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<document id="D718E871E478732B97BEDBBD44B231FD" ID-CLB-Dataset="3916" ID-DOI="10.11646/zootaxa.3879.1.1" ID-GBIF-Dataset="4b976ba3-1a7b-4dd5-8afb-630b7cc8a695" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949051" ID-ZooBank="6423894B-97D9-4286-ABB9-D4AF072B57FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623702926238" checkinUser="felipe" docAuthor="Borkent, Art" docDate="2014" docId="027587C9BD613026FD4E199F4801E6A4" docLanguage="en" docName="zt03879p327.pdf.imf" docOrigin="Zootaxa 3879 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dibezzia Kieffer" docType="treatment" docVersion="10" lastPageNumber="81" masterDocId="FE4CFFB1BD2E3076FFF81A154D70E204" masterDocTitle="The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera" masterLastPageNumber="327" masterPageNumber="1" pageNumber="80" updateTime="1699015668130" updateUser="ExternalLinkService">
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<mods:title id="AAD3186F5B657604300F7EA91B6ECBFB">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<treatment id="027587C9BD613026FD4E199F4801E6A4" ID-DOI="http://doi.org/10.5281/zenodo.5593021" ID-GBIF-Taxon="189161045" ID-Zenodo-Dep="5593021" LSID="urn:lsid:plazi:treatment:027587C9BD613026FD4E199F4801E6A4" httpUri="http://treatment.plazi.org/id/027587C9BD613026FD4E199F4801E6A4" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<subSubSection id="C2C66554BD613039FD4E199F4E0DE1A0" box="[694,893,906,932]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="8A6336DFBD613039FD4E199F4E0DE1A0" blockId="79.[495,1092,906,966]" box="[694,893,906,932]" pageId="79" pageNumber="80">
<taxonomicName id="4DDC4D5CBD613039FD4E199F4E0DE1A0" ID-CoL="44FR" authority="Kieffer" authorityName="Kieffer" box="[694,893,906,932]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD613039FD4E199F4E0DE1A0" bold="true" box="[694,893,906,932]" pageId="79" pageNumber="80">
<emphasis id="B8A8EACDBD613039FD4E199F4E6CE1A0" bold="true" box="[694,796,906,932]" italics="true" pageId="79" pageNumber="80">Dibezzia</emphasis>
Kieffer
</emphasis>
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<subSubSection id="C2C66554BD613039FE1719B84934E1C2" box="[495,1092,941,966]" pageId="79" pageNumber="80" type="description">
<paragraph id="8A6336DFBD613039FE1719B84934E1C2" blockId="79.[495,1092,906,966]" box="[495,1092,941,966]" pageId="79" pageNumber="80">
(
<figureCitation id="12E72A5ABD613039FE0F19B84F18E1C2" box="[503,616,941,966]" captionStart="FIGURE 12" captionStartId="257.[151,250,1853,1875]" captionTargetBox="[215,1366,217,1823]" captionTargetId="figure-122@257.[203,1398,210,1827]" captionTargetPageId="257" captionText="FIGURE 12. Habitus. A. Pellucidomyia leei, male, in dorsal view (from Debenham 1970). B. Dibezzia prominens, sex unknown, in dorsal view (form Mayer 1934c). C. Macropeza albitarsis, male, in dorsal view (from Szadziewski &amp; Dominiak, 2007, modified). D. Mallochohelea munda, male, in ventral view (from Szadziewski et al. 1997). E. Sphaeromias fasciata, sex unknown, in dorsal view (from Mayer 1934a). F. Bezzia glabra, female, in ventral view (from Thomsen 1937). G. Palpomyia sp., male, in dorsal view (from Grogan &amp; Wirth 1979). H. Paryphoconus oliveirai, female, in ventral view (modified from Ronderos et al. 2007)." figureDoi="http://doi.org/10.5281/zenodo.4949100" httpUri="https://zenodo.org/record/4949100/files/figure.png" pageId="79" pageNumber="80">Figs. 12B</figureCitation>
,
<figureCitation id="12E72A5ABD613039FD8B19B84FD5E1C2" box="[627,677,941,966]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="79" pageNumber="80">21A</figureCitation>
,
<figureCitation id="12E72A5ABD613039FD4919B84F91E1C2" box="[689,737,941,966]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">26E</figureCitation>
,
<figureCitation id="12E72A5ABD613039FD1519B84E66E1C2" box="[749,790,941,966]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="79" pageNumber="80">30J</figureCitation>
,
<figureCitation id="12E72A5ABD613039FCD919B84E24E1C2" box="[801,852,941,966]" captionStart="FIGURE 38" captionStartId="283.[151,250,1936,1958]" captionTargetBox="[195,1395,204,1921]" captionTargetId="figure-66@283.[186,1417,186,1925]" captionTargetPageId="283" captionText="FIGURE 38. Legs, wings and apices of the antennae, in ventral view. A. Dibezzia debenhamae (reconstructed from distorted specimen). B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea sp. (from Ottawa, ON, Canada). D. Macropeza natalensis. E. Macropeza sp. (from Laos). F. Mallochohelea caudellii." figureDoi="http://doi.org/10.5281/zenodo.4949194" httpUri="https://zenodo.org/record/4949194/files/figure.png" pageId="79" pageNumber="80">38A</figureCitation>
,
<figureCitation id="12E72A5ABD613039FCA719B84EFFE1C2" box="[863,911,941,966]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="79" pageNumber="80">45G</figureCitation>
,
<figureCitation id="12E72A5ABD613039FC6219B84EBEE1C2" box="[922,974,941,966]" captionStart="FIGURE 51" captionStartId="296.[151,250,1944,1966]" captionTargetBox="[184,1397,192,1912]" captionTargetId="figure-64@296.[174,1406,183,1922]" captionTargetPageId="296" captionText="FIGURE 51. Metathoraces and tergites 1, in dorsal view. A. Dibezzia debenhamae. B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea magnipennis. D. Macropeza natalensis. E. Mallochohelea caudellii. F. Probezzia sabroskyi." figureDoi="http://doi.org/10.5281/zenodo.4949230" httpUri="https://zenodo.org/record/4949230/files/figure.png" pageId="79" pageNumber="80">51A</figureCitation>
,
<figureCitation id="12E72A5ABD613039FC2119B8497AE1C2" box="[985,1034,941,966]" captionStart="FIGURE 64" captionStartId="309.[151,250,1979,2001]" captionTargetBox="[171,1416,189,1957]" captionTargetId="figure-49@309.[161,1426,183,1967]" captionTargetPageId="309" captionText="FIGURE 64. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Pellucidomyia leei. B. Dibezzia debenhamae (reconstructed from distorted and dirty specimen). C. Hebetula tonnoiri." figureDoi="http://doi.org/10.5281/zenodo.4949278" httpUri="https://zenodo.org/record/4949278/files/figure.png" pageId="79" pageNumber="80">64B</figureCitation>
,
<figureCitation id="12E72A5ABD613039FBED19B8494BE1C2" box="[1045,1083,941,966]" captionStart="FIGURE 75" captionStartId="320.[151,250,1792,1814]" captionTargetBox="[220,1359,193,1769]" captionTargetId="figure-191@320.[210,1379,193,1773]" captionTargetPageId="320" captionText="FIGURE 75. Abdominal segments 9. A. Parabezzia huberti, female, in ventral view (from Grogan &amp; Wirth 1977a). B. Echinohelea lanei (from Wirth 1994b). C. Clinohelea bimaculata, female, in ventral view (from Wirth &amp; Grogan 1979). D. Heteromyia wokei, male, in dorsal and ventral view (from Wirth &amp; Grogan 1977). E. Heteromyia wokei, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). F. Heteromyia clavata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). G. Pellucidomyia leei, male, in dorsal view (from Elson-Harris &amp; Kettle 1986). H. Pellucidomyia geari, male, in ventral view (from de Meillon &amp; Wirth 1981). I. Dibezzia debenhamae, male, in dorsal and ventral view (shagreen not shown). J. Hebetula tonnoiri, female, in dorsal view (from Elson-Harris 1987). K. Jenkinshelea albaria, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Jenkinshelea albaria, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949320" httpUri="https://zenodo.org/record/4949320/files/figure.png" pageId="79" pageNumber="80">75I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD613039FF6F19E04E91E636" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8A6336DFBD613039FF6F19E04E91E636" blockId="79.[151,1437,1013,2010]" pageId="79" pageNumber="80">
<emphasis id="B8A8EACDBD613039FF6F19E04C44E60A" bold="true" box="[151,308,1013,1038]" pageId="79" pageNumber="80">DIAGNOSIS</emphasis>
: Only pupa of
<taxonomicName id="4DDC4D5CBD613039FE0819E04FDFE60A" authorityName="Borkent" authorityYear="2014" box="[496,687,1013,1038]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with a well-developed prothoracic extension (
<figureCitation id="12E72A5ABD613039FB2B19E0484AE60A" box="[1235,1338,1013,1038]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">Fig. 26E</figureCitation>
) and an elongate seta (D-5-IX) on the base of the terminal process (
<figureCitation id="12E72A5ABD613039FCD71E0C4ED0E636" box="[815,928,1049,1074]" captionStart="FIGURE 12" captionStartId="257.[151,250,1853,1875]" captionTargetBox="[215,1366,217,1823]" captionTargetId="figure-122@257.[203,1398,210,1827]" captionTargetPageId="257" captionText="FIGURE 12. Habitus. A. Pellucidomyia leei, male, in dorsal view (from Debenham 1970). B. Dibezzia prominens, sex unknown, in dorsal view (form Mayer 1934c). C. Macropeza albitarsis, male, in dorsal view (from Szadziewski &amp; Dominiak, 2007, modified). D. Mallochohelea munda, male, in ventral view (from Szadziewski et al. 1997). E. Sphaeromias fasciata, sex unknown, in dorsal view (from Mayer 1934a). F. Bezzia glabra, female, in ventral view (from Thomsen 1937). G. Palpomyia sp., male, in dorsal view (from Grogan &amp; Wirth 1979). H. Paryphoconus oliveirai, female, in ventral view (modified from Ronderos et al. 2007)." figureDoi="http://doi.org/10.5281/zenodo.4949100" httpUri="https://zenodo.org/record/4949100/files/figure.png" pageId="79" pageNumber="80">Figs. 12B</figureCitation>
,
<figureCitation id="12E72A5ABD613039FC531E0C4EA5E636" box="[939,981,1049,1074]" captionStart="FIGURE 75" captionStartId="320.[151,250,1792,1814]" captionTargetBox="[220,1359,193,1769]" captionTargetId="figure-191@320.[210,1379,193,1773]" captionTargetPageId="320" captionText="FIGURE 75. Abdominal segments 9. A. Parabezzia huberti, female, in ventral view (from Grogan &amp; Wirth 1977a). B. Echinohelea lanei (from Wirth 1994b). C. Clinohelea bimaculata, female, in ventral view (from Wirth &amp; Grogan 1979). D. Heteromyia wokei, male, in dorsal and ventral view (from Wirth &amp; Grogan 1977). E. Heteromyia wokei, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). F. Heteromyia clavata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). G. Pellucidomyia leei, male, in dorsal view (from Elson-Harris &amp; Kettle 1986). H. Pellucidomyia geari, male, in ventral view (from de Meillon &amp; Wirth 1981). I. Dibezzia debenhamae, male, in dorsal and ventral view (shagreen not shown). J. Hebetula tonnoiri, female, in dorsal view (from Elson-Harris 1987). K. Jenkinshelea albaria, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Jenkinshelea albaria, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949320" httpUri="https://zenodo.org/record/4949320/files/figure.png" pageId="79" pageNumber="80">75I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD613026FF3F1E28498FE3F1" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="description">
<paragraph id="8A6336DFBD613026FF3F1E28498FE3F1" blockId="79.[151,1437,1013,2010]" lastBlockId="80.[151,1436,151,1184]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="B8A8EACDBD613039FF3F1E284CFAE652" bold="true" box="[199,394,1085,1110]" pageId="79" pageNumber="80">DESCRIPTION</emphasis>
: Habitus as in
<figureCitation id="12E72A5ABD613039FDC81E284FE5E652" box="[560,661,1085,1110]" captionStart="FIGURE 12" captionStartId="257.[151,250,1853,1875]" captionTargetBox="[215,1366,217,1823]" captionTargetId="figure-122@257.[203,1398,210,1827]" captionTargetPageId="257" captionText="FIGURE 12. Habitus. A. Pellucidomyia leei, male, in dorsal view (from Debenham 1970). B. Dibezzia prominens, sex unknown, in dorsal view (form Mayer 1934c). C. Macropeza albitarsis, male, in dorsal view (from Szadziewski &amp; Dominiak, 2007, modified). D. Mallochohelea munda, male, in ventral view (from Szadziewski et al. 1997). E. Sphaeromias fasciata, sex unknown, in dorsal view (from Mayer 1934a). F. Bezzia glabra, female, in ventral view (from Thomsen 1937). G. Palpomyia sp., male, in dorsal view (from Grogan &amp; Wirth 1979). H. Paryphoconus oliveirai, female, in ventral view (modified from Ronderos et al. 2007)." figureDoi="http://doi.org/10.5281/zenodo.4949100" httpUri="https://zenodo.org/record/4949100/files/figure.png" pageId="79" pageNumber="80">Fig. 12B</figureCitation>
. Total length =
<quantity id="4D249B3ABD613039FCBB1E284ED7E651" box="[835,935,1085,1110]" metricMagnitude="-3" metricUnit="m" metricValue="4.03" pageId="79" pageNumber="80" unit="mm" value="4.03">4.03 mm</quantity>
(
<quantity id="4D249B3ABD613039FC4D1E28497CE651" box="[949,1036,1085,1110]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="79" pageNumber="80" unit="mm" value="5.5">5.5 mm</quantity>
from
<bibRefCitation id="EE4D4B2EBD613039FBA81E2B4999E652" author="Mayer" box="[1104,1257,1085,1110]" firstAuthor="Mayer" pageId="79" pageNumber="80" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">Mayer 1934c</bibRefCitation>
). Without larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of midleg, wing (as in
<figureCitation id="12E72A5ABD613039FA9D1E744DB8E69A" captionStart="FIGURE 16" captionStartId="261.[151,250,1897,1919]" captionTargetBox="[172,1416,199,1871]" captionTargetId="figure-85@261.[172,1416,193,1876]" captionTargetPageId="261" captionText="FIGURE 16. Anterior portion of heads. A. Parabezzia huberti, in ventral view (distorted slide specimen). B. Parabezzia sp. (5 km. E. Danby, Vermont, USA), ventrolateral view. C. Clinohelea curriei, in ventral view. D. Heteromyia clavata, in ventral view, dorsal apotome not present. E. Heteromyia wokei, in ventral, slightly lateral view. F. Pellucidomyia leei, in ventral view, dorsal apotome not present." figureDoi="http://doi.org/10.5281/zenodo.4949118" httpUri="https://zenodo.org/record/4949118/files/figure.png" pageId="79" pageNumber="80">Figs. 16B</figureCitation>
,
<figureCitation id="12E72A5ABD613039FF2E1E904C79E69A" box="[214,265,1157,1182]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 13, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="79" pageNumber="80">33B</figureCitation>
). Ecdysial tear uncertain anteriorly; present along prothoracic extension.
<emphasis id="B8A8EACDBD613039FBAE1E9049E6E69A" bold="true" box="[1110,1174,1157,1182]" pageId="79" pageNumber="80">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD613039FA961E904DBAE6C6" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="79" pageNumber="80">Fig. 21A</figureCitation>
), with partial ventral line of weakness, without dorsomedial tubercle, without central dome; dorsolateral cephalic sclerite (as in
<figureCitation id="12E72A5ABD613039FE621ED84C8FE6E2" box="[410,511,1229,1254]" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="79" pageNumber="80">Fig. 13H</figureCitation>
) fused to scutum, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD613039FEDE1EE44CFBE70E" box="[294,395,1265,1290]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">Fig. 26E</figureCitation>
) with mandible well-developed, lacinia absent; palpus extending posterior to posterolateral margin of labium; labium entire (not divided medially); apex of antenna (
<figureCitation id="12E72A5ABD613039FC291F004946E72A" box="[977,1078,1301,1326]" captionStart="FIGURE 38" captionStartId="283.[151,250,1936,1958]" captionTargetBox="[195,1395,204,1921]" captionTargetId="figure-66@283.[186,1417,186,1925]" captionTargetPageId="283" captionText="FIGURE 38. Legs, wings and apices of the antennae, in ventral view. A. Dibezzia debenhamae (reconstructed from distorted specimen). B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea sp. (from Ottawa, ON, Canada). D. Macropeza natalensis. E. Macropeza sp. (from Laos). F. Mallochohelea caudellii." figureDoi="http://doi.org/10.5281/zenodo.4949194" httpUri="https://zenodo.org/record/4949194/files/figure.png" pageId="79" pageNumber="80">Fig. 38A</figureCitation>
) posterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum), narrowed posteriorly; sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD613039FF671F484C60E772" box="[159,272,1373,1398]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="79" pageNumber="80">Fig. 21A</figureCitation>
)—1 elongate seta, 1 campaniform sensillum; dorsolateral cephalic sclerite sensilla—1 seta, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD613039FD9F1F944FBAE79E" box="[615,714,1409,1434]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">Fig. 26E</figureCitation>
)—2 slender setae; oculars (
<figureCitation id="12E72A5ABD613039FBF11F94491DE79E" box="[1033,1133,1409,1434]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">Fig. 26E</figureCitation>
)—2 setae, 1 campaniform sensillum.
<emphasis id="B8A8EACDBD613039FEEC1FB04C1DE7BA" bold="true" box="[276,365,1445,1470]" pageId="79" pageNumber="80">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD613039FD741FB04F83E7BA" box="[652,755,1445,1470]" captionStart="FIGURE 26" captionStartId="271.[151,250,1955,1977]" captionTargetBox="[190,1393,203,1919]" captionTargetId="figure-66@271.[189,1408,203,1924]" captionTargetPageId="271" captionText="FIGURE 26. Posterior portion of heads, in ventral view. A. Echinohelea lanei (reconstructed from distorted specimen). B. Clinohelea curriei. C. Heteromyia wokei. D. Pellucidomyia leei. E. Dibezzia debenhamae (reconstructed from distorted specimen). F. Hebetula tonnoiri. G. Jenkinshelea magnipennis H. Macropeza bayeri." figureDoi="http://doi.org/10.5281/zenodo.4949162" httpUri="https://zenodo.org/record/4949162/files/figure.png" pageId="79" pageNumber="80">Fig. 26E</figureCitation>
) wide, well-developed, extending from palpus but narrow dorsolaterally, extent to antenna uncertain; mesonotum without tubercles, not extending posteromedially, not dividing metathorax medially (
<figureCitation id="12E72A5ABD613039FDFA1FF84F01E402" box="[514,625,1517,1542]" captionStart="FIGURE 51" captionStartId="296.[151,250,1944,1966]" captionTargetBox="[184,1397,192,1912]" captionTargetId="figure-64@296.[174,1406,183,1922]" captionTargetPageId="296" captionText="FIGURE 51. Metathoraces and tergites 1, in dorsal view. A. Dibezzia debenhamae. B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea magnipennis. D. Macropeza natalensis. E. Mallochohelea caudellii. F. Probezzia sabroskyi." figureDoi="http://doi.org/10.5281/zenodo.4949230" httpUri="https://zenodo.org/record/4949230/files/figure.png" pageId="79" pageNumber="80">Fig. 51A</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD613039FC9D1FF84EA0E402" box="[869,976,1517,1542]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="79" pageNumber="80">Fig. 45G</figureCitation>
) length/width = 3.113.61, moderately elongate, with blunt apex, somewhat flattened apically, with pores closely abutting at apex of respiratory organ, arranged in single row, outer surface with plates with very fine spicules, without pedicel, base with elongate posteromedial apodeme, membranous base of respiratory organ short, tracheal tube straight to slightly curved along length, with spirals restricted to base, plates to 3/4 length; wing (
<figureCitation id="12E72A5ABD613039FC581C68497CE492" box="[928,1036,1661,1686]" captionStart="FIGURE 38" captionStartId="283.[151,250,1936,1958]" captionTargetBox="[195,1395,204,1921]" captionTargetId="figure-66@283.[186,1417,186,1925]" captionTargetPageId="283" captionText="FIGURE 38. Legs, wings and apices of the antennae, in ventral view. A. Dibezzia debenhamae (reconstructed from distorted specimen). B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea sp. (from Ottawa, ON, Canada). D. Macropeza natalensis. E. Macropeza sp. (from Laos). F. Mallochohelea caudellii." figureDoi="http://doi.org/10.5281/zenodo.4949194" httpUri="https://zenodo.org/record/4949194/files/figure.png" pageId="79" pageNumber="80">Fig. 38A</figureCitation>
) without apical tubercle or angle, separated medially by fore-, midlegs; halter apex and hind leg (as in
<figureCitation id="12E72A5ABD613039FC371CB44948E4BE" box="[975,1080,1697,1722]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 13, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="79" pageNumber="80">Fig. 33A</figureCitation>
) broadly abutting; halter apex abutting anterolateral knob-like extension of tergite 2; legs (
<figureCitation id="12E72A5ABD613039FCCA1CD04EE8E4DA" box="[818,920,1733,1758]" captionStart="FIGURE 38" captionStartId="283.[151,250,1936,1958]" captionTargetBox="[195,1395,204,1921]" captionTargetId="figure-66@283.[186,1417,186,1925]" captionTargetPageId="283" captionText="FIGURE 38. Legs, wings and apices of the antennae, in ventral view. A. Dibezzia debenhamae (reconstructed from distorted specimen). B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea sp. (from Ottawa, ON, Canada). D. Macropeza natalensis. E. Macropeza sp. (from Laos). F. Mallochohelea caudellii." figureDoi="http://doi.org/10.5281/zenodo.4949194" httpUri="https://zenodo.org/record/4949194/files/figure.png" pageId="79" pageNumber="80">Fig. 38A</figureCitation>
) with lateral margin of foreleg near midlength of wing evenly curved; hind leg visible at lateral margin of wing (as in
<figureCitation id="12E72A5ABD613039FC411CFC496FE506" box="[953,1055,1769,1794]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 13, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="79" pageNumber="80">Fig. 33C</figureCitation>
); with apex of foreleg moderately anterior to apex of midleg; apex of hind leg abutting apex of midleg laterally; sensilla: anteromedials—2 elongate setae (as in
<figureCitation id="12E72A5ABD613039FED81D244CC9E54E" box="[288,441,1841,1866]" captionStart="FIGURE 31" captionStartId="276.[151,250,1885,1907]" captionTargetBox="[177,1424,178,1864]" captionTargetId="figure-106@276.[177,1424,178,1864]" captionTargetPageId="276" captionText="FIGURE 31. Dorsal setae from left side of thoraces; anteromedial setae. A. Homohelea delanoe. B. Leehelea hispida. C. Sphaeromias longipennis. D. Xenohelea galatea. E. Bezzia nobilis. F. Clastrieromyia dycei. G. Pachyhelea pachymera. H. Palpomyia jonesi. I. Phaenobezzia mashonensis. J. Paryphoconus oliveirai. K. Echinohelea lanei, partial lateral view (distorted slide specimen). L. Palpomyia (JAD685), dorsal view. M. Palpomyia (JAD685), dorsolateral view. N. Probezzia seminigra, dorsal view of left side." figureDoi="http://doi.org/10.5281/zenodo.4949178" httpUri="https://zenodo.org/record/4949178/files/figure.png" pageId="79" pageNumber="80">Figs. 31LM</figureCitation>
); anterolaterals—1 moderately long seta; dorsal setae (
<figureCitation id="12E72A5ABD613039FBD41D2449FBE54E" box="[1068,1163,1841,1866]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="79" pageNumber="80">Fig. 30J</figureCitation>
)—D-1-T, D-2-T, D-4-T setae, D-5-T broken or absent, D-3-T campaniform sensillum, D-3-T lateral to D-4-T; supraalar 2—campaniform sensillum; metathoracics (
<figureCitation id="12E72A5ABD613039FE431D6C4F50E596" box="[443,544,1913,1938]" captionStart="FIGURE 51" captionStartId="296.[151,250,1944,1966]" captionTargetBox="[184,1397,192,1912]" captionTargetId="figure-64@296.[174,1406,183,1922]" captionTargetPageId="296" captionText="FIGURE 51. Metathoraces and tergites 1, in dorsal view. A. Dibezzia debenhamae. B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea magnipennis. D. Macropeza natalensis. E. Mallochohelea caudellii. F. Probezzia sabroskyi." figureDoi="http://doi.org/10.5281/zenodo.4949230" httpUri="https://zenodo.org/record/4949230/files/figure.png" pageId="79" pageNumber="80">Fig. 51A</figureCitation>
)—1 campaniform sensillum; M-3-T distant from margin of metathorax (at least 1/3 length of metathorax).
<emphasis id="B8A8EACDBD613039FE2B1D884F37E5B2" bold="true" box="[467,583,1949,1974]" pageId="79" pageNumber="80">Abdomen</emphasis>
: without pigmentation pattern, segment 2 as wide or slightly wider than segment 3, segments with undivided, thin to thick setae, with rounded to pointed, short to moderately elongate tubercles, tergites or sternites entire, each without membranous disc; segment 9 (
<figureCitation id="12E72A5ABD7E3026FBCA1A8249FEE2B4" box="[1074,1166,151,176]" captionStart="FIGURE 75" captionStartId="320.[151,250,1792,1814]" captionTargetBox="[220,1359,193,1769]" captionTargetId="figure-191@320.[210,1379,193,1773]" captionTargetPageId="320" captionText="FIGURE 75. Abdominal segments 9. A. Parabezzia huberti, female, in ventral view (from Grogan &amp; Wirth 1977a). B. Echinohelea lanei (from Wirth 1994b). C. Clinohelea bimaculata, female, in ventral view (from Wirth &amp; Grogan 1979). D. Heteromyia wokei, male, in dorsal and ventral view (from Wirth &amp; Grogan 1977). E. Heteromyia wokei, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). F. Heteromyia clavata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). G. Pellucidomyia leei, male, in dorsal view (from Elson-Harris &amp; Kettle 1986). H. Pellucidomyia geari, male, in ventral view (from de Meillon &amp; Wirth 1981). I. Dibezzia debenhamae, male, in dorsal and ventral view (shagreen not shown). J. Hebetula tonnoiri, female, in dorsal view (from Elson-Harris 1987). K. Jenkinshelea albaria, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Jenkinshelea albaria, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949320" httpUri="https://zenodo.org/record/4949320/files/figure.png" pageId="80" pageNumber="81">Fig. 75I</figureCitation>
) not strongly modified, terminal processes closely approximated basally, each projecting nearly posteriorly, elongate, slender, tapering to pointed apex; sensilla: tergite 1 (
<figureCitation id="12E72A5ABD7E3026FDFC1ACA4F1AE2FC" box="[516,618,223,248]" captionStart="FIGURE 51" captionStartId="296.[151,250,1944,1966]" captionTargetBox="[184,1397,192,1912]" captionTargetId="figure-64@296.[174,1406,183,1922]" captionTargetPageId="296" captionText="FIGURE 51. Metathoraces and tergites 1, in dorsal view. A. Dibezzia debenhamae. B. Hebetula tonnoiri (reconstructed from distorted specimen). C. Jenkinshelea magnipennis. D. Macropeza natalensis. E. Mallochohelea caudellii. F. Probezzia sabroskyi." figureDoi="http://doi.org/10.5281/zenodo.4949230" httpUri="https://zenodo.org/record/4949230/files/figure.png" pageId="80" pageNumber="81">Fig. 51A</figureCitation>
) with 6 setae, 2 campaniform sensilla, including 3 lateral sensilla, D-2-I, D-3-I closely approximated, D-7-I situated anteriorly near D-3-I; segment 4 (
<figureCitation id="12E72A5ABD7E3026FBFC1B114917E319" box="[1028,1127,260,285]" captionStart="FIGURE 64" captionStartId="309.[151,250,1979,2001]" captionTargetBox="[171,1416,189,1957]" captionTargetId="figure-49@309.[161,1426,183,1967]" captionTargetPageId="309" captionText="FIGURE 64. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Pellucidomyia leei. B. Dibezzia debenhamae (reconstructed from distorted and dirty specimen). C. Hebetula tonnoiri." figureDoi="http://doi.org/10.5281/zenodo.4949278" httpUri="https://zenodo.org/record/4949278/files/figure.png" pageId="80" pageNumber="81">Fig. 64B</figureCitation>
)—D-2-IV, D-3-IV short to moderately elongate setae, D-2-IV on short tubercle; D-8-IV, D-9-IV moderately elongate setae, D-5-IV, D-7-IV absent (or perhaps not visible); D-8-IV, D-9-IV on elongate, single tubercle, posterior dorsal sensilla in transverse row, arranged medially to laterally: D-4-IV, D-8-IV, D-9-IV; L-1-IV short seta without tubercle, just anterior of base of tubercle with L-2-IV; L-2-IV, L-3-IV, L-4-IV moderately elongate setae on elongate tubercles, V-5-IV, V-6- IV, V-7-IV moderately elongate setae, without tubercles, all closely approximated; segment 8 without D-3-VIII, without L-1-VIII; segment 9 (
<figureCitation id="12E72A5ABD7E3026FE1E1BC94F4EE3F1" box="[486,574,476,501]" captionStart="FIGURE 75" captionStartId="320.[151,250,1792,1814]" captionTargetBox="[220,1359,193,1769]" captionTargetId="figure-191@320.[210,1379,193,1773]" captionTargetPageId="320" captionText="FIGURE 75. Abdominal segments 9. A. Parabezzia huberti, female, in ventral view (from Grogan &amp; Wirth 1977a). B. Echinohelea lanei (from Wirth 1994b). C. Clinohelea bimaculata, female, in ventral view (from Wirth &amp; Grogan 1979). D. Heteromyia wokei, male, in dorsal and ventral view (from Wirth &amp; Grogan 1977). E. Heteromyia wokei, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). F. Heteromyia clavata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1977). G. Pellucidomyia leei, male, in dorsal view (from Elson-Harris &amp; Kettle 1986). H. Pellucidomyia geari, male, in ventral view (from de Meillon &amp; Wirth 1981). I. Dibezzia debenhamae, male, in dorsal and ventral view (shagreen not shown). J. Hebetula tonnoiri, female, in dorsal view (from Elson-Harris 1987). K. Jenkinshelea albaria, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Jenkinshelea albaria, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949320" httpUri="https://zenodo.org/record/4949320/files/figure.png" pageId="80" pageNumber="81">Fig. 75I</figureCitation>
)—with D-5-IX elongate seta, D-6-IX campaniform sensillum.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD7E3026FF3F1BEA499BE064" pageId="80" pageNumber="81" type="distribution">
<paragraph id="8A6336DFBD7E3026FF3F1BEA499BE064" blockId="80.[151,1436,151,1184]" pageId="80" pageNumber="81">
<emphasis id="B8A8EACDBD7E3026FF3F1BEA4F16E01C" bold="true" box="[199,614,511,536]" pageId="80" pageNumber="81">DISTRIBUTION AND HABITAT</emphasis>
: The genus
<taxonomicName id="4DDC4D5CBD7E3026FD071BEA4E11E01C" box="[767,865,511,536]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FD071BEA4E11E01C" box="[767,865,511,536]" italics="true" pageId="80" pageNumber="81">Dibezzia</emphasis>
</taxonomicName>
is known from five species in the Oriental and Afrotropical Regions (
<taxonomicName id="4DDC4D5CBD7E3026FE6B18314F42E039" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[403,562,548,573]" genus="Borkent" pageId="80" pageNumber="92" rank="genus" status="gen. nov.">
<bibRefCitation id="EE4D4B2EBD7E3026FE6B18314F42E039" author="Borkent" box="[403,562,548,573]" firstAuthor="Borkent" pageId="80" pageNumber="81" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
</taxonomicName>
, additional species below). The two species were reared from either &quot;bamboo peat&quot; (
<bibRefCitation id="EE4D4B2EBD7E3026FF2718524CE6E064" author="Johannsen" box="[223,406,583,608]" firstAuthor="Johannsen" pageId="80" pageNumber="81" pagination="403 - 448" refId="ref186650" refString="Johannsen, O. A. (1932) Ceratopogoninae from the Malayan subregion of the Dutch East Indies. Archiv fur Hydrobiologie, Supplement, 9, 403 - 448, pls. 4 - 8." type="journal article" year="1932">Johannsen 1932</bibRefCitation>
) in
<collectingCountry id="F2CB764FBD7E3026FE3A18524F5FE064" box="[450,559,583,608]" name="Indonesia" pageId="80" pageNumber="81">Indonesia</collectingCountry>
or a tree hole in
<collectingCountry id="F2CB764FBD7E3026FD1518524E2FE064" box="[749,863,583,608]" name="Singapore" pageId="80" pageNumber="81">Singapore</collectingCountry>
(
<bibRefCitation id="EE4D4B2EBD7E3026FC94185249AEE064" author="Wirth &amp; Ratanaworabhan" box="[876,1246,583,608]" firstAuthor="Wirth" pageId="80" pageNumber="81" pagination="287 - 295" refId="ref200234" refString="Wirth, W. W. &amp; Ratanaworabhan, N. C. (1981 a) The Oriental species of the genus Dibezzia Kieffer (Diptera Ceratopogonidae). Proceedings of the Entomological Society of Washington, 83, 287 - 295." type="journal article" year="1981" yearSuffix="a">Wirth &amp; Ratanaworabhan 1981a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD7E3026FF3F18794EFCE679" pageId="80" pageNumber="81" type="discussion">
<paragraph id="8A6336DFBD7E3026FF3F18794942E1E9" blockId="80.[151,1436,151,1184]" pageId="80" pageNumber="81">
<emphasis id="B8A8EACDBD7E3026FF3F18794F5BE081" bold="true" box="[199,555,620,645]" pageId="80" pageNumber="81">TAXONOMIC DISCUSSION</emphasis>
: The pupae of this genus are known from two species (
<tableCitation id="C75E0364BD7E3026FB6E18794865E081" box="[1174,1301,620,645]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" pageId="80" pageNumber="81" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 23</tableCitation>
). Originally described as
<taxonomicName id="4DDC4D5CBD7E3026FED4189A4F16E0A3" box="[300,614,655,680]" class="Insecta" family="Ceratopogonidae" genus="Johannsenomyia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="B8A8EACDBD7E3026FED4189A4F16E0A3" box="[300,614,655,680]" italics="true" pageId="80" pageNumber="81">Johannsenomyia prominens</emphasis>
</taxonomicName>
by
<bibRefCitation id="EE4D4B2EBD7E3026FD6C189A4E2FE0AC" author="Johannsen" box="[660,863,655,680]" firstAuthor="Johannsen" pageId="80" pageNumber="81" pagination="403 - 448" refId="ref186650" refString="Johannsen, O. A. (1932) Ceratopogoninae from the Malayan subregion of the Dutch East Indies. Archiv fur Hydrobiologie, Supplement, 9, 403 - 448, pls. 4 - 8." type="journal article" year="1932">Johannsen (1932)</bibRefCitation>
,
<bibRefCitation id="EE4D4B2EBD7E3026FC9518854931E0AC" author="Mayer" box="[877,1089,655,680]" firstAuthor="Mayer" page="244" pageId="80" pageNumber="81" pagination="205 - 288" refId="ref190389" refString="Mayer, K. (1934 a) Die Metamorphose der Ceratopogonidae (Dipt.). Ein Beitrag zur Morphologie, Systematik. Okologie und Biologie der Jugendstadien dieser Dipterenfamilie. Archiv fur Naturgeschicthe, 3, 205 - 288." type="journal article" year="1934" yearSuffix="a">Mayer (1934a:244</bibRefCitation>
,
<bibRefCitation id="EE4D4B2EBD7E3026FBB7189A49E4E0AC" author="Mayer" box="[1103,1172,655,680]" firstAuthor="Mayer" pageId="80" pageNumber="81" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">1934c</bibRefCitation>
) described the pupa of this Indonesian species as (?)
<taxonomicName id="4DDC4D5CBD7E3026FE1E18A14F94E0C8" box="[486,740,692,717]" class="Insecta" family="Ceratopogonidae" genus="Homohelea" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="B8A8EACDBD7E3026FE1E18A14F94E0C8" box="[486,740,692,717]" italics="true" pageId="80" pageNumber="81">Homohelea prominens</emphasis>
</taxonomicName>
, based on the pupal exuviae from which the male
<typeStatus id="5567887DBD7E3026FAE318A1480DE0C9" box="[1307,1405,692,717]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
of this species was reared.
<bibRefCitation id="EE4D4B2EBD7E3026FE4F18C24F3DE0F4" author="Wirth" box="[439,589,727,752]" firstAuthor="Wirth" pageId="80" pageNumber="81" pagination="346 - 388" refId="ref199123" refString="Wirth, W. W. (1973) Family Ceratopogonidae. In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A Catalog of the Oriental Region. Vol. 1. Suborder Nematocera. University of Hawaii, Honolulu, pp. 346 - 388" type="book chapter" year="1973">Wirth (1973)</bibRefCitation>
placed the species in
<taxonomicName id="4DDC4D5CBD7E3026FCA118C24971E0F4" box="[857,1025,727,752]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FCA118C24971E0F4" box="[857,1025,727,752]" italics="true" pageId="80" pageNumber="81">Mallochohelea</emphasis>
</taxonomicName>
as a new combination but did not identify it as such or give any justification for doing so and this is where it has remained since. The descriptions by
<bibRefCitation id="EE4D4B2EBD7E3026FF6F19354C4CE13C" author="Mayer" box="[151,316,799,824]" firstAuthor="Mayer" pageId="80" pageNumber="81" pagination="205 - 288" refId="ref190389" refString="Mayer, K. (1934 a) Die Metamorphose der Ceratopogonidae (Dipt.). Ein Beitrag zur Morphologie, Systematik. Okologie und Biologie der Jugendstadien dieser Dipterenfamilie. Archiv fur Naturgeschicthe, 3, 205 - 288." type="journal article" year="1934" yearSuffix="a">Mayer (1934a</bibRefCitation>
,
<bibRefCitation id="EE4D4B2EBD7E3026FEB3190A4CE3E13C" author="Mayer" box="[331,403,799,824]" firstAuthor="Mayer" pageId="80" pageNumber="81" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">1934c</bibRefCitation>
), however, clearly indicate the presence of the elongate seta at the base of the terminal process, a feature of
<taxonomicName id="4DDC4D5CBD7E3026FE6C19514C86E159" box="[404,502,836,861]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FE6C19514C86E159" box="[404,502,836,861]" italics="true" pageId="80" pageNumber="81">Dibezzia</emphasis>
</taxonomicName>
and unique within the
<taxonomicName id="4DDC4D5CBD7E3026FCEA19514EA5E159" box="[786,981,836,861]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Ceratopogoninae">Ceratopogoninae</taxonomicName>
. In addition,
<bibRefCitation id="EE4D4B2EBD7E3026FB8F19514853E159" author="Mayer" box="[1143,1315,836,861]" firstAuthor="Mayer" pageId="80" pageNumber="81" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">Mayer (1934c)</bibRefCitation>
noted the presence of &quot;breiten Schuppen&quot; (broad scales) on the surface of the respiratory organ, which is what the fine scales appear as in
<taxonomicName id="4DDC4D5CBD7E3026FEDB19984CA2E1A1" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1981" box="[291,466,908,933]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="debenhamae">
<emphasis id="B8A8EACDBD7E3026FEDB19984CA2E1A1" box="[291,466,908,933]" italics="true" pageId="80" pageNumber="81">D. debenhamae</emphasis>
</taxonomicName>
. This feature is unique within the
<taxonomicName id="4DDC4D5CBD7E3026FCA919994E96E1A1" box="[849,998,908,933]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Heteromyiini">Heteromyiini</taxonomicName>
+
<taxonomicName id="4DDC4D5CBD7E3026FBFC199949EBE1A1" box="[1028,1179,908,933]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Sphaeromiini">Sphaeromiini</taxonomicName>
<emphasis id="B8A8EACDBD7E3026FB5A199849AFE1A1" box="[1186,1247,908,933]" italics="true" pageId="80" pageNumber="81">s. lat.</emphasis>
+
<taxonomicName id="4DDC4D5CBD7E3026FB04199948F5E1A1" box="[1276,1413,908,933]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Palpomyiini">Palpomyiini</taxonomicName>
+
<taxonomicName id="4DDC4D5CBD7E3026FF6F19BA4C50E1CC" box="[151,288,943,968]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Stenoxenini">Stenoxenini</taxonomicName>
. I therefore remove the name from
<taxonomicName id="4DDC4D5CBD7E3026FD5419BA4E24E1CC" box="[684,852,943,968]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FD5419BA4E24E1CC" box="[684,852,943,968]" italics="true" pageId="80" pageNumber="81">Mallochohelea</emphasis>
</taxonomicName>
and consider it a member of
<taxonomicName id="4DDC4D5CBD7E3026FB6319BA498DE1CC" box="[1179,1277,943,968]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FB6319BA498DE1CC" box="[1179,1277,943,968]" italics="true" pageId="80" pageNumber="81">Dibezzia</emphasis>
</taxonomicName>
as follows:
<taxonomicName id="4DDC4D5CBD7E3026FF6F19C14F0BE1E9" authority="(Johannsen) 1932: 435" baseAuthorityName="Johannsen" baseAuthorityPageNumber="435" baseAuthorityYear="1932" box="[151,635,980,1005]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="B8A8EACDBD7E3026FF6F19C14C05E1E8" box="[151,373,980,1005]" italics="true" pageId="80" pageNumber="81">Dibezzia prominens</emphasis>
(Johannsen) 1932: 435
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD7E3026FD7319C14E37E1E9" box="[651,839,980,1005]" class="Insecta" family="Ceratopogonidae" genus="Johannsenomyia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD7E3026FD7319C14E37E1E9" box="[651,839,980,1005]" italics="true" pageId="80" pageNumber="81">Johannsenomyia</emphasis>
</taxonomicName>
).
<emphasis id="B8A8EACDBD7E3026FCA719C14942E1E9" bold="true" box="[863,1074,980,1005]" pageId="80" pageNumber="81">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD7E3026FF3F19E24EFCE679" blockId="80.[151,1436,151,1184]" pageId="80" pageNumber="81">
<taxonomicName id="4DDC4D5CBD7E3026FF3F19E24CDCE60B" box="[199,428,1015,1040]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="B8A8EACDBD7E3026FF3F19E24CDCE60B" box="[199,428,1015,1040]" italics="true" pageId="80" pageNumber="81">Dibezzia prominens</emphasis>
</taxonomicName>
, as illustrated by
<bibRefCitation id="EE4D4B2EBD7E3026FD6919ED4E32E614" author="Mayer" box="[657,834,1015,1040]" firstAuthor="Mayer" pageId="80" pageNumber="81" pagination="166 - 202" refId="ref190459" refString="Mayer, K. (1934 c) Ceratopogoniden-Metamorphosen (C. Intermediae und C. Vermiformes) der Deutschen Limnologischen Sunda-Expedition. Archiv fur Hydrobiologie, Supplement 13, 166 - 202." type="journal article" year="1934" yearSuffix="c">Mayer (1934c)</bibRefCitation>
, has D-5-IV represented as a small campaniform sensillum or perhaps a very small seta and D-7-IV located posterolateral to D-3-IV (like other members of
<taxonomicName id="4DDC4D5CBD7E3026FF6F1E2A4C5CE65C" box="[151,300,1087,1112]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Heteromyiini">Heteromyiini</taxonomicName>
+
<taxonomicName id="4DDC4D5CBD7E3026FEB41E2A4C93E65C" box="[332,483,1087,1112]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Sphaeromiini">Sphaeromiini</taxonomicName>
<emphasis id="B8A8EACDBD7E3026FE131E544F59E65C" box="[491,553,1087,1112]" italics="true" pageId="80" pageNumber="81">s. lat.</emphasis>
+
<taxonomicName id="4DDC4D5CBD7E3026FDB11E2A4FA2E65C" box="[585,722,1087,1112]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Palpomyiini">Palpomyiini</taxonomicName>
+
<taxonomicName id="4DDC4D5CBD7E3026FD0A1E2A4E0BE65C" box="[754,891,1087,1112]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Stenoxenini">Stenoxenini</taxonomicName>
; see character 64). I could not find either on the single, dirty, and distorted specimen of
<taxonomicName id="4DDC4D5CBD7E3026FDA81E704E70E679" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1981" box="[592,768,1124,1149]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="debenhamae">
<emphasis id="B8A8EACDBD7E3026FDA81E704E70E679" box="[592,768,1124,1149]" italics="true" pageId="80" pageNumber="81">D. debenhamae</emphasis>
</taxonomicName>
I examined.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD7E3026FF3F1E924801E6A4" box="[199,1393,1159,1184]" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="8A6336DFBD7E3026FF3F1E924801E6A4" blockId="80.[151,1436,151,1184]" box="[199,1393,1159,1184]" pageId="80" pageNumber="81">
<emphasis id="B8A8EACDBD7E3026FF3F1E924F71E6A4" bold="true" box="[199,513,1159,1184]" pageId="80" pageNumber="81">MATERIAL EXAMINED</emphasis>
:
<taxonomicName id="4DDC4D5CBD7E3026FDE81E9C4FCFE6A4" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1981" box="[528,703,1159,1184]" class="Insecta" family="Ceratopogonidae" genus="Dibezzia" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="debenhamae">
<emphasis id="B8A8EACDBD7E3026FDE81E9C4FCFE6A4" box="[528,703,1159,1184]" italics="true" pageId="80" pageNumber="81">D. debenhamae</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD7E3026FC561E92497CE6A4" box="[942,1036,1159,1184]" pageId="80" pageNumber="81" type="allotype">allotype</typeStatus>
),
<collectingCountry id="F2CB764FBD7E3026FBD81E9249E4E6A4" box="[1056,1172,1159,1184]" name="Singapore" pageId="80" pageNumber="81">Singapore</collectingCountry>
,
<date id="FE62101FBD7E3026FB671E92498FE6A4" box="[1183,1279,1159,1184]" pageId="80" pageNumber="81" value="1959-06">VI-1959</date>
(USNM).
</paragraph>
</subSubSection>
</treatment>
</document>