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<document id="060FAB503A63A1AFEF16DAC035ABB8EB" ID-DOI="10.5281/zenodo.3713503" ID-GBIF-Dataset="145c534d-f4eb-4099-9779-02ff529be895" ID-ISSN="1942-1354" ID-Zenodo-Dep="3713503" ID-ZooBank="FD766FC7-F2E5-47D1-96CE-9FED2AF7F483" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584472512751" checkinUser="felipe" docAuthor="Flowers, R. Wills" docDate="2018" docId="026C87B4F33EFFFFFF15FC3EFA39606A" docLanguage="en" docName="InsectaMundi.0672.1-43.pdf.imf" docOrigin="Insecta Mundi 672" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Beltia angustomarginata Flowers 2018, new combination" docType="treatment" docVersion="5" lastPageNumber="7" masterDocId="FE55FFCCF339FFF7FFD5FFB9FF99643F" masterDocTitle="A review of the genus Beltia Jacoby (Chrysomelidae: Eumolpinae: Eumolpini), with descriptions of fourteen new species from Costa Rica, Panama, and northwestern South America" masterLastPageNumber="43" masterPageNumber="1" pageNumber="6" updateTime="1698729393675" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="C7EE24B511E9C77E846F2783963A85BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BE1F0A621DDBF53E1F98BA1E54578F1F">
<mods:title id="D5A4A49DB68D20250064730C4744BCC7">A review of the genus Beltia Jacoby (Chrysomelidae: Eumolpinae: Eumolpini), with descriptions of fourteen new species from Costa Rica, Panama, and northwestern South America</mods:title>
</mods:titleInfo>
<mods:name id="AC1BD3957737145E2F6BBC77C58B2CF9" type="personal">
<mods:role id="0735CDAAC4387C04F646119C34EBABF4">
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<mods:namePart id="45C8D34DF0B1AFD667DD3C85B6E6C8E2">Flowers, R. Wills</mods:namePart>
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<mods:typeOfResource id="903CFBD55137172F6D78EF80769D7D31">text</mods:typeOfResource>
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<mods:title id="67C8C89AF15E86ACBD4701E52E038F85">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="B153560377D740A687262A05A36EDAC0">
<mods:date id="D221E5AF9D7AF26142801F877F9DF8C8">2018</mods:date>
<mods:detail id="CC2BBA8F350C0718A2FC22058499EFD7" type="pubDate">
<mods:number id="80AA8E20455981596E14408BD6362866">2018-11-30</mods:number>
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<mods:detail id="AEBF62E1CCB721403970FAE96D3FFEC0" type="issue">
<mods:number id="5720B9B36C84274B013622D8D7CBA6C4">672</mods:number>
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<mods:classification id="5D7E365430EB13A54308658BC15E3F9C">journal article</mods:classification>
<mods:identifier id="00B5E61EBF94ABEA43B53C949B05EBE4" type="DOI">10.5281/zenodo.3713503</mods:identifier>
<mods:identifier id="93C6E2CD93166934CB93B03A0309F86D" type="GBIF-Dataset">145c534d-f4eb-4099-9779-02ff529be895</mods:identifier>
<mods:identifier id="1DFE919A647FFE6EF26FF76CE21AF778" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="6D95E814005AF71E6D4892DDB40B09D3" type="Zenodo-Dep">3713503</mods:identifier>
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<treatment id="026C87B4F33EFFFFFF15FC3EFA39606A" ID-DOI="http://doi.org/10.5281/zenodo.3717390" ID-GBIF-Taxon="162629087" ID-Zenodo-Dep="3717390" LSID="urn:lsid:plazi:treatment:026C87B4F33EFFFFFF15FC3EFA39606A" httpUri="http://treatment.plazi.org/id/026C87B4F33EFFFFFF15FC3EFA39606A" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<subSubSection id="C2DF6529F33EFFF0FF15FC3EFC3367A0" box="[192,938,903,928]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8A7A36A2F33EFFF0FF15FC3EFC3367A0" blockId="7.[192,938,903,928]" box="[192,938,903,928]" pageId="7" pageNumber="6">
<heading id="D13281CEF33EFFF0FF15FC3EFC3367A0" bold="true" box="[192,938,903,928]" fontSize="10" level="2" pageId="7" pageNumber="6" reason="3">
<emphasis id="B8B1EAB0F33EFFF0FF15FC3EFC3367A0" bold="true" box="[192,938,903,928]" pageId="7" pageNumber="6">
<taxonomicName id="4DC54D21F33EFFF0FF15FC3EFD3667A0" authority="(Bechyne)" authorityName="Flowers" authorityYear="2018" baseAuthorityName="Bechyne" baseAuthorityYear="2018" box="[192,687,903,927]" class="Insecta" family="Chrysomelidae" genus="Beltia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="angustomarginata" status="comb. nov.">
<emphasis id="B8B1EAB0F33EFFF0FF15FC3EFD8267A0" bold="true" box="[192,539,903,927]" italics="true" pageId="7" pageNumber="6">Beltia angustomarginata</emphasis>
(Bechynĕ)
</taxonomicName>
,
<taxonomicNameLabel id="A38257CBF33EFFF0FD68FC31FC3367A0" box="[701,938,903,928]" pageId="7" pageNumber="6" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F33EFFF0FF15FC01FD8667EF" box="[192,543,952,976]" pageId="7" pageNumber="6" type="description">
<paragraph id="8A7A36A2F33EFFF0FF15FC01FD8667EF" blockId="7.[192,543,952,976]" box="[192,543,952,976]" pageId="7" pageNumber="6">
<figureCitation id="12FE2A27F33EFFF0FF15FC01FEE967EF" box="[192,368,952,976]" captionStart="Figures 9–17" captionStartId="33.[192,287,1820,1842]" captionTargetBox="[220,1417,236,1764]" captionTargetId="figure@33.[214,1417,213,1786]" captionTargetPageId="33" captionText="Figures 9–17. Structures of Beltia. 9) Profemur, B. nicaraguensis. 10) Profemur, B. tisingalita. 11) Pronotum, B. angustomarginata. 12) Pronotum, B. ledesmae. 13) En-face view of median lobe, B. angustomarginata. 14) Apex of abdomen, B. placidula. 15) Concave prosternum (arrow), B. ledesmae. 16) Head and pronotum, B. napoensis. 17) Male abdomen with seta patches (arrow), B. herreri." figureDoi="http://doi.org/10.5281/zenodo.3713507" httpUri="https://zenodo.org/record/3713507/files/figure.png" pageId="7" pageNumber="6">Figures 11, 13</figureCitation>
,
<figureCitation id="12FE2A27F33EFFF0FEA8FC01FE0467EF" box="[381,413,952,976]" captionStart="Figures 18–26" captionStartId="34.[192,287,1820,1842]" captionTargetBox="[212,1410,225,1758]" captionTargetId="figure@34.[204,1428,213,1789]" captionTargetPageId="34" captionText="Figures 18–26. Beltia, dorsal habitus. 18) Beltia angustomarginata, male. 19) Beltia awapita, male. 20) Beltia chiriquensis, female. 21) Beltia confusa, male. 22) Beltia gorgona, female. 23) Beltia herreri, male. 24–26) Beltia ledesmae, females. 24) Monochrome specimen. 25) Bicolored specimen. 26) Same female as (25) photographed with diffuser." figureDoi="http://doi.org/10.5281/zenodo.3713509" httpUri="https://zenodo.org/record/3713509/files/figure.png" pageId="7" pageNumber="6">18</figureCitation>
,
<figureCitation id="12FE2A27F33EFFF0FE7FFC01FE5367EF" box="[426,458,952,976]" captionStart="Figures 42–49" captionStartId="37.[192,287,1850,1872]" captionTargetBox="[261,1391,300,1811]" captionTargetId="figure@37.[225,1408,290,1816]" captionTargetPageId="37" captionText="Figures 42–49. Beltia, male genitalia: a) lateral view of median lobe; b) endophallic apical sclerites; c) en-face view of median lobe. 42) Beltia angustomarginata. 43) Beltia awapita. 44) Beltia chiriquensis. 45) Beltia confusa. 46) Beltia gorgona. 47) Beltia herreri. 48) Beltia ledesmae. 49) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713515" httpUri="https://zenodo.org/record/3713515/files/figure.png" pageId="7" pageNumber="6">42</figureCitation>
,
<figureCitation id="12FE2A27F33EFFF0FE03FC01FE6F67EF" box="[470,502,952,976]" captionStart="Figures 60–66" captionStartId="40.[192,287,1847,1869]" captionTargetBox="[222,1412,261,1803]" captionTargetId="figure@40.[192,1440,247,1814]" captionTargetPageId="40" captionText="Figures 60–66. Beltia, female genitalia: a) ovipositor, ventral view; b) spermatheca. 60) Beltia angustomarginata. 61) Beltia chiriquensis. 62) Beltia gorgona. 63) Beltia herreri. 64) Beltia ledesmae. 65) Beltia napoensis. 66) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713521" httpUri="https://zenodo.org/record/3713521/files/figure.png" pageId="7" pageNumber="6">60</figureCitation>
,
<figureCitation id="12FE2A27F33EFFF0FDD7FC01FD8667EF" box="[514,543,952,976]" captionStart="Figures 80–82" captionStartId="44.[192,286,1848,1870]" captionTargetBox="[203,1427,215,1809]" captionTargetId="figure@44.[203,1429,211,1815]" captionTargetPageId="44" captionText="Figures 80–82. Distribution maps for Beltia spp.: Nicaragua, Costa Rica, and Panama. 80) Beltia angustomarginata (squares), B. osa (triangles), B. vacilona (circles). 81) Beltia chiriquensis (circles), B. herreri (stars), B. nicaraguensis (hexagons). 82) Beltia sanchezae (square), B. tilarana (inverted triangles), B. tisingalita (stars)." figureDoi="http://doi.org/10.5281/zenodo.3713529" httpUri="https://zenodo.org/record/3713529/files/figure.png" pageId="7" pageNumber="6">80</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F33EFFF0FF15FC5FFA396026" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="8A7A36A2F33EFFF0FF15FC5FFA396026" blockId="7.[192,1440,998,1049]" pageId="7" pageNumber="6">
<treatmentCitationGroup id="AAD5118CF33EFFF0FF15FC5FFA396026" pageId="7" pageNumber="6">
<treatmentCitation id="0B6410B3F33EFFF0FF15FC5FFCFC67C4" author="Bechyne, J." box="[192,869,998,1020]" page="123" pageId="7" pageNumber="6" year="1953">
<taxonomicName id="4DC54D21F33EFFF0FF15FC5FFCFC67C4" authority="Bechyne 1953: 123" authorityName="Bechyne" authorityPageNumber="123" authorityYear="1953" box="[192,869,998,1020]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="placidula" subSpecies="angustomarginata">
<emphasis id="B8B1EAB0F33EFFF0FF15FC5FFD1067C4" box="[192,649,998,1019]" italics="true" pageId="7" pageNumber="6">Colaspoides placidula angustomarginata</emphasis>
<bibRefCitation id="EE544B53F33EFFF0FD47FC5FFCFC67C4" author="Bechyne, J." box="[658,869,998,1020]" pageId="7" pageNumber="6" pagination="26 - 303" refId="ref24974" refString="Bechyne, J. 1953. Katalog der neotropischen eumolpiden (Col. Phytoph. Chrysomeloidea). Entomologische Arbeiten aus dem Museum G. Frey 4: 26 - 303." type="journal article" year="1953">Bechynĕ 1953: 123</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0B6410B3F33EFFF0FBB3FC5FFAA567C4" author="Bechyne, J." box="[1126,1340,998,1020]" page="279" pageId="7" pageNumber="6" year="1953">
<bibRefCitation id="EE544B53F33EFFF0FBB3FC5FFAA567C4" author="Bechyne, J." box="[1126,1340,998,1020]" pageId="7" pageNumber="6" pagination="26 - 303" refId="ref24974" refString="Bechyne, J. 1953. Katalog der neotropischen eumolpiden (Col. Phytoph. Chrysomeloidea). Entomologische Arbeiten aus dem Museum G. Frey 4: 26 - 303." type="journal article" year="1953">Bechynĕ 1953: 279</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0B6410B3F33EFFF0FA9DFC5FFED16027" author="Flowers, R. W." page="31" pageId="7" pageNumber="6" year="1996">
<bibRefCitation id="EE544B53F33EFFF0FA9DFC5FFED16027" author="Flowers, R. W." pageId="7" pageNumber="6" pagination="1 - 60" refId="ref25104" refString="Flowers, R. W. 1996. La subfamilia Eumolpinae (Coleoptera: Chrysomelidae) en America Central. Revista de Biologia Tropical, Publicacion Especial 2: 1 - 60." type="journal article" year="1996">Flowers 1996: 31</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="557E8800F33EFFF0FE87FBBAFE2F6026" box="[338,438,1027,1049]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
deposited in NHMUK, photos of
<typeStatus id="557E8800F33EFFF0FCF4FBBAFCE66026" box="[801,895,1027,1049]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
seen. Type locality:
<collectingCountry id="F2D27632F33EFFF0FB8AFBBAFB276026" box="[1119,1214,1027,1049]" name="Panama" pageId="7" pageNumber="6">Panamá</collectingCountry>
: Gatun,
<date id="FE7B1062F33EFFF0FACEFBBAFA056026" box="[1307,1436,1027,1049]" pageId="7" pageNumber="6" value="1925-02-01" valueMax="1925-02-03" valueMin="1925-02-01">1–3.II.1925</date>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F33EFFFFFF15FB88FAF4665A" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="description">
<paragraph id="8A7A36A2F33EFFF0FF15FB88FAA760B6" blockId="7.[192,1440,1073,1161]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15FB88FE136076" bold="true" box="[192,394,1073,1097]" pageId="7" pageNumber="6">Redescription.</emphasis>
Male (
<figureCitation id="12FE2A27F33EFFF0FE34FB88FDA06076" box="[481,569,1073,1097]" captionStart="Figures 18–26" captionStartId="34.[192,287,1820,1842]" captionTargetBox="[212,1410,225,1758]" captionTargetId="figure@34.[204,1428,213,1789]" captionTargetPageId="34" captionText="Figures 18–26. Beltia, dorsal habitus. 18) Beltia angustomarginata, male. 19) Beltia awapita, male. 20) Beltia chiriquensis, female. 21) Beltia confusa, male. 22) Beltia gorgona, female. 23) Beltia herreri, male. 24–26) Beltia ledesmae, females. 24) Monochrome specimen. 25) Bicolored specimen. 26) Same female as (25) photographed with diffuser." figureDoi="http://doi.org/10.5281/zenodo.3713509" httpUri="https://zenodo.org/record/3713509/files/figure.png" pageId="7" pageNumber="6">Fig. 18</figureCitation>
). Body ovate, dorsally convex; length
<quantity id="4D3D9B47F33EFFF0FBDEFB88FBF26076" box="[1035,1131,1073,1097]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="6" unit="mm" value="6.1">6.1 mm</quantity>
. Head, pronotum, elytra, and underside metallic dark blue with strong purple reflexion; antennae testaceous with apical four antennomeres darker, legs dark reddish brown, central parts of femora shining greenish blue.
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15FB18FD906126" blockId="7.[192,1440,1185,1305]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15FB18FE896086" bold="true" box="[192,272,1185,1209]" pageId="7" pageNumber="6">Head.</emphasis>
Clypeus densely punctate, punctures separated by distance subequal to their diameters. Frontoclypeal suture distinct. Frons strongly punctate, surface between punctures smooth, punctures separated by distance greater than their diameters; vertex with a distinct median impressed line; antennal calli weakly differentiated.
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15FA88FC7561F6" blockId="7.[192,1440,1329,1481]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15FA88FEB56176" bold="true" box="[192,300,1329,1353]" pageId="7" pageNumber="6">Thorax.</emphasis>
Prothorax distinctly wider than long, L/W = 0.5 (
<figureCitation id="12FE2A27F33EFFF0FC5BFA88FC7D6176" box="[910,996,1329,1353]" captionStart="Figures 9–17" captionStartId="33.[192,287,1820,1842]" captionTargetBox="[220,1417,236,1764]" captionTargetId="figure@33.[214,1417,213,1786]" captionTargetPageId="33" captionText="Figures 9–17. Structures of Beltia. 9) Profemur, B. nicaraguensis. 10) Profemur, B. tisingalita. 11) Pronotum, B. angustomarginata. 12) Pronotum, B. ledesmae. 13) En-face view of median lobe, B. angustomarginata. 14) Apex of abdomen, B. placidula. 15) Concave prosternum (arrow), B. ledesmae. 16) Head and pronotum, B. napoensis. 17) Male abdomen with seta patches (arrow), B. herreri." figureDoi="http://doi.org/10.5281/zenodo.3713507" httpUri="https://zenodo.org/record/3713507/files/figure.png" pageId="7" pageNumber="6">Fig. 11</figureCitation>
,
<figureCitation id="12FE2A27F33EFFF0FC27FA88FB886176" box="[1010,1041,1329,1353]" captionStart="Figures 18–26" captionStartId="34.[192,287,1820,1842]" captionTargetBox="[212,1410,225,1758]" captionTargetId="figure@34.[204,1428,213,1789]" captionTargetPageId="34" captionText="Figures 18–26. Beltia, dorsal habitus. 18) Beltia angustomarginata, male. 19) Beltia awapita, male. 20) Beltia chiriquensis, female. 21) Beltia confusa, male. 22) Beltia gorgona, female. 23) Beltia herreri, male. 24–26) Beltia ledesmae, females. 24) Monochrome specimen. 25) Bicolored specimen. 26) Same female as (25) photographed with diffuser." figureDoi="http://doi.org/10.5281/zenodo.3713509" httpUri="https://zenodo.org/record/3713509/files/figure.png" pageId="7" pageNumber="6">18</figureCitation>
); disc regularly, finely punctate, punctures separated by distance greater than their diameters; surface between punctures glossy, with numerous punctulae. Prosternum smooth, sparsely punctate, glabrous; posterior margin of intercoxal process slightly concave, width of intercoxal process 1.1× diameter of procoxa. Metasternum smooth in center, finely wrinkled laterally; metepisternum finely alutaceous.
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15FA58FD2A6206" blockId="7.[192,1440,1505,1593]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15FA58FE8761C6" bold="true" box="[192,286,1505,1529]" pageId="7" pageNumber="6">Elytra.</emphasis>
Evenly punctate, punctures separated by distance greater than their diameters, surface between punctures with small punctulae; width across humeri 1.1× width across pronotum. Basal calli weakly developed, postbasal depression shallow.
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15F9E8FBAB6296" blockId="7.[192,1440,1617,1706]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15F9E8FED06256" bold="true" box="[192,329,1617,1641]" pageId="7" pageNumber="6">Abdomen.</emphasis>
Sterna with numerous short setae, and long setae across middle third at rear margin of segments III–VI, surface of segments alutaceous. Terga with a row of distinct spicules along base of segments II and III and laterally on IV; surface of V and VI shagreened.
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15F978FDDE6306" blockId="7.[192,1441,1729,1849]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15F978FED162E6" bold="true" box="[192,328,1729,1753]" pageId="7" pageNumber="6">Genitalia.</emphasis>
Median lobe in lateral view strongly curved, with lateral flanges (
<figureCitation id="12FE2A27F33EFFF0FBBAF978FB4C62E6" box="[1135,1237,1729,1753]" captionStart="Figures 42–49" captionStartId="37.[192,287,1850,1872]" captionTargetBox="[261,1391,300,1811]" captionTargetId="figure@37.[225,1408,290,1816]" captionTargetPageId="37" captionText="Figures 42–49. Beltia, male genitalia: a) lateral view of median lobe; b) endophallic apical sclerites; c) en-face view of median lobe. 42) Beltia angustomarginata. 43) Beltia awapita. 44) Beltia chiriquensis. 45) Beltia confusa. 46) Beltia gorgona. 47) Beltia herreri. 48) Beltia ledesmae. 49) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713515" httpUri="https://zenodo.org/record/3713515/files/figure.png" pageId="7" pageNumber="6">Fig. 42a</figureCitation>
); in en-face view “arrowhead”-shaped (
<figureCitation id="12FE2A27F33EFFF0FE1DF958FDB562C6" box="[456,556,1761,1785]" captionStart="Figures 42–49" captionStartId="37.[192,287,1850,1872]" captionTargetBox="[261,1391,300,1811]" captionTargetId="figure@37.[225,1408,290,1816]" captionTargetPageId="37" captionText="Figures 42–49. Beltia, male genitalia: a) lateral view of median lobe; b) endophallic apical sclerites; c) en-face view of median lobe. 42) Beltia angustomarginata. 43) Beltia awapita. 44) Beltia chiriquensis. 45) Beltia confusa. 46) Beltia gorgona. 47) Beltia herreri. 48) Beltia ledesmae. 49) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713515" httpUri="https://zenodo.org/record/3713515/files/figure.png" pageId="7" pageNumber="6">Fig. 42c</figureCitation>
), lateral flanges strongly sclerotized, apicolateral margins convergent to a small median triangular projection; apex of endophallus with a patch of strong spicules and a small curved apical sclerite (
<figureCitation id="12FE2A27F33EFFF0FE01F898FDA26306" box="[468,571,1825,1849]" captionStart="Figures 42–49" captionStartId="37.[192,287,1850,1872]" captionTargetBox="[261,1391,300,1811]" captionTargetId="figure@37.[225,1408,290,1816]" captionTargetPageId="37" captionText="Figures 42–49. Beltia, male genitalia: a) lateral view of median lobe; b) endophallic apical sclerites; c) en-face view of median lobe. 42) Beltia angustomarginata. 43) Beltia awapita. 44) Beltia chiriquensis. 45) Beltia confusa. 46) Beltia gorgona. 47) Beltia herreri. 48) Beltia ledesmae. 49) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713515" httpUri="https://zenodo.org/record/3713515/files/figure.png" pageId="7" pageNumber="6">Fig. 42b</figureCitation>
).
</paragraph>
<paragraph id="8A7A36A2F33EFFF0FF15F8E8FC0A63B6" blockId="7.[192,1439,1873,1929]" pageId="7" pageNumber="6">
<emphasis id="B8B1EAB0F33EFFF0FF15F8E8FEB56356" bold="true" box="[192,300,1873,1897]" pageId="7" pageNumber="6">Female.</emphasis>
Length
<quantity id="4D3D9B47F33EFFF0FE5AF8E8FD876356" box="[399,542,1873,1897]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.6" metricValueMin="6.1" pageId="7" pageNumber="6" unit="mm" value="6.85" valueMax="7.6" valueMin="6.1">6.1–7.6 mm</quantity>
; color similar to male but purple reflex stronger; antennae testaceous, with apical three antennomeres piceous; color of legs as in male.
</paragraph>
<paragraph id="8A7A36A2F331FFFFFF15FF64FDB264CA" blockId="8.[192,555,221,245]" box="[192,555,221,245]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FF64FE8964CA" bold="true" box="[192,272,221,245]" pageId="8" pageNumber="7">Head.</emphasis>
Punctation as in male.
</paragraph>
<paragraph id="8A7A36A2F331FFFFFF15FEB4FA39657A" blockId="8.[192,1440,269,325]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FEB4FEB3651A" bold="true" box="[192,298,269,293]" pageId="8" pageNumber="7">Thorax.</emphasis>
Pronotum with proportions slightly less transverse than male, punctation as in male. Surface of prosternum with short whitish setae, posterior margin less concave. Metasternum and elytra as in male.
</paragraph>
<paragraph id="8A7A36A2F331FFFFFF15FEE4FCA465EA" blockId="8.[192,1440,349,470]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FEE4FED0654A" bold="true" box="[192,329,349,373]" pageId="8" pageNumber="7">Abdomen.</emphasis>
Sterna with scattered whitish setae, apex of sternum VII weakly crenulate, with apical, shallow, V-shaped emargination and a short, subapical transverse costa. Terga heavily sclerotized, surface alutaceous; a row of spicules along base of tergum II, a short basal row on either side of midline on terga III and IV; surface of V and VI shagreened.
</paragraph>
<paragraph id="8A7A36A2F331FFFFFF15FE54FAF4665A" blockId="8.[192,1441,493,613]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FE54FED1663A" bold="true" box="[192,328,493,517]" pageId="8" pageNumber="7">Genitalia.</emphasis>
Segments VIII–XI forming an elongate (L/W = 5.0) ovipositor (
<figureCitation id="12FE2A27F331FFFFFB97FE54FB31663A" box="[1090,1192,493,517]" captionStart="Figures 60–66" captionStartId="40.[192,287,1847,1869]" captionTargetBox="[222,1412,261,1803]" captionTargetId="figure@40.[192,1440,247,1814]" captionTargetPageId="40" captionText="Figures 60–66. Beltia, female genitalia: a) ovipositor, ventral view; b) spermatheca. 60) Beltia angustomarginata. 61) Beltia chiriquensis. 62) Beltia gorgona. 63) Beltia herreri. 64) Beltia ledesmae. 65) Beltia napoensis. 66) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713521" httpUri="https://zenodo.org/record/3713521/files/figure.png" pageId="8" pageNumber="7">Fig. 60a</figureCitation>
). Sternum VIII with a long, wire-like basal apodeme, slightly widened at apex; segment IX covered with minute spicules; hemisternites with elongate basal rods; baculum distinct, elongate; gonocoxae distinctly longer than wide. Spermatheca (
<figureCitation id="12FE2A27F331FFFFFE6FFDF4FD87665A" box="[442,542,589,613]" captionStart="Figures 60–66" captionStartId="40.[192,287,1847,1869]" captionTargetBox="[222,1412,261,1803]" captionTargetId="figure@40.[192,1440,247,1814]" captionTargetPageId="40" captionText="Figures 60–66. Beltia, female genitalia: a) ovipositor, ventral view; b) spermatheca. 60) Beltia angustomarginata. 61) Beltia chiriquensis. 62) Beltia gorgona. 63) Beltia herreri. 64) Beltia ledesmae. 65) Beltia napoensis. 66) Beltia nicaraguensis." figureDoi="http://doi.org/10.5281/zenodo.3713521" httpUri="https://zenodo.org/record/3713521/files/figure.png" pageId="8" pageNumber="7">Fig. 60b</figureCitation>
) with bulb-like receptacle, distinctly constricted at union with pump.
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F331FFFFFF15FDC4FAE866CA" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="8A7A36A2F331FFFFFF15FDC4FAE866CA" blockId="8.[192,1440,636,758]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FDC4FE7066AA" bold="true" box="[192,489,637,661]" pageId="8" pageNumber="7">Specimens examined.</emphasis>
(
<specimenCount id="9CC3FD2BF331FFFFFE2CFDC4FD8566AA" box="[505,540,637,661]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
,
<specimenCount id="9CC3FD2BF331FFFFFDFCFDC4FDD266AA" box="[553,587,637,661]" pageId="8" pageNumber="7" type="female">4♀</specimenCount>
)
<materialsCitation id="3AAD3CFFF331FFFFFD89FDC4FDCC668A" ID-GBIF-Occurrence="2577335588" collectingDate="2012-02-17" collectingDateMax="2012-02-21" collectingDateMin="2012-02-17" collectionCode="II-, FSCA" collectorName="Col. &amp; J. B. Heppner" country="Panama" location="Sierra Llorona Lodge" municipality="Colon" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Colon">
<collectingCountry id="F2D27632F331FFFFFD89FDC4FD4366AA" box="[604,730,637,661]" name="Panama" pageId="8" pageNumber="7">
<collectingRegion id="4801F840F331FFFFFD89FDC4FD4366AA" box="[604,730,637,661]" country="Panama" name="Panama" pageId="8" pageNumber="7">PANAMA</collectingRegion>
</collectingCountry>
:
<emphasis id="B8B1EAB0F331FFFFFD31FDC5FC1F66AA" bold="true" box="[740,902,636,661]" pageId="8" pageNumber="7">
<collectingRegion id="4801F840F331FFFFFD31FDC5FC1F66AA" box="[740,902,636,661]" country="Panama" name="Colon" pageId="8" pageNumber="7">Colón Prov.</collectingRegion>
</emphasis>
(
<specimenCount id="9CC3FD2BF331FFFFFC43FDC4FC2166AA" box="[918,952,637,661]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
)
<collectingMunicipality id="6A1EACD8F331FFFFFC1DFDC4FB8866AA" box="[968,1041,637,661]" pageId="8" pageNumber="7">
<collectingRegion id="4801F840F331FFFFFC1DFDC4FB8866AA" box="[968,1041,637,661]" country="Panama" name="Colon" pageId="8" pageNumber="7">Colón</collectingRegion>
</collectingMunicipality>
,
<location id="8F1A6079F331FFFFFBC8FDC4FABD66AA" LSID="urn:lsid:plazi:treatment:026C87B4F33EFFFFFF15FC3EFA39606A:8F1A6079F331FFFFFBC8FDC4FABD66AA" box="[1053,1316,637,661]" country="Panama" municipality="Colon" name="Sierra Llorona Lodge" pageId="8" pageNumber="7" stateProvince="Colon">Sierra Llorona Lodge</location>
,
<date id="FE7B1062F331FFFFFAFAFDC4FF67668A" pageId="8" pageNumber="7" value="2012-02-17" valueMax="2012-02-21" valueMin="2012-02-17">
<collectingDate id="EE3FE98AF331FFFFFAFAFDC4FF67668A" pageId="8" pageNumber="7" value="2012-02-17" valueMax="2012-02-21" valueMin="2012-02-17">
17–21-
<collectionCode id="ECD4AE67F331FFFFFA54FDC4FA3966AA" box="[1409,1440,637,661]" pageId="8" pageNumber="7">II-</collectionCode>
2012
</collectingDate>
</date>
,
<collectorName id="27305374F331FFFFFEDEFD24FEA6668A" box="[267,319,669,693]" pageId="8" pageNumber="7">Col.</collectorName>
:
<collectorName id="27305374F331FFFFFE9EFD27FE76668A" box="[331,495,670,693]" pageId="8" pageNumber="7">J.B. Heppner</collectorName>
(
<collectionCode id="ECD4AE67F331FFFFFDD5FD24FDD4668A" box="[512,589,669,693]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="8" pageNumber="7">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3AAD3CFFF331FFFFFDB7FD25FD3C66EA" ID-GBIF-Occurrence="2577335593" collectingDate="1987-01-05" collectingDateMax="1987-01-09" collectingDateMin="1987-01-05" collectionCode="USNM" collectorName="Windopans &amp; H. Wolda &amp; Gpo." country="Panama" location="Barro Colorado Island" pageId="8" pageNumber="7" specimenCount="3" specimenCount-female="3" stateProvince="Panama">
<emphasis id="B8B1EAB0F331FFFFFDB7FD25FCBF668A" bold="true" box="[610,806,668,693]" pageId="8" pageNumber="7">
<collectingRegion id="4801F840F331FFFFFDB7FD25FCBF668A" box="[610,806,668,693]" country="Panama" name="Panama" pageId="8" pageNumber="7">
<collectingCountry id="F2D27632F331FFFFFDB7FD25FD4A668B" box="[610,723,668,692]" name="Panama" pageId="8" pageNumber="7">Panamá</collectingCountry>
Prov.
</collectingRegion>
</emphasis>
(
<specimenCount id="9CC3FD2BF331FFFFFCECFD24FCCE668A" box="[825,855,669,693]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
)
<location id="8F1A6079F331FFFFFCBEFD27FBE7668A" LSID="urn:lsid:plazi:treatment:026C87B4F33EFFFFFF15FC3EFA39606A:8F1A6079F331FFFFFCBEFD27FBE7668A" box="[875,1150,669,693]" country="Panama" name="Barro Colorado Island" pageId="8" pageNumber="7" stateProvince="Panama">
Barro Colorado
<collectingCountry id="F2D27632F331FFFFFBE5FD24FBE7668A" box="[1072,1150,669,693]" name="Iceland" pageId="8" pageNumber="7">Island</collectingCountry>
</location>
<date id="FE7B1062F331FFFFFB53FD24FA9F668A" box="[1158,1286,669,693]" pageId="8" pageNumber="7" value="1987-01-05" valueMax="1987-01-09" valueMin="1987-01-05">
<collectingDate id="EE3FE98AF331FFFFFB53FD24FA9F668A" box="[1158,1286,669,693]" pageId="8" pageNumber="7" value="1987-01-05" valueMax="1987-01-09" valueMin="1987-01-05">5–9.I.1987</collectingDate>
</date>
<collectorName id="27305374F331FFFFFADAFD24FA05668A" box="[1295,1436,669,693]" pageId="8" pageNumber="7">Windopans</collectorName>
, forest
<collectorName id="27305374F331FFFFFEDAFD07FE1E66EA" box="[271,391,701,725]" pageId="8" pageNumber="7">H. Wolda</collectorName>
,
<collectorName id="27305374F331FFFFFE40FD04FE5766EA" box="[405,462,701,725]" pageId="8" pageNumber="7">Gpo.</collectorName>
4B (
<collectionCode id="ECD4AE67F331FFFFFDD4FD04FDC366EA" box="[513,602,701,725]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="8" pageNumber="7">USNM</collectionCode>
); (
<specimenCount id="9CC3FD2BF331FFFFFDAFFD04FD0266EA" box="[634,667,701,725]" pageId="8" pageNumber="7" type="female">2♀</specimenCount>
)
</materialsCitation>
<materialsCitation id="3AAD3CFFF331FFFFFD7BFD04FE4B66CA" ID-GBIF-Occurrence="2577335579" collectingDate="1992-05-24" collectingDateMax="1992-06-03" collectingDateMin="1992-05-24" collectionCode="USNM" collectorName="El Llano &amp; J. E. Wappes" country="Panama" location="Carti rd." pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Panama">
<collectorName id="27305374F331FFFFFD7BFD04FC8566EA" box="[686,796,701,725]" pageId="8" pageNumber="7">El Llano</collectorName>
,
<location id="8F1A6079F331FFFFFCFFFD04FC0F66EA" LSID="urn:lsid:plazi:treatment:026C87B4F33EFFFFFF15FC3EFA39606A:8F1A6079F331FFFFFCFFFD04FC0F66EA" box="[810,918,701,725]" country="Panama" name="Carti rd." pageId="8" pageNumber="7" stateProvince="Panama">Carti rd.</location>
km
<date id="FE7B1062F331FFFFFC04FD04FB9766EA" box="[977,1038,701,725]" pageId="8" pageNumber="7" value="1992-05">8–11</date>
,
<date id="FE7B1062F331FFFFFBC8FD04FA8166EA" box="[1053,1304,701,725]" pageId="8" pageNumber="7" value="1992-05-24" valueMax="1992-06-03" valueMin="1992-05-24">
<collectingDate id="EE3FE98AF331FFFFFBC8FD04FA8166EA" box="[1053,1304,701,725]" pageId="8" pageNumber="7" value="1992-05-24" valueMax="1992-06-03" valueMin="1992-05-24">24 May–3 Jun 1992</collectingDate>
</date>
, 1100′
<collectorName id="27305374F331FFFFFABAFD07FEB866CA" pageId="8" pageNumber="7">J.E. Wappes</collectorName>
(
<collectionCode id="ECD4AE67F331FFFFFEE5FD64FE1066CA" box="[304,393,733,757]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="8" pageNumber="7">USNM</collectionCode>
); (
<specimenCount id="9CC3FD2BF331FFFFFE7DFD64FE5F66CA" box="[424,454,733,757]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
)
</materialsCitation>
<materialsCitation id="3AAD3CFFF331FFFFFE0CFD65FAF466CA" ID-GBIF-Occurrence="2577335587" box="[473,1389,732,758]" collectingDate="1978-05-13" collectionCode="FSCA" collectorName="LB O'Brien &amp; Marshall" country="Panama" location="Cerro Campana" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Panama">
2700′,
<location id="8F1A6079F331FFFFFDFDFD64FD7766CA" LSID="urn:lsid:plazi:treatment:026C87B4F33EFFFFFF15FC3EFA39606A:8F1A6079F331FFFFFDFDFD64FD7766CA" box="[552,750,733,757]" country="Panama" name="Cerro Campana" pageId="8" pageNumber="7" stateProvince="Panama">Cerro Campana</location>
,
<date id="FE7B1062F331FFFFFD2FFD67FC0766CA" box="[762,926,733,758]" pageId="8" pageNumber="7" value="1978-05-13">
<collectingDate id="EE3FE98AF331FFFFFD2FFD67FC0766CA" box="[762,926,733,758]" pageId="8" pageNumber="7" value="1978-05-13">May 13, 1978</collectingDate>
</date>
CW&amp;
<collectorName id="27305374F331FFFFFC3FFD64FBED66CA" box="[1002,1140,733,757]" pageId="8" pageNumber="7">LB O’Brien</collectorName>
&amp;
<collectorName id="27305374F331FFFFFB4DFD64FA9E66CA" box="[1176,1287,733,757]" pageId="8" pageNumber="7">Marshall</collectorName>
(
<collectionCode id="ECD4AE67F331FFFFFAC2FD64FAFC66CA" box="[1303,1381,733,757]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="8" pageNumber="7">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F331FFFFFF15FCB4FD6F675A" pageId="8" pageNumber="7" type="diagnosis">
<paragraph id="8A7A36A2F331FFFFFF15FCB4FD6F675A" blockId="8.[192,1440,781,870]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FCB4FED4671A" bold="true" box="[192,333,781,805]" pageId="8" pageNumber="7">Diagnosis.</emphasis>
Males of
<taxonomicName id="4DC54D21F331FFFFFE69FCB4FD70671B" authorityName="Flowers" authorityYear="2018" baseAuthorityName="Bechyne" baseAuthorityYear="2018" box="[444,745,781,805]" class="Insecta" family="Chrysomelidae" genus="Beltia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="angustomarginata">
<emphasis id="B8B1EAB0F331FFFFFE69FCB4FD70671B" box="[444,745,781,805]" italics="true" pageId="8" pageNumber="7">Beltia angustomarginata</emphasis>
</taxonomicName>
can be recognized by the wide pronotum and “arrowhead”- shaped apex of the median lobe. Females have a less obviously transverse pronotum, but share with the male a strong purple reflex on the upper body.
</paragraph>
</subSubSection>
<subSubSection id="C2DF6529F331FFFFFF15FCC4FA39606A" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8A7A36A2F331FFFFFF15FCC4FA39606A" blockId="8.[192,1440,893,1109]" pageId="8" pageNumber="7">
<emphasis id="B8B1EAB0F331FFFFFF15FCC4FEDA67AA" bold="true" box="[192,323,893,917]" pageId="8" pageNumber="7">Remarks.</emphasis>
<bibRefCitation id="EE544B53F331FFFFFE9FFCC4FD9F67AA" author="Bechyne, J." box="[330,518,893,917]" pageId="8" pageNumber="7" pagination="26 - 303" refId="ref24974" refString="Bechyne, J. 1953. Katalog der neotropischen eumolpiden (Col. Phytoph. Chrysomeloidea). Entomologische Arbeiten aus dem Museum G. Frey 4: 26 - 303." type="journal article" year="1953">Bechynĕ (1953)</bibRefCitation>
described
<taxonomicName id="4DC54D21F331FFFFFD5CFCC4FC1467AB" authorityName="Flowers" authorityYear="2018" baseAuthorityName="Bechyne" baseAuthorityYear="2018" box="[649,909,893,916]" class="Insecta" family="Chrysomelidae" genus="Beltia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="angustomarginata">
<emphasis id="B8B1EAB0F331FFFFFD5CFCC4FC1467AB" box="[649,909,893,916]" italics="true" pageId="8" pageNumber="7">B. angustomarginata</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="4DC54D21F331FFFFFBA6FCC4FA9D67AA" box="[1139,1284,893,917]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B8B1EAB0F331FFFFFBA6FCC4FA9D67AA" box="[1139,1284,893,917]" italics="true" pageId="8" pageNumber="7">Colaspoides</emphasis>
</taxonomicName>
(now
<taxonomicName id="4DC54D21F331FFFFFA9BFCC4FA0E67AA" authorityName="Jacoby" authorityYear="1881" box="[1358,1431,893,917]" class="Insecta" family="Chrysomelidae" genus="Beltia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B8B1EAB0F331FFFFFA9BFCC4FA0E67AA" box="[1358,1431,893,917]" italics="true" pageId="8" pageNumber="7">Beltia</emphasis>
</taxonomicName>
)
<taxonomicName id="4DC54D21F331FFFFFF15FC24FEAD678A" box="[192,308,925,949]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="placidula">
<emphasis id="B8B1EAB0F331FFFFFF15FC24FEAD678A" box="[192,308,925,949]" italics="true" pageId="8" pageNumber="7">placidula</emphasis>
</taxonomicName>
, which is so far known only from a small area of
<collectingCountry id="F2D27632F331FFFFFC58FC27FC52678A" box="[909,971,926,949]" name="Peru" pageId="8" pageNumber="7">Peru</collectingCountry>
, based on a narrower pronotal margin and deeper postbasal depression. The structure and punctation of the pronotum in these two forms are different; additionally, in all specimens of
<taxonomicName id="4DC54D21F331FFFFFD27FC64FC1167CA" box="[754,904,989,1013]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="placidula">
<emphasis id="B8B1EAB0F331FFFFFD27FC64FC1167CA" box="[754,904,989,1013]" italics="true" pageId="8" pageNumber="7">B. placidula</emphasis>
</taxonomicName>
the legs are entirely dark fulvous to brown, whereas all known specimens of
<taxonomicName id="4DC54D21F331FFFFFD98FC44FCC9602B" authorityName="Flowers" authorityYear="2018" baseAuthorityName="Bechyne" baseAuthorityYear="2018" box="[589,848,1021,1044]" class="Insecta" family="Chrysomelidae" genus="Beltia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="angustomarginata">
<emphasis id="B8B1EAB0F331FFFFFD98FC44FCC9602B" box="[589,848,1021,1044]" italics="true" pageId="8" pageNumber="7">B. angustomarginata</emphasis>
</taxonomicName>
have metallic legs. This, along with the fact that both forms seem to have limited and disjunct distributions, appears to me to justify considering them distinct species. So far, this species (including the
<typeStatus id="557E8800F331FFFFFCD3FB84FCA3606A" box="[774,826,1085,1109]" pageId="8" pageNumber="7">type</typeStatus>
) has been collected only in central
<collectingCountry id="F2D27632F331FFFFFB1BFB87FAAB606A" box="[1230,1330,1086,1109]" name="Panama" pageId="8" pageNumber="7">Panama</collectingCountry>
(
<figureCitation id="12FE2A27F331FFFFFA95FB84FA0A606A" box="[1344,1427,1085,1109]" captionStart="Figures 80–82" captionStartId="44.[192,286,1848,1870]" captionTargetBox="[203,1427,215,1809]" captionTargetId="figure@44.[203,1429,211,1815]" captionTargetPageId="44" captionText="Figures 80–82. Distribution maps for Beltia spp.: Nicaragua, Costa Rica, and Panama. 80) Beltia angustomarginata (squares), B. osa (triangles), B. vacilona (circles). 81) Beltia chiriquensis (circles), B. herreri (stars), B. nicaraguensis (hexagons). 82) Beltia sanchezae (square), B. tilarana (inverted triangles), B. tisingalita (stars)." figureDoi="http://doi.org/10.5281/zenodo.3713529" httpUri="https://zenodo.org/record/3713529/files/figure.png" pageId="8" pageNumber="7">Fig. 80</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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