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<document id="1711294BECAE4B64A0B7DE0A3A445223" ID-CLB-Dataset="6001" ID-DOI="10.11646/zootaxa.4980.2.8" ID-GBIF-Dataset="13ae2f6e-362f-4d58-a581-d3842dacfa6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4889248" ID-ZooBank="C55A445F-76EC-460A-B259-98AF744ECFFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622538088690" checkinUser="plazi" docAuthor="Sudhikumar, Ambalaparambil Vasu" docDate="2021" docId="024F878359677F30C5FD7A94FAB622D6" docLanguage="en" docName="zootaxa.4980.2.8.pdf" docOrigin="Zootaxa 4980 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Klimakodesmus bilobocaudatus Sudhikumar 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="380" masterDocId="FE76FFFB59647F37C56A7A4BFFC12550" masterDocTitle="The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae)" masterLastPageNumber="382" masterPageNumber="373" pageNumber="376" updateTime="1698995773394" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="77C491A7D1CD58AC5399EC0590E2BDE6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B3AA98E4F54B54BC72209C40688230B">
<mods:title id="1DC2FB84B5D02044D25254C6EE583C62">The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="942B4F83A7BB9778B75D6D6C316B96C1" type="personal">
<mods:role id="F4795D8C3E745808092444E2F5BD0F2C">
<mods:roleTerm id="03ECFB19021F0A7CFFB32957088F635F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C8868E8F8F110EBC8A94C37FC4E9E28">Sudhikumar, Ambalaparambil Vasu</mods:namePart>
<mods:nameIdentifier id="1BCDD4FB8361A5C753BC103E2F2C2539" type="ORCID">0000-0002-4479-4995</mods:nameIdentifier>
<mods:affiliation id="F3471AAC1427FA2036C887929CF07272">Centre for Animal Taxonomy and Ecology (CATE), Christ College, Irinjalakuda, Kerala, India &amp; spidersudhi @ gmail. com; https: // orcid. org / 0000 - 0002 - 4479 - 4995</mods:affiliation>
<mods:nameIdentifier id="B31849381422CA5E8800AC3473A50ED0" type="email">spidersudhi@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BDB77F2D6B7DF0DCF83E077FE037396F">text</mods:typeOfResource>
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<mods:titleInfo id="6BA53B8047123CD5018BBB6F9AEDFCED">
<mods:title id="001E0A3E809C56C9FDB092650E777691">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B7FBAE14CE47328BD469C251D03BD940">
<mods:date id="1C7FA4D801979AFA59305EB49A9CE589">2021</mods:date>
<mods:detail id="AB51B780699734512EA3417F245EBA6A" type="pubDate">
<mods:number id="67ED5B35FF8E3187B810EA61A8E08A64">2021-06-01</mods:number>
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<mods:number id="02C162719A0638C7B6D601EC24BC3B03">4980</mods:number>
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<mods:number id="B24F6C62052091A807F843AB76E0310B">2</mods:number>
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<mods:classification id="D43356C875F209CEB4B91A767098C1BC">journal article</mods:classification>
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<mods:identifier id="DF2F6AEF3C0D439631AA3BAD50147E3C" type="DOI">10.11646/zootaxa.4980.2.8</mods:identifier>
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<mods:identifier id="BEF6DAA99079BF873606BC4837441E8E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="024F878359677F30C5FD7A94FAB622D6" ID-DOI="http://doi.org/10.5281/zenodo.4986346" ID-GBIF-Taxon="182958732" ID-Zenodo-Dep="4986346" LSID="urn:lsid:plazi:treatment:024F878359677F30C5FD7A94FAB622D6" httpUri="http://treatment.plazi.org/id/024F878359677F30C5FD7A94FAB622D6" lastPageId="7" lastPageNumber="380" pageId="3" pageNumber="376">
<subSubSection id="C2FC651E59677F34C5FD7A94FDBD25A9" box="[151,636,223,250]" pageId="3" pageNumber="376" type="nomenclature">
<paragraph id="8A59369559677F34C5FD7A94FDBD25A9" blockId="3.[151,636,223,286]" box="[151,636,223,250]" pageId="3" pageNumber="376">
<heading id="D11181F959677F34C5FD7A94FDBD25A9" bold="true" box="[151,636,223,250]" fontSize="11" level="1" pageId="3" pageNumber="376" reason="1">
<emphasis id="B892EA8759677F34C5FD7A94FDBD25A9" bold="true" box="[151,636,223,250]" pageId="3" pageNumber="376">
<taxonomicName id="4DE64D1659677F34C5FD7A94FDD625AA" authority="Sudhikumar, 2021" authorityName="Sudhikumar" authorityYear="2021" box="[151,535,223,250]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="bilobocaudatus" status="sp. nov.">
<emphasis id="B892EA8759677F34C5FD7A94FDD625AA" bold="true" box="[151,535,223,250]" italics="true" pageId="3" pageNumber="376">Klimakodesmus bilobocaudatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3A157FC59677F34C7747A94FDBD25A9" box="[542,636,223,249]" pageId="3" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2FC651E59677F34C5FD7B48FEC6244E" box="[151,263,259,286]" pageId="3" pageNumber="376" type="description">
<paragraph id="8A59369559677F34C5FD7B48FEC6244E" blockId="3.[151,636,223,286]" box="[151,263,259,286]" pageId="3" pageNumber="376">
<figureCitation id="12DD2A1059677F34C5FD7B48FEC6244E" box="[151,263,259,286]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStart-2="FIGURES 8–15" captionStartId-0="4.[151,264,1877,1901]" captionStartId-1="5.[151,264,1579,1603]" captionStartId-2="6.[151,264,1898,1922]" captionTargetBox-0="[151,1436,237,1854]" captionTargetBox-1="[185,1402,181,1555]" captionTargetBox-2="[191,1396,345,1875]" captionTargetId-0="figure-55@4.[151,1436,237,1854]" captionTargetId-1="figure-17@5.[185,1402,181,1555]" captionTargetId-2="figure-109@6.[191,1396,345,1875]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." captionText-1="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." captionText-2="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.4889250" figureDoi-1="http://doi.org/10.5281/zenodo.4889252" figureDoi-2="http://doi.org/10.5281/zenodo.4889254" httpUri-0="https://zenodo.org/record/4889250/files/figure.png" httpUri-1="https://zenodo.org/record/4889252/files/figure.png" httpUri-2="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Figs 1–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C2FC651E59677F34C5FD7B00FF0824A5" pageId="3" pageNumber="376" type="materials_examined">
<paragraph id="8A59369559677F34C5FD7B00FC7524FD" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
<emphasis id="B892EA8759677F34C5FD7B00FE422435" bold="true" box="[151,387,331,357]" pageId="3" pageNumber="376">Material examined.</emphasis>
<materialsCitation id="3A8E3CC859677F34C4E77B00FE5024DA" ID-GBIF-Occurrence="3305365301" collectingDate="2021-01-27" collectionCode="ZMUM" collectorName="M. D. Aswathy" country="India" elevation="5" latitude="11.9673605" location="Kerala state" longLatPrecision="1" longitude="75.29628" pageId="3" pageNumber="376" specimenCount="11" specimenCount-adult="3" specimenCount-subadult="8" stateProvince="Kerala" typeStatus="holotype">
Adult male
<typeStatus id="555D883759677F34C77C7B00FDB92435" box="[534,632,331,357]" pageId="3" pageNumber="376" type="holotype">holotype</typeStatus>
(CATE-5302),
<specimenCount id="9CE0FD1C59677F34C6477B00FCBD2435" box="[813,892,331,357]" pageId="3" pageNumber="376" type="adult">2 adult</specimenCount>
male and
<specimenCount id="9CE0FD1C59677F34C6927B00FBAE2435" box="[1016,1135,331,357]" pageId="3" pageNumber="376" type="subadult">6 subadult</specimenCount>
female
<typeStatus id="555D883759677F34C1A57B07FAFB2436" box="[1231,1338,332,358]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
(CATE- 5302B–CATE-5302I),
</materialsCitation>
<materialsCitation id="3A8E3CC859677F34C4F27B24FC7524FD" ID-GBIF-Occurrence="3305365302" collectingDate="2021-01-27" collectionCode="ZMUM" collectorName="M. D. Aswathy" country="India" elevation="5" latitude="11.9673605" location="Kerala state" longLatPrecision="1" longitude="75.29628" pageId="3" pageNumber="376" specimenCount="11" specimenCount-adult="3" specimenCount-subadult="8" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="9CE0FD1C59677F34C4F27B24FE2424D9" box="[408,485,367,393]" pageId="3" pageNumber="376" type="adult">1 adult</specimenCount>
male and
<specimenCount id="9CE0FD1C59677F34C7337B24FD0C24D9" box="[601,717,367,393]" pageId="3" pageNumber="376" type="subadult">2 subadult</specimenCount>
female
<typeStatus id="555D883759677F34C64D7B3BFC5324DA" box="[807,914,368,394]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ECF7AE5059677F34C6C87B24FC3E24D9" box="[930,1023,367,393]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="3" pageNumber="376">ZMUM</collectionCode>
),
<collectingCountry id="F2F1760559677F34C1787B24FB8F24D9" box="[1042,1102,367,393]" name="India" pageId="3" pageNumber="376">India</collectingCountry>
,
<collectingRegion id="4822F87759677F34C1337B24FB2024DA" box="[1113,1249,367,394]" country="India" name="Kerala" pageId="3" pageNumber="376">Kerala state</collectingRegion>
, Kannur district,
<geoCoordinate id="EFD2505259677F34C5FD7BD8FEE824FE" box="[151,297,403,430]" degrees="11" direction="north" minutes="58" orientation="latitude" pageId="3" pageNumber="376" precision="1" seconds="2.5" value="11.9673605">N11°58’2.5”</geoCoordinate>
,
<geoCoordinate id="EFD2505259677F34C45E7BD8FE1324FE" box="[308,466,403,430]" degrees="75" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="376" precision="1" seconds="46.6" value="75.29628">E75°17’46.6”</geoCoordinate>
,
<elevation id="01CBD1A659677F34C4B77BD8FD8424FD" box="[477,581,403,430]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="376" unit="m" value="5.0">
<quantity id="4D1E9B7059677F34C4B77BD8FDC924FE" box="[477,520,403,430]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="376" unit="m" value="5.0">5 m</quantity>
a.s.l.
</elevation>
,
<date id="FE58105559677F34C73B7BD8FD1224FD" box="[593,723,403,429]" pageId="3" pageNumber="376" value="2021-01-27">
<collectingDate id="EE1CE9BD59677F34C73B7BD8FD1224FD" box="[593,723,403,429]" pageId="3" pageNumber="376" value="2021-01-27">27/01/2021</collectingDate>
</date>
,
<collectorName id="2713534359677F34C7B57BDFFC4224FD" box="[735,899,403,430]" pageId="3" pageNumber="376">M.D. Aswathy</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8A59369559677F34C5AD7BFCFF0824A5" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
<emphasis id="B892EA8759677F34C5AD7BFCFECC2481" bold="true" box="[199,269,439,465]" pageId="3" pageNumber="376">Name</emphasis>
. To emphasize a caudally relatively deeply bilobed dorsal keel (
<collectionCode id="ECF7AE5059677F34C6BB7BFCFC3C2481" box="[977,1021,439,465]" country="USA" name="Pratt Museum" pageId="3" pageNumber="376">PM</collectionCode>
) on ring
<quantity id="4D1E9B7059677F34C1357BFCFB562481" box="[1119,1175,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="3" pageNumber="376" unit="in" value="19.0">19 in</quantity>
the new species; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C2FC651E59677F34C5AD7BB4FD6C2645" pageId="3" pageNumber="376" type="diagnosis">
<paragraph id="8A59369559677F34C5AD7BB4FD6C2645" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
<emphasis id="B892EA8759677F34C5AD7BB4FE832749" bold="true" box="[199,322,511,537]" pageId="3" pageNumber="376">Diagnosis.</emphasis>
The new species differs clearly from
<taxonomicName id="4DE64D1659677F34C7B5784BFC9B2749" authorityName="Carl" authorityYear="1932" box="[735,858,511,537]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="B892EA8759677F34C7B5784BFC9B2749" box="[735,858,511,537]" italics="true" pageId="3" pageNumber="376">K. gravelyi</emphasis>
</taxonomicName>
, the only other congener to be accepted (
<bibRefCitation id="EE774B6459677F34C0767BB4FA512749" author="Carl, J." box="[1308,1424,511,537]" pageId="3" pageNumber="376" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl 1932</bibRefCitation>
), by the more regularly tuberculate collum (cf.
<figureCitation id="12DD2A1059677F34C7E67868FD11276E" box="[652,720,547,574]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 5</figureCitation>
and
<figureCitation id="12DD2A1059677F34C66E7868FCE3276E" box="[772,802,547,574]" captionStart="FIGURES 16–21" captionStartId="7.[151,264,1722,1746]" captionTargetBox="[151,1416,209,1678]" captionTargetId="figure-17@7.[151,1436,181,1694]" captionTargetPageId="7" captionText="FIGURES 16–21. Klimakodesmus gravelyi Carl, 1932, male holotype, after Carl (1932). 16. Collum, dorsal view. 17. Anterior part of body, lateral view. 18 &amp; 19. Posterior part of body, dorsal and lateral views, respectively. 20 &amp; 21. Left and right gonopods, caudal and oral views, respectively. Abbreviations: cx, coxite; lo, lateral lobe; sl, solenomere. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="3" pageNumber="376">16</figureCitation>
), the lower PM and DL which are mostly inclined caudad (
<emphasis id="B892EA8759677F34C5CB7803FF7E2731" box="[161,191,584,609]" italics="true" pageId="3" pageNumber="376">vs.</emphasis>
inclined forward until ring 16, cf.
<figureCitation id="12DD2A1059677F34C728780CFD4B2732" box="[578,650,583,610]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 7</figureCitation>
,
<figureCitation id="12DD2A1059677F34C7FC780CFD652731" box="[662,676,583,609]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">8</figureCitation>
and
<figureCitation id="12DD2A1059677F34C7B0780CFD392731" box="[730,760,583,609]" captionStart="FIGURES 16–21" captionStartId="7.[151,264,1722,1746]" captionTargetBox="[151,1416,209,1678]" captionTargetId="figure-17@7.[151,1436,181,1694]" captionTargetPageId="7" captionText="FIGURES 16–21. Klimakodesmus gravelyi Carl, 1932, male holotype, after Carl (1932). 16. Collum, dorsal view. 17. Anterior part of body, lateral view. 18 &amp; 19. Posterior part of body, dorsal and lateral views, respectively. 20 &amp; 21. Left and right gonopods, caudal and oral views, respectively. Abbreviations: cx, coxite; lo, lateral lobe; sl, solenomere. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="3" pageNumber="376">17</figureCitation>
), the caudally more deeply bilobed dorsal keel (PM) on ring 19 (
<emphasis id="B892EA8759677F34C5A97827FF2327D5" box="[195,226,620,645]" italics="true" pageId="3" pageNumber="376">vs.</emphasis>
indistinctly bilobulate, cf.
<figureCitation id="12DD2A1059677F34C7677820FDBD27D5" box="[525,636,619,645]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Figs 9, 10</figureCitation>
and
<figureCitation id="12DD2A1059677F34C7D87820FD1127D5" box="[690,720,619,645]" captionStart="FIGURES 16–21" captionStartId="7.[151,264,1722,1746]" captionTargetBox="[151,1416,209,1678]" captionTargetId="figure-17@7.[151,1436,181,1694]" captionTargetPageId="7" captionText="FIGURES 16–21. Klimakodesmus gravelyi Carl, 1932, male holotype, after Carl (1932). 16. Collum, dorsal view. 17. Anterior part of body, lateral view. 18 &amp; 19. Posterior part of body, dorsal and lateral views, respectively. 20 &amp; 21. Left and right gonopods, caudal and oral views, respectively. Abbreviations: cx, coxite; lo, lateral lobe; sl, solenomere. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="3" pageNumber="376">18</figureCitation>
), the laterally bi- (rings 3 and 4) or trilobate (remaining postcollum rings) paraterga (
<emphasis id="B892EA8759677F34C4FA78DBFE6627F9" box="[400,423,656,681]" italics="true" pageId="3" pageNumber="376">vs</emphasis>
. mostly bilobate, trilobate only on ring 2, cf.
<figureCitation id="12DD2A1059677F34C6DC78C4FC3E27FA" box="[950,1023,655,682]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 5</figureCitation>
,
<figureCitation id="12DD2A1059677F34C16778C4FBF827F9" box="[1037,1081,655,681]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">8, 9</figureCitation>
and
<figureCitation id="12DD2A1059677F34C11E78C4FB7D27F9" box="[1140,1212,655,681]" captionStart="FIGURES 16–21" captionStartId="7.[151,264,1722,1746]" captionTargetBox="[151,1416,209,1678]" captionTargetId="figure-17@7.[151,1436,181,1694]" captionTargetPageId="7" captionText="FIGURES 16–21. Klimakodesmus gravelyi Carl, 1932, male holotype, after Carl (1932). 16. Collum, dorsal view. 17. Anterior part of body, lateral view. 18 &amp; 19. Posterior part of body, dorsal and lateral views, respectively. 20 &amp; 21. Left and right gonopods, caudal and oral views, respectively. Abbreviations: cx, coxite; lo, lateral lobe; sl, solenomere. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="3" pageNumber="376">17–19</figureCitation>
), and the gonopods showing subspherical coxites (
<emphasis id="B892EA8759677F34C49978FFFDD3279D" box="[499,530,692,717]" italics="true" pageId="3" pageNumber="376">vs.</emphasis>
squarish) and subcontiguous, apically subunciform and crossing each other (
<emphasis id="B892EA8759677F34C01478FFFA5D279D" box="[1406,1436,692,717]" italics="true" pageId="3" pageNumber="376">vs.</emphasis>
divergent and suberect) telopodites, each of the latter additionally reinforced with a mesobasally thickened wall (
<emphasis id="B892EA8759677F34C0147893FA5D27A1" box="[1406,1436,728,753]" italics="true" pageId="3" pageNumber="376">vs.</emphasis>
a laterobasal lobe, cf.
<figureCitation id="12DD2A1059677F34C4E778B0FDCB2646" box="[397,522,763,790]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Figs 12–15</figureCitation>
and
<figureCitation id="12DD2A1059677F34C72B78B0FD5E2645" box="[577,671,763,789]" captionStart="FIGURES 16–21" captionStartId="7.[151,264,1722,1746]" captionTargetBox="[151,1416,209,1678]" captionTargetId="figure-17@7.[151,1436,181,1694]" captionTargetPageId="7" captionText="FIGURES 16–21. Klimakodesmus gravelyi Carl, 1932, male holotype, after Carl (1932). 16. Collum, dorsal view. 17. Anterior part of body, lateral view. 18 &amp; 19. Posterior part of body, dorsal and lateral views, respectively. 20 &amp; 21. Left and right gonopods, caudal and oral views, respectively. Abbreviations: cx, coxite; lo, lateral lobe; sl, solenomere. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="3" pageNumber="376">20 &amp; 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2FC651E59677F30C5AD7954FAB622D6" lastPageId="7" lastPageNumber="380" pageId="3" pageNumber="376" type="description">
<paragraph id="8A59369559677F34C5AD7954FD4D26BD" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
<emphasis id="B892EA8759677F34C5AD7954FE982669" bold="true" box="[199,345,799,825]" pageId="3" pageNumber="376">Description.</emphasis>
Length ca.
<quantity id="4D1E9B7059677F34C48A7954FD86266A" box="[480,583,799,826]" metricMagnitude="-3" metricUnit="m" metricValue="3.68" pageId="3" pageNumber="376" unit="mm" value="3.68">3.68 mm</quantity>
, width of midbody segments 0.10 and
<quantity id="4D1E9B7059677F34C6957954FBA5266A" box="[1023,1124,799,826]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="376" unit="mm" value="0.14">0.14 mm</quantity>
on pro- and metazonae, respectively (
<typeStatus id="555D883759677F34C4737908FE43260E" box="[281,386,835,862]" pageId="3" pageNumber="376" type="holotype">holotype</typeStatus>
). Length of adult male
<typeStatus id="555D883759677F34C7E5790FFD3B260E" box="[655,762,836,862]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
ca. 3.0–
<quantity id="4D1E9B7059677F34C6347908FC76260E" box="[862,951,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="376" unit="mm" value="3.8">3.8 mm</quantity>
, that of subadult female
<typeStatus id="555D883759677F34C1BE790FFAFE260E" box="[1236,1343,836,862]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
<quantity id="4D1E9B7059677F34C0227908FF0726D2" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="3" pageNumber="376" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2–3.5 mm</quantity>
. Width of adult male
<typeStatus id="555D883759677F34C4D67923FDE626D2" box="[444,551,872,898]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
ca.
<quantity id="4D1E9B7059677F34C732792CFD3F26D2" box="[600,766,871,898]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="3" pageNumber="376" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
, that of subadult female
<typeStatus id="555D883759677F34C17D7923FB4326D2" box="[1047,1154,872,898]" pageId="3" pageNumber="376" type="paratype">paratypes</typeStatus>
<quantity id="4D1E9B7059677F34C1E0792CFAF026D2" box="[1162,1329,871,898]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="3" pageNumber="376" unit="mm" value="0.45999999999999996" valueMax="0.48" valueMin="0.44">0.44–0.48 mm</quantity>
. Live colouration generally pink; antennae, legs and venter translucent, lighter than dorsum (
<figureCitation id="12DD2A1059677F34C12679C0FB5226F5" box="[1100,1171,907,933]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1877,1901]" captionTargetBox="[151,1436,237,1854]" captionTargetId="figure-55@4.[151,1436,237,1854]" captionTargetPageId="4" captionText="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="3" pageNumber="376">Figs 1</figureCitation>
,
<figureCitation id="12DD2A1059677F34C1CB79C7FB6E26F6" box="[1185,1199,908,934]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">7</figureCitation>
&amp;
<figureCitation id="12DD2A1059677F34C1BE79C0FB2526F5" box="[1236,1252,907,933]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">9</figureCitation>
). Colouration in alcohol after 2 months of preservation pale brown to pinkish brown, legs and antennae faded to translucent (
<figureCitation id="12DD2A1059677F34C00079E4FF0526BD" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStartId-0="4.[151,264,1877,1901]" captionStartId-1="5.[151,264,1579,1603]" captionTargetBox-0="[151,1436,237,1854]" captionTargetBox-1="[185,1402,181,1555]" captionTargetId-0="figure-55@4.[151,1436,237,1854]" captionTargetId-1="figure-17@5.[185,1402,181,1555]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." captionText-1="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.4889250" figureDoi-1="http://doi.org/10.5281/zenodo.4889252" httpUri-0="https://zenodo.org/record/4889250/files/figure.png" httpUri-1="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 2–6</figureCitation>
,
<figureCitation id="12DD2A1059677F34C5A57998FEE126BD" box="[207,288,979,1005]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">8 &amp; 10</figureCitation>
). No earth crust visible on body.
</paragraph>
<paragraph id="8A59369559677F34C5AD79B3FC1D2141" blockId="3.[151,1437,331,2014]" box="[199,988,1015,1042]" pageId="3" pageNumber="376">Most characters as in the generic description above, except as follows.</paragraph>
<paragraph id="8A59369559677F34C5AD7E50FEAA23ED" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
Body with 20 segments (likely in both sexes, see below) (
<figureCitation id="12DD2A1059677F34C60B7E50FC222165" box="[865,995,1051,1077]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1877,1901]" captionTargetBox="[151,1436,237,1854]" captionTargetId="figure-55@4.[151,1436,237,1854]" captionTargetPageId="4" captionText="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="3" pageNumber="376">Figs 2 &amp; 4</figureCitation>
). In width, head &lt;&lt; collum &lt;segment 2=3 &lt;4–6 &lt;7–9 &lt;10–15, body gradually tapering towards telson thereafter. Head circular, labral region moderately setose, epicranial suture rather deep, vertex microgranulate (
<figureCitation id="12DD2A1059677F34C6E87E28FC06212E" box="[898,967,1123,1150]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Fig. 6</figureCitation>
). Interantennal isthmus nearly as wide as diameter of antennal socket. Antennae short, weakly clavate,
<emphasis id="B892EA8759677F34C6277EC3FC5321F1" box="[845,914,1160,1185]" italics="true" pageId="3" pageNumber="376">in situ</emphasis>
reaching body segment/ring 3 when stretched ventrolaterally. In length, antennomere 7&lt;3&lt;1&lt;6&lt;4&lt;2&lt;&lt;5; antennomeres 5–7 each with a more or less compact apicodorsal group of bacilliform sensilla (
<figureCitation id="12DD2A1059677F34C7037E84FD2021BA" box="[617,737,1231,1258]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 6 &amp; 7</figureCitation>
). Collum flabellate, completely covering the head from above; anterior margin with 5+5 distinct, equal, moderately incised and rounded lobulations; central part domed, with 2+2 and 3+3 prominent and rounded tubercles arranged in two transverse rows (
<figureCitation id="12DD2A1059677F34C6987F5CFB982061" box="[1010,1113,1303,1329]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStartId-0="4.[151,264,1877,1901]" captionStartId-1="5.[151,264,1579,1603]" captionTargetBox-0="[151,1436,237,1854]" captionTargetBox-1="[185,1402,181,1555]" captionTargetId-0="figure-55@4.[151,1436,237,1854]" captionTargetId-1="figure-17@5.[185,1402,181,1555]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." captionText-1="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.4889250" figureDoi-1="http://doi.org/10.5281/zenodo.4889252" httpUri-0="https://zenodo.org/record/4889250/files/figure.png" httpUri-1="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 2–7</figureCitation>
). Paraterga set low (at about half of midbody height), subhorizontal to faintly declivous. Dorsum strongly arched, its outline smoothly extending onto paraterga (
<figureCitation id="12DD2A1059677F34C41C7F14FE7F2029" box="[374,446,1375,1401]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1877,1901]" captionTargetBox="[151,1436,237,1854]" captionTargetId="figure-55@4.[151,1436,237,1854]" captionTargetPageId="4" captionText="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="3" pageNumber="376">Figs 3</figureCitation>
&amp;
<figureCitation id="12DD2A1059677F34C48F7F2BFE34202A" box="[485,501,1376,1402]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">7</figureCitation>
). Prozonae, as well as deep and thin strictures between pro- and metazonae finely alveolate. Postcollum metaterga with strongly differentiated dorsal tuberculations: 2+2 longitudinal crests/rows of mostly 2+2 fused tubercles, those before ring 19 slightly inclined caudad, but PM on ring 19 abruptly modified into a mid-dorsal, strongly elongate, conspicuous and caudally clearly bilobate ridge overhanging and concealing a short epiproct (
<figureCitation id="12DD2A1059677F34C4557FA4FE652359" box="[319,420,1519,1545]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1877,1901]" captionTargetBox="[151,1436,237,1854]" captionTargetId="figure-55@4.[151,1436,237,1854]" captionTargetPageId="4" captionText="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="3" pageNumber="376">Figs 2–4</figureCitation>
,
<figureCitation id="12DD2A1059677F34C4C57FA4FE2B2359" box="[431,490,1519,1545]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">8–10</figureCitation>
). Paraterga broad, thin, rather irregularly tuberculate near bases and clearly lobulated laterally, with three rounded and moderately incised lobulations on rings 2–19, regardless of cylindrical porosteles located between the 2
<superScript id="7D939BDD59677F34C4E17C7CFE5A2315" attach="left" box="[395,411,1591,1605]" fontSize="6" pageId="3" pageNumber="376">nd</superScript>
and 3
<superScript id="7D939BDD59677F34C48B7C7CFE2E2315" attach="left" box="[481,495,1591,1605]" fontSize="6" pageId="3" pageNumber="376">rd</superScript>
lobulations on rings 5, 7, 9, 10, 12, 13, 15 and 16 (
<figureCitation id="12DD2A1059677F34C1477C7CFB632301" box="[1069,1186,1591,1617]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStart-2="FIGURES 8–15" captionStartId-0="4.[151,264,1877,1901]" captionStartId-1="5.[151,264,1579,1603]" captionStartId-2="6.[151,264,1898,1922]" captionTargetBox-0="[151,1436,237,1854]" captionTargetBox-1="[185,1402,181,1555]" captionTargetBox-2="[191,1396,345,1875]" captionTargetId-0="figure-55@4.[151,1436,237,1854]" captionTargetId-1="figure-17@5.[185,1402,181,1555]" captionTargetId-2="figure-109@6.[191,1396,345,1875]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." captionText-1="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." captionText-2="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.4889250" figureDoi-1="http://doi.org/10.5281/zenodo.4889252" figureDoi-2="http://doi.org/10.5281/zenodo.4889254" httpUri-0="https://zenodo.org/record/4889250/files/figure.png" httpUri-1="https://zenodo.org/record/4889252/files/figure.png" httpUri-2="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Figs 2–10</figureCitation>
); only paraterga 3 and 4 with last lobulation in a clearly caudolateral position (
<figureCitation id="12DD2A1059677F34C6667C10FCB52326" box="[780,884,1627,1654]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1579,1603]" captionTargetBox="[185,1402,181,1555]" captionTargetId="figure-17@5.[185,1402,181,1555]" captionTargetPageId="5" captionText="FIGURES 5–7. Klimakodesmus bilobocaudatus sp. nov. Anterior parts of body, dorsal, ventral and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="3" pageNumber="376">Figs 5–7</figureCitation>
). Epiproct (
<figureCitation id="12DD2A1059677F34C6927C10FB882325" box="[1016,1097,1627,1653]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Fig. 10</figureCitation>
) finger-shaped, tip bent down and divided into two round bulbs. Hypoproct (
<figureCitation id="12DD2A1059677F34C7F27C34FD2923C9" box="[664,744,1663,1689]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Fig. 10</figureCitation>
) semi-circular, with 1+1 strong setae borne on minute, caudal, paramedian knobs.
</paragraph>
<paragraph id="8A59369559677F34C5AD7C83FD332255" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="376">
Legs with neither adenostyles nor tarsal brushes (
<figureCitation id="12DD2A1059677F34C7827C8CFCF923B1" box="[744,824,1735,1761]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Fig. 11</figureCitation>
). In length, tarsi&gt; femora&gt; prefemora&gt; tibiae&gt; coxae&gt; postfemora; claws simple, slightly curved ventrally.
</paragraph>
<paragraph id="8A59369559677F33C5AD7D44FBB22585" blockId="3.[151,1437,331,2014]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="377" pageId="3" pageNumber="376">
Gonopods (
<figureCitation id="12DD2A1059677F34C4237D44FE09227A" box="[329,456,1807,1834]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 &amp; 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 &amp; 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="3" pageNumber="376">Figs 12–15</figureCitation>
) relatively simple, placed inside a transversely oblong-oval gonopodal aperture, the latter with thin and slightly elevated caudal and lateral margins;
<emphasis id="B892EA8759677F34C6267D7FFC50221D" box="[844,913,1844,1869]" italics="true" pageId="3" pageNumber="376">in situ</emphasis>
, both gonopods strongly exposed, held mostly parallel to one another, with only tips crossing each other. Coxites (
<emphasis id="B892EA8759677F34C6EE7D1CFC5F2221" bold="true" box="[900,926,1879,1905]" pageId="3" pageNumber="376">cx</emphasis>
) voluminous, but moderately enlarged, gonocoel thus being relatively shallow; cannulae simple. Telopodites directed caudally, each unipartite and subtriangular, long and slender, gradually attenuating distad and acuminate apically, each represented solely by a solenomere (
<emphasis id="B892EA8759677F34C0EA7DD4FA5222E9" bold="true" box="[1408,1427,1951,1977]" pageId="3" pageNumber="376">sl</emphasis>
) and supporting a seminal groove running along lateral margin to the end of a laterally directed and unciform apical part, and additionally reinforced basally through a distinctly thickened caudomesal wall of a prominent, membranous, mesal lobe (
<emphasis id="B892EA8759607F33C4097AF0FEB82585" bold="true" box="[355,377,187,213]" pageId="4" pageNumber="377">lo</emphasis>
); prefemorite as usual, relatively short, densely and strongly setose.
</paragraph>
<caption id="DE99661D59607F33C5FD7D1EFE9222E5" ID-DOI="http://doi.org/10.5281/zenodo.4889250" ID-Zenodo-Dep="4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="4" pageNumber="377" startId="4.[151,264,1877,1901]" targetBox="[151,1436,237,1854]" targetPageId="4">
<paragraph id="8A59369559607F33C5FD7D1EFE9222E5" blockId="4.[151,1436,1877,1974]" pageId="4" pageNumber="377">
<emphasis id="B892EA8759607F33C5FD7D1EFEFD223D" bold="true" box="[151,316,1877,1901]" pageId="4" pageNumber="377">FIGURES 1–4.</emphasis>
<taxonomicName id="4DE64D1659607F33C4297D1EFD46223D" authority="Sudhikumar, 2021" authorityName="Sudhikumar" authorityYear="2021" box="[323,647,1877,1901]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="bilobocaudatus" status="sp. nov.">
<emphasis id="B892EA8759607F33C4297D1EFD46223D" box="[323,647,1877,1901]" italics="true" pageId="4" pageNumber="377">Klimakodesmus bilobocaudatus</emphasis>
</taxonomicName>
<emphasis id="B892EA8759607F33C7E57D1EFD37223D" bold="true" box="[655,758,1877,1901]" pageId="4" pageNumber="377">
<taxonomicNameLabel id="A3A157FC59607F33C7E57D1EFD20223D" box="[655,737,1877,1901]" pageId="4" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
1
</emphasis>
. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale).
<emphasis id="B892EA8759607F33C4D97D32FE0122C1" bold="true" box="[435,448,1913,1937]" pageId="4" pageNumber="377">2</emphasis>
. Adult male paratype, dorsal view.
<emphasis id="B892EA8759607F33C6407D32FCF622C1" bold="true" box="[810,823,1913,1937]" pageId="4" pageNumber="377">3</emphasis>
. Subadult female paratype, lateral view.
<emphasis id="B892EA8759607F33C1BD7D32FB2522C1" bold="true" box="[1239,1252,1913,1937]" pageId="4" pageNumber="377">4</emphasis>
. Adult male paratype, ventral view.
</paragraph>
</caption>
<caption id="DE99661D59617F32C5FD7C60FF162337" ID-DOI="http://doi.org/10.5281/zenodo.4889252" ID-Zenodo-Dep="4889252" httpUri="https://zenodo.org/record/4889252/files/figure.png" pageId="5" pageNumber="378" startId="5.[151,264,1579,1603]" targetBox="[185,1402,181,1555]" targetPageId="5">
<paragraph id="8A59369559617F32C5FD7C60FF162337" blockId="5.[151,1436,1579,1639]" pageId="5" pageNumber="378">
<emphasis id="B892EA8759617F32C5FD7C60FEFF2313" bold="true" box="[151,318,1579,1603]" pageId="5" pageNumber="378">FIGURES 5–7.</emphasis>
<taxonomicName id="4DE64D1659617F32C4227C60FD4F2313" authority="Sudhikumar, 2021" authorityName="Sudhikumar" authorityYear="2021" box="[328,654,1579,1603]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="bilobocaudatus" status="sp. nov.">
<emphasis id="B892EA8759617F32C4227C60FD4F2313" box="[328,654,1579,1603]" italics="true" pageId="5" pageNumber="378">Klimakodesmus bilobocaudatus</emphasis>
</taxonomicName>
<emphasis id="B892EA8759617F32C7F27C60FD2D2313" bold="true" box="[664,748,1579,1603]" pageId="5" pageNumber="378">
<taxonomicNameLabel id="A3A157FC59617F32C7F27C60FD2D2313" box="[664,748,1579,1603]" pageId="5" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Anterior parts of body, dorsal, ventral and lateral views, respectively.
</paragraph>
</caption>
<paragraph id="8A59369559617F32C5AD7CDEFCB4224C" blockId="5.[151,1437,1685,2036]" pageId="5" pageNumber="378">
<emphasis id="B892EA8759617F32C5AD7CDEFE4F23E0" bold="true" box="[199,398,1685,1712]" pageId="5" pageNumber="378">Ecological notes.</emphasis>
This new species was found under rotting wood near a lowland swamp overgrown with
<taxonomicName id="4DE64D1659617F32C0057CDDFF1E2383" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="378" phylum="Tracheophyta" rank="genus">
<emphasis id="B892EA8759617F32C0057CDDFF1E2383" italics="true" pageId="5" pageNumber="378">Myristica</emphasis>
</taxonomicName>
forest in the
<collectingCountry id="F2F1760559617F32C4047CF1FE032384" box="[366,450,1722,1748]" name="India" pageId="5" pageNumber="378">Kannur</collectingCountry>
district, northern
<collectingRegion id="4822F87759617F32C7ED7CF2FD152383" box="[647,724,1721,1747]" country="India" name="Kerala" pageId="5" pageNumber="378">Kerala</collectingRegion>
,
<collectingCountry id="F2F1760559617F32C7B57CF2FCDD2383" box="[735,796,1721,1747]" name="India" pageId="5" pageNumber="378">India</collectingCountry>
. The habitat area of approximately 20 ha is a sacred grove protected by local people. In contrast,
<taxonomicName id="4DE64D1659617F32C72B7C95FD7F23A7" authorityName="Carl" authorityYear="1932" box="[577,702,1757,1783]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="B892EA8759617F32C72B7C95FD7F23A7" box="[577,702,1757,1783]" italics="true" pageId="5" pageNumber="378">K. gravelyi</emphasis>
</taxonomicName>
came from high-altitude areas (
<quantity id="4D1E9B7059617F32C14A7C96FB7D23A8" box="[1056,1212,1757,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="5" pageNumber="378" unit="m" value="1300.0" valueMax="1600.0" valueMin="1000.0">1000–1600 m</quantity>
a.s.l.) in the state of
<collectingRegion id="4822F87759617F32C5FD7D4AFEDD224B" box="[151,284,1793,1819]" country="India" name="Tamil Nadu" pageId="5" pageNumber="378">Tamil Nadu</collectingRegion>
(
<bibRefCitation id="EE774B6459617F32C4407D4AFE5E224C" author="Carl, J." box="[298,415,1793,1820]" pageId="5" pageNumber="378" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl 1932</bibRefCitation>
), Western Ghats, southern
<collectingCountry id="F2F1760559617F32C7A17D4AFCC4224B" box="[715,773,1793,1819]" name="India" pageId="5" pageNumber="378">India</collectingCountry>
(
<figureCitation id="12DD2A1059617F32C67E7D4AFCA9224C" box="[788,872,1793,1820]" captionStart="FIGURE 22" captionStartId="8.[151,250,1116,1140]" captionTargetBox="[151,1436,181,1092]" captionTargetId="figure-24@8.[151,1436,181,1092]" captionTargetPageId="8" captionText="FIGURE 22. Distribution of the genus Klimakodesmus: K. gravelyi in Tamil Nadu state (orange dots), and K. bilobocaudatus sp. nov. in Kerala state (red dot)." figureDoi="http://doi.org/10.5281/zenodo.4889258" httpUri="https://zenodo.org/record/4889258/files/figure.png" pageId="5" pageNumber="378">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8A59369559617F31C5AD7D6DFC4B2411" blockId="5.[151,1437,1685,2036]" lastBlockId="6.[151,1437,151,321]" lastPageId="6" lastPageNumber="379" pageId="5" pageNumber="378">
It seems noteworthy that all females collected at the
<typeStatus id="555D883759617F32C6727D6DFC892210" box="[792,840,1830,1856]" pageId="5" pageNumber="378">type</typeStatus>
locality in January appear to be subadults, each with 19 body segments. This has been proven by dissecting the vulvae which have always been found underdeveloped (amorphous ovoid clods devoid of setae), and an anterior, ventral, transverse, marginal ridge on ring 3 missing. This agrees well with the common wisdom that only adult males in
<taxonomicName id="4DE64D1659617F32C6F17DDAFBEE22FB" authorityName="Latreille" authorityYear="1802" box="[923,1071,1937,1963]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="378" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
, but never adult females, can be composed of a lesser number of body rings. Usually, in the course of teloanamorphosis characteristic of the order
<taxonomicName id="4DE64D1659617F32C5FD7D92FEEA22A3" authorityName="Latreille" authorityYear="1802" box="[151,299,2009,2035]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="378" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
, both the males and females of
<taxonomicName id="4DE64D1659617F32C7F37D92FCEB22A3" authorityName="Latreille" authorityYear="1802" box="[665,810,2009,2035]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="378" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
show the same number of body rings, either 20 or 19, more rarely 18, exceptionally up to 40 (
<bibRefCitation id="EE774B6459627F31C7317ADCFCE225E2" author="Shear, W. A. &amp; Ferreira, R. L. &amp; Iniesta, L. F. M. &amp; Marek, P." box="[603,803,151,178]" pageId="6" pageNumber="379" pagination="371 - 390" refId="ref5254" refString="Shear, W. A., Ferreira, R. L., Iniesta, L. F. M. &amp; Marek, P. (2016) A millipede missing link: Dobrodesmidae, a remarkable new polydesmidan millipede family from Brazil with supernumerary rings (Diplopoda, Polydesmida), and the establishment of a new suborder Dobrodesmidea. Zootaxa, 4178 (3), 371 - 390. https: // doi. org / 10.11646 / zootaxa. 4178.3.4" type="journal article" year="2016">
Shear
<emphasis id="B892EA8759627F31C7CF7ADCFD1F25E1" box="[677,734,151,177]" italics="true" pageId="6" pageNumber="379">et al.</emphasis>
2016
</bibRefCitation>
). When there is a difference in polydesmidan body segment counts, the adult males typically have one segment less and nearly always remain stable per sex (
<bibRefCitation id="EE774B6459627F31C0687AF0FF1525A9" author="Enghoff, H. &amp; Golovatch, S. I. &amp; Short, M. &amp; Stoev, P. &amp; Wesener, T." pageId="6" pageNumber="379" pagination="363 - 453" refId="ref4523" refString="Enghoff, H., Golovatch, S. I., Short, M., Stoev, P. &amp; Wesener, T. (2015) Diplopoda - taxonomic overview. In: Minelli, A. (Ed.) Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda. 2 (16). Brill, Leiden and Boston, pp. 363 - 453. https: // doi. org / 10.1163 / 9789004188273 _ 017" type="book chapter" year="2015">
Enghoff
<emphasis id="B892EA8759627F31C00F7AF0FA5C2585" box="[1381,1437,187,213]" italics="true" pageId="6" pageNumber="379">et al.</emphasis>
2015
</bibRefCitation>
). Only exceptionally, as is the case of
<taxonomicName id="4DE64D1659627F31C7147A94FBC125A9" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" box="[638,1024,223,249]" class="Diplopoda" family="Ammodesmidae" genus="Ammodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="granum">
<emphasis id="B892EA8759627F31C7147A94FCB725A8" box="[638,886,223,249]" italics="true" pageId="6" pageNumber="379">Ammodesmus granum</emphasis>
Cook, 1896
</taxonomicName>
and
<taxonomicName id="4DE64D1659627F31C15C7A94FDE4244E" authority="VandenSpiegel &amp; Golovatch, 2015" authorityName="VandenSpiegel &amp; Golovatch" authorityYear="2015" class="Diplopoda" family="Ammodesmidae" genus="Ammodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B892EA8759627F31C15C7A94FB1025A8" box="[1078,1233,223,249]" italics="true" pageId="6" pageNumber="379">A. congoensis</emphasis>
<bibRefCitation id="EE774B6459627F31C1B27A94FE90244D" author="VandenSpiegel, D. &amp; Golovatch, S. I." pageId="6" pageNumber="379" pagination="1 - 7" refId="ref5420" refString="VandenSpiegel, D. &amp; Golovatch, S. I. (2015) A new millipede of the family Ammodesmidae found in central Africa (Diplopoda, Polydesmida). ZooKeys, 483, 1 - 7. https: // doi. org / 10.3897 / zookeys. 483.9150" type="journal article" year="2015">VandenSpiegel &amp; Golovatch, 2015</bibRefCitation>
(Ammodesmidae)
</taxonomicName>
, from western or central Africa, respectively, can both sexes vary in the number (18 or 19) of body rings (
<bibRefCitation id="EE774B6459627F31C4D87B6CFCF72411" author="VandenSpiegel, D. &amp; Golovatch, S. I." box="[434,822,295,321]" pageId="6" pageNumber="379" pagination="1 - 17" refId="ref5373" refString="VandenSpiegel, D. &amp; Golovatch, S. I. (2012) The millipede family Ammodesmidae (Diplopoda, Polydesmida) in western Africa. ZooKeys, 221, 1 - 17. https: // doi. org / 10.3897 / zookeys. 221.3739" type="journal article" year="2012">VandenSpiegel &amp; Golovatch 2012</bibRefCitation>
, 2015).
</paragraph>
<caption id="DE99661D59627F31C5FD7D21FACA22BE" ID-DOI="http://doi.org/10.5281/zenodo.4889254" ID-Zenodo-Dep="4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="6" pageNumber="379" startId="6.[151,264,1898,1922]" targetBox="[191,1396,345,1875]" targetPageId="6">
<paragraph id="8A59369559627F31C5FD7D21FACA22BE" blockId="6.[151,1437,1898,2030]" pageId="6" pageNumber="379">
<emphasis id="B892EA8759627F31C5FD7D21FE8822D2" bold="true" box="[151,329,1898,1922]" pageId="6" pageNumber="379">FIGURES 8–15.</emphasis>
<taxonomicName id="4DE64D1659627F31C43B7D21FD5422D2" authority="Sudhikumar, 2021" authorityName="Sudhikumar" authorityYear="2021" box="[337,661,1898,1922]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="bilobocaudatus" status="sp. nov.">
<emphasis id="B892EA8759627F31C43B7D21FD5422D2" box="[337,661,1898,1922]" italics="true" pageId="6" pageNumber="379">Klimakodesmus bilobocaudatus</emphasis>
</taxonomicName>
<emphasis id="B892EA8759627F31C7F77D21FCC422D2" bold="true" box="[669,773,1898,1922]" pageId="6" pageNumber="379">
<taxonomicNameLabel id="A3A157FC59627F31C7F77D21FD3122D2" box="[669,752,1898,1922]" pageId="6" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
8
</emphasis>
. Posterior half of body (subadult female paratype), lateral view.
<emphasis id="B892EA8759627F31C5FD7DC5FF1E22F6" bold="true" box="[151,223,1934,1958]" pageId="6" pageNumber="379">9 &amp; 10</emphasis>
. Posterior part of body (adult male paratype), dorsal and ventral views, respectively.
<emphasis id="B892EA8759627F31C12B7DC5FB9822F6" bold="true" box="[1089,1113,1934,1958]" pageId="6" pageNumber="379">11</emphasis>
. Midbody leg (adult male paratype), lateral view.
<emphasis id="B892EA8759627F31C43D7DF9FE57229A" bold="true" box="[343,406,1970,1994]" pageId="6" pageNumber="379">12–14</emphasis>
. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively.
<emphasis id="B892EA8759627F31C1B07DF9FB32229A" bold="true" box="[1242,1267,1970,1994]" pageId="6" pageNumber="379">15</emphasis>
. Right gonopod, caudal view.
<emphasis id="B892EA8759627F31C4767D9DFE7422BE" bold="true" box="[284,437,2006,2030]" pageId="6" pageNumber="379">Abbreviations</emphasis>
:
<emphasis id="B892EA8759627F31C4A87D9DFE1B22BE" bold="true" box="[450,474,2006,2030]" pageId="6" pageNumber="379">cx</emphasis>
, coxite;
<emphasis id="B892EA8759627F31C7587D9DFD8722BE" bold="true" box="[562,582,2006,2030]" pageId="6" pageNumber="379">lo</emphasis>
, mesal lobe;
<emphasis id="B892EA8759627F31C7A77D9DFD1F22BE" bold="true" box="[717,734,2006,2030]" pageId="6" pageNumber="379">sl</emphasis>
, solenomere.
<emphasis id="B892EA8759627F31C6067D9DFC5922BE" bold="true" box="[876,920,2006,2030]" pageId="6" pageNumber="379">NB:</emphasis>
Figures
<emphasis id="B892EA8759627F31C69A7D9DFB8522BE" bold="true" box="[1008,1092,2006,2030]" pageId="6" pageNumber="379">14 &amp; 15</emphasis>
drawn not to scale.
</paragraph>
</caption>
<caption id="DE99661D59637F30C5FD7CF1FA8E224A" ID-DOI="http://doi.org/10.5281/zenodo.4889256" ID-Zenodo-Dep="4889256" httpUri="https://zenodo.org/record/4889256/files/figure.png" pageId="7" pageNumber="380" startId="7.[151,264,1722,1746]" targetBox="[151,1416,209,1678]" targetPageId="7">
<paragraph id="8A59369559637F30C5FD7CF1FA8E224A" blockId="7.[151,1436,1722,1818]" pageId="7" pageNumber="380">
<emphasis id="B892EA8759637F30C5FD7CF1FE952382" bold="true" box="[151,340,1722,1746]" pageId="7" pageNumber="380">FIGURES 16–21.</emphasis>
<taxonomicName id="4DE64D1659637F30C4307CF1FD062382" authority="Carl, 1932" authorityName="Carl" authorityYear="1932" box="[346,711,1722,1746]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="380" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="B892EA8759637F30C4307CF1FD922382" box="[346,595,1722,1746]" italics="true" pageId="7" pageNumber="380">Klimakodesmus gravelyi</emphasis>
<bibRefCitation id="EE774B6459637F30C7337CF1FD062382" author="Carl, J." box="[601,711,1722,1746]" pageId="7" pageNumber="380" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl, 1932</bibRefCitation>
</taxonomicName>
, male holotype, after
<bibRefCitation id="EE774B6459637F30C6C87CF1FBDA2382" author="Carl, J." box="[930,1051,1722,1746]" pageId="7" pageNumber="380" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl (1932)</bibRefCitation>
.
<emphasis id="B892EA8759637F30C14E7CF1FBFC2382" bold="true" box="[1060,1085,1722,1746]" pageId="7" pageNumber="380">16</emphasis>
. Collum, dorsal view.
<emphasis id="B892EA8759637F30C0487CF1FAFA2382" bold="true" box="[1314,1339,1722,1746]" pageId="7" pageNumber="380">17</emphasis>
. Anterior part of body, lateral view.
<emphasis id="B892EA8759637F30C4F47C95FE3023A6" bold="true" box="[414,497,1758,1782]" pageId="7" pageNumber="380">18 &amp; 19</emphasis>
. Posterior part of body, dorsal and lateral views, respectively.
<emphasis id="B892EA8759637F30C10E7C95FB7923A6" bold="true" box="[1124,1208,1758,1782]" pageId="7" pageNumber="380">20 &amp; 21</emphasis>
. Left and right gonopods, caudal and oral views, respectively.
<emphasis id="B892EA8759637F30C75F7D49FD0F224A" bold="true" box="[565,718,1794,1818]" pageId="7" pageNumber="380">Abbreviations</emphasis>
:
<emphasis id="B892EA8759637F30C7B67D49FD35224A" bold="true" box="[732,756,1794,1818]" pageId="7" pageNumber="380">cx</emphasis>
, coxite;
<emphasis id="B892EA8759637F30C6217D49FC9E224A" bold="true" box="[843,863,1794,1818]" pageId="7" pageNumber="380">lo</emphasis>
, lateral lobe;
<emphasis id="B892EA8759637F30C6807D49FC3A224A" bold="true" box="[1002,1019,1794,1818]" pageId="7" pageNumber="380">sl</emphasis>
, solenomere. Drawn not to scale.
</paragraph>
</caption>
<paragraph id="8A59369559637F30C5AD7D02FAB622D6" blockId="7.[151,1437,1864,1927]" pageId="7" pageNumber="380">
In
<taxonomicName id="4DE64D1659637F30C58C7D03FE322232" box="[230,499,1864,1890]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="380" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B892EA8759637F30C58C7D03FE562232" box="[230,407,1864,1890]" italics="true" pageId="7" pageNumber="380">Klimakodesmus</emphasis>
species
</taxonomicName>
, however, the situation is ordinary, as both sexes (must) equally have 20 body segments. The absence of adult females in the
<typeStatus id="555D883759637F30C7117D26FD6A22D7" box="[635,683,1901,1927]" pageId="7" pageNumber="380">type</typeStatus>
series seems best to be accounted for by seasonal factors alone.
</paragraph>
</subSubSection>
</treatment>
</document>
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