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<document ID-DOI="http://doi.org/10.5281/zenodo.3850345" ID-GBIF-Dataset="d3e9578f-3031-417e-8f1a-1bcec3bfda48" ID-GBIF-Taxon="164167623" ID-ISSN="2468-872X" ID-Zenodo-Dep="3850345" ID-ZooBank="urn:lsid:zoobank.org:pub:DDD13523-E480-47C3-94BC-089E40AB3579" approvalRequired="55" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="53" checkinTime="1589833990572" checkinUser="felipe" docAuthor="Davenport, Chris &amp; Grimaldi, Nicolas" docDate="2019" docId="0389BB02A52A7255FF0A0A0E577EEEF1" docLanguage="en" docName="SugapaDigital.11.2.53-67.pdf.imf" docOrigin="Sugapa Digital 11 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6f628e0e" docStyleName="sugapadigital.2019.journal_article" docTitle="Delias leucias subsp. leucias leucias" docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="59" masterDocId="FFB0C37AA52F7253FFB70E5D5275EF7B" masterDocTitle="A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae)" masterLastPageNumber="67" masterPageNumber="53" pageId="5" pageNumber="58" updateTime="1643459741562" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A new subspecies of Delias clathrata Rothschild, 1904 and records of other Delias species occurring on Mount Hagen, Papua New Guinea (Lepidoptera: Pieridae, Pierinae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davenport, Chris</mods:namePart>
<mods:affiliation>Tynaherrick, Drumnadrochit, Inverness-shire, UK Plaza Damas, 60 Jalan Sri Hartamas 1, Kuala Lumpur</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Grimaldi, Nicolas</mods:namePart>
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<mods:title>Sugapa Digital</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-02-03</mods:number>
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<mods:detail type="volume">
<mods:number>11</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="DOI">10.19269/sugapa2019.11(2).01</mods:identifier>
<mods:identifier type="GBIF-Dataset">d3e9578f-3031-417e-8f1a-1bcec3bfda48</mods:identifier>
<mods:identifier type="ISSN">2468-872X</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3850345" ID-GBIF-Taxon="164167623" ID-Zenodo-Dep="3850345" LSID="urn:lsid:plazi:treatment:0389BB02A52A7255FF0A0A0E577EEEF1" httpUri="http://treatment.plazi.org/id/0389BB02A52A7255FF0A0A0E577EEEF1" lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
<heading bold="true" box="[189,861,1107,1142]" fontSize="14" level="2" pageId="5" pageNumber="58" reason="2">
<subSubSection box="[189,718,1107,1142]" pageId="5" pageNumber="58" type="nomenclature">
<paragraph blockId="5.[189,1444,1107,1886]" box="[189,718,1107,1142]" pageId="5" pageNumber="58">
<emphasis bold="true" box="[189,718,1107,1142]" pageId="5" pageNumber="58">
<taxonomicName authority="leucias" box="[189,512,1107,1142]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="leucias" subSpecies="leucias">
<emphasis bold="true" box="[189,512,1107,1142]" italics="true" pageId="5" pageNumber="58">Delias leucias leucias</emphasis>
</taxonomicName>
<collectingCountry box="[521,630,1107,1142]" name="Jordan" pageId="5" pageNumber="58">Jordan</collectingCountry>
, 1912
</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[726,861,1111,1141]" pageId="5" pageNumber="58" type="description">
<paragraph blockId="5.[189,1444,1107,1886]" box="[726,861,1111,1141]" pageId="5" pageNumber="58">
(
<figureCitation box="[733,854,1111,1141]" captionStart-0="Figs 9-14" captionStart-1="Figs 15-16" captionStartId-0="9.[381,423,1729,1754]" captionStartId-1="9.[381,423,1859,1884]" captionTargetBox-0="[332,1303,233,944]" captionTargetBox-1="[323,1313,979,1687]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionText-0="Figs 9-14. Delias leucias leucias Jordan, 1912. 9. ♂, PNG, Mt Hagen, Pap Creek, 2500m (CD), dorso; 10. Idem, verso; 11. ♀, PNG, Mt Hagen, Murmur Pass, 2850m (CD), dorso; 12. Idem, verso; 13. ♂, Indonesia, Papua, Star Mts, Ok Tedi (CD), a. dorso, b. verso; 14. ♀, Indonesia, Papua, Ilu (KSP), a. dorso, b. verso." captionText-1="Figs 15-16. Delias leucias huonensis Talbot, 1928. 15. ♂ holotype, PNG, Rawlinson Mts (NHMUK), dorso; 16. Idem, verso." figureDoi-0="http://doi.org/10.5281/zenodo.3832857" figureDoi-1="http://doi.org/10.5281/zenodo.3832859" httpUri-0="https://zenodo.org/record/3832857/files/figure.png" httpUri-1="https://zenodo.org/record/3832859/files/figure.png" pageId="5" pageNumber="58">Figs 9-16</figureCitation>
)
</paragraph>
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</heading>
<subSubSection lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58" type="discussion">
<paragraph blockId="5.[189,1444,1107,1886]" pageId="5" pageNumber="58">
The subspecific identity of
<taxonomicName authorityName="Jordan" authorityYear="1912" box="[550,678,1152,1182]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="leucias">
<emphasis box="[550,678,1152,1182]" italics="true" pageId="5" pageNumber="58">D. leucias</emphasis>
</taxonomicName>
from the central mountains of
<collectingCountry box="[1105,1360,1152,1183]" name="Papua New Guinea" pageId="5" pageNumber="58">Papua New Guinea</collectingCountry>
has a confusing history, having been assigned to the nominate race (
<typeStatus box="[1053,1113,1191,1221]" pageId="5" pageNumber="58">type</typeStatus>
locality: Mt Goliath) by DAbrera (1978), and subsequently to subspecies
<emphasis box="[861,990,1230,1260]" italics="true" pageId="5" pageNumber="58">huonensis</emphasis>
by later authors such as
<bibRefCitation author="Parsons, M. J." pageId="5" pageNumber="58" refId="ref4591" refString="Parsons, M. J., 1998. The butterflies of Papua New Guinea: their systematics and biology. Academic Press, London: 736 pp. + 104 pls." type="book" year="1998">Parsons (1998)</bibRefCitation>
,
<bibRefCitation author="Yagishita, A. &amp; Nakano, S. &amp; Morita, S." box="[293,587,1269,1299]" pageId="5" pageNumber="58" refId="ref4697" refString="Yagishita, A., Nakano, S. &amp; Morita, S., 1993. An illustrated list of the genus Delias Hubner of the World. 2 volumes, Khepera, Tokyo; xiv, 384, 409, Vi pp." type="book" year="1993">Yagishita et al. (1993)</bibRefCitation>
and Van Mastrigt (2003).
<bibRefCitation author="Lachlan, R. B." box="[963,1157,1269,1299]" pageId="5" pageNumber="58" pagination="71 - 96" refId="ref4188" refString="Lachlan, R. B., 2000. New species, subspecies and records of Delias Hubner (Lepidoptera: Pieridae) from the upper Ok Tedi and Tari regions, Papua New Guinea. Aust. Entom. 27 (3): 71 - 96." type="journal article" year="2000">Lachlan (2000)</bibRefCitation>
reports collection of
<taxonomicName authority="leucias" box="[189,466,1308,1338]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="leucias" subSpecies="leucias">
<emphasis box="[189,466,1308,1338]" italics="true" pageId="5" pageNumber="58">Delias leucias leucias</emphasis>
</taxonomicName>
at Ok Tedi,
<collectingRegion box="[634,869,1308,1338]" country="Papua New Guinea" name="Western" pageId="5" pageNumber="58">Western Province</collectingRegion>
and
<taxonomicName authorityName="Talbot" authorityYear="1928" box="[940,1207,1308,1338]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="leucias" subSpecies="huonensis">
<emphasis box="[940,1207,1308,1338]" italics="true" pageId="5" pageNumber="58">D. leucias huonensis</emphasis>
</taxonomicName>
at Tari,
<collectingRegion country="Papua New Guinea" name="Southern Highlands" pageId="5" pageNumber="58">Southern Highlands</collectingRegion>
.
<bibRefCitation author="Gotts, R. I. C. &amp; Ginn, S. G." box="[346,617,1347,1378]" pageId="5" pageNumber="58" pagination="49 - 58" refId="ref4144" refString="Gotts, R. I. C. &amp; Ginn, S. G., 2004. New subspecies of Delias Hubner (Lepidoptera: Pieridae) from West Papua. Aust. Entom. 31 (2): 49 - 58" type="journal article" year="2004">Gotts &amp; Ginn (2004)</bibRefCitation>
described a new subspecies,
<taxonomicName authorityName="Jordan" authorityYear="1912" box="[1018,1145,1347,1377]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="leucias">
D.
<emphasis box="[1057,1145,1347,1377]" italics="true" pageId="5" pageNumber="58">leucias</emphasis>
</taxonomicName>
<emphasis box="[1156,1233,1347,1377]" italics="true" pageId="5" pageNumber="58">torini̡</emphasis>
from the Snow Mts, west of the nominate range, and discussed variation in
<taxonomicName authorityName="Jordan" authorityYear="1912" box="[1031,1162,1386,1416]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="leucias">
<emphasis box="[1031,1162,1386,1416]" italics="true" pageId="5" pageNumber="58">D. leucias</emphasis>
</taxonomicName>
across New
<collectingCountry box="[1342,1440,1386,1417]" name="Guinea-Bissau" pageId="5" pageNumber="58">Guinea</collectingCountry>
, mentioning the differences between the Kerowagi (Mt Wilhelm) population and the nominate subspecies from the Star Mountains. They considered the taxon
<taxonomicName authority="Okano, 1989" authorityName="Okano" authorityYear="1989" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="weiskei" subSpecies="sayuriae">
<emphasis box="[1078,1336,1464,1494]" italics="true" pageId="5" pageNumber="58">D. weiskei sayuriae</emphasis>
Okano, 1989
</taxonomicName>
, recorded only from Mt Wilhelm, to be a variation of
<taxonomicName authorityName="Jordan" authorityYear="1912" box="[957,1046,1504,1534]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="leucias">
<emphasis box="[957,1046,1504,1534]" italics="true" pageId="5" pageNumber="58">leucias</emphasis>
</taxonomicName>
rather than a subspecies of
<taxonomicName authority="Ribbe, 1900" authorityName="Ribbe" authorityYear="1900" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="weiskei">
<emphasis italics="true" pageId="5" pageNumber="58">D. weiskei</emphasis>
Ribbe, 1900
</taxonomicName>
.
</paragraph>
<paragraph blockId="5.[189,1444,1107,1886]" pageId="5" pageNumber="58">
<taxonomicName authority="Talbot, 1928" authorityName="Talbot" authorityYear="1928" box="[189,628,1582,1612]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="leucias" subSpecies="huonensis">
<emphasis box="[189,453,1582,1612]" italics="true" pageId="5" pageNumber="58">D. leucias huonensis</emphasis>
<bibRefCitation author="Talbot, G." box="[464,628,1582,1612]" pageId="5" pageNumber="58" pagination="1 - 259" refId="ref4665" refString="Talbot, G., 1928 - 1937. A Monograph of the Pierine Genus Delias. British Museum, London Parts 1 - 5; pp. 1 - 259." type="book chapter" year="1928">Talbot, 1928</bibRefCitation>
</taxonomicName>
was described from a single male collected by Keysser in the isolated Rawlinson Mountains on the Huon peninsula in north east New
<collectingCountry box="[1208,1305,1621,1652]" name="Guinea-Bissau" pageId="5" pageNumber="58">Guinea</collectingCountry>
. The
<taxonomicName pageId="5" pageNumber="58" rank="subSpecies" subSpecies="remains">race remains</taxonomicName>
rare in collections and apparently in nature; William Brandt did not record the species during his expedition to the Rawlinson Mts in 1968.
</paragraph>
<paragraph blockId="5.[189,1444,1107,1886]" pageId="5" pageNumber="58">
Specimens from Mt Hagen are similar to those from Mt Wilhelm &amp; the
<collectingRegion box="[1131,1364,1738,1768]" country="Papua New Guinea" name="Eastern Highlands" pageId="5" pageNumber="58">Eastern Highlands</collectingRegion>
, both can be distinguished
<emphasis box="[484,883,1777,1807]" italics="true" pageId="5" pageNumber="58">in the majority of individuals</emphasis>
from the nominate subspecies - by the complete separation of the pale hindwing patch from the white costal border by the red and black subapical bands, and a more orange coloration of the forewing underside - and from ssp
</paragraph>
<paragraph blockId="6.[189,1443,208,394]" pageId="6" pageNumber="59">
<emphasis box="[189,318,208,238]" italics="true" pageId="6" pageNumber="59">huonensis</emphasis>
by the narrower black margins on the both surfaces and the more elongated hind wing patch. Although these characters are consistent in the majority of specimens from central
<collectingCountry box="[189,251,285,316]" name="Papua New Guinea" pageId="6" pageNumber="59">PNG</collectingCountry>
, populations of the nominate
<taxonomicName box="[658,789,286,316]" pageId="6" pageNumber="59" rank="subSpecies" subSpecies="from">race from</taxonomicName>
various locations within Indonesian Papua show clinal development of the same features on a west - east axis.
<taxonomicName authorityName="Jordan" authorityYear="1912" box="[1005,1180,325,355]" class="Insecta" family="Pieridae" genus="Delias" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="leucias">
<emphasis box="[1005,1180,325,355]" italics="true" pageId="6" pageNumber="59">Delias leucias</emphasis>
</taxonomicName>
from central
<collectingCountry box="[1357,1415,325,356]" name="Papua New Guinea" pageId="6" pageNumber="59">PNG</collectingCountry>
is considered to represent a recognizable geographic form of the nominate subspecies.
</paragraph>
</subSubSection>
</treatment>
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