213 lines
28 KiB
XML
213 lines
28 KiB
XML
<document id="291EBA7BB9C7D98DAEC322A167E1B2F6" ID-CLB-Dataset="6001" ID-DOI="10.11646/zootaxa.4980.2.8" ID-GBIF-Dataset="13ae2f6e-362f-4d58-a581-d3842dacfa6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4889248" ID-ZooBank="C55A445F-76EC-460A-B259-98AF744ECFFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622538088690" checkinUser="plazi" docAuthor="Sudhikumar, Ambalaparambil Vasu" docDate="2021" docId="024F878359667F35C5FD7B24FCBA22FF" docLanguage="en" docName="zootaxa.4980.2.8.pdf" docOrigin="Zootaxa 4980 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Klimakodesmus Carl 1932" docType="treatment" docVersion="7" lastPageNumber="375" masterDocId="FE76FFFB59647F37C56A7A4BFFC12550" masterDocTitle="The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae)" masterLastPageNumber="382" masterPageNumber="373" pageNumber="375" updateTime="1698995773394" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="27C7E186A57127A6EADA7A981E94B3F0">
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<mods:title id="ED3EADA94795EDFDF38672250F368703">The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae)</mods:title>
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<mods:name id="74782DFF8F3C2E4CF00A7B9B2C753971" type="personal">
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<mods:roleTerm id="117E577AC2D5AB2A8C6403285191A7A8">Author</mods:roleTerm>
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<mods:namePart id="E72EA091CFBCB5E8B56CE0FFAC7C94A2">Sudhikumar, Ambalaparambil Vasu</mods:namePart>
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<mods:nameIdentifier id="A909046AECE8A5D4EE03496EB1FEDD68" type="ORCID">0000-0002-4479-4995</mods:nameIdentifier>
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<mods:affiliation id="F565DD931DCD5EDA081DCFAF8E3845D0">Centre for Animal Taxonomy and Ecology (CATE), Christ College, Irinjalakuda, Kerala, India & spidersudhi @ gmail. com; https: // orcid. org / 0000 - 0002 - 4479 - 4995</mods:affiliation>
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<mods:nameIdentifier id="B41CA29E7E72C02B924E09700F1373EE" type="email">spidersudhi@gmail.com</mods:nameIdentifier>
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<mods:title id="8C236B2A9CE0463FBE9396D241D3A4A6">Zootaxa</mods:title>
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<mods:part id="444CA56D73B0FFA8F21EAD54A8F36DD8">
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<mods:date id="F56858AD68BD31A99016FF7B53ABA032">2021</mods:date>
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<mods:number id="3927FF9068FC087032523A9CBF3567C0">2021-06-01</mods:number>
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<mods:number id="EFAD1C89527B7F5DEC6750F847574F04">4980</mods:number>
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<mods:detail id="C897DFDFAA602066CCDD11FFFCFEA7F0" type="issue">
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<mods:number id="B31480FD7349128A6DB63CA86246A5AE">2</mods:number>
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<mods:identifier id="0AEA54EF89A5FF9FD7A808AA2197AE52" type="DOI">10.11646/zootaxa.4980.2.8</mods:identifier>
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<mods:identifier id="57E70489C827907F4C34E9ED5BFD0028" type="GBIF-Dataset">13ae2f6e-362f-4d58-a581-d3842dacfa6e</mods:identifier>
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<treatment id="024F878359667F35C5FD7B24FCBA22FF" ID-DOI="http://doi.org/10.5281/zenodo.4986344" ID-GBIF-Taxon="182958731" ID-Zenodo-Dep="4986344" LSID="urn:lsid:plazi:treatment:024F878359667F35C5FD7B24FCBA22FF" httpUri="http://treatment.plazi.org/id/024F878359667F35C5FD7B24FCBA22FF" lastPageNumber="375" pageId="2" pageNumber="375">
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<subSubSection id="C2FC651E59667F35C5FD7B24FDF824D9" box="[151,569,367,394]" pageId="2" pageNumber="375" type="nomenclature">
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<paragraph id="8A59369559667F35C5FD7B24FDF824D9" blockId="2.[151,569,367,394]" box="[151,569,367,394]" pageId="2" pageNumber="375">
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<heading id="D11181F959667F35C5FD7B24FDF824D9" bold="true" box="[151,569,367,394]" fontSize="11" level="1" pageId="2" pageNumber="375" reason="1">
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<emphasis id="B892EA8759667F35C5FD7B24FDF824D9" bold="true" box="[151,569,367,394]" pageId="2" pageNumber="375">
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Genus
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<taxonomicName id="4DE64D1659667F35C5847B24FDF824D9" ID-CoL="7PM5S" authority="Carl, 1932" authorityName="Carl" authorityYear="1932" box="[238,569,367,394]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
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<emphasis id="B892EA8759667F35C5847B24FE6F24DA" bold="true" box="[238,430,367,394]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
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<bibRefCitation id="EE774B6459667F35C4DC7B24FDF824D9" author="Carl, J." box="[438,569,367,393]" pageId="2" pageNumber="375" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl, 1932</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C2FC651E59667F35C5FD7BFFFCED249C" box="[151,812,436,460]" pageId="2" pageNumber="375" type="type_taxon">
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<paragraph id="8A59369559667F35C5FD7BFFFCED249C" blockId="2.[151,812,436,460]" box="[151,812,436,460]" pageId="2" pageNumber="375">
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Type-species:
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<taxonomicName id="4DE64D1659667F35C4427BFFFD57249C" ID-CoL="3R9LZ" authority="Carl, 1932" authorityName="Carl" authorityYear="1932" box="[296,662,436,460]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="gravelyi">
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<emphasis id="B892EA8759667F35C4427BFFFDE3249C" box="[296,546,436,460]" italics="true" pageId="2" pageNumber="375">Klimakodesmus gravelyi</emphasis>
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<bibRefCitation id="EE774B6459667F35C7437BFFFD57249C" author="Carl, J." box="[553,662,436,460]" pageId="2" pageNumber="375" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl, 1932</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C2FC651E59667F35C5FD7BBDFD7F201A" pageId="2" pageNumber="375" type="description">
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<paragraph id="8A59369559667F35C5FD7BBDFCE4275E" blockId="2.[151,805,502,526]" box="[151,805,502,526]" pageId="2" pageNumber="375">
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Other species included:
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<taxonomicName id="4DE64D1659667F35C4E17BBDFD0F275E" authority="Sudhikumar, 2021" authorityName="Sudhikumar" authorityYear="2021" box="[395,718,502,526]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bilobocaudatus" status="sp. nov.">
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<emphasis id="B892EA8759667F35C4E17BBDFD0F275E" box="[395,718,502,526]" italics="true" pageId="2" pageNumber="375">Klimakodesmus bilobocaudatus</emphasis>
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</taxonomicName>
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<emphasis id="B892EA8759667F35C7BF7BBDFCE4275E" bold="true" box="[725,805,502,526]" pageId="2" pageNumber="375">
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<taxonomicNameLabel id="A3A157FC59667F35C7BF7BBDFCE4275E" box="[725,805,502,526]" pageId="2" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8A59369559667F35C5FD7877FBFD21C7" blockId="2.[151,1437,572,1967]" pageId="2" pageNumber="375">
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<emphasis id="B892EA8759667F35C5FD7877FEA62706" bold="true" box="[151,359,572,598]" pageId="2" pageNumber="375">Brief description.</emphasis>
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This genus shows a body shape and an ornamentation pattern typical of
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<taxonomicName id="4DE64D1659667F35C1FA7877FAFE2706" authorityName="Silvestri" authorityYear="1896" box="[1168,1343,572,598]" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
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, characterized by 20 segments/rings (19+T) both in the male and female; a flabellate collum is strongly domed and tuberculate, completely covering the head from above, with 5+5 distinct, equal, clearly incised and rounded lobulations at a subhorizontal anterior margin; the antennae are short, two basal antennomeres of each antenna being sunken inside a distinct transverse groove, and antennomere 5 being much longer and thicker than the 6
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<superScript id="7D939BDD59667F35C1087887FBAE278A" attach="none" box="[1122,1135,716,730]" fontSize="6" pageId="2" pageNumber="375">th</superScript>
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; a cerotegument crust with microvilli covers most of the dorsal surface of the collum and following metaterga, the limbus being microlobulate and microspiculate caudally; body rings are strongly arched dorsally, each with 2+2 longitudinal rows of basically 2+2 or 3+3 larger and fused tubercles/lobes (paramedian, PM, and dorsolateral, DL), both rows largely representing higher bilobed keels mostly slightly inclined either forward or caudad, but PM always being higher than DL. PM of only ring 19 is abruptly and unusually strongly elongate and more or less clearly bilobate caudally, conspicuously overhanging and concealing a short epiproct from above (
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<figureCitation id="12DD2A1059667F35C67F79EFFCBB26EF" box="[789,890,932,959]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1877,1901]" captionTargetBox="[151,1436,237,1854]" captionTargetId="figure-55@4.[151,1436,237,1854]" captionTargetPageId="4" captionText="FIGURES 1–4. Klimakodesmus bilobocaudatus sp. nov. 1. Natural microhabitat (rotting wood with a colony on the surface, photographed not to scale). 2. Adult male paratype, dorsal view. 3. Subadult female paratype, lateral view. 4. Adult male paratype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4889250" httpUri="https://zenodo.org/record/4889250/files/figure.png" pageId="2" pageNumber="375">Figs 2–4</figureCitation>
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,
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<figureCitation id="12DD2A1059667F35C6EE79EEFC7E26EF" box="[900,959,933,959]" captionStart="FIGURES 8–15" captionStartId="6.[151,264,1898,1922]" captionTargetBox="[191,1396,345,1875]" captionTargetId="figure-109@6.[191,1396,345,1875]" captionTargetPageId="6" captionText="FIGURES 8–15. Klimakodesmus bilobocaudatus sp. nov. 8. Posterior half of body (subadult female paratype), lateral view. 9 & 10. Posterior part of body (adult male paratype), dorsal and ventral views, respectively. 11. Midbody leg (adult male paratype), lateral view. 12–14. Both gonopods (adult male paratype), caudal, oral and caudal views, respectively. 15. Right gonopod, caudal view. Abbreviations: cx, coxite; lo, mesal lobe; sl, solenomere. NB: Figures 14 & 15 drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.4889254" httpUri="https://zenodo.org/record/4889254/files/figure.png" pageId="2" pageNumber="375">8–10</figureCitation>
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). Neither intercalary (i) series of grains nor antero- (Am), nor truly caudomarginal lobulations (Cm) are distinguished. The pore formula is normal: ozopores 5, 7, 9, 10, 12, 13, 15 and 16 are borne on distinct porosteles,
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<emphasis id="B892EA8759667F35C65E79A6FC922156" box="[820,851,1005,1030]" italics="true" pageId="2" pageNumber="375">vs.</emphasis>
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17, 18 and 19 which open flush on the surface. The paraterga are set low (at about half the height of midbody metazonae), slightly declivous to subhorizontal, leaving the dorsum very convex, moderately bi- or trilobate laterally regardless of a cylindrical porostele or ozopore located just before or upon the ultimate lobulation, respectively. The legs are short and stout, rather sparsely setose, the prefemur bearing a particularly strong seta distomesally, the tibia a distodorsal one.
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</paragraph>
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<paragraph id="8A59369559667F35C5AD7EEBFD7F201A" blockId="2.[151,1437,572,1967]" pageId="2" pageNumber="375">
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The gonopods are fairly simple,
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<emphasis id="B892EA8759667F35C7527EEAFDBF21EA" box="[568,638,1185,1210]" italics="true" pageId="2" pageNumber="375">in situ</emphasis>
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either slightly diverging or distally crossing each other; each coxite is moderately enlarged, hemispherical or squarish, microgranulate and microsetose laterally, and concave medially, the gonocoel thus being moderately deep; the cannula is as usual, simple, moderately long and strongly unciform. The telopodite is simple, slender, unipartite, represented by a very strong, moderately long, but mostly exposed, gradually attenuating and acuminate solenomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="C2FC651E59667F35C5AD7F1FFC65224F" pageId="2" pageNumber="375" type="diagnosis">
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<paragraph id="8A59369559667F35C5AD7F1FFC292316" blockId="2.[151,1437,572,1967]" pageId="2" pageNumber="375">
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<emphasis id="B892EA8759667F35C5AD7F1FFEFB203E" bold="true" box="[199,314,1364,1390]" pageId="2" pageNumber="375">Diagnosis</emphasis>
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.
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<taxonomicName id="4DE64D1659667F35C4267F1FFE3C203E" authorityName="Carl" authorityYear="1932" box="[332,509,1364,1390]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
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<emphasis id="B892EA8759667F35C4267F1FFE3C203E" box="[332,509,1364,1390]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
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</taxonomicName>
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seems to be particularly similar to
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<taxonomicName id="4DE64D1659667F35C6F57F1FFBF4203E" authorityName="Pocock" authorityYear="1892" box="[927,1077,1364,1390]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
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<emphasis id="B892EA8759667F35C6F57F1FFBF4203E" box="[927,1077,1364,1390]" italics="true" pageId="2" pageNumber="375">Pyrgodesmus</emphasis>
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</taxonomicName>
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in most somatic characters, as correctly noted by
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<bibRefCitation id="EE774B6459667F35C4007F33FE2F20C3" author="Carl, J." box="[362,494,1400,1427]" pageId="2" pageNumber="375" pagination="411 - 529" refId="ref4416" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl (1932)</bibRefCitation>
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, but both genera are distinct primarily in gonopodal structure: clearly hypertrophied gonocoxites and “blade-like” telopodites (
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<bibRefCitation id="EE774B6459667F35C7067FD7FCC120E7" author="Pocock, R. I." box="[620,768,1436,1463]" pageId="2" pageNumber="375" pagination="131 - 174" refId="ref5168" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural His- tory Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock 1892</bibRefCitation>
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)
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<emphasis id="B892EA8759667F35C6657FD6FCEF20E6" box="[783,814,1437,1462]" italics="true" pageId="2" pageNumber="375">vs.</emphasis>
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moderately incrassate, apparently smaller, subspherical or squarish gonocoxites, each of which supports a strong and unipartite telopodite represented solely by a slender, strongly exposed, increasingly attenuating and acuminate solenomere. In addition, PM crests, including the one on the penultimate ring that overhangs and fully conceals the epiproct from above, are much higher and stronger, whereas DL reduced, in
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<taxonomicName id="4DE64D1659667F35C4C37C67FDFE2316" authorityName="Pocock" authorityYear="1892" box="[425,575,1580,1606]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
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<emphasis id="B892EA8759667F35C4C37C67FDFE2316" box="[425,575,1580,1606]" italics="true" pageId="2" pageNumber="375">Pyrgodesmus</emphasis>
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</taxonomicName>
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compared to
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<taxonomicName id="4DE64D1659667F35C7B27C67FC292316" box="[728,1000,1580,1606]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="undefined-a">
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<emphasis id="B892EA8759667F35C7B27C67FC482316" box="[728,905,1580,1606]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
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species.
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</taxonomicName>
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</paragraph>
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<paragraph id="8A59369559667F35C5AD7C1AFC65224F" blockId="2.[151,1437,572,1967]" pageId="2" pageNumber="375">
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A similar, remarkably strong, massive and largely exposed solenomere taking up most of the gonopod telopodite is also observed in the genus
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<taxonomicName id="4DE64D1659667F35C4B67C3FFC4E23DF" authority="Miyosi, 1957" authorityName="Miyosi" authorityYear="1957" box="[476,911,1652,1679]" class="Diplopoda" family="Pyrgodesmidae" genus="Pseudocatapyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
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<emphasis id="B892EA8759667F35C4B67C3FFD3023DE" box="[476,753,1652,1678]" italics="true" pageId="2" pageNumber="375">Pseudocatapyrgodesmus</emphasis>
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<bibRefCitation id="EE774B6459667F35C79D7C3FFC4E23DF" author="Miyosi, Y." box="[759,911,1652,1679]" pageId="2" pageNumber="375" pagination="264 - 268" refId="ref5112" refString="Miyosi, Y. (1957) Beitrage zur Kenntnis japanischer Myriopoden. 20. Aufsatz: Uber eine neue Gattung von Diplopoda, eine neue Art und eine neue Unterart von Chilopoda. Zoological Magazine, Tokyo, 66 (6), 264 - 268. [in Japanese, with English abstract]" type="journal article" year="1957">Miyosi, 1957</bibRefCitation>
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</taxonomicName>
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, with
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<taxonomicName id="4DE64D1659667F35C6B87C3EFB2323DF" authority="Miyosi, 1957" authorityName="Miyosi" authorityYear="1957" box="[978,1250,1652,1679]" class="Diplopoda" family="Pyrgodesmidae" genus="Pseudocatapyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="glaucus">
|
||
<emphasis id="B892EA8759667F35C6B87C3EFB8523DE" box="[978,1092,1652,1678]" italics="true" pageId="2" pageNumber="375">P. glaucus</emphasis>
|
||
<bibRefCitation id="EE774B6459667F35C1207C3FFB2323DF" author="Miyosi, Y." box="[1098,1250,1652,1679]" pageId="2" pageNumber="375" pagination="264 - 268" refId="ref5112" refString="Miyosi, Y. (1957) Beitrage zur Kenntnis japanischer Myriopoden. 20. Aufsatz: Uber eine neue Gattung von Diplopoda, eine neue Art und eine neue Unterart von Chilopoda. Zoological Magazine, Tokyo, 66 (6), 264 - 268. [in Japanese, with English abstract]" type="journal article" year="1957">Miyosi, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="555D883759667F35C07F7C3EFA8423DF" box="[1301,1349,1653,1679]" pageId="2" pageNumber="375">type</typeStatus>
|
||
species from Honshu,
|
||
<collectingCountry id="F2F1760559667F35C4507CD2FEBB23E3" box="[314,378,1689,1715]" name="Japan" pageId="2" pageNumber="375">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EE774B6459667F35C4E37CD3FDDC23E3" author="Miyosi, Y." box="[393,541,1688,1715]" pageId="2" pageNumber="375" pagination="264 - 268" refId="ref5112" refString="Miyosi, Y. (1957) Beitrage zur Kenntnis japanischer Myriopoden. 20. Aufsatz: Uber eine neue Gattung von Diplopoda, eine neue Art und eine neue Unterart von Chilopoda. Zoological Magazine, Tokyo, 66 (6), 264 - 268. [in Japanese, with English abstract]" type="journal article" year="1957">Miyosi 1957</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="4DE64D1659667F35C70A7CD2FA9A23E3" authority="Golovatch, Semenyuk, VandenSpiegel & Anichkin. 2011" authorityName="Golovatch, Semenyuk, VandenSpiegel & Anichkin." authorityYear="2011" box="[608,1371,1688,1715]" class="Diplopoda" family="Pyrgodesmidae" genus="Pseudocatapyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="B892EA8759667F35C70A7CD2FD1323E2" box="[608,722,1688,1714]" italics="true" pageId="2" pageNumber="375">P. pulcher</emphasis>
|
||
Golovatch, Semenyuk, VandenSpiegel & Anichkin. 2011
|
||
</taxonomicName>
|
||
, from southern
|
||
<collectingCountry id="F2F1760559667F35C5977CF7FE9D2386" box="[253,348,1724,1750]" name="Vietnam" pageId="2" pageNumber="375">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EE774B6459667F35C4037CF7FD9F2387" author="Golovatch, S. I. & Semenyuk, I. I. & VandenSpiegel, D. & Anichkin, A. E." box="[361,606,1724,1751]" pageId="2" pageNumber="375" pagination="1 - 9" refId="ref4848" refString="Golovatch, S. I., Semenyuk, I. I., VandenSpiegel, D. & Anichkin, A. E. (2011) Three new species of the millipede family Pyrgodesmidae from Nam Cat Tien National Park, southern Vietnam (Diplopoda: Polydesmida). Arthropoda Selecta, 20 (1), 1 - 9. [https: // kmkjournals. com / upload / PDF / ArthropodaSelecta / 20 / 20 _ 1 % 20001 _ 009 % 20 Golovatch. pdf] https: // doi. org / 10.15298 / arthsel. 20.1.01" type="journal article" year="2011">
|
||
Golovatch
|
||
<emphasis id="B892EA8759667F35C48F7CF6FDDC2386" box="[485,541,1724,1750]" italics="true" pageId="2" pageNumber="375">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). Yet their gonopods are considerably more elaborate and clearly branched,
|
||
<emphasis id="B892EA8759667F35C5FD7CAAFF7723AA" box="[151,182,1761,1786]" italics="true" pageId="2" pageNumber="375">vs.</emphasis>
|
||
relatively very simple and unipartite in
|
||
<taxonomicName id="4DE64D1659667F35C7E97CABFCF523AA" authorityName="Carl" authorityYear="1932" box="[643,820,1760,1786]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B892EA8759667F35C7E97CABFCF523AA" box="[643,820,1760,1786]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
|
||
</taxonomicName>
|
||
. They are the gonopods of
|
||
<taxonomicName id="4DE64D1659667F35C1187CAAFB2723AA" box="[1138,1254,1760,1786]" class="Diplopoda" family="Pyrgodesmidae" genus="Pseudocatapyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="B892EA8759667F35C1187CAAFB2723AA" box="[1138,1254,1760,1786]" italics="true" pageId="2" pageNumber="375">P. pulcher</emphasis>
|
||
</taxonomicName>
|
||
that seem to be especially similar to those of
|
||
<taxonomicName id="4DE64D1659667F35C48A7D4FFC61224F" authority="(Golovatch et al. 2011)" baseAuthorityName="Golovatch" baseAuthorityYear="2011" box="[480,928,1796,1823]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B892EA8759667F35C48A7D4FFD50224E" box="[480,657,1796,1822]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
|
||
(
|
||
<bibRefCitation id="EE774B6459667F35C7CA7D4FFC59224F" author="Golovatch, S. I. & Semenyuk, I. I. & VandenSpiegel, D. & Anichkin, A. E." box="[672,920,1796,1823]" pageId="2" pageNumber="375" pagination="1 - 9" refId="ref4848" refString="Golovatch, S. I., Semenyuk, I. I., VandenSpiegel, D. & Anichkin, A. E. (2011) Three new species of the millipede family Pyrgodesmidae from Nam Cat Tien National Park, southern Vietnam (Diplopoda: Polydesmida). Arthropoda Selecta, 20 (1), 1 - 9. [https: // kmkjournals. com / upload / PDF / ArthropodaSelecta / 20 / 20 _ 1 % 20001 _ 009 % 20 Golovatch. pdf] https: // doi. org / 10.15298 / arthsel. 20.1.01" type="journal article" year="2011">
|
||
Golovatch
|
||
<emphasis id="B892EA8759667F35C6777D4EFC97224E" box="[797,854,1796,1822]" italics="true" pageId="2" pageNumber="375">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2FC651E59667F35C5AD7D63FCBA22FF" pageId="2" pageNumber="375" type="discussion">
|
||
<paragraph id="8A59369559667F35C5AD7D63FCBA22FF" blockId="2.[151,1437,572,1967]" pageId="2" pageNumber="375">
|
||
The strong, flattened, dorsal crest PM on the penultimate ring that overhangs and conceals the epiproct from above, however conspicuous, is unique to neither
|
||
<taxonomicName id="4DE64D1659667F35C7A07D07FCBA2236" box="[714,891,1868,1894]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B892EA8759667F35C7A07D07FCBA2236" box="[714,891,1868,1894]" italics="true" pageId="2" pageNumber="375">Klimakodesmus</emphasis>
|
||
</taxonomicName>
|
||
nor
|
||
<taxonomicName id="4DE64D1659667F35C6DA7D07FB672236" box="[944,1190,1868,1894]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B892EA8759667F35C6DA7D07FB872236" box="[944,1094,1868,1894]" italics="true" pageId="2" pageNumber="375">Pyrgodesmus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
For instance, several, but not all of the Afrotropical species of the genera
|
||
<taxonomicName id="4DE64D1659667F35C7A17D3BFBEE22DB" authority="Silvestri, 1927" authorityName="Silvestri" authorityYear="1927" box="[715,1071,1904,1931]" class="Diplopoda" family="Pyrgodesmidae" genus="Monachodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B892EA8759667F35C7A17D3BFC4622DA" box="[715,903,1904,1930]" italics="true" pageId="2" pageNumber="375">Monachodesmus</emphasis>
|
||
Silvestri, 1927
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4DE64D1659667F35C1387D3BFA9A22DA" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" box="[1106,1371,1904,1930]" class="Diplopoda" family="Pyrgodesmidae" genus="Udodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B892EA8759667F35C1387D3BFB1422DA" box="[1106,1237,1904,1930]" italics="true" pageId="2" pageNumber="375">Udodesmus</emphasis>
|
||
Cook, 1896
|
||
</taxonomicName>
|
||
show the same or very similar conditions (
|
||
<bibRefCitation id="EE774B6459667F35C75A7DDFFCE622FF" author="Golovatch, S. I. & Nzoko Fiemapong, A. R. & VandenSpiegel, D." box="[560,807,1940,1967]" pageId="2" pageNumber="375" pagination="387 - 400" refId="ref4660" refString="Golovatch, S. I., Nzoko Fiemapong, A. R. & VandenSpiegel, D. (2015) Notes on Afrotropical Pyrgodesmidae, 2 (Diplopoda: Polydesmida). Arthropoda Selecta, 24 (4), 387 - 400. [https: // kmkjournals. com / upload / PDF / ArthropodaSelecta / 24 / 24 _ 4 _ 387 _ 400 _ Golovatch _ et _ al _ for _ Inet. pdf] https: // doi. org / 10.15298 / arthsel. 24.4.02" type="journal article" year="2015">
|
||
Golovatch
|
||
<emphasis id="B892EA8759667F35C7C77DDEFD2722FE" box="[685,742,1940,1966]" italics="true" pageId="2" pageNumber="375">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EE774B6459667F35C6587DDEFCAE22FF" author="Golovatch, S. I. & Nzoko Fiemapong, A. R. & VandenSpiegel, D." box="[818,879,1941,1967]" pageId="2" pageNumber="375" pagination="175 - 215" refId="ref4754" refString="Golovatch, S. I., Nzoko Fiemapong, A. R. & VandenSpiegel, D. (2017) Notes on Afrotropical Pyrgodesmidae, 3 (Diplopoda: Polydesmida). Arthropoda Selecta, 26 (3), 175 - 215. [https: // kmkjournals. com / upload / PDF / ArthropodaSelecta / 26 / 26 _ 3 _ 175 _ 215 _ Golovatch _ et _ al _ for _ Inet. pdf] https: // doi. org / 10.15298 / arthsel. 26.3.01" type="journal article" year="2017">2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |