treatments-xml/data/02/75/87/027587C9BD04305DFD6E1B7A4E47E583.xml

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<mods:title id="6010E5CD24457606EFA760AFD7C31557">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<treatment id="027587C9BD04305DFD6E1B7A4E47E583" ID-DOI="http://doi.org/10.5281/zenodo.5592972" ID-GBIF-Taxon="189161068" ID-Zenodo-Dep="5592972" LSID="urn:lsid:plazi:treatment:027587C9BD04305DFD6E1B7A4E47E583" httpUri="http://treatment.plazi.org/id/027587C9BD04305DFD6E1B7A4E47E583" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<taxonomicName id="4DDC4D5CBD04305CFD6E1B7A4EECE38E" ID-CoL="8YQBD" authority="Kieffer" authorityName="Kieffer" box="[662,924,367,394]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="B8A8EACDBD04305CFD6E1B7A4E4CE38D" bold="true" box="[662,828,367,393]" italics="true" pageId="42" pageNumber="43">Atrichopogon</emphasis>
Kieffer
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<subSubSection id="C2C66554BD04305CFE6E1B8749EDE3AF" box="[406,1181,402,427]" pageId="42" pageNumber="43" type="description">
<paragraph id="8A6336DFBD04305CFE6E1B8749EDE3AF" blockId="42.[406,1181,367,427]" box="[406,1181,402,427]" pageId="42" pageNumber="43">
(
<figureCitation id="12E72A5ABD04305CFE661B874F5BE3AF" box="[414,555,402,427]" captionStart="FIGURE 11" captionStartId="256.[151,250,1845,1867]" captionTargetBox="[213,1374,203,1806]" captionTargetId="figure-145@256.[191,1386,193,1810]" captionTargetPageId="256" captionText="FIGURE 11. Habitus. A. Forcipomyia (Thyridomyia) rugosa, male, in dorsal view (from Chan &amp; LeRoux 1970). B. Forcipomyia (Pterobosca) crinume, female, in ventral view (from Tokunaga 1932). C. Forcipomyia (Lasiohelea) rugosa, male, in dorsal view (from Saunders 1964). D. Atrichopogon hexastichus, male, in dorsal view (from Nielsen 1951). E. Atrichopogon wirthi, female, in dorsal view (from Chan &amp; Linley 1988). F. Atrichopogon muelleri, sex unknown, in dorsal view, with larval exuviae on posterior of abdomen (from Szadziewski et al. 1997). G. Dasyhelea vericolor, sex unknown, in dorsal view (from Szadziewski et al. 1997). H. Schizonyxhelea bulla, sex unknown, in dorsal view (from Thomsen 1937)." figureDoi="http://doi.org/10.5281/zenodo.4949094" httpUri="https://zenodo.org/record/4949094/files/figure.png" pageId="42" pageNumber="43">Figs. 11D–F</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFDCE1B874F15E3AF" box="[566,613,402,427]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="42" pageNumber="43">14E</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFD891B874FD1E3AF" box="[625,673,402,427]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="42" pageNumber="43">18E</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFD541B874FACE3AF" box="[684,732,402,427]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="42" pageNumber="43">23E</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFD1F1B874E67E3AF" box="[743,791,402,427]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="42" pageNumber="43">29G</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFCDA1B874E25E3AF" box="[802,853,402,427]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="42" pageNumber="43">32D</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFC981B874EE3E3AF" box="[864,915,402,427]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="42" pageNumber="43">34D</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFC671B874E9AE3AF" box="[927,1002,402,427]" captionStart="FIGURE 42" captionStartId="287.[151,250,1674,1696]" captionTargetBox="[241,1365,180,1637]" captionTargetId="figure-223@287.[219,1376,176,1661]" captionTargetPageId="287" captionText="FIGURE 42. Respiratory organs, most in dorsal view, Fig. 42B in lateral view, pattern on tracheal tubes shown only in some. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Austroconops mcmillani (from Borkent &amp; Craig 2004). C. Leptoconops americanus (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar (from Clastrier &amp; Wirth 1978). E. Forcipomyia (Euprojoannisia) wirthi (from Saunders 1957). F. Forcipomyia (Euprojoannisia) longispina (from Saunders 1957). G. Forcipomyia (Warmkea) malayae (from Saunders 1957). H. Forcipomyia (Caloforcipomyia) caerulea (from Saunders 1957). I. Forcipomyia (Caloforcipomyia) varicolor (from Saunders 1957). J. Forcipomyia (Metaforcipomyia) cerifera (from Saunders 1957). K. Forcipomyia (Forcipomyia) genualis (from Saunders 1957, as raleighi). L. Forcipomyia (Forcipomyia) nigra (from Saunders 1924). M. Forcipomyia (Forcipomyia) ciliata (from Saunders 1924). N. Forcipomyia (Phytohelea) bromelicola (from Saunders 1925, (as an Apelma)). O. Forcipomyia (Synthyridomyia) acidicola (Synthyridomyia) (from Saunders 1924). P. Forcipomyia (Trichohelea) crinita (from Saunders 1964). Q. Forcipomyia (Lasiohelea) quasicornuta (from Saunders 1964). R. Forcipomyia (Lasiohelea) cornuta (from Saunders 1964). S. Atrichopogon wirthi (from Chan &amp; Linley 1988). T. Atrichopogon thienemanni (from Lenz 1934)." figureDoi="http://doi.org/10.5281/zenodo.4949204" httpUri="https://zenodo.org/record/4949204/files/figure.png" pageId="42" pageNumber="43">42S–T</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFC0D1B874954E3AF" box="[1013,1060,402,427]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="42" pageNumber="43">47E</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFBC81B874912E3AF" box="[1072,1122,402,427]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="42" pageNumber="43">56B</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFB951B8749E5E3AF" box="[1133,1173,402,427]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="42" pageNumber="43">72J</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD04305CFF6F1BCF48EDE3F7" box="[151,1437,474,499]" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8A6336DFBD04305CFF6F1BCF48EDE3F7" blockId="42.[151,1437,474,2010]" box="[151,1437,474,499]" pageId="42" pageNumber="43">
<emphasis id="B8A8EACDBD04305CFF6F1BCF4C44E3F7" bold="true" box="[151,308,474,499]" pageId="42" pageNumber="43">DIAGNOSIS</emphasis>
: Only pupa of 
<taxonomicName id="4DDC4D5CBD04305CFE241BCF4FEBE3F7" authorityName="Borkent" authorityYear="2014" box="[476,667,474,499]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with terminal process with at least some spicules directed anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD04305CFF3F1BEB4CF7E507" pageId="42" pageNumber="43" type="description">
<paragraph id="8A6336DFBD04305CFF3F1BEB4CF7E507" blockId="42.[151,1437,474,2010]" pageId="42" pageNumber="43">
<emphasis id="B8A8EACDBD04305CFF3F1BEB4CFAE013" bold="true" box="[199,394,510,535]" pageId="42" pageNumber="43">DESCRIPTION</emphasis>
: Habitus as in 
<figureCitation id="12E72A5ABD04305CFDC51BEB4FBDE013" box="[573,717,510,535]" captionStart="FIGURE 11" captionStartId="256.[151,250,1845,1867]" captionTargetBox="[213,1374,203,1806]" captionTargetId="figure-145@256.[191,1386,193,1810]" captionTargetPageId="256" captionText="FIGURE 11. Habitus. A. Forcipomyia (Thyridomyia) rugosa, male, in dorsal view (from Chan &amp; LeRoux 1970). B. Forcipomyia (Pterobosca) crinume, female, in ventral view (from Tokunaga 1932). C. Forcipomyia (Lasiohelea) rugosa, male, in dorsal view (from Saunders 1964). D. Atrichopogon hexastichus, male, in dorsal view (from Nielsen 1951). E. Atrichopogon wirthi, female, in dorsal view (from Chan &amp; Linley 1988). F. Atrichopogon muelleri, sex unknown, in dorsal view, with larval exuviae on posterior of abdomen (from Szadziewski et al. 1997). G. Dasyhelea vericolor, sex unknown, in dorsal view (from Szadziewski et al. 1997). H. Schizonyxhelea bulla, sex unknown, in dorsal view (from Thomsen 1937)." figureDoi="http://doi.org/10.5281/zenodo.4949094" httpUri="https://zenodo.org/record/4949094/files/figure.png" pageId="42" pageNumber="43">Figs. 11D–F</figureCitation>
. Total length = 
<quantity id="4D249B3ABD04305CFC7F1BEB495FE012" box="[903,1071,510,535]" metricMagnitude="-3" metricUnit="m" metricValue="2.425" metricValueMax="3.44" metricValueMin="1.41" pageId="42" pageNumber="43" unit="mm" value="2.425" valueMax="3.44" valueMin="1.41">1.41–3.44 mm</quantity>
. With (
<figureCitation id="12E72A5ABD04305CFB7E1BEB4999E013" box="[1158,1257,510,535]" captionStart="FIGURE 11" captionStartId="256.[151,250,1845,1867]" captionTargetBox="[213,1374,203,1806]" captionTargetId="figure-145@256.[191,1386,193,1810]" captionTargetPageId="256" captionText="FIGURE 11. Habitus. A. Forcipomyia (Thyridomyia) rugosa, male, in dorsal view (from Chan &amp; LeRoux 1970). B. Forcipomyia (Pterobosca) crinume, female, in ventral view (from Tokunaga 1932). C. Forcipomyia (Lasiohelea) rugosa, male, in dorsal view (from Saunders 1964). D. Atrichopogon hexastichus, male, in dorsal view (from Nielsen 1951). E. Atrichopogon wirthi, female, in dorsal view (from Chan &amp; Linley 1988). F. Atrichopogon muelleri, sex unknown, in dorsal view, with larval exuviae on posterior of abdomen (from Szadziewski et al. 1997). G. Dasyhelea vericolor, sex unknown, in dorsal view (from Szadziewski et al. 1997). H. Schizonyxhelea bulla, sex unknown, in dorsal view (from Thomsen 1937)." figureDoi="http://doi.org/10.5281/zenodo.4949094" httpUri="https://zenodo.org/record/4949094/files/figure.png" pageId="42" pageNumber="43">Fig. 11F</figureCitation>
) larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of face (
<figureCitation id="12E72A5ABD04305CFB7218374982E03F" box="[1162,1266,546,571]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="42" pageNumber="43">Fig. 14E</figureCitation>
). Ecdysial tear anterior or slightly medial to base of antenna (
<figureCitation id="12E72A5ABD04305CFD5E18534E66E05B" box="[678,790,582,607]" captionStart="FIGURE 14" captionStartId="259.[151,250,1923,1945]" captionTargetBox="[152,1436,177,1909]" captionTargetId="figure-97@259.[151,1436,175,1914]" captionTargetPageId="259" captionText="FIGURE 14. Anterior portion of heads. A. Austroconops mcmillani, in ventral view, dorsal apotome not present. B. Leptoconops bequaerti, in anterior view (distorted slide specimen). C. Forcipomyia (Phytohelea) sp. in anterior view (Florida, USA). D. Forcipomyia (Phytohelea) sp. in anteroventral view (Florida, USA). E. Atrichopogon sp., in anterior view (from 6.5 km NW Enderby, BC, Canada). F. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949108" httpUri="https://zenodo.org/record/4949108/files/figure.png" pageId="42" pageNumber="43">Figs. 14E</figureCitation>
, 
<figureCitation id="12E72A5ABD04305CFCDB18534E26E05B" box="[803,854,582,607]" captionStart="FIGURE 79" captionStartId="324.[151,250,1861,1883]" captionTargetBox="[192,1396,201,1846]" captionTargetId="figure-148@324.[184,1406,195,1847]" captionTargetPageId="324" captionText="FIGURE 79. Evolution of ecdysial sutures and splits in pupal exuviae of Ceratopogonidae, head and anterior portion of thorax in ventral view. Numbers refer to character states of characters 2 and 8 in text. Red indicates ecdysial splits. A. Leptoconopinae. B. Forcipomyiinae. C. Dasyheleinae. D. Most Ceratopogonini. E. Most Ceratopogonini, Clinohelea. F. Allohelea, Atyphohelea, Parabezzia. G. Heteromyia, Pellucidomyia, Hebetula. H. most Sphaeromiini s. lat., most Palpomyiini, Paryphoconus; some Bezzia are nearly completed fused (as in Fig. 79I), with only the dorsal apotome separate, along with an ecdysial split along the base of the antenna and the dorsal thoracic split. I. Jenkinshelea, Macropeza, Mallochohelea, Probezzia, Neobezzia." figureDoi="http://doi.org/10.5281/zenodo.4949336" httpUri="https://zenodo.org/record/4949336/files/figure.png" pageId="42" pageNumber="43">79B</figureCitation>
); along posterolateral portion of eye. 
<emphasis id="B8A8EACDBD04305CFAF918534831E05B" bold="true" box="[1281,1345,582,607]" pageId="42" pageNumber="43">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD04305CFEE8187F4C0EE087" box="[272,382,618,643]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="42" pageNumber="43">Fig. 18E</figureCitation>
), without ventral line of weakness, with dorsomedial tubercle, without central dome; dorsolateral cephalic sclerite (as in 
<figureCitation id="12E72A5ABD04305CFDD6189B4FE2E0A3" box="[558,658,654,679]" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="42" pageNumber="43">Fig. 13B</figureCitation>
) separated from scutum by thin cuticle, separate from scutum upon emergence, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD04305CFB7818A74993E0CF" box="[1152,1251,690,715]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="42" pageNumber="43">Fig. 23E</figureCitation>
) with mandible, lacinia well-developed, not overlapping apically; palpus extending posterior to posterolateral margin of labium; labium separated medially by labrum, in some by hypopharynx; apex of antenna (
<figureCitation id="12E72A5ABD04305CFBB918EF49D9E117" box="[1089,1193,762,787]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="42" pageNumber="43">Fig. 34D</figureCitation>
) anterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum); sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD04305CFAFD190B481AE133" box="[1285,1386,798,823]" captionStart="FIGURE 18" captionStartId="263.[151,250,1850,1872]" captionTargetBox="[164,1422,191,1811]" captionTargetId="figure-119@263.[151,1436,185,1842]" captionTargetPageId="263" captionText="FIGURE 18. Dorsal apotomes, anterior view. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Leptoconops bequaerti. C. Forcipomyia (Phytohelea) n. sp. near F. antiguensis. D. Forcipomyia (Euprojoannisia) sp. (undescribed, from Peru). E. Atrichopogon sp. (undescribed, from Costa Rica). F. Dasyhelea insons (from Mayer 1934c). G. Dasyhelea tersa (from Mayer 1934c). H. Dasyhelea assimilis (from Mayer 1934c). I. Dasyhelea pseudocincta (from Waugh &amp; Wirth 1976, modified). J. Dasyhelea pseudoincisurata (from Waugh &amp; Wirth 1976, modified). K. Dasyhelea messersmithi (from Waugh &amp; Wirth 1976, modified). L. Dasyhelea atlantis (from Waugh &amp; Wirth 1976, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949124" httpUri="https://zenodo.org/record/4949124/files/figure.png" pageId="42" pageNumber="43">Fig. 18E</figureCitation>
)—1 moderate to elongate seta; dorsolateral cephalic sclerite sensilla—1 short seta or 1 very short or elongate seta, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD04305CFD8C19734FAFE17B" box="[628,735,870,895]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="42" pageNumber="43">Fig. 23E</figureCitation>
)—1 seta, 1 campaniform sensillum; oculars (
<figureCitation id="12E72A5ABD04305CFAF91973481BE17B" box="[1281,1387,870,895]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="42" pageNumber="43">Fig. 23E</figureCitation>
)—1 campaniform sensillum. 
<emphasis id="B8A8EACDBD04305CFE40199F4F61E1A7" bold="true" box="[440,529,906,931]" pageId="42" pageNumber="43">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD04305CFCCC199F4EECE1A7" box="[820,924,906,931]" captionStart="FIGURE 23" captionStartId="268.[151,250,1947,1969]" captionTargetBox="[188,1386,225,1936]" captionTargetId="figure-81@268.[180,1399,219,1940]" captionTargetPageId="268" captionText="FIGURE 23. Posterior portion of heads, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Forcipomyia (Phytohelea) n. sp. near antiguensis. E. Atrichopogon sp. (from St. Pierre de Wakefield, Quebec, Canada). F. Dasyhelea paracincta. G. Dasyhelea pseudoincisurata." figureDoi="http://doi.org/10.5281/zenodo.4949144" httpUri="https://zenodo.org/record/4949144/files/figure.png" pageId="42" pageNumber="43">Fig. 23E</figureCitation>
) absent; mesonotum with long, bifurcating tubercles, extending posteromedially, completely dividing metathorax medially (
<figureCitation id="12E72A5ABD04305CFBE419BB49F2E1C3" box="[1052,1154,942,967]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="42" pageNumber="43">Fig. 47E</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD04305CFA9C19BB4DC4E1EF" captionStart="FIGURE 42" captionStartId="287.[151,250,1674,1696]" captionTargetBox="[241,1365,180,1637]" captionTargetId="figure-223@287.[219,1376,176,1661]" captionTargetPageId="287" captionText="FIGURE 42. Respiratory organs, most in dorsal view, Fig. 42B in lateral view, pattern on tracheal tubes shown only in some. A. Austroconops mcmillani (from Borkent &amp; Craig 2004). B. Austroconops mcmillani (from Borkent &amp; Craig 2004). C. Leptoconops americanus (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar (from Clastrier &amp; Wirth 1978). E. Forcipomyia (Euprojoannisia) wirthi (from Saunders 1957). F. Forcipomyia (Euprojoannisia) longispina (from Saunders 1957). G. Forcipomyia (Warmkea) malayae (from Saunders 1957). H. Forcipomyia (Caloforcipomyia) caerulea (from Saunders 1957). I. Forcipomyia (Caloforcipomyia) varicolor (from Saunders 1957). J. Forcipomyia (Metaforcipomyia) cerifera (from Saunders 1957). K. Forcipomyia (Forcipomyia) genualis (from Saunders 1957, as raleighi). L. Forcipomyia (Forcipomyia) nigra (from Saunders 1924). M. Forcipomyia (Forcipomyia) ciliata (from Saunders 1924). N. Forcipomyia (Phytohelea) bromelicola (from Saunders 1925, (as an Apelma)). O. Forcipomyia (Synthyridomyia) acidicola (Synthyridomyia) (from Saunders 1924). P. Forcipomyia (Trichohelea) crinita (from Saunders 1964). Q. Forcipomyia (Lasiohelea) quasicornuta (from Saunders 1964). R. Forcipomyia (Lasiohelea) cornuta (from Saunders 1964). S. Atrichopogon wirthi (from Chan &amp; Linley 1988). T. Atrichopogon thienemanni (from Lenz 1934)." figureDoi="http://doi.org/10.5281/zenodo.4949204" httpUri="https://zenodo.org/record/4949204/files/figure.png" pageId="42" pageNumber="43">Figs. 42</figureCitation>
S-T) length/width = 1.25–3.50, variable, knob-like to elongate, with posterior basal swelling or lobe, somewhat flattened laterally, with pores closely abutting or slightly separated at apex of respiratory organ, arranged in single straight or curved row or two widely spaced rows, outer surface smooth or with a few wrinkles, with welldeveloped, flattened pedicel, base without posteromedial apodeme, membranous base of respiratory organ short, tracheal tube straight to slightly curved along length, surface smooth; wing (
<figureCitation id="12E72A5ABD04305CFBEA1E77490CE67F" box="[1042,1148,1122,1147]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="42" pageNumber="43">Fig. 34D</figureCitation>
) without apical tubercle, separated medially by fore-, midlegs; halter apex and hind leg (
<figureCitation id="12E72A5ABD04305CFC8C1E934EACE69B" box="[884,988,1158,1183]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="42" pageNumber="43">Fig. 32D</figureCitation>
) broadly separate; halter apex abutting anterolateral edge of tergite 2; legs (
<figureCitation id="12E72A5ABD04305CFDBE1EBF4FC0E6C7" box="[582,688,1194,1219]" captionStart="FIGURE 34" captionStartId="279.[151,250,1931,1953]" captionTargetBox="[195,1385,213,1901]" captionTargetId="figure-69@279.[187,1395,201,1905]" captionTargetPageId="279" captionText="FIGURE 34. Legs, wings and apices of the antennae, in ventral view. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Euprojoannisia) sp. (from Peru). D. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). E. Dasyhelea paracincta. F. Culicoides sp. (from 3 km E. Carp, Ontario, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949186" httpUri="https://zenodo.org/record/4949186/files/figure.png" pageId="42" pageNumber="43">Fig. 34D</figureCitation>
) with lateral margin of foreleg near midlength of wing slightly sinuous to evenly curved; hind leg visible at lateral margin of wing (
<figureCitation id="12E72A5ABD04305CFC541EDB4967E6E3" box="[940,1047,1230,1255]" captionStart="FIGURE 32" captionStartId="277.[151,250,1887,1909]" captionTargetBox="[174,1419,183,1869]" captionTargetId="figure-104@277.[173,1420,183,1869]" captionTargetPageId="277" captionText="FIGURE 32. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Austroconops mcmillani. B. Leptoconops linleyi. C. Forcipomyia (Phytohelea) sp. (Florida, USA). D. Atrichopogon sp. (from 6.5 km NW Enderby, BC, Canada). E. Dasyhelea sp., in anteroventral view (Herman Lake, 4 km SE Eagle Bay, BC, Canada). F. Dasyhelea oppresa. G. Culicoides denticulatus. H. Ceratopogon nr. abstrusus. I. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). J. Alluaudomyia bella. K. Baeodasymyia michaeli. L. Stilobezzia antennalis." figureDoi="http://doi.org/10.5281/zenodo.4949182" httpUri="https://zenodo.org/record/4949182/files/figure.png" pageId="42" pageNumber="43">Fig. 32D</figureCitation>
); with apex of foreleg moderately anterior to apex of midleg; apex of hind leg abutting apex of midleg laterally; sensilla: anteromedials—1 campaniform sensillum; anterolaterals—1 short seta, 1 campaniform sensillum; dorsal setae (
<figureCitation id="12E72A5ABD04305CFB3D1F03485EE72B" box="[1221,1326,1302,1327]" captionStart="FIGURE 29" captionStartId="274.[151,250,1822,1844]" captionTargetBox="[165,1429,184,1785]" captionTargetId="figure-146@274.[165,1430,181,1801]" captionTargetPageId="274" captionText="FIGURE 29. Dorsal setae from left side of thoraces. A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Leptoconops linleyi. D. Forcipomyia (Euprojoannisia) sp. (from Peru). E. Forcipomyia (Forcipomyia) bipunctata. F. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. G. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). H. Dasyhelea paracincta. I. Dasyhelea atlantis. J. Dasyhelea pseudoincisurata. K. Culicoides haematopotus. L. Culicoides phlebotomus. M. Paradasyhelea albipunctata. N. Ceratopogon boomerangus. O. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). P. Alluaudomyia needhami. Q. Baeodasymyia christopheri (from Borkent &amp; Craig 1999). R. Austrohelea campbellensis. S. Stilobezzia antennalis. T. Schizonyxhelea forattinii (from Borkent 2000a)." figureDoi="http://doi.org/10.5281/zenodo.4949172" httpUri="https://zenodo.org/record/4949172/files/figure.png" pageId="42" pageNumber="43">Fig. 29G</figureCitation>
)—D-1-T, D-2-T setae, D-3-T campaniform sensillum; D-2-T, D-3-T on single tubercle in some, or closely associated; supraalar 2—campaniform sensillum present or absent; metathoracics (
<figureCitation id="12E72A5ABD04305CFC4E1F4B4967E773" box="[950,1047,1374,1399]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="42" pageNumber="43">Fig. 47E</figureCitation>
)—2 campaniform sensilla; M-3-T near anterior margin of metathorax. 
<emphasis id="B8A8EACDBD04305CFDC71F974FC3E79F" bold="true" box="[575,691,1410,1435]" pageId="42" pageNumber="43">Abdomen</emphasis>
: without pigmentation pattern, segment 2 as wide or somewhat wider than segment 3, segments with undivided, thin to thick setae, without tubercles or segments 1–5 to 1–8 with branched or setaceous elongate tubercles, tubercles rounded to pointed, tergites or sternites entire, each without membranous disc; segment 9 (
<figureCitation id="12E72A5ABD04305CFDFB1FFB4F15E403" box="[515,613,1518,1543]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="42" pageNumber="43">Fig. 72J</figureCitation>
) variable in shape, terminal processes widely separated basally, each projecting posterolaterally, tapering near apex to point, with at least some spicules directed anteriorly; sensilla: tergite 1 (
<figureCitation id="12E72A5ABD04305CFEFD1C234C1AE44B" box="[261,362,1590,1615]" captionStart="FIGURE 47" captionStartId="292.[151,250,1952,1974]" captionTargetBox="[209,1375,191,1853]" captionTargetId="figure-74@292.[201,1383,188,1927]" captionTargetPageId="292" captionText="FIGURE 47. Metathoraces and tergites 1 (in dorsal view). A. Austroconops mcmillani. B. Leptoconops kerteszi. C. Forcipomyia (Forcipomyia) bipunctata. D. Forcipomyia (Phytohelea) n. sp. nr. antiguensis. E. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). F. Dasyhelea pseudoincisuata." figureDoi="http://doi.org/10.5281/zenodo.4949218" httpUri="https://zenodo.org/record/4949218/files/figure.png" pageId="42" pageNumber="43">Fig. 47E</figureCitation>
) with 3 setae, 2 campaniform sensilla, including 2 lateral sensilla, D-3-I absent, D-7-I absent; segment 4 (
<figureCitation id="12E72A5ABD04305CFEEF1C4F4C0AE477" box="[279,378,1626,1651]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="42" pageNumber="43">Fig. 56B</figureCitation>
)—D-2-IV moderately elongate seta, without tubercle, D-3-IV absent; D-4-IV present; D-5-IV, D-7-IV, D-8-IV, D-9-IV absent; D-4-IV without tubercle, L-1-IV slender or stout, short to elongate seta present or absent, V-5-IV, V-6-IV present or absent, V-7-IV small seta, none on tubercles; segment 8 without D-3-VIII, without L-1-VIII; segment 9 (
<figureCitation id="12E72A5ABD04305CFE121CD34F37E4DB" box="[490,583,1734,1759]" captionStart="FIGURE 72" captionStartId="317.[151,250,1765,1787]" captionTargetBox="[228,1373,193,1750]" captionTargetId="figure-192@317.[227,1378,193,1750]" captionTargetPageId="317" captionText="FIGURE 72. Abdominal segments 9. A. Austroconops mcmillani, male, in dorsal and ventral view (from Borkent &amp; Craig 2004). B. Leptoconops whitseli, male, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). C. Leptoconops whitseli, female, in dorsal and ventral view (from Clastrier &amp; Wirth 1978). D. Leptoconops asilomar, male, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). E. Leptoconops asilomar, female, in dorsal and ventral view (modified from Clastrier &amp; Wirth 1978). F. Forcipomyia anabaenae, female, in dorsal view (from Chan &amp; Saunders 1965). G. Forcipomyia eques, male, in ventral view (from Chan &amp; LeRoux 1965). H. Forcipomyia rugosa, female, in dorsal view (from Chan &amp; LeRoux 1970). I. Forcipomyia brevis, female, in dorsal view (from Mayer 1934a). J. Atrichopogon wirthi, male, in dorsal view, arrow indicating anteriorly directed spines (from Chan &amp; Linley 1988). K. Dasyhelea contigua, female, in ventral view (from Mayer 1934c). L. Dasyhelea grata, male, in ventral view (from Mayer 1934c)." figureDoi="http://doi.org/10.5281/zenodo.4949308" httpUri="https://zenodo.org/record/4949308/files/figure.png" pageId="42" pageNumber="43">Fig. 72J</figureCitation>
)—D-5-IX campaniform sensillum, possibly D-6-IX present but inadequate specimens examined.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD04305CFF3F1D1B4E77E5B2" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8A6336DFBD04305CFF3F1D1B4E77E5B2" blockId="42.[151,1437,474,2010]" pageId="42" pageNumber="43">
<emphasis id="B8A8EACDBD04305CFF3F1D1B4F2EE523" bold="true" box="[199,606,1806,1831]" pageId="42" pageNumber="43">DISTRIBUTION AND HABITAT</emphasis>
: The genus 
<taxonomicName id="4DDC4D5CBD04305CFD131D1B4EF5E523" box="[747,901,1806,1831]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD04305CFD131D1B4EF5E523" box="[747,901,1806,1831]" italics="true" pageId="42" pageNumber="43">Atrichopogon</emphasis>
</taxonomicName>
is known from 521 species from every Region worldwide (
<taxonomicName id="4DDC4D5CBD04305CFEE51D274CCDE54F" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[285,445,1842,1867]" genus="Borkent" pageId="42" pageNumber="92" rank="genus" status="gen. nov.">
<bibRefCitation id="EE4D4B2EBD04305CFEE51D274CCDE54F" author="Borkent" box="[285,445,1842,1867]" firstAuthor="Borkent" pageId="42" pageNumber="43" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
</taxonomicName>
). Immatures occur in aquatic to terrestrial habitats, generally in shaded areas on algae or mosses present on or at the edge of ponds, marshes and streams, or over wet soil, rock surfaces and wet wood. Larvae and pupae are also found under bark of logs, on rotting leaves, and other moist microhabitats in forests where fungi and microorganisms are available for food.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD04305DFF3F1DD44973E153" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8A6336DFBD04305DFF3F1DD44EA3E2FC" blockId="42.[151,1437,474,2010]" lastBlockId="43.[151,1437,151,1927]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B8A8EACDBD04305CFF3F1DD44F5DE5DE" bold="true" box="[199,557,1985,2010]" pageId="42" pageNumber="43">TAXONOMIC DISCUSSION</emphasis>
: The pupae of only 39 species of 
<taxonomicName id="4DDC4D5CBD04305CFC4F1DD44921E5DE" box="[951,1105,1985,2010]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD04305CFC4F1DD44921E5DE" box="[951,1105,1985,2010]" italics="true" pageId="42" pageNumber="43">Atrichopogon</emphasis>
</taxonomicName>
have been described (
<tableCitation id="C75E0364BD04305DFAA91DD44DB2E2B4" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 2–3</tableCitation>
) and these different significantly in numbers of details. Indeed, the morphology of both larvae and pupae are so diverse that previous workers have suggested that the immatures would provide a better means of distinguishing the species (
<bibRefCitation id="EE4D4B2EBD05305DFED81ACA4CC8E2FC" author="Nielsen" box="[288,440,223,248]" firstAuthor="Nielsen" pageId="43" pageNumber="44" pagination="1 - 95" refId="ref191812" refString="Nielsen, A. (1951) Contributions to the metamorphosis and biology of the genus Atrichopogon Kieffer (Diptera, Ceratopogonidae) with remarks on the evolution and taxonomy of the genus. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 6 (6), 1 - 95, pls. 1 - 2." type="journal article" year="1951">Nielsen 1951</bibRefCitation>
, 
<bibRefCitation id="EE4D4B2EBD05305DFE3C1AF54FA0E2FC" author="Ewen &amp; Saunders" box="[452,720,223,248]" firstAuthor="Ewen" pageId="43" pageNumber="44" pagination="671 - 724" refId="ref182623" refString="Ewen, A. B. &amp; Saunders, L. G. (1958) Contributions toward a revision of the genus Atrichopogon based on characters of all stages (Diptera, Heleidae). Canadian Journal of Zoology, 36, 671 - 724. http: // dx. doi. org / 10.1139 / z 58 - 061" type="journal article" year="1958">Ewen &amp; Saunders 1958</bibRefCitation>
, 
<bibRefCitation id="EE4D4B2EBD05305DFD231ACA4EB6E2FC" author="Chan &amp; Linley" box="[731,966,223,248]" firstAuthor="Chan" pageId="43" pageNumber="44" pagination="186 - 201" refId="ref179312" refString="Chan, K. L. &amp; Linley, J. R. (1988) Description of Atrichopogon wirthi new species (Diptera: Ceratopogonidae) from leaves of the water lettuce (Pistia stratiotes) in Florida. Florida Entomologist, 71, 186 - 201." type="journal article" year="1988">Chan &amp; Linley 1988</bibRefCitation>
).
</paragraph>
<paragraph id="8A6336DFBD05305DFF3F1B164C6CE3CA" blockId="43.[151,1437,151,1927]" pageId="43" pageNumber="44">
<bibRefCitation id="EE4D4B2EBD05305DFF3F1B164C9FE318" author="Szadziewski" box="[199,495,259,284]" etAl="et al." firstAuthor="Szadziewski" pageId="43" pageNumber="44" pagination="181 - 190" refId="ref197198" refString="Szadziewski, R., Gilka, W. &amp; Anthon, H. (1995) Immature stages of two European species of the subgenus Meloehelea (Diptera: Ceratopogonidae), with keys to the European subgenera of Atrichopogon. Entomologica Scandinavica, 26, 181 - 190. http: // dx. doi. org / 10.1163 / 187631295 X 00189" type="journal article" year="1995">Szadziewski et al. (1995)</bibRefCitation>
provided a key to European subgenera and 
<bibRefCitation id="EE4D4B2EBD05305DFC061B1649FCE318" author="Lenz" box="[1022,1164,259,284]" firstAuthor="Lenz" pageId="43" pageNumber="44" refId="ref189178" refString="Lenz, F. (1934) B. Die Metamorphose der Heleidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. Vol. 3. Stuttgart, pp. 95 - 128." type="book" year="1934">Lenz (1934)</bibRefCitation>
gave a key to the few European species previously described as pupae. 
<bibRefCitation id="EE4D4B2EBD05305DFD2B1B324E89E33B" author="Ewen &amp; Saunders" box="[723,1017,294,319]" firstAuthor="Ewen" pageId="43" pageNumber="44" pagination="671 - 724" refId="ref182623" refString="Ewen, A. B. &amp; Saunders, L. G. (1958) Contributions toward a revision of the genus Atrichopogon based on characters of all stages (Diptera, Heleidae). Canadian Journal of Zoology, 36, 671 - 724. http: // dx. doi. org / 10.1139 / z 58 - 061" type="journal article" year="1958">Ewen &amp; Saunders (1958)</bibRefCitation>
described 19 species from the New World but did not include a key (in spite of obvious differences between taxa). 
<bibRefCitation id="EE4D4B2EBD05305DFBFD1B5F49DCE367" author="Nielsen" box="[1029,1196,330,355]" firstAuthor="Nielsen" pageId="43" pageNumber="44" pagination="1 - 95" refId="ref191812" refString="Nielsen, A. (1951) Contributions to the metamorphosis and biology of the genus Atrichopogon Kieffer (Diptera, Ceratopogonidae) with remarks on the evolution and taxonomy of the genus. Det Kongelige Danske Videnskabernes Selskab, Biologiske Skrifter, 6 (6), 1 - 95, pls. 1 - 2." type="journal article" year="1951">Nielsen (1951)</bibRefCitation>
described six species from 
<collectingCountry id="F2CB764FBD05305DFF211B7B4C31E383" box="[217,321,366,391]" name="Denmark" pageId="43" pageNumber="44">Denmark</collectingCountry>
in well-illustrated and detailed study, including their behaviour and biology. His phylogenetic interpretation of the genus included pupal structures. 
<bibRefCitation id="EE4D4B2EBD05305DFD111B874ED2E3AF" author="Thomsen" box="[745,930,402,427]" firstAuthor="Thomsen" pageId="43" pageNumber="44" pagination="57 - 80" refId="ref197679" refString="Thomsen, L. C. (1937) Aquatic Diptera Part V. Ceratopogonidae. Cornell University Agricultural Experiment Station Memoir, 210, 57 - 80, pls. 10 - 18." type="journal article" year="1937">Thomsen (1937)</bibRefCitation>
gave a brief key to the three Nearctic species she studied.
</paragraph>
<paragraph id="8A6336DFBD05305DFF3F1BCC4EA6E03D" blockId="43.[151,1437,151,1927]" pageId="43" pageNumber="44">
The description of 
<taxonomicName id="4DDC4D5CBD05305DFE581BCF4F50E3F6" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[416,544,473,498]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="websteri">
<emphasis id="B8A8EACDBD05305DFE581BCF4F50E3F6" box="[416,544,473,498]" italics="true" pageId="43" pageNumber="44">A. websteri</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EE4D4B2EBD05305DFDB61BCC4E79E3F6" author="Thomsen" box="[590,777,473,498]" firstAuthor="Thomsen" pageId="43" pageNumber="44" pagination="57 - 80" refId="ref197679" refString="Thomsen, L. C. (1937) Aquatic Diptera Part V. Ceratopogonidae. Cornell University Agricultural Experiment Station Memoir, 210, 57 - 80, pls. 10 - 18." type="journal article" year="1937">Thomsen (1937)</bibRefCitation>
suggests it was misidentified. As illustrated, the elongate respiratory organ, the lack of lateral abdominal tubercles and the terminal processes closely approximated medially and with posteriorly directed spicules suggest it is actually a 
<taxonomicName id="4DDC4D5CBD05305DFCB818374EBFE03D" box="[832,975,546,569]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD05305DFCB818374EBFE03D" box="[832,975,546,569]" italics="true" pageId="43" pageNumber="44">Forcipomyia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8A6336DFBD05305DFF3F18504973E153" blockId="43.[151,1437,151,1927]" pageId="43" pageNumber="44">
Pupae of 
<taxonomicName id="4DDC4D5CBD05305DFECA18514CBCE059" box="[306,460,580,605]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD05305DFECA18514CBCE059" box="[306,460,580,605]" italics="true" pageId="43" pageNumber="44">Atrichopogon</emphasis>
</taxonomicName>
have a reduced number of thoracic sensilla, making the exact naming of these unsure. D-3-T, as a unique campaniform sensillum, is likely homologous to that in other 
<taxonomicName id="4DDC4D5CBD05305DFBDD187D4998E085" authorityName="Borkent" authorityYear="2014" box="[1061,1256,616,641]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
. The abdominal chaetotaxy is also strongly reduced and the remaining sensilla vary in position, making identification of specific sensilla (as in 
<figureCitation id="12E72A5ABD05305DFEC218BA4CEEE0CC" box="[314,414,687,712]" captionStart="FIGURE 56" captionStartId="301.[151,250,1953,1975]" captionTargetBox="[221,1382,193,1855]" captionTargetId="figure-49@301.[213,1395,188,1927]" captionTargetPageId="301" captionText="FIGURE 56. Abdominal segment four, in dorsal and ventral view (shagreen not shown). A. Forcipomyia dominicana. B. Atrichopogon sp. (from Lake Opinicon, Ontario, Canada). C. Dasyhelea paracincta." figureDoi="http://doi.org/10.5281/zenodo.4949250" httpUri="https://zenodo.org/record/4949250/files/figure.png" pageId="43" pageNumber="44">Fig. 56B</figureCitation>
) often uncertain. Some species have one or two more sensilla on abdominal segment four (e.g. 
<taxonomicName id="4DDC4D5CBD05305DFF2818C14C16E0E8" authorityName="Ewen" authorityYear="1958" box="[208,358,723,748]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8A8EACDBD05305DFF2818C14C16E0E8" box="[208,358,723,748]" italics="true" pageId="43" pageNumber="44">A. maculosus</emphasis>
</taxonomicName>
has two lateral setae) and the presence of heavy shagreen in many species makes it particularly difficult to discern sensilla when these are small. In 
<taxonomicName id="4DDC4D5CBD05305DFD1418ED4E09E114" baseAuthorityName="de Meijere" baseAuthorityYear="1907" box="[748,889,759,784]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B8A8EACDBD05305DFD1418ED4E09E114" box="[748,889,759,784]" italics="true" pageId="43" pageNumber="44">A. jacobsoni</emphasis>
</taxonomicName>
, sensillum D-4-IV identified here appears to be homologous to an elongate seta. Furthermore, homologies are further confused by the failure of some descriptions to distinguish tubercles with or without sensilla (e.g. 
<bibRefCitation id="EE4D4B2EBD05305DFD11192A4E87E153" author="Ewen &amp; Saunders" box="[745,1015,830,855]" firstAuthor="Ewen" pageId="43" pageNumber="44" pagination="671 - 724" refId="ref182623" refString="Ewen, A. B. &amp; Saunders, L. G. (1958) Contributions toward a revision of the genus Atrichopogon based on characters of all stages (Diptera, Heleidae). Canadian Journal of Zoology, 36, 671 - 724. http: // dx. doi. org / 10.1139 / z 58 - 061" type="journal article" year="1958">Ewen &amp; Saunders 1958</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD05305DFF3F19774E47E583" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8A6336DFBD05305DFF3F19774E47E583" blockId="43.[151,1437,151,1927]" pageId="43" pageNumber="44">
<emphasis id="B8A8EACDBD05305DFF3F19774F76E17F" bold="true" box="[199,518,866,891]" pageId="43" pageNumber="44">MATERIAL EXAMINED</emphasis>
: 
<taxonomicName id="4DDC4D5CBD05305DFDE319764FFBE17F" authorityName="Ewen &amp; Saunders" authorityYear="1958" box="[539,651,866,891]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="B8A8EACDBD05305DFDE319764FFBE17F" box="[539,651,866,891]" italics="true" pageId="43" pageNumber="44">A. bifidus</emphasis>
</taxonomicName>
: 
<specimenCount id="9CDAFD56BD05305DFD5819774F80E17E" box="[672,752,866,891]" pageId="43" pageNumber="44" type="pupa">1 pupa</specimenCount>
(of 
<typeStatus id="5567887DBD05305DFCD219774EFFE17F" box="[810,911,866,891]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), Nictheroy, 
<collectingCountry id="F2CB764FBD05305DFBD119774900E17F" box="[1065,1136,866,891]" name="Brazil" pageId="43" pageNumber="44">Brazil</collectingCountry>
, 
<date id="FE62101FBD05305DFB791977487FE17F" box="[1153,1295,866,891]" pageId="43" pageNumber="44" value="1923-07-31">31-VII-1923</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFA7C19764C52E19B" authorityName="Ewen &amp; Saunders" authorityYear="1958" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="caribbeanus">
<emphasis id="B8A8EACDBD05305DFA7C19764C52E19B" italics="true" pageId="43" pageNumber="44">A. caribbeanus</emphasis>
</taxonomicName>
: 2 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFDEF19934FF4E19B" box="[535,644,902,927]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
) 
<collectingCountry id="F2CB764FBD05305DFD6C19934F9AE19B" box="[660,746,902,927]" name="Trinidad and Tobago" pageId="43" pageNumber="44">Tobago</collectingCountry>
, 
<collectingCountry id="F2CB764FBD05305DFD0F19934E96E19B" box="[759,998,902,927]" name="Trinidad and Tobago" pageId="43" pageNumber="44">Tobago and Trinidad</collectingCountry>
, 
<date id="FE62101FBD05305DFC0B1993491BE19B" box="[1011,1131,902,927]" pageId="43" pageNumber="44" value="1953-05-31">31-V-1953</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFB211993480DE19B" authorityName="Ewen" authorityYear="1958" box="[1241,1405,902,927]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B8A8EACDBD05305DFB211993480DE19B" box="[1241,1405,902,927]" italics="true" pageId="43" pageNumber="44">A. corpulentus</emphasis>
</taxonomicName>
: 1 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFE8F19BF4CACE1C7" box="[375,476,938,963]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), Nanaimo, BC, 
<collectingCountry id="F2CB764FBD05305DFD5419BF4E73E1C7" box="[684,771,938,963]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFCEE19BF4ED3E1C7" box="[790,931,938,963]" pageId="43" pageNumber="44" value="1926-07-22">22-VII-1926</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFBE419BF49E7E1C6" authorityName="Ewen" authorityYear="1958" box="[1052,1175,938,962]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="crinitus">
<emphasis id="B8A8EACDBD05305DFBE419BF49E7E1C6" box="[1052,1175,938,962]" italics="true" pageId="43" pageNumber="44">A. crinitus</emphasis>
</taxonomicName>
: 2 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFF6F19D84C77E1E2" box="[151,263,973,998]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), Nanaimo, BC, 
<collectingCountry id="F2CB764FBD05305DFE3C19D84F6BE1E2" box="[452,539,973,998]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFDDF19D84FC4E1E2" box="[551,692,973,998]" pageId="43" pageNumber="44" value="1926-07-22">22-VII-1926</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFCD919DB4EF4E1E2" authorityName="Ewen" authorityYear="1958" box="[801,900,973,998]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="flavus">
<emphasis id="B8A8EACDBD05305DFCD919DB4EF4E1E2" box="[801,900,973,998]" italics="true" pageId="43" pageNumber="44">A. flavus</emphasis>
</taxonomicName>
: 2 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFB8F19D84997E1E2" box="[1143,1255,973,998]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), Beaver Creek, Saskatoon, 
<collectingRegion id="4818F83DBD05305DFED819E44CB1E60E" box="[288,449,1009,1034]" country="Canada" name="Saskatchewan" pageId="43" pageNumber="44">Saskatchewan</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFE2C19E44F5BE60E" box="[468,555,1009,1034]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFDC619E44FDBE60E" box="[574,683,1009,1034]" pageId="43" pageNumber="44" value="1955-10-08">8-X-1955</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFCDC19E74ED7E60E" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[804,935,1009,1034]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="fusculus">
<emphasis id="B8A8EACDBD05305DFCDC19E74ED7E60E" box="[804,935,1009,1034]" italics="true" pageId="43" pageNumber="44">A. fusculus</emphasis>
</taxonomicName>
: 2 pupal exuviae, Pike Lake, Saskatoon, 
<collectingRegion id="4818F83DBD05305DFF6F1E004C49E62A" box="[151,313,1045,1070]" country="Canada" name="Saskatchewan" pageId="43" pageNumber="44">Saskatchewan</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFEBD1E004CECE62A" box="[325,412,1045,1070]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFE511E004F78E62A" box="[425,520,1045,1070]" pageId="43" pageNumber="44" value="1956-09">IX-1956</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFD8E1E034FADE62A" baseAuthorityName="Meigen" baseAuthorityYear="1804" box="[630,733,1045,1070]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B8A8EACDBD05305DFD8E1E034FADE62A" box="[630,733,1045,1070]" italics="true" pageId="43" pageNumber="44">A. fuscus</emphasis>
</taxonomicName>
: 5 pupal exuviae, no locality, coll. by Goetghebuer (so likely western Europe) (CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFE421E2C4F47E655" authorityName="Boesel" authorityYear="1973" box="[442,567,1081,1105]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B8A8EACDBD05305DFE421E2C4F47E655" box="[442,567,1081,1105]" italics="true" pageId="43" pageNumber="44">A. geminus</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4DDC4D5CBD05305DFD9E1E2C4FC8E656" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[614,696,1081,1106]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="B8A8EACDBD05305DFD9E1E2C4FC8E656" box="[614,696,1081,1106]" italics="true" pageId="43" pageNumber="44">A. levis</emphasis>
</taxonomicName>
): 3 pupal exuviae, Put-in-Bay, 
<collectingRegion id="4818F83DBD05305DFBE01E2C4923E656" box="[1048,1107,1081,1106]" country="United States of America" name="Ohio" pageId="43" pageNumber="44">Ohio</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFBA71E2C49E7E656" box="[1119,1175,1081,1106]" name="United States of America" pageId="43" pageNumber="44">USA</collectingCountry>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFAFA1E2C48E4E656" authorityName="Ewen" authorityYear="1958" box="[1282,1428,1081,1106]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="humicolus">
<emphasis id="B8A8EACDBD05305DFAFA1E2C48E4E656" box="[1282,1428,1081,1106]" italics="true" pageId="43" pageNumber="44">A. humicolus</emphasis>
</taxonomicName>
: 1 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFE7C1E494C99E671" box="[388,489,1116,1141]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), Saskatoon, 
<collectingRegion id="4818F83DBD05305DFD7B1E494E54E671" box="[643,804,1116,1141]" country="Canada" name="Saskatchewan" pageId="43" pageNumber="44">Saskatchewan</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFCCB1E494EFAE671" box="[819,906,1116,1141]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFC601E49497FE671" box="[920,1039,1116,1141]" pageId="43" pageNumber="44" value="1955-09-09">9-IX-1955</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFB871E484843E670" authorityName="Ewen" authorityYear="1958" box="[1151,1331,1117,1140]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="inconspicuus">
<emphasis id="B8A8EACDBD05305DFB871E484843E670" box="[1151,1331,1117,1140]" italics="true" pageId="43" pageNumber="44">A. inconspicuus</emphasis>
</taxonomicName>
: 4 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFEE41E954CFCE69D" box="[284,396,1152,1177]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), Saskatoon, 
<collectingRegion id="4818F83DBD05305DFDE61E954FB0E69D" box="[542,704,1152,1177]" country="Canada" name="Saskatchewan" pageId="43" pageNumber="44">Saskatchewan</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFD321E954E51E69D" box="[714,801,1152,1177]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFCD41E954EA5E69D" box="[812,981,1152,1177]" pageId="43" pageNumber="44" value="1955-09-17" valueMax="1955-09-24" valueMin="1955-09-17">17-24-IX-1955</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFBC71E9449C6E69D" authorityName="Ewen &amp; Saunders" authorityYear="1958" box="[1087,1206,1152,1177]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="incultus">
<emphasis id="B8A8EACDBD05305DFBC71E9449C6E69D" box="[1087,1206,1152,1177]" italics="true" pageId="43" pageNumber="44">A. incultus</emphasis>
</taxonomicName>
: 1 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFF6F1EB14D8CE6B9" box="[151,252,1188,1213]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), Siquirres, 
<collectingCountry id="F2CB764FBD05305DFE791EB14C8EE6B9" box="[385,510,1188,1213]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD05305DFDF11EB14FFEE6B9" box="[521,654,1188,1213]" pageId="43" pageNumber="44" value="1956-06-18">18-VI-1956</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFD001EB04EF4E6B9" baseAuthorityName="de Meijere" baseAuthorityYear="1907" box="[760,900,1188,1213]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B8A8EACDBD05305DFD001EB04EF4E6B9" box="[760,900,1188,1213]" italics="true" pageId="43" pageNumber="44">A. jacobsoni</emphasis>
</taxonomicName>
: 1 pupal exuviae, Batu Caves, Cavern A, 
<collectingRegion id="4818F83DBD05305DFAA11EB14D83E6DB" country="Malaysia" name="Kuala Lumpur" pageId="43" pageNumber="44">Kuala Lumpur</collectingRegion>
, Selanger, 
<collectingCountry id="F2CB764FBD05305DFE8D1ED24CAEE6E4" box="[373,478,1223,1248]" name="Malaysia" pageId="43" pageNumber="44">Malaysia</collectingCountry>
, 
<date id="FE62101FBD05305DFE151ED24F0AE6E4" box="[493,634,1223,1248]" pageId="43" pageNumber="44" value="1960-12-27">27-XII-1960</date>
(ANIC); 
<specimenCount id="9CDAFD56BD05305DFD081ED24E32E6DB" box="[752,834,1223,1248]" pageId="43" pageNumber="44" type="pupa">1 pupa</specimenCount>
, 1 pupal exuviae, as previous locality, 
<date id="FE62101FBD05305DFAF71ED248EBE6E4" box="[1295,1435,1223,1248]" pageId="43" pageNumber="44" value="1960-12-27">27-XII-1960</date>
(USNM). 
<taxonomicName id="4DDC4D5CBD05305DFEF01EF94CECE700" authorityName="Ewen" authorityYear="1958" box="[264,412,1259,1284]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8A8EACDBD05305DFEF01EF94CECE700" box="[264,412,1259,1284]" italics="true" pageId="43" pageNumber="44">A. maculosus</emphasis>
</taxonomicName>
: 
<specimenCount id="9CDAFD56BD05305DFE531EFE4F74E707" box="[427,516,1259,1284]" pageId="43" pageNumber="44" type="pupa">4 pupae</specimenCount>
, 5 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFD151EFE4E2DE700" box="[749,861,1259,1284]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), Beaver Creek, Saskatoon, 
<collectingRegion id="4818F83DBD05305DFB6D1EFE4847E700" box="[1173,1335,1259,1284]" country="Canada" name="Saskatchewan" pageId="43" pageNumber="44">Saskatchewan</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFAB91EFE48E8E700" box="[1345,1432,1259,1284]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFF6F1F1A4C22E72C" box="[151,338,1295,1320]" pageId="43" pageNumber="44" value="1955-08-25" valueMax="1955-08-28" valueMin="1955-08-25">25-28-VIII-1955</date>
(CNCI); 1 pupal exuviae, Patuxent Wildlife Refuge, Prince George’s County, 
<collectingRegion id="4818F83DBD05305DFB3B1F1A4843E72C" box="[1219,1331,1295,1320]" country="United States of America" name="Maryland" pageId="43" pageNumber="44">Maryland</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFAC61F1A4809E72C" box="[1342,1401,1295,1320]" name="United States of America" pageId="43" pageNumber="44">USA</collectingCountry>
, 
<date id="FE62101FBD05305DFA7D1F1A4C78E748" pageId="43" pageNumber="44" value="1979-08-01">1- VIII-1979</date>
(USNM); 2 pupal exuviae, as previous locality, 
<date id="FE62101FBD05305DFCCF1F264EBDE748" box="[823,973,1331,1356]" pageId="43" pageNumber="44" value="1979-08-10">10-VIII-1979</date>
(USNM); 1 pupal exuviae, 
<quantity id="4D249B3ABD05305DFAE81F264834E748" box="[1296,1348,1331,1356]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="43" pageNumber="44" unit="mi" value="5.0">5 mi</quantity>
NW of Davidsburg, York County, 
<collectingRegion id="4818F83DBD05305DFE3F1F434F2FE76B" box="[455,607,1366,1391]" country="United States of America" name="Pennsylvania" pageId="43" pageNumber="44">Pennsylvania</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFD941F434FD7E76B" box="[620,679,1366,1391]" name="United States of America" pageId="43" pageNumber="44">USA</collectingCountry>
, 
<date id="FE62101FBD05305DFD4C1F434E43E76B" box="[692,819,1366,1391]" pageId="43" pageNumber="44" value="1961-07-04">4-VII-1961</date>
(USNM). 
<taxonomicName id="4DDC4D5CBD05305DFC561F424957E76B" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[942,1063,1367,1391]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B8A8EACDBD05305DFC561F424957E76B" box="[942,1063,1367,1391]" italics="true" pageId="43" pageNumber="44">A. minutus</emphasis>
</taxonomicName>
: 4 pupal exuviae, Lit. Abington, Cambs., 
<collectingCountry id="F2CB764FBD05305DFF031F6F4C2BE797" box="[251,347,1402,1427]" name="United Kingdom" pageId="43" pageNumber="44">England</collectingCountry>
, 
<collectingCountry id="F2CB764FBD05305DFE901F6F4C8FE797" box="[360,511,1402,1427]" name="United Kingdom" pageId="43" pageNumber="44">Great Britain</collectingCountry>
, 
<date id="FE62101FBD05305DFDF51F6F4FD3E797" box="[525,675,1402,1427]" pageId="43" pageNumber="44" value="1924-08-27">27-VIII-1924</date>
(CNCI); 2 pupal exuviae, Truro, 
<collectingCountry id="F2CB764FBD05305DFBDD1F6F490CE797" box="[1061,1148,1402,1427]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFB701F6F486FE797" box="[1160,1311,1402,1427]" pageId="43" pageNumber="44" value="1925-08-10">10-VIII-1925</date>
(CNCI); 8 pupal exuviae, Nanaimo, BC, 
<collectingCountry id="F2CB764FBD05305DFE1F1F8B4F4EE7B3" box="[487,574,1438,1463]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFDB01F8B4FA5E7B3" box="[584,725,1438,1463]" pageId="43" pageNumber="44" value="1926-07-22">22-VII-1926</date>
(CNCI); 11 pupal exuviae, Victoria, BC, 
<collectingCountry id="F2CB764FBD05305DFB5C1F8B498BE7B3" box="[1188,1275,1438,1463]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFAFE1F8B48EBE7B3" box="[1286,1435,1438,1463]" pageId="43" pageNumber="44" value="1947-08-26">26-VIII-1947</date>
(CNCI); 14 pupal exuviae, previous locality, 
<date id="FE62101FBD05305DFD5B1FD74E41E7DF" box="[675,817,1474,1499]" pageId="43" pageNumber="44" value="1948-07-20">20-VII-1948</date>
(CNCI); 11 pupal exuviae, previous locality, 
<date id="FE62101FBD05305DFABF1FD74DBFE7FA" pageId="43" pageNumber="44" value="1948-07-12">12-VII- 1948</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFEC31FF34CB0E7FA" baseAuthorityName="Ewen &amp; Saunders" baseAuthorityYear="1958" box="[315,448,1509,1534]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B8A8EACDBD05305DFEC31FF34CB0E7FA" box="[315,448,1509,1534]" italics="true" pageId="43" pageNumber="44">A. obscurus</emphasis>
</taxonomicName>
: 
<specimenCount id="9CDAFD56BD05305DFE281FF04F6EE7F9" box="[464,542,1509,1534]" pageId="43" pageNumber="44" type="pupa">1 pupa</specimenCount>
, 1 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFCF61FF04E03E7FA" box="[782,883,1509,1534]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), 
<collectingRegion id="4818F83DBD05305DFC7E1FF34E8DE7FA" box="[902,1021,1510,1534]" country="Puerto Rico" name="Mayaguez" pageId="43" pageNumber="44">Mayaguez</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFBF21FF349E3E7FA" box="[1034,1171,1509,1534]" name="Puerto Rico" pageId="43" pageNumber="44">Puerto Rico</collectingCountry>
, 
<date id="FE62101FBD05305DFB671FF04868E7FA" box="[1183,1304,1509,1534]" pageId="43" pageNumber="44" value="1953-02-25">25-II-1953</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFA7C1FF34C76E425" baseAuthorityName="Ewen &amp; Saunders" baseAuthorityYear="1958" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="remigatus">
<emphasis id="B8A8EACDBD05305DFA7C1FF34C76E425" italics="true" pageId="43" pageNumber="44">A. remigatus</emphasis>
</taxonomicName>
: 1 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFE061C1C4F13E426" box="[510,611,1545,1570]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
), Petropolis, 
<collectingCountry id="F2CB764FBD05305DFD011C1C4E30E426" box="[761,832,1545,1570]" name="Brazil" pageId="43" pageNumber="44">Brazil</collectingCountry>
, 
<date id="FE62101FBD05305DFCB51C1C4EABE426" box="[845,987,1545,1570]" pageId="43" pageNumber="44" value="1923-07-27">27-VII-1923</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFBB01C1F49A6E426" baseAuthorityName="Ewen &amp; Saunders" baseAuthorityYear="1958" box="[1096,1238,1545,1570]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="B8A8EACDBD05305DFBB01C1F49A6E426" box="[1096,1238,1545,1570]" italics="true" pageId="43" pageNumber="44">A. saundersi</emphasis>
</taxonomicName>
: 4 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFF3C1C384C44E442" box="[196,308,1581,1606]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), 
<collectingRegion id="4818F83DBD05305DFEB41C3B4CB3E442" box="[332,451,1582,1606]" country="Puerto Rico" name="Mayaguez" pageId="43" pageNumber="44">Mayaguez</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFE2B1C3B4F10E442" box="[467,608,1581,1606]" name="Puerto Rico" pageId="43" pageNumber="44">Puerto Rico</collectingCountry>
, 
<date id="FE62101FBD05305DFD891C384F83E442" box="[625,755,1581,1606]" pageId="43" pageNumber="44" value="1953-03-27">27-III-1953</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFC901C3B4967E442" baseAuthorityName="Ewen &amp; Saunders" baseAuthorityYear="1948" box="[872,1047,1581,1606]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B8A8EACDBD05305DFC901C3B4967E442" box="[872,1047,1581,1606]" italics="true" pageId="43" pageNumber="44">A. tuberculatus</emphasis>
</taxonomicName>
: 2 pupal exuviae (of 
<typeStatus id="5567887DBD05305DFAD81C3848E0E442" box="[1312,1424,1581,1606]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
), Macaras, 
<collectingCountry id="F2CB764FBD05305DFEF11C444C1BE46E" box="[265,363,1617,1642]" name="Trinidad and Tobago" pageId="43" pageNumber="44">Trinidad</collectingCountry>
, 
<date id="FE62101FBD05305DFE841C444F79E46E" box="[380,521,1617,1642]" pageId="43" pageNumber="44" value="1957-07-14">14-VII-1957</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFD871C444E7CE46D" authorityName="Goetghebuer" authorityYear="1922" box="[639,780,1617,1641]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="winnertzi">
<emphasis id="B8A8EACDBD05305DFD871C444E7CE46D" box="[639,780,1617,1641]" italics="true" pageId="43" pageNumber="44">A. winnertzi</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4DDC4D5CBD05305DFCBD1C444E8BE46E" authorityName="Kieffer" authorityYear="1922" box="[837,1019,1617,1642]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="meloesugans">
<emphasis id="B8A8EACDBD05305DFCBD1C444E8BE46E" box="[837,1019,1617,1642]" italics="true" pageId="43" pageNumber="44">A. meloesugans</emphasis>
</taxonomicName>
): 12 pupal exuviae, Strelly, Notts., 
<collectingCountry id="F2CB764FBD05305DFF6F1C614D87E489" box="[151,247,1652,1677]" name="United Kingdom" pageId="43" pageNumber="44">England</collectingCountry>
, 
<collectingCountry id="F2CB764FBD05305DFEFD1C614CEEE489" box="[261,414,1652,1677]" name="United Kingdom" pageId="43" pageNumber="44">Great Britain</collectingCountry>
, 
<date id="FE62101FBD05305DFE531C614F51E489" box="[427,545,1652,1677]" pageId="43" pageNumber="44" value="1922-11-03">3-XI-1922</date>
(CNCI). 
<taxonomicName id="4DDC4D5CBD05305DFD691C604F85E489" box="[657,757,1652,1677]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="B8A8EACDBD05305DFD691C604F85E489" box="[657,757,1652,1677]" italics="true" pageId="43" pageNumber="44">A. wirthi</emphasis>
</taxonomicName>
: 2 pupal exuviae, Chinese Farm, Ft. Pierce, Florida, 
<collectingCountry id="F2CB764FBD05305DFAA51C6148E8E489" box="[1373,1432,1652,1677]" name="United States of America" pageId="43" pageNumber="44">USA</collectingCountry>
, 
<date id="FE62101FBD05305DFF6F1C8D4D8FE4B5" box="[151,255,1688,1713]" pageId="43" pageNumber="44" value="1987-07">VII-1987</date>
(CNCI). 
<emphasis id="B8A8EACDBD05305DFE941C8C4C0DE4B4" box="[364,381,1689,1712]" italics="true" pageId="43" pageNumber="44">A</emphasis>
. nr. 
<taxonomicName id="4DDC4D5CBD05305DFE491C8D4F54E4B5" authorityName="Ewen" authorityYear="1958" box="[433,548,1688,1713]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="humicolus">
<emphasis id="B8A8EACDBD05305DFE491C8D4F54E4B5" box="[433,548,1688,1713]" italics="true" pageId="43" pageNumber="44">humicolus</emphasis>
</taxonomicName>
: no locality given, 
<date id="FE62101FBD05305DFCF91C8D4E24E4B5" box="[769,852,1688,1713]" pageId="43" pageNumber="44" value="1953-05">V-1953</date>
(USNM). 
<emphasis id="B8A8EACDBD05305DFC371C8C4E90E4B4" box="[975,992,1689,1712]" italics="true" pageId="43" pageNumber="44">A</emphasis>
. (
<emphasis id="B8A8EACDBD05305DFC001C8D49D0E4B5" box="[1016,1184,1688,1713]" italics="true" pageId="43" pageNumber="44">Lophomyidium</emphasis>
) sp.: 1 pupal exuviae, orilla Rio Dantes, Parque Nacional Barbilla, 
<collectingRegion id="4818F83DBD05305DFD621CA94F85E4D1" box="[666,757,1724,1749]" country="Costa Rica" name="Cartago" pageId="43" pageNumber="44">Cartago</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFCFA1CA94EF1E4D1" box="[770,897,1724,1749]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD05305DFC771CA94978E4D1" box="[911,1032,1724,1749]" pageId="43" pageNumber="44" value="2006-02-10">10-II-2006</date>
(CNCI). 
<emphasis id="B8A8EACDBD05305DFB8E1CA849F7E4D0" box="[1142,1159,1725,1748]" italics="true" pageId="43" pageNumber="44">A</emphasis>
. sp.: 4 pupal exuviae, 2 pupal exuviae (in glycerin), 
<quantity id="4D249B3ABD05305DFE1D1CCA4F47E4FC" box="[485,567,1759,1784]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="43" pageNumber="44" unit="km" value="6.5">6.5 km</quantity>
NW of Enderby, 
<collectingRegion id="4818F83DBD05305DFCF61CCA4EA6E4FC" box="[782,982,1759,1784]" country="Canada" name="British Columbia" pageId="43" pageNumber="44">British Columbia</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFC1E1CCA494DE4FC" box="[998,1085,1759,1784]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFBB41CCA49A1E4FC" box="[1100,1233,1759,1784]" pageId="43" pageNumber="44" value="1990-06-27">27-VI-1990</date>
(CNCI); 1 pupal exuviae, Lake Opinicon, 
<collectingRegion id="4818F83DBD05305DFE4E1D164F7EE518" box="[438,526,1795,1820]" country="Canada" name="Ontario" pageId="43" pageNumber="44">Ontario</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFDE31D164F01E518" box="[539,625,1795,1820]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, coll. 
<date id="FE62101FBD05305DFD401D164E37E518" box="[696,839,1795,1820]" pageId="43" pageNumber="44" value="1966-07-11">11-VII-1966</date>
, emerged 
<date id="FE62101FBD05305DFC441D164923E518" box="[956,1107,1795,1820]" pageId="43" pageNumber="44" value="1966-08-29">29-VIII-1966</date>
(CNCI); 1 pupal exuviae, as previous locality, 
<date id="FE62101FBD05305DFE921D324C87E544" box="[362,503,1831,1856]" pageId="43" pageNumber="44" value="1966-12-12">12-XII-1966</date>
(CNCI); 1 pupal exuviae, as previous locality, 
<date id="FE62101FBD05305DFBCE1D3249BCE544" box="[1078,1228,1831,1856]" pageId="43" pageNumber="44" value="1966-08-25">25-VIII-1966</date>
(CNCI); 2 pupal exuviae, 
<collectingCountry id="F2CB764FBD05305DFF041D5E4C14E560" box="[252,356,1867,1892]" name="Saint Pierre and Miquelon" pageId="43" pageNumber="44">St. Pierre</collectingCountry>
de Wakefield, 
<collectingRegion id="4818F83DBD05305DFDF51D5E4F15E560" box="[525,613,1867,1892]" country="Canada" name="Quebec" pageId="43" pageNumber="44">Quebec</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFD881D5E4FB7E560" box="[624,711,1867,1892]" name="Canada" pageId="43" pageNumber="44">Canada</collectingCountry>
, 
<date id="FE62101FBD05305DFD2A1D5E4E26E560" box="[722,854,1867,1892]" pageId="43" pageNumber="44" value="1964-06-25">25-VI-1964</date>
(CNCI); 5 pupal exuviae, orilla Rio Dantes, Parque Nacional Barbilla, 
<collectingRegion id="4818F83DBD05305DFE931D7B4CB6E583" box="[363,454,1902,1927]" country="Costa Rica" name="Cartago" pageId="43" pageNumber="44">Cartago</collectingRegion>
, 
<collectingCountry id="F2CB764FBD05305DFE2A1D7B4F3EE583" box="[466,590,1902,1927]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
, 
<date id="FE62101FBD05305DFDA21D7B4FA3E583" box="[602,723,1902,1927]" pageId="43" pageNumber="44" value="2006-02-10">10-II-2006</date>
(CNCI).
</paragraph>
</subSubSection>
</treatment>
</document>