treatments-xml/data/02/4F/DF/024FDF7CFFAAE41CFF7B55B1FE1BFF24.xml
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<document id="3D38B663DB04A9C9189D659B911F13F3" ID-CLB-Dataset="3126" ID-DOI="10.11646/zootaxa.5072.6.4" ID-GBIF-Dataset="ec1c8bca-c18b-49cb-b4c1-ce7e33180d29" ID-ISSN="1175-5326" ID-Zenodo-Dep="5751653" ID-ZooBank="1CFF5E1E-7213-4B7B-A24F-953EA25B87D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638523388918" checkinUser="plazi" docAuthor="Han, Wu, Liu, Jie, Luo, Yifan &amp; Tang, Hongqu" docDate="2021" docId="024FDF7CFFAAE41CFF7B55B1FE1BFF24" docLanguage="en" docName="zootaxa.5072.6.4.pdf" docOrigin="Zootaxa 5072 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kribiodosis Kieffer 1921" docType="treatment" docVersion="7" lastPageNumber="563" masterDocId="FE76A704FFA9E418FFEC5703FF9CFFF1" masterDocTitle="No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses" masterLastPageNumber="574" masterPageNumber="560" pageNumber="563" updateTime="1699471950998" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1695A80142C3BBBE72489B6989BE1DCF">No longer endemic to Africa: Kribiodosis Kieffer, 1921 (Diptera, Chironomidae) new to Oriental China with a phylogeny and expanded adult generic diagnoses</mods:title>
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<mods:namePart id="8528D8091F222C0A7A3832328AD935D4">Han, Wu</mods:namePart>
<mods:affiliation id="6CF4C651E52EFE0479C7DDE117DF19F8">State Key Laboratory of Lake Science and Environment, Nanjing Institute of Geography and Limnology, Chinese Academy of Science, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="F373FE79C5FE41D9D6A9BF2F1BB5B27D">Liu, Jie</mods:namePart>
<mods:affiliation id="497227C50AD1A3DF8C3BFEEE9A99CF22">Guangzhou Medical University, Guangzhou 510632, China &amp; Life Science and Technology College, Jinan University, Guangzhou 510632, China</mods:affiliation>
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<mods:namePart id="12D32E603839BABF10D91AD6B4FB57AD">Luo, Yifan</mods:namePart>
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<mods:affiliation id="D5CFEC927275B84D400594CE1FF0C727">Life Science and Technology College, Jinan University, Guangzhou 510632, China &amp; lyf 1999 @ stu 2020. jnu. edu. cn; https: // orcid. org / 0000 - 0002 - 8805 - 3590</mods:affiliation>
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<mods:namePart id="07EDAF7A30DC3BFEBE2AF5553352BD23">Tang, Hongqu</mods:namePart>
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<mods:affiliation id="E7BF7B24C53F33867AACCA7123E10D6C">Life Science and Technology College, Jinan University, Guangzhou 510632, China &amp; Tang: thqtang @ jnu. edu. cn; townningt @ gmail. com; https: // orcid. org / 0000 - 0002 - 4957 - 4617</mods:affiliation>
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Genus
<taxonomicName id="4DE615E9FFAAE41BFF0255B1FD83FD3D" ID-CoL="595W" authority="Kieffer, 1921" authorityName="Kieffer" authorityYear="1921" box="[238,543,690,717]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="genus">
<emphasis id="B892B278FFAAE41BFF0255B1FEEBFD3C" bold="true" box="[238,375,690,717]" italics="true" pageId="3" pageNumber="563">Kribiodosis</emphasis>
<bibRefCitation id="EE77139BFFAAE41BFE9355B1FD83FD3D" author="Kieffer, J. J." box="[383,543,690,716]" pageId="3" pageNumber="563" pagination="1 - 56" refId="ref5440" refString="Kieffer, J. J. (1921) Chironomides de l'Afrique Equatoriale (1 e partie). Annales de la Societe entomologique de France, 90, 1 - 56." type="journal article" year="1921">Kieffer, 1921</bibRefCitation>
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<paragraph id="8A596E6AFFAAE41BFF7B55F8FBE8FCE4" blockId="3.[151,1140,762,825]" box="[151,1140,762,789]" pageId="3" pageNumber="563">
<typeStatus id="555DD0C8FFAAE41BFF7B55F8FF53FCE4" box="[151,207,763,789]" pageId="3" pageNumber="563">Type</typeStatus>
species:
<taxonomicName id="4DE615E9FFAAE41BFED955F9FD28FCE5" authority="Kieffer, 1921" authorityName="Kieffer" authorityYear="1921" box="[309,692,762,788]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B892B278FFAAE41BFED955F9FD84FCE5" box="[309,536,762,788]" italics="true" pageId="3" pageNumber="563">Kribiodosis fasciata</emphasis>
<bibRefCitation id="EE77139BFFAAE41BFDF355F9FD28FCE5" author="Kieffer, J. J." box="[543,692,762,788]" pageId="3" pageNumber="563" pagination="1 - 56" refId="ref5440" refString="Kieffer, J. J. (1921) Chironomides de l'Afrique Equatoriale (1 e partie). Annales de la Societe entomologique de France, 90, 1 - 56." type="journal article" year="1921">Kieffer, 1921</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4DE615E9FFAAE41BFD3755F9FBF1FCE4" authority="Kieffer, 1921" authorityName="Kieffer" authorityYear="1921" box="[731,1133,762,789]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="clavigera">
<emphasis id="B892B278FFAAE41BFD3755F9FC53FCE5" box="[731,975,762,788]" italics="true" pageId="3" pageNumber="563">Kribiodosis clavigera</emphasis>
<bibRefCitation id="EE77139BFFAAE41BFC3A55F9FBF1FCE4" author="Kieffer, J. J." box="[982,1133,762,789]" pageId="3" pageNumber="563" pagination="1 - 56" refId="ref5440" refString="Kieffer, J. J. (1921) Chironomides de l'Afrique Equatoriale (1 e partie). Annales de la Societe entomologique de France, 90, 1 - 56." type="journal article" year="1921">Kieffer, 1921</bibRefCitation>
</taxonomicName>
]
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Other included species:
<taxonomicName id="4DE615E9FFAAE41BFE49541DFD6EFCC8" authority="Tang" authorityName="Han &amp; Liu &amp; Luo &amp; Tang" authorityYear="2021" box="[421,754,798,825]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="cantonensis" status="sp. nov.">
<emphasis id="B892B278FFAAE41BFE49541DFD2FFCC9" box="[421,691,798,824]" italics="true" pageId="3" pageNumber="563">Kribiodosis cantonensis</emphasis>
Tang
</taxonomicName>
<emphasis id="B892B278FFAAE41BFD14541DFCA4FCC9" bold="true" box="[760,824,798,824]" pageId="3" pageNumber="563">
<taxonomicNameLabel id="A3A10F03FFAAE41BFD14541DFCA4FCC9" box="[760,824,798,824]" pageId="3" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF7B5465FA06FC71" blockId="3.[151,1437,870,2013]" box="[151,1434,870,897]" pageId="3" pageNumber="563">
These two species conformed largely to the generic diagnosis of
<bibRefCitation id="EE77139BFFAAE41BFC815464FBBFFC70" author="Freeman, P." box="[877,1059,870,897]" pageId="3" pageNumber="563" pagination="261 - 363" refId="ref5250" refString="Freeman, P. (1958) A study of the Chironomidae (Diptera) of Africa South of the Sahara. Part IV. Bulletin of the British Museum (Nature History) Entomology, 6 (11), 261 - 363. https: // doi. org / 10.5962 / bhl. part. 17110" type="journal article" year="1958">Freeman (1958)</bibRefCitation>
, with the following emendations.
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<paragraph id="8A596E6AFFAAE41BFF2B5489FB1DFBE1" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5489FE97FC55" bold="true" box="[199,267,906,932]" pageId="3" pageNumber="563">Male.</emphasis>
<emphasis id="B892B278FFAAE41BFEF85489FE13FC55" box="[276,399,906,932]" italics="true" pageId="3" pageNumber="563">Coloration</emphasis>
. Thorax generally brown in color, with variable extent and depth of pigment, scutal vittae dark or dark brown. Wing slightly brown, membrane weakly punctate. Leg brown or dark brown, with pale bands at the apices of all femorae and tibiae.Abdomen generally yellow, with banded intersegmental conjunctives on TI-VII. Segment VIII and IX usually wholly dark brown, gonocoxite and gonostylus pale yellow.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5319FEFDFB8C" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5319FE98FBC5" box="[199,260,1050,1076]" italics="true" pageId="3" pageNumber="563">Head</emphasis>
. Antenna with 13 segments (if counting as 12-segmented, then the boundary of last segmentation is indistinct). Frontal tubercle absent. Palp segment normal, the terminal segment usually no longer than the preterminal segment.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5385FF6CFB18" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5385FE84FB51" box="[199,280,1158,1184]" italics="true" pageId="3" pageNumber="563">Thorax</emphasis>
. With scutal tubercle (
<taxonomicName id="4DE615E9FFAAE41BFDF75384FD5CFB51" authority="Han &amp; Liu &amp; Luo &amp; Tang, 2021" authorityName="Han &amp; Liu &amp; Luo &amp; Tang" authorityYear="2021" box="[539,704,1159,1184]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="cantonensis" status="sp. nov.">
<emphasis id="B892B278FFAAE41BFDF75384FD5CFB51" box="[539,704,1159,1184]" italics="true" pageId="3" pageNumber="563">K. cantonensis</emphasis>
</taxonomicName>
<emphasis id="B892B278FFAAE41BFD255385FC95FB51" bold="true" box="[713,777,1158,1184]" pageId="3" pageNumber="563">
<taxonomicNameLabel id="A3A10F03FFAAE41BFD255385FC95FB51" box="[713,777,1158,1184]" pageId="3" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or without (
<taxonomicName id="4DE615E9FFAAE41BFC4D5384FBB0FB51" authorityName="Kieffer" authorityYear="1921" box="[929,1068,1158,1184]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="clavigera">
<emphasis id="B892B278FFAAE41BFC4D5384FBB0FB51" box="[929,1068,1158,1184]" italics="true" pageId="3" pageNumber="563">K. clavigera</emphasis>
</taxonomicName>
) a tubercle. Acrostichals absent, dorsocentrals and scutellars few (usually no more than five). Prealars and supraalars both absent. Humeral pits present.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B53F0FD70FA89" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B53F0FF60FAFD" box="[199,252,1267,1292]" italics="true" pageId="3" pageNumber="563">Legs</emphasis>
. Fore tibia with slender round scale bearing thin and pointed spur. Tibial combs of mid and hind tibiae separate (
<taxonomicName id="4DE615E9FFAAE41BFEEF5214FE12FAC1" authorityName="Kieffer" authorityYear="1921" box="[259,398,1302,1328]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="clavigera">
<emphasis id="B892B278FFAAE41BFEEF5214FE12FAC1" box="[259,398,1302,1328]" italics="true" pageId="3" pageNumber="563">K. clavigera</emphasis>
</taxonomicName>
) or fused (
<taxonomicName id="4DE615E9FFAAE41BFDE55214FD31FAC1" authority="Han &amp; Liu &amp; Luo &amp; Tang, 2021" authorityName="Han &amp; Liu &amp; Luo &amp; Tang" authorityYear="2021" box="[521,685,1303,1328]" class="Insecta" family="Chironomidae" genus="Kribiodosis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="cantonensis" status="sp. nov.">
<emphasis id="B892B278FFAAE41BFDE55214FD31FAC1" box="[521,685,1303,1328]" italics="true" pageId="3" pageNumber="563">K. cantonensis</emphasis>
</taxonomicName>
<emphasis id="B892B278FFAAE41BFD585215FD6FFAC1" bold="true" box="[692,755,1302,1328]" pageId="3" pageNumber="563">
<taxonomicNameLabel id="A3A10F03FFAAE41BFD585215FD6FFAC1" box="[692,755,1302,1328]" pageId="3" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); if separated, with only one hooked long spur in the smaller comb. Middle section of fourth tarsus of male fore leg with three dark brown pseudospurs, longer than the diameter of fourth tarsus, projecting inwardly. Pulvilli distinct.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5280FE6FFA15" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5280FE9DFA6D" box="[199,257,1411,1436]" italics="true" pageId="3" pageNumber="563">Wing</emphasis>
. Membrane without setae or microtrichia.Anal lobe absent. Costa ending abruptly at apex of R
<subScript id="16626C2FFFAAE41BFAFE5297FAB7FA53" attach="left" box="[1298,1323,1428,1442]" fontSize="6" pageId="3" pageNumber="563">4+5</subScript>
, proximal to wing apex; R
<subScript id="16626C2FFFAAE41BFEA452BAFEFDFA36" attach="left" box="[328,353,1465,1479]" fontSize="6" pageId="3" pageNumber="563">2+3</subScript>
ending near proximal 1/41/5 between R
<subScript id="16626C2FFFAAE41BFCC352BAFCABFA36" attach="left" box="[815,823,1465,1479]" fontSize="6" pageId="3" pageNumber="563">1</subScript>
and R
<subScript id="16626C2FFFAAE41BFC6C52BBFC06FA37" attach="left" box="[896,922,1464,1478]" fontSize="6" pageId="3" pageNumber="563">4+5</subScript>
. FCu clearly distal to RM. R bare, R
<subScript id="16626C2FFFAAE41BFADE52BAFAA6FA36" attach="left" box="[1330,1338,1465,1479]" fontSize="6" pageId="3" pageNumber="563">1</subScript>
and R
<subScript id="16626C2FFFAAE41BFA6F52BBFA01FA37" attach="left" box="[1411,1437,1464,1478]" fontSize="6" pageId="3" pageNumber="563">4+5</subScript>
with a few setae. Squama bare.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B52EDFD62F9F9" blockId="3.[151,1437,870,2013]" box="[199,766,1518,1545]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B52EDFEADF9F9" box="[199,305,1518,1544]" italics="true" pageId="3" pageNumber="563">Abdomen</emphasis>
. Segment VIII clearly tapered anteriorly.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5110FE20F94C" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5110FECAF9DD" box="[199,342,1555,1580]" italics="true" pageId="3" pageNumber="563">Hypopygium</emphasis>
. Anal tergite bands strong, meeting in median area. T IX with 12 median anal setae and 48 marginal setae on each side. Anal point short, parallel-sided, distally rounded. Superior volsella thin triangular, with narrow apex, inner margin with 23 basal setae. Inferior volsella setose, apex with 1012 strong setae, extending beyond apex of anal point. Gonostylus relatively short, with a slightly triangular apex, inner margin with 68 nearly equally spaced long setae.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B51C5FE83F8F5" blockId="3.[151,1437,870,2013]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B51C5FEB9F911" bold="true" box="[199,293,1734,1760]" pageId="3" pageNumber="563">Female.</emphasis>
<emphasis id="B892B278FFAAE41BFEC151C5FE34F911" box="[301,424,1734,1760]" italics="true" pageId="3" pageNumber="563">Coloration</emphasis>
. As male except wing somewhat smoky, and abdominal segments IVVIII nearly wholly dark brown.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B500DFCEAF8D9" blockId="3.[151,1437,870,2013]" box="[199,886,1806,1833]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B500DFE98F8D9" box="[199,260,1806,1832]" italics="true" pageId="3" pageNumber="563">Head</emphasis>
. Antenna with 4 or 5 segments. Frontal tubercle absent.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5031FE74F8BD" blockId="3.[151,1437,870,2013]" box="[199,488,1842,1869]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5031FE84F8BD" box="[199,280,1842,1868]" italics="true" pageId="3" pageNumber="563">Thorax</emphasis>
and
<emphasis id="B892B278FFAAE41BFEA35030FE19F8BD" box="[335,389,1843,1868]" italics="true" pageId="3" pageNumber="563">wing</emphasis>
as male.
</paragraph>
<paragraph id="8A596E6AFFAAE41BFF2B5054FBBAF881" blockId="3.[151,1437,870,2013]" box="[199,1062,1878,1905]" pageId="3" pageNumber="563">
<emphasis id="B892B278FFAAE41BFF2B5054FF60F881" box="[199,252,1879,1904]" italics="true" pageId="3" pageNumber="563">Legs</emphasis>
without pseudospurs on the fore tarsomere 4 on fore leg. Color as male.
</paragraph>
<paragraph id="8A596E6AFFAAE41CFF2B5079FE1BFF24" blockId="3.[151,1437,870,2013]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="564" pageId="3" pageNumber="563">Genitalia. S VIII with 1416 setae arising from the large pale pits on each side. Gonocoxapodeme straight, fused medially. Notum distinctly longer than the rami. Dorsomesal lobe prominent, expanded posterior medially; ventrolateral lobe absent or reduced into a narrow strip. Apodeme lobe lamellar, with slightly expanded apex. Labium apparently bare. Postgenital plate triangular. T IX with setae biserial, the upper row with 2 setae, and the lower with 68 setae.</paragraph>
</subSubSection>
</treatment>
</document>