treatments-xml/data/02/75/87/027587C9BD14304DFDB81E2F4C43E6EC.xml

<document id="99A657B79B6B1EBB246A5CB782FE24B0" ID-CLB-Dataset="3916" ID-DOI="10.11646/zootaxa.3879.1.1" ID-GBIF-Dataset="4b976ba3-1a7b-4dd5-8afb-630b7cc8a695" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949051" ID-ZooBank="6423894B-97D9-4286-ABB9-D4AF072B57FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623702926238" checkinUser="felipe" docAuthor="Borkent, Art" docDate="2014" docId="027587C9BD14304DFDB81E2F4C43E6EC" docLanguage="en" docName="zt03879p327.pdf.imf" docOrigin="Zootaxa 3879 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Baeodasymyia Clastrier &amp; Raccurt" docType="treatment" docVersion="12" lastPageNumber="60" masterDocId="FE4CFFB1BD2E3076FFF81A154D70E204" masterDocTitle="The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera" masterLastPageNumber="327" masterPageNumber="1" pageNumber="59" updateTime="1699015668130" updateUser="ExternalLinkService">
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<mods:title id="569C7024FE7B25F55C768DDCF837EC67">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<mods:namePart id="D52C9F1BBD55CD1CFA9DF84175459D0B">Borkent, Art</mods:namePart>
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<treatment id="027587C9BD14304DFDB81E2F4C43E6EC" ID-DOI="http://doi.org/10.5281/zenodo.5589813" ID-GBIF-Taxon="189161062" ID-Zenodo-Dep="5589813" LSID="urn:lsid:plazi:treatment:027587C9BD14304DFDB81E2F4C43E6EC" httpUri="http://treatment.plazi.org/id/027587C9BD14304DFDB81E2F4C43E6EC" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<subSubSection id="C2C66554BD14304CFDB81E2F4E82E651" box="[576,1010,1082,1109]" pageId="58" pageNumber="59" type="nomenclature">
<paragraph id="8A6336DFBD14304CFDB81E2F4E82E651" blockId="58.[468,1118,1082,1142]" box="[576,1010,1082,1109]" pageId="58" pageNumber="59">
<taxonomicName id="4DDC4D5CBD14304CFDB81E2F4E82E651" ID-CoL="62FHH" authority="Clastrier &amp; Raccurt" authorityName="Clastrier &amp; Raccurt" box="[576,1010,1082,1109]" class="Insecta" family="Ceratopogonidae" genus="Baeodasymyia" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD14304CFDB81E2F4E82E651" bold="true" box="[576,1010,1082,1109]" pageId="58" pageNumber="59">
<emphasis id="B8A8EACDBD14304CFDB81E2F4F9EE650" bold="true" box="[576,750,1082,1108]" italics="true" pageId="58" pageNumber="59">Baeodasymyia</emphasis>
Clastrier &amp; Raccurt
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD14304CFE2C1E48492EE672" box="[468,1118,1117,1142]" pageId="58" pageNumber="59" type="description">
<paragraph id="8A6336DFBD14304CFE2C1E48492EE672" blockId="58.[468,1118,1082,1142]" box="[468,1118,1117,1142]" pageId="58" pageNumber="59">
(
<figureCitation id="12E72A5ABD14304CFE251E484F4AE672" box="[477,570,1117,1142]" captionStart="FIGURE 2" captionStartId="247.[151,250,1829,1851]" captionTargetBox="[194,1399,181,1811]" captionTargetId="figure-139@247.[194,1399,181,1811]" captionTargetPageId="247" captionText="FIGURE 2. Pupae in unusual habitats, preservation and relative sizes. A. Forcipomyia pupae, Rancho Grande, Venezuela; photo by Steve Marshall. B. Forcipomyia pupae under bark of log of Pinus sylvestris, Poland; photo by Marek W. Kozlowski. C. Pupae of Mallochohelea sp. on emergent plants at edge of Bolean Lk., 6 km NE Falkland, British Columbia. D. Palpomyia sp. showing mounting technique on pin, 8 km E. Sicamous, British Columbia. E. Permanent slide mount of dissected adult and pupal exuviae of Mallochohelea albibasis. F. Pupal exuviae, from left to right, of Baeodasymyia michaeli, Culicoides denticulatus and Sphaeromias longipennis showing size differences within the family. Total length of Baeodasymyia michaeli = 1.2 mm, of Sphaeromias = 7.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4949061" httpUri="https://zenodo.org/record/4949061/files/figure.png" pageId="58" pageNumber="59">Figs. 2F</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFDBD1E484F08E672" box="[581,632,1117,1142]" captionStart="FIGURE 19" captionStartId="264.[151,250,1782,1804]" captionTargetBox="[215,1399,189,1744]" captionTargetId="figure-155@264.[176,1410,183,1764]" captionTargetPageId="264" captionText="FIGURE 19. Dorsal apotomes, anterior view. A. Culicoides circumscriptus (from Kettle &amp; Lawson 1952). B. Culicoides cubitalis (from Kettle &amp; Lawson 1952). C. Culicoides truncorum (from Kettle &amp; Lawson 1952). D. Culicoides chiopterus (from Jamnback 1965). E. Paradasyhelea minuta (from Kettle &amp; Elson 1975a). F. Paradasyhelea albipunctata (from Elson- Harris &amp; Kettle 1985a). G. Paradasyhelea reyei (from Elson-Harris &amp; Kettle 1985a). H. Ceratopogon boomerangus (from Borkent &amp; Grogan 1995). I. Ceratopogon abstrusus (from Borkent &amp; Grogan 1995). J. Ceratopogon arcanus (from Borkent &amp; Grogan 1995). K. Brachypogon artemis (from Debenham 1991). L. Alluaudomyia appendiculata (from Elson-Harris &amp; Kettle 1985b). M. Alluaudomyia fumosipennis (from Elson-Harris &amp; Kettle 1985b). N. Baeodasymyia michaeli (from Borkent &amp; Craig 1999). O. Austrohelea campbellensis. P. Stilobezzia rabelloi (from Borkent &amp; Craig 2001)." figureDoi="http://doi.org/10.5281/zenodo.4949130" httpUri="https://zenodo.org/record/4949130/files/figure.png" pageId="58" pageNumber="59">19N</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFD7C1E484FC0E672" box="[644,688,1117,1142]" captionStart="FIGURE 24" captionStartId="269.[151,250,1949,1971]" captionTargetBox="[195,1401,210,1925]" captionTargetId="figure-63@269.[186,1405,208,1929]" captionTargetPageId="269" captionText="FIGURE 24. Posterior portion of heads, in ventral view. A. Culicoides guttipennis. B. Paradasyhelea albipunctata. C. Ceratopogon arcanus. D. Brachypogon sp. (Flag Glade, Pocahontas Co., West Virginia, USA). E. Alluaudomyia needhami. F. Baeodasymyia michaeli. G. Austrohelea campbellensis." figureDoi="http://doi.org/10.5281/zenodo.4949152" httpUri="https://zenodo.org/record/4949152/files/figure.png" pageId="58" pageNumber="59">24F</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFD431E484F9EE672" box="[699,750,1117,1142]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="58" pageNumber="59">29Q</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFD011E484E5DE672" box="[761,813,1117,1142]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="58" pageNumber="59">32K</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFCCF1E484E17E672" box="[823,871,1117,1142]" captionStart="FIGURE 35" captionStartId="280.[151,250,1920,1942]" captionTargetBox="[207,1370,219,1888]" captionTargetId="figure-83@280.[197,1391,206,1892]" captionTargetPageId="280" captionText="FIGURE 35. Legs, wings and apices of the antennae, in ventral view. A. Paradasyhelea sp. (from Borlase Stream, Lake Rotoiti, New Zealand). B. Ceratopogon sp. nr. abstrusus. C. Brachypogon sp. (from Spanish Lake, 6 km E. Falkland, BC, Canada). D. Alluaudomyia needhami. E. Baeodasymyia michaeli. F. Austrohelea campbellensis (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949188" httpUri="https://zenodo.org/record/4949188/files/figure.png" pageId="58" pageNumber="59">35E</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFC8B1E484ED4E672" box="[883,932,1117,1142]" captionStart="FIGURE 44" captionStartId="289.[151,250,1730,1752]" captionTargetBox="[238,1376,198,1685]" captionTargetId="figure-184@289.[205,1387,192,1716]" captionTargetPageId="289" captionText="FIGURE 44. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Alluaudomyia latipennis (from Debenham 1971). B. Alluaudomyia footei (from Wirth &amp; Grogan 1981). C. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). D. Austrohelea campbellensis. E. Stilobezzia antennalis (from Wirth &amp; Grogan 1981, modified). F. Stilobezzia glauca (from Wirth &amp; Grogan 1981). G. Stilobezzia ochracea (from Kettle &amp; Lawson 1952). H. Stilobezzia diversa (from Wirth &amp; Grogan 1981). I. Stilobezzia rabelloi (from Borkent &amp; Craig 2001). J. Stilobezzia navaiae (from Wirth &amp; Grogan 1981). K. Stilobezzia lutea (from Wirth &amp; Grogan 1981). L. Schizonyxhelea bulla (from Wirth &amp; Grogan 1981, as a Stilobezzia). M. Schizonyxhelea forattinii (from Borkent 2000a). N. Serromyia femorata (from Kettle &amp; Lawson 1952). O. Allohelea japonica. P.Monohelea hirsuta (from Wirth &amp; Grogan 1981, modified). Q. Atyphohelea macroneura (from Borkent 1998). R. Parabezzia alexanderi (from Grogan &amp; Wirth 1977a). S. Parabezzia huberti (from Grogan &amp; Wirth 1977a). T. Echinohelea lanei (from Wirth 1994b)." figureDoi="http://doi.org/10.5281/zenodo.4949208" httpUri="https://zenodo.org/record/4949208/files/figure.png" pageId="58" pageNumber="59">44C</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFC571E484EABE672" box="[943,987,1117,1142]" captionStart="FIGURE 48" captionStartId="293.[151,250,1946,1968]" captionTargetBox="[193,1408,188,1923]" captionTargetId="figure-61@293.[189,1420,188,1927]" captionTargetPageId="293" captionText="FIGURE 48. Metathoraces and tergites 1, in dorsal view. A. Culicoides haematopotus. B. Paradasyhelea minuta. C. Ceratopogon sp. nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia needhami. F. Baeodasymyia michaeli." figureDoi="http://doi.org/10.5281/zenodo.4949222" httpUri="https://zenodo.org/record/4949222/files/figure.png" pageId="58" pageNumber="59">48F</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFC1F1E48496AE672" box="[999,1050,1117,1142]" captionStart="FIGURE 59" captionStartId="304.[151,250,1975,1997]" captionTargetBox="[203,1416,215,1938]" captionTargetId="figure-57@304.[194,1426,209,1948]" captionTargetPageId="304" captionText="FIGURE 59. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). B. Austrohelea campbellensis. C. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949258" httpUri="https://zenodo.org/record/4949258/files/figure.png" pageId="58" pageNumber="59">59A</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFBDD1E484927E672" box="[1061,1111,1117,1142]" captionStart="FIGURE 74" captionStartId="319.[151,250,1737,1759]" captionTargetBox="[224,1348,183,1717]" captionTargetId="figure-195@319.[222,1356,183,1717]" captionTargetPageId="319" captionText="FIGURE 74. Abdominal segments 9. A. Baeodasymyia michaeli, female, in dorsal view (modified from Borkent &amp; Craig 1999). B. Austrohelea campbellensis, female, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). C. Stilobezzia lutea, male, in dorsal and ventral view (from Wirth &amp; Grogan 1981). D. Stilobezzia lutea, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). E. Stilobezzia rabelloi, male, in ventral view (from Borkent &amp; Craig 2001). F. Schizonyxhelea bulla, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). G. Schizonyxhelea forattinii, female, in ventral view (from Borkent 2000a). H. Serromyia sp., female (in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). I. Allohelea japonica, female, in dorsal and ventral view (shagreen not shown). J. Monohelea obscura, male, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). K. Atyphohelea macroneura, female, in dorsal view (from Borkent 1998). L. Parabezzia huberti, male, in ventral view (from Grogan &amp; Wirth 1977a)." figureDoi="http://doi.org/10.5281/zenodo.4949316" httpUri="https://zenodo.org/record/4949316/files/figure.png" pageId="58" pageNumber="59">74A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD14304CFF6F1EB04EC9E776" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8A6336DFBD14304CFF6F1EB04EC9E776" blockId="58.[151,1437,1189,2006]" pageId="58" pageNumber="59">
<emphasis id="B8A8EACDBD14304CFF6F1EB04C44E6BA" bold="true" box="[151,308,1189,1214]" pageId="58" pageNumber="59">DIAGNOSIS</emphasis>
: Only pupa of 
<taxonomicName id="4DDC4D5CBD14304CFE271EB04FEEE6BA" authorityName="Borkent" authorityYear="2014" box="[479,670,1189,1214]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with the respiratory organ elongate and with three distinct sections: a cylindrical, smooth basal portion, a cylindrical middle portion with annulations for most of its distal length, and a compressed, rounded or concave apical portion (
<figureCitation id="12E72A5ABD14304CFD311EF84E44E702" box="[713,820,1261,1286]" captionStart="FIGURE 44" captionStartId="289.[151,250,1730,1752]" captionTargetBox="[238,1376,198,1685]" captionTargetId="figure-184@289.[205,1387,192,1716]" captionTargetPageId="289" captionText="FIGURE 44. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Alluaudomyia latipennis (from Debenham 1971). B. Alluaudomyia footei (from Wirth &amp; Grogan 1981). C. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). D. Austrohelea campbellensis. E. Stilobezzia antennalis (from Wirth &amp; Grogan 1981, modified). F. Stilobezzia glauca (from Wirth &amp; Grogan 1981). G. Stilobezzia ochracea (from Kettle &amp; Lawson 1952). H. Stilobezzia diversa (from Wirth &amp; Grogan 1981). I. Stilobezzia rabelloi (from Borkent &amp; Craig 2001). J. Stilobezzia navaiae (from Wirth &amp; Grogan 1981). K. Stilobezzia lutea (from Wirth &amp; Grogan 1981). L. Schizonyxhelea bulla (from Wirth &amp; Grogan 1981, as a Stilobezzia). M. Schizonyxhelea forattinii (from Borkent 2000a). N. Serromyia femorata (from Kettle &amp; Lawson 1952). O. Allohelea japonica. P.Monohelea hirsuta (from Wirth &amp; Grogan 1981, modified). Q. Atyphohelea macroneura (from Borkent 1998). R. Parabezzia alexanderi (from Grogan &amp; Wirth 1977a). S. Parabezzia huberti (from Grogan &amp; Wirth 1977a). T. Echinohelea lanei (from Wirth 1994b)." figureDoi="http://doi.org/10.5281/zenodo.4949208" httpUri="https://zenodo.org/record/4949208/files/figure.png" pageId="58" pageNumber="59">Fig. 44C</figureCitation>
); also unique with a short foreleg, with its apex well anterior of the apex of the wing (
<figureCitation id="12E72A5ABD14304CFDF41F044F02E72E" box="[524,626,1297,1322]" captionStart="FIGURE 35" captionStartId="280.[151,250,1920,1942]" captionTargetBox="[207,1370,219,1888]" captionTargetId="figure-83@280.[197,1391,206,1892]" captionTargetPageId="280" captionText="FIGURE 35. Legs, wings and apices of the antennae, in ventral view. A. Paradasyhelea sp. (from Borlase Stream, Lake Rotoiti, New Zealand). B. Ceratopogon sp. nr. abstrusus. C. Brachypogon sp. (from Spanish Lake, 6 km E. Falkland, BC, Canada). D. Alluaudomyia needhami. E. Baeodasymyia michaeli. F. Austrohelea campbellensis (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949188" httpUri="https://zenodo.org/record/4949188/files/figure.png" pageId="58" pageNumber="59">Fig. 35E</figureCitation>
); also unique in being very small, with total body length 
<quantity id="4D249B3ABD14304CFB011F0448EDE72D" box="[1273,1437,1297,1322]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.19" metricValueMin="0.91" pageId="58" pageNumber="59" unit="mm" value="1.05" valueMax="1.19" valueMin="0.91">0.91–1.19 mm</quantity>
(only 
<taxonomicName id="4DDC4D5CBD14304CFF251F204CFAE74A" authorityName="Clastrier. Study" authorityYear="1984" box="[221,394,1333,1358]" class="Insecta" family="Ceratopogonidae" genus="Schizonyxhelea" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD14304CFF251F204CFAE74A" box="[221,394,1333,1358]" italics="true" pageId="58" pageNumber="59">Schizonyxhelea</emphasis>
</taxonomicName>
are similarly small, with a body length 
<quantity id="4D249B3ABD14304CFC921F204964E749" box="[874,1044,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="2.16" metricValueMin="1.19" pageId="58" pageNumber="59" unit="mm" value="1.675" valueMax="2.16" valueMin="1.19">1.19–2.16 mm</quantity>
; however, the unknown pupae of 
<taxonomicName id="4DDC4D5CBD14304CFF6F1F4C4C6EE776" authorityName="Grogan &amp; Wirth" authorityYear="1980" box="[151,286,1369,1394]" class="Insecta" family="Ceratopogonidae" genus="Nannohelea" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD14304CFF6F1F4C4C6EE776" box="[151,286,1369,1394]" italics="true" pageId="58" pageNumber="59">Nannohelea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4DDC4D5CBD14304CFED41F4C4CD2E776" authorityName="Wirth &amp; Blanton" authorityYear="1970" box="[300,418,1369,1394]" class="Insecta" family="Ceratopogonidae" genus="Baeohelea" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD14304CFED41F4C4CD2E776" box="[300,418,1369,1394]" italics="true" pageId="58" pageNumber="59">Baeohelea</emphasis>
</taxonomicName>
and some others will certainly be equally tiny).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD14304DFF3F1F684EBCE159" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
<paragraph id="8A6336DFBD14304DFF3F1F684EBCE159" blockId="58.[151,1437,1189,2006]" lastBlockId="59.[151,1436,151,1256]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<emphasis id="B8A8EACDBD14304CFF3F1F684CFAE792" bold="true" box="[199,394,1405,1430]" pageId="58" pageNumber="59">DESCRIPTION</emphasis>
: Total length = 
<quantity id="4D249B3ABD14304CFDBD1F684F9DE791" box="[581,749,1405,1430]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.19" metricValueMin="0.91" pageId="58" pageNumber="59" unit="mm" value="1.05" valueMax="1.19" valueMin="0.91">0.91–1.19 mm</quantity>
. Without larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of face, in some specimens against wing (as in 
<figureCitation id="12E72A5ABD14304CFB2C1FB4483BE7BE" box="[1236,1355,1441,1466]" captionStart="FIGURE 15" captionStartId="260.[151,250,1905,1927]" captionTargetBox="[165,1428,181,1886]" captionTargetId="figure-103@260.[165,1429,181,1891]" captionTargetPageId="260" captionText="FIGURE 15. Anterior portion of heads. A. Culicoides denticulatus, in anteroventral view. B. Ceratopogon nr. abstrusus, in anterior view. C. Brachypogon sp., in anterior view (from 6 km E. Falkland, BC, Canada). D. Stilobezzia antennalis, in ventral view, dorsal apotome not present. E. Allohelea japonica, in ventral view, dorsal apotome not present (distorted slide specimen). F. Atyphohelea macroneura, in ventral view, right side of head, dorsal apotome not present." figureDoi="http://doi.org/10.5281/zenodo.4949112" httpUri="https://zenodo.org/record/4949112/files/figure.png" pageId="58" pageNumber="59">Figs. 15D</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFAA31FB448E0E7BE" box="[1371,1424,1441,1466]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="58" pageNumber="59">32K</figureCitation>
). Ecdysial tear medial to base of antenna (as in 
<figureCitation id="12E72A5ABD14304CFD6E1FD04E74E7DA" box="[662,772,1477,1502]" captionStart="FIGURE 15" captionStartId="260.[151,250,1905,1927]" captionTargetBox="[165,1428,181,1886]" captionTargetId="figure-103@260.[165,1429,181,1891]" captionTargetPageId="260" captionText="FIGURE 15. Anterior portion of heads. A. Culicoides denticulatus, in anteroventral view. B. Ceratopogon nr. abstrusus, in anterior view. C. Brachypogon sp., in anterior view (from 6 km E. Falkland, BC, Canada). D. Stilobezzia antennalis, in ventral view, dorsal apotome not present. E. Allohelea japonica, in ventral view, dorsal apotome not present (distorted slide specimen). F. Atyphohelea macroneura, in ventral view, right side of head, dorsal apotome not present." figureDoi="http://doi.org/10.5281/zenodo.4949112" httpUri="https://zenodo.org/record/4949112/files/figure.png" pageId="58" pageNumber="59">Figs. 15B</figureCitation>
, 
<figureCitation id="12E72A5ABD14304CFCE81FD04E35E7DA" box="[784,837,1477,1502]" captionStart="FIGURE 79" captionStartId="324.[151,250,1861,1883]" captionTargetBox="[192,1396,201,1846]" captionTargetId="figure-148@324.[184,1406,195,1847]" captionTargetPageId="324" captionText="FIGURE 79. Evolution of ecdysial sutures and splits in pupal exuviae of Ceratopogonidae, head and anterior portion of thorax in ventral view. Numbers refer to character states of characters 2 and 8 in text. Red indicates ecdysial splits. A. Leptoconopinae. B. Forcipomyiinae. C. Dasyheleinae. D. Most Ceratopogonini. E. Most Ceratopogonini, Clinohelea. F. Allohelea, Atyphohelea, Parabezzia. G. Heteromyia, Pellucidomyia, Hebetula. H. most Sphaeromiini s. lat., most Palpomyiini, Paryphoconus; some Bezzia are nearly completed fused (as in Fig. 79I), with only the dorsal apotome separate, along with an ecdysial split along the base of the antenna and the dorsal thoracic split. I. Jenkinshelea, Macropeza, Mallochohelea, Probezzia, Neobezzia." figureDoi="http://doi.org/10.5281/zenodo.4949336" httpUri="https://zenodo.org/record/4949336/files/figure.png" pageId="58" pageNumber="59">79D</figureCitation>
); along prothoracic extension. 
<emphasis id="B8A8EACDBD14304CFB641FD049ACE7DA" bold="true" box="[1180,1244,1477,1502]" pageId="58" pageNumber="59">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD14304CFF671FFC4C7DE406" box="[159,269,1513,1538]" captionStart="FIGURE 19" captionStartId="264.[151,250,1782,1804]" captionTargetBox="[215,1399,189,1744]" captionTargetId="figure-155@264.[176,1410,183,1764]" captionTargetPageId="264" captionText="FIGURE 19. Dorsal apotomes, anterior view. A. Culicoides circumscriptus (from Kettle &amp; Lawson 1952). B. Culicoides cubitalis (from Kettle &amp; Lawson 1952). C. Culicoides truncorum (from Kettle &amp; Lawson 1952). D. Culicoides chiopterus (from Jamnback 1965). E. Paradasyhelea minuta (from Kettle &amp; Elson 1975a). F. Paradasyhelea albipunctata (from Elson- Harris &amp; Kettle 1985a). G. Paradasyhelea reyei (from Elson-Harris &amp; Kettle 1985a). H. Ceratopogon boomerangus (from Borkent &amp; Grogan 1995). I. Ceratopogon abstrusus (from Borkent &amp; Grogan 1995). J. Ceratopogon arcanus (from Borkent &amp; Grogan 1995). K. Brachypogon artemis (from Debenham 1991). L. Alluaudomyia appendiculata (from Elson-Harris &amp; Kettle 1985b). M. Alluaudomyia fumosipennis (from Elson-Harris &amp; Kettle 1985b). N. Baeodasymyia michaeli (from Borkent &amp; Craig 1999). O. Austrohelea campbellensis. P. Stilobezzia rabelloi (from Borkent &amp; Craig 2001)." figureDoi="http://doi.org/10.5281/zenodo.4949130" httpUri="https://zenodo.org/record/4949130/files/figure.png" pageId="58" pageNumber="59">Fig. 19N</figureCitation>
), without ventral line of weakness, with dorsomedial tubercle, without central dome; dorsolateral cephalic sclerite (as in 
<figureCitation id="12E72A5ABD14304CFE591C184F76E422" box="[417,518,1549,1574]" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="58" pageNumber="59">Fig. 13B</figureCitation>
) separated from scutum by thin cuticle, separate from scutum upon emergence, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD14304CFBD91C2449F6E44E" box="[1057,1158,1585,1610]" captionStart="FIGURE 24" captionStartId="269.[151,250,1949,1971]" captionTargetBox="[195,1401,210,1925]" captionTargetId="figure-63@269.[186,1405,208,1929]" captionTargetPageId="269" captionText="FIGURE 24. Posterior portion of heads, in ventral view. A. Culicoides guttipennis. B. Paradasyhelea albipunctata. C. Ceratopogon arcanus. D. Brachypogon sp. (Flag Glade, Pocahontas Co., West Virginia, USA). E. Alluaudomyia needhami. F. Baeodasymyia michaeli. G. Austrohelea campbellensis." figureDoi="http://doi.org/10.5281/zenodo.4949152" httpUri="https://zenodo.org/record/4949152/files/figure.png" pageId="58" pageNumber="59">Fig. 24F</figureCitation>
) with mandible, lacinia absent; palpus extending posterior to posterolateral margin of labium; labium entire (not divided medially); apex of antenna (
<figureCitation id="12E72A5ABD14304CFEF81C6C4C15E496" box="[256,357,1657,1682]" captionStart="FIGURE 35" captionStartId="280.[151,250,1920,1942]" captionTargetBox="[207,1370,219,1888]" captionTargetId="figure-83@280.[197,1391,206,1892]" captionTargetPageId="280" captionText="FIGURE 35. Legs, wings and apices of the antennae, in ventral view. A. Paradasyhelea sp. (from Borlase Stream, Lake Rotoiti, New Zealand). B. Ceratopogon sp. nr. abstrusus. C. Brachypogon sp. (from Spanish Lake, 6 km E. Falkland, BC, Canada). D. Alluaudomyia needhami. E. Baeodasymyia michaeli. F. Austrohelea campbellensis (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949188" httpUri="https://zenodo.org/record/4949188/files/figure.png" pageId="58" pageNumber="59">Fig. 35E</figureCitation>
) anterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum); sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD14304CFE2A1C884F4BE4B2" box="[466,571,1693,1718]" captionStart="FIGURE 19" captionStartId="264.[151,250,1782,1804]" captionTargetBox="[215,1399,189,1744]" captionTargetId="figure-155@264.[176,1410,183,1764]" captionTargetPageId="264" captionText="FIGURE 19. Dorsal apotomes, anterior view. A. Culicoides circumscriptus (from Kettle &amp; Lawson 1952). B. Culicoides cubitalis (from Kettle &amp; Lawson 1952). C. Culicoides truncorum (from Kettle &amp; Lawson 1952). D. Culicoides chiopterus (from Jamnback 1965). E. Paradasyhelea minuta (from Kettle &amp; Elson 1975a). F. Paradasyhelea albipunctata (from Elson- Harris &amp; Kettle 1985a). G. Paradasyhelea reyei (from Elson-Harris &amp; Kettle 1985a). H. Ceratopogon boomerangus (from Borkent &amp; Grogan 1995). I. Ceratopogon abstrusus (from Borkent &amp; Grogan 1995). J. Ceratopogon arcanus (from Borkent &amp; Grogan 1995). K. Brachypogon artemis (from Debenham 1991). L. Alluaudomyia appendiculata (from Elson-Harris &amp; Kettle 1985b). M. Alluaudomyia fumosipennis (from Elson-Harris &amp; Kettle 1985b). N. Baeodasymyia michaeli (from Borkent &amp; Craig 1999). O. Austrohelea campbellensis. P. Stilobezzia rabelloi (from Borkent &amp; Craig 2001)." figureDoi="http://doi.org/10.5281/zenodo.4949130" httpUri="https://zenodo.org/record/4949130/files/figure.png" pageId="58" pageNumber="59">Fig. 19N</figureCitation>
)—1 elongate seta, 1 campaniform sensillum; dorsolateral cephalic sclerite sensilla—2 setae, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD14304CFCC51CD44EEDE4DE" box="[829,925,1729,1754]" captionStart="FIGURE 24" captionStartId="269.[151,250,1949,1971]" captionTargetBox="[195,1401,210,1925]" captionTargetId="figure-63@269.[186,1405,208,1929]" captionTargetPageId="269" captionText="FIGURE 24. Posterior portion of heads, in ventral view. A. Culicoides guttipennis. B. Paradasyhelea albipunctata. C. Ceratopogon arcanus. D. Brachypogon sp. (Flag Glade, Pocahontas Co., West Virginia, USA). E. Alluaudomyia needhami. F. Baeodasymyia michaeli. G. Austrohelea campbellensis." figureDoi="http://doi.org/10.5281/zenodo.4949152" httpUri="https://zenodo.org/record/4949152/files/figure.png" pageId="58" pageNumber="59">Fig. 24F</figureCitation>
)—not visible; oculars (
<figureCitation id="12E72A5ABD14304CFB5C1CD44874E4DE" box="[1188,1284,1729,1754]" captionStart="FIGURE 24" captionStartId="269.[151,250,1949,1971]" captionTargetBox="[195,1401,210,1925]" captionTargetId="figure-63@269.[186,1405,208,1929]" captionTargetPageId="269" captionText="FIGURE 24. Posterior portion of heads, in ventral view. A. Culicoides guttipennis. B. Paradasyhelea albipunctata. C. Ceratopogon arcanus. D. Brachypogon sp. (Flag Glade, Pocahontas Co., West Virginia, USA). E. Alluaudomyia needhami. F. Baeodasymyia michaeli. G. Austrohelea campbellensis." figureDoi="http://doi.org/10.5281/zenodo.4949152" httpUri="https://zenodo.org/record/4949152/files/figure.png" pageId="58" pageNumber="59">Fig. 24F</figureCitation>
)—2 elongate setae. 
<emphasis id="B8A8EACDBD14304CFF1D1CF04C4EE4FA" bold="true" box="[229,318,1765,1790]" pageId="58" pageNumber="59">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD14304CFD911CF04FA3E4FA" box="[617,723,1765,1790]" captionStart="FIGURE 24" captionStartId="269.[151,250,1949,1971]" captionTargetBox="[195,1401,210,1925]" captionTargetId="figure-63@269.[186,1405,208,1929]" captionTargetPageId="269" captionText="FIGURE 24. Posterior portion of heads, in ventral view. A. Culicoides guttipennis. B. Paradasyhelea albipunctata. C. Ceratopogon arcanus. D. Brachypogon sp. (Flag Glade, Pocahontas Co., West Virginia, USA). E. Alluaudomyia needhami. F. Baeodasymyia michaeli. G. Austrohelea campbellensis." figureDoi="http://doi.org/10.5281/zenodo.4949152" httpUri="https://zenodo.org/record/4949152/files/figure.png" pageId="58" pageNumber="59">Fig. 24F</figureCitation>
) wide, well-developed, extending from palpus to antenna; mesonotum with short tubercles, extending posteromedially, completely dividing metathorax medially (
<figureCitation id="12E72A5ABD14304CFAD21D1C48FFE526" box="[1322,1423,1801,1826]" captionStart="FIGURE 48" captionStartId="293.[151,250,1946,1968]" captionTargetBox="[193,1408,188,1923]" captionTargetId="figure-61@293.[189,1420,188,1927]" captionTargetPageId="293" captionText="FIGURE 48. Metathoraces and tergites 1, in dorsal view. A. Culicoides haematopotus. B. Paradasyhelea minuta. C. Ceratopogon sp. nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia needhami. F. Baeodasymyia michaeli." figureDoi="http://doi.org/10.5281/zenodo.4949222" httpUri="https://zenodo.org/record/4949222/files/figure.png" pageId="58" pageNumber="59">Fig. 48F</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD14304CFE971D384CA8E542" box="[367,472,1837,1862]" captionStart="FIGURE 44" captionStartId="289.[151,250,1730,1752]" captionTargetBox="[238,1376,198,1685]" captionTargetId="figure-184@289.[205,1387,192,1716]" captionTargetPageId="289" captionText="FIGURE 44. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Alluaudomyia latipennis (from Debenham 1971). B. Alluaudomyia footei (from Wirth &amp; Grogan 1981). C. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). D. Austrohelea campbellensis. E. Stilobezzia antennalis (from Wirth &amp; Grogan 1981, modified). F. Stilobezzia glauca (from Wirth &amp; Grogan 1981). G. Stilobezzia ochracea (from Kettle &amp; Lawson 1952). H. Stilobezzia diversa (from Wirth &amp; Grogan 1981). I. Stilobezzia rabelloi (from Borkent &amp; Craig 2001). J. Stilobezzia navaiae (from Wirth &amp; Grogan 1981). K. Stilobezzia lutea (from Wirth &amp; Grogan 1981). L. Schizonyxhelea bulla (from Wirth &amp; Grogan 1981, as a Stilobezzia). M. Schizonyxhelea forattinii (from Borkent 2000a). N. Serromyia femorata (from Kettle &amp; Lawson 1952). O. Allohelea japonica. P.Monohelea hirsuta (from Wirth &amp; Grogan 1981, modified). Q. Atyphohelea macroneura (from Borkent 1998). R. Parabezzia alexanderi (from Grogan &amp; Wirth 1977a). S. Parabezzia huberti (from Grogan &amp; Wirth 1977a). T. Echinohelea lanei (from Wirth 1994b)." figureDoi="http://doi.org/10.5281/zenodo.4949208" httpUri="https://zenodo.org/record/4949208/files/figure.png" pageId="58" pageNumber="59">Fig. 44C</figureCitation>
) length/width = 3.75–6.67, elongate, with three distinct sections: cylindrical basal portion smooth, cylindrical middle portion with annulations for most of distal length, compressed apical portion rounded or concave apically, apical portion somewhat flattened, with pores closely abutting or slightly separated at apex of respiratory organ, arranged in single row, with one additional, more basal pore, outer surface with annulations, without other surface modifications, with moderately elongate pedicel, base without posteromedial apodeme, membranous base of respiratory organ short, tracheal tube straight to slightly curved along length, with spirals restricted to base; wing (
<figureCitation id="12E72A5ABD15304DFE051AA94F2FE2D1" box="[509,607,188,213]" captionStart="FIGURE 35" captionStartId="280.[151,250,1920,1942]" captionTargetBox="[207,1370,219,1888]" captionTargetId="figure-83@280.[197,1391,206,1892]" captionTargetPageId="280" captionText="FIGURE 35. Legs, wings and apices of the antennae, in ventral view. A. Paradasyhelea sp. (from Borlase Stream, Lake Rotoiti, New Zealand). B. Ceratopogon sp. nr. abstrusus. C. Brachypogon sp. (from Spanish Lake, 6 km E. Falkland, BC, Canada). D. Alluaudomyia needhami. E. Baeodasymyia michaeli. F. Austrohelea campbellensis (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949188" httpUri="https://zenodo.org/record/4949188/files/figure.png" pageId="59" pageNumber="60">Fig. 35E</figureCitation>
) with slight angle lateral to apex of hind leg, separated medially by fore-, midlegs; halter apex and hind leg (
<figureCitation id="12E72A5ABD15304DFDDF1ACA4FFEE2FC" box="[551,654,223,248]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="59" pageNumber="60">Fig. 32K</figureCitation>
) broadly abutting; halter apex extending posteriorly to 1/3 length of tergite 2; legs (
<figureCitation id="12E72A5ABD15304DFEBA1B114CD6E319" box="[322,422,260,285]" captionStart="FIGURE 35" captionStartId="280.[151,250,1920,1942]" captionTargetBox="[207,1370,219,1888]" captionTargetId="figure-83@280.[197,1391,206,1892]" captionTargetPageId="280" captionText="FIGURE 35. Legs, wings and apices of the antennae, in ventral view. A. Paradasyhelea sp. (from Borlase Stream, Lake Rotoiti, New Zealand). B. Ceratopogon sp. nr. abstrusus. C. Brachypogon sp. (from Spanish Lake, 6 km E. Falkland, BC, Canada). D. Alluaudomyia needhami. E. Baeodasymyia michaeli. F. Austrohelea campbellensis (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949188" httpUri="https://zenodo.org/record/4949188/files/figure.png" pageId="59" pageNumber="60">Fig. 35E</figureCitation>
) with lateral margin of foreleg near midlength of wing evenly curved; hind leg visible at lateral margin of wing (
<figureCitation id="12E72A5ABD15304DFE5F1B324F61E344" box="[423,529,295,320]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="59" pageNumber="60">Fig. 32K</figureCitation>
); with apex of foreleg far anterior to apex of midleg; apex of hind leg abutting apex of midleg laterally; sensilla: anteromedials—1 seta; anterolaterals—2 setae; dorsal setae (
<figureCitation id="12E72A5ABD15304DFB3F1B59485EE361" box="[1223,1326,332,357]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="59" pageNumber="60">Fig. 29Q</figureCitation>
)—D-1-T, D-2-T, D-4-T setae, D-3-T campaniform sensillum, D-5-T absent; D-1-T, D-2-T on single tubercle, D-3-T posteromedial to D-4-T; supraalar 2—campaniform sensillum; metathoracics (
<figureCitation id="12E72A5ABD15304DFBED1B81490AE3A9" box="[1045,1146,404,429]" captionStart="FIGURE 48" captionStartId="293.[151,250,1946,1968]" captionTargetBox="[193,1408,188,1923]" captionTargetId="figure-61@293.[189,1420,188,1927]" captionTargetPageId="293" captionText="FIGURE 48. Metathoraces and tergites 1, in dorsal view. A. Culicoides haematopotus. B. Paradasyhelea minuta. C. Ceratopogon sp. nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia needhami. F. Baeodasymyia michaeli." figureDoi="http://doi.org/10.5281/zenodo.4949222" httpUri="https://zenodo.org/record/4949222/files/figure.png" pageId="59" pageNumber="60">Fig. 48F</figureCitation>
)—1 seta, 2 campaniform sensilla; M-3-T near anterior margin of metathorax. 
<emphasis id="B8A8EACDBD15304DFD251BA24E21E3D4" bold="true" box="[733,849,439,464]" pageId="59" pageNumber="60">Abdomen</emphasis>
: without pigmentation pattern, segment 2 as wide or slightly wider than segment 3, segments with undivided, thin to thick setae, with slightly rounded to pointed, or serrate short tubercles, tergites or sternites entire, each without membranous disc; segment 9 (
<figureCitation id="12E72A5ABD15304DFB001BEA4814E01C" box="[1272,1380,511,536]" captionStart="FIGURE 74" captionStartId="319.[151,250,1737,1759]" captionTargetBox="[224,1348,183,1717]" captionTargetId="figure-195@319.[222,1356,183,1717]" captionTargetPageId="319" captionText="FIGURE 74. Abdominal segments 9. A. Baeodasymyia michaeli, female, in dorsal view (modified from Borkent &amp; Craig 1999). B. Austrohelea campbellensis, female, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). C. Stilobezzia lutea, male, in dorsal and ventral view (from Wirth &amp; Grogan 1981). D. Stilobezzia lutea, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). E. Stilobezzia rabelloi, male, in ventral view (from Borkent &amp; Craig 2001). F. Schizonyxhelea bulla, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). G. Schizonyxhelea forattinii, female, in ventral view (from Borkent 2000a). H. Serromyia sp., female (in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). I. Allohelea japonica, female, in dorsal and ventral view (shagreen not shown). J. Monohelea obscura, male, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). K. Atyphohelea macroneura, female, in dorsal view (from Borkent 1998). L. Parabezzia huberti, male, in ventral view (from Grogan &amp; Wirth 1977a)." figureDoi="http://doi.org/10.5281/zenodo.4949316" httpUri="https://zenodo.org/record/4949316/files/figure.png" pageId="59" pageNumber="60">Fig. 74A</figureCitation>
) not strongly modified but with dorsolateral row of spines at near midlength, bordering shallow posteromedial concavity, terminal processes closely approximated basally, each projecting posterodorsolaterally, tapering to pointed apex; sensilla: tergite 1 (
<figureCitation id="12E72A5ABD15304DFDFC18794F15E081" box="[516,613,620,645]" captionStart="FIGURE 48" captionStartId="293.[151,250,1946,1968]" captionTargetBox="[193,1408,188,1923]" captionTargetId="figure-61@293.[189,1420,188,1927]" captionTargetPageId="293" captionText="FIGURE 48. Metathoraces and tergites 1, in dorsal view. A. Culicoides haematopotus. B. Paradasyhelea minuta. C. Ceratopogon sp. nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia needhami. F. Baeodasymyia michaeli." figureDoi="http://doi.org/10.5281/zenodo.4949222" httpUri="https://zenodo.org/record/4949222/files/figure.png" pageId="59" pageNumber="60">Fig. 48F</figureCitation>
) with 6 setae, 1 campaniform sensillum, including 3 lateral sensilla, D-2- I, D-3-I closely approximated, D-7-I situated posteriorly near D-8-I; segment 4 (
<figureCitation id="12E72A5ABD15304DFBDA189A49F8E0AC" box="[1058,1160,655,680]" captionStart="FIGURE 59" captionStartId="304.[151,250,1975,1997]" captionTargetBox="[203,1416,215,1938]" captionTargetId="figure-57@304.[194,1426,209,1948]" captionTargetPageId="304" captionText="FIGURE 59. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). B. Austrohelea campbellensis. C. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949258" httpUri="https://zenodo.org/record/4949258/files/figure.png" pageId="59" pageNumber="60">Fig. 59A</figureCitation>
)—D-2-IV, D-3-IV short setae on serrate tubercles; D-5-IV barely visible short seta, D-8-IV short seta, D-9-IV absent; D-5-IV, D-4-IV, D-7- IV, D-8-IV on short, serrate tubercles, posterior dorsal sensilla in transverse row, arranged medially to laterally: D- 5-IV, D-4-IV, D-7-IV, D-8-IV; L-1-IV absent, L-2-IV, L-3-IV short setae on pointed tubercles, L-4-IV thick seta on serrate tubercle, V-5-IV, V-6-IV, V-7-IV short setae on serrate tubercles; segment 8 without D-3-VIII, without L-1- VIII; segment 9 (
<figureCitation id="12E72A5ABD15304DFEAF19514CCCE159" box="[343,444,836,861]" captionStart="FIGURE 74" captionStartId="319.[151,250,1737,1759]" captionTargetBox="[224,1348,183,1717]" captionTargetId="figure-195@319.[222,1356,183,1717]" captionTargetPageId="319" captionText="FIGURE 74. Abdominal segments 9. A. Baeodasymyia michaeli, female, in dorsal view (modified from Borkent &amp; Craig 1999). B. Austrohelea campbellensis, female, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). C. Stilobezzia lutea, male, in dorsal and ventral view (from Wirth &amp; Grogan 1981). D. Stilobezzia lutea, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). E. Stilobezzia rabelloi, male, in ventral view (from Borkent &amp; Craig 2001). F. Schizonyxhelea bulla, female, in dorsal and ventral view (from Wirth &amp; Grogan 1981). G. Schizonyxhelea forattinii, female, in ventral view (from Borkent 2000a). H. Serromyia sp., female (in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). I. Allohelea japonica, female, in dorsal and ventral view (shagreen not shown). J. Monohelea obscura, male, in dorsal and ventral view (reconstructed from distorted specimen, shagreen not shown). K. Atyphohelea macroneura, female, in dorsal view (from Borkent 1998). L. Parabezzia huberti, male, in ventral view (from Grogan &amp; Wirth 1977a)." figureDoi="http://doi.org/10.5281/zenodo.4949316" httpUri="https://zenodo.org/record/4949316/files/figure.png" pageId="59" pageNumber="60">Fig. 74A</figureCitation>
)—with D-5-IX, D-6-IX campaniform sensilla.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD15304DFF3F19724C32E1CC" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8A6336DFBD15304DFF3F19724C32E1CC" blockId="59.[151,1436,151,1256]" pageId="59" pageNumber="60">
<emphasis id="B8A8EACDBD15304DFF3F19724F2DE184" bold="true" box="[199,605,871,896]" pageId="59" pageNumber="60">DISTRIBUTION AND HABITAT</emphasis>
: The genus 
<taxonomicName id="4DDC4D5CBD15304DFD1319724EFFE184" box="[747,911,871,896]" class="Insecta" family="Ceratopogonidae" genus="Baeodasymyia" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD15304DFD1319724EFFE184" box="[747,911,871,896]" italics="true" pageId="59" pageNumber="60">Baeodasymyia</emphasis>
</taxonomicName>
is known from five species in the Neotropical Region (
<taxonomicName id="4DDC4D5CBD15304DFF0E19994CE6E1A1" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[246,406,908,933]" genus="Borkent" pageId="59" pageNumber="92" rank="genus" status="gen. nov.">
<bibRefCitation id="EE4D4B2EBD15304DFF0E19994CE6E1A1" author="Borkent" box="[246,406,908,933]" firstAuthor="Borkent" pageId="59" pageNumber="60" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
</taxonomicName>
). The immatures of two species have been reared from small springs in 
<collectingCountry id="F2CB764FBD15304DFB4019994842E1A1" box="[1208,1330,908,933]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
(
<bibRefCitation id="EE4D4B2EBD15304DFAB819994C46E1CC" author="Borkent &amp; Craig" firstAuthor="Borkent" pageId="59" pageNumber="60" pagination="1 - 26" refId="ref177390" refString="Borkent, A. &amp; Craig, D. A. (1999) A revision of the Neotropical genus Baeodasymyia Clastrier &amp; Raccurt (Diptera: Ceratopogonidae) with a discussion of their phylogenetic relationships. American Museum Novitates, 3274, 1 - 26." type="journal article" year="1999">
<taxonomicName id="4DDC4D5CBD15304DFAB8199948ECE1A1" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[1344,1436,908,933]" genus="Borkent" pageId="59" pageNumber="92" rank="genus" status="gen. nov.">Borkent</taxonomicName>
&amp; Craig 1999
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD15304DFF3F19C14F01E631" pageId="59" pageNumber="60" type="discussion">
<paragraph id="8A6336DFBD15304DFF3F19C14F01E631" blockId="59.[151,1436,151,1256]" pageId="59" pageNumber="60">
<emphasis id="B8A8EACDBD15304DFF3F19C14F5AE1E9" bold="true" box="[199,554,980,1005]" pageId="59" pageNumber="60">TAXONOMIC DISCUSSION</emphasis>
: 
<bibRefCitation id="EE4D4B2EBD15304DFDC119C14E34E1E9" author="Borkent &amp; Craig" box="[569,836,980,1005]" firstAuthor="Borkent" pageId="59" pageNumber="60" pagination="1 - 26" refId="ref177390" refString="Borkent, A. &amp; Craig, D. A. (1999) A revision of the Neotropical genus Baeodasymyia Clastrier &amp; Raccurt (Diptera: Ceratopogonidae) with a discussion of their phylogenetic relationships. American Museum Novitates, 3274, 1 - 26." type="journal article" year="1999">
<taxonomicName id="4DDC4D5CBD15304DFDC119C14FE3E1E9" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[569,659,980,1005]" genus="Borkent" pageId="59" pageNumber="92" rank="genus" status="gen. nov.">Borkent</taxonomicName>
&amp; Craig (1999)
</bibRefCitation>
described the pupae of the only two species known in this stage (
<tableCitation id="C75E0364BD15304DFEE519E24CD3E614" box="[285,419,1015,1040]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" pageId="59" pageNumber="60" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 2–3</tableCitation>
). Pupae of 
<taxonomicName id="4DDC4D5CBD15304DFDCE19E24FAAE614" box="[566,730,1015,1040]" class="Insecta" family="Ceratopogonidae" genus="Baeodasymyia" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD15304DFDCE19E24FAAE614" box="[566,730,1015,1040]" italics="true" pageId="59" pageNumber="60">Baeodasymyia</emphasis>
</taxonomicName>
are lacking thoracic sensillum D-5-T and two sensilla on abdominal segment 4: D-9-IV and L-1-IV.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD15304DFF3F1E2A4C43E6EC" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="8A6336DFBD15304DFF3F1E2A4C43E6EC" blockId="59.[151,1436,151,1256]" pageId="59" pageNumber="60">
<emphasis id="B8A8EACDBD15304DFF3F1E2A4F71E65C" bold="true" box="[199,513,1087,1112]" pageId="59" pageNumber="60">MATERIAL EXAMINED</emphasis>
: 
<taxonomicName id="4DDC4D5CBD15304DFDE91E544FC9E65C" authorityName="Borkent" authorityYear="1999" box="[529,697,1087,1112]" class="Insecta" family="Ceratopogonidae" genus="Baeodasymyia" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="christopheri">
<emphasis id="B8A8EACDBD15304DFDE91E544FC9E65C" box="[529,697,1087,1112]" italics="true" pageId="59" pageNumber="60">B. christopheri</emphasis>
</taxonomicName>
: 1 pupal exuviae (of 
<typeStatus id="5567887DBD15304DFC521E2A497FE65C" box="[938,1039,1087,1112]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
), Atenas, 
<collectingCountry id="F2CB764FBD15304DFB861E2A498BE65C" box="[1150,1275,1087,1112]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD15304DFAFF1E2A4803E65C" box="[1287,1395,1087,1112]" pageId="59" pageNumber="60" value="1993-10-05">5-X-1993</date>
(as larva, reared) (CNCI); 
<specimenCount id="9CDAFD56BD15304DFE6F1E714C93E67F" box="[407,483,1124,1149]" pageId="59" pageNumber="60" type="pupa">1 pupa</specimenCount>
(of 
<typeStatus id="5567887DBD15304DFDE81E714F05E679" box="[528,629,1124,1149]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
), 
<quantity id="4D249B3ABD15304DFD701E714FB1E679" box="[648,705,1124,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="km" value="2.0">2 km</quantity>
NE Tarcoles, 
<collectingCountry id="F2CB764FBD15304DFC981E714EACE679" box="[864,988,1124,1149]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD15304DFC101E714906E679" box="[1000,1142,1124,1149]" pageId="59" pageNumber="60" value="1993-12-17">17-XII-1993</date>
(as larva, reared) (CNCI). 
<taxonomicName id="4DDC4D5CBD15304DFF6F1E9C4C68E6A4" authorityName="Borkent" authorityYear="1999" box="[151,280,1159,1184]" class="Insecta" family="Ceratopogonidae" genus="Baeodasymyia" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B8A8EACDBD15304DFF6F1E9C4C68E6A4" box="[151,280,1159,1184]" italics="true" pageId="59" pageNumber="60">B. michaeli</emphasis>
</taxonomicName>
: 
<specimenCount id="9CDAFD56BD15304DFED11E924CF3E69B" box="[297,387,1159,1184]" pageId="59" pageNumber="60" type="pupa">3 pupae</specimenCount>
(of 
<typeStatus id="5567887DBD15304DFE4E1E924F56E6A4" box="[438,550,1159,1184]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
), 9 pupal exuviae (of 
<typeStatus id="5567887DBD15304DFCDC1E924EE4E6A4" box="[804,916,1159,1184]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
), 1 pupal exuviae, Atenas, 
<collectingCountry id="F2CB764FBD15304DFB341E92483BE6A4" box="[1228,1355,1159,1184]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD15304DFAA11E924DBFE6C1" pageId="59" pageNumber="60" value="1993-10-25">25-X- 1993</date>
(as larvae, reared) (CNCI, 
<collectionCode id="ECCDAE1ABD15304DFDF01EB94F20E6C1" box="[520,592,1196,1221]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="59" pageNumber="60">INBC</collectionCode>
); 1 pupal exuviae (in glycerin), 
<quantity id="4D249B3ABD15304DFC451EB94E88E6C1" box="[957,1016,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="km" value="2.0">2 km</quantity>
NE of Tarcoles, 
<collectingCountry id="F2CB764FBD15304DFB431EB9484AE6C1" box="[1211,1338,1196,1221]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD15304DFABE1EB94DBFE6EC" pageId="59" pageNumber="60" value="1993-12-17">17-XII- 1993</date>
(CNCI).
</paragraph>
</subSubSection>
</treatment>
</document>