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<mods:title id="D875089281902ACA4722E08DA35A528C">Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)</mods:title>
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<mods:namePart id="AE3C7A93A7E7081B466788A434F91C7C">A, Contreras-Félix Gerardo</mods:namePart>
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<emphasis id="B8FAEAFAFF841963FF163D32FEF9FEDA" bold="true" box="[151,350,295,321]" italics="true" pageId="32" pageNumber="33">Vaejovis darwini</emphasis>
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<figureCitation id="12B52A6DFF841963FF163D5FFEA5FEF8" box="[151,258,330,355]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="32" pageNumber="33">Figures 4</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FE8F3D5FFE9DFEF8" box="[270,314,330,355]" captionStart="FIGURE 13" captionStartId="22.[151,250,989,1011]" captionTargetBox="[151,1435,193,968]" captionTargetId="figure@22.[151,1435,193,968]" captionTargetPageId="22" captionText="FIGURE 13. Comparison in shapes on carapace in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659119/files/figure.png" pageId="32" pageNumber="33">13d</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FEC73D5EFED7FEF8" box="[326,368,331,355]" captionStart="FIGURE 14" captionStartId="23.[151,250,1558,1580]" captionTargetBox="[340,1236,345,1537]" captionTargetId="figure@23.[340,1247,337,1537]" captionTargetPageId="23" captionText="FIGURE 14. Comparison of the different shapes in the caudal gland on species within the “mexicanus” group (each caudal gland, if present, is highlighted in red): a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov." httpUri="https://zenodo.org/record/2659121/files/figure.png" pageId="32" pageNumber="33">14c</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FEFD3D5FFE0EFEF8" box="[380,425,330,355]" captionStart="FIGURE 15" captionStartId="24.[151,250,1398,1420]" captionTargetBox="[340,1247,195,1374]" captionTargetId="figure@24.[340,1247,195,1374]" captionTargetPageId="24" captionText="FIGURE 15. Comparison in shapes and trichobothrial patterns in the pedipalp femur, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659123/files/figure.png" pageId="32" pageNumber="33">15d</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FE353D5FFE46FEF8" box="[436,481,330,355]" captionStart="FIGURE 16" captionStartId="25.[151,250,1480,1502]" captionTargetBox="[340,1247,258,1458]" captionTargetId="figure@25.[340,1247,258,1458]" captionTargetPageId="25" captionText="FIGURE 16. Comparison in shapes and trichobothrial patterns in the pedipalp patella, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659125/files/figure.png" pageId="32" pageNumber="33">16d</figureCitation>
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<figureCitation id="12B52A6DFF841963FE6D3D5FFDBEFEF8" box="[492,537,330,355]" captionStart="FIGURE 17" captionStartId="26.[151,250,1585,1607]" captionTargetBox="[335,1252,335,1567]" captionTargetId="figure@26.[335,1252,335,1567]" captionTargetPageId="26" captionText="FIGURE 17. Comparison in shapes and trichobothrial patterns in the pedipalp chela of males in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659127/files/figure.png" pageId="32" pageNumber="33">17d</figureCitation>
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<figureCitation id="12B52A6DFF841963FDA53D5FFDF6FEF8" box="[548,593,330,355]" captionStart="FIGURE 18" captionStartId="27.[151,250,1747,1769]" captionTargetBox="[340,1247,589,1726]" captionTargetId="figure@27.[340,1247,589,1726]" captionTargetPageId="27" captionText="FIGURE 18. Comparison in the different shapes and trichobothrial patterns in the pedipalp chela of females, that selected species within “mexicanus” group present: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659129/files/figure.png" pageId="32" pageNumber="33">18d</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FDDD3D5FFD2EFEF8" box="[604,649,330,355]" captionStart="FIGURE 19" captionStartId="29.[151,250,1805,1827]" captionTargetBox="[319,1255,877,1784]" captionTargetId="figure@29.[319,1267,877,1784]" captionTargetPageId="29" captionText="FIGURE 19. Comparison on the shapes in the hemispermatophore in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659131/files/figure.png" pageId="32" pageNumber="33">19d</figureCitation>
, 
<figureCitation id="12B52A6DFF841963FD153D5EFD17FEF8" box="[660,688,331,355]" captionStart="FIGURE 22" captionStartId="32.[151,250,1619,1641]" captionTargetBox="[337,1237,690,1597]" captionTargetId="figure@32.[337,1250,690,1597]" captionTargetPageId="32" captionText="FIGURE 22. Habtus of Vaejovis darwini holotype (♂): a—dorsal view; b—ventral view. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2659137/files/figure.png" pageId="32" pageNumber="33">22</figureCitation>
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<emphasis id="B8FAEAFAFF841963FF163D86FEE6FE33" box="[151,321,402,424]" italics="true" pageId="32" pageNumber="33">Vaejovis darwini</emphasis>
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: 556
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–559, 567–570, Fig: 37–43, 54.
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<emphasis id="B8FAEAFAFF841963FF163DC0FEE2FE75" bold="true" box="[151,325,469,494]" pageId="32" pageNumber="33">Type material.</emphasis>
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: 
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<collectorName id="277B533EFF841963FBC93DECFB40FD89" box="[1096,1255,505,530]" pageId="32" pageNumber="33">C. Santibáñez</collectorName>
and 
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. 
<collectorName id="277B533EFF841963FEC73E08FE67FDAD" box="[326,448,541,566]" pageId="32" pageNumber="33">Examined.</collectorName>
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</paragraph>
<paragraph id="8A3136E8FF841963FF463E57FE21FDE5" blockId="32.[151,1437,469,638]" pageId="32" pageNumber="33">
<materialsCitation id="3AE63CB5FF841963FF463E57FBAEFDC1" ID-GBIF-Occurrence="2237797874" box="[199,1033,577,602]" collectingDate="2006-06-27" collectionCode="CNAN-T" collectorName="Cols. &amp; O. Francke &amp; G. Villegas &amp; H. Montano &amp; C. Santibanez &amp; A. Valdez. &amp; Examined." country="Mexico" elevation="845" latitude="16.35" location="San Pedro Juchatengo" longLatPrecision="5359" longitude="-97.1" municipality="Distrito de Juquila" pageId="32" pageNumber="33" specimenCode="CNAN-T0389" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
<typeStatus id="5535884AFF841963FF463E57FE9EFDC1" box="[199,313,578,602]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9C88FD61FF841963FEC33E54FECCFDC2" box="[322,363,577,601]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
and 
<specimenCount id="9C88FD61FF841963FE213E54FE6DFDC2" box="[416,458,577,601]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
, same data as holotype (
<specimenCode id="DA289E93FF841963FD583E54FC25FDC2" box="[729,898,577,601]" collectionCode="CNAN-T" pageId="32" pageNumber="33">CNAN-T0389</specimenCode>
); Examined
</materialsCitation>
. 
<materialsCitation id="3AE63CB5FF841963FB923E54FE25FDE5" ID-GBIF-Occurrence="2237797795" collectingDate="2006-06-27" collectionCode="AMNH" collectorName="Cols. &amp; O. Francke &amp; G. Villegas &amp; H. Montano &amp; C. Santibanez &amp; A. Valdez. &amp; Examined." country="Mexico" elevation="845" latitude="16.35" location="San Pedro Juchatengo" longLatPrecision="5359" longitude="-97.1" municipality="Distrito de Juquila" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
<specimenCount id="9C88FD61FF841963FB923E54FB9BFDC2" box="[1043,1084,577,601]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
and 
<specimenCount id="9C88FD61FF841963FBF03E54FB3DFDC2" box="[1137,1178,577,601]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="EC9FAE2DFF841963FF1E3E73FF5CFDE5" box="[159,251,614,638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="32" pageNumber="33">AMNH</collectionCode>
). Examined
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DEF16660FF841963FF163A46FB7EF9F2" box="[151,1241,1619,1641]" httpUri="https://zenodo.org/record/2659137/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1619,1641]" targetBox="[337,1237,690,1597]" targetPageId="32">
<paragraph id="8A3136E8FF841963FF163A46FB7EF9F2" blockId="32.[151,1241,1619,1641]" box="[151,1241,1619,1641]" pageId="32" pageNumber="33">
<emphasis id="B8FAEAFAFF841963FF163A46FE87F9F2" bold="true" box="[151,288,1619,1641]" pageId="32" pageNumber="33">FIGURE 22.</emphasis>
Habtus of 
<taxonomicName id="4D8E4D6BFF841963FE0E3A41FD9EF9F2" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[399,569,1619,1641]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="darwini">
<emphasis id="B8FAEAFAFF841963FE0E3A41FD9EF9F2" box="[399,569,1619,1641]" italics="true" pageId="32" pageNumber="33">Vaejovis darwini</emphasis>
</taxonomicName>
holotype (♂): 
<emphasis id="B8FAEAFAFF841963FD4E3A46FD52F9F2" bold="true" box="[719,757,1619,1641]" pageId="32" pageNumber="33">a—</emphasis>
dorsal view; 
<emphasis id="B8FAEAFAFF841963FCF63A46FC39F9F2" bold="true" box="[887,926,1619,1641]" pageId="32" pageNumber="33">b—</emphasis>
ventral view. Scale bar = 1 cm.
</paragraph>
</caption>
<subSubSection id="C2946563FF841962FF463A85FB5DFD83" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8A3136E8FF841962FF463A85FE61FE36" blockId="32.[151,1437,1680,2029]" lastBlockId="33.[151,1436,151,645]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B8FAEAFAFF841963FF463A85FE9DF932" bold="true" box="[199,314,1680,1705]" pageId="32" pageNumber="33">Diagnosis</emphasis>
(modified from 
<bibRefCitation id="EE1F4B19FF841963FE773A85FC2FF933" author="Santibanez-Lopez, C. E. &amp; Francke, O. F." box="[502,904,1680,1705]" pageId="32" pageNumber="33" pagination="555 - 571" refId="ref65257" refString="Santibanez-Lopez, C. E. &amp; Francke, O. F. (2010) New and poorly known species of the mexicanus group of the genus Vaejovis (Scorpiones: Vaejovidae) from Oaxaca, Mexico. Journal of Arachnology, 38, 555 - 571. https: // doi. org / 10.1636 / Ha 09 - 64.1" type="journal article" year="2010">Santibáñez-López &amp; Francke, 2010</bibRefCitation>
): Small scorpions, adults 
<quantity id="4D769B0DFF841963FB2C3A85FA82F933" box="[1197,1317,1680,1705]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="32" pageNumber="33" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23–26 mm</quantity>
long (Fig: 22-a, b). Carapace (Fig: 13-d), shorter than metasomal segment V (Carapace L/MS-V 0.85±0.01 [0.86]), longer than the femur (Carapace L/Femur L 1.19±0.08 [1.3]); with anterior margin slightly concave. Mesosomal tergite VII with two median carinae, noticeably granular; beginning on second third of segment, composed by a single line of granules. Pectinal teeth 17–17 [15–16]; metasomal sternite V on males with a diffuse posterior, whitish patch almost absent; sternite VII with a pair of feeble carinae only present on medial third of segment and 11–12 setae. Metasomal segment I slightly wider than long, almost square in dorsal view (MSI L/ 
<geoCoordinate id="EFBA502FFF841963FB213B7DFB5EF81B" box="[1184,1273,1896,1920]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="555" value="-0.83">W 0.83</geoCoordinate>
±0.02 [0.85]); intercarinal spaces on segments I–IV sparsely granular; segment V thin, conspicuously longer than wide (MS-V L/ 
<geoCoordinate id="EFBA502FFF841963FF163BA4FF7CF852" box="[151,219,1968,1993]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5555" value="-2.5">W 2.5</geoCoordinate>
±0.24 [2.81]) and as wide as deep (W/D 1±0.07 [1]). Vesicle long and rounded (Fig: 14-d) (L/ 
<geoCoordinate id="EFBA502FFF841963FA863BA5FAFCF853" box="[1287,1371,1968,1992]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="555" value="-1.83">W 1.83</geoCoordinate>
±0.03 [1.5]) and broad, slightly wider than deep (W/D 1.19± 0.03 [1.33]), ventrally with two light-colored longitudinal median bands, conspicuously depressed; vesicle with ventral surface granular; dorsal surface flat with a tiny circle on posterior half representing the vesicular gland. Pedipalp femur (Fig: 15-d) more than three times longer than wide on males, slightly shorter on females (L/ 
<geoCoordinate id="EFBA502FFF851962FD453CF5FCBAFF63" box="[708,797,224,248]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="555" value="-3.29">W 3.29</geoCoordinate>
±0.04 [2.72]); wider than deep (W/D 1.4±0.2 [1.37]). Pedipalp patella (Fig: 16-d) more than three times longer than wide in both sexes (L/ 
<geoCoordinate id="EFBA502FFF851962FBEC3D11FB13FE87" box="[1133,1204,260,284]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="5555" value="-3.3">W 3.3</geoCoordinate>
±0.03 [3.16]); wider than deep (W/D 1.36 ±0.14 [1.2]), with dorsointernal carina granulated. Pedipalp chela manus two times longer than wide (Fig: 17, 18-d)(L/ 
<geoCoordinate id="EFBA502FFF851962FE5A3D59FD95FEFF" box="[475,562,332,357]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="555" value="-1.97">W 1.97</geoCoordinate>
±0.11 [2.06]), slightly deeper than wide (W/D 0.94±0.06 [0.88]); the dorsal prosubmedian carina granulated; fixed finger shorter than chela manus (CM L/FF 1.25±0.06 [1.2]). Dentate margins of fingers straight.
</paragraph>
<paragraph id="8A3136E8FF851962FF463DA2FD17FE6E" blockId="33.[151,1436,151,645]" pageId="33" pageNumber="34">
Hemispermatophore (Fig: 19-d): Lamelliform (measurements: TL 2.5; LL 1.7; 
<geoCoordinate id="EFBA502FFF851962FBD33DADFB3AFE4B" box="[1106,1181,440,464]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="5555" value="-0.4">W 0.4</geoCoordinate>
); capsular region with median lobe sclerotized, two basal hooks lobed.
</paragraph>
<paragraph id="8A3136E8FF851962FF463DEAFB5DFD83" blockId="33.[151,1436,151,645]" box="[199,1274,511,536]" pageId="33" pageNumber="34">Full morphometric variation is given in tables 13 and 14. Setae variation is given in tables 9–12.</paragraph>
</subSubSection>
<subSubSection id="C2946563FF851962FF463E31FEA2FD1E" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8A3136E8FF851962FF463E31FB74FDA7" blockId="33.[151,1436,151,645]" box="[199,1235,548,573]" pageId="33" pageNumber="34">
<emphasis id="B8FAEAFAFF851962FF463E31FEC7FDA6" bold="true" box="[199,352,548,573]" pageId="33" pageNumber="34">Distribution.</emphasis>
<collectingRegion id="484AF80AFF851962FEE63E31FE19FDA7" box="[359,446,548,572]" country="Mexico" name="Oaxaca" pageId="33" pageNumber="34">Oaxaca</collectingRegion>
, 
<collectingCountry id="F2997678FF851962FE483E31FD85FDA6" box="[457,546,548,573]" name="Mexico" pageId="33" pageNumber="34">México</collectingCountry>
. This species is known only from the 
<typeStatus id="5535884AFF851962FC483E30FC5EFDA6" box="[969,1017,549,573]" pageId="33" pageNumber="34">type</typeStatus>
locality (
<figureCitation id="12B52A6DFF851962FBE23E31FB61FDA7" box="[1123,1222,548,573]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="33" pageNumber="34">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="8A3136E8FF851962FF463E52FEA2FD1E" blockId="33.[151,1436,151,645]" pageId="33" pageNumber="34">
<emphasis id="B8FAEAFAFF851962FF463E52FE2DFDFB" bold="true" box="[199,394,583,608]" pageId="33" pageNumber="34">Natural history.</emphasis>
This species inhabits the rocky slopes along a disturbed riparian community at 
<quantity id="4D769B0DFF851962FAD13E52FA3BFDC4" box="[1360,1436,583,608]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="33" pageNumber="34" unit="m" value="845.0">845 m</quantity>
elevation.
</paragraph>
</subSubSection>
</treatment>
</document>