957A82AFFDCA467FF3DFD8EFDD37D4D" bold="true" box="[151,603,512,538]" pageId="28" pageNumber="29">
<taxonomicName id="4C230FBBFFDCA467FF3DFD8EFE7E7D4D" box="[151,502,512,538]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FF3DFD8EFE7E7D4D" bold="true" box="[151,502,512,538]" italics="true" pageId="28" pageNumber="29">Trogolaphysa paracarpenteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2641551FFDCA467FE57FD8EFDD37D4D" box="[509,603,512,538]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C7438FFDCA467FF3DFDACFE267D6C" blockId="28.[151,603,512,571]" box="[151,430,546,571]" pageId="28" pageNumber="29">
<figureCitation id="131868BDFFDCA467FF3DFDACFF6C7D6C" box="[151,228,546,571]" captionStart="FIGURE 4" captionStartId="10.[151,249,1596,1618]" captionTargetBox="[160,1421,198,1561]" captionTargetId="figure@10.[151,1436,193,1575]" captionTargetPageId="10" captionText="FIGURE 4. Trogolaphysa spp., habitus. A, T. ocellata sp. nov. lateral view; B, T. ocellata sp. nov. dorsal view of body; C, T. balteata sp. nov., lateral view; D, T. balteata sp. nov., detail of abdomen 4 color pattern; E, T. paracarpenteri sp. nov.; F, T. octosetosa sp. nov." httpUri="https://zenodo.org/record/289748/files/figure.png" pageId="28" pageNumber="29">Figs. 4</figureCitation>
E, 16A–H
<tableCitation id="C6A14183FFDCA467FEF6FDACFE267D6C" box="[348,430,546,571]" captionStart="TABLE 4" captionStartId="29.[125,210,612,634]" captionTargetBox="[125,1822,669,1596]" captionTargetPageId="29" captionText="TABLE 4. Diagnostic table of Neotropical species of Trogolaphysa with 6 – 8 eyes and two (A 0, A 2) dorsal anterior head macrochaetae. Mc = macrochaeta." httpUri="http://table.plazi.org/id/DF5C24B0FFDDA466FFD7FDEAF9287D2D" pageId="28" pageNumber="29" tableUuid="DF5C24B0FFDDA466FFD7FDEAF9287D2D">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33927B3FFDCA467FF3DFDE4FCF17DD5" box="[151,889,618,643]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B9C7438FFDCA467FF3DFDE4FCF17DD5" blockId="28.[151,1437,618,2011]" box="[151,889,618,643]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF3DFDE4FE967DD4" bold="true" box="[151,286,618,643]" pageId="28" pageNumber="29">Etymology.</emphasis>
The epithet refers to the similarity with
<taxonomicName id="4C230FBBFFDCA467FD48FDE2FCF97DD4" box="[738,881,620,643]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FD48FDE2FCF97DD4" box="[738,881,620,643]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33927B3FFDCA467FF6DFD00FD707DB8" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B9C7438FFDCA467FF6DFD00FD707DB8" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFD00FE3E7DF0" bold="true" box="[199,438,654,679]" pageId="28" pageNumber="29">Material examined.</emphasis>
<typeStatus id="5498CA9AFFDCA467FE68FD00FDA27DF0" box="[450,554,654,679]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male in preparation,
<collectingCountry id="F33434A8FFDCA467FC9AFD01FC2A7DF0" box="[816,930,655,679]" name="Mexico" pageId="28" pageNumber="29">MEXICO</collectingCountry>
, Chiapas, Ocosingo Valley, Finca El Real, floodplain debris,
<date id="FF9D52F8FFDCA467FED9FD3CFD8B7D9C" box="[371,515,690,715]" pageId="28" pageNumber="29" value="1950-07-01" valueMax="1950-07-07" valueMin="1950-07-01">1–7.vii.1950</date>
, C. &amp; M. Goodnight and J. Stannard, coll.,
<collectionCode id="ED32ECFDFFDCA467FB84FD3DFBF87D9C" LSID="urn:lsid:biocol.org:col:34797" box="[1070,1136,691,715]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="28" pageNumber="29">INHS</collectionCode>
579,830; 2
<typeStatus id="5498CA9AFFDCA467FAA6FD3DFAFF7D9C" box="[1292,1399,691,715]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
in preparation and 20, mostly small juveniles, in alcohol.
</paragraph>
</subSubSection>
<subSubSection id="C33927B3FFDCA467FF6DFD74FE8C7950" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9C7438FFDCA467FF6DFD74FDBC7C45" blockId="28.[151,1437,618,2011]" box="[199,564,762,787]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFD74FED17C44" bold="true" box="[199,345,762,787]" pageId="28" pageNumber="29">Description.</emphasis>
Size up to
<quantity id="4CDBD9DDFFDCA467FE7CFD75FDA77C45" box="[470,559,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="mm" value="1.9">1.9 mm</quantity>
.
</paragraph>
<paragraph id="8B9C7438FFDCA467FF6DFC90FC707C60" blockId="28.[151,1437,618,2011]" box="[199,1016,798,823]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFC90FEE27C60" bold="true" box="[199,362,798,823]" pageId="28" pageNumber="29">Color pattern</emphasis>
. Uniformly blue, sometimes with lighter stripes (
<figureCitation id="131868BDFFDCA467FC3EFC90FC507C60" box="[916,984,798,823]" captionStart="FIGURE 4" captionStartId="10.[151,249,1596,1618]" captionTargetBox="[160,1421,198,1561]" captionTargetId="figure@10.[151,1436,193,1575]" captionTargetPageId="10" captionText="FIGURE 4. Trogolaphysa spp., habitus. A, T. ocellata sp. nov. lateral view; B, T. ocellata sp. nov. dorsal view of body; C, T. balteata sp. nov., lateral view; D, T. balteata sp. nov., detail of abdomen 4 color pattern; E, T. paracarpenteri sp. nov.; F, T. octosetosa sp. nov." httpUri="https://zenodo.org/record/289748/files/figure.png" pageId="28" pageNumber="29">Fig. 4</figureCitation>
E).
</paragraph>
<paragraph id="8B9C7438FFDCA467FF6DFCCCFF6A7B64" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFCCCFE867C0C" bold="true" box="[199,270,834,859]" pageId="28" pageNumber="29">Head.</emphasis>
Antennae up to 2.9x as long as head diagonal. Ant. 4 subapical sense organ weakly capitate, almost reaching upper margin of pit; guard sensillum not seen. Main sensilla in Ant. 3 sense organ globose (
<figureCitation id="131868BDFFDCA467FA8DFCE8FAF37C28" box="[1319,1403,870,895]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
C). Eyes 8+8, eyes G and H greatly reduced; eye well with 4–5 ciliate chaetae (
<figureCitation id="131868BDFFDCA467FC54FC04FBD97CF4" box="[1022,1105,906,931]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
A). Head dorsally (
<figureCitation id="131868BDFFDCA467FA87FC04FAF67CF4" box="[1325,1406,906,931]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
A) with 2 anterior (A0, A2) and 2 posterior (Pa5, Pm3) macrochaetae. Pre-labral chaetae ciliate. Distal margin of labrum with 1+1 spines with partially fused base (
<figureCitation id="131868BDFFDCA467FD6DFC5CFC927CBC" box="[711,794,978,1003]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
B). Labial triangle as M1M2rEL1L2A1–5: chaeta r short, thick and smooth; all other posterior chaetae long and ciliate. Post-labial field with 5–6 chaetae between columns I and
<collectingCountry id="F33434A8FFDCA467FF6CFB95FF567B64" box="[198,222,1051,1075]" name="Iceland" pageId="28" pageNumber="29">O</collectingCountry>
.
</paragraph>
<paragraph id="8B9C7438FFDCA467FF6DFBB0FD917BB0" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFBB0FE837B00" bold="true" box="[199,267,1086,1111]" pageId="28" pageNumber="29">Body.</emphasis>
Macrochaeta formula as 60/0243+0+6. Mesothorax with 1 anterior (a5) and 5 posterior (p3 complex) macrochaetae (
<figureCitation id="131868BDFFDCA467FEEAFBECFE1A7B2C" box="[320,402,1122,1147]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
E). Metathorax without macrochaetae. Abd. 1 chaeta a6 obscured, apparently absent. Abd. 4 (
<figureCitation id="131868BDFFDCA467FF35FB08FF757BC8" box="[159,253,1158,1183]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Figs. 16</figureCitation>
G–H) with pseudopore in field posterior to bothriotrix T4; with inner macrochaetae A3, A5 and B5; macrochaeta A5 closer to pseudopore than to macrochaeta B5; laterally with 6 macrochaetae; posterior chaetae present but actual number unclear.
</paragraph>
<paragraph id="8B9C7438FFDCA467FF6DFB7CFEF27ACD" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFB7CFE8E7A5C" bold="true" box="[199,262,1266,1291]" pageId="28" pageNumber="29">Legs.</emphasis>
Trochanteral organ with at least 15 chaetae. Metathoracic claw complex as in
<figureCitation id="131868BDFFDCA467FB1BFB7CFB777A5C" box="[1201,1279,1266,1291]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig 16</figureCitation>
F; tenent hair spatulate, 0.92x as long as unguiculus; unguis with 3 inner teeth; basal paired teeth drawn out, needle-shape, one basal tooth longer than other; unpaired tooth as long or shorter than shortest basal tooth; inner teeth spread as 38%, 42%, 64%; dorsal tooth small, barely projecting; lateral teeth inconspicuous. Unguiculus lanceolate, smooth or very weakly serrate.
</paragraph>
<paragraph id="8B9C7438FFDCA467FF6DFA28FE8C7950" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DFA28FEA47AE8" bold="true" box="[199,300,1446,1471]" pageId="28" pageNumber="29">Furcula.</emphasis>
Dens with 2 rows of ciliate spines: inner row with 21 spines; outer row with 17 spines. Mucro with 4 teeth, basal teeth sequential (
<figureCitation id="131868BDFFDCA467FE70FA44FDA37AB4" box="[474,555,1482,1507]" captionStart="FIGURE 16" captionStartId="30.[151,249,1937,1959]" captionTargetBox="[225,1349,202,1890]" captionTargetId="figure@30.[206,1370,193,1905]" captionTargetPageId="30" captionText="FIGURE 16. Trogolaphysa paracapenteri sp. nov. A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual." httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
D); mucro 2.1x as long as distal end of dens and 0.96x as long as inner edge of hind legs.
</paragraph>
</subSubSection>
<subSubSection id="C33927B3FFDCA464FF6DF99CFC127FAF" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B9C7438FFDCA467FF6DF99CFDB6788C" blockId="28.[151,1437,618,2011]" pageId="28" pageNumber="29">
<emphasis id="B957A82AFFDCA467FF6DF99CFEB3797C" bold="true" box="[199,315,1554,1579]" pageId="28" pageNumber="29">Remarks.</emphasis>
<taxonomicName id="4C230FBBFFDCA467FEE9F99CFD06797C" box="[323,654,1554,1579]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FEE9F99CFD06797C" box="[323,654,1554,1579]" italics="true" pageId="28" pageNumber="29">Trogolaphysa paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FD3FF99CFD78797C" bold="true" box="[661,752,1554,1579]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FD3FF99CFD78797C" box="[661,752,1554,1579]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only New World member of the genus with 6–8 eyes, inner labral papilla fused into a short pedicel and mesothorax p3 complex with 5 macrochaetae. This species differs from
<taxonomicName id="4C230FBBFFDCA467FF7EF9D2FEEB7924" box="[212,355,1628,1651]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FF7EF9D2FEEB7924" box="[212,355,1628,1651]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
in the original sense of
<bibRefCitation id="EFB209C9FFDCA467FDDAF9D4FC8A7924" author="Denis" box="[624,770,1626,1651]" pageId="28" pageNumber="29" refString="Denis, J. R. (1925) Sur les Collemboles du museum de Paris (2 e partie). Annales de la Societe Entomologique de France, 94, 261 - 289." type="journal article" year="1925">Denis (1925)</bibRefCitation>
in color pattern (Denis species is yellowish brown whereas
<taxonomicName id="4C230FBBFFDCA467FF3DF9F1FED679C1" box="[151,350,1663,1686]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FF3DF9F1FED679C1" box="[151,350,1663,1686]" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FECCF9F0FE4A79C0" bold="true" box="[358,450,1662,1687]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FECCF9F0FE4A79C0" box="[358,450,1662,1687]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is blue), however there are so many gaps in the 1925 description of
<taxonomicName id="4C230FBBFFDCA467FB7DF9F1FAE179C1" box="[1239,1385,1663,1686]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FB7DF9F1FAE179C1" box="[1239,1385,1663,1686]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
that additional meaningful comparisons are impossible. The new species is also similar to specimens identified by
<bibRefCitation id="EFB209C9FFDCA467FF3DF948FEBB7988" author="Yoshii" box="[151,307,1734,1759]" pageId="28" pageNumber="29" refString="Yoshii, R. (1988) Paronellid Collembola from caves of Central and South America collected by P. Strinati. Revue suisse Zoologie, 95, 449 - 459. http: // dx. doi. org / 10.5962 / bhl. part. 82022" type="journal article" year="1988">Yoshii (1988)</bibRefCitation>
as
<taxonomicName id="4C230FBBFFDCA467FEF7F949FE787989" box="[349,496,1735,1758]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FEF7F949FE787989" box="[349,496,1735,1758]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
. The two forms share the color pattern and anterior head macrochaetae (the only aspect of the dorsal chaetotaxy described by Yoshii), but differ in the ornamentation of pre-labral chaetae (smooth in
<taxonomicName id="4C230FBBFFDCA467FF1DF881FEC27871" box="[183,330,1807,1830]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FF1DF881FEC27871" box="[183,330,1807,1830]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
, ciliate in
<taxonomicName id="4C230FBBFFDCA467FE61F881FD1B7871" box="[459,659,1807,1830]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FE61F881FD1B7871" box="[459,659,1807,1830]" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FD34F880FD737870" bold="true" box="[670,763,1806,1831]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FD34F880FD737870" box="[670,763,1806,1831]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), shape of distal labral spines (separated in
<taxonomicName id="4C230FBBFFDCA467FAABF881FA1C7871" box="[1281,1428,1807,1830]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FAABF881FA1C7871" box="[1281,1428,1807,1830]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
, basally fused in
<taxonomicName id="4C230FBBFFDCA467FECFF8BAFDB9781C" box="[357,561,1844,1867]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FECFF8BAFDB9781C" box="[357,561,1844,1867]" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FDEAF8BCFD29781C" bold="true" box="[576,673,1842,1867]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FDEAF8BCFD29781C" box="[576,673,1842,1867]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), shape of ungual teeth (subequal in
<taxonomicName id="4C230FBBFFDCA467FBDFF8BAFB61781C" box="[1141,1257,1844,1867]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FBDFF8BAFB61781C" box="[1141,1257,1844,1867]" italics="true" pageId="28" pageNumber="29">carpenteri</emphasis>
</taxonomicName>
, one larger in
<taxonomicName id="4C230FBBFFDCA467FF3DF8D9FEC97839" box="[151,321,1879,1902]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FF3DF8D9FEC97839" box="[151,321,1879,1902]" italics="true" pageId="28" pageNumber="29">paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FEE7F8D8FE257838" bold="true" box="[333,429,1878,1903]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FEE7F8D8FE257838" box="[333,429,1878,1903]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and shape of unguiculus (narrow lanceolate in
<taxonomicName id="4C230FBBFFDCA467FC52F8D9FB077839" box="[1016,1167,1879,1902]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDCA467FC52F8D9FB077839" box="[1016,1167,1879,1902]" italics="true" pageId="28" pageNumber="29">T. carpenteri</emphasis>
</taxonomicName>
, wide lanceolate in
<taxonomicName id="4C230FBBFFDCA467FA2DF8D9FEC978C4" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FA2DF8D9FEC978C4" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FEEDF8F4FE1778C4" bold="true" box="[327,415,1914,1939]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FEEDF8F4FE1778C4" box="[327,415,1914,1939]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The new species is also similar to
<taxonomicName id="4C230FBBFFDCA467FC9BF8F2FB2F78C4" authority="Mari Mutt, 1987" authorityName="Mari Mutt" authorityYear="1987" box="[817,1191,1914,1939]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B957A82AFFDCA467FC9BF8F2FC5B78C4" box="[817,979,1914,1939]" italics="true" pageId="28" pageNumber="29">T. subterranea</emphasis>
(
<bibRefCitation id="EFB209C9FFDCA467FC4BF8F4FB1778C4" author="Mari" box="[993,1183,1914,1939]" pageId="28" pageNumber="29" refString="Mari Mutt, J. (1987) Puerto Rican species of Paronellidae (Insecta: Collembola). Caribbean Journal of Science, 23, 400 - 416." type="journal article" year="1987">
<collectingCountry id="F33434A8FFDCA467FC4BF8F4FB9178C4" box="[993,1049,1914,1939]" name="Serbia" pageId="28" pageNumber="29">Mari</collectingCountry>
Mutt, 1987
</bibRefCitation>
)
</taxonomicName>
, but this species lacks distal labral spines (present in
<taxonomicName id="4C230FBBFFDCA467FE50F811FD4A78E1" box="[506,706,1951,1974]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FE50F811FD4A78E1" box="[506,706,1951,1974]" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FD61F810FCAF78E0" bold="true" box="[715,807,1950,1975]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FD61F810FCAF78E0" box="[715,807,1950,1975]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and the paired ungual teeth are keel-shaped (needleshaped in
<taxonomicName id="4C230FBBFFDCA467FEA2F84AFE46788C" box="[264,462,1988,2011]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDCA467FEA2F84AFE46788C" box="[264,462,1988,2011]" italics="true" pageId="28" pageNumber="29">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDCA467FE7EF84CFDA6788C" bold="true" box="[468,558,1986,2011]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A2641551FFDCA467FE7EF84CFDA6788C" box="[468,558,1986,2011]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<caption id="DF5C24B0FFDDA466FFD7FDEAF9287D2D" ID-Table-UUID="DF5C24B0FFDDA466FFD7FDEAF9287D2D" box="[125,1696,612,634]" httpUri="http://table.plazi.org/id/DF5C24B0FFDDA466FFD7FDEAF9287D2D" pageId="29" pageNumber="30" targetBox="[125,1822,669,1596]" targetIsTable="true" targetPageId="29">
<paragraph id="8B9C7438FFDDA466FFD7FDEAF9287D2D" blockId="29.[125,1696,612,634]" box="[125,1696,612,634]" pageId="29" pageNumber="30">
<emphasis id="B957A82AFFDDA466FFD7FDEAFF637D2D" bold="true" box="[125,235,612,634]" pageId="29" pageNumber="30">TABLE 4.</emphasis>
Diagnostic table of Neotropical species of
<taxonomicName id="4C230FBBFFDDA466FD0AFDEAFCA57D2D" box="[672,813,612,634]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B957A82AFFDDA466FD0AFDEAFCA57D2D" box="[672,813,612,634]" italics="true" pageId="29" pageNumber="30">Trogolaphysa</emphasis>
</taxonomicName>
with 6–8 eyes and two (A0, A2) dorsal anterior head macrochaetae. Mc= macrochaeta.
</paragraph>
</caption>
<paragraph id="8B9C7438FFDDA466FFD7FD13F97F796B" pageId="29" pageNumber="30">
<table id="F9238698FFDD5B84FFD7FD13F896796B" box="[125,1822,669,1596]" gridcols="8" gridrows="8" pageId="29" pageNumber="30">
<tr id="3513767AFFDD5B84FFD7FD13F8967C15" box="[125,1822,669,834]" gridrow="0" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FD13FE197C15" box="[125,401,669,834]" gridcol="0" gridrow="0" pageId="29" pageNumber="30">
Characters Species
<taxonomicName id="4C230FBBFFDDA466FFD7FC9EFEBD7C15" authority="Mari Mutt, 1987" authorityName="Mari Mutt" authorityYear="1987" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B957A82AFFDDA466FFD7FC9EFE997C71" box="[125,273,784,806]" italics="true" pageId="29" pageNumber="30">T. subterranea</emphasis>
(Mari Mutt, 1987)
</taxonomicName>
</th>
<th id="76C21F06FFDD5B84FE62FD13FDAB7C15" box="[456,547,669,834]" gridcol="1" gridrow="0" pageId="29" pageNumber="30">Prelabral Chaetae ciliate</th>
<th id="76C21F06FFDD5B84FDFAFD13FD327C15" box="[592,698,669,834]" gridcol="2" gridrow="0" pageId="29" pageNumber="30">Labral Spines absent</th>
<th id="76C21F06FFDD5B84FCACFD13FCEA7C15" box="[774,866,669,834]" gridcol="3" gridrow="0" pageId="29" pageNumber="30">Thorax 2 Mc p3 group 6</th>
<th id="76C21F06FFDD5B84FC24FD13FC627C15" box="[910,1002,669,834]" gridcol="4" gridrow="0" pageId="29" pageNumber="30">Thorax 3 Mc 0</th>
<th id="76C21F06FFDD5B84FBBEFD13FA517C15" box="[1044,1497,669,834]" gridcol="5" gridrow="0" pageId="29" pageNumber="30">Abdomen 4 Inner Shape Position Inner Ungual Ungual Basal Mc Teeth Teeth Ungual Teeth 3 3 keel basal A3,A5,B5 half</th>
<th id="76C21F06FFDD5B84F9B6FD13F9087C15" box="[1564,1664,669,834]" gridcol="6" gridrow="0" pageId="29" pageNumber="30">Position Unpaired Ungual Tooth middle quarter</th>
<th id="76C21F06FFDD5B84F912FD13F8967C15" box="[1720,1822,669,834]" gridcol="7" gridrow="0" pageId="29" pageNumber="30">Mucro Teeth 4 sequential</th>
</tr>
<tr id="3513767AFFDD5B84FFD7FCEAF8967CE4" box="[125,1822,868,947]" gridrow="1" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FCEAFE197CE4" box="[125,401,868,947]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FCEBFE897C2D" box="[125,257,869,890]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="octosetosa" status="sp. nov.">
<emphasis id="B957A82AFFDDA466FFD7FCEBFE897C2D" box="[125,257,869,890]" italics="true" pageId="29" pageNumber="30">T. octosetosa</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDDA466FFD7FC0FFF477CC0" bold="true" box="[125,207,897,919]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2641551FFDDA466FFD7FC0FFF477CC0" box="[125,207,897,919]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76C21F06FFDD5B84FE62FCEAFDAB7CE4" box="[456,547,868,947]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">weakly ciliate</td>
<td id="76C21F06FFDD5B84FDFAFCEAFD327CE4" box="[592,698,868,947]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">separated</td>
<td id="76C21F06FFDD5B84FCACFCEAFCEA7CE4" box="[774,866,868,947]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">5</td>
<td id="76C21F06FFDD5B84FC24FCEAFC627CE4" box="[910,1002,868,947]" gridcol="4" gridrow="1" pageId="29" pageNumber="30">3</td>
<td id="76C21F06FFDD5B84FBBEFCEAFA517CE4" box="[1044,1497,868,947]" gridcol="5" gridrow="1" pageId="29" pageNumber="30">3 4 keel middle A3,A5,B5 quarter</td>
<td id="76C21F06FFDD5B84F9B6FCEAF9087CE4" box="[1564,1664,868,947]" gridcol="6" gridrow="1" pageId="29" pageNumber="30">middle quarter distal half</td>
<td id="76C21F06FFDD5B84F912FCEAF8967CE4" box="[1720,1822,868,947]" gridcol="7" gridrow="1" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FC5BF8967B74" box="[125,1822,981,1059]" gridrow="2" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FC5BFE197B74" box="[125,401,981,1059]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FC58FF7B7CBC" box="[125,243,981,1003]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="palaciosi" status="sp. nov.">
<emphasis id="B957A82AFFDDA466FFD7FC58FF7B7CBC" box="[125,243,981,1003]" italics="true" pageId="29" pageNumber="30">T. palaciosi</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDDA466FFD7FC7CFF477B5F" bold="true" box="[125,207,1010,1032]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2641551FFDDA466FFD7FC7CFF477B5F" box="[125,207,1010,1032]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76C21F06FFDD5B84FE62FC5BFDAB7B74" box="[456,547,981,1059]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">smooth</td>
<td id="76C21F06FFDD5B84FDFAFC5BFD327B74" box="[592,698,981,1059]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">absent</td>
<td id="76C21F06FFDD5B84FCACFC5BFCEA7B74" box="[774,866,981,1059]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">3</td>
<td id="76C21F06FFDD5B84FC24FC5BFC627B74" box="[910,1002,981,1059]" gridcol="4" gridrow="2" pageId="29" pageNumber="30">1-2</td>
<td id="76C21F06FFDD5B84FBBEFC5BFA517B74" box="[1044,1497,981,1059]" gridcol="5" gridrow="2" pageId="29" pageNumber="30">4 2-3 keel basal A3,A5,B4,B5 quarter</td>
<td id="76C21F06FFDD5B84F9B6FC5BF9087B74" box="[1564,1664,981,1059]" gridcol="6" gridrow="2" pageId="29" pageNumber="30">absent or middle quarter</td>
<td id="76C21F06FFDD5B84F912FC5BF8967B74" box="[1720,1822,981,1059]" gridcol="7" gridrow="2" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FBC8F8967BC3" box="[125,1822,1094,1172]" gridrow="3" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FBC8FE197BC3" box="[125,401,1094,1172]" gridcol="0" gridrow="3" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FBC9FE837B0B" box="[125,267,1094,1116]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dimorphica" status="sp. nov.">
<emphasis id="B957A82AFFDDA466FFD7FBC9FE837B0B" box="[125,267,1094,1116]" italics="true" pageId="29" pageNumber="30">T. dimorphica</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDDA466FFD7FBEDFF477B2E" bold="true" box="[125,207,1123,1145]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2641551FFDDA466FFD7FBEDFF477B2E" box="[125,207,1123,1145]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
form 2
</th>
<td id="76C21F06FFDD5B84FE62FBC8FDAB7BC3" box="[456,547,1094,1172]" gridcol="1" gridrow="3" pageId="29" pageNumber="30">ciliate</td>
<td id="76C21F06FFDD5B84FDFAFBC8FD327BC3" box="[592,698,1094,1172]" gridcol="2" gridrow="3" pageId="29" pageNumber="30">separated unusually short</td>
<td id="76C21F06FFDD5B84FCACFBC8FCEA7BC3" box="[774,866,1094,1172]" gridcol="3" gridrow="3" pageId="29" pageNumber="30">5</td>
<td id="76C21F06FFDD5B84FC24FBC8FC627BC3" box="[910,1002,1094,1172]" gridcol="4" gridrow="3" pageId="29" pageNumber="30">3</td>
<td id="76C21F06FFDD5B84FBBEFBC8FA517BC3" box="[1044,1497,1094,1172]" gridcol="5" gridrow="3" pageId="29" pageNumber="30">4 3 keel middle A3,A5,B4,B5 quarter</td>
<td id="76C21F06FFDD5B84F9B6FBC8F9087BC3" box="[1564,1664,1094,1172]" gridcol="6" gridrow="3" pageId="29" pageNumber="30">distal half</td>
<td id="76C21F06FFDD5B84F912FBC8F8967BC3" box="[1720,1822,1094,1172]" gridcol="7" gridrow="3" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FB39F8967BBD" box="[125,1822,1207,1258]" gridrow="4" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FB39FE197BBD" box="[125,401,1207,1258]" gridcol="0" gridrow="4" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FB39FEB87B9B" box="[125,304,1207,1228]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDDA466FFD7FB39FEB87B9B" box="[125,304,1207,1228]" italics="true" pageId="29" pageNumber="30">T. paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDDA466FFD7FB5AFF477BBD" bold="true" box="[125,207,1236,1258]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A2641551FFDDA466FFD7FB5AFF477BBD" box="[125,207,1236,1258]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76C21F06FFDD5B84FE62FB39FDAB7BBD" box="[456,547,1207,1258]" gridcol="1" gridrow="4" pageId="29" pageNumber="30">ciliate</td>
<td id="76C21F06FFDD5B84FDFAFB39FD327BBD" box="[592,698,1207,1258]" gridcol="2" gridrow="4" pageId="29" pageNumber="30">base fused wide</td>
<td id="76C21F06FFDD5B84FCACFB39FCEA7BBD" box="[774,866,1207,1258]" gridcol="3" gridrow="4" pageId="29" pageNumber="30">5</td>
<td id="76C21F06FFDD5B84FC24FB39FC627BBD" box="[910,1002,1207,1258]" gridcol="4" gridrow="4" pageId="29" pageNumber="30">0</td>
<td id="76C21F06FFDD5B84FBBEFB39FA517BBD" box="[1044,1497,1207,1258]" gridcol="5" gridrow="4" pageId="29" pageNumber="30">3 3 needle middle A3,A5,B5 quarter</td>
<td id="76C21F06FFDD5B84F9B6FB39F9087BBD" box="[1564,1664,1207,1258]" gridcol="6" gridrow="4" pageId="29" pageNumber="30">distal half</td>
<td id="76C21F06FFDD5B84F912FB39F8967BBD" box="[1720,1822,1207,1258]" gridcol="7" gridrow="4" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FA82F8967A69" box="[125,1822,1292,1342]" gridrow="5" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FA82FE197A69" box="[125,401,1292,1342]" gridcol="0" gridrow="5" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FA82FECF7A69" authority="(Denis, 1925) sensu Yoshii (1988)" authorityName="Yoshii" authorityYear="1988" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis id="B957A82AFFDDA466FFD7FA82FE887A76" box="[125,256,1292,1313]" italics="true" pageId="29" pageNumber="30">T. carpenteri</emphasis>
(Denis, 1925)
<emphasis id="B957A82AFFDDA466FFD7FAA6FF3D7A6A" box="[125,181,1320,1341]" italics="true" pageId="29" pageNumber="30">sensu</emphasis>
Yoshii (1988)
</taxonomicName>
</th>
<td id="76C21F06FFDD5B84FE62FA82FDAB7A69" box="[456,547,1292,1342]" gridcol="1" gridrow="5" pageId="29" pageNumber="30">smooth</td>
<td id="76C21F06FFDD5B84FDFAFA82FD327A69" box="[592,698,1292,1342]" gridcol="2" gridrow="5" pageId="29" pageNumber="30">separated</td>
<td id="76C21F06FFDD5B84FCACFA82FCEA7A69" box="[774,866,1292,1342]" gridcol="3" gridrow="5" pageId="29" pageNumber="30">?</td>
<td id="76C21F06FFDD5B84FC24FA82FC627A69" box="[910,1002,1292,1342]" gridcol="4" gridrow="5" pageId="29" pageNumber="30">?</td>
<td id="76C21F06FFDD5B84FBBEFA82FA517A69" box="[1044,1497,1292,1342]" gridcol="5" gridrow="5" pageId="29" pageNumber="30">? 3 keel middle quarter</td>
<td id="76C21F06FFDD5B84F9B6FA82F9087A69" box="[1564,1664,1292,1342]" gridcol="6" gridrow="5" pageId="29" pageNumber="30">distal half</td>
<td id="76C21F06FFDD5B84F912FA82F8967A69" box="[1720,1822,1292,1342]" gridcol="7" gridrow="5" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FAEEF8967AF8" box="[125,1822,1376,1455]" gridrow="6" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FAEEFE197AF8" box="[125,401,1376,1455]" gridcol="0" gridrow="6" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FAEFFECA7AF8" authority="Palacios-Vargas, Ojeda &amp; Christiansen, 1985" authorityName="Palacios-Vargas, Ojeda &amp; Christiansen" authorityYear="1985" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="B957A82AFFDDA466FFD7FAEFFF517A21" box="[125,217,1376,1398]" italics="true" pageId="29" pageNumber="30">T. relicta</emphasis>
(Palacios-Vargas, Ojeda &amp; Christiansen, 1985)
</taxonomicName>
</th>
<td id="76C21F06FFDD5B84FE62FAEEFDAB7AF8" box="[456,547,1376,1455]" gridcol="1" gridrow="6" pageId="29" pageNumber="30">ciliate</td>
<td id="76C21F06FFDD5B84FDFAFAEEFD327AF8" box="[592,698,1376,1455]" gridcol="2" gridrow="6" pageId="29" pageNumber="30">base fused wide</td>
<td id="76C21F06FFDD5B84FCACFAEEFCEA7AF8" box="[774,866,1376,1455]" gridcol="3" gridrow="6" pageId="29" pageNumber="30">5</td>
<td id="76C21F06FFDD5B84FC24FAEEFC627AF8" box="[910,1002,1376,1455]" gridcol="4" gridrow="6" pageId="29" pageNumber="30">0</td>
<td id="76C21F06FFDD5B84FBBEFAEEFA517AF8" box="[1044,1497,1376,1455]" gridcol="5" gridrow="6" pageId="29" pageNumber="30">2 3 keel middle A3,B5 quarter</td>
<td id="76C21F06FFDD5B84F9B6FAEEF9087AF8" box="[1564,1664,1376,1455]" gridcol="6" gridrow="6" pageId="29" pageNumber="30">distal half</td>
<td id="76C21F06FFDD5B84F912FAEEF8967AF8" box="[1720,1822,1376,1455]" gridcol="7" gridrow="6" pageId="29" pageNumber="30">4 sequential</td>
</tr>
<tr id="3513767AFFDD5B84FFD7FA5FF896796B" box="[125,1822,1489,1596]" gridrow="7" pageId="29" pageNumber="30">
<th id="76C21F06FFDD5B84FFD7FA5FFE19796B" box="[125,401,1489,1596]" gridcol="0" gridrow="7" pageId="29" pageNumber="30">
<taxonomicName id="4C230FBBFFDDA466FFD7FA5CFED37954" authority="Thibaud &amp; Najt, 1988" authorityName="Thibaud &amp; Najt" authorityYear="1988" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cotopaxiana">
<emphasis id="B957A82AFFDDA466FFD7FA5CFE9B7AB0" box="[125,275,1490,1511]" italics="true" pageId="29" pageNumber="30">T. cotopaxiana</emphasis>
Thibaud &amp; Najt, 1988
</taxonomicName>
</th>
<td id="76C21F06FFDD5B84FE62FA5FFDAB796B" box="[456,547,1489,1596]" gridcol="1" gridrow="7" pageId="29" pageNumber="30">ciliate</td>
<td id="76C21F06FFDD5B84FDFAFA5FFD32796B" box="[592,698,1489,1596]" gridcol="2" gridrow="7" pageId="29" pageNumber="30">absent?</td>
<td id="76C21F06FFDD5B84FCACFA5FFCEA796B" box="[774,866,1489,1596]" gridcol="3" gridrow="7" pageId="29" pageNumber="30">3</td>
<td id="76C21F06FFDD5B84FC24FA5FFC62796B" box="[910,1002,1489,1596]" gridcol="4" gridrow="7" pageId="29" pageNumber="30">0</td>
<td id="76C21F06FFDD5B84FBBEFA5FFA51796B" box="[1044,1497,1489,1596]" gridcol="5" gridrow="7" pageId="29" pageNumber="30">3 4 keel basal A3, A5, B5 half</td>
<td id="76C21F06FFDD5B84F9B6FA5FF908796B" box="[1564,1664,1489,1596]" gridcol="6" gridrow="7" pageId="29" pageNumber="30">distal half</td>
<td id="76C21F06FFDD5B84F912FA5FF896796B" box="[1720,1822,1489,1596]" gridcol="7" gridrow="7" pageId="29" pageNumber="30">5 middle teeth paired</td>
</tr>
</table>
</paragraph>
<caption id="DF5C24B0FFDEA465FF3DF81FFD5E78B3" httpUri="https://zenodo.org/record/289760/files/figure.png" pageId="30" pageNumber="31" targetBox="[225,1349,202,1890]" targetPageId="30">
<paragraph id="8B9C7438FFDEA465FF3DF81FFD5E78B3" blockId="30.[151,1436,1937,2020]" pageId="30" pageNumber="31">
<emphasis id="B957A82AFFDEA465FF3DF81FFE9778F1" bold="true" box="[151,287,1937,1959]" pageId="30" pageNumber="31">FIGURE 16.</emphasis>
<taxonomicName id="4C230FBBFFDEA465FE8FF81FFDCD78F0" box="[293,581,1937,1959]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="paracapenteri" status="sp. nov.">
<emphasis id="B957A82AFFDEA465FE8FF81FFDCD78F0" box="[293,581,1937,1959]" italics="true" pageId="30" pageNumber="31">Trogolaphysa paracapenteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2641551FFDEA465FDE6F81FFD1478F1" box="[588,668,1937,1958]" pageId="30" pageNumber="31" rank="species">
<emphasis id="B957A82AFFDEA465FDE6F81FFD1478F1" bold="true" box="[588,668,1937,1958]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
A, Dorsal chaetotaxy of head; B, Distal labral spines; C, Sensillum of third antennal segment sense organ; D, Mucro; E, Mesothoracic chaetotaxy; F, Hind claw complex; G, Abdomen 4 macrochaetae; H, Abdomen 4 outer macrochaetae in a different individual.
</paragraph>
</caption>
<paragraph id="8B9C7438FFDFA464FF6DFF19FC127FAF" blockId="31.[151,1436,151,248]" pageId="31" pageNumber="32">
<taxonomicName id="4C230FBBFFDFA464FF6DFF19FD9C7FE7" box="[199,532,151,176]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="paracarpenteri" status="sp. nov.">
<emphasis id="B957A82AFFDFA464FF6DFF19FD9C7FE7" box="[199,532,151,176]" italics="true" pageId="31" pageNumber="32">Trogolaphysa paracarpenteri</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDFA464FDB4FF19FDF47FE7" bold="true" box="[542,636,151,176]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A2641551FFDFA464FDB4FF19FDF47FE7" box="[542,636,151,176]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the same color pattern as
<taxonomicName id="4C230FBBFFDFA464FC4FFF17FBF27FE7" box="[997,1146,153,176]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="octosetosa" status="sp. nov.">
<emphasis id="B957A82AFFDFA464FC4FFF17FBF27FE7" box="[997,1146,153,176]" italics="true" pageId="31" pageNumber="32">T. octosetosa</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDFA464FB2EFF19FB697FE7" bold="true" box="[1156,1249,151,176]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A2641551FFDFA464FB2EFF19FB697FE7" box="[1156,1249,151,176]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the differences between these two species, and all other species with 6–8 eyes and 2 anterior head macrochaetae are detailed in
<tableCitation id="C6A14183FFDFA464FF3DFF51FF627FAF" box="[151,234,223,248]" captionStart="TABLE 4" captionStartId="29.[125,210,612,634]" captionTargetBox="[125,1822,669,1596]" captionTargetPageId="29" captionText="TABLE 4. Diagnostic table of Neotropical species of Trogolaphysa with 6 – 8 eyes and two (A 0, A 2) dorsal anterior head macrochaetae. Mc = macrochaeta." httpUri="http://table.plazi.org/id/DF5C24B0FFDDA466FFD7FDEAF9287D2D" pageId="31" pageNumber="32" tableUuid="DF5C24B0FFDDA466FFD7FDEAF9287D2D">Table 4</tableCitation>
and discussed in the remarks to
<taxonomicName id="4C230FBBFFDFA464FDFCFF6FFD6F7FAF" box="[598,743,225,248]" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="octosetosa" status="sp. nov.">
<emphasis id="B957A82AFFDFA464FDFCFF6FFD6F7FAF" box="[598,743,225,248]" italics="true" pageId="31" pageNumber="32">T. octosetosa</emphasis>
</taxonomicName>
<emphasis id="B957A82AFFDFA464FD45FF51FCC07FAF" bold="true" box="[751,840,223,248]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A2641551FFDFA464FD45FF51FCC07FAF" box="[751,840,223,248]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
below.
</paragraph>
</subSubSection>
</treatment>
</document>
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