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de
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Verschelde
<emphasis id="B6DDEACE5A4FFFF8FED98356FEE7FCF9" box="[287,338,816,841]" italics="true" pageId="18" pageNumber="169">et al</emphasis>
. (1998)
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proposed a taxonomic revision together with an emended generic diagnosis and a list of valid species of
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<emphasis id="B6DDEACE5A4FFFF8FEB2833EFE48FCC1" box="[372,509,856,881]" italics="true" pageId="18" pageNumber="169">Desmodora</emphasis>
</taxonomicName>
. The list is supplemented in the NeMys database (
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).
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<emphasis id="B6DDEACE5A4FFFF8FF5183DBFE66FC67" bold="true" box="[151,467,957,983]" italics="true" pageId="18" pageNumber="169">Desmodora striatocephala</emphasis>
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<figureCitation id="1C922A595A4FFFF8FF518386FEACFC4A" box="[151,281,992,1018]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="19.[151,244,1610,1631]" captionStartId-1="20.[151,244,1900,1921]" captionTargetBox-0="[182,1416,183,1609]" captionTargetBox-1="[151,1412,187,1901]" captionTargetId-0="figure@19.[151,1436,177,1612]" captionTargetId-1="figure@20.[151,1436,178,1901]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURE 10. Desmodora striatocephala sp. nov., holotype male. A: total view; B: anterior body; C: posterior end. Scale bars: A 100 ì m; B, C 50 µ m." captionText-1="FIGURE 11. Desmodora striatocephala sp. nov., details of the holotype male. A: Anterior end; B: cloacal region. Scale bars: 10 µ m." httpUri-0="https://zenodo.org/record/183785/files/figure.png" httpUri-1="https://zenodo.org/record/183786/files/figure.png" pageId="18" pageNumber="169">Figs 1011</figureCitation>
</paragraph>
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<paragraph id="841636DC5A4FFFF8FF518456FB77FBFA" blockId="18.[151,1437,1071,2018]" box="[151,1218,1071,1098]" pageId="18" pageNumber="169">
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material
</emphasis>
:
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<emphasis id="B6DDEACE5A4FFFF8FE8E8449FE0FFBF9" bold="true" box="[328,442,1071,1097]" pageId="18" pageNumber="169">Holotype</emphasis>
</typeStatus>
male.
<typeStatus id="5B12887E5A4FFFF8FDCF8456FDF6FBFA" box="[521,579,1072,1098]" pageId="18" pageNumber="169">Type</typeStatus>
specimen deposited in the
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<collectionCode id="E2B8AE195A4FFFF8FC478456FC73FBFA" box="[897,966,1072,1098]" pageId="18" pageNumber="169">DIVA</collectionCode>
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nematode collection.
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<paragraph id="841636DC5A4FFFF8FF00843EFDCDFB2A" blockId="18.[151,1437,1071,2018]" pageId="18" pageNumber="169">
<emphasis id="B6DDEACE5A4FFFF8FF00843EFEE8FBC1" box="[198,349,1112,1137]" italics="true" pageId="18" pageNumber="169">
<typeStatus id="5B12887E5A4FFFF8FF00843EFF48FBC1" box="[198,253,1112,1137]" pageId="18" pageNumber="169">Type</typeStatus>
locality
</emphasis>
:
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<collectionCode id="E2B8AE195A4FFFF8FEAB843EFE07FBC2" box="[365,434,1112,1138]" pageId="18" pageNumber="169">DIVA</collectionCode>
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I, Meteor 48/1, Station 325/4, Multicorer 8: South­east Atlantic Ocean,
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&amp;
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, depth
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.
<date id="F017101C5A4FFFF8FE1E84E6FDC1FB2A" box="[472,628,1152,1178]" pageId="18" pageNumber="169" value="2000-07-14">14 July 2000</date>
.
</paragraph>
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<emphasis id="B6DDEACE5A4FFFF8FF0084CEFEF6FB71" box="[198,323,1192,1217]" italics="true" pageId="18" pageNumber="169">Etymology</emphasis>
: The species name reflects fine cross striation of the head.
</paragraph>
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<subSubSection id="CCB365575A4FFFFFFF0084B6FD93FE42" lastPageId="21" lastPageNumber="172" pageId="18" pageNumber="169" type="description">
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<emphasis id="B6DDEACE5A4FFFF8FF0084B6FEEDFB59" box="[198,344,1232,1257]" italics="true" pageId="18" pageNumber="169">Description.</emphasis>
Body cylindrical, of brownish­yellow colouration. Anterior end truncate. Somatic cuticle finely but distinctly annulated (just posterior to the cephalic capsule, nine annules for 10
<emphasis id="B6DDEACE5A4FFFF8FB76849EFB0AFAA1" box="[1200,1215,1272,1297]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m on convex body side and twelve annules on the opposite concave side; in the midbody, 1720 annules for 10
<emphasis id="B6DDEACE5A4FFFF8FB128546FB56FA89" box="[1236,1251,1312,1337]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), with no lateral differentiation.
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Body length 1009
<emphasis id="B6DDEACE5A4FFFF8FE598516FE1BFA39" box="[415,430,1392,1417]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, a 34.8, b 7.6, c 7.4. Body diameter at the level of: cephalic setae 16
<emphasis id="B6DDEACE5A4FFFF8FB368516FB4AFA39" box="[1264,1279,1392,1417]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, amphidial fovea 21
<emphasis id="B6DDEACE5A4FFFF8FEC285FEFEA6FA01" box="[260,275,1432,1457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, nerve ring 29
<emphasis id="B6DDEACE5A4FFFF8FE1885FEFE58FA01" box="[478,493,1432,1457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, cardia 29
<emphasis id="B6DDEACE5A4FFFF8FD4185FEFD23FA01" box="[647,662,1432,1457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, midbody 29
<emphasis id="B6DDEACE5A4FFFF8FC9485FEFCD4FA01" box="[850,865,1432,1457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m, cloaca 24
<emphasis id="B6DDEACE5A4FFFF8FC3985FEFBBBFA01" box="[1023,1038,1432,1457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m.
</paragraph>
<paragraph id="841636DC5A4FFFF8FF0085A6FD60F9E2" blockId="18.[151,1437,1071,2018]" pageId="18" pageNumber="169">
Labial region slightly drawn inward and therefore inner and outer labial sensilla not discernible. There are four short cephalic setae 3
<emphasis id="B6DDEACE5A4FFFF8FE08858EFE68F9B1" box="[462,477,1512,1537]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m long at anterior cephalic capsule. Cephalic capsule 18
<emphasis id="B6DDEACE5A4FFFF8FBB3858EFB31F9B1" box="[1141,1156,1512,1537]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m long, shaped by thick cuticle with very fine traces of fused annules; the annules of the cephalic capsule by a factor of 1.52 wider and much less distinct than postcapsular annules.
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Amphidial fovea very large and occupies nearly the whole lateral surface of the cephalic capsule. Amphidial fovea loop­shaped, slightly longitudinally oval and narrowed to its anterior end. Exit site of the amphidial nerve is thus unusually situated at the anterior end of the fovea where descending and ascending branches contact with one another. Amphidial fovea width 16
<emphasis id="B6DDEACE5A4FFFF8FC4286BEFC26F941" box="[900,915,1752,1777]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m or 76% c.b.d; distance from the cephalic apex to the amphidial fovea 34
<emphasis id="B6DDEACE5A4FFFF8FDD18766FD93F8A9" box="[535,550,1792,1817]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m.
</paragraph>
<paragraph id="841636DC5A4FFFF8FF00874EFE3DF8DA" blockId="18.[151,1437,1071,2018]" pageId="18" pageNumber="169">Somatic setae arranged in longitudinal rows distinct in the pharyngeal region and becoming irregular in the intestinal region.</paragraph>
<paragraph id="841636DC5A4FFFF8FF00871EFECFF852" blockId="18.[151,1437,1071,2018]" pageId="18" pageNumber="169">Cheilostoma indistinctly discernible because of the dense pigmentation. The subsequent part of the buccal cavity narrow, equal to the cephalic capsule. Walls of the buccal cavity and small but distinct dorsal tooth very weakly sclerotized.</paragraph>
<caption id="D0D666545A4EFFF9FF51862CFE81F9CD" httpUri="https://zenodo.org/record/183785/files/figure.png" pageId="19" pageNumber="170" targetBox="[182,1416,183,1609]" targetPageId="19">
<paragraph id="841636DC5A4EFFF9FF51862CFE81F9CD" blockId="19.[151,1436,1610,1661]" pageId="19" pageNumber="170">
<emphasis id="B6DDEACE5A4EFFF9FF51862CFEA6F9EF" bold="true" box="[151,275,1610,1631]" pageId="19" pageNumber="170">FIGURE 10</emphasis>
.
<taxonomicName id="43A94D5F5A4EFFF9FED9862DFDAAF9D0" box="[287,543,1611,1632]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="19" pageNumber="170" phylum="Nematoda" rank="species" species="striatocephala" status="sp. nov.">
<emphasis id="B6DDEACE5A4EFFF9FED9862DFDAAF9D0" box="[287,543,1611,1632]" italics="true" pageId="19" pageNumber="170">Desmodora striatocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4EFFF9FDE0862CFDC6F9EF" box="[550,627,1610,1631]" pageId="19" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male.
<emphasis id="B6DDEACE5A4EFFF9FCD4862CFC96F9EF" bold="true" box="[786,803,1610,1631]" pageId="19" pageNumber="170">A</emphasis>
: total view;
<emphasis id="B6DDEACE5A4EFFF9FC5B862CFC18F9EF" bold="true" box="[925,941,1610,1631]" pageId="19" pageNumber="170">B</emphasis>
: anterior body;
<emphasis id="B6DDEACE5A4EFFF9FB8E862CFBECF9EF" bold="true" box="[1096,1113,1610,1631]" pageId="19" pageNumber="170">C</emphasis>
: posterior end. Scale bars: A 100
<emphasis id="B6DDEACE5A4EFFF9FF51860EFF2BF9CD" box="[151,158,1640,1661]" italics="true" pageId="19" pageNumber="170">ì</emphasis>
m; B, C 50
<emphasis id="B6DDEACE5A4EFFF9FECA860EFEADF9CD" box="[268,280,1640,1661]" italics="true" pageId="19" pageNumber="170">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="841636DC5A4EFFF9FF0086ABFB39F887" blockId="19.[151,1437,1741,2007]" pageId="19" pageNumber="170">
Pharynx slender, evenly muscular, with sinuous cuticular lining of the internal lumen. Posteriorly, the pharynx widened into a clear roundish bulb. Pharynx width at the level of the nerve ring 13
<emphasis id="B6DDEACE5A4EFFF9FB048690FB64F8BF" box="[1218,1233,1782,1807]" italics="true" pageId="19" pageNumber="170">µ</emphasis>
m; posterior bulb 24
<emphasis id="B6DDEACE5A4EFFF9FF7D8778FF7FF887" box="[187,202,1822,1847]" italics="true" pageId="19" pageNumber="170">µ</emphasis>
m wide and 25
<emphasis id="B6DDEACE5A4EFFF9FE478778FE25F887" box="[385,400,1822,1847]" italics="true" pageId="19" pageNumber="170">µ</emphasis>
m long. Nerve ring hardly visible. No indication of a renette cell.
</paragraph>
<paragraph id="841636DC5A4EFFF9FF008723FB26F8EF" blockId="19.[151,1437,1741,2007]" box="[198,1171,1861,1887]" pageId="19" pageNumber="170">Cardia short, trapezium­shaped. Intestine with yellow­brownish inclusion droplets.</paragraph>
<paragraph id="841636DC5A4EFFF9FF00870BFE57F867" blockId="19.[151,1437,1741,2007]" pageId="19" pageNumber="170">Testis singular, outstretched, situated ventrally to the intestine. Spermatozoa relatively large, of uncertain shape, with granular content. Vas deferens divided into alternate regions made up of cells with transparent and coarsely granulated content.</paragraph>
<caption id="D0D666545A49FFFEFF51870AFA22F832" box="[151,1431,1900,1922]" httpUri="https://zenodo.org/record/183786/files/figure.png" pageId="20" pageNumber="171" targetBox="[151,1412,187,1901]" targetPageId="20">
<paragraph id="841636DC5A49FFFEFF51870AFA22F832" blockId="20.[151,1431,1900,1922]" box="[151,1431,1900,1922]" pageId="20" pageNumber="171">
<emphasis id="B6DDEACE5A49FFFEFF51870AFEA4F831" bold="true" box="[151,273,1900,1921]" pageId="20" pageNumber="171">FIGURE 11</emphasis>
.
<taxonomicName id="43A94D5F5A49FFFEFEDA870BFDA8F832" box="[284,541,1901,1922]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="20" pageNumber="171" phylum="Nematoda" rank="species" species="striatocephala" status="sp. nov.">
<emphasis id="B6DDEACE5A49FFFEFEDA870BFDA8F832" box="[284,541,1901,1922]" italics="true" pageId="20" pageNumber="171">Desmodora striatocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A49FFFEFDE5870BFDC5F832" box="[547,624,1901,1922]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
, details of the holotype male.
<emphasis id="B6DDEACE5A49FFFEFC49870AFC15F831" bold="true" box="[911,928,1900,1921]" pageId="20" pageNumber="171">A</emphasis>
: Anterior end;
<emphasis id="B6DDEACE5A49FFFEFBF5870AFBF6F831" bold="true" box="[1075,1091,1900,1921]" pageId="20" pageNumber="171">B</emphasis>
: cloacal region. Scale bars: 10
<emphasis id="B6DDEACE5A49FFFEFAA9870BFACEF832" box="[1391,1403,1901,1922]" italics="true" pageId="20" pageNumber="171">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="841636DC5A48FFFFFF0080FEFDDEFEB2" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">
Spicules arched, distally pointed and proximally widened into shovel­like knobs, Gubernaculum paired, as two curved plates nearly parallel to the distal half of the spicules. Spicules length 29
<emphasis id="B6DDEACE5A48FFFFFB5580A6FB17FF69" box="[1171,1186,192,217]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m (chord) and 40
<emphasis id="B6DDEACE5A48FFFFFAB280A6FA36FF69" box="[1396,1411,192,217]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m (arch); gubernaculum plate 15
<emphasis id="B6DDEACE5A48FFFFFDC6808EFDBAFEB1" box="[512,527,232,257]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m long.
</paragraph>
<paragraph id="841636DC5A48FFFFFF008176FB2DFECA" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">Precloacal ventral cuticle thickened and transparent. About five midventral supplementary organs inserted in the precloacal cuticle. Supplementary organ presents an intracuticular canal with a small papilla on the surface of the cuticle. The supplementary organs decrease in size from the cloaca anteriad.</paragraph>
<paragraph id="841636DC5A48FFFFFF0081EEFD93FE42" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">
Tail elongate conical, relatively slender, with terminal spinnerete tube. Tail length 5.32 anal diameters long. There a few pre­ and postanal subventral setae on the posterior body; their length up to 4.5
<emphasis id="B6DDEACE5A48FFFFFAD281D6FA96FE79" box="[1300,1315,432,457]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m, greater than those of the intestinal region.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A48FFFFFF008266FA8AFBFA" pageId="21" pageNumber="172" type="diagnosis">
<paragraph id="841636DC5A48FFFFFF008266FF66FCEA" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">
<emphasis id="B6DDEACE5A48FFFFFF008266FE88FDA9" box="[198,317,512,537]" italics="true" pageId="21" pageNumber="172">Diagnosis</emphasis>
. Body length 1009
<emphasis id="B6DDEACE5A48FFFFFDE38266FD81FDA9" box="[549,564,512,537]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m, a 34.8, c 7.38. There are four short cephalic setae 3
<emphasis id="B6DDEACE5A48FFFFFB7C8266FB7CFDA9" box="[1210,1225,512,537]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m long at anterior cephalic capsule. Cephalic capsule 18
<emphasis id="B6DDEACE5A48FFFFFDA8824EFDC8FDF1" box="[622,637,552,577]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m long, shaped by thick cuticle with traces of fused annules; the annules of the cephalic capsule by a factor of 1.52 wider and much less distinct than postcapsular annules. Amphidial fovea very large and occupies nearly the whole lateral surface of the cephalic capsule. Amphidial fovea loop­shaped, slightly longitudinally oval and narrowed to its anterior end. Amphidial fovea width 16
<emphasis id="B6DDEACE5A48FFFFFF5182AEFF13FD51" box="[151,166,712,737]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m or 76% c.b.d. Precloacal ventral cuticle thickened and transparent. About five midventral supplementary organs inserted in the precloacal cuticle. Supplementary organ present as an intracuticular canal with a small papilla on the surface of the cuticle. Tail elongate conical, relatively slender. Tail length 5.32 anal diameters long.
</paragraph>
<paragraph id="841636DC5A48FFFFFF00830EFA8AFBFA" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">
<emphasis id="B6DDEACE5A48FFFFFF00830EFE73FC31" box="[198,454,872,897]" italics="true" pageId="21" pageNumber="172">Differential diagnosis</emphasis>
:
<taxonomicName id="43A94D5F5A48FFFFFE11830EFCA4FC31" box="[471,785,872,897]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephala" status="sp. nov.">
<emphasis id="B6DDEACE5A48FFFFFE11830EFCA4FC31" box="[471,785,872,897]" italics="true" pageId="21" pageNumber="172">Desmodora striatocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A48FFFFFCDF830EFCC1FC32" box="[793,884,872,898]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by very fine but distinct cross striation of the cephalic capsule. Some related desmodorid genera such as
<taxonomicName id="43A94D5F5A48FFFFFBCC83F6FA22FC1A" authority="Stekhoven, 1950" authorityName="Stekhoven" authorityYear="1950" box="[1034,1431,912,938]" class="Adenophorea" family="Desmodoridae" genus="Paradesmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A48FFFFFBCC83F6FB73FC19" box="[1034,1222,912,937]" italics="true" pageId="21" pageNumber="172">Paradesmodora</emphasis>
Stekhoven, 1950
</taxonomicName>
,
<taxonomicName id="43A94D5F5A48FFFFFF5183DEFDBAFC62" authority="Blome, 1982" authorityName="Blome" authorityYear="1982" box="[151,527,952,978]" class="Adenophorea" family="Desmodoridae" genus="Echinodesmodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A48FFFFFF5183DEFED9FC61" box="[151,364,952,977]" italics="true" pageId="21" pageNumber="172">Echinodesmodora</emphasis>
Blome, 1982
</taxonomicName>
and
<taxonomicName id="43A94D5F5A48FFFFFD8D83DEFC05FC62" authority="Blome, 1982" authorityName="Blome" authorityYear="1982" box="[587,944,952,978]" class="Adenophorea" family="Desmodoridae" genus="Stygodesmodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A48FFFFFD8D83DEFCB8FC61" box="[587,781,952,977]" italics="true" pageId="21" pageNumber="172">Stygodesmodora</emphasis>
Blome, 1982
</taxonomicName>
are also characterized by a cross­striated periamphidial cuticle hardly distinguishable from that of the neck region. However, the cephalic region of
<taxonomicName id="43A94D5F5A48FFFFFAB98386FEE2FB91" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephalata">
<emphasis id="B6DDEACE5A48FFFFFAB98386FEE2FB91" italics="true" pageId="21" pageNumber="172">D. striatocephalata</emphasis>
</taxonomicName>
is clearly different in having a thickened cuticle with very fine transversal grooves thus forming a cephalic capsule set off the neck region with thinner cuticle and sharp grooves between annules.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A48FFFFFF00843EFD5BF99A" pageId="21" pageNumber="172" type="description">
<paragraph id="841636DC5A48FFFFFF00843EFD5BF99A" blockId="21.[151,1437,152,1578]" pageId="21" pageNumber="172">
<taxonomicName id="43A94D5F5A48FFFFFF00843EFDB7FBC1" box="[198,514,1112,1137]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephala" status="sp. nov.">
<emphasis id="B6DDEACE5A48FFFFFF00843EFDB7FBC1" box="[198,514,1112,1137]" italics="true" pageId="21" pageNumber="172">Desmodora striatocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A48FFFFFDCF843EFDD0FBC2" box="[521,613,1112,1138]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
is well characterized by a combination of very large amphidial fovea occupying nearly the entire lateral surface of the cephalic capsule and thickened midventral preanal cuticle with a few supplementary papillae inserted therein in males. The new species shares this set of features with
<taxonomicName id="43A94D5F5A48FFFFFF5184B6FE73FB5A" authority="Inglis, 1963" authorityName="Inglis" authorityYear="1963" box="[151,454,1232,1258]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="cuddlesae">
<emphasis id="B6DDEACE5A48FFFFFF5184B6FE84FB59" box="[151,305,1232,1257]" italics="true" pageId="21" pageNumber="172">D. cuddlesae</emphasis>
Inglis, 1963
</taxonomicName>
and to some lesser degree with
<taxonomicName id="43A94D5F5A48FFFFFC8284B6FBEBFB5A" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[836,1118,1232,1258]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="inflexa">
<emphasis id="B6DDEACE5A48FFFFFC8284B6FC02FB59" box="[836,951,1232,1257]" italics="true" pageId="21" pageNumber="172">D. inflexa</emphasis>
Wieser, 1954
</taxonomicName>
.
<taxonomicName id="43A94D5F5A48FFFFFBAD84B6FA8DFB59" box="[1131,1336,1232,1257]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephala" status="sp. nov.">
<emphasis id="B6DDEACE5A48FFFFFBAD84B6FA8DFB59" box="[1131,1336,1232,1257]" italics="true" pageId="21" pageNumber="172">D. striatocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A48FFFFFA8784B6FA29FB5A" box="[1345,1436,1232,1258]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="43A94D5F5A48FFFFFEED849EFE70FAA1" box="[299,453,1272,1297]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="cuddlesae">
<emphasis id="B6DDEACE5A48FFFFFEED849EFE70FAA1" box="[299,453,1272,1297]" italics="true" pageId="21" pageNumber="172">D. cuddlesae</emphasis>
</taxonomicName>
by the shorter body (1009
<emphasis id="B6DDEACE5A48FFFFFCC3849EFCA1FAA1" box="[773,788,1272,1297]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m
<emphasis id="B6DDEACE5A48FFFFFCF2849EFCCAFAA1" box="[820,895,1272,1297]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
17601980
<emphasis id="B6DDEACE5A48FFFFFBD5849EFB97FAA1" box="[1043,1058,1272,1297]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m), relatively longer tail (c 7.38
<emphasis id="B6DDEACE5A48FFFFFF518546FF57FA89" box="[151,226,1312,1337]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
13.514.1, c 5.32
<emphasis id="B6DDEACE5A48FFFFFE0E8546FDA6FA89" box="[456,531,1312,1337]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
2.652.75), absolutely smaller but relatively bigger amphidial fovea (16
<emphasis id="B6DDEACE5A48FFFFFAB28546FA36FA89" box="[1396,1411,1312,1337]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m wide and 76% c.b.d.
<emphasis id="B6DDEACE5A48FFFFFE56852EFE6EFAD1" box="[400,475,1352,1377]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
<quantity id="43519B395A48FFFFFE24852EFDEEFAD2" box="[482,603,1352,1378]" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="3.7" metricValueMin="2.6" pageId="21" pageNumber="172" unit="m" value="31.5" valueMax="37.0" valueMin="26.0">2637 m</quantity>
and 59% c.b.d.), about five
<emphasis id="B6DDEACE5A48FFFFFC6D852EFC43FAD1" box="[939,1014,1352,1377]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
twenty­nine preanal supplementary papillae in the ventral thickened transparent cuticle.
<taxonomicName id="43A94D5F5A48FFFFFCD28516FC50FA39" box="[788,997,1392,1417]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephala">
<emphasis id="B6DDEACE5A48FFFFFCD28516FC50FA39" box="[788,997,1392,1417]" italics="true" pageId="21" pageNumber="172">D. striatocephala</emphasis>
</taxonomicName>
differs from
<taxonomicName id="43A94D5F5A48FFFFFB4C8516FAB5FA39" box="[1162,1280,1392,1417]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="inflexa">
<emphasis id="B6DDEACE5A48FFFFFB4C8516FAB5FA39" box="[1162,1280,1392,1417]" italics="true" pageId="21" pageNumber="172">D. inflexa</emphasis>
</taxonomicName>
with shorter body (1009
<emphasis id="B6DDEACE5A48FFFFFEE385FEFE81FA01" box="[293,308,1432,1457]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m
<emphasis id="B6DDEACE5A48FFFFFE9285FEFE2AFA01" box="[340,415,1432,1457]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
2450
<emphasis id="B6DDEACE5A48FFFFFE2F85FEFE4DFA01" box="[489,504,1432,1457]" italics="true" pageId="21" pageNumber="172">µ</emphasis>
m), relatively thicker body (a 34.8
<emphasis id="B6DDEACE5A48FFFFFC5285FEFC6AFA01" box="[916,991,1432,1457]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
53.7), about five
<emphasis id="B6DDEACE5A48FFFFFB7785FEFB49FA01" box="[1201,1276,1432,1457]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
fourteen preanal supplementary papillae in the ventral thickened transparent cuticle and shorter tail (c 7.38
<emphasis id="B6DDEACE5A48FFFFFB3785A6FA89FA69" box="[1265,1340,1472,1497]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
25.8, c 5.32
<emphasis id="B6DDEACE5A48FFFFFF15858EFEABF9B1" box="[211,286,1512,1537]" italics="true" pageId="21" pageNumber="172">versus</emphasis>
2.00). Other
<taxonomicName id="43A94D5F5A48FFFFFE78858EFDF2F9B1" box="[446,583,1512,1537]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A48FFFFFE78858EFDF2F9B1" box="[446,583,1512,1537]" italics="true" pageId="21" pageNumber="172">Desmodora</emphasis>
</taxonomicName>
species with ventrally thickened preanal cuticles are much more distinguished from
<taxonomicName id="43A94D5F5A48FFFFFEFC8676FDB2F999" box="[314,519,1552,1577]" class="Adenophorea" family="Desmodoridae" genus="Desmodora" kingdom="Animalia" order="Desmodorida" pageId="21" pageNumber="172" phylum="Nematoda" rank="species" species="striatocephala">
<emphasis id="B6DDEACE5A48FFFFFEFC8676FDB2F999" box="[314,519,1552,1577]" italics="true" pageId="21" pageNumber="172">D. striatocephala</emphasis>
</taxonomicName>
in other characters.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="95AAB87A331BD923B07E9401524BD80C">Three new species of free­living nematodes from the South­East Atlantic Abyss (DIVA I Expedition) *</mods:title>
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<emphasis id="B6DDEACE5A51FFE6FF5184C1FE9AFB71" bold="true" box="[151,303,1191,1217]" italics="true" pageId="12" pageNumber="163">Pomponema</emphasis>
Cobb, 1917
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The genus was revised by
<bibRefCitation id="E0384B2D5A51FFE6FE1D8494FD13FABC" author="Lorenzen" box="[475,678,1266,1292]" pageId="12" pageNumber="173" refString="Lorenzen, S. (1972) Die Nematodenfauna im Verklappungsgebiet fur Industrieabwasser nordwestlich von Helgoland. III. Cyatholaimidae, mit einer Revision von Pomponema Cobb 1917. Veroffentlichungen des Instituts fur Meeresforschungen in Bremerhaven, 13, 285 - 306." type="journal article" year="1972">Lorenzen (1972)</bibRefCitation>
. An emended generic diagnosis of
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<emphasis id="B6DDEACE5A51FFE6FB948495FB51FABC" box="[1106,1252,1267,1292]" italics="true" pageId="12" pageNumber="163">Pomponema</emphasis>
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is provided by
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. According to NeMys (
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), twenty­nine
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<emphasis id="B6DDEACE5A51FFE6FBAC857DFB48FA84" box="[1130,1277,1307,1332]" italics="true" pageId="12" pageNumber="163">Pomponema</emphasis>
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species have been described up until recently.
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<emphasis id="B6DDEACE5A51FFE6FF5185DCFDBAFA64" bold="true" box="[151,527,1466,1492]" italics="true" pageId="12" pageNumber="163">Pomponema proximamphidum</emphasis>
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<figureCitation id="1C922A595A51FFE6FF5185BBFF48FA47" box="[151,253,1501,1527]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="14.[151,244,1971,1992]" captionStartId-1="15.[151,244,1654,1675]" captionStartId-2="17.[151,244,1927,1948]" captionTargetBox-0="[187,1388,351,1948]" captionTargetBox-1="[151,1377,185,1637]" captionTargetBox-2="[164,1436,189,1913]" captionTargetId-0="figure@14.[151,1436,340,1962]" captionTargetId-1="figure@15.[151,1436,165,1663]" captionTargetId-2="figure@17.[151,1436,177,1924]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="17" captionText-0="FIGURE 7. Pomponema proximamphidum sp. nov., anterior ends. A: male N 1 (holotype), depicted right row of denticles; B: male N 3 (paratype), surface view; C: male N 3 (paratype), optical section, depicted left row of denticles. Scale bars: 10 µ m." captionText-1="FIGURE 8. Pomponema proximamphidum sp. nov., details of the holotype male. A: anterior body; B: posterior end; C: copulatory apparatus. Scale bars: A, B 50 µm; C 10 µm." captionText-2="FIGURE 9. Pomponema proximamphidum sp. nov., details. A: male N 1 (holotype), entire; B: male N 3 (paratype), cloacal region. Scale bars: A 100 µ m; B 10 µ m." httpUri-0="https://zenodo.org/record/183782/files/figure.png" httpUri-1="https://zenodo.org/record/183783/files/figure.png" httpUri-2="https://zenodo.org/record/183784/files/figure.png" pageId="12" pageNumber="163">Figs 79</figureCitation>
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<typeStatus id="5B12887E5A51FFE6FF51864BFF79F9F6" box="[151,204,1581,1606]" pageId="12" pageNumber="163">Type</typeStatus>
material:
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<emphasis id="B6DDEACE5A51FFE6FE8B864BFE0AF9F7" bold="true" box="[333,447,1581,1607]" pageId="12" pageNumber="163">Holotype</emphasis>
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male N1, two
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males NN2 &amp; 3 mounted in glycerin on glass slides.
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specimens are deposited in the
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nematode collection.
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locality
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:
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I, Meteor 48/1, Station 346/2, Multicorer 3: South­east Atlantic Ocean,
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&amp;
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, depth
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.
<date id="F017101C5A51FFE6FE1E86C3FDC1F90F" box="[472,628,1701,1727]" pageId="12" pageNumber="163" value="2000-07-27">27 July 2000</date>
.
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<emphasis id="B6DDEACE5A51FFE6FF0086ABFEF6F956" box="[198,323,1741,1766]" italics="true" pageId="12" pageNumber="163">Etymology</emphasis>
: The species name reflects the position of amphids close to the labial position.
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<emphasis id="B6DDEACE5A51FFE6FF008693FE1EF8BE" box="[198,427,1781,1806]" italics="true" pageId="12" pageNumber="163">Morphometric data</emphasis>
: Table 3.
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<emphasis id="B6DDEACE5A51FFE6FF00877BFEE4F886" box="[198,337,1821,1846]" italics="true" pageId="12" pageNumber="163">Description</emphasis>
. Body cylindrical, slender. Cuticle annulated and punctated with dots. Cuticle heterogeneous: thick, light­refracting, brownish in colour cuticle with sharp annules in the anterior body while thinner, lighter in colour cuticle with less sharply defined annules from about 0.4 pharynx length onwards. Anterior end characterized by large, distinct dots arranged in regular transversal rows becoming smaller towards the midpharynx and fusing thereafter into transversal lines. Lateral differentiations of the body cuticle begin just posterior to the amphidial fovea. Lateral differentiation expressed in a pattern of a sequence of paired big light­refractive dots sharply set off other lateromedian dots of the respective transversal row. The big dots in pairs joined by narrow dots. Space between big lateral dots about 7
<emphasis id="B6DDEACE5A50FFE7FCD8808EFC98FEB1" box="[798,813,232,257]" italics="true" pageId="13" pageNumber="164">µ</emphasis>
m at the level of the nerve ring and 4
<emphasis id="B6DDEACE5A50FFE7FB20808EFB40FEB1" box="[1254,1269,232,257]" italics="true" pageId="13" pageNumber="164">µ</emphasis>
m on the midbody.
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<emphasis id="B6DDEACE5A50FFE7FF518107FEB6FEC9" bold="true" box="[151,259,353,377]" pageId="13" pageNumber="164">TABLE 3</emphasis>
. Morphometry of
<taxonomicName id="43A94D5F5A50FFE7FE0E8104FDF8FEC9" box="[456,589,354,377]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="13" pageNumber="164" phylum="Nematoda" rank="genus" status="sp. nov.">
<emphasis id="B6DDEACE5A50FFE7FE0E8104FDF8FEC9" box="[456,589,354,377]" italics="true" pageId="13" pageNumber="164">Pomponema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A50FFE7FD928107FD13FECA" box="[596,678,352,378]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="841636DC5A50FFE7FECA81F9FB92FE04" blockId="13.[268,1370,415,585]" box="[268,1063,415,436]" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FECA81F9FB92FE04" bold="true" box="[268,1063,415,436]" pageId="13" pageNumber="164">Character Specimens &amp; labels</emphasis>
</paragraph>
<paragraph id="841636DC5A50FFE7FD8181ACFAFCFE6F" blockId="13.[268,1370,415,585]" box="[583,1353,458,479]" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FD8181ACFAFCFE6F" bold="true" box="[583,1353,458,479]" pageId="13" pageNumber="164">
1 (
<typeStatus id="5B12887E5A50FFE7FD9981ACFD0EFE6F" box="[607,699,458,479]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
)* 2 (
<typeStatus id="5B12887E5A50FFE7FC6381ACFBB1FE6F" box="[933,1028,458,479]" pageId="13" pageNumber="164" type="paratype">paratype</typeStatus>
) 3 (
<typeStatus id="5B12887E5A50FFE7FB2281ACFAF6FE6F" box="[1252,1347,458,479]" pageId="13" pageNumber="164" type="paratype">paratype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="841636DC5A50FFE7FD818192FAF3FDB9" blockId="13.[268,1370,415,585]" box="[583,1350,500,521]" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FD818192FAF3FDB9" bold="true" box="[583,1350,500,521]" pageId="13" pageNumber="164">Box A N3(7) Box A N9(3) Box A N9(4)</emphasis>
</paragraph>
<paragraph id="841636DC5A50FFE7FDF28272FAEFFD99" blockId="13.[268,1370,415,585]" box="[564,1370,532,553]" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FDF28272FAEFFD99" bold="true" box="[564,1370,532,553]" pageId="13" pageNumber="164">
<taxonomicName id="43A94D5F5A50FFE7FDF28272FDC7FD99" box="[564,626,532,553]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="13" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1,
<taxonomicName id="43A94D5F5A50FFE7FCB08272FC01FD99" box="[886,948,532,553]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="13" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1,
<taxonomicName id="43A94D5F5A50FFE7FB728272FB44FD99" box="[1204,1265,532,553]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="13" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1,
</emphasis>
</paragraph>
<paragraph id="841636DC5A50FFE7FD828252FDDAF898" blockId="13.[268,1370,415,585]" lastBlockId="13.[151,796,1770,1866]" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FD828252FAFDFDF9" bold="true" box="[580,1352,564,585]" pageId="13" pageNumber="164">346/2, MuC3 346/2, MuC3 346/2, MuC3</emphasis>
*The specimen was slightly flattened while being measured. **Anteriormost posteriormost supplement
</paragraph>
<paragraph id="841636DC5A50FFE7FF638238FAACF976" pageId="13" pageNumber="164">
<table id="F6A9C47C5A500015FF638238FAEFF976" box="[165,1370,606,1734]" gridcols="4" gridrows="27" pageId="13" pageNumber="164">
<tr id="3A99349E5A500015FF638238FAEFFDC4" box="[165,1370,606,628]" gridrow="0" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638238FEE4FDC4" box="[165,337,606,628]" gridcol="0" gridrow="0" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638238FF00FDC3" bold="true" box="[165,181,606,627]" pageId="13" pageNumber="164">L</emphasis>
</th>
<th id="79485DE25A500015FD9F8238FD01FDC4" box="[601,692,606,628]" gridcol="1" gridrow="0" pageId="13" pageNumber="164">911</th>
<th id="79485DE25A500015FC778238FC54FDC4" box="[945,993,606,628]" gridcol="2" gridrow="0" pageId="13" pageNumber="164">1138</th>
<th id="79485DE25A500015FB2D8238FAEFFDC4" box="[1259,1370,606,628]" gridcol="3" gridrow="0" pageId="13" pageNumber="164">1203</th>
</tr>
<tr id="3A99349E5A500015FF6382EFFAEFFD2E" box="[165,1370,649,670]" gridrow="1" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6382EFFEE4FD2E" box="[165,337,649,670]" gridcol="0" gridrow="1" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6382EFFF04FD2E" bold="true" box="[165,177,649,670]" pageId="13" pageNumber="164">a</emphasis>
</th>
<td id="79485DE25A500015FD9F82EFFD01FD2E" box="[601,692,649,670]" gridcol="1" gridrow="1" pageId="13" pageNumber="164">27.6</td>
<td id="79485DE25A500015FC7782EFFC54FD2E" box="[945,993,649,670]" gridcol="2" gridrow="1" pageId="13" pageNumber="164">34</td>
<td id="79485DE25A500015FB2D82EFFAEFFD2E" box="[1259,1370,649,670]" gridcol="3" gridrow="1" pageId="13" pageNumber="164">?</td>
</tr>
<tr id="3A99349E5A500015FF6382D2FAEFFD79" box="[165,1370,692,713]" gridrow="2" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6382D2FEE4FD79" box="[165,337,692,713]" gridcol="0" gridrow="2" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6382D2FF07FD79" bold="true" box="[165,178,692,713]" pageId="13" pageNumber="164">b</emphasis>
</th>
<td id="79485DE25A500015FD9F82D2FD01FD79" box="[601,692,692,713]" gridcol="1" gridrow="2" pageId="13" pageNumber="164">4.67</td>
<td id="79485DE25A500015FC7782D2FC54FD79" box="[945,993,692,713]" gridcol="2" gridrow="2" pageId="13" pageNumber="164">5.83</td>
<td id="79485DE25A500015FB2D82D2FAEFFD79" box="[1259,1370,692,713]" gridcol="3" gridrow="2" pageId="13" pageNumber="164">6.17</td>
</tr>
<tr id="3A99349E5A500015FF6382B8FAEFFD44" box="[165,1370,734,756]" gridrow="3" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6382B8FEE4FD44" box="[165,337,734,756]" gridcol="0" gridrow="3" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6382B8FF05FD43" bold="true" box="[165,176,734,755]" pageId="13" pageNumber="164">c</emphasis>
</th>
<td id="79485DE25A500015FD9F82B8FD01FD44" box="[601,692,734,756]" gridcol="1" gridrow="3" pageId="13" pageNumber="164">4.00</td>
<td id="79485DE25A500015FC7782B8FC54FD44" box="[945,993,734,756]" gridcol="2" gridrow="3" pageId="13" pageNumber="164">6.03</td>
<td id="79485DE25A500015FB2D82B8FAEFFD44" box="[1259,1370,734,756]" gridcol="3" gridrow="3" pageId="13" pageNumber="164">5.87</td>
</tr>
<tr id="3A99349E5A500015FF63836FFAEFFCAE" box="[165,1370,777,798]" gridrow="4" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF63836FFEE4FCAE" box="[165,337,777,798]" gridcol="0" gridrow="4" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF63836FFF4AFCAE" bold="true" box="[165,255,777,798]" pageId="13" pageNumber="164">diam.c.s.</emphasis>
</th>
<td id="79485DE25A500015FD9F836FFD01FCAE" box="[601,692,777,798]" gridcol="1" gridrow="4" pageId="13" pageNumber="164">13</td>
<td id="79485DE25A500015FC77836FFC54FCAE" box="[945,993,777,798]" gridcol="2" gridrow="4" pageId="13" pageNumber="164">14</td>
<td id="79485DE25A500015FB2D836FFAEFFCAE" box="[1259,1370,777,798]" gridcol="3" gridrow="4" pageId="13" pageNumber="164">14</td>
</tr>
<tr id="3A99349E5A500015FF638355FAEFFCF9" box="[165,1370,819,841]" gridrow="5" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638355FEE4FCF9" box="[165,337,819,841]" gridcol="0" gridrow="5" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638355FEB0FCF8" bold="true" box="[165,261,819,840]" pageId="13" pageNumber="164">diam.am.</emphasis>
</th>
<td id="79485DE25A500015FD9F8355FD01FCF9" box="[601,692,819,841]" gridcol="1" gridrow="5" pageId="13" pageNumber="164">19</td>
<td id="79485DE25A500015FC778355FC54FCF9" box="[945,993,819,841]" gridcol="2" gridrow="5" pageId="13" pageNumber="164">19</td>
<td id="79485DE25A500015FB2D8355FAEFFCF9" box="[1259,1370,819,841]" gridcol="3" gridrow="5" pageId="13" pageNumber="164">20</td>
</tr>
<tr id="3A99349E5A500015FF638338FAEFFCC3" box="[165,1370,862,883]" gridrow="6" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638338FEE4FCC3" box="[165,337,862,883]" gridcol="0" gridrow="6" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638338FEB6FCC3" bold="true" box="[165,259,862,883]" pageId="13" pageNumber="164">diam.n.r.</emphasis>
</th>
<td id="79485DE25A500015FD9F8338FD01FCC3" box="[601,692,862,883]" gridcol="1" gridrow="6" pageId="13" pageNumber="164">38</td>
<td id="79485DE25A500015FC778338FC54FCC3" box="[945,993,862,883]" gridcol="2" gridrow="6" pageId="13" pageNumber="164">29</td>
<td id="79485DE25A500015FB2D8338FAEFFCC3" box="[1259,1370,862,883]" gridcol="3" gridrow="6" pageId="13" pageNumber="164">30</td>
</tr>
<tr id="3A99349E5A500015FF6383EEFAEFFC2D" box="[165,1370,904,925]" gridrow="7" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6383EEFEE4FC2D" box="[165,337,904,925]" gridcol="0" gridrow="7" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6383EEFF49FC2D" bold="true" box="[165,252,904,925]" pageId="13" pageNumber="164">diam.ca.</emphasis>
</th>
<td id="79485DE25A500015FD9F83EEFD01FC2D" box="[601,692,904,925]" gridcol="1" gridrow="7" pageId="13" pageNumber="164">41</td>
<td id="79485DE25A500015FC7783EEFC54FC2D" box="[945,993,904,925]" gridcol="2" gridrow="7" pageId="13" pageNumber="164">32</td>
<td id="79485DE25A500015FB2D83EEFAEFFC2D" box="[1259,1370,904,925]" gridcol="3" gridrow="7" pageId="13" pageNumber="164">29</td>
</tr>
<tr id="3A99349E5A500015FF6383D5FAEFFC78" box="[165,1370,947,968]" gridrow="8" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6383D5FEE4FC78" box="[165,337,947,968]" gridcol="0" gridrow="8" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6383D5FEAFFC78" bold="true" box="[165,282,947,968]" pageId="13" pageNumber="164">diam.midb.</emphasis>
</th>
<td id="79485DE25A500015FD9F83D5FD01FC78" box="[601,692,947,968]" gridcol="1" gridrow="8" pageId="13" pageNumber="164">47</td>
<td id="79485DE25A500015FC7783D5FC54FC78" box="[945,993,947,968]" gridcol="2" gridrow="8" pageId="13" pageNumber="164">33.5</td>
<td id="79485DE25A500015FB2D83D5FAEFFC78" box="[1259,1370,947,968]" gridcol="3" gridrow="8" pageId="13" pageNumber="164">?</td>
</tr>
<tr id="3A99349E5A500015FF6383BBFAEFFC43" box="[165,1370,989,1011]" gridrow="9" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6383BBFEE4FC43" box="[165,337,989,1011]" gridcol="0" gridrow="9" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6383BBFF7EFC42" bold="true" box="[165,203,989,1010]" pageId="13" pageNumber="164">a.d.</emphasis>
</th>
<td id="79485DE25A500015FD9F83BBFD01FC43" box="[601,692,989,1011]" gridcol="1" gridrow="9" pageId="13" pageNumber="164">27</td>
<td id="79485DE25A500015FC7783BBFC54FC43" box="[945,993,989,1011]" gridcol="2" gridrow="9" pageId="13" pageNumber="164">24</td>
<td id="79485DE25A500015FB2D83BBFAEFFC43" box="[1259,1370,989,1011]" gridcol="3" gridrow="9" pageId="13" pageNumber="164">24</td>
</tr>
<tr id="3A99349E5A500015FF63846EFAEFFBAD" box="[165,1370,1032,1053]" gridrow="10" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF63846EFEE4FBAD" box="[165,337,1032,1053]" gridcol="0" gridrow="10" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF63846EFF66FBAD" bold="true" box="[165,211,1032,1053]" pageId="13" pageNumber="164">o.l.s.</emphasis>
</th>
<td id="79485DE25A500015FD9F846EFD01FBAD" box="[601,692,1032,1053]" gridcol="1" gridrow="10" pageId="13" pageNumber="164">2+5.5***</td>
<td id="79485DE25A500015FC77846EFC54FBAD" box="[945,993,1032,1053]" gridcol="2" gridrow="10" pageId="13" pageNumber="164">2+5</td>
<td id="79485DE25A500015FB2D846EFAEFFBAD" box="[1259,1370,1032,1053]" gridcol="3" gridrow="10" pageId="13" pageNumber="164">2+5</td>
</tr>
<tr id="3A99349E5A500015FF638454FAEFFBF8" box="[165,1370,1074,1096]" gridrow="11" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638454FEE4FBF8" box="[165,337,1074,1096]" gridcol="0" gridrow="11" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638454FF70FBF7" bold="true" box="[165,197,1074,1095]" pageId="13" pageNumber="164">c.s.</emphasis>
</th>
<td id="79485DE25A500015FD9F8454FD01FBF8" box="[601,692,1074,1096]" gridcol="1" gridrow="11" pageId="13" pageNumber="164">4.5</td>
<td id="79485DE25A500015FC778454FC54FBF8" box="[945,993,1074,1096]" gridcol="2" gridrow="11" pageId="13" pageNumber="164">4</td>
<td id="79485DE25A500015FB2D8454FAEFFBF8" box="[1259,1370,1074,1096]" gridcol="3" gridrow="11" pageId="13" pageNumber="164">4</td>
</tr>
<tr id="3A99349E5A500015FF63843BFAEFFBC2" box="[165,1370,1117,1138]" gridrow="12" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF63843BFEE4FBC2" box="[165,337,1117,1138]" gridcol="0" gridrow="12" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF63843BFF57FBC2" bold="true" box="[165,226,1117,1138]" pageId="13" pageNumber="164">am.w.</emphasis>
</th>
<td id="79485DE25A500015FD9F843BFD01FBC2" box="[601,692,1117,1138]" gridcol="1" gridrow="12" pageId="13" pageNumber="164">9</td>
<td id="79485DE25A500015FC77843BFC54FBC2" box="[945,993,1117,1138]" gridcol="2" gridrow="12" pageId="13" pageNumber="164">10.5</td>
<td id="79485DE25A500015FB2D843BFAEFFBC2" box="[1259,1370,1117,1138]" gridcol="3" gridrow="12" pageId="13" pageNumber="164">11.5</td>
</tr>
<tr id="3A99349E5A500015FF6384E1FAEFFB2D" box="[165,1370,1159,1181]" gridrow="13" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6384E1FEE4FB2D" box="[165,337,1159,1181]" gridcol="0" gridrow="13" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6384E1FEA5FB2C" bold="true" box="[165,272,1159,1180]" pageId="13" pageNumber="164">am.w. (%)</emphasis>
</th>
<td id="79485DE25A500015FD9F84E1FD01FB2D" box="[601,692,1159,1181]" gridcol="1" gridrow="13" pageId="13" pageNumber="164">59</td>
<td id="79485DE25A500015FC7784E1FC54FB2D" box="[945,993,1159,1181]" gridcol="2" gridrow="13" pageId="13" pageNumber="164">55</td>
<td id="79485DE25A500015FB2D84E1FAEFFB2D" box="[1259,1370,1159,1181]" gridcol="3" gridrow="13" pageId="13" pageNumber="164">57.5</td>
</tr>
<tr id="3A99349E5A500015FF6384D4FAEFFB77" box="[165,1370,1202,1223]" gridrow="14" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6384D4FEE4FB77" box="[165,337,1202,1223]" gridcol="0" gridrow="14" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6384D4FF61FB77" bold="true" box="[165,212,1202,1223]" pageId="13" pageNumber="164">st.w.</emphasis>
</th>
<td id="79485DE25A500015FD9F84D4FD01FB77" box="[601,692,1202,1223]" gridcol="1" gridrow="14" pageId="13" pageNumber="164">8</td>
<td id="79485DE25A500015FC7784D4FC54FB77" box="[945,993,1202,1223]" gridcol="2" gridrow="14" pageId="13" pageNumber="164">8</td>
<td id="79485DE25A500015FB2D84D4FAEFFB77" box="[1259,1370,1202,1223]" gridcol="3" gridrow="14" pageId="13" pageNumber="164">6</td>
</tr>
<tr id="3A99349E5A500015FF6384BBFAEFFB42" box="[165,1370,1245,1266]" gridrow="15" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6384BBFEE4FB42" box="[165,337,1245,1266]" gridcol="0" gridrow="15" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6384BBFF7CFB42" bold="true" box="[165,201,1245,1266]" pageId="13" pageNumber="164">st.l.</emphasis>
</th>
<td id="79485DE25A500015FD9F84BBFD01FB42" box="[601,692,1245,1266]" gridcol="1" gridrow="15" pageId="13" pageNumber="164">27</td>
<td id="79485DE25A500015FC7784BBFC54FB42" box="[945,993,1245,1266]" gridcol="2" gridrow="15" pageId="13" pageNumber="164">24</td>
<td id="79485DE25A500015FB2D84BBFAEFFB42" box="[1259,1370,1245,1266]" gridcol="3" gridrow="15" pageId="13" pageNumber="164">25</td>
</tr>
<tr id="3A99349E5A500015FF638561FAEFFAAC" box="[165,1370,1287,1308]" gridrow="16" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638561FEE4FAAC" box="[165,337,1287,1308]" gridcol="0" gridrow="16" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638561FF5DFAAC" bold="true" box="[165,232,1287,1308]" pageId="13" pageNumber="164">cheil.l.</emphasis>
</th>
<td id="79485DE25A500015FD9F8561FD01FAAC" box="[601,692,1287,1308]" gridcol="1" gridrow="16" pageId="13" pageNumber="164">8</td>
<td id="79485DE25A500015FC778561FC54FAAC" box="[945,993,1287,1308]" gridcol="2" gridrow="16" pageId="13" pageNumber="164">8</td>
<td id="79485DE25A500015FB2D8561FAEFFAAC" box="[1259,1370,1287,1308]" gridcol="3" gridrow="16" pageId="13" pageNumber="164">6</td>
</tr>
<tr id="3A99349E5A500015FF638557FAEFFAF7" box="[165,1370,1329,1351]" gridrow="17" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638557FEE4FAF7" box="[165,337,1329,1351]" gridcol="0" gridrow="17" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638557FF41FAF6" bold="true" box="[165,244,1329,1350]" pageId="13" pageNumber="164">dors.t.l.</emphasis>
</th>
<td id="79485DE25A500015FD9F8557FD01FAF7" box="[601,692,1329,1351]" gridcol="1" gridrow="17" pageId="13" pageNumber="164">17</td>
<td id="79485DE25A500015FC778557FC54FAF7" box="[945,993,1329,1351]" gridcol="2" gridrow="17" pageId="13" pageNumber="164">19</td>
<td id="79485DE25A500015FB2D8557FAEFFAF7" box="[1259,1370,1329,1351]" gridcol="3" gridrow="17" pageId="13" pageNumber="164">18</td>
</tr>
<tr id="3A99349E5A500015FF63853AFAEFFAC1" box="[165,1370,1372,1393]" gridrow="18" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF63853AFEE4FAC1" box="[165,337,1372,1393]" gridcol="0" gridrow="18" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF63853AFEBCFAC1" bold="true" box="[165,265,1372,1393]" pageId="13" pageNumber="164">
dis.
<taxonomicName id="43A94D5F5A50FFE7FF0E853AFEBCFAC1" box="[200,265,1372,1393]" pageId="13" pageNumber="164" rank="variety" variety="pore">v.pore</taxonomicName>
</emphasis>
</th>
<td id="79485DE25A500015FD9F853AFD01FAC1" box="[601,692,1372,1393]" gridcol="1" gridrow="18" pageId="13" pageNumber="164">54</td>
<td id="79485DE25A500015FC77853AFC54FAC1" box="[945,993,1372,1393]" gridcol="2" gridrow="18" pageId="13" pageNumber="164">60</td>
<td id="79485DE25A500015FB2D853AFAEFFAC1" box="[1259,1370,1372,1393]" gridcol="3" gridrow="18" pageId="13" pageNumber="164">58</td>
</tr>
<tr id="3A99349E5A500015FF6385E0FAEFFA2C" box="[165,1370,1414,1436]" gridrow="19" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6385E0FEE4FA2C" box="[165,337,1414,1436]" gridcol="0" gridrow="19" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6385E0FEBBFA2B" bold="true" box="[165,270,1414,1435]" pageId="13" pageNumber="164">spic.chord</emphasis>
</th>
<td id="79485DE25A500015FD9F85E0FD01FA2C" box="[601,692,1414,1436]" gridcol="1" gridrow="19" pageId="13" pageNumber="164">33</td>
<td id="79485DE25A500015FC7785E0FC54FA2C" box="[945,993,1414,1436]" gridcol="2" gridrow="19" pageId="13" pageNumber="164">33</td>
<td id="79485DE25A500015FB2D85E0FAEFFA2C" box="[1259,1370,1414,1436]" gridcol="3" gridrow="19" pageId="13" pageNumber="164">35</td>
</tr>
<tr id="3A99349E5A500015FF6385D7FAEFFA76" box="[165,1370,1457,1478]" gridrow="20" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6385D7FEE4FA76" box="[165,337,1457,1478]" gridcol="0" gridrow="20" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6385D7FF41FA76" bold="true" box="[165,244,1457,1478]" pageId="13" pageNumber="164">spic.arc</emphasis>
</th>
<td id="79485DE25A500015FD9F85D7FD01FA76" box="[601,692,1457,1478]" gridcol="1" gridrow="20" pageId="13" pageNumber="164">41</td>
<td id="79485DE25A500015FC7785D7FC54FA76" box="[945,993,1457,1478]" gridcol="2" gridrow="20" pageId="13" pageNumber="164">46</td>
<td id="79485DE25A500015FB2D85D7FAEFFA76" box="[1259,1370,1457,1478]" gridcol="3" gridrow="20" pageId="13" pageNumber="164">44.5</td>
</tr>
<tr id="3A99349E5A500015FF6385BAFAEFFA41" box="[165,1370,1500,1521]" gridrow="21" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6385BAFEE4FA41" box="[165,337,1500,1521]" gridcol="0" gridrow="21" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6385BAFF6AFA41" bold="true" box="[165,223,1500,1521]" pageId="13" pageNumber="164">gub.l.</emphasis>
</th>
<td id="79485DE25A500015FD9F85BAFD01FA41" box="[601,692,1500,1521]" gridcol="1" gridrow="21" pageId="13" pageNumber="164">21</td>
<td id="79485DE25A500015FC7785BAFC54FA41" box="[945,993,1500,1521]" gridcol="2" gridrow="21" pageId="13" pageNumber="164">19</td>
<td id="79485DE25A500015FB2D85BAFAEFFA41" box="[1259,1370,1500,1521]" gridcol="3" gridrow="21" pageId="13" pageNumber="164">17</td>
</tr>
<tr id="3A99349E5A500015FF638660FAEFF9AB" box="[165,1370,1542,1563]" gridrow="22" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638660FEE4F9AB" box="[165,337,1542,1563]" gridcol="0" gridrow="22" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638660FF43F9AB" bold="true" box="[165,246,1542,1563]" pageId="13" pageNumber="164">suppl.n.</emphasis>
</th>
<td id="79485DE25A500015FD9F8660FD01F9AB" box="[601,692,1542,1563]" gridcol="1" gridrow="22" pageId="13" pageNumber="164">14</td>
<td id="79485DE25A500015FC778660FC54F9AB" box="[945,993,1542,1563]" gridcol="2" gridrow="22" pageId="13" pageNumber="164">13</td>
<td id="79485DE25A500015FB2D8660FAEFF9AB" box="[1259,1370,1542,1563]" gridcol="3" gridrow="22" pageId="13" pageNumber="164">13</td>
</tr>
<tr id="3A99349E5A500015FF638657FAEFF9F6" box="[165,1370,1585,1606]" gridrow="23" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF638657FEE4F9F6" box="[165,337,1585,1606]" gridcol="0" gridrow="23" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF638657FF45F9F6" bold="true" box="[165,240,1585,1606]" pageId="13" pageNumber="164">suppl.l.</emphasis>
</th>
<td id="79485DE25A500015FD9F8657FD01F9F6" box="[601,692,1585,1606]" gridcol="1" gridrow="23" pageId="13" pageNumber="164">76**</td>
<td id="79485DE25A500015FC778657FC54F9F6" box="[945,993,1585,1606]" gridcol="2" gridrow="23" pageId="13" pageNumber="164">87</td>
<td id="79485DE25A500015FB2D8657FAEFF9F6" box="[1259,1370,1585,1606]" gridcol="3" gridrow="23" pageId="13" pageNumber="164">10 ­ 7</td>
</tr>
<tr id="3A99349E5A500015FF63863DFAEFF9C1" box="[165,1370,1627,1649]" gridrow="24" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF63863DFEE4F9C1" box="[165,337,1627,1649]" gridcol="0" gridrow="24" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF63863DFF02F9C0" bold="true" box="[165,183,1627,1648]" pageId="13" pageNumber="164">c</emphasis>
</th>
<td id="79485DE25A500015FD9F863DFD01F9C1" box="[601,692,1627,1649]" gridcol="1" gridrow="24" pageId="13" pageNumber="164">8.12</td>
<td id="79485DE25A500015FC77863DFC54F9C1" box="[945,993,1627,1649]" gridcol="2" gridrow="24" pageId="13" pageNumber="164">7.45</td>
<td id="79485DE25A500015FB2D863DFAEFF9C1" box="[1259,1370,1627,1649]" gridcol="3" gridrow="24" pageId="13" pageNumber="164">7.89</td>
</tr>
<tr id="3A99349E5A500015FF6386E0FAEFF92B" box="[165,1370,1670,1691]" gridrow="25" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6386E0FEE4F92B" box="[165,337,1670,1691]" gridcol="0" gridrow="25" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6386E0FEE4F92B" bold="true" box="[165,337,1670,1691]" pageId="13" pageNumber="164">dist.tail part (%)</emphasis>
</th>
<td id="79485DE25A500015FD9F86E0FD01F92B" box="[601,692,1670,1691]" gridcol="1" gridrow="25" pageId="13" pageNumber="164">62</td>
<td id="79485DE25A500015FC7786E0FC54F92B" box="[945,993,1670,1691]" gridcol="2" gridrow="25" pageId="13" pageNumber="164">52</td>
<td id="79485DE25A500015FB2D86E0FAEFF92B" box="[1259,1370,1670,1691]" gridcol="3" gridrow="25" pageId="13" pageNumber="164">52</td>
</tr>
<tr id="3A99349E5A500015FF6386D6FAEFF976" box="[165,1370,1712,1734]" gridrow="26" pageId="13" pageNumber="164">
<th id="79485DE25A500015FF6386D6FEE4F976" box="[165,337,1712,1734]" gridcol="0" gridrow="26" pageId="13" pageNumber="164">
<emphasis id="B6DDEACE5A50FFE7FF6386D6FEBBF975" bold="true" box="[165,270,1712,1733]" pageId="13" pageNumber="164">termin sp.</emphasis>
</th>
<td id="79485DE25A500015FD9F86D6FD01F976" box="[601,692,1712,1734]" gridcol="1" gridrow="26" pageId="13" pageNumber="164">?</td>
<td id="79485DE25A500015FC7786D6FC54F976" box="[945,993,1712,1734]" gridcol="2" gridrow="26" pageId="13" pageNumber="164">3.5</td>
<td id="79485DE25A500015FB2D86D6FAEFF976" box="[1259,1370,1712,1734]" gridcol="3" gridrow="26" pageId="13" pageNumber="164">34</td>
</tr>
</table>
</paragraph>
<paragraph id="841636DC5A50FFE7FF518755FDB5F8FA" blockId="13.[151,796,1770,1866]" box="[151,512,1843,1866]" pageId="13" pageNumber="164">***lateral + lateromedian sensilla</paragraph>
<paragraph id="841636DC5A50FFE4FF0087EDFE43FEB2" blockId="13.[151,1437,1931,2037]" lastBlockId="14.[151,1436,152,258]" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164">Inner labial sensilla indistinct. Outer labial and cephalic sensilla constitute together a common circle. Six outer labial setae differ notably to one another: lateral setae much shorter than lateromedian outer labial setae. Four cephalic setae a little shorter than the outer labial lateromedian setae. Cervical and somatic setae not found. There are few fuzzy cuticular pores in the pharyngeal region and no visible pores farther posteriad. Amphidial fovea very large multispiral groove, ventrally coiled, with about five turns. It is half shifted to the apical surface of the head.</paragraph>
<caption id="D0D666545A53FFE4FF5187D5FB6AF856" httpUri="https://zenodo.org/record/183782/files/figure.png" pageId="14" pageNumber="165" targetBox="[187,1388,351,1948]" targetPageId="14">
<paragraph id="841636DC5A53FFE4FF5187D5FB6AF856" blockId="14.[151,1435,1971,2022]" pageId="14" pageNumber="165">
<emphasis id="B6DDEACE5A53FFE4FF5187D5FEB3F878" bold="true" box="[151,262,1971,1992]" pageId="14" pageNumber="165">FIGURE 7</emphasis>
.
<taxonomicName id="43A94D5F5A53FFE4FED787D2FD82F879" box="[273,567,1972,1993]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="14" pageNumber="165" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A53FFE4FED787D2FD82F879" box="[273,567,1972,1993]" italics="true" pageId="14" pageNumber="165">Pomponema proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A53FFE4FDFB87D2FD3CF879" box="[573,649,1972,1993]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
, anterior ends.
<emphasis id="B6DDEACE5A53FFE4FCDC87D5FC9EF878" bold="true" box="[794,811,1971,1992]" pageId="14" pageNumber="165">A</emphasis>
: male N1 (holotype), depicted right row of denticles;
<emphasis id="B6DDEACE5A53FFE4FAE887D5FA8BF878" bold="true" box="[1326,1342,1971,1992]" pageId="14" pageNumber="165">B</emphasis>
: male N3 (paratype), surface view;
<emphasis id="B6DDEACE5A53FFE4FE4A87B7FE28F856" bold="true" box="[396,413,2001,2022]" pageId="14" pageNumber="165">C</emphasis>
: male N3 (paratype), optical section, depicted left row of denticles. Scale bars: 10
<emphasis id="B6DDEACE5A53FFE4FB7187B7FB76F856" box="[1207,1219,2001,2022]" italics="true" pageId="14" pageNumber="165">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="D0D666545A52FFE5FF518610FDF2F918" httpUri="https://zenodo.org/record/183783/files/figure.png" pageId="15" pageNumber="166" targetBox="[151,1377,185,1637]" targetPageId="15">
<paragraph id="841636DC5A52FFE5FF518610FDF2F918" blockId="15.[151,1436,1654,1704]" pageId="15" pageNumber="166">
<emphasis id="B6DDEACE5A52FFE5FF518610FEB2F93B" bold="true" box="[151,263,1654,1675]" pageId="15" pageNumber="166">FIGURE 8</emphasis>
.
<taxonomicName id="43A94D5F5A52FFE5FED38610FD88F93B" box="[277,573,1654,1675]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="15" pageNumber="166" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A52FFE5FED38610FD88F93B" box="[277,573,1654,1675]" italics="true" pageId="15" pageNumber="166">Pomponema proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A52FFE5FD828610FD27F93B" box="[580,658,1654,1675]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
, details of the holotype male.
<emphasis id="B6DDEACE5A52FFE5FC7D8610FC79F93B" bold="true" box="[955,972,1654,1675]" pageId="15" pageNumber="166">A</emphasis>
: anterior body;
<emphasis id="B6DDEACE5A52FFE5FBAC8610FBCFF93B" bold="true" box="[1130,1146,1654,1675]" pageId="15" pageNumber="166">B</emphasis>
: posterior end;
<emphasis id="B6DDEACE5A52FFE5FAD08610FA92F93B" bold="true" box="[1302,1319,1654,1675]" pageId="15" pageNumber="166">C</emphasis>
: copulatory apparatus. Scale bars: A, B 50 µm; C 10 µm.
</paragraph>
</caption>
<paragraph id="841636DC5A52FFE5FF0086BFFC81F8DB" blockId="15.[151,1437,1753,2019]" pageId="15" pageNumber="166">Buccal cavity differentiated into two compartments. Cheilostoma shaped as a truncate cone; its walls armoured with twelve stick­like ternary (three­storyed) rugae. Stegostoma provided with a big and acute dorsal tooth as well as two smaller subventral acute teeth. At the level of each side of the subventral teeth, a lateroventral row of six to eight tiny two­storyed dentlicles.</paragraph>
<paragraph id="841636DC5A52FFFAFF00871FFEE6FF02" blockId="15.[151,1437,1753,2019]" lastBlockId="16.[151,1437,152,2018]" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">Pharynx evenly muscular throughout its length, slightly widening posteriorly but not forming a well defined bulb. Nerve ring well visible at the midpharynx. Terminal ampulla of the dorsal pharyngeal gland discernible at the base of the dorsal tooth. Cardia indistinct. There are rather large swallowed particles inside the intestine lumen.</paragraph>
<paragraph id="841636DC5A4DFFFAFF0080A6FDAEFEB2" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">Renette ampulla transparent and voluminous; ventral pore situated at a distance of about one third pharynx length from the anterior end.</paragraph>
<paragraph id="841636DC5A4DFFFAFF008176FC72FECA" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">Reproductive system diorchic. Both anterior outstretched and posterior reflected testes situated to the left of the intestine in all the specimens. Vas deferens consists of several parts (from anterior end caudad): medium granulated, finely granulated, homogeneous, and coarsely granulated.</paragraph>
<paragraph id="841636DC5A4DFFFAFF0081EEFDDBFE42" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">Spicules short, arched, distally pointed and proximally widened. Gubernaculum as paired slightly curved plate distally adjoined to spicules. There is a small funnel­like structure with an indented sclerotized rim at each lateral side of the spicule distal tip.</paragraph>
<paragraph id="841636DC5A4DFFFAFF008266FF48FD22" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">Thirteen to fourteen midventral preanal supplementary organs. Each organ consists of a stout cylindrical body and a flat cover with a central opening. The supplementary organs may or may not be elevated above the body cuticle depending on the body curvature. There a small midventral sensilla just anterior to the cloacal opening.</paragraph>
<paragraph id="841636DC5A4DFFFAFF0082C6FBFAFD52" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">Tail relatively long, consisting of proximal conical and slender cylindrical parts. A short terminal spinneret tube at the tail end. There are a few small setae laterally on the conical part.</paragraph>
</subSubSection>
<subSubSection id="CCB365575A4DFFFAFF008296FE8DFA02" pageId="16" pageNumber="167" type="diagnosis">
<paragraph id="841636DC5A4DFFFAFF008296FF71FAA2" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">
<emphasis id="B6DDEACE5A4DFFFAFF008296FE88FCB9" box="[198,317,752,777]" italics="true" pageId="16" pageNumber="167">Diagnosis</emphasis>
.
<collectingCountry id="FCBE764C5A4DFFFAFE8B8296FE3CFCBA" box="[333,393,752,778]" name="Malta" pageId="16" pageNumber="167">Male</collectingCountry>
body length 9111203
<emphasis id="B6DDEACE5A4DFFFAFD628296FD06FCB9" box="[676,691,752,777]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m. Cuticle heterogeneous: anteriormost cuticle thicker, brownish, with sharp annules in the anterior body while more posterior cuticle thinner, light, with less sharply defined annules. Lateral differentiation expressed in a pattern of a sequence of paired light­refractive dots sharply set off other much smaller lateromedian dots. Inner labial sensilla as conical papillae. Six outer labial sensilla differ notably to one another: labial sensilla as minute setae much shorter than lateromedian sensilla looking like true short setae. Four cephalic setae a little shorter than outer labial lateromedian setae. Amphidial fovea very large, multispiral, with about five turns; half shifted to the apical surface of the head. Stegostoma armoured by a big acute dorsal tooth, two smaller subventral acute teeth and two lateroventral rows of denticles at the level of the subventral teeth. Ventral pore situated at a distance of about one third pharynx length from the anterior end. Diorchic, posterior testis reflexed. Spicules short, arched, distally pointed and proximally widened. Gubernaculum as paired slightly curved plate with a small funnel­like structure with an indented sclerotized rim at each lateral side of the spicule distal tip. Thirteen to fourteen midventral preanal supplementary organs. Tail length 7.458.12 anal diameters; its distal cylindrical part 5262% of the entire tail.
</paragraph>
<paragraph id="841636DC5A4DFFFAFF008546FE8DFA02" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">
<emphasis id="B6DDEACE5A4DFFFAFF008546FE7DFA89" box="[198,456,1312,1337]" italics="true" pageId="16" pageNumber="167">Differential diagnosis</emphasis>
:
<taxonomicName id="43A94D5F5A4DFFFAFE1C8546FD72FA89" box="[474,711,1312,1337]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4DFFFAFE1C8546FD72FA89" box="[474,711,1312,1337]" italics="true" pageId="16" pageNumber="167">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4DFFFAFD168546FC98FA8A" box="[720,813,1312,1338]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
share the subapical position of the amphidial fovea and distinct lateral differentiation of the somatic cuticle with five other species:
<taxonomicName id="43A94D5F5A4DFFFAFBFD852EFACEFAD2" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[1083,1403,1352,1378]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="concinnum">
<emphasis id="B6DDEACE5A4DFFFAFBFD852EFB62FAD1" box="[1083,1239,1352,1377]" italics="true" pageId="16" pageNumber="167">P. concinnum</emphasis>
Wieser, 1954
</taxonomicName>
,
<taxonomicName id="43A94D5F5A4DFFFAFA40852EFE44FA3A" authority="Gourbault, 1980" authorityName="Gourbault" authorityYear="1980" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="corniculata">
<emphasis id="B6DDEACE5A4DFFFAFA40852EFEABFA39" italics="true" pageId="16" pageNumber="167">P. corniculata</emphasis>
Gourbault, 1980
</taxonomicName>
,
<taxonomicName id="43A94D5F5A4DFFFAFDC68516FCAAFA3A" authority="Cobb, 1917" authorityName="Cobb" authorityYear="1917" box="[512,799,1392,1418]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="mirabile">
<emphasis id="B6DDEACE5A4DFFFAFDC68516FD30FA39" box="[512,645,1392,1417]" italics="true" pageId="16" pageNumber="167">P. mirabile</emphasis>
Cobb, 1917
</taxonomicName>
,
<taxonomicName id="43A94D5F5A4DFFFAFCF68516FB0AFA3A" authority="Filipjev, 1922" authorityName="Filipjev" authorityYear="1922" box="[816,1215,1392,1418]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="multipapillatum">
<emphasis id="B6DDEACE5A4DFFFAFCF68516FBB9FA39" box="[816,1036,1392,1417]" italics="true" pageId="16" pageNumber="167">P. multipapillatum</emphasis>
Filipjev, 1922
</taxonomicName>
and
<taxonomicName id="43A94D5F5A4DFFFAFB398516FE81FA02" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="stomachor">
<emphasis id="B6DDEACE5A4DFFFAFB398516FA2EFA39" box="[1279,1435,1392,1417]" italics="true" pageId="16" pageNumber="167">P. stomachor</emphasis>
Wieser, 1954
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A4DFFF8FF0085A6FF70FDDA" lastPageId="18" lastPageNumber="169" pageId="16" pageNumber="167" type="description">
<paragraph id="841636DC5A4DFFFAFF0085A6FB52F99A" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">
<taxonomicName id="43A94D5F5A4DFFFAFF0085A6FE05FA69" box="[198,432,1472,1497]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4DFFFAFF0085A6FE05FA69" box="[198,432,1472,1497]" italics="true" pageId="16" pageNumber="167">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4DFFFAFE7185A6FDA4FA6A" box="[439,529,1472,1498]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="43A94D5F5A4DFFFAFD6C85A6FCF3FA69" box="[682,838,1472,1497]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="concinnum">
<emphasis id="B6DDEACE5A4DFFFAFD6C85A6FCF3FA69" box="[682,838,1472,1497]" italics="true" pageId="16" pageNumber="167">P. concinnum</emphasis>
</taxonomicName>
(
<collectingCountry id="FCBE764C5A4DFFFAFC9385A6FC2CFA6A" box="[853,921,1472,1498]" name="Chile" pageId="16" pageNumber="167">Chile</collectingCountry>
,
<quantity id="43519B395A4DFFFAFC6385A6FBB8FA6A" box="[933,1037,1472,1498]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="16" pageNumber="167" unit="m" value="75.0" valueMax="80.0" valueMin="70.0">7080 m</quantity>
) with the index c (7.458.12
<emphasis id="B6DDEACE5A4DFFFAFAAB85A6FF08F9B1" italics="true" pageId="16" pageNumber="167">versus</emphasis>
6), longer posterior cylindrical part of the tail (5262%
<emphasis id="B6DDEACE5A4DFFFAFC8A858EFC22F9B1" box="[844,919,1512,1537]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
41%), relative width of the amphidial fovea (5559% c.b.d.
<emphasis id="B6DDEACE5A4DFFFAFE978676FE29F999" box="[337,412,1552,1577]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
40% c.b.d.), lesser number of supplementary organs (1314
<emphasis id="B6DDEACE5A4DFFFAFBA18676FB07F999" box="[1127,1202,1552,1577]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
20).
</paragraph>
<paragraph id="841636DC5A4DFFFAFF00865EFD9BF8AA" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">
<taxonomicName id="43A94D5F5A4DFFFAFF00865EFE06F9E1" box="[198,435,1592,1617]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4DFFFAFF00865EFE06F9E1" box="[198,435,1592,1617]" italics="true" pageId="16" pageNumber="167">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4DFFFAFE7B865EFDAFF9E2" box="[445,538,1592,1618]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="43A94D5F5A4DFFFAFD7A865EFCD6F9E1" box="[700,867,1592,1617]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="corniculata">
<emphasis id="B6DDEACE5A4DFFFAFD7A865EFCD6F9E1" box="[700,867,1592,1617]" italics="true" pageId="16" pageNumber="167">P. corniculata</emphasis>
</taxonomicName>
, a geographically close species (
<collectingCountry id="FCBE764C5A4DFFFAFB29865EFAF2F9E2" box="[1263,1351,1592,1618]" name="Angola" pageId="16" pageNumber="167">Angola</collectingCountry>
Basin,
<quantity id="43519B395A4DFFFAFF518606FE89F9CA" box="[151,316,1632,1658]" metricMagnitude="3" metricUnit="m" metricValue="2.8389999999999995" metricValueMax="3.6149999999999998" metricValueMin="2.063" pageId="16" pageNumber="167" unit="m" value="2839.0" valueMax="3615.0" valueMin="2063.0">20633615 m</quantity>
) with much longer tail (index c 4.006.03
<emphasis id="B6DDEACE5A4DFFFAFC848606FC38F9C9" box="[834,909,1632,1657]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
11.413.5 and c 7.458.12
<emphasis id="B6DDEACE5A4DFFFAFB208606FA84F9C9" box="[1254,1329,1632,1657]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
2.32.5) with longer posterior cylindrical part of the tail (5262%
<emphasis id="B6DDEACE5A4DFFFAFC8186EEFC27F911" box="[839,914,1672,1697]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
21%), more anterior position of the ventral pore at about half the distance from the cephalic apex to the nerve ring
<emphasis id="B6DDEACE5A4DFFFAFC1586D6FBABF979" box="[979,1054,1712,1737]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
just at the nerve ring (5460
<emphasis id="B6DDEACE5A4DFFFAFAB286D6FA36F979" box="[1396,1411,1712,1737]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m and 105
<emphasis id="B6DDEACE5A4DFFFAFF3A86BEFEBEF941" box="[252,267,1752,1777]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m, respectively), bigger amphidial fovea (width 911.5
<emphasis id="B6DDEACE5A4DFFFAFC5986BEFC1BF941" box="[927,942,1752,1777]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m
<emphasis id="B6DDEACE5A4DFFFAFC0986BEFBAFF941" box="[975,1050,1752,1777]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
7
<emphasis id="B6DDEACE5A4DFFFAFBFE86BEFBF2F941" box="[1080,1095,1752,1777]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m), lesser number of supplementary organs (1314
<emphasis id="B6DDEACE5A4DFFFAFE6B8766FE4DF8A9" box="[429,504,1792,1817]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
20).
</paragraph>
<paragraph id="841636DC5A4DFFFAFF00874EFC6EF852" blockId="16.[151,1437,152,2018]" pageId="16" pageNumber="167">
The original description of
<emphasis id="B6DDEACE5A4DFFFAFDC0874EFD37F8F1" box="[518,642,1832,1857]" italics="true" pageId="16" pageNumber="167">P. m i r a b i l e</emphasis>
(no locality indicated by Cobb) misses some significant details and any indication on locality, hence the relation of the new species to
<emphasis id="B6DDEACE5A4DFFFAFC7D8736FB8CF8D9" box="[955,1081,1872,1897]" italics="true" pageId="16" pageNumber="167">P. m i r a b i l e</emphasis>
remains unclear. However
<taxonomicName id="43A94D5F5A4DFFFAFA408736FED0F821" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4DFFFAFA408736FED0F821" italics="true" pageId="16" pageNumber="167">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4DFFFAFEA8871EFE7EF822" box="[366,459,1912,1938]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
is distinctly separated from
<taxonomicName id="43A94D5F5A4DFFFAFCE3871EFC1DF821" box="[805,936,1912,1937]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="167" phylum="Nematoda" rank="species" species="mirabile">
<emphasis id="B6DDEACE5A4DFFFAFCE3871EFC1DF821" box="[805,936,1912,1937]" italics="true" pageId="16" pageNumber="167">P. mirabile</emphasis>
</taxonomicName>
in the body length (9111203
<emphasis id="B6DDEACE5A4DFFFAFAD9871EFA9BF821" box="[1311,1326,1912,1937]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m
<emphasis id="B6DDEACE5A4DFFFAFA97871EFA29F821" box="[1361,1436,1912,1937]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
&gt;1800
<emphasis id="B6DDEACE5A4DFFFAFF2D87C6FF4FF809" box="[235,250,1952,1977]" italics="true" pageId="16" pageNumber="167">µ</emphasis>
m), smooth versus jonted setae of the cephalic ring, smaller amphidial fovea (5559% c.b.d.
<emphasis id="B6DDEACE5A4DFFFAFA9787C6FA29F809" box="[1361,1436,1952,1977]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
93% c.b.d.), lesser number of supplementary organs (1314
<emphasis id="B6DDEACE5A4DFFFAFC9C87AEFC10F851" box="[858,933,1992,2017]" italics="true" pageId="16" pageNumber="167">versus</emphasis>
20).
</paragraph>
<caption id="D0D666545A4CFFFBFF5187E1FE7AF80D" httpUri="https://zenodo.org/record/183784/files/figure.png" pageId="17" pageNumber="168" targetBox="[164,1436,189,1913]" targetPageId="17">
<paragraph id="841636DC5A4CFFFBFF5187E1FE7AF80D" blockId="17.[151,1437,1927,1981]" pageId="17" pageNumber="168">
<emphasis id="B6DDEACE5A4CFFFBFF5187E1FEBDF82C" bold="true" box="[151,264,1927,1948]" pageId="17" pageNumber="168">FIGURE 9</emphasis>
.
<taxonomicName id="43A94D5F5A4CFFFBFED087EEFD8BF82D" box="[278,574,1928,1949]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="17" pageNumber="168" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4CFFFBFED087EEFD8BF82D" box="[278,574,1928,1949]" italics="true" pageId="17" pageNumber="168">Pomponema proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4CFFFBFD8187EEFD20F82D" box="[583,661,1928,1949]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
, details.
<emphasis id="B6DDEACE5A4CFFFBFD2B87E1FD4BF82C" bold="true" box="[749,766,1927,1948]" pageId="17" pageNumber="168">A</emphasis>
: male N1 (holotype), entire;
<emphasis id="B6DDEACE5A4CFFFBFBE787E1FB84F82C" bold="true" box="[1057,1073,1927,1948]" pageId="17" pageNumber="168">B</emphasis>
: male N3 (paratype), cloacal region. Scale bars: A 100
<emphasis id="B6DDEACE5A4CFFFBFE8087CEFEE7F80D" box="[326,338,1960,1981]" italics="true" pageId="17" pageNumber="168">µ</emphasis>
m; B 10
<emphasis id="B6DDEACE5A4CFFFBFE6187CEFE06F80D" box="[423,435,1960,1981]" italics="true" pageId="17" pageNumber="168">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="841636DC5A4FFFF8FF0080FEFBD3FECA" blockId="18.[151,1437,152,618]" pageId="18" pageNumber="169">
<taxonomicName id="43A94D5F5A4FFFF8FF0080FEFE2AFF01" box="[198,415,152,177]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="169" phylum="Nematoda" rank="species" species="multipapillatum">
<emphasis id="B6DDEACE5A4FFFF8FF0080FEFE2AFF01" box="[198,415,152,177]" italics="true" pageId="18" pageNumber="169">P. multipapillatum</emphasis>
</taxonomicName>
is known from two thorough descriptions of Filipjev (1922, Black Sea, sublittoral silt) and
<bibRefCitation id="E0384B2D5A4FFFF8FF0F80A6FCEDFF6A" author="Lorenzen" box="[201,856,192,218]" pageId="18" pageNumber="173" refString="Lorenzen, S. (1972) Die Nematodenfauna im Verklappungsgebiet fur Industrieabwasser nordwestlich von Helgoland. III. Cyatholaimidae, mit einer Revision von Pomponema Cobb 1917. Veroffentlichungen des Instituts fur Meeresforschungen in Bremerhaven, 13, 285 - 306." type="journal article" year="1972" yearSuffix="m">
Lorenzen (1972,
<collectingCountry id="FCBE764C5A4FFFF8FE5480A6FDB9FF6A" box="[402,524,192,218]" name="Germany" pageId="18" pageNumber="169">Helgoland</collectingCountry>
in the North Sea,
<quantity id="43519B395A4FFFF8FD2580A6FCFAFF6A" box="[739,847,192,218]" metricMagnitude="1" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="18" pageNumber="169" unit="m" value="32.0" valueMax="34.0" valueMin="30.0">3034 m</quantity>
)
</bibRefCitation>
.
<taxonomicName id="43A94D5F5A4FFFF8FCA480A6FBF8FF69" box="[866,1101,192,217]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="169" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4FFFF8FCA480A6FBF8FF69" box="[866,1101,192,217]" italics="true" pageId="18" pageNumber="169">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4FFFF8FB9280A6FB1AFF6A" box="[1108,1199,192,218]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="43A94D5F5A4FFFF8FA8F80A6FE92FEB1" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="169" phylum="Nematoda" rank="species" species="multipapillatum">
<emphasis id="B6DDEACE5A4FFFF8FA8F80A6FE92FEB1" italics="true" pageId="18" pageNumber="169">P. multipapillatum</emphasis>
</taxonomicName>
with much longer tail (index c 4.006.03
<emphasis id="B6DDEACE5A4FFFF8FCE0808EFCC4FEB1" box="[806,881,232,257]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
7.310 and c 7.458.12
<emphasis id="B6DDEACE5A4FFFF8FB6E808EFB46FEB1" box="[1192,1267,232,257]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
3.74.5) with longer posterior cylindrical part of the tail (5262%
<emphasis id="B6DDEACE5A4FFFF8FD3E8176FCF6FE99" box="[760,835,272,297]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
2844%), position of the ventral pore (respectively 5460
<emphasis id="B6DDEACE5A4FFFF8FF21815EFF43FEE1" box="[231,246,312,337]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m and 1538
<emphasis id="B6DDEACE5A4FFFF8FE5F815EFE1DFEE1" box="[409,424,312,337]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m from the cephalic apex to the ventral pore), shorter setae of the cephalic ring (24.5
<emphasis id="B6DDEACE5A4FFFF8FF518106FF13FEC9" box="[151,166,352,377]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m
<emphasis id="B6DDEACE5A4FFFF8FF008106FEA4FEC9" box="[198,273,352,377]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
48
<emphasis id="B6DDEACE5A4FFFF8FE8D8106FEEFFEC9" box="[331,346,352,377]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), lesser number of supplementary organs (1314
<emphasis id="B6DDEACE5A4FFFF8FC7F8106FBB1FEC9" box="[953,1028,352,377]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
1519).
</paragraph>
<paragraph id="841636DC5A4FFFF8FF0081EEFF70FDDA" blockId="18.[151,1437,152,618]" pageId="18" pageNumber="169">
<taxonomicName id="43A94D5F5A4FFFF8FF0081EEFE04FE11" box="[198,433,392,417]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="169" phylum="Nematoda" rank="species" species="proximamphidum" status="sp. nov.">
<emphasis id="B6DDEACE5A4FFFF8FF0081EEFE04FE11" box="[198,433,392,417]" italics="true" pageId="18" pageNumber="169">P. proximamphidum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A4FFFF8FE7F81EEFDA1FE12" box="[441,532,392,418]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="43A94D5F5A4FFFF8FD6981EEFCFDFE11" box="[687,840,392,417]" class="Adenophorea" family="Cyatholaimidae" genus="Pomponema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="169" phylum="Nematoda" rank="species" species="stomachor">
<emphasis id="B6DDEACE5A4FFFF8FD6981EEFCFDFE11" box="[687,840,392,417]" italics="true" pageId="18" pageNumber="169">P. stomachor</emphasis>
</taxonomicName>
(
<collectingCountry id="FCBE764C5A4FFFF8FC9E81EEFC29FE12" box="[856,924,392,418]" name="Chile" pageId="18" pageNumber="169">Chile</collectingCountry>
, tidal exposed sand) by smaller body (911 1203
<emphasis id="B6DDEACE5A4FFFF8FF1B81D6FF59FE79" box="[221,236,432,457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m
<emphasis id="B6DDEACE5A4FFFF8FED681D6FEEEFE79" box="[272,347,432,457]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
19502170
<emphasis id="B6DDEACE5A4FFFF8FE3381D6FDB1FE79" box="[501,516,432,457]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), longer tail (index c 46.03
<emphasis id="B6DDEACE5A4FFFF8FC4A81D6FC62FE79" box="[908,983,432,457]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
7.18, c 7.458.12
<emphasis id="B6DDEACE5A4FFFF8FB1F81D6FA91FE79" box="[1241,1316,432,457]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
7), much shorter setae of the cephalic crown (25.5
<emphasis id="B6DDEACE5A4FFFF8FD5381BEFD11FE41" box="[661,676,472,497]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m
<emphasis id="B6DDEACE5A4FFFF8FD0081BEFCA4FE41" box="[710,785,472,497]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
318
<emphasis id="B6DDEACE5A4FFFF8FC9881BEFCD8FE41" box="[862,877,472,497]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), smaller amphidial fovea (911.5
<emphasis id="B6DDEACE5A4FFFF8FAED81BEFAC3FE41" box="[1323,1398,472,497]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
20
<emphasis id="B6DDEACE5A4FFFF8FF518266FF13FDA9" box="[151,166,512,537]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), shorter distance from the cephalic apex to the ventral excretory pore (width 5460
<emphasis id="B6DDEACE5A4FFFF8FB758266FB77FDA9" box="[1203,1218,512,537]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m
<emphasis id="B6DDEACE5A4FFFF8FB258266FA9BFDA9" box="[1251,1326,512,537]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
110150
<emphasis id="B6DDEACE5A4FFFF8FF51824EFF13FDF1" box="[151,166,552,577]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), smaller spicules (3344.5
<emphasis id="B6DDEACE5A4FFFF8FDCC824EFDACFDF1" box="[522,537,552,577]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m
<emphasis id="B6DDEACE5A4FFFF8FDFF824EFD31FDF1" box="[569,644,552,577]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
75
<emphasis id="B6DDEACE5A4FFFF8FD76824EFD0AFDF1" box="[688,703,552,577]" italics="true" pageId="18" pageNumber="169">µ</emphasis>
m), lesser number of supplementary organs (1314
<emphasis id="B6DDEACE5A4FFFF8FAD8824EFADCFDF1" box="[1310,1385,552,577]" italics="true" pageId="18" pageNumber="169">versus</emphasis>
15 19).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,501 @@
<document id="B1CFB74F5A1FB98A3C9DA9D24D58C628" ID-DOI="10.5281/zenodo.183775" ID-GBIF-Dataset="1208b1c5-c358-4a24-9ee8-7246d1494b9a" ID-ISSN="1175­5326" ID-Zenodo-Dep="183775" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459928395076" checkinUser="plazi" docAuthor="Tchesunov, Alexei V." docDate="2008" docId="0C0087CA5A5BFFE6FF518650FD2BFBFA" docLanguage="en" docName="zt01866p174.pdf" docOrigin="Zootaxa 1866" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Paracyatholaimus diva Tchesunov, 2008, sp. nov." docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="F039FFB25A5DFFEAFFC68066FFB5FFB0" masterDocTitle="Three new species of free­living nematodes from the South­East Atlantic Abyss (DIVA I Expedition) *" masterLastPageNumber="174" masterPageNumber="151" pageNumber="157" updateTime="1698229710615" updateUser="plazi">
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<mods:title id="49B08089A20F5F898F39EE2DBBF7FE75">Three new species of free­living nematodes from the South­East Atlantic Abyss (DIVA I Expedition) *</mods:title>
</mods:titleInfo>
<mods:name id="6C9DFE9C63D37DEEDD8BFC8CE1310333" type="personal">
<mods:role id="2E5C5988CFF37B7962ECC93D83E6D296">
<mods:roleTerm id="E18A2C282A0F54FBD6B8A08D2BD53154">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="414CF6747EE2C63229A2467BC0FF494F">Tchesunov, Alexei V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5A549BF9AD058A30837055487C22A782">text</mods:typeOfResource>
<mods:relatedItem id="12C8E983B87FB7E922EB45996B5DB9E9" type="host">
<mods:titleInfo id="A6EF8BDD2391D628B96E152B41F2B8AA">
<mods:title id="91BC89A02A68C432D0E36D3D417E4C08">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6192C2720AA31ECD28135E26CA342ECC">
<mods:date id="EAD62E49F2CA5B7BB69141D686D68346">2008</mods:date>
<mods:detail id="232D9779D1336E1E36B31B1938838F05" type="volume">
<mods:number id="B17F52A8FCEB6F59A958D3EFDDA117E9">1866</mods:number>
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<mods:start id="6F9FF8F6D2A9FACCA27EEF0D70E7ADBC">151</mods:start>
<mods:end id="624B4F21B88FC67AD6A4DAF2D5643251">174</mods:end>
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<mods:classification id="F93CE71F07E74F593DC7F205E3AD3A15">journal article</mods:classification>
<mods:identifier id="3584BCF04D2B0A4A82F2B79CC08B3052" type="DOI">10.5281/zenodo.183775</mods:identifier>
<mods:identifier id="20ED95AAA4F7280C521DB9E5BA6529F3" type="GBIF-Dataset">1208b1c5-c358-4a24-9ee8-7246d1494b9a</mods:identifier>
<mods:identifier id="5B40C39CA6EB0099A1CEBBC65AFAAECD" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="AA19E9E434F1C365515A485633889949" type="Zenodo-Dep">183775</mods:identifier>
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<treatment id="0C0087CA5A5BFFE6FF518650FD2BFBFA" ID-DOI="http://doi.org/10.5281/zenodo.6231736" ID-GBIF-Taxon="119374537" ID-Zenodo-Dep="6231736" LSID="urn:lsid:plazi:treatment:0C0087CA5A5BFFE6FF518650FD2BFBFA" httpUri="http://treatment.plazi.org/id/0C0087CA5A5BFFE6FF518650FD2BFBFA" lastPageId="12" lastPageNumber="163" pageId="6" pageNumber="157">
<subSubSection id="CCB365575A5BFFECFF518650FF49F9C3" pageId="6" pageNumber="157" type="nomenclature">
<paragraph id="841636DC5A5BFFECFF518650FDA4F9E0" blockId="6.[151,529,1590,1651]" box="[151,529,1590,1616]" pageId="6" pageNumber="157">
<heading id="DF5E81B05A5BFFECFF518650FDA4F9E0" bold="true" box="[151,529,1590,1616]" fontSize="11" level="1" pageId="6" pageNumber="157" reason="1">
<emphasis id="B6DDEACE5A5BFFECFF518650FDA4F9E0" bold="true" box="[151,529,1590,1616]" pageId="6" pageNumber="157">
<taxonomicName id="43A94D5F5A5BFFECFF518650FE18F9E0" box="[151,429,1590,1616]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="6" pageNumber="157" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A5BFFECFF518650FE18F9E0" bold="true" box="[151,429,1590,1616]" italics="true" pageId="6" pageNumber="157">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A5BFFECFE758650FDA4F9E0" box="[435,529,1590,1616]" pageId="6" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="841636DC5A5BFFECFF51863FFF49F9C3" blockId="6.[151,529,1590,1651]" box="[151,252,1625,1651]" pageId="6" pageNumber="157">
<figureCitation id="1C922A595A5BFFECFF51863FFF49F9C3" box="[151,252,1625,1651]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="8.[151,244,2005,2026]" captionStartId-1="9.[151,244,1438,1459]" captionStartId-2="10.[151,244,1870,1891]" captionStartId-3="11.[151,244,1769,1790]" captionTargetBox-0="[184,1372,428,2004]" captionTargetBox-1="[185,1400,185,1414]" captionTargetBox-2="[185,1393,189,1846]" captionTargetBox-3="[152,1205,183,1768]" captionTargetId-0="figure@8.[151,1436,402,2004]" captionTargetId-1="figure@9.[151,1436,171,1440]" captionTargetId-2="figure@10.[151,1436,170,1876]" captionTargetId-3="figure@11.[151,1436,162,1785]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURE 3. Paracyatholaimus diva sp. nov., entire. A: male N 1 (holotype); B: female (paratype). Scale bars: 100 µ m." captionText-1="FIGURE 4. Paracyatholaimus diva sp. nov., anterior ends. A: male N 1 (holotype); B: female (paratype). Scale bars: 10 µ m." captionText-2="FIGURE 5. Paracyatholaimus diva sp. nov., details. A: male N 1 (holotype), anterior body; B: male N 1 (holotype), posterior end; C: female (paratype), posterior end. Scale bars: 50 µ m." captionText-3="FIGURE 6. Paracyatholaimus diva sp. nov., cloacal region of the male N 1 (holotype). Scale bar: 10 µ m." httpUri-0="https://zenodo.org/record/183778/files/figure.png" httpUri-1="https://zenodo.org/record/183779/files/figure.png" httpUri-2="https://zenodo.org/record/183780/files/figure.png" httpUri-3="https://zenodo.org/record/183781/files/figure.png" pageId="6" pageNumber="157">Figs 36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5BFFECFF5186CFFDCDF8D6" pageId="6" pageNumber="157" type="materials_examined">
<paragraph id="841636DC5A5BFFECFF5186CFFD94F8A3" blockId="6.[151,1437,1704,1974]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF5186CFFE8DF972" box="[151,312,1705,1730]" italics="true" pageId="6" pageNumber="157">
<typeStatus id="5B12887E5A5BFFECFF5186CFFF7BF972" box="[151,206,1705,1730]" pageId="6" pageNumber="157">Type</typeStatus>
material
</emphasis>
:
<emphasis id="B6DDEACE5A5BFFECFE8686CEFE09F972" bold="true" box="[320,444,1704,1730]" pageId="6" pageNumber="157">
<typeStatus id="5B12887E5A5BFFECFE8686CEFE02F972" box="[320,439,1704,1730]" pageId="6" pageNumber="157" type="holotype">Holotype</typeStatus>
:
</emphasis>
male 1, two
<typeStatus id="5B12887E5A5BFFECFD9E86CFFD08F973" box="[600,701,1705,1731]" pageId="6" pageNumber="157" type="paratype">paratype</typeStatus>
males 2 &amp; 3 and one
<typeStatus id="5B12887E5A5BFFECFC0286CFFB9CF973" box="[964,1065,1705,1731]" pageId="6" pageNumber="157" type="paratype">paratype</typeStatus>
female mounted in glycerin on glass slides.
<typeStatus id="5B12887E5A5BFFECFEEA86B7FED3F95B" box="[300,358,1745,1771]" pageId="6" pageNumber="157">Type</typeStatus>
specimens deposited in the
<taxonomicName id="43A94D5F5A5BFFECFD7586B7FD4DF95B" box="[691,760,1745,1771]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="6" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<collectionCode id="E2B8AE195A5BFFECFD7586B7FD4DF95B" box="[691,760,1745,1771]" pageId="6" pageNumber="157">DIVA</collectionCode>
</taxonomicName>
nematode collection DZMB­Senckenberg, Südstrand 44, 26382 Wilhelmshaven,
<collectingCountry id="FCBE764C5A5BFFECFE6B869FFDA9F8A3" box="[429,540,1785,1811]" name="Germany" pageId="6" pageNumber="157">Germany</collectingCountry>
.
</paragraph>
<paragraph id="841636DC5A5BFFECFF008747FDCDF8D6" blockId="6.[151,1437,1704,1974]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF008747FEE8F88A" box="[198,349,1825,1850]" italics="true" pageId="6" pageNumber="157">
<typeStatus id="5B12887E5A5BFFECFF008747FF48F88A" box="[198,253,1825,1850]" pageId="6" pageNumber="157">Type</typeStatus>
locality
</emphasis>
:
<taxonomicName id="43A94D5F5A5BFFECFEAB8747FE07F88B" box="[365,434,1825,1851]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="6" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<collectionCode id="E2B8AE195A5BFFECFEAB8747FE07F88B" box="[365,434,1825,1851]" pageId="6" pageNumber="157">DIVA</collectionCode>
</taxonomicName>
I, Meteor 48/1, Station 325/4, multicorer 8: South­east Atlantic Ocean,
<geoCoordinate id="E19D501B5A5BFFECFACD8747FAC8F88B" box="[1291,1405,1825,1851]" direction="south" orientation="latitude" pageId="6" pageNumber="157" precision="92" value="-19.971666">19°58.3'S</geoCoordinate>
&amp;
<geoCoordinate id="E19D501B5A5BFFECFF51872AFEA9F8D6" box="[151,284,1868,1894]" direction="east" orientation="longitude" pageId="6" pageNumber="157" precision="92" value="2.9966667">002°59.8'E</geoCoordinate>
, depth
<quantity id="43519B395A5BFFECFEB7872AFE78F8D6" box="[369,461,1868,1894]" metricMagnitude="3" metricUnit="m" metricValue="5.45" pageId="6" pageNumber="157" unit="m" value="5450.0">5450 m</quantity>
.
<date id="F017101C5A5BFFECFE1E872AFDC1F8D6" box="[472,628,1868,1894]" pageId="6" pageNumber="157" value="2000-07-14">14 July 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5BFFECFF008712FDADF806" pageId="6" pageNumber="157" type="etymology">
<paragraph id="841636DC5A5BFFECFF008712FC77F83E" blockId="6.[151,1437,1704,1974]" box="[198,962,1908,1934]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF008712FEF6F83D" box="[198,323,1908,1933]" italics="true" pageId="6" pageNumber="157">Etymology</emphasis>
: the species name is derived from the acronym
<taxonomicName id="43A94D5F5A5BFFECFCB38712FC0AF83E" box="[885,959,1908,1934]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="6" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
.
</paragraph>
<paragraph id="841636DC5A5BFFECFF0087FAFDADF806" blockId="6.[151,1437,1704,1974]" box="[198,536,1948,1974]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF0087FAFE1EF805" box="[198,427,1948,1973]" italics="true" pageId="6" pageNumber="157">Morphometric data</emphasis>
: Table 2.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5AFFE1FF5180FFFBAEF82B" lastPageId="11" lastPageNumber="162" pageId="7" pageNumber="158" type="description">
<paragraph id="841636DC5A5AFFEDFF5180FFFCACFF02" blockId="7.[151,793,152,178]" box="[151,793,152,178]" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF5180FFFEB6FF01" bold="true" box="[151,259,153,177]" pageId="7" pageNumber="158">TABLE 2</emphasis>
. Morphometry of
<taxonomicName id="43A94D5F5A5AFFEDFE0E80FCFD75FF01" box="[456,704,154,177]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="7" pageNumber="158" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A5AFFEDFE0E80FCFD75FF01" box="[456,704,154,177]" italics="true" pageId="7" pageNumber="158">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A5AFFEDFD0180FFFCACFF02" box="[711,793,152,178]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="841636DC5A5AFFEDFEC380B1FF5FF96B" blockId="7.[261,1033,215,236]" lastBlockId="7.[151,1437,1609,2035]" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFEC380B1FBBCFF5C" bold="true" box="[261,1033,215,236]" pageId="7" pageNumber="158">Label Specimens &amp; labels</emphasis>
<emphasis id="B6DDEACE5A5AFFEDFF00862FFEE4F9D2" box="[198,337,1609,1634]" italics="true" pageId="7" pageNumber="158">Description</emphasis>
. Body cylindrical. Cuticle transversally annulated and punctated. The dots (punctations) about the same size along the body and closely arranged in transversal rows. No clear lateral differentiation in size and position of the dots. The dots become a bit more distinct and convex to the amphid and to the anal region.
</paragraph>
<paragraph id="841636DC5A5AFFEDFF6381F0FA85FA4E" pageId="7" pageNumber="158">
<table id="F6A9C47C5A5A0015FF638164FACAFA4E" box="[165,1407,258,1534]" gridcols="5" gridrows="8" pageId="7" pageNumber="158">
<tr id="3A99349E5A5A0015FF638164FACAFE66" box="[165,1407,258,470]" gridrow="0" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF638164FEE4FE66" box="[165,337,258,470]" gridcol="0" gridrow="0" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF6381F0FF04FE66" bold="true" box="[165,181,406,470]" pageId="7" pageNumber="158">L a</emphasis>
</th>
<th id="79485DE25A5A0015FE148164FDCCFE66" box="[466,633,258,470]" gridcol="1" gridrow="0" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFE2A8164FDCCFE31" bold="true" pageId="7" pageNumber="158">
1 (holotype) Box B N 55(6)
<taxonomicName id="43A94D5F5A5AFFEDFE15812AFDA5FED1" box="[467,528,332,353]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="7" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1, st. 325/4, MuC 8
</emphasis>
936 24.6
</th>
<th id="79485DE25A5A0015FD128164FCCEFE66" box="[724,891,258,470]" gridcol="2" gridrow="0" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFD2A8164FCCEFE31" bold="true" pageId="7" pageNumber="158">
2 (paratype) Box B N 55(7)
<taxonomicName id="43A94D5F5A5AFFEDFD13812AFCA7FED1" box="[725,786,332,353]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="7" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1, st. 325/4, MuC 8
</emphasis>
1120 23.8
</th>
<th id="79485DE25A5A0015FC1C8164FB34FE66" box="[986,1153,258,470]" gridcol="3" gridrow="0" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFC378164FB34FE31" bold="true" pageId="7" pageNumber="158">
3 (paratype) Box B N 56(4)
<taxonomicName id="43A94D5F5A5AFFEDFC1D812AFBADFED1" box="[987,1048,332,353]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="7" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1, st. 325/4, MuC 8
</emphasis>
1176 28
</th>
<th id="79485DE25A5A0015FB1E8164FACAFE66" box="[1240,1407,258,470]" gridcol="4" gridrow="0" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFB3E8164FACAFE31" bold="true" pageId="7" pageNumber="158">
(paratype) Box B N 59(5)
<taxonomicName id="43A94D5F5A5AFFEDFB1E812AFAA0FED1" box="[1240,1301,332,353]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="7" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">DIVA</taxonomicName>
I, M 48/1, st. 325/4, MuC 8
</emphasis>
1147 26.1
</th>
</tr>
<tr id="3A99349E5A5A0015FF63818AFACAFD30" box="[165,1407,492,640]" gridrow="1" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF63818AFEE4FD30" box="[165,337,492,640]" gridcol="0" gridrow="1" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF63818AFF4AFD30" bold="true" pageId="7" pageNumber="158">b c V (%) diam.c.s.</emphasis>
</th>
<td id="79485DE25A5A0015FE14818AFDCCFD30" box="[466,633,492,640]" gridcol="1" gridrow="1" pageId="7" pageNumber="158">6.65 7.45 ­ 17</td>
<td id="79485DE25A5A0015FD12818AFCCEFD30" box="[724,891,492,640]" gridcol="2" gridrow="1" pageId="7" pageNumber="158">7.00 8.75 ­ ~17</td>
<td id="79485DE25A5A0015FC1C818AFB34FD30" box="[986,1153,492,640]" gridcol="3" gridrow="1" pageId="7" pageNumber="158">7.35 8.86 ­ 19</td>
<td id="79485DE25A5A0015FB1E818AFACAFD30" box="[1240,1407,492,640]" gridcol="4" gridrow="1" pageId="7" pageNumber="158">6.23 8.64 46 18</td>
</tr>
<tr id="3A99349E5A5A0015FF6382F0FACAFD65" box="[165,1407,662,725]" gridrow="2" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF6382F0FEE4FD65" box="[165,337,662,725]" gridcol="0" gridrow="2" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF6382F0FEB6FD65" bold="true" pageId="7" pageNumber="158">diam.am. diam.n.r.</emphasis>
</th>
<td id="79485DE25A5A0015FE1482F0FDCCFD65" box="[466,633,662,725]" gridcol="1" gridrow="2" pageId="7" pageNumber="158">23.5 34</td>
<td id="79485DE25A5A0015FD1282F0FCCEFD65" box="[724,891,662,725]" gridcol="2" gridrow="2" pageId="7" pageNumber="158">? 31</td>
<td id="79485DE25A5A0015FC1C82F0FB34FD65" box="[986,1153,662,725]" gridcol="3" gridrow="2" pageId="7" pageNumber="158">22 39</td>
<td id="79485DE25A5A0015FB1E82F0FACAFD65" box="[1240,1407,662,725]" gridcol="4" gridrow="2" pageId="7" pageNumber="158">30 32</td>
</tr>
<tr id="3A99349E5A5A0015FF63828DFACAFCB0" box="[165,1407,747,768]" gridrow="3" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF63828DFEE4FCB0" box="[165,337,747,768]" gridcol="0" gridrow="3" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF63828DFF49FCB0" bold="true" box="[165,252,747,768]" pageId="7" pageNumber="158">diam.ca.</emphasis>
</th>
<td id="79485DE25A5A0015FE14828DFDCCFCB0" box="[466,633,747,768]" gridcol="1" gridrow="3" pageId="7" pageNumber="158">38</td>
<td id="79485DE25A5A0015FD12828DFCCEFCB0" box="[724,891,747,768]" gridcol="2" gridrow="3" pageId="7" pageNumber="158">40</td>
<td id="79485DE25A5A0015FC1C828DFB34FCB0" box="[986,1153,747,768]" gridcol="3" gridrow="3" pageId="7" pageNumber="158">41</td>
<td id="79485DE25A5A0015FB1E828DFACAFCB0" box="[1240,1407,747,768]" gridcol="4" gridrow="3" pageId="7" pageNumber="158">39</td>
</tr>
<tr id="3A99349E5A5A0015FF638370FACAFC4F" box="[165,1407,790,1023]" gridrow="4" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF638370FEE4FC4F" box="[165,337,790,1023]" gridcol="0" gridrow="4" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF638326FEA5FC4F" bold="true" pageId="7" pageNumber="158">a.d. c.s.+o.l.s. c.s.+o.l.s. (%) am.w. am.w. (%)</emphasis>
</th>
<td id="79485DE25A5A0015FE148370FDCCFC4F" box="[466,633,790,1023]" gridcol="1" gridrow="4" pageId="7" pageNumber="158">38 33 3+3.5 17.5+20.5 12.5 53</td>
<td id="79485DE25A5A0015FD128370FCCEFC4F" box="[724,891,790,1023]" gridcol="2" gridrow="4" pageId="7" pageNumber="158">47 39????</td>
<td id="79485DE25A5A0015FC1C8370FB34FC4F" box="[986,1153,790,1023]" gridcol="3" gridrow="4" pageId="7" pageNumber="158">42 40 3.5+4 18.5+21 14 64</td>
<td id="79485DE25A5A0015FB1E8370FACAFC4F" box="[1240,1407,790,1023]" gridcol="4" gridrow="4" pageId="7" pageNumber="158">44 30 3.5+5 19.5+27.5 8 26.5</td>
</tr>
<tr id="3A99349E5A5A0015FF638473FACAFB4E" box="[165,1407,1045,1278]" gridrow="5" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF638473FEE4FB4E" box="[165,337,1045,1278]" gridcol="0" gridrow="5" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF638473FEBBFB4E" bold="true" pageId="7" pageNumber="158">dis.am. st.w. st.l. cheil.l. dors.t.l. spic.chord</emphasis>
</th>
<td id="79485DE25A5A0015FE148473FDCCFB4E" box="[466,633,1045,1278]" gridcol="1" gridrow="5" pageId="7" pageNumber="158">11.5 7 16.5 5 12 34</td>
<td id="79485DE25A5A0015FD128473FCCEFB4E" box="[724,891,1045,1278]" gridcol="2" gridrow="5" pageId="7" pageNumber="158">????? 36</td>
<td id="79485DE25A5A0015FC1C8473FB34FB4E" box="[986,1153,1045,1278]" gridcol="3" gridrow="5" pageId="7" pageNumber="158">11 8 15 3.5 11 34</td>
<td id="79485DE25A5A0015FB1E8473FACAFB4E" box="[1240,1407,1045,1278]" gridcol="4" gridrow="5" pageId="7" pageNumber="158">4 8 18 3.5 12 ­</td>
</tr>
<tr id="3A99349E5A5A0015FF638572FACAFAE3" box="[165,1407,1300,1363]" gridrow="6" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF638572FEE4FAE3" box="[165,337,1300,1363]" gridcol="0" gridrow="6" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF638572FF6AFAE3" bold="true" pageId="7" pageNumber="158">spic.arc gub.l.</emphasis>
</th>
<td id="79485DE25A5A0015FE148572FDCCFAE3" box="[466,633,1300,1363]" gridcol="1" gridrow="6" pageId="7" pageNumber="158">45 33</td>
<td id="79485DE25A5A0015FD128572FCCEFAE3" box="[724,891,1300,1363]" gridcol="2" gridrow="6" pageId="7" pageNumber="158">51 29</td>
<td id="79485DE25A5A0015FC1C8572FB34FAE3" box="[986,1153,1300,1363]" gridcol="3" gridrow="6" pageId="7" pageNumber="158">46.5 31</td>
<td id="79485DE25A5A0015FB1E8572FACAFAE3" box="[1240,1407,1300,1363]" gridcol="4" gridrow="6" pageId="7" pageNumber="158">­ ­</td>
</tr>
<tr id="3A99349E5A5A0015FF63850FFACAFA4E" box="[165,1407,1385,1534]" gridrow="7" pageId="7" pageNumber="158">
<th id="79485DE25A5A0015FF63850FFEE4FA4E" box="[165,337,1385,1534]" gridcol="0" gridrow="7" pageId="7" pageNumber="158">
<emphasis id="B6DDEACE5A5AFFEDFF63850FFEBBFA4D" bold="true" pageId="7" pageNumber="158">suppl.l. c dist.tail part (%) termin sp.</emphasis>
</th>
<td id="79485DE25A5A0015FE14850FFDCCFA4E" box="[466,633,1385,1534]" gridcol="1" gridrow="7" pageId="7" pageNumber="158">3 3.14 59 7.5</td>
<td id="79485DE25A5A0015FD12850FFCCEFA4E" box="[724,891,1385,1534]" gridcol="2" gridrow="7" pageId="7" pageNumber="158">? 3.02 41 8</td>
<td id="79485DE25A5A0015FC1C850FFB34FA4E" box="[986,1153,1385,1534]" gridcol="3" gridrow="7" pageId="7" pageNumber="158">? 3.43 45 7</td>
<td id="79485DE25A5A0015FB1E850FFACAFA4E" box="[1240,1407,1385,1534]" gridcol="4" gridrow="7" pageId="7" pageNumber="158">­ 5.08 49.8?</td>
</tr>
</table>
</paragraph>
<paragraph id="841636DC5A5AFFEDFF00868FFED3F87B" blockId="7.[151,1437,1609,2035]" pageId="7" pageNumber="158">Anterior end truncate. Mouth opening surrounded by unclear conical lips with hardly visible inner labial sensilla. Outer labial and cephalic sensilla constitute a jointed circle of 6+4 short setae. Four cephalic setae slightly smaller than six outer labial setae. Both outer labial and cephalic sensilla seem to be two­jointed, with the proximal joint thicker and at least twice as long as the distal joint. However, the jointed setae are often difficult to recognize in this species. The jointed nature of the outer labial setae is more evident in the female than in the males.</paragraph>
<paragraph id="841636DC5A5AFFE2FF0087BFFBE0FEB2" blockId="7.[151,1437,1609,2035]" lastBlockId="8.[151,1437,152,378]" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158">Amphidial fovea large, round or slightly transversally oval in outline, multispirally coiled with about six turns in the males and four turns in the female, situated at a distance from the apex. Amphidial fovea of the female notably smaller than that of the male. Cervical setae about four in number, situated laterally posterior to the amphid. There are very few small setae scarcely distributed along the body.</paragraph>
<paragraph id="841636DC5A55FFE2FF008176FB3EFECA" blockId="8.[151,1437,152,378]" pageId="8" pageNumber="159">
Cuticular pores indistinct in the pharyngeal region but become very obviously crater­shaped more posteriorly Cuticular pores arranged in two sublateral rows from the level of the cardia posteriad. There are 21 lateroventral pores and ~37 laterodorsal pores on the left body side of the
<typeStatus id="5B12887E5A55FFE2FC1A8106FBF6FECA" box="[988,1091,352,378]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
male.
</paragraph>
<caption id="D0D666545A55FFE2FF5187B3FAB2F85B" box="[151,1287,2005,2027]" httpUri="https://zenodo.org/record/183778/files/figure.png" pageId="8" pageNumber="159" targetBox="[184,1372,428,2004]" targetPageId="8">
<paragraph id="841636DC5A55FFE2FF5187B3FAB2F85B" blockId="8.[151,1287,2005,2027]" box="[151,1287,2005,2027]" pageId="8" pageNumber="159">
<emphasis id="B6DDEACE5A55FFE2FF5187B3FEB3F85A" bold="true" box="[151,262,2005,2026]" pageId="8" pageNumber="159">FIGURE 3</emphasis>
.
<taxonomicName id="43A94D5F5A55FFE2FED487B0FE44F85B" box="[274,497,2006,2027]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="8" pageNumber="159" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A55FFE2FED487B0FE44F85B" box="[274,497,2006,2027]" italics="true" pageId="8" pageNumber="159">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A55FFE2FE3187B0FDF1F85B" box="[503,580,2006,2027]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
, entire.
<emphasis id="B6DDEACE5A55FFE2FD4987B3FD15F85A" bold="true" box="[655,672,2005,2026]" pageId="8" pageNumber="159">A</emphasis>
: male N1 (holotype);
<emphasis id="B6DDEACE5A55FFE2FCB387B3FC30F85A" bold="true" box="[885,901,2005,2026]" pageId="8" pageNumber="159">B</emphasis>
: female (paratype). Scale bars: 100
<emphasis id="B6DDEACE5A55FFE2FB2687B0FB59F85B" box="[1248,1260,2006,2027]" italics="true" pageId="8" pageNumber="159">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="D0D666545A54FFE3FF5185F8FAF4FA04" box="[151,1345,1438,1460]" httpUri="https://zenodo.org/record/183779/files/figure.png" pageId="9" pageNumber="160" targetBox="[185,1400,185,1414]" targetPageId="9">
<paragraph id="841636DC5A54FFE3FF5185F8FAF4FA04" blockId="9.[151,1345,1438,1460]" box="[151,1345,1438,1460]" pageId="9" pageNumber="160">
<emphasis id="B6DDEACE5A54FFE3FF5185F8FEB3FA03" bold="true" box="[151,262,1438,1459]" pageId="9" pageNumber="160">FIGURE 4</emphasis>
.
<taxonomicName id="43A94D5F5A54FFE3FED485F9FE44FA04" box="[274,497,1439,1460]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="9" pageNumber="160" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A54FFE3FED485F9FE44FA04" box="[274,497,1439,1460]" italics="true" pageId="9" pageNumber="160">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A54FFE3FE3185F9FDF1FA04" box="[503,580,1439,1460]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
, anterior ends.
<emphasis id="B6DDEACE5A54FFE3FD1385F8FD53FA03" bold="true" box="[725,742,1438,1459]" pageId="9" pageNumber="160">A</emphasis>
: male N1 (holotype);
<emphasis id="B6DDEACE5A54FFE3FC7D85F8FC7EFA03" bold="true" box="[955,971,1438,1459]" pageId="9" pageNumber="160">B</emphasis>
: female (paratype). Scale bars: 10
<emphasis id="B6DDEACE5A54FFE3FADC85F9FA93FA04" box="[1306,1318,1439,1460]" italics="true" pageId="9" pageNumber="160">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="841636DC5A54FFE3FF008582FB2DF9FE" blockId="9.[151,1438,1508,2014]" pageId="9" pageNumber="160">Cheilostoma cup­shaped, its walls reinforced by triangular rugae apparently twelve in number. A big pointed sclerotized tooth on the dorsal side of the pharyngeal portion of the buccal cavity. There are no smaller opposite teeth, but obscure subventral cuticular hardenings in the buccal cavity.</paragraph>
<paragraph id="841636DC5A54FFE3FF00863AFB89F976" blockId="9.[151,1438,1508,2014]" pageId="9" pageNumber="160">Pharynx evenly muscular throughout its length, widening posteriorly but not forming a true bulb. Internal cuticular lining twisted and folded. Cardia poorly visible. Nerve ring hardly discernible in some specimens. Renette cell not found except an oval body just posterior to the cardia ventrally.</paragraph>
<paragraph id="841636DC5A54FFE3FF0086B2FCBEF86E" blockId="9.[151,1438,1508,2014]" pageId="9" pageNumber="160">Didelphic, ovaries antidromously reflected; anterior ovary situated subventrally to the right of the intestine, posterior ovary subventrally to the left of the intestine. Diorchic; anterior testis outstretched, posterior smaller testis reflexed. Anterior testis situated to the right of the intestine, posterior testis to the left of the intestine. Spicules short, weak, sharply angularly bent, distally pointed, proximally with wide knobs. Gubernaculum paired, composed of two slightly sigmoid­curved platelets parallel to the distal portion of the spicules. Six supplementary organs arranged in a midventral preanal row. They look like thick conical setae one third protruded of cuticle and directed aslant anteriad.</paragraph>
<caption id="D0D666545A57FFE0FF518728FD3FF831" httpUri="https://zenodo.org/record/183780/files/figure.png" pageId="10" pageNumber="161" targetBox="[185,1393,189,1846]" targetPageId="10">
<paragraph id="841636DC5A57FFE0FF518728FD3FF831" blockId="10.[151,1436,1870,1921]" pageId="10" pageNumber="161">
<emphasis id="B6DDEACE5A57FFE0FF518728FEB2F8D3" bold="true" box="[151,263,1870,1891]" pageId="10" pageNumber="161">FIGURE 5</emphasis>
.
<taxonomicName id="43A94D5F5A57FFE0FED58729FE46F8D4" box="[275,499,1871,1892]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="10" pageNumber="161" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A57FFE0FED58729FE46F8D4" box="[275,499,1871,1892]" italics="true" pageId="10" pageNumber="161">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A57FFE0FE3F8729FDF3F8D4" box="[505,582,1871,1892]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
, details.
<emphasis id="B6DDEACE5A57FFE0FD5D8728FD19F8D3" bold="true" box="[667,684,1870,1891]" pageId="10" pageNumber="161">A</emphasis>
: male N1 (holotype), anterior body;
<emphasis id="B6DDEACE5A57FFE0FBD48728FB97F8D3" bold="true" box="[1042,1058,1870,1891]" pageId="10" pageNumber="161">B</emphasis>
: male N1 (holotype), posterior end;
<emphasis id="B6DDEACE5A57FFE0FA428728FA20F8D3" bold="true" box="[1412,1429,1870,1891]" pageId="10" pageNumber="161">C</emphasis>
: female (paratype), posterior end. Scale bars: 50
<emphasis id="B6DDEACE5A57FFE0FDA5870AFDDAF831" box="[611,623,1900,1921]" italics="true" pageId="10" pageNumber="161">µ</emphasis>
m.
</paragraph>
</caption>
<caption id="D0D666545A56FFE1FF51868FFB3CF94E" box="[151,1161,1769,1790]" httpUri="https://zenodo.org/record/183781/files/figure.png" pageId="11" pageNumber="162" targetBox="[152,1205,183,1768]" targetPageId="11">
<paragraph id="841636DC5A56FFE1FF51868FFB3CF94E" blockId="11.[151,1161,1769,1790]" box="[151,1161,1769,1790]" pageId="11" pageNumber="162">
<emphasis id="B6DDEACE5A56FFE1FF51868FFEB3F94E" bold="true" box="[151,262,1769,1790]" pageId="11" pageNumber="162">FIGURE 6</emphasis>
.
<taxonomicName id="43A94D5F5A56FFE1FED4868FFE44F94E" box="[274,497,1769,1790]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="11" pageNumber="162" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A56FFE1FED4868FFE44F94E" box="[274,497,1769,1790]" italics="true" pageId="11" pageNumber="162">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A56FFE1FE31868FFDF1F94E" box="[503,580,1769,1790]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
, cloacal region of the male N1 (holotype). Scale bar: 10
<emphasis id="B6DDEACE5A56FFE1FBA4868FFBDBF94E" box="[1122,1134,1769,1790]" italics="true" pageId="11" pageNumber="162">µ</emphasis>
m.
</paragraph>
</caption>
<paragraph id="841636DC5A56FFE1FF00873FFBAEF82B" blockId="11.[151,1437,1881,2027]" pageId="11" pageNumber="162">Tail of moderate length, consists of proximal conical and distal slender cylindrical portions, with delicate terminal spinnerete tube. There are a few short lateroventral setae on the tail.</paragraph>
</subSubSection>
<subSubSection id="CCB365575A56FFE6FF0087CFFAFBFE7A" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162" type="diagnosis">
<paragraph id="841636DC5A56FFE6FF0087CFFB93FE9A" blockId="11.[151,1437,1881,2027]" lastBlockId="12.[151,1436,152,1098]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
<emphasis id="B6DDEACE5A56FFE1FF0087CFFE88F872" box="[198,317,1961,1986]" italics="true" pageId="11" pageNumber="162">Diagnosis</emphasis>
:
<taxonomicName id="43A94D5F5A56FFE1FE8887CFFD90F872" box="[334,549,1961,1986]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="11" pageNumber="162" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A56FFE1FE8887CFFD90F872" box="[334,549,1961,1986]" italics="true" pageId="11" pageNumber="162">Paracyatholaimus</emphasis>
</taxonomicName>
. Body length 9361176
<emphasis id="B6DDEACE5A56FFE1FC8D87CFFCEFF872" box="[843,858,1961,1986]" italics="true" pageId="11" pageNumber="162">µ</emphasis>
m. No lateral differentiation in the cuticle. Outer labial and cephalic setae about 3.55
<emphasis id="B6DDEACE5A56FFE1FD9587B7FDD7F85A" box="[595,610,2001,2026]" italics="true" pageId="11" pageNumber="162">µ</emphasis>
m. Amphidial fovea multispiral with six turns in males and four turns in females situated at the level of the dorsal tooth. Buccal cavity armoured with a prominent dorsal tooth and no subventral teeth. Tail consists of anterior conical and posterior slender cylindrical half portions. Diorchic, posterior testis reflexed. Spicules sharply bent, distally pointed, proximally knobbed. Gubernaculum distally narrowed and toothless. Six preanal midventral setose supplementary organs.
</paragraph>
<paragraph id="841636DC5A51FFE6FF00815EFAFBFE7A" blockId="12.[151,1436,152,1098]" pageId="12" pageNumber="163">
<emphasis id="B6DDEACE5A51FFE6FF00815EFE70FEE1" box="[198,453,312,337]" italics="true" pageId="12" pageNumber="163">Differential diagnosis</emphasis>
: The new species resembles
<taxonomicName id="43A94D5F5A51FFE6FCD9815EFAA6FEE2" authority="Gerlach, 1964" authorityName="Gerlach" authorityYear="1964" box="[799,1299,312,338]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="rotundus">
<emphasis id="B6DDEACE5A51FFE6FCD9815EFBD1FEE1" box="[799,1124,312,337]" italics="true" pageId="12" pageNumber="163">Paracyatholaimus rotundus</emphasis>
Gerlach, 1964
</taxonomicName>
closer than other
<taxonomicName id="43A94D5F5A51FFE6FF1C8106FE04FEC9" box="[218,433,352,377]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A51FFE6FF1C8106FE04FEC9" box="[218,433,352,377]" italics="true" pageId="12" pageNumber="163">Paracyatholaimus</emphasis>
</taxonomicName>
species.
<taxonomicName id="43A94D5F5A51FFE6FDDC8106FD28FEC9" box="[538,669,352,377]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="rotundus">
<emphasis id="B6DDEACE5A51FFE6FDDC8106FD28FEC9" box="[538,669,352,377]" italics="true" pageId="12" pageNumber="163">P. rotundus</emphasis>
</taxonomicName>
is known from only one male specimen from the Red Sea.
<emphasis id="B6DDEACE5A51FFE6FA8B8106FA2DFEC9" box="[1357,1432,352,377]" italics="true" pageId="12" pageNumber="163">P. d i v a</emphasis>
sp. nov. differs from
<taxonomicName id="43A94D5F5A51FFE6FE5381EEFDAEFE11" box="[405,539,392,417]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="rotundus">
<emphasis id="B6DDEACE5A51FFE6FE5381EEFDAEFE11" box="[405,539,392,417]" italics="true" pageId="12" pageNumber="163">P. rotundus</emphasis>
</taxonomicName>
by a slightly longer body (9361176
<emphasis id="B6DDEACE5A51FFE6FC2281EEFC46FE11" box="[996,1011,392,417]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FBD081EEFBD4FE11" box="[1046,1121,392,417]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
705
<emphasis id="B6DDEACE5A51FFE6FB6681EEFB1AFE11" box="[1184,1199,392,417]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), tail shape (with clearly narrowed posterior half
<emphasis id="B6DDEACE5A51FFE6FDCE81D6FDE6FE79" box="[520,595,432,457]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
conical), six
<emphasis id="B6DDEACE5A51FFE6FD3781D6FC89FE79" box="[753,828,432,457]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
four supplementary organs of another shape.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A51FFE6FF0081BEFD2BFBFA" pageId="12" pageNumber="163" type="description">
<paragraph id="841636DC5A51FFE6FF0081BEFD2BFBFA" blockId="12.[151,1436,152,1098]" pageId="12" pageNumber="163">
<emphasis id="B6DDEACE5A51FFE6FF0081BEFEA1FE41" box="[198,276,472,497]" italics="true" pageId="12" pageNumber="163">P. d i v a</emphasis>
sp. nov. shares large multispiral amphidial fovea with 56 turns with
<taxonomicName id="43A94D5F5A51FFE6FBA481BEFF61FDAA" authority="Andrassy, 1973" authorityName="Andrassy" authorityYear="1973" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="botosaneanui">
<emphasis id="B6DDEACE5A51FFE6FBA481BEFAAAFE41" box="[1122,1311,472,497]" italics="true" pageId="12" pageNumber="163">P. botosaneanui</emphasis>
<bibRefCitation id="E0384B2D5A51FFE6FAEE81BEFF61FDAA" author="Andrassy" pageId="12" pageNumber="173" refString="Andrassy, I. (1973) Nematoden aus Strand- und Hohlenbiotopen von Kuba. Acta Zoologica Academiae Scientarum Hungaricae, 19, 233 - 270." type="journal article" year="1973">Andrássy, 1973</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="43A94D5F5A51FFE6FF198266FD91FDAA" authority="Gerlach, 1964" authorityName="Gerlach" authorityYear="1964" box="[223,548,512,538]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="duplicatus">
<emphasis id="B6DDEACE5A51FFE6FF198266FEC3FDA9" box="[223,374,512,537]" italics="true" pageId="12" pageNumber="163">P. duplicatus</emphasis>
Gerlach, 1964
</taxonomicName>
,
<taxonomicName id="43A94D5F5A51FFE6FDF68266FC04FDAA" authority="Gerlach, 1955" authorityName="Gerlach" authorityYear="1955" box="[560,945,512,538]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="paucipapillatus">
<emphasis id="B6DDEACE5A51FFE6FDF68266FCB7FDA9" box="[560,770,512,537]" italics="true" pageId="12" pageNumber="163">P. paucipapillatus</emphasis>
<bibRefCitation id="E0384B2D5A51FFE6FCCF8266FC04FDAA" author="Gerlach" box="[777,945,512,538]" pageId="12" pageNumber="173" refString="Gerlach, S. A. (1955) Zur Kenntnis der freilebenden marinen Nematoden von San Salvador. Zeitschrift fur Wissenschaftliche Zoologie, 158, 249 - 303." type="journal article" year="1955">Gerlach, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="43A94D5F5A51FFE6FC7A8266FACEFDAA" authority="Wieser &amp; Hopper, 1967" authorityName="Wieser &amp; Hopper" authorityYear="1967" box="[956,1403,512,538]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="pugettensis">
<emphasis id="B6DDEACE5A51FFE6FC7A8266FBE9FDA9" box="[956,1116,512,537]" italics="true" pageId="12" pageNumber="163">P. pugettensis</emphasis>
<bibRefCitation id="E0384B2D5A51FFE6FBA58266FACEFDAA" author="Wieser" box="[1123,1403,512,538]" pageId="12" pageNumber="174" refString="Wieser, W. &amp; Hopper, B. (1967) Marine nematodes of the east coast of North America. I. Florida. Bulletin of the Museum for Comparative Zoology 135 (5), 239 - 344." type="journal article" year="1967">Wieser &amp; Hopper, 1967</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="43A94D5F5A51FFE6FA408266FE7FFDF2" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" class="Arachnida" family="Dictynidae" genus="Marilynia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A51FFE6FA408266FEA1FDF1" italics="true" pageId="12" pageNumber="163">P. quadriseta</emphasis>
(Wieser, 1954)
</taxonomicName>
and
<taxonomicName id="43A94D5F5A51FFE6FDC3824EFC46FDF2" authority="Sharma &amp; Vincx, 1982" authorityName="Sharma &amp; Vincx" authorityYear="1982" box="[517,1011,552,578]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="vancouverensis">
<emphasis id="B6DDEACE5A51FFE6FDC3824EFD62FDF1" box="[517,727,552,577]" italics="true" pageId="12" pageNumber="163">P. vancouverensis</emphasis>
<bibRefCitation id="E0384B2D5A51FFE6FD18824EFC46FDF2" author="Sharma" box="[734,1011,552,578]" pageId="12" pageNumber="174" refString="Sharma, J. &amp; Vincx, M. (1982) Cyatholaimidae (Nematoda) from the Canadian Pacific Coast. Canadian Journal of Zoology, 60, 271 - 280." type="journal article" year="1982">Sharma &amp; Vincx, 1982</bibRefCitation>
</taxonomicName>
.
<emphasis id="B6DDEACE5A51FFE6FC38824EFBFEFDF1" box="[1022,1099,552,577]" italics="true" pageId="12" pageNumber="163">P. d i v a</emphasis>
sp. nov. differs from
<emphasis id="B6DDEACE5A51FFE6FA89824EFEBAFDD9" italics="true" pageId="12" pageNumber="163">P. b o t ­ osaneanui</emphasis>
by slightly a thicker body (a 23.828
<emphasis id="B6DDEACE5A51FFE6FD1C8236FC90FDD9" box="[730,805,592,617]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
3133), index c (7.458.86
<emphasis id="B6DDEACE5A51FFE6FBBF8236FB71FDD9" box="[1145,1220,592,617]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
13.515), smaller cephalic setae (34
<emphasis id="B6DDEACE5A51FFE6FE44821EFE24FD21" box="[386,401,632,657]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FE74821EFE48FD21" box="[434,509,632,657]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
1012
<emphasis id="B6DDEACE5A51FFE6FD90821EFDD0FD21" box="[598,613,632,657]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), longer spicules (3436
<emphasis id="B6DDEACE5A51FFE6FC6A821EFC0EFD21" box="[940,955,632,657]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FC1A821EFB92FD21" box="[988,1063,632,657]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
2628
<emphasis id="B6DDEACE5A51FFE6FB47821EFB25FD21" box="[1153,1168,632,657]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m) and six
<emphasis id="B6DDEACE5A51FFE6FADE821EFAD6FD21" box="[1304,1379,632,657]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
four supplementary organs of another shape; from
<taxonomicName id="43A94D5F5A51FFE6FD7082C6FCF8FD09" box="[694,845,672,697]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="duplicatus">
<emphasis id="B6DDEACE5A51FFE6FD7082C6FCF8FD09" box="[694,845,672,697]" italics="true" pageId="12" pageNumber="163">P. duplicatus</emphasis>
</taxonomicName>
by shorter body (9361176
<emphasis id="B6DDEACE5A51FFE6FB5B82C6FB19FD09" box="[1181,1196,672,697]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FB0B82C6FAADFD09" box="[1229,1304,672,697]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
1485
<emphasis id="B6DDEACE5A51FFE6FAA582C6FAC7FD09" box="[1379,1394,672,697]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), smaller cephalic setae (34
<emphasis id="B6DDEACE5A51FFE6FE1D82AEFE5FFD51" box="[475,490,712,737]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FDCF82AEFDE1FD51" box="[521,596,712,737]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
78
<emphasis id="B6DDEACE5A51FFE6FD4882AEFD28FD51" box="[654,669,712,737]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), relatively shorter nearly conical tail (c 3.023.43
<emphasis id="B6DDEACE5A51FFE6FAD182AEFAD7FD51" box="[1303,1378,712,737]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
5.5 6.5) and arrangement of supplementary organs (three distinct pairs in
<taxonomicName id="43A94D5F5A51FFE6FC158296FBDEFCB9" box="[979,1131,752,777]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="duplicatus">
<emphasis id="B6DDEACE5A51FFE6FC158296FBDEFCB9" box="[979,1131,752,777]" italics="true" pageId="12" pageNumber="163">P. duplicatus</emphasis>
</taxonomicName>
); from
<taxonomicName id="43A94D5F5A51FFE6FB0E8296FA2EFCB9" box="[1224,1435,752,777]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="paucipapillatus">
<emphasis id="B6DDEACE5A51FFE6FB0E8296FA2EFCB9" box="[1224,1435,752,777]" italics="true" pageId="12" pageNumber="163">P. paucipapillatus</emphasis>
</taxonomicName>
by shorter body (9361176
<emphasis id="B6DDEACE5A51FFE6FE27837EFE45FC81" box="[481,496,792,817]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FDD7837EFDE9FC81" box="[529,604,792,817]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
<date id="F017101C5A51FFE6FDA2837EFD5CFC82" box="[612,745,792,818]" pageId="12" pageNumber="163" value="1445" valueMax="1860">14451860</date>
<emphasis id="B6DDEACE5A51FFE6FD37837EFCB5FC81" box="[753,768,792,817]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), index c (7.458.86
<emphasis id="B6DDEACE5A51FFE6FBDF837EFBD1FC81" box="[1049,1124,792,817]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
13.914.1), much smaller cephalic setae (34
<emphasis id="B6DDEACE5A51FFE6FE408326FE20FCE9" box="[390,405,832,857]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FE7E8326FDB6FCE9" box="[440,515,832,857]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
815
<emphasis id="B6DDEACE5A51FFE6FD948326FDD4FCE9" box="[594,609,832,857]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m) and six
<emphasis id="B6DDEACE5A51FFE6FD2B8326FC8DFCE9" box="[749,824,832,857]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
two supplementary organs; from
<taxonomicName id="43A94D5F5A51FFE6FB178326FAC0FCE9" box="[1233,1397,832,857]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="pugettensis">
<emphasis id="B6DDEACE5A51FFE6FB178326FAC0FCE9" box="[1233,1397,832,857]" italics="true" pageId="12" pageNumber="163">P. pugettensis</emphasis>
</taxonomicName>
by shorter body (9361176
<emphasis id="B6DDEACE5A51FFE6FE79830EFE7BFC31" box="[447,462,872,897]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FE37830EFD89FC31" box="[497,572,872,897]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
13102230
<emphasis id="B6DDEACE5A51FFE6FD15830EFD57FC31" box="[723,738,872,897]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), smaller cephalic setae (34
<emphasis id="B6DDEACE5A51FFE6FBA1830EFBC3FC31" box="[1127,1142,872,897]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FB5E830EFB56FC31" box="[1176,1251,872,897]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
59
<emphasis id="B6DDEACE5A51FFE6FAE5830EFA87FC31" box="[1315,1330,872,897]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), relatively shorter tail (c 3.023.43
<emphasis id="B6DDEACE5A51FFE6FDDA83F6FDD2FC19" box="[540,615,912,937]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
5.56.5) and six
<emphasis id="B6DDEACE5A51FFE6FC8683F6FC3EFC19" box="[832,907,912,937]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
four to five supplementary organs; from
<taxonomicName id="43A94D5F5A51FFE6FA4083F6FEA1FC61" class="Arachnida" family="Dictynidae" genus="Marilynia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A51FFE6FA4083F6FEA1FC61" italics="true" pageId="12" pageNumber="163">P. quadriseta</emphasis>
</taxonomicName>
by smaller cephalic setae (34
<emphasis id="B6DDEACE5A51FFE6FD4B83DEFD29FC61" box="[653,668,952,977]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FD7883DEFCBCFC61" box="[702,777,952,977]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
89
<emphasis id="B6DDEACE5A51FFE6FC8083DEFCE0FC61" box="[838,853,952,977]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m) and relatively shorter tail (c3.023.43
<emphasis id="B6DDEACE5A51FFE6FA9783DEFA29FC61" box="[1361,1436,952,977]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
6.3); from
<taxonomicName id="43A94D5F5A51FFE6FED28386FE51FC49" box="[276,484,992,1017]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="12" pageNumber="163" phylum="Nematoda" rank="species" species="vancouverensis">
<emphasis id="B6DDEACE5A51FFE6FED28386FE51FC49" box="[276,484,992,1017]" italics="true" pageId="12" pageNumber="163">P. vancouverensis</emphasis>
</taxonomicName>
by shorter body (9361176
<emphasis id="B6DDEACE5A51FFE6FCEA8386FC8EFC49" box="[812,827,992,1017]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m
<emphasis id="B6DDEACE5A51FFE6FC9D8386FC13FC49" box="[859,934,992,1017]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
18901960
<emphasis id="B6DDEACE5A51FFE6FBF18386FBF3FC49" box="[1079,1094,992,1017]" italics="true" pageId="12" pageNumber="163">µ</emphasis>
m), relatively thicker body (a 23.828
<emphasis id="B6DDEACE5A51FFE6FF3B846EFEFDFB91" box="[253,328,1032,1057]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
3435), wider amphidial fovea (5363%
<emphasis id="B6DDEACE5A51FFE6FCF7846EFCC9FB91" box="[817,892,1032,1057]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
42% c.b.d.) and relatively shorter tail (c 7.45 8.86
<emphasis id="B6DDEACE5A51FFE6FF178456FEA9FBF9" box="[209,284,1072,1097]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
12.116, c 3.023.43
<emphasis id="B6DDEACE5A51FFE6FDEA8456FDC2FBF9" box="[556,631,1072,1097]" italics="true" pageId="12" pageNumber="163">versus</emphasis>
5).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,900 @@
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<mods:title id="EAD2F5F73E431C6FC3A4C734B0EFC1CE">Three new species of free­living nematodes from the South­East Atlantic Abyss (DIVA I Expedition) *</mods:title>
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<treatment id="0C0087CA5A5FFFECFF5180BBFB93FEFF" ID-DOI="http://doi.org/10.5281/zenodo.6231734" ID-GBIF-Taxon="119374535" ID-Zenodo-Dep="6231734" LSID="urn:lsid:plazi:treatment:0C0087CA5A5FFFECFF5180BBFB93FEFF" httpUri="http://treatment.plazi.org/id/0C0087CA5A5FFFECFF5180BBFB93FEFF" lastPageId="6" lastPageNumber="157" pageId="2" pageNumber="153">
<subSubSection id="CCB365575A5FFFE8FF5180BBFE70FEA7" pageId="2" pageNumber="153" type="nomenclature">
<paragraph id="841636DC5A5FFFE8FF5180BBFDF9FF47" blockId="2.[151,588,151,279]" box="[151,588,221,247]" pageId="2" pageNumber="153">
<heading id="DF5E81B05A5FFFE8FF5180BBFDF9FF47" bold="true" box="[151,588,221,247]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
<taxonomicName id="43A94D5F5A5FFFE8FF5180BBFDF9FF47" authority="Micoletzky, 1923" authorityName="Micoletzky" authorityYear="1923" box="[151,588,221,247]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5FFFE8FF5180BBFDF9FF47" bold="true" box="[151,588,221,247]" pageId="2" pageNumber="153">
<emphasis id="B6DDEACE5A5FFFE8FF5180BBFEC6FF47" bold="true" box="[151,371,221,247]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus</emphasis>
Micoletzky, 1923
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518166FE70FEA7" blockId="2.[151,588,151,279]" box="[151,453,256,279]" pageId="2" pageNumber="153">
<bibRefCitation id="E0384B2D5A5FFFE8FF518166FE87FEA7" author="Wieser" box="[151,306,256,279]" pageId="2" pageNumber="174" refString="Wieser, W. (1954 a) Free-living marine nematodes. II. Chromadoroidea. Lunds Universitets Arsskrift. N. F. Avd. 2, 50 (16), 1 - 148." type="journal article" year="1954" yearSuffix="a">Wieser, 1954a</bibRefCitation>
: 26, emended
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF51812EFED1FDE2" pageId="2" pageNumber="153" type="diagnosis">
<paragraph id="841636DC5A5FFFE8FF51812EFEBBFED1" blockId="2.[151,1437,328,594]" box="[151,270,328,353]" pageId="2" pageNumber="153">
<emphasis id="B6DDEACE5A5FFFE8FF51812EFEBBFED1" box="[151,270,328,353]" italics="true" pageId="2" pageNumber="153">Diagnosis</emphasis>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF008116FED1FDE2" blockId="2.[151,1437,328,594]" pageId="2" pageNumber="153">Cuticle punctated, lateral differentiation hardly developed. Six outer labial sensilla + four cephalic sensilla setose, in a single circle. Amphidial fovea multispiral. Cheilostoma armoured with twelve rugae; distinct dorsal tooth, often supplemented with smaller subventral teeth and occasionally other denticles in the stegostoma. Precloacal ventromedian supplements as setae­like organs half inserted into the body. Gubernaculum hardly dilated at the distal end and devoid of denticles or serrations. Tail conical or with more or less slender cylindrical distal portion.</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF5182EEFE9DFD11" box="[151,296,648,673]" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="841636DC5A5FFFE8FF5182EEFE9DFD11" blockId="2.[151,658,648,714]" box="[151,296,648,673]" pageId="2" pageNumber="153">
<emphasis id="B6DDEACE5A5FFFE8FF5182EEFE9DFD11" box="[151,296,648,673]" italics="true" pageId="2" pageNumber="153">
<typeStatus id="5B12887E5A5FFFE8FF5182EEFF79FD11" box="[151,204,648,673]" pageId="2" pageNumber="153">Type</typeStatus>
species
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF0082D6FB09FC22" pageId="2" pageNumber="153" type="reference_group">
<paragraph id="841636DC5A5FFFE8FF0082D6FD27FD7A" blockId="2.[151,658,648,714]" box="[198,658,688,714]" pageId="2" pageNumber="153">
<taxonomicName id="43A94D5F5A5FFFE8FF0082D6FD3BFD7A" authority="Butschli, 1874" authorityName="Butschli" authorityYear="1874" box="[198,654,688,714]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="dubiosus">
<emphasis id="B6DDEACE5A5FFFE8FF0082D6FE6DFD79" box="[198,472,688,713]" italics="true" pageId="2" pageNumber="153">Cyatholaimus dubiosus</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FE1982D6FD3BFD7A" author="Butschli" box="[479,654,688,714]" pageId="2" pageNumber="173" refString="Butschli, O. (1874) Zur Kenntnis der freilebenden Nematoden, insbesondere der des Kieler Hafens. Abhandlungen der Senckenbergischen Naturforschen Gesellschaft, 9, 236 - 292." type="journal article" year="1874">Bütschli, 1874</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518366FECBFCA9" blockId="2.[151,1439,768,2034]" box="[151,382,768,793]" pageId="2" pageNumber="153">
<emphasis id="B6DDEACE5A5FFFE8FF518366FECBFCA9" box="[151,382,768,793]" italics="true" pageId="2" pageNumber="153">List of valid species</emphasis>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF51834EFB3FFCF2" blockId="2.[151,1439,768,2034]" box="[151,1162,808,834]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF51834EFB3FFCF2" box="[151,1162,808,834]" pageId="2" pageNumber="153">
1
<taxonomicName id="43A94D5F5A5FFFE8FF01834EFD23FCF2" authority="Kreis, 1963" authorityName="Kreis" authorityYear="1963" box="[199,662,808,834]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="arcticus">
<emphasis id="B6DDEACE5A5FFFE8FF01834EFDB4FCF1" box="[199,513,808,833]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus arcticus</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FDCC834EFD23FCF2" author="Kreis" box="[522,662,808,834]" pageId="2" pageNumber="173" refString="Kreis, H. (1963) Marine Nematoda. Zoology of Iceland, 2 (14), 1 - 68." type="journal article" year="1963">Kreis, 1963</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD67834EFCE3FCF2" author="Kreis" box="[673,854,808,834]" page="39" pageId="2" pageNumber="153" year="1963">
<bibRefCitation id="E0384B2D5A5FFFE8FD67834EFC9BFCF2" author="Kreis" box="[673,814,808,834]" pageId="2" pageNumber="173" refString="Kreis, H. (1963) Marine Nematoda. Zoology of Iceland, 2 (14), 1 - 68." type="journal article" year="1963">Kreis, 1963</bibRefCitation>
: 39
</treatmentCitation>
40, fig. 20 AC;
<collectingCountry id="FCBE764C5A5FFFE8FBEA834EFB33FCF2" box="[1068,1158,808,834]" name="Iceland" pageId="2" pageNumber="153">Iceland</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518336FA9AFCDA" blockId="2.[151,1439,768,2034]" box="[151,1327,848,874]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF518336FA9AFCDA" box="[151,1327,848,874]" pageId="2" pageNumber="153">
2
<taxonomicName id="43A94D5F5A5FFFE8FF018336FCB1FCDA" authority="Andrassy, 1973" authorityName="Andrassy" authorityYear="1973" box="[199,772,848,874]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="botosaneanui">
<emphasis id="B6DDEACE5A5FFFE8FF018336FDF6FCD9" box="[199,579,848,873]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus botosaneanui</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FD8C8336FCB1FCDA" author="Andrassy" box="[586,772,848,874]" pageId="2" pageNumber="173" refString="Andrassy, I. (1973) Nematoden aus Strand- und Hohlenbiotopen von Kuba. Acta Zoologica Academiae Scientarum Hungaricae, 19, 233 - 270." type="journal article" year="1973">Andrássy, 1973</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FCC98336FBB6FCDA" author="Andrassy" box="[783,1027,848,874]" page="259" pageId="2" pageNumber="153" year="1973">
<bibRefCitation id="E0384B2D5A5FFFE8FCC98336FC7FFCDA" author="Andrassy" box="[783,970,848,874]" pageId="2" pageNumber="173" refString="Andrassy, I. (1973) Nematoden aus Strand- und Hohlenbiotopen von Kuba. Acta Zoologica Academiae Scientarum Hungaricae, 19, 233 - 270." type="journal article" year="1973">Andrássy, 1973</bibRefCitation>
: 259
</treatmentCitation>
261, Abb. 9 AC;
<collectingCountry id="FCBE764C5A5FFFE8FB2F8336FA9EFCDA" box="[1257,1323,848,874]" name="Cuba" pageId="2" pageNumber="153">Cuba</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF51831EFB09FC22" blockId="2.[151,1439,768,2034]" box="[151,1212,888,914]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF51831EFB09FC22" box="[151,1212,888,914]" pageId="2" pageNumber="153">
3
<taxonomicName id="43A94D5F5A5FFFE8FF01831EFD09FC22" authority="Gerlach, 1953" authorityName="Gerlach" authorityYear="1953" box="[199,700,888,914]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="chilensis">
<emphasis id="B6DDEACE5A5FFFE8FF01831EFDB9FC21" box="[199,524,888,913]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus chilensis</emphasis>
Gerlach, 1953
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD0E831EFC1CFC22" author="Gerlach" box="[712,937,888,914]" page="19" pageId="2" pageNumber="153" year="1953">
<bibRefCitation id="E0384B2D5A5FFFE8FD0E831EFC37FC22" author="Gerlach" box="[712,898,888,914]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1953 b) Freilebende marine Nematoden aus dem Kustengrundwasser und aus dem Brackwasser der chilenischen Kuste. Lunds Universitets Arsskrift. N. F. Avd. 2, 49 (10), 1 - 37." type="journal article" year="1953" yearSuffix="b">Gerlach, 1953b</bibRefCitation>
: 19
</treatmentCitation>
21, Abb. 9 ae;
<collectingCountry id="FCBE764C5A5FFFE8FBB2831EFB0DFC22" box="[1140,1208,888,914]" name="Chile" pageId="2" pageNumber="153">Chile</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF5183C6FD50FC0A" box="[151,741,928,954]" pageId="2" pageNumber="153" type="description">
<paragraph id="841636DC5A5FFFE8FF5183C6FD50FC0A" blockId="2.[151,1439,768,2034]" box="[151,741,928,954]" pageId="2" pageNumber="153">
4
<taxonomicName id="43A94D5F5A5FFFE8FF0183C6FE6DFC09" box="[199,472,928,953]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="diva" status="sp. nov.">
<emphasis id="B6DDEACE5A5FFFE8FF0183C6FE6DFC09" box="[199,472,928,953]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus diva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADEE57B55A5FFFE8FE1983C6FD8CFC0A" box="[479,569,928,954]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
Present paper.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF5183AEFAF8FBEA" pageId="2" pageNumber="153" type="reference_group">
<paragraph id="841636DC5A5FFFE8FF5183AEFE38FB82" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5183AEFE38FB82" pageId="2" pageNumber="153">
5
<taxonomicName id="43A94D5F5A5FFFE8FF0183AEFD6EFC52" authority="Butschli, 1874" authorityName="Butschli" authorityYear="1874" box="[199,731,968,994]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="dubiosus">
<emphasis id="B6DDEACE5A5FFFE8FF0183AEFDBAFC51" box="[199,527,968,993]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus dubiosus</emphasis>
(
<bibRefCitation id="E0384B2D5A5FFFE8FDD983AEFD67FC52" author="Butschli" box="[543,722,968,994]" pageId="2" pageNumber="173" refString="Butschli, O. (1874) Zur Kenntnis der freilebenden Nematoden, insbesondere der des Kieler Hafens. Abhandlungen der Senckenbergischen Naturforschen Gesellschaft, 9, 236 - 292." type="journal article" year="1874">Bütschli, 1874</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD2183AEFC70FC52" author="Butschli" box="[743,965,968,994]" page="48" pageId="2" pageNumber="153" year="1874">
<bibRefCitation id="E0384B2D5A5FFFE8FD2183AEFC2DFC52" author="Butschli" box="[743,920,968,994]" pageId="2" pageNumber="173" refString="Butschli, O. (1874) Zur Kenntnis der freilebenden Nematoden, insbesondere der des Kieler Hafens. Abhandlungen der Senckenbergischen Naturforschen Gesellschaft, 9, 236 - 292." type="journal article" year="1874">Bütschli, 1874</bibRefCitation>
: 48
</treatmentCitation>
, fig. 31 a, b (=
<taxonomicName id="43A94D5F5A5FFFE8FBB083AEFA3FFC51" box="[1142,1418,968,993]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="dubiosus">
<emphasis id="B6DDEACE5A5FFFE8FBB083AEFA3FFC51" box="[1142,1418,968,993]" italics="true" pageId="2" pageNumber="153">Cyatholaimus dubiosus</emphasis>
</taxonomicName>
); Kiel Bay.
<treatmentCitation id="050810CD5A5FFFE8FE838396FDE0FBBA" author="Micoletzky" box="[325,597,1008,1034]" page="377" pageId="2" pageNumber="153" year="1922">
<bibRefCitation id="E0384B2D5A5FFFE8FE838396FDAFFBBA" author="Micoletzky" box="[325,538,1008,1034]" pageId="2" pageNumber="173" refString="Micoletzky, H. (1922) Die freilebenden Erdnematoden. Archiv fur Naturgeschichte, 87 A, 1 - 650." type="journal article" year="1922">Micoletzky, 1922</bibRefCitation>
: 377
</treatmentCitation>
(to subgenus
<taxonomicName id="43A94D5F5A5FFFE8FCC28396FC6EFBB9" box="[772,987,1008,1033]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5FFFE8FCC28396FC6EFBB9" box="[772,987,1008,1033]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus</emphasis>
</taxonomicName>
).
<treatmentCitation id="050810CD5A5FFFE8FC318396FB71FBBA" author="Meyl" box="[1015,1220,1008,1034]" page="428" pageId="2" pageNumber="153" year="1954">
<bibRefCitation id="E0384B2D5A5FFFE8FC318396FB32FBBA" author="Meyl" box="[1015,1159,1008,1034]" pageId="2" pageNumber="173" refString="Meyl, A. H. (1954) Beitrage zur Kenntnis der Nematodenfauna vulkanisch erhitzter Biotope. III. Mitteilung. Nematoden aus der Mischungszone strandnaher, heisser Suβwasserquellen mit dem Meerwasser auf der Insel Ischia. Zeitschrift fur Morphologie und Okologie der Tiere, 42, 421 - 448." type="journal article" year="1954">Meyl, 1954</bibRefCitation>
: 428
</treatmentCitation>
429, Abb. 2 af; (Mediterranean).
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518426FAF8FBEA" blockId="2.[151,1439,768,2034]" box="[151,1357,1088,1114]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF518426FAF8FBEA" box="[151,1357,1088,1114]" pageId="2" pageNumber="153">
6
<taxonomicName id="43A94D5F5A5FFFE8FF018426FD65FBEA" authority="Gerlach, 1964" authorityName="Gerlach" authorityYear="1964" box="[199,720,1088,1114]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="duplicatus">
<emphasis id="B6DDEACE5A5FFFE8FF018426FDAAFBE9" box="[199,543,1088,1113]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus duplicatus</emphasis>
Gerlach, 1964
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD1D8426FC0BFBEA" author="Gerlach" box="[731,958,1088,1114]" page="77" pageId="2" pageNumber="153" year="1964">
<bibRefCitation id="E0384B2D5A5FFFE8FD1D8426FC20FBEA" author="Gerlach" box="[731,917,1088,1114]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1964 b) Neue Cyatholaimidae (Nematoda Chromadorida) von den Malediven. Veroffentlichungen des Instituts fur Meeresforschungen in Bremerhaven, 9, 70 - 78." type="journal article" year="1964" yearSuffix="b">Gerlach, 1964b</bibRefCitation>
: 77
</treatmentCitation>
78, Abb. 5 ad;
<collectingCountry id="FCBE764C5A5FFFE8FB4F8426FAFCFBEA" box="[1161,1353,1088,1114]" name="Maldives" pageId="2" pageNumber="153">Maldive Islands</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE8FF51840EFB3EFB32" box="[151,1163,1128,1154]" pageId="2" pageNumber="153" type="description">
<paragraph id="841636DC5A5FFFE8FF51840EFB3EFB32" blockId="2.[151,1439,768,2034]" box="[151,1163,1128,1154]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF51840EFB3EFB32" box="[151,1163,1128,1154]" pageId="2" pageNumber="153">
7
<taxonomicName id="43A94D5F5A5FFFE8FF01840EFD0FFB32" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[199,698,1128,1154]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="helicellus">
<emphasis id="B6DDEACE5A5FFFE8FF01840EFDA3FB31" box="[199,534,1128,1153]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus helicellus</emphasis>
Wieser, 1954
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD00840EFC29FB32" author="Wieser" box="[710,924,1128,1154]" page="27" pageId="2" pageNumber="153" year="1954">
<bibRefCitation id="E0384B2D5A5FFFE8FD00840EFCC7FB32" author="Wieser" box="[710,882,1128,1154]" pageId="2" pageNumber="174" refString="Wieser, W. (1954 a) Free-living marine nematodes. II. Chromadoroidea. Lunds Universitets Arsskrift. N. F. Avd. 2, 50 (16), 1 - 148." type="journal article" year="1954" yearSuffix="a">Wieser, 1954a</bibRefCitation>
: 27
</treatmentCitation>
, fig. 107 ac;
<collectingCountry id="FCBE764C5A5FFFE8FB85840EFB32FB32" box="[1091,1159,1128,1154]" name="Chile" pageId="2" pageNumber="153">Chile</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5FFFE9FF5184F6FB63FDAA" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="reference_group">
<paragraph id="841636DC5A5FFFE8FF5184F6FE10FA92" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5184F6FE10FA92" pageId="2" pageNumber="153">
8
<taxonomicName id="43A94D5F5A5FFFE8FF0184F6FD84FB19" box="[199,561,1168,1193]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="intermedius">
<emphasis id="B6DDEACE5A5FFFE8FF0184F6FD84FB19" box="[199,561,1168,1193]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus intermedius</emphasis>
</taxonomicName>
(de
<bibRefCitation id="E0384B2D5A5FFFE8FDA184F6FD58FB1A" author="Man" box="[615,749,1168,1194]" pageId="2" pageNumber="173" refString="de Man, J. G. (1880) Die einheimischen, frei in der reinen Erde und im suβen Wasser lebenden Nematoden monographisch bearbeitet. Vorlaufiger Bericht und descriptiv-systematischer Theil. Tijdschrift der Nederlandsche dierkundige Vereeniging, 5, 1 - 104." type="journal article" year="1880">
<collectingCountry id="FCBE764C5A5FFFE8FDA184F6FD15FB1A" box="[615,672,1168,1194]" name="Isle of Man" pageId="2" pageNumber="153">Man</collectingCountry>
, 1880
</bibRefCitation>
).
<collectingCountry id="FCBE764C5A5FFFE8FCC484F6FC91FB1A" box="[770,804,1168,1194]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="153">De</collectingCountry>
<treatmentCitation id="050810CD5A5FFFE8FCEB84F6FC6EFB1A" author="Man" box="[813,987,1168,1194]" page="16" pageId="2" pageNumber="153" year="1880">
<bibRefCitation id="E0384B2D5A5FFFE8FCEB84F6FC07FB1A" author="Man" box="[813,946,1168,1194]" pageId="2" pageNumber="173" refString="de Man, J. G. (1880) Die einheimischen, frei in der reinen Erde und im suβen Wasser lebenden Nematoden monographisch bearbeitet. Vorlaufiger Bericht und descriptiv-systematischer Theil. Tijdschrift der Nederlandsche dierkundige Vereeniging, 5, 1 - 104." type="journal article" year="1880">
<collectingCountry id="FCBE764C5A5FFFE8FCEB84F6FCD3FB1A" box="[813,870,1168,1194]" name="Isle of Man" pageId="2" pageNumber="153">Man</collectingCountry>
, 1880
</bibRefCitation>
: 16
</treatmentCitation>
17 (=
<taxonomicName id="43A94D5F5A5FFFE8FBEF84F6FAEAFB19" box="[1065,1375,1168,1193]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="intermedius">
<emphasis id="B6DDEACE5A5FFFE8FBEF84F6FAEAFB19" box="[1065,1375,1168,1193]" italics="true" pageId="2" pageNumber="153">Cyatholaimus intermedius</emphasis>
</taxonomicName>
);
<collectingCountry id="FCBE764C5A5FFFE8FABC84F6FA29FB1A" box="[1402,1436,1168,1194]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="153">De</collectingCountry>
<treatmentCitation id="050810CD5A5FFFE8FF0184DEFECDFB62" author="Man" box="[199,376,1208,1234]" page="52" pageId="2" pageNumber="153" year="1884">
<bibRefCitation id="E0384B2D5A5FFFE8FF0184DEFEF9FB62" author="Man" box="[199,332,1208,1234]" pageId="2" pageNumber="173" refString="de Man, J. G. (1884) Die frei in der reinen Erde und im suβen Wasser lebenden Nematoden der niederlandischen Fauna. Eine Systematisch-faunistische Monographie, Leiden 1884, 206 pp." type="book" year="1884">
<collectingCountry id="FCBE764C5A5FFFE8FF0184DEFEB5FB62" box="[199,256,1208,1234]" name="Isle of Man" pageId="2" pageNumber="153">Man</collectingCountry>
, 1884
</bibRefCitation>
: 52
</treatmentCitation>
54, fig. 2525f (=
<taxonomicName id="43A94D5F5A5FFFE8FD9D84DEFC24FB61" box="[603,913,1208,1233]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="intermedius">
<emphasis id="B6DDEACE5A5FFFE8FD9D84DEFC24FB61" box="[603,913,1208,1233]" italics="true" pageId="2" pageNumber="153">Cyatholaimus intermedius</emphasis>
</taxonomicName>
); Kattegat.
<treatmentCitation id="050810CD5A5FFFE8FBE784DEFAB1FB62" author="Gerlach" box="[1057,1284,1208,1234]" page="21" pageId="2" pageNumber="153" year="1953">
<bibRefCitation id="E0384B2D5A5FFFE8FBE784DEFB6EFB62" author="Gerlach" box="[1057,1243,1208,1234]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1953 a) Die Nematodenfauna der Uferzonen und des Kustengrundwassers am finnischen Meeresbusen. Acta Zoologica Fennica, 73, 1 - 32." type="journal article" year="1953" yearSuffix="a">Gerlach, 1953a</bibRefCitation>
: 21
</treatmentCitation>
22, Abb. 5; Baltic Sea.
<treatmentCitation id="050810CD5A5FFFE8FE8A8486FD98FB4A" author="Gerlach" box="[332,557,1248,1274]" page="127" pageId="2" pageNumber="153" year="1965">
<bibRefCitation id="E0384B2D5A5FFFE8FE8A8486FE43FB4A" author="Gerlach" box="[332,502,1248,1274]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1965) Freilebende Meeresnematoden aus der Gezeitenzone von Spitzbergen. Veroffentlichungen des Instituts fur Meeresforschungen in Bremerhaven, 9, 109 - 172." type="journal article" year="1965">Gerlach, 1965</bibRefCitation>
: 127
</treatmentCitation>
128, Abb. 9 ac; Spitzbergen.
<treatmentCitation id="050810CD5A5FFFE8FC5B8486FBC3FB4A" author="Bussau" box="[925,1142,1248,1274]" page="170" pageId="2" pageNumber="153" year="1990">
<bibRefCitation id="E0384B2D5A5FFFE8FC5B8486FB8AFB4A" author="Bussau" box="[925,1087,1248,1274]" pageId="2" pageNumber="173" refString="Bussau, C. (1990) Freilebende Nematoden aus Kustendunen und angrenzenden Biotopen der deutschen und danischen Kusten. I. Gesamtuberblick und Chromadorida (Nematoda). Zoologischer Anzeiger (Jena), 225 (3 / 4), 161 - 188." type="journal article" year="1990">Bussau, 1990</bibRefCitation>
: 170
</treatmentCitation>
172, Abb. 3 AC; North Sea, coastal dunes.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518556FBA7FAC2" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF518556FBA7FAC2" pageId="2" pageNumber="153">
9
<taxonomicName id="43A94D5F5A5FFFE8FF018556FC02FAFA" authority="Coomans, Vincx &amp; Decraemer, 1985" authorityName="Coomans, Vincx &amp; Decraemer" authorityYear="1985" box="[199,951,1328,1354]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="lewisi">
<emphasis id="B6DDEACE5A5FFFE8FF018556FE58FAF9" box="[199,493,1328,1353]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus lewisi</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FE308556FC02FAFA" author="Coomans" box="[502,951,1328,1354]" pageId="2" pageNumber="173" refString="Coomans, A., Vincx, M. &amp; Decraemer, W. (1985) Nematodes from a freshwater pool on a coral island in the Solomon Islands. Hydrobiologia, 123, 265 - 281." type="journal article" year="1985">Coomans, Vincx &amp; Decraemer, 1985</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FC008556FABDFAFA" author="Coomans" box="[966,1288,1328,1354]" page="268" pageId="2" pageNumber="153" year="1985">
<bibRefCitation id="E0384B2D5A5FFFE8FC008556FB7EFAFA" author="Coomans" box="[966,1227,1328,1354]" pageId="2" pageNumber="173" refString="Coomans, A., Vincx, M. &amp; Decraemer, W. (1985) Nematodes from a freshwater pool on a coral island in the Solomon Islands. Hydrobiologia, 123, 265 - 281." type="journal article" year="1985">Coomans et al., 1985</bibRefCitation>
: 268
</treatmentCitation>
, fig. 2 AF;
<collectingCountry id="FCBE764C5A5FFFE8FF01853EFE3BFAC2" box="[199,398,1368,1394]" name="Solomon Islands" pageId="2" pageNumber="153">Solomon Islands</collectingCountry>
, freshwater pool on a coral island; no males described.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5185E6FB6BFA2A" blockId="2.[151,1439,768,2034]" box="[151,1246,1408,1434]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5185E6FB6BFA2A" box="[151,1246,1408,1434]" pageId="2" pageNumber="153">
10
<taxonomicName id="43A94D5F5A5FFFE8FF0185E6FD02FA2A" authority="Gerlach, 1956" authorityName="Gerlach" authorityYear="1956" box="[199,695,1408,1434]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="occultus">
<emphasis id="B6DDEACE5A5FFFE8FF0185E6FDB2FA29" box="[199,519,1408,1433]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus occultus</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FDC885E6FD02FA2A" author="Gerlach" box="[526,695,1408,1434]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1956) Diagnosen neuer Nematoden aus der Kieler Bucht. Kieler Meeresforschungen, 12, 85 - 109." type="journal article" year="1956">Gerlach, 1956</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD0585E6FC23FA2A" author="Gerlach" box="[707,918,1408,1434]" page="91" pageId="2" pageNumber="153" year="1956">
<bibRefCitation id="E0384B2D5A5FFFE8FD0585E6FCDBFA2A" author="Gerlach" box="[707,878,1408,1434]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1956) Diagnosen neuer Nematoden aus der Kieler Bucht. Kieler Meeresforschungen, 12, 85 - 109." type="journal article" year="1956">Gerlach, 1956</bibRefCitation>
: 91
</treatmentCitation>
92, Abb. 29 ac; Kiel Bay.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5185CEFAD6FA72" blockId="2.[151,1439,768,2034]" box="[151,1379,1448,1474]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5185CEFAD6FA72" box="[151,1379,1448,1474]" pageId="2" pageNumber="153">
11
<taxonomicName id="43A94D5F5A5FFFE8FF0185CEFCB9FA72" authority="Gerlach, 1955" authorityName="Gerlach" authorityYear="1955" box="[199,780,1448,1474]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="paucipapillatus">
<emphasis id="B6DDEACE5A5FFFE8FF0185CEFDEEFA71" box="[199,603,1448,1473]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus paucipapillatus</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FDA585CEFCB9FA72" author="Gerlach" box="[611,780,1448,1474]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1955) Zur Kenntnis der freilebenden marinen Nematoden von San Salvador. Zeitschrift fur Wissenschaftliche Zoologie, 158, 249 - 303." type="journal article" year="1955">Gerlach, 1955</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FCDE85CEFC4EFA72" author="Gerlach" box="[792,1019,1448,1474]" page="265" pageId="2" pageNumber="153" year="1955">
<bibRefCitation id="E0384B2D5A5FFFE8FCDE85CEFC77FA72" author="Gerlach" box="[792,962,1448,1474]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1955) Zur Kenntnis der freilebenden marinen Nematoden von San Salvador. Zeitschrift fur Wissenschaftliche Zoologie, 158, 249 - 303." type="journal article" year="1955">Gerlach, 1955</bibRefCitation>
: 265
</treatmentCitation>
267, Abb. 7 ae;
<collectingCountry id="FCBE764C5A5FFFE8FB1585CEFAEAFA72" box="[1235,1375,1448,1474]" name="El Salvador" pageId="2" pageNumber="153">El Salvador</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5185B6FCFFF9A2" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5185B6FCFFF9A2" pageId="2" pageNumber="153">
12
<taxonomicName id="43A94D5F5A5FFFE8FF0185B6FD61FA5A" authority="Riemann, 1966" authorityName="Riemann" authorityYear="1966" box="[199,724,1488,1514]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="pentodon">
<emphasis id="B6DDEACE5A5FFFE8FF0185B6FDA1FA59" box="[199,532,1488,1513]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus pentodon</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FDDA85B6FD61FA5A" author="Riemann" box="[540,724,1488,1514]" pageId="2" pageNumber="174" refString="Riemann, F. (1966) Die interstitielle Fauna im Elbe-Astuar. Verbreitung und Systematik // Archiv fur Hydrobiologie, Supplement Band 31 (1 / 2), 1 - 279." type="journal article" year="1966">Riemann, 1966</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD2785B6FC66FA5A" author="Riemann" box="[737,979,1488,1514]" page="125" pageId="2" pageNumber="153" year="1966">
<bibRefCitation id="E0384B2D5A5FFFE8FD2785B6FC2FFA5A" author="Riemann" box="[737,922,1488,1514]" pageId="2" pageNumber="174" refString="Riemann, F. (1966) Die interstitielle Fauna im Elbe-Astuar. Verbreitung und Systematik // Archiv fur Hydrobiologie, Supplement Band 31 (1 / 2), 1 - 279." type="journal article" year="1966">Riemann, 1966</bibRefCitation>
: 125
</treatmentCitation>
127, Abb. 31 ae; North Sea.
<treatmentCitation id="050810CD5A5FFFE8FA8F85B6FE0DF9A2" author="Platt" page="282" pageId="2" pageNumber="153" year="1988">
<bibRefCitation id="E0384B2D5A5FFFE8FA8F85B6FECBF9A2" author="Platt" pageId="2" pageNumber="174" refString="Platt, H. M. &amp; Warwick, R. M. (1988) Free-living marine nematodes. Part II. British Chromadorids. Pictorial keys to world genera and notes for the identification of British species. Synopses of the British Fauna, 38. Publushed for the Linnean Society of London and the Estuarine and Brackish Water Sciences Association. By E. G. Brill IDr. W. Backhuys. Leiden, New York, KObenhavn, Koln. 502 pp." type="book" year="1988">Platt &amp; Warwick, 1988</bibRefCitation>
: 282
</treatmentCitation>
, fig. 128; East and West
<collectingCountry id="FCBE764C5A5FFFE8FD1A859EFCF2F9A2" box="[732,839,1528,1554]" name="United Kingdom" pageId="2" pageNumber="153">Scotland</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF518646FB44F98A" blockId="2.[151,1439,768,2034]" box="[151,1265,1568,1594]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF518646FB44F98A" box="[151,1265,1568,1594]" pageId="2" pageNumber="153">
13
<taxonomicName id="43A94D5F5A5FFFE8FF018646FCE2F98A" authority="Wieser &amp; Hopper, 1967: 268" authorityName="Wieser &amp; Hopper" authorityPageNumber="268" authorityYear="1967" box="[199,855,1568,1594]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="pesavis">
<emphasis id="B6DDEACE5A5FFFE8FF018646FE4EF989" box="[199,507,1568,1593]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus pesavis</emphasis>
<treatmentCitation id="050810CD5A5FFFE8FDC58646FCE2F98A" author="Wieser" box="[515,855,1568,1594]" page="268" pageId="2" pageNumber="153" year="1967">
<bibRefCitation id="E0384B2D5A5FFFE8FDC58646FCABF98A" author="Wieser" box="[515,798,1568,1594]" pageId="2" pageNumber="174" refString="Wieser, W. &amp; Hopper, B. (1967) Marine nematodes of the east coast of North America. I. Florida. Bulletin of the Museum for Comparative Zoology 135 (5), 239 - 344." type="journal article" year="1967">Wieser &amp; Hopper, 1967</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
269, pl. XVI, fig. 32 ae; Florida.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF51862EFDCBF902" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF51862EFDCBF902" pageId="2" pageNumber="153">
14
<taxonomicName id="43A94D5F5A5FFFE8FF01862EFD68F9D2" authority="Butschli, 1874" authorityName="Butschli" authorityYear="1874" box="[199,733,1608,1634]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="proximus">
<emphasis id="B6DDEACE5A5FFFE8FF01862EFDA7F9D1" box="[199,530,1608,1633]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus proximus</emphasis>
(
<bibRefCitation id="E0384B2D5A5FFFE8FDE4862EFD61F9D2" author="Butschli" box="[546,724,1608,1634]" pageId="2" pageNumber="173" refString="Butschli, O. (1874) Zur Kenntnis der freilebenden Nematoden, insbesondere der des Kieler Hafens. Abhandlungen der Senckenbergischen Naturforschen Gesellschaft, 9, 236 - 292." type="journal article" year="1874">Bütschli, 1874</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD2F862EFC70F9D2" author="Butschli" box="[745,965,1608,1634]" page="49" pageId="2" pageNumber="153" year="1874">
<bibRefCitation id="E0384B2D5A5FFFE8FD2F862EFC2CF9D2" author="Butschli" box="[745,921,1608,1634]" pageId="2" pageNumber="173" refString="Butschli, O. (1874) Zur Kenntnis der freilebenden Nematoden, insbesondere der des Kieler Hafens. Abhandlungen der Senckenbergischen Naturforschen Gesellschaft, 9, 236 - 292." type="journal article" year="1874">Bütschli, 1874</bibRefCitation>
: 49
</treatmentCitation>
, fig. 30 ab (=
<taxonomicName id="43A94D5F5A5FFFE8FBB5862EFA3FF9D1" box="[1139,1418,1608,1633]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="proximus">
<emphasis id="B6DDEACE5A5FFFE8FBB5862EFA3FF9D1" box="[1139,1418,1608,1633]" italics="true" pageId="2" pageNumber="153">Cyatholaimus proximus</emphasis>
</taxonomicName>
); Kiel Bay.
<collectingCountry id="FCBE764C5A5FFFE8FEF88616FED5F93A" box="[318,352,1648,1674]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="153">De</collectingCountry>
<treatmentCitation id="050810CD5A5FFFE8FEAE8616FD90F93A" author="Man" box="[360,549,1648,1674]" page="239" pageId="2" pageNumber="153" year="1922">
<bibRefCitation id="E0384B2D5A5FFFE8FEAE8616FE59F93A" author="Man" box="[360,492,1648,1674]" pageId="2" pageNumber="173" refString="de Man, J. G. (1922) Vrijlevende Nematoden. In: H. C. Redeke (Ed.), Flora en Fauna der Zuiderzee, Te Helder (C. de Boer), pp. 214 - 261." type="book chapter" year="1922">
<collectingCountry id="FCBE764C5A5FFFE8FEAE8616FE15F93A" box="[360,416,1648,1674]" name="Isle of Man" pageId="2" pageNumber="153">Man</collectingCountry>
, 1922
</bibRefCitation>
: 239
</treatmentCitation>
240, fig. 28 a, b (=
<taxonomicName id="43A94D5F5A5FFFE8FCD68616FB93F939" box="[784,1062,1648,1673]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="proximus">
<emphasis id="B6DDEACE5A5FFFE8FCD68616FB93F939" box="[784,1062,1648,1673]" italics="true" pageId="2" pageNumber="153">Cyatholaimus proximus</emphasis>
</taxonomicName>
); North Sea.
<treatmentCitation id="050810CD5A5FFFE8FB008616FF46F902" author="Micoletzky" page="140" pageId="2" pageNumber="153" year="1924">
<bibRefCitation id="E0384B2D5A5FFFE8FB008616FA2DF93A" author="Micoletzky" box="[1222,1432,1648,1674]" lastPageId="2" lastPageNumber="174" pageId="2" pageNumber="173" refString="Micoletzky, H. (1924) Letzter Bericht uber freilebende Nematoden aus Suez. Sitzungsberichten der Akademie der Wissenschaften in Wien, Mathematische-naturwissenschaft Klasse, Abteilung 1, 133, 137 - 179." type="journal article" year="1924">Micoletzky, 1924</bibRefCitation>
: 140
</treatmentCitation>
(to subgenus
<taxonomicName id="43A94D5F5A5FFFE8FE5086FEFDD8F901" box="[406,621,1688,1713]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5FFFE8FE5086FEFDD8F901" box="[406,621,1688,1713]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5186A6FD94F89A" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5186A6FD94F89A" bridged="true" pageId="2" pageNumber="153">
15
<taxonomicName id="43A94D5F5A5FFFE8FF0186A6FCC5F96A" authority="Wieser &amp; Hopper, 1967" authorityName="Wieser &amp; Hopper" authorityYear="1967" box="[199,880,1728,1754]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="pugettensis">
<emphasis id="B6DDEACE5A5FFFE8FF0186A6FD85F969" box="[199,560,1728,1753]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus pugettensis</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FDF986A6FCC5F96A" author="Wieser" box="[575,880,1728,1754]" pageId="2" pageNumber="174" refString="Wieser, W. &amp; Hopper, B. (1967) Marine nematodes of the east coast of North America. I. Florida. Bulletin of the Museum for Comparative Zoology 135 (5), 239 - 344." type="journal article" year="1967">Wieser &amp; Hopper, 1967</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FC4486A6FBEDF96A" author="Wieser" box="[898,1112,1728,1754]" page="37" pageId="2" pageNumber="153" year="1959">
<bibRefCitation id="E0384B2D5A5FFFE8FC4486A6FB9DF96A" author="Wieser" box="[898,1064,1728,1754]" pageId="2" pageNumber="174" refString="Wieser, W. (1959) Free-living nematodes and other small invertebrates of Puget Sound beaches. Seattle (University of Washington Press), 179 pp." type="book" year="1959">Wieser, 1959</bibRefCitation>
: 37
</treatmentCitation>
38, fig. 35 ac (=
<taxonomicName id="43A94D5F5A5FFFE8FA8886A6FD39F8B2" authority="Wieser, 1959" authorityName="Wieser" authorityYear="1959" class="Adenophorea" family="Cyatholaimidae" genus="Longicyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A5FFFE8FA8886A6FE5DF8B1" italics="true" pageId="2" pageNumber="153">Longicyatholaimus quadriseta</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FE36868EFD39F8B2" author="Wieser" box="[496,652,1768,1794]" pageId="2" pageNumber="174" refString="Wieser, W. (1959) Free-living nematodes and other small invertebrates of Puget Sound beaches. Seattle (University of Washington Press), 179 pp." type="book" year="1959">Wieser, 1959</bibRefCitation>
</taxonomicName>
, nec
<taxonomicName id="43A94D5F5A5FFFE8FD0F868EFBB3F8B2" authority="Wieser 1954" authorityName="Wieser" authorityYear="1954" box="[713,1030,1768,1794]" class="Adenophorea" family="Cyatholaimidae" genus="Longicyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A5FFFE8FD0F868EFCD3F8B1" box="[713,870,1768,1793]" italics="true" pageId="2" pageNumber="153">L. quadriseta</emphasis>
Wieser 1954
</taxonomicName>
); Puget Sound, Washington,
<collectingCountry id="FCBE764C5A5FFFE8FA9C868EFA2CF8B2" box="[1370,1433,1768,1794]" name="United States of America" pageId="2" pageNumber="153">USA</collectingCountry>
.
<treatmentCitation id="050810CD5A5FFFE8FF018776FDA8F89A" author="Wieser" box="[199,541,1808,1834]" page="265" pageId="2" pageNumber="153" year="1967">
<bibRefCitation id="E0384B2D5A5FFFE8FF018776FE56F89A" author="Wieser" box="[199,483,1808,1834]" pageId="2" pageNumber="174" refString="Wieser, W. &amp; Hopper, B. (1967) Marine nematodes of the east coast of North America. I. Florida. Bulletin of the Museum for Comparative Zoology 135 (5), 239 - 344." type="journal article" year="1967">Wieser &amp; Hopper, 1967</bibRefCitation>
: 265
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF51875EFC10F8CA" blockId="2.[151,1439,768,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF51875EFC10F8CA" pageId="2" pageNumber="153">
16
<taxonomicName id="43A94D5F5A5FFFE8FF01875EFD44F8E2" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[199,753,1848,1874]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A5FFFE8FF01875EFD9CF8E1" box="[199,553,1848,1873]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus quadriseta</emphasis>
(Wieser, 1954)
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FCC2875EFC5DF8E2" author="Wieser" box="[772,1000,1848,1874]" page="13" pageId="2" pageNumber="153" year="1954">
<bibRefCitation id="E0384B2D5A5FFFE8FCC2875EFC03F8E2" author="Wieser" box="[772,950,1848,1874]" pageId="2" pageNumber="174" refString="Wieser, W. (1954 a) Free-living marine nematodes. II. Chromadoroidea. Lunds Universitets Arsskrift. N. F. Avd. 2, 50 (16), 1 - 148." type="journal article" year="1954" yearSuffix="a">Wieser, 1954a</bibRefCitation>
: 13
</treatmentCitation>
, fig. 98 a, b (=
<taxonomicName id="43A94D5F5A5FFFE8FB7D875EFEF1F8C9" class="Adenophorea" family="Cyatholaimidae" genus="Longicyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A5FFFE8FB7D875EFEF1F8C9" italics="true" pageId="2" pageNumber="153">Longicyatholaimus quadriseta</emphasis>
</taxonomicName>
);
<collectingCountry id="FCBE764C5A5FFFE8FE9B8706FE15F8CA" box="[349,416,1888,1914]" name="Chile" pageId="2" pageNumber="153">Chile</collectingCountry>
.
<treatmentCitation id="050810CD5A5FFFE8FE6A8706FDCDF8CA" author="Hopper" box="[428,632,1888,1914]" page="69" pageId="2" pageNumber="153" year="1972">
<bibRefCitation id="E0384B2D5A5FFFE8FE6A8706FDE5F8CA" author="Hopper" box="[428,592,1888,1914]" pageId="2" pageNumber="173" refString="Hopper, B. E. (1972) Free-living marine nematodes from Biscayne Bay, Florida IV. Cyatholaimidae: On the occurence of Marylina n. gen. and Longicyatholaimus Micoletzky, 1924 in Biscayne Bay, with a description of L. longicaudatus (De Man, 1876) from the type locality. Zoologischer Anzeiger (Jena), 189, 64 - 88." type="journal article" year="1972">Hopper, 1972</bibRefCitation>
: 69
</treatmentCitation>
(=
<taxonomicName id="43A94D5F5A5FFFE8FD5C8706FC26F8C9" box="[666,915,1888,1913]" class="Arachnida" family="Dictynidae" genus="Marilynia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A5FFFE8FD5C8706FC26F8C9" box="[666,915,1888,1913]" italics="true" pageId="2" pageNumber="153">Marilynia quadriseta</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5187EEFB05F812" blockId="2.[151,1439,768,2034]" box="[151,1200,1928,1954]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5187EEFB05F812" box="[151,1200,1928,1954]" pageId="2" pageNumber="153">
17
<taxonomicName id="43A94D5F5A5FFFE8FF0187EEFD09F812" authority="Gerlach, 1964" authorityName="Gerlach" authorityYear="1964" box="[199,700,1928,1954]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="rotundus">
<emphasis id="B6DDEACE5A5FFFE8FF0187EEFDBEF811" box="[199,523,1928,1953]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus rotundus</emphasis>
Gerlach, 1964
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD0E87EEFC1FF812" author="Gerlach" box="[712,938,1928,1954]" page="25" pageId="2" pageNumber="153" year="1964">
<bibRefCitation id="E0384B2D5A5FFFE8FD0E87EEFC35F812" author="Gerlach" box="[712,896,1928,1954]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1964 a) Freilebende Nematoden aus dem Roten Meer. Kieler Meeresforschungen, 20 (Sonderheft), 18 - 34." type="journal article" year="1964" yearSuffix="a">Gerlach, 1964a</bibRefCitation>
: 25
</treatmentCitation>
, Abb. 2 de; Red Sea.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5187D6FB0AF87A" blockId="2.[151,1439,768,2034]" box="[151,1215,1968,1994]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5187D6FB0AF87A" box="[151,1215,1968,1994]" pageId="2" pageNumber="153">
18
<taxonomicName id="43A94D5F5A5FFFE8FF0187D6FD15F87A" authority="Gerlach, 1967" authorityName="Gerlach" authorityYear="1967" box="[199,672,1968,1994]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="saradi">
<emphasis id="B6DDEACE5A5FFFE8FF0187D6FE5AF879" box="[199,495,1968,1993]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus saradi</emphasis>
<bibRefCitation id="E0384B2D5A5FFFE8FE3187D6FD15F87A" author="Gerlach" box="[503,672,1968,1994]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1967) Freilebende Meeres-Nematoden von den Sarso-Inseln (Rotes Meer). Meteor-Forschungergebnisse (D), 2, 19 - 43." type="journal article" year="1967">Gerlach, 1967</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD6A87D6FC35F87A" author="Gerlach" box="[684,896,1968,1994]" page="30" pageId="2" pageNumber="153" year="1967">
<bibRefCitation id="E0384B2D5A5FFFE8FD6A87D6FCE3F87A" author="Gerlach" box="[684,854,1968,1994]" pageId="2" pageNumber="173" refString="Gerlach, S. A. (1967) Freilebende Meeres-Nematoden von den Sarso-Inseln (Rotes Meer). Meteor-Forschungergebnisse (D), 2, 19 - 43." type="journal article" year="1967">Gerlach, 1967</bibRefCitation>
: 30
</treatmentCitation>
31, Abb. 15 ae; Red Sea.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5FFFE8FF5187BEFAF0F842" blockId="2.[151,1439,768,2034]" box="[151,1349,2008,2034]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="A4B911F25A5FFFE8FF5187BEFAF0F842" box="[151,1349,2008,2034]" pageId="2" pageNumber="153">
19
<taxonomicName id="43A94D5F5A5FFFE8FF0187BEFD0EF842" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[199,699,2008,2034]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="2" pageNumber="153" phylum="Nematoda" rank="species" species="separatus">
<emphasis id="B6DDEACE5A5FFFE8FF0187BEFDA3F841" box="[199,534,2008,2033]" italics="true" pageId="2" pageNumber="153">Paracyatholaimus separatus</emphasis>
Wieser, 1954
</taxonomicName>
.
<treatmentCitation id="050810CD5A5FFFE8FD0187BEFC1EF842" author="Wieser" box="[711,939,2008,2034]" page="194" pageId="2" pageNumber="153" year="1954">
<bibRefCitation id="E0384B2D5A5FFFE8FD0187BEFCC1F842" author="Wieser" box="[711,884,2008,2034]" pageId="2" pageNumber="174" refString="Wieser, W. (1954 b) Beitrage zur Kenntnis der Nematoden submariner Hohlen. Ergebnisse der osterreichischen Tyrrhenia-Expedition 1952, Teil II. Osterreichische zoologische Zeitschrift, 5, 172 - 230." type="journal article" year="1954" yearSuffix="b">Wieser, 1954b</bibRefCitation>
: 194
</treatmentCitation>
196, Abb. 12 ac; Mediterranean.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF5180FEFA84FF02" blockId="3.[151,1437,152,538]" box="[151,1329,152,178]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="A4B911F25A5EFFE9FF5180FEFA84FF02" box="[151,1329,152,178]" pageId="3" pageNumber="154">
20
<taxonomicName id="43A94D5F5A5EFFE9FF0180FEFD76FF02" authority="Jensen, 1985" authorityName="Jensen" authorityYear="1985" box="[199,707,152,178]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="spinulosus">
<emphasis id="B6DDEACE5A5EFFE9FF0180FEFD95FF01" box="[199,544,152,177]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus spinulosus</emphasis>
<bibRefCitation id="E0384B2D5A5EFFE9FDEE80FEFD76FF02" author="Jensen" box="[552,707,152,178]" pageId="3" pageNumber="173" refString="Jensen, P. (1985) The nematode fauna in the sulphide-rich brine seep and adjacent bottoms of the East Flower Garden, NW Gulf of Mexico. I. Chromadorida. Zoologica Scripta, 14 (4), 247 - 263." type="journal article" year="1985">Jensen, 1985</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5EFFE9FD0880FEFC14FF02" author="Jensen" box="[718,929,152,178]" page="253" pageId="3" pageNumber="154" year="1985">
<bibRefCitation id="E0384B2D5A5EFFE9FD0880FEFCDFFF02" author="Jensen" box="[718,874,152,178]" pageId="3" pageNumber="173" refString="Jensen, P. (1985) The nematode fauna in the sulphide-rich brine seep and adjacent bottoms of the East Flower Garden, NW Gulf of Mexico. I. Chromadorida. Zoologica Scripta, 14 (4), 247 - 263." type="journal article" year="1985">Jensen, 1985</bibRefCitation>
: 253
</treatmentCitation>
254, fig. 7 AF; Gulf of
<collectingCountry id="FCBE764C5A5EFFE9FB0980FEFA98FF02" box="[1231,1325,152,178]" name="Mexico" pageId="3" pageNumber="154">Mexico</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF5180A6FB20FF6A" blockId="3.[151,1437,152,538]" box="[151,1173,192,218]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="A4B911F25A5EFFE9FF5180A6FB20FF6A" box="[151,1173,192,218]" pageId="3" pageNumber="154">
21
<taxonomicName id="43A94D5F5A5EFFE9FF0880A6FD2FFF6A" authority="Wieser, 1954" authorityName="Wieser" authorityYear="1954" box="[206,666,192,218]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="ternus">
<emphasis id="B6DDEACE5A5EFFE9FF0880A6FE40FF69" box="[206,501,192,217]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus ternus</emphasis>
Wieser, 1954
</taxonomicName>
.
<treatmentCitation id="050810CD5A5EFFE9FD6380A6FCCCFF6A" author="Wieser" box="[677,889,192,218]" page="27" pageId="3" pageNumber="154" year="1954">
<bibRefCitation id="E0384B2D5A5EFFE9FD6380A6FCE4FF6A" author="Wieser" box="[677,849,192,218]" pageId="3" pageNumber="174" refString="Wieser, W. (1954 a) Free-living marine nematodes. II. Chromadoroidea. Lunds Universitets Arsskrift. N. F. Avd. 2, 50 (16), 1 - 148." type="journal article" year="1954" yearSuffix="a">Wieser, 1954a</bibRefCitation>
: 27
</treatmentCitation>
28, fig. 108 ae;
<collectingCountry id="FCBE764C5A5EFFE9FB8B80A6FB25FF6A" box="[1101,1168,192,218]" name="Chile" pageId="3" pageNumber="154">Chile</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF51808EFC5FFE7A" blockId="3.[151,1437,152,538]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="A4B911F25A5EFFE9FF51808EFC5FFE7A" bridged="true" pageId="3" pageNumber="154">
22
<taxonomicName id="43A94D5F5A5EFFE9FF01808EFD77FEB2" authority="Cobb, 1914" authorityName="Cobb" authorityYear="1914" box="[199,706,232,258]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="truncatus">
<emphasis id="B6DDEACE5A5EFFE9FF01808EFDA6FEB1" box="[199,531,232,257]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus truncatus</emphasis>
(
<bibRefCitation id="E0384B2D5A5EFFE9FDE2808EFD1AFEB2" author="Cobb" box="[548,687,232,258]" pageId="3" pageNumber="173" refString="Cobb, N. A. (1914) The North American free-living fresh-water nematodes. Transactions of the American Microscopical Society, 33, 69 - 119 (reprinted in Contributions to a Science of Nematology (Baltimore), 2, 35 - 99)." type="journal article" year="1914">Cobb, 1914</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="050810CD5A5EFFE9FD09808EFC33FEB2" author="Cobb" box="[719,902,232,258]" page="60" pageId="3" pageNumber="154" year="1914">
<bibRefCitation id="E0384B2D5A5EFFE9FD09808EFCE9FEB2" author="Cobb" box="[719,860,232,258]" pageId="3" pageNumber="173" refString="Cobb, N. A. (1914) The North American free-living fresh-water nematodes. Transactions of the American Microscopical Society, 33, 69 - 119 (reprinted in Contributions to a Science of Nematology (Baltimore), 2, 35 - 99)." type="journal article" year="1914">Cobb, 1914</bibRefCitation>
: 60
</treatmentCitation>
62, fig. 17 (=
<taxonomicName id="43A94D5F5A5EFFE9FBE9808EFAFDFEB1" box="[1071,1352,232,257]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="truncatus">
<emphasis id="B6DDEACE5A5EFFE9FBE9808EFAFDFEB1" box="[1071,1352,232,257]" italics="true" pageId="3" pageNumber="154">Cyatholaimus truncatus</emphasis>
</taxonomicName>
); Florida, fresh or brackish water.
<treatmentCitation id="050810CD5A5EFFE9FDD58176FCA9FE9A" author="Micoletzky" box="[531,796,272,298]" page="377" pageId="3" pageNumber="154" year="1922">
<bibRefCitation id="E0384B2D5A5EFFE9FDD58176FD50FE9A" author="Micoletzky" box="[531,741,272,298]" pageId="3" pageNumber="173" refString="Micoletzky, H. (1922) Die freilebenden Erdnematoden. Archiv fur Naturgeschichte, 87 A, 1 - 650." type="journal article" year="1922">Micoletzky, 1922</bibRefCitation>
: 377
</treatmentCitation>
(to subgenus
<taxonomicName id="43A94D5F5A5EFFE9FC068176FB22FE99" box="[960,1175,272,297]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FC068176FB22FE99" box="[960,1175,272,297]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
). Finding of
<bibRefCitation id="E0384B2D5A5EFFE9FAF48176FEA1FEE2" author="Riemann" pageId="3" pageNumber="174" refString="Riemann, F. (1970) Freilebende Nematoden aus dem Grenzbereich Meer-Suβ-Wasser in Kolumbien, Sudamerika. Ver offentlichungen des Instituts fur Meeresforschungen in Bremerhaven, 12, 365 - 412." type="journal article" year="1970">Riemann (1970)</bibRefCitation>
in a similar habitat at the Caribbean coast of
<collectingCountry id="FCBE764C5A5EFFE9FCEE815EFC29FEE2" box="[808,924,312,338]" name="Colombia" pageId="3" pageNumber="154">Colombia</collectingCountry>
may be doubtful because of the lesser male body length (1050
<emphasis id="B6DDEACE5A5EFFE9FE618106FE75FEC9" box="[423,448,352,377]" italics="true" pageId="3" pageNumber="154">vs</emphasis>
1600
<emphasis id="B6DDEACE5A5EFFE9FDCC8106FDACFEC9" box="[522,537,352,377]" italics="true" pageId="3" pageNumber="154">µ</emphasis>
m), strong conical inner labial papillae, other pattern of four preanal supplementary organs (both posteriormost and penultimate organs are situated at the level of spicules while those of the
<typeStatus id="5B12887E5A5EFFE9FE9181D6FE3FFE7A" box="[343,394,432,458]" pageId="3" pageNumber="154">type</typeStatus>
specimen are separated by much greater distances).
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF5181BEFAFAFE42" blockId="3.[151,1437,152,538]" box="[151,1359,472,498]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="A4B911F25A5EFFE9FF5181BEFAFAFE42" box="[151,1359,472,498]" pageId="3" pageNumber="154">
23
<taxonomicName id="43A94D5F5A5EFFE9FF0181BEFC1EFE42" authority="Sharma &amp; Vincx, 1982: 276" authorityName="Sharma &amp; Vincx" authorityPageNumber="276" authorityYear="1982" box="[199,939,472,498]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="vancouverensis">
<emphasis id="B6DDEACE5A5EFFE9FF0181BEFDECFE41" box="[199,601,472,497]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus vancouverensis</emphasis>
<treatmentCitation id="050810CD5A5EFFE9FDA681BEFC1EFE42" author="Sharma" box="[608,939,472,498]" page="276" pageId="3" pageNumber="154" year="1982">
<bibRefCitation id="E0384B2D5A5EFFE9FDA681BEFCC6FE42" author="Sharma" box="[608,883,472,498]" pageId="3" pageNumber="174" refString="Sharma, J. &amp; Vincx, M. (1982) Cyatholaimidae (Nematoda) from the Canadian Pacific Coast. Canadian Journal of Zoology, 60, 271 - 280." type="journal article" year="1982">Sharma &amp; Vincx, 1982</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
278, fig. 1523; British
<collectingCountry id="FCBE764C5A5EFFE9FB1581BEFAFEFE42" box="[1235,1355,472,498]" name="United States of America" pageId="3" pageNumber="154">Columbia</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF518266FB63FDAA" blockId="3.[151,1437,152,538]" box="[151,1238,512,538]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="A4B911F25A5EFFE9FF518266FB63FDAA" box="[151,1238,512,538]" pageId="3" pageNumber="154">
24
<taxonomicName id="43A94D5F5A5EFFE9FF018266FD01FDAA" authority="Gerlach, 1957" authorityName="Gerlach" authorityYear="1957" box="[199,692,512,538]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="vitraeus">
<emphasis id="B6DDEACE5A5EFFE9FF018266FDB6FDA9" box="[199,515,512,537]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus vitraeus</emphasis>
<bibRefCitation id="E0384B2D5A5EFFE9FDCD8266FD01FDAA" author="Gerlach" box="[523,692,512,538]" pageId="3" pageNumber="173" refString="Gerlach, S. A. (1957) Marine Nematoden aus dem Mangrove-Gebiet von Cananeia (Brasilianische Meeres-Nematoden III). Akademie der Wissenschaften und Literatur, Mainz. Abhandlungen der Mathemathik-naturwissenschaftlichen Klasse, 5, 129 - 176." type="journal article" year="1957">Gerlach, 1957</bibRefCitation>
</taxonomicName>
.
<treatmentCitation id="050810CD5A5EFFE9FD068266FC11FDAA" author="Gerlach" box="[704,932,512,538]" page="140" pageId="3" pageNumber="154" year="1957">
<bibRefCitation id="E0384B2D5A5EFFE9FD068266FCDFFDAA" author="Gerlach" box="[704,874,512,538]" pageId="3" pageNumber="173" refString="Gerlach, S. A. (1957) Marine Nematoden aus dem Mangrove-Gebiet von Cananeia (Brasilianische Meeres-Nematoden III). Akademie der Wissenschaften und Literatur, Mainz. Abhandlungen der Mathemathik-naturwissenschaftlichen Klasse, 5, 129 - 176." type="journal article" year="1957">Gerlach, 1957</bibRefCitation>
: 140
</treatmentCitation>
141, Abb. 21 lo;
<collectingCountry id="FCBE764C5A5EFFE9FB408266FB67FDAA" box="[1158,1234,512,538]" name="Brazil" pageId="3" pageNumber="154">Brazil</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5EFFE9FF518234FB14FB9F" pageId="3" pageNumber="154" type="discussion">
<paragraph id="841636DC5A5EFFE9FF518234FC33FDDB" blockId="3.[151,1437,594,1471]" box="[151,902,594,620]" pageId="3" pageNumber="154">
Remarks to species composition of the genus
<taxonomicName id="43A94D5F5A5EFFE9FD698234FC33FDDB" box="[687,902,594,619]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FD698234FC33FDDB" box="[687,902,594,619]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="841636DC5A5EFFE9FF00821CFC50FD54" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
Four species originally assigned to
<taxonomicName id="43A94D5F5A5EFFE9FDB1821CFCFBFD23" box="[631,846,634,659]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FDB1821CFCFBFD23" box="[631,846,634,659]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
, i.e.
<emphasis id="B6DDEACE5A5EFFE9FC49821CFBE2FD23" box="[911,1111,634,659]" italics="true" pageId="3" pageNumber="154">choanolaimoides</emphasis>
Stekhoven, 1942,
<emphasis id="B6DDEACE5A5EFFE9FAFB821CFA2EFD23" box="[1341,1435,634,659]" italics="true" pageId="3" pageNumber="154">effilatus</emphasis>
Stekhoven, 1946,
<emphasis id="B6DDEACE5A5EFFE9FEAD82C4FE04FD0B" box="[363,433,674,699]" italics="true" pageId="3" pageNumber="154">major</emphasis>
Kreis, 1928 and
<emphasis id="B6DDEACE5A5EFFE9FDBA82C4FCB4FD0B" box="[636,769,674,699]" italics="true" pageId="3" pageNumber="154">tyrrhenicus</emphasis>
Brunetti, 1949, were later transferred to other genera by various authors (
<bibRefCitation id="E0384B2D5A5EFFE9FE9C82ACFD3EFD54" author="Gerlach" box="[346,651,714,740]" pageId="3" pageNumber="173" refString="Gerlach, S. A. &amp; Riemann, F. (1973) The Bremerhaven checklist of aquatic nematodes. A catalogue of Nematoda Adenophorea excluding the Dorylaimida. Part 1. Veroffentlichungen des Instituts fur Meeresforschung in Bremerhaven, Supplement 4, Heft 1, 404 pp." type="book" year="1973">Gerlach &amp; Riemann 1973</bibRefCitation>
) and are not considered here.
</paragraph>
<paragraph id="841636DC5A5EFFE9FF008294FE46FC3E" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
Twenty­seven species of
<taxonomicName id="43A94D5F5A5EFFE9FDD68294FD4EFCBE" box="[528,763,754,782]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FDD68294FD4EFCBE" box="[528,763,754,782]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
are enumerated in the NeMys (
<bibRefCitation id="E0384B2D5A5EFFE9FB5B8294FA21FCBE" author="Deprez" box="[1181,1428,754,782]" pageId="3" pageNumber="173" refString="Deprez, T. and oth. (2005) NeMys. World Wide Web electronic publication. Available from: http: // nemys. ugent. be (Version 8 / 2007)." type="journal article" year="2005">Deprez et al. 2005</bibRefCitation>
) where
<taxonomicName id="43A94D5F5A5EFFE9FF28837BFDDEFC89" authority="Allgen, 1935" authorityName="Allgen" authorityYear="1935" box="[238,619,797,825]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="parasaveljevi">
<emphasis id="B6DDEACE5A5EFFE9FF28837BFE0FFC89" box="[238,442,797,825]" italics="true" pageId="3" pageNumber="154">P. parasaveljevi</emphasis>
Allgén, 1935
</taxonomicName>
is designated
<emphasis id="B6DDEACE5A5EFFE9FCE2837BFBA1FC89" box="[804,1044,797,825]" italics="true" pageId="3" pageNumber="154">species inquirenda</emphasis>
. Here four more other species are considered as
<emphasis id="B6DDEACE5A5EFFE9FEBA832EFDCEFCD4" box="[380,635,840,868]" italics="true" pageId="3" pageNumber="154">species inquirendae</emphasis>
because of insufficient
<typeStatus id="5B12887E5A5EFFE9FC68832EFC50FCD4" box="[942,997,840,868]" pageId="3" pageNumber="154">type</typeStatus>
material lacking adult stages and/ or incomplete descriptions:
</paragraph>
<paragraph id="841636DC5A5EFFE9FF0083FBFB14FB9F" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FF0083FBFD4DFC07" authority="Hoeppli et Chu, 1932" authorityName="Hoeppli et Chu" authorityYear="1932" box="[198,760,925,951]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="chungsani">
<emphasis id="B6DDEACE5A5EFFE9FF0083FBFE5FFC06" box="[198,490,925,950]" italics="true" pageId="3" pageNumber="154">Cyatholaimus chungsani</emphasis>
Hoeppli et Chu, 1932
</taxonomicName>
;
<collectingCountry id="FCBE764C5A5EFFE9FCC383FBFCFAFC07" box="[773,847,925,951]" name="China" pageId="3" pageNumber="154">China</collectingCountry>
. The species was shifted to
<taxonomicName id="43A94D5F5A5EFFE9FB5983FBFAC3FC06" box="[1183,1398,925,950]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FB5983FBFAC3FC06" box="[1183,1398,925,950]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
by
<bibRefCitation id="E0384B2D5A5EFFE9FF5183A3FE9CFC6F" author="Meyl" box="[151,297,965,991]" pageId="3" pageNumber="173" refString="Meyl, A. H. (1954) Beitrage zur Kenntnis der Nematodenfauna vulkanisch erhitzter Biotope. III. Mitteilung. Nematoden aus der Mischungszone strandnaher, heisser Suβwasserquellen mit dem Meerwasser auf der Insel Ischia. Zeitschrift fur Morphologie und Okologie der Tiere, 42, 421 - 448." type="journal article" year="1954">Meyl (1954)</bibRefCitation>
and, missing in the NeMys list, is here considered as
<emphasis id="B6DDEACE5A5EFFE9FC5A83A3FBC2FC6E" box="[924,1143,965,990]" italics="true" pageId="3" pageNumber="154">species inquirenda</emphasis>
because of the very poor original description even lacking details actually indicating its affiliation with
<taxonomicName id="43A94D5F5A5EFFE9FBF2838BFABEFBB6" box="[1076,1291,1005,1030]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FBF2838BFABEFBB6" box="[1076,1291,1005,1030]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
. This is one of three species ascribed to
<taxonomicName id="43A94D5F5A5EFFE9FE1D8473FD07FB9E" box="[475,690,1045,1070]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FE1D8473FD07FB9E" box="[475,690,1045,1070]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
which were found in fresh­water habitats.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5EFFE9FF00845BFEE5FB47" pageId="3" pageNumber="154" type="reference_group">
<paragraph id="841636DC5A5EFFE9FF00845BFB9AFBCF" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FF00845BFCEBFBE7" authority="(Cobb, 1898) Micoletzky, 1924" authorityName="(Cobb, 1898) Micoletzky" authorityYear="1924" box="[198,862,1085,1111]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="exilis">
<emphasis id="B6DDEACE5A5EFFE9FF00845BFE56FBE6" box="[198,483,1085,1110]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus exilis</emphasis>
(Cobb, 1898)
<bibRefCitation id="E0384B2D5A5EFFE9FD56845BFCEBFBE7" author="Micoletzky" box="[656,862,1085,1111]" lastPageId="3" lastPageNumber="174" pageId="3" pageNumber="173" refString="Micoletzky, H. (1924) Letzter Bericht uber freilebende Nematoden aus Suez. Sitzungsberichten der Akademie der Wissenschaften in Wien, Mathematische-naturwissenschaft Klasse, Abteilung 1, 133, 137 - 179." type="journal article" year="1924">Micoletzky, 1924</bibRefCitation>
</taxonomicName>
(=
<emphasis id="B6DDEACE5A5EFFE9FC4E845BFBF5FBE6" box="[904,1088,1085,1110]" italics="true" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FC4E845BFB9FFBE6" box="[904,1066,1085,1110]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">Cyatholaimus</taxonomicName>
e
</emphasis>
. Cobb, 1898);
<collectingCountry id="FCBE764C5A5EFFE9FB32845BFAD6FBE7" box="[1268,1379,1085,1111]" name="Australia" pageId="3" pageNumber="154">Australia</collectingCountry>
. The original description is based on only a female specimen and lacks illustrations.
</paragraph>
<paragraph id="841636DC5A5EFFE9FF0084EBFEE5FB47" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FF0084EBFC70FB17" authority="(Allgen, 1935) Wieser, 1954" authorityName="(Allgen, 1935) Wieser" authorityYear="1954" box="[198,965,1165,1191]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="oistospiculoides">
<emphasis id="B6DDEACE5A5EFFE9FF0084EBFDD1FB16" box="[198,612,1165,1190]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus oistospiculoides</emphasis>
(Allgén, 1935) Wieser, 1954
</taxonomicName>
(=
<emphasis id="B6DDEACE5A5EFFE9FC3284EBFB69FB16" box="[1012,1244,1165,1190]" italics="true" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FC3284EBFB77FB16" box="[1012,1218,1165,1190]" class="Adenophorea" family="Cyatholaimidae" genus="Paracanthonchus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">Paracanthonchus</taxonomicName>
o
</emphasis>
. Allgén, 1935);
<collectingCountry id="FCBE764C5A5EFFE9FF5184D3FF40FB7F" box="[151,245,1205,1231]" name="Norway" pageId="3" pageNumber="154">Norway</collectingCountry>
Sea. The original description and drawings based on a single male specimen are too poor to determine the genus.
</paragraph>
</subSubSection>
<subSubSection id="CCB365575A5EFFECFF008563FB93FEFF" lastPageId="6" lastPageNumber="157" pageId="3" pageNumber="154" type="description">
<paragraph id="841636DC5A5EFFE9FF008563FEE7FAF7" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FF008563FD5DFAAF" authority="Allgen, 1935" authorityName="Allgen" authorityYear="1935" box="[198,744,1285,1311]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="parasaveljevi">
<emphasis id="B6DDEACE5A5EFFE9FF008563FDF6FAAE" box="[198,579,1285,1310]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus parasaveljevi</emphasis>
Allgén, 1935
</taxonomicName>
;
<collectingCountry id="FCBE764C5A5EFFE9FD328563FCE7FAAF" box="[756,850,1285,1311]" name="Norway" pageId="3" pageNumber="154">Norway</collectingCountry>
Sea.
<bibRefCitation id="E0384B2D5A5EFFE9FC528563FBFCFAAF" author="Wieser" box="[916,1097,1285,1311]" pageId="3" pageNumber="174" refString="Wieser, W. (1954 a) Free-living marine nematodes. II. Chromadoroidea. Lunds Universitets Arsskrift. N. F. Avd. 2, 50 (16), 1 - 148." type="journal article" year="1954" yearSuffix="a">Wieser (1954a)</bibRefCitation>
qualified the species as
<emphasis id="B6DDEACE5A5EFFE9FAAD8563FEF9FAF6" italics="true" pageId="3" pageNumber="154">species inquirenda</emphasis>
.
</paragraph>
<paragraph id="841636DC5A5EFFE9FF008533FD10FA0F" blockId="3.[151,1437,594,1471]" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FF008533FC08FADF" authority="(Allgen, 1951) Wieser, 1954" authorityName="(Allgen, 1951) Wieser" authorityYear="1954" box="[198,957,1365,1391]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="species" species="tenuispiculum">
<emphasis id="B6DDEACE5A5EFFE9FF008533FDE5FADE" box="[198,592,1365,1390]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus tenuispiculum</emphasis>
(Allgén, 1951) Wieser, 1954
</taxonomicName>
(=
<emphasis id="B6DDEACE5A5EFFE9FC348533FB63FADE" box="[1010,1238,1365,1390]" italics="true" pageId="3" pageNumber="154">
<taxonomicName id="43A94D5F5A5EFFE9FC348533FB74FADE" box="[1010,1217,1365,1390]" class="Adenophorea" family="Cyatholaimidae" genus="Paracanthonchus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">Paracanthonchus</taxonomicName>
t
</emphasis>
. Allgén, 1951); Hawaii. The original description and drawings are based on a single male specimen and lacks many details such as amphids and length of anterior setae.
</paragraph>
<paragraph id="841636DC5A5EFFE9FF518593FC63F9BF" blockId="3.[151,1435,1525,1871]" box="[151,982,1525,1551]" pageId="3" pageNumber="154">
Illustrated guide for species of
<taxonomicName id="43A94D5F5A5EFFE9FDC58593FD6FF9BE" box="[515,730,1525,1550]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FDC58593FD6FF9BE" box="[515,730,1525,1550]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
(
<figureCitation id="1C922A595A5EFFE9FD2F8593FC86F9BF" box="[745,819,1525,1551]" captionStart="FIGURE 1" captionStartId="4.[151,244,1916,1937]" captionTargetBox="[190,1405,184,1901]" captionTargetId="figure@4.[163,1423,168,1921]" captionTargetPageId="4" captionText="FIGURE 1. Pictorial guide for valid species of Paracyatholaimus. Part 1, beginning. Simplified images of species come from the published descriptions: arcticus — Kreis, 1963; botosaneanui — Andrássy, 1973; chilensis — Gerlach, 1953 b; diva — orig.; dubiosus — Meyl, 1954; duplicatus — Gerlach, 1964 b; helicellus — Wieser, 1954 a; intermedius — Gerlach, 1953 a; lewisi — Coomans et al., 1985; occultus — Gerlach, 1956; paucipapillatus — Gerlach, 1955; pentodon — Riemann, 1966." httpUri="https://zenodo.org/record/183776/files/figure.png" pageId="3" pageNumber="154">Figs 1</figureCitation>
&amp;
<figureCitation id="1C922A595A5EFFE9FC9E8593FCDCF9BF" box="[856,873,1525,1551]" captionStart="FIGURE 2" captionStartId="5.[151,244,1911,1932]" captionTargetBox="[182,1346,182,1910]" captionTargetId="figure@5.[163,1423,163,1916]" captionTargetPageId="5" captionText="FIGURE 2. Pictorial guide for valid species of Paracyatholaimus. Part 2, ending. Simplified images of species come from the published descriptions: pesavis — Wieser &amp; Hopper, 1967; proximus — de Man, 1922; pugettensis — Wieser, 1959; quadriseta — Wieser, 1954 a; rotundus — Gerlach, 1964 a; saradi — Gerlach, 1967; separatus — Wieser, 1954 b; spinulosus — Jensen, 1985; ternus — Wieser, 1954 a; truncatus — Cobb, 1914; vancouverensis — Sharma &amp; Vincx, 1982; vitraeus — Gerlach, 1957." httpUri="https://zenodo.org/record/183777/files/figure.png" pageId="3" pageNumber="154">2</figureCitation>
, Table 1)
</paragraph>
<paragraph id="841636DC5A5EFFE9FF00867BFF48F8FF" blockId="3.[151,1435,1525,1871]" pageId="3" pageNumber="154">
The guide is a set of simplified images (
<figureCitation id="1C922A595A5EFFE9FD6F867BFD40F987" box="[681,757,1565,1591]" captionStart="FIGURE 1" captionStartId="4.[151,244,1916,1937]" captionTargetBox="[190,1405,184,1901]" captionTargetId="figure@4.[163,1423,168,1921]" captionTargetPageId="4" captionText="FIGURE 1. Pictorial guide for valid species of Paracyatholaimus. Part 1, beginning. Simplified images of species come from the published descriptions: arcticus — Kreis, 1963; botosaneanui — Andrássy, 1973; chilensis — Gerlach, 1953 b; diva — orig.; dubiosus — Meyl, 1954; duplicatus — Gerlach, 1964 b; helicellus — Wieser, 1954 a; intermedius — Gerlach, 1953 a; lewisi — Coomans et al., 1985; occultus — Gerlach, 1956; paucipapillatus — Gerlach, 1955; pentodon — Riemann, 1966." httpUri="https://zenodo.org/record/183776/files/figure.png" pageId="3" pageNumber="154">Figs 1</figureCitation>
&amp;
<figureCitation id="1C922A595A5EFFE9FCD8867BFC9BF987" box="[798,814,1565,1591]" captionStart="FIGURE 2" captionStartId="5.[151,244,1911,1932]" captionTargetBox="[182,1346,182,1910]" captionTargetId="figure@5.[163,1423,163,1916]" captionTargetPageId="5" captionText="FIGURE 2. Pictorial guide for valid species of Paracyatholaimus. Part 2, ending. Simplified images of species come from the published descriptions: pesavis — Wieser &amp; Hopper, 1967; proximus — de Man, 1922; pugettensis — Wieser, 1959; quadriseta — Wieser, 1954 a; rotundus — Gerlach, 1964 a; saradi — Gerlach, 1967; separatus — Wieser, 1954 b; spinulosus — Jensen, 1985; ternus — Wieser, 1954 a; truncatus — Cobb, 1914; vancouverensis — Sharma &amp; Vincx, 1982; vitraeus — Gerlach, 1957." httpUri="https://zenodo.org/record/183777/files/figure.png" pageId="3" pageNumber="154">2</figureCitation>
) constructed in some accordance with the approved practice of
<bibRefCitation id="E0384B2D5A5EFFE9FEDF8623FE17F9EF" author="Platt" box="[281,418,1605,1631]" pageId="3" pageNumber="174" refString="Platt, H. M. (1984) Pictorial taxonomic keys: their construction and use for the identification of freeliving marine nematodes. Cahiers de Biologie Marine, 25, 83 - 91." type="journal article" year="1984">Platt (1984)</bibRefCitation>
and others (e.g., Platt &amp; Warwick, 1998). Species of
<taxonomicName id="43A94D5F5A5EFFE9FBCB8623FB51F9EE" box="[1037,1252,1605,1630]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5EFFE9FBCB8623FB51F9EE" box="[1037,1252,1605,1630]" italics="true" pageId="3" pageNumber="154">Paracyatholaimus</emphasis>
</taxonomicName>
differ from one another in rather fine morphological details concerning mainly length of anterior setae, number of turns in the multispiral amphidial fovea, supplementary organs and copulatory apparatus. The species can neither be disposed in a morphological continuum nor grouped in any evident morphological clusters. Therefore, it is not easy to develop a convenient polytomous or dichotomous key. I arrange the images of species in alphabetical order. An important component of the guide is Table 1, where the most important morphometric data are summarized.
</paragraph>
<caption id="D0D666545A59FFEEFF51871AFC3AF841" httpUri="https://zenodo.org/record/183776/files/figure.png" pageId="4" pageNumber="155" targetBox="[190,1405,184,1901]" targetPageId="4">
<paragraph id="841636DC5A59FFEEFF51871AFC3AF841" blockId="4.[151,1436,1916,2033]" pageId="4" pageNumber="155">
<emphasis id="B6DDEACE5A59FFEEFF51871AFEB3F821" bold="true" box="[151,262,1916,1937]" pageId="4" pageNumber="155">FIGURE 1</emphasis>
. Pictorial guide for valid species of
<taxonomicName id="43A94D5F5A59FFEEFD9B871AFCB8F821" box="[605,781,1916,1937]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A59FFEEFD9B871AFCB8F821" box="[605,781,1916,1937]" italics="true" pageId="4" pageNumber="155">Paracyatholaimus</emphasis>
</taxonomicName>
. Part 1, beginning. Simplified images of species come from the published descriptions:
<taxonomicName id="43A94D5F5A59FFEEFE9087FAFE17F801" box="[342,418,1948,1969]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="arcticus">
<emphasis id="B6DDEACE5A59FFEEFE9087FAFE17F801" box="[342,418,1948,1969]" italics="true" pageId="4" pageNumber="155">arcticus</emphasis>
</taxonomicName>
<emphasis id="B6DDEACE5A59FFEEFE6487FAFE0FF801" bold="true" box="[418,442,1948,1969]" pageId="4" pageNumber="155"></emphasis>
Kreis, 1963;
<taxonomicName id="43A94D5F5A59FFEEFDF087FAFD02F801" box="[566,695,1948,1969]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="botosaneanui">
<emphasis id="B6DDEACE5A59FFEEFDF087FAFD02F801" box="[566,695,1948,1969]" italics="true" pageId="4" pageNumber="155">botosaneanui</emphasis>
</taxonomicName>
—Andrássy, 1973;
<taxonomicName id="43A94D5F5A59FFEEFCB687FAFC71F801" box="[880,964,1948,1969]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="chilensis">
<emphasis id="B6DDEACE5A59FFEEFCB687FAFC71F801" box="[880,964,1948,1969]" italics="true" pageId="4" pageNumber="155">chilensis</emphasis>
</taxonomicName>
—Gerlach, 1953b;
<taxonomicName id="43A94D5F5A59FFEEFBBA87FAFB10F801" box="[1148,1189,1948,1969]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="diva">
<emphasis id="B6DDEACE5A59FFEEFBBA87FAFB10F801" box="[1148,1189,1948,1969]" italics="true" pageId="4" pageNumber="155">diva</emphasis>
</taxonomicName>
—orig.;
<taxonomicName id="43A94D5F5A59FFEEFB3087FAFAFEF801" box="[1270,1355,1948,1969]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="dubiosus">
<emphasis id="B6DDEACE5A59FFEEFB3087FAFAFEF801" box="[1270,1355,1948,1969]" italics="true" pageId="4" pageNumber="155">dubiosus</emphasis>
</taxonomicName>
—Meyl, 1954;
<taxonomicName id="43A94D5F5A59FFEEFF1287DAFE8DF861" box="[212,312,1980,2001]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="duplicatus">
<emphasis id="B6DDEACE5A59FFEEFF1287DAFE8DF861" box="[212,312,1980,2001]" italics="true" pageId="4" pageNumber="155">duplicatus</emphasis>
</taxonomicName>
—Gerlach, 1964b;
<taxonomicName id="43A94D5F5A59FFEEFE3787DAFDF8F861" box="[497,589,1980,2001]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="helicellus">
<emphasis id="B6DDEACE5A59FFEEFE3787DAFDF8F861" box="[497,589,1980,2001]" italics="true" pageId="4" pageNumber="155">helicellus</emphasis>
</taxonomicName>
—Wieser, 1954a;
<taxonomicName id="43A94D5F5A59FFEEFD3C87DAFCDEF861" box="[762,875,1980,2001]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="intermedius">
<emphasis id="B6DDEACE5A59FFEEFD3C87DAFCDEF861" box="[762,875,1980,2001]" italics="true" pageId="4" pageNumber="155">intermedius</emphasis>
</taxonomicName>
—Gerlach, 1953a;
<taxonomicName id="43A94D5F5A59FFEEFBE587DAFBEEF861" box="[1059,1115,1980,2001]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="lewisi">
<emphasis id="B6DDEACE5A59FFEEFBE587DAFBEEF861" box="[1059,1115,1980,2001]" italics="true" pageId="4" pageNumber="155">lewisi</emphasis>
</taxonomicName>
—Coomans et al., 1985;
<taxonomicName id="43A94D5F5A59FFEEFA8A87DAFA29F861" box="[1356,1436,1980,2001]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="occultus">
<emphasis id="B6DDEACE5A59FFEEFA8A87DAFA29F861" box="[1356,1436,1980,2001]" italics="true" pageId="4" pageNumber="155">occultus</emphasis>
</taxonomicName>
—Gerlach, 1956;
<taxonomicName id="43A94D5F5A59FFEEFE8587BAFE6DF841" box="[323,472,2012,2033]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="paucipapillatus">
<emphasis id="B6DDEACE5A59FFEEFE8587BAFE6DF841" box="[323,472,2012,2033]" italics="true" pageId="4" pageNumber="155">paucipapillatus</emphasis>
</taxonomicName>
—Gerlach, 1955;
<taxonomicName id="43A94D5F5A59FFEEFD4387BAFD6BF841" box="[645,734,2012,2033]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="4" pageNumber="155" phylum="Nematoda" rank="species" species="pentodon">
<emphasis id="B6DDEACE5A59FFEEFD4387BAFD6BF841" box="[645,734,2012,2033]" italics="true" pageId="4" pageNumber="155">pentodon</emphasis>
</taxonomicName>
—Riemann, 1966.
</paragraph>
</caption>
<caption id="D0D666545A58FFEFFF518711FBE0F85C" httpUri="https://zenodo.org/record/183777/files/figure.png" pageId="5" pageNumber="156" targetBox="[182,1346,182,1910]" targetPageId="5">
<paragraph id="841636DC5A58FFEFFF518711FBE0F85C" blockId="5.[151,1436,1911,2029]" pageId="5" pageNumber="156">
<emphasis id="B6DDEACE5A58FFEFFF518711FEB2F83C" bold="true" box="[151,263,1911,1932]" pageId="5" pageNumber="156">FIGURE 2</emphasis>
. Pictorial guide for valid species of
<taxonomicName id="43A94D5F5A58FFEFFDAC871EFCAFF83D" box="[618,794,1912,1933]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A58FFEFFDAC871EFCAFF83D" box="[618,794,1912,1933]" italics="true" pageId="5" pageNumber="156">Paracyatholaimus</emphasis>
</taxonomicName>
. Part 2, ending. Simplified images of species come from the published descriptions:
<taxonomicName id="43A94D5F5A58FFEFFE9A87FEFE16F81D" box="[348,419,1944,1965]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="pesavis">
<emphasis id="B6DDEACE5A58FFEFFE9A87FEFE16F81D" box="[348,419,1944,1965]" italics="true" pageId="5" pageNumber="156">pesavis</emphasis>
</taxonomicName>
—Wieser &amp; Hopper, 1967;
<taxonomicName id="43A94D5F5A58FFEFFD7187FEFCBAF81D" box="[695,783,1944,1965]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="proximus">
<emphasis id="B6DDEACE5A58FFEFFD7187FEFCBAF81D" box="[695,783,1944,1965]" italics="true" pageId="5" pageNumber="156">proximus</emphasis>
</taxonomicName>
—de Man, 1922;
<taxonomicName id="43A94D5F5A58FFEFFC0787FEFB98F81D" box="[961,1069,1944,1965]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="pugettensis">
<emphasis id="B6DDEACE5A58FFEFFC0787FEFB98F81D" box="[961,1069,1944,1965]" italics="true" pageId="5" pageNumber="156">pugettensis</emphasis>
</taxonomicName>
—Wieser, 1959;
<taxonomicName id="43A94D5F5A58FFEFFB1387FEFA8EF81D" box="[1237,1339,1944,1965]" class="Arachnida" family="Dictynidae" genus="Marilynia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="quadriseta">
<emphasis id="B6DDEACE5A58FFEFFB1387FEFA8EF81D" box="[1237,1339,1944,1965]" italics="true" pageId="5" pageNumber="156">quadriseta</emphasis>
</taxonomicName>
—Wieser, 1954a;
<taxonomicName id="43A94D5F5A58FFEFFF2787DEFE80F87D" box="[225,309,1976,1997]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="rotundus">
<emphasis id="B6DDEACE5A58FFEFFF2787DEFE80F87D" box="[225,309,1976,1997]" italics="true" pageId="5" pageNumber="156">rotundus</emphasis>
</taxonomicName>
—Gerlach, 1964a;
<taxonomicName id="43A94D5F5A58FFEFFE3587DEFD85F87D" box="[499,560,1976,1997]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="saradi">
<emphasis id="B6DDEACE5A58FFEFFE3587DEFD85F87D" box="[499,560,1976,1997]" italics="true" pageId="5" pageNumber="156">saradi</emphasis>
</taxonomicName>
—Gerlach, 1967;
<taxonomicName id="43A94D5F5A58FFEFFD2487DEFC8AF87D" box="[738,831,1976,1997]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="separatus">
<emphasis id="B6DDEACE5A58FFEFFD2487DEFC8AF87D" box="[738,831,1976,1997]" italics="true" pageId="5" pageNumber="156">separatus</emphasis>
</taxonomicName>
—Wieser, 1954b;
<taxonomicName id="43A94D5F5A58FFEFFC3287DEFBECF87D" box="[1012,1113,1976,1997]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="spinulosus">
<emphasis id="B6DDEACE5A58FFEFFC3287DEFBECF87D" box="[1012,1113,1976,1997]" italics="true" pageId="5" pageNumber="156">spinulosus</emphasis>
</taxonomicName>
—Jensen, 1985;
<taxonomicName id="43A94D5F5A58FFEFFAC687DEFA89F87D" box="[1280,1340,1976,1997]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="ternus">
<emphasis id="B6DDEACE5A58FFEFFAC687DEFA89F87D" box="[1280,1340,1976,1997]" italics="true" pageId="5" pageNumber="156">ternus</emphasis>
</taxonomicName>
—Wieser, 1954a;
<taxonomicName id="43A94D5F5A58FFEFFF1887BEFE8CF85D" box="[222,313,2008,2029]" class="Adenophorea" family="Cyatholaimidae" genus="Cyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="truncatus">
<emphasis id="B6DDEACE5A58FFEFFF1887BEFE8CF85D" box="[222,313,2008,2029]" italics="true" pageId="5" pageNumber="156">truncatus</emphasis>
</taxonomicName>
—Cobb, 1914;
<taxonomicName id="43A94D5F5A58FFEFFE0B87BEFDD5F85D" box="[461,608,2008,2029]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="vancouverensis">
<emphasis id="B6DDEACE5A58FFEFFE0B87BEFDD5F85D" box="[461,608,2008,2029]" italics="true" pageId="5" pageNumber="156">vancouverensis</emphasis>
</taxonomicName>
—Sharma &amp; Vincx, 1982;
<taxonomicName id="43A94D5F5A58FFEFFCA787BEFC1BF85D" box="[865,942,2008,2029]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="156" phylum="Nematoda" rank="species" species="vitraeus">
<emphasis id="B6DDEACE5A58FFEFFCA787BEFC1BF85D" box="[865,942,2008,2029]" italics="true" pageId="5" pageNumber="156">vitraeus</emphasis>
</taxonomicName>
—Gerlach, 1957.
</paragraph>
</caption>
<paragraph id="841636DC5A5BFFECFF5180FFFCCCFF02" blockId="6.[151,889,152,178]" box="[151,889,152,178]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF5180FFFEB6FF01" bold="true" box="[151,259,153,177]" pageId="6" pageNumber="157">TABLE 1</emphasis>
. Morphometry of valid
<taxonomicName id="43A94D5F5A5BFFECFDC280FCFD72FF01" box="[516,711,154,177]" class="Adenophorea" family="Cyatholaimidae" genus="Paracyatholaimus" kingdom="Animalia" order="Desmodorida" pageId="6" pageNumber="157" phylum="Nematoda" rank="genus">
<emphasis id="B6DDEACE5A5BFFECFDC280FCFD72FF01" box="[516,711,154,177]" italics="true" pageId="6" pageNumber="157">Paracyatholaimus</emphasis>
</taxonomicName>
species (males).
</paragraph>
<paragraph id="841636DC5A5BFFECFF1580B1FC01FF57" blockId="6.[211,1412,210,335]" box="[211,948,210,236]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFF1580B1FC01FF57" bold="true" box="[211,948,210,236]" pageId="6" pageNumber="157">Species Characters</emphasis>
</paragraph>
<paragraph id="841636DC5A5BFFECFE51809CFA31FEBF" blockId="6.[211,1412,210,335]" box="[407,1412,250,274]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFE51809CFCBBFEA2" bold="true" box="[407,782,250,274]" pageId="6" pageNumber="157">L a c o.l.s.,</emphasis>
μ
<emphasis id="B6DDEACE5A5BFFECFCE5809CFA31FEBF" bold="true" box="[803,1412,250,274]" pageId="6" pageNumber="157">m am.w., m, amphid, c spic., m suppl. n.</emphasis>
</paragraph>
<paragraph id="841636DC5A5BFFECFC9D817CFB93FE9F" blockId="6.[211,1412,210,335]" box="[859,1062,282,303]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFC9D817CFB93FE9F" bold="true" box="[859,1062,282,303]" pageId="6" pageNumber="157">% c.b.d. number</emphasis>
</paragraph>
<paragraph id="841636DC5A5BFFECFC13815CFB93FEFF" blockId="6.[211,1412,210,335]" box="[981,1062,314,335]" pageId="6" pageNumber="157">
<emphasis id="B6DDEACE5A5BFFECFC13815CFB93FEFF" bold="true" box="[981,1062,314,335]" pageId="6" pageNumber="157">of turns</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
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<mods:title id="8FE790DDA4668A11377BFFB460C6633C">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1</mods:title>
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<treatment id="0C00A4CC8DE4B75865383C55076F8CE4" ID-DOI="http://doi.org/10.5281/zenodo.11330329" ID-GBIF-Taxon="231535682" ID-Zenodo-Dep="11330329" LSID="urn:lsid:plazi:treatment:0C00A4CC8DE4B75865383C55076F8CE4" httpUri="http://treatment.plazi.org/id/0C00A4CC8DE4B75865383C55076F8CE4" lastPageNumber="444" pageNumber="444" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13801695">
<subSubSection id="E2A1574CEF0953B5006E3945F7B507EB" pageNumber="444" type="nomenclature">
<paragraph id="BF7BAC7C85C93B7A76DACDCF9FF7C32D" pageNumber="444">
<taxonomicName id="1012A1047B311D0053B0684D532E15FC" ID-CoL="7VW4R" authority="Thomas 1903" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="444" phylum="Chordata" rank="species" species="thersites" subGenus="Xiphonycteris">
<emphasis id="420DB2570C01B68440822738B5140BA4" italics="true" pageNumber="444">Mops (Xiphonycteris) thersites</emphasis>
Thomas 1903
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="FC1496119EFAEC6B8B033B67B0CE5C47" pageNumber="444" type="reference_group">
<paragraph id="B3B80B466380552E70A6E182AFE3CE1A" pageNumber="444">
<treatmentCitation id="7AD71C82A9E212798E12863BCC468C0A" author="Thomas" authority="Thomas 1903" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Molossidae" genus="Mops" journal="Ann. Mag. Nat. Hist., ser. 7" kingdom="Animalia" order="Chiroptera" page="634" pageNumber="444" phylum="Chordata" rank="species" species="thersites" subGenus="Xiphonycteris" title="Mops (Xiphonycteris) thersites" volume="12" year="1903">
<taxonomicName id="852494953C54F7CCBE655568812BCBF1" ID-CoL="7VW4R" authority="Thomas 1903" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="444" phylum="Chordata" rank="species" species="thersites" subGenus="Xiphonycteris">
<emphasis id="63EC12ECC802EB1673FB66EDA0B33D9E" italics="true" pageNumber="444">Mops (Xiphonycteris) thersites</emphasis>
<bibRefCitation id="9208C10AF3CA594AE4E835AC7D45A97B" author="Thomas, O." refId="ref352582" refString="Thomas, O. 1903. Notes on Neotropical mammals of the genera Felis, Hapale, Oryzomys, Akodon, and Ctenomys, with descriptions of new species. Annals and Magazine of Natural History, ser. 7, 12: 234 - 243." year="1903">Thomas 1903</bibRefCitation>
</taxonomicName>
,
<bibCitation id="209D61470FB01CE1784FBACE2DFF5FAE" author="Thomas" journal="Ann. Mag. Nat. Hist., ser. 7" pageNumber="444" pagination="634" title="Mops (Xiphonycteris) thersites" volume="12" year="1903">Ann. Mag. Nat. Hist., ser. 7, 12: 634</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="729A746D8BE32DC4BC01922AEE9279A7" pageNumber="444" type="materials_examined">
<paragraph id="97ADEE4815143B922C1844B224E1DB51" pageNumber="444">
<emphasis id="5E4A21E879AEBB44E943F5C75A092CC7" inLineHeading="true" pageNumber="444">Type Locality:</emphasis>
<materialsCitation id="1D30DDB25666431A604471798FE416BC" ID-GBIF-Occurrence="4867302332" country="Cameroon" pageNumber="444">
<collectingCountry id="FA5378438A71C375DD6B7C8AD937C359" name="Cameroon">Cameroon</collectingCountry>
, Efulen.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6A07D696AA0985784557E4FBFD0D5432" pageNumber="444" type="vernacular_names">
<paragraph id="75FF42F09F2BA42278BC01D3D14CD6E5" pageNumber="444">
<emphasis id="07BA9F06306580E3AED8FA460F2DB918" inLineHeading="true" pageNumber="444">Vernacular Names:</emphasis>
<vernacularName id="E22C28407DE895D6D2954CE19DCAB092" language="eng" pageNumber="444">Railer Free-tailed Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="08295796E5A92CB97D0A840C540097E3" pageNumber="444" type="synonym_list">
<paragraph id="B4C486C8316782238984E60BE49B6C5E" pageNumber="444">
<emphasis id="10C5F76078467E7DC17590E64135D045" inLineHeading="true" pageNumber="444">Synonyms:</emphasis>
<taxonomicName id="9E85F7AC2167CA1862543184F0890239" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="444" phylum="Chordata" rank="species" species="occipitalis" subGenus="Xiphonycteris">
<emphasis id="502DAC96E2A63E40708AF0AA0D4C7B1C" italics="true" pageNumber="444">Mops (Xiphonycteris) occipitalis</emphasis>
J. A. Allen 1917
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F7C584A6B4D074D8BA4C05283DA0B06C" pageNumber="444" type="distribution">
<paragraph id="AA830F59A90D206A3A4ABF1B17EF0331" pageNumber="444">
<emphasis id="C8D038B9534B05F3BF6E10AC1423D5EA" inLineHeading="true" pageNumber="444">Distribution:</emphasis>
<collectingCountry id="ECA60F6A6CC2107EB4BF505BBD9E0C36" name="Sierra Leone">Sierra Leone</collectingCountry>
to
<collectingCountry id="817CE78D35BFA14235F047C3072F2B3A" name="Rwanda">Rwanda</collectingCountry>
; Bioko (
<collectingCountry id="1A405A09B28C9E8AB73E54C3D2C2AED1" name="Equatorial Guinea">Equatorial Guinea</collectingCountry>
); perhaps
<collectingCountry id="C024E5F353C23D41C5FD81524DC3AE57" name="Mozambique">Mozambique</collectingCountry>
and
<collectingCountry id="F803C16FA7ED6A54C3F286749AA3588C" name="Tanzania">Zanzibar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D679EFFB7FD02A191BF24D5E5CEF1DBF" pageNumber="444" type="conservation">
<paragraph id="29EE5D4E2860230C2DD04D898FD3B05C" pageNumber="444">
<emphasis id="242B85C23AF62D14F3AA7F15E2E83FD3" inLineHeading="true" pageNumber="444">Conservation:</emphasis>
<collectionCode id="8279CA461178E76DD1C88B31297A5EC0">IUCN</collectionCode>
2003 and
<collectionCode id="02C579A43220C4FC5D9F955B469A289F">IUCN</collectionCode>
/
<collectionCode id="64CDF46021AF3DF4E87748D337132EC3" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="D3E40167D403FAE7E7CA60344E741770" pageNumber="444" type="discussion">
<paragraph id="E6BA030F284368579F3BF03556846D88" pageNumber="444">
<emphasis id="93A1B45BFC9CCB08B71E59B0970EC7B7" inLineHeading="true" pageNumber="444">Discussion:</emphasis>
Subgenus
<taxonomicName id="3ABDE2393F9536E619456947F5899AA1" authorityName="Dollman" authorityYear="1911" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="subGenus" subGenus="Xiphonycteris">
<emphasis id="B9A9D407349A9A2A6EC1789043688008" italics="true" pageNumber="444">Xiphonycteris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,59 @@
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<taxonomicName id="D8F0B1A02FAED1277707A911D955725F" ID-CoL="TP5K" authority="Thomson, 1888" authorityName="Thomson" authorityYear="1888" class="Insecta" family="Ichneumonidae" genus="Charitopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charitopes clausus" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="clausus">Charitopes clausus (Thomson, 1888)</taxonomicName>
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<taxonomicName id="FB49AB8E2BBE628A05FFAC39D09D9A68" class="Insecta" family="Ichneumonidae" genus="Hemiteles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemiteles clausus" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="clausus">Hemiteles clausus</taxonomicName>
Thomson, 1888
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<paragraph id="E8E4ED8952CD4A899B127F28DE46CF66" pageId="0" pageNumber="9042">England, Scotland, Ireland, Isle of Man</paragraph>
</subSubSection>
<subSubSection id="E297D51419FE8B853F7DB7F2407FEE7A" pageId="0" pageNumber="9042" type="notes">
<paragraph id="2867E71A9F2945544591ABED300FB32A" pageId="0" pageNumber="9042">Notes</paragraph>
<paragraph id="E4B18EB0576C48034291065C1549FC1B" pageId="0" pageNumber="9042">
added by
<bibRefCitation id="487E000ED0F44CD544EA45FB214AD7DF" author="Townes, H." journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="9042" pagination="1 - 281" title="Revisions of twenty genera of Gelini (Ichneumonidae)" volume="35" year="1983">Townes (1983)</bibRefCitation>
</paragraph>
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@ -0,0 +1,289 @@
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<mods:namePart>Liebherr, James K.</mods:namePart>
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<treatment ID-GBIF-Taxon="182754103" LSID="urn:lsid:zoobank.org:act:9985664D-8441-4E08-BE2F-AA1AE24BE8C2" httpUri="http://treatment.plazi.org/id/0C00EF7E1F3C5F19B3D09B6A6653C622" lastPageNumber="229" pageId="0" pageNumber="229">
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<taxonomicName LSID="http://zoobank.org/9985664D-8441-4E08-BE2F-AA1AE24BE8C2" authority="Liebherr, 2021" authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa" status="sp. nov.">Paratachys aaa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="229">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">Figures 3D-F</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paratachys spp., left female gonocoxa, ventral view A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) C P. aaa; Hawai'i I.; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: afs, apical fringe seta; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite; gc 2, apical gonocoxite; les, lateral ensiform setae." figureDoi="10.3897/zookeys.1044.59674.figure4" httpUri="https://binary.pensoft.net/fig/553680" pageId="0" pageNumber="229">, 4C</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Paratachys aaa female reproductive tract, ventral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: bc, bursa copulatrix; co, common oviduct; sd, spermathecal duct; sg, spermathecal gland; sp, spermatheca. Schematic drawing breaks spermathecal duct at &quot; A &quot; to accommodate drawing to page size. Estimated length of spermathecal duct 1.7 mm." figureDoi="10.3897/zookeys.1044.59674.figure10" httpUri="https://binary.pensoft.net/fig/553686" pageId="0" pageNumber="229">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributional records of Paratachys aaa. Abbreviations: H, Hilo; K, caldera of Kilauea Volcano; ML, summit caldera of Mauna Loa Volcano." figureDoi="10.3897/zookeys.1044.59674.figure11" httpUri="https://binary.pensoft.net/fig/553687" pageId="0" pageNumber="229">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="type material">
<paragraph pageId="0" pageNumber="229">Type material.</paragraph>
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Holotype</emphasis>
male (BPBM): HI: Hawaii Mountain View / Kazumura Cave 22-VII- / 1971 F. G. Howarth / 600' Inside lava tube cave / on slime on wall // BBM-00301 // HOLOTYPE ♂ / Paratachys / aaa / J. K. Liebherr 2020 (black-margined red label).
</paragraph>
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Allotypic paratype</emphasis>
female (BPBM): HI: Hawaii Mountain View / Kazumura Cave 25-VII- / 1971 F. G. Howarth / 200' Inside lava tube cave // BBM-00302 // ALLOTYPE ♀ / Paratachys / aaa / J. K. Liebherr 2020 (black-margined red label).
</paragraph>
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" pageId="0" pageNumber="229">
<emphasis italics="true" pageId="0" pageNumber="229">Paratypes</emphasis>
.
<normalizedToken originalValue="Hawaii">Hawai'i</normalizedToken>
I.: Hamakua District
</emphasis>
: Pohakuloa Military Training Area, Bobcat Trail, Cave 10265DE, deep zone, 1650 m el., 30-xii-1994, Howarth (BPBM, 3).
<emphasis bold="true" pageId="0" pageNumber="229">Kau District</emphasis>
: Ocean View, Kipuka Kanohina System, Cordwinder Natural Bridge, upslope, 23-xi-2018, Hackell/ Porter/ Hudson/ Katz, lot HI00776 (UHIM, 1), Kona Mala Driveway Entr., 22-xi-2016, M. Slay/ C. Slay/ Porter, lot HI00094 (BPBM, 1; UHIM, 2), 21-xi-2017, M. Slay/ C. Slay/ Gracanin/ Hackell/ A. Engel/ S. Engel/ Porter, lot HI00179 (CUIC, 1), 25-xi-2017, C. Slay/ A. Engel/ Hackell, lot HI00301 (UHIM, 3), Kula Kai Caverns, Chocolate Factory, 22-xi-2017, Porter/ S. Engel/ A. Engel/ Bosted, lot HI00227 (UHIM, 1), Menehune Entrance, 11-xi-2018, Chong/ Hudson/ Porter/ Thomson, lot HI00508 (UHIM, 1), 20-xi-2018, M. Slay/ S. Engel/ Katz/ Taylor (UHIM, 3),
<normalizedToken originalValue="Wilsons">Wilson's</normalizedToken>
Big Room, 20-xi-2018, C. Slay/ Engel/ Katz/ Taylor, HI00673, lot HI00619 (UHIM, 2), Xanadu XD 1, 23-xi-2016, M. Slay/ C. Slay/ Porter, lot HI00142 (UHIM, 1), Xanadu Extension 1, 23-xi-2018, M. Slay/ C. Slay/ Yelverton/ Gunter/ Gracanin, lot HI00755 (UHIM, 1).
<emphasis bold="true" pageId="0" pageNumber="229">Puna District</emphasis>
: Kaimu, Burn Cave, deep zone, site 4, 260 m el., 16-iii-1994, Howarth/ Miller (BPBM, 1), pitfall trap L-8, 260 m el., 16-19-iii-1994, Howarth/ Miller (BPBM, 1), pitfall trap M-5, 16-19-iii-1994, Howarth/ Miller (BPBM, 2); Mountain View,
<normalizedToken originalValue="Cows">Cow's</normalizedToken>
Eye Entrance, 22-iii-2019, Hudson/ Gracanin/ Hackell/ C. Slay/ M. Slay, lot HI01050 (UHIM, 2), lot HI01061 (BPBM, 3), D Road Cave, 20-iii-2019, Porter/ Chong/ Engel/ Hudson/ Hackell, lot HI00973 (UHIM, 2), lot HI00988 (BPBM, 2),
<normalizedToken originalValue="Eppersons">Epperson's</normalizedToken>
Cave, 20-iii-2019, C. Slay, M. Slay, Engel, Gracanin, lot HI 00949 (UHIM, 1), Keala Cave Entrance, 19-iii-2019, A. Engel, S. Engel, Hudson, Hackell, M. Slay, C. Slay, Porter, Gracanin, lot HI 00911 (UHIM, 1), Kazumura Cave, lava tube cave, 200' inside lava tube cave, 25-vii-1971, Howarth (BPBM, 1 [male genitalia missing]), Weldon Sheldon entrance, downslope, 21-xi-2018, A. Engel/ Hudson/ Taylor, HI00713 (CUIC, 2), upslope, 21-xi-2018, C. Slay/ Yelverton/ Gunter/ Gracanin/ Hackell, HI00722 (UHIM, 2), 21-iii-2019, Chong, Hudson, A. Engel, S. Engel, Gracanin, lot HI01017 (UHIM, 1); Pahoa, Pahoa Cave, deep zone, 180 m el., 15-iii-1994, Howarth/ Miller (BPBM, 1), on bait, 18-iii-1994, Howarth/ Miller (BPBM, 1).
<emphasis bold="true" pageId="0" pageNumber="229">South Hilo District</emphasis>
: Kaumana Cave, 10-xi-2018, Chong, Hudson, Porter, Thomson, Lot HI00499 (UHIM, 1), 19-iii-2019, Engel/ Hackell/ Gracanin, Lot HI00925 (UHIM, 1), Porter/ Engel/ Hudson, Lot 00942, (CUIC, 1; UHIM, 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="diagnosis">
<paragraph pageId="0" pageNumber="229">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="229">
This is a small-bodied, pallid
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys</emphasis>
</taxonomicName>
with thin, translucent cuticle, and an iridescent sheen to the elytra due to the elongate transverse microsculpture; standardized body length 1.9-2.2 mm. The eyes are small but somewhat variable, with from 3-5 ommatidia crossed by a horizontal diameter of the eye, and from 4-6 ommatidia crossed by a vertical diameter (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9C-F</figureCitation>
); OR = 1.14-1.19. The elytra are broad, with broadly rounded humeri and broad, nearly truncate apices. The elytral apical recurrent groove is broadly, shallowly impressed. Sutural interneur 1 (i1) of each elytron is well impressed, with the two interneurs bracketing an elevated, callous-like elytral suture, whereas interneurs 2 and 3 are broadly, shallowly impressed to obsolete on the disc, with the outer interneurs absent except for the configuration of interneur 8 (i8) characteristic of
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.626.10033" author="Boyd, OF" journalOrPublisher="ZooKeys" pageId="0" pageNumber="229" pagination="87 - 123" refId="B16" refString="Boyd, OF, Erwin, TL, 2016. Taxonomic review of the New World Tachyina (Coleoptera, Carabidae): descriptions of new genera, subgenera, and species, with an updated key to the subtribe in the Americas. ZooKeys 626: 87 - 123, DOI: https://doi.org/10.3897/zookeys.626.10033" title="Taxonomic review of the New World Tachyina (Coleoptera, Carabidae): descriptions of new genera, subgenera, and species, with an updated key to the subtribe in the Americas." url="https://doi.org/10.3897/zookeys.626.10033" volume="626" year="2016">Boyd and Erwin 2016</bibRefCitation>
: fig. 1A).
</paragraph>
<caption doi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" pageId="0" pageNumber="229">Figure 9.</emphasis>
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys aaa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="229">A</emphasis>
male, dorsal habitus view; Kau, Kona Mala, 21-xi-2017, lot HI00179
<emphasis bold="true" pageId="0" pageNumber="229">B</emphasis>
mentum and associated mouthparts, ventral view, of above male specimen
<emphasis bold="true" pageId="0" pageNumber="229">C</emphasis>
larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21-xi-2018, lot HI00722
<emphasis bold="true" pageId="0" pageNumber="229">D</emphasis>
moderately sized ovoid compound eye of male, lateral view; also lot HI00722
<emphasis bold="true" pageId="0" pageNumber="229">E</emphasis>
smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16-19-iii-1994
<emphasis bold="true" pageId="0" pageNumber="229">F</emphasis>
small compound eye of male, lateral view; Kaimu, Burn Cave, 16-iii-1994
<emphasis bold="true" pageId="0" pageNumber="229">G</emphasis>
stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21-xi-2017, lot HI00179.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="description">
<paragraph pageId="0" pageNumber="229">Description.</paragraph>
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Head</emphasis>
narrow, ocular lobes nearly flattened anterad the genae in dorsal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9A</figureCitation>
); frontal grooves narrow mesad anterior supraorbital seta, broadest between anterior eye margins, and extended to lateral reaches of frontoclypeal suture; antennae moderately elongate, antennomere 9 length twice diameter; mandibles of moderate length, mandibles elongate, distance from dorsal condyle to apex 1.6
<normalizedToken originalValue="×">x</normalizedToken>
distance from condyle to anterolateral labral margin; labrum transverse, anterior margin evenly emarginate across width, six-setose; penultimate maxillary palpomere broadly spindle-shaped, apical palpomere a narrow spindle (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9A</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Prothorax</emphasis>
variably transverse, MPW/PL = 1.33-1.46, with lateral margins distinctly sinuate anterad the acute, projected hind angles, MPW/BPW = 1.18-1.27, the sinuosity of the lateral margins accentuating the cordate appearance more than that represented by measuring pronotal width over the hind angles; pronotal median base depressed, its juncture with disc smooth, basal margin extended medially to form a collar that extends posterad the concave margins posterad the laterobasal depressions; laterobasal depression deepest as a lateral extension of the discal-median base juncture that is directed to the concave basal margin mesad hind angles; pronotal median impression finely depressed, narrow, pronotal disc flattened each side of midline; anterior transverse impression obsolete medially, broadly, shallowly extended laterally to narrowly rounded, moderately projected front angles.
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Elytra</emphasis>
subquadrate, short, humeri broad, EL/MEW = 1.38-1.47; basal groove present laterad position of third interneur on interval distance laterad the parascutellar seta Ed1, groove convexly joined to lateral marginal depression; lateral marginal depression moderately reflexed, of equal breadth from seta Eo3 to subapical sinuation; broader Eo elytral setae elongate, e.g., seta Eo2 length = 0.7 mm, Eo9 length = 0.8 mm (Pahoa Cave, 15-iii-1994, ♂, BPBM).
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Pterothorax</emphasis>
moderately elongate; mesepisternal depression smooth posterad juncture with mesosternum; metepisternum elongate, lateral length 1.6
<normalizedToken originalValue="×">x</normalizedToken>
maximal width; metathoracic flight wings vestigial, the alae reduced to broad-based stenopterous flaps that extend nearly to posterior margin of first abdominal tergite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9G</figureCitation>
) or to position of seta Eo4 when viewed through elytron, vestiges of radial, medial and cubital veins visible in the wing membrane.
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Abdomen</emphasis>
with one seta each side of apical ventrite in males, two seta each side of ventrite in females. Microsculpture evident on all somites; frons covered with shallow transverse mesh, sculpticells isodiametric and cuticular surface rougher in broad frontal grooves; pronotal disc glossy with elongate transverse sculpticells producing silvery sheen, median base with distinct transverse lines medially, and irregular isodiametric sculpticells laterally; elytra iridescent due to mix of elongate transverse-mesh and transverse-line microsculpture, the transverse lines denser laterally; abdominal ventrites glossy laterally with shallow, swirling elongate-mesh microsculpture.
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Pelage</emphasis>
present on head, prothorax, elytra, pterothorax, abdominal ventrites and legs; pelage on head, pronotal disc, and elytra comprising microsetae separated by distances subequal to setal length (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9C-F</figureCitation>
), the microsetae linearly arrayed along elytral interneurs as well as along ommatidial margins of the eyes (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paratachys aaa A male, dorsal habitus view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179 B mentum and associated mouthparts, ventral view, of above male specimen C larger ovoid compound eye of female, lateral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00722 D moderately sized ovoid compound eye of male, lateral view; also lot HI 00722 E smaller rounded compound eye of male, lateral view; Kaimu, Burn Cave, 16 - 19 - iii- 1994 F small compound eye of male, lateral view; Kaimu, Burn Cave, 16 - iii- 1994 G stenopterous left metathoracic flight wing of male (elytron removed), dorsal view; Kau, Kona Mala, 21 - xi- 2017, lot HI 00179." figureDoi="10.3897/zookeys.1044.59674.figure9" httpUri="https://binary.pensoft.net/fig/553685" pageId="0" pageNumber="229">9B</figureCitation>
); prosternum and mesosternum covered with sparse pelage medially; pelage of short microsetae present on abdomen in middle of ventrite 2 between metalegs, and progressively more broadly on ventrites 3-6 (absent dorsad arc of metaleg movement); anterior and posterior surface of prothoracic femora sparsely covered with microsetae, anterior (ventral) surfaces of meso- and metathoracic legs more densely covered with elongate microsetae, the setal bases situated more closely than microsetal lengths, trochanters and coxae similarly covered with microsetae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Coloration</emphasis>
pale; vertex rufous to rufo-flavous, clypeus rufo-flavous, labrum flavous; antennomeres flavous, outer antennomeres 4-11 with slight smoky cast; maxillary and labial palps flavous; pronotum rufo-flavous; elytra rufo-flavous, elytral lateral marginal depression narrowly flavous medially; elytral epipleuron rufo-flavous, concolorous with thoracic and abdominal ventrites; legs flavous from trochanters outward; pro- and mesocoxae concolorous with outer leg segments, metacoxae rufo-flavous to match thoracic ventrites.
<emphasis bold="true" italics="true" pageId="0" pageNumber="229">Legs</emphasis>
with only basal male protarsomere bearing a blunt, antero-apical process.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="variation">
<paragraph pageId="0" pageNumber="229">Variation.</paragraph>
<paragraph pageId="0" pageNumber="229">A series of five individuals from Kaumana Cave, South Hilo District, include the largest individuals observed for this species. Their standardized body lengths range 2.1-2.2 mm, whereas all other individuals from caves in Hamakua, Puna, and Kau District range 1.9-2.1 mm standardized body length. The Kaumana specimens are also more heavily melanized, though their rufo-brunneous coloration is concolorous with the darkest individuals from the other localities. As neither of these attributes are diagnostic, and variation in male genitalia (below) does not support recognition of the Kaumana Cave individuals as representatives of a distinct species, all individuals described here are considered conspecific.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="male genitalia">
<paragraph pageId="0" pageNumber="229">Male genitalia.</paragraph>
<paragraph pageId="0" pageNumber="229">
Aedeagal median lobe straight to slightly expanded near midlength, dorsal surface straight near midlength, evenly narrowed to rounded tip that is densely covered with sensilla (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3D-F</figureCitation>
); flagellar complex broadly hemi-ovoid, with parallel cuticular ridges emanating from densely sclerotized ventrobasal margin; narrow, strap-like right paramere with three setae on narrowly rounded apex; basally broad left paramere elongate, the parallel-sided apical portion with broadly rounded, three-setose apex. There is some variability observable among aedeagal dissections with the dissection of a beetle from Kazumura Cave (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3D</figureCitation>
) somewhat broader at midlength, and the aedeagus of a male from Kaumana Cave (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3F</figureCitation>
) more gracile than the male from Kau (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3E</figureCitation>
). The Kaumana male is larger than the other two (2.2 mm vs. 2.0 mm) consistent with the slightly longer aedeagus observed in that dissection (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3F</figureCitation>
vs. Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paratachys spp., male aedeagus, i. e., median lobe and associated smaller right paramere and larger left paramere (right lateral view) A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. arcanicola lectotype slide; &quot; Tachys arcanicola Blackburn Type ♂ E. C. Z. (permanent slide, Euparal?, left paramere missing; NHMUK). &quot; C P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) D-F P. aaa; Hawai'i I D Puna, Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC) E Kau, Ocean View, Kipuka Kanohina System, 21 - xi- 2017, lot HI 00179 (CUIC) F South Hilo, Kaumana Cave, 19 - iii- 2019, lot HI 00942 (CUIC); disassociated left paramere below, ectal view." figureDoi="10.3897/zookeys.1044.59674.figure3" httpUri="https://binary.pensoft.net/fig/553679" pageId="0" pageNumber="229">3D, E</figureCitation>
). The position of the flagellar complex within the aedeagal shaft is to be discounted, as the entire internal sac is eversible, and its position in repose would vary depending on the level of saccal inversion.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="female reproductive tract">
<paragraph pageId="0" pageNumber="229">Female reproductive tract.</paragraph>
<paragraph pageId="0" pageNumber="229">
Bursa copulatrix short, broad, with spermathecal duct entering near base of common oviduct (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Paratachys aaa female reproductive tract, ventral view; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: bc, bursa copulatrix; co, common oviduct; sd, spermathecal duct; sg, spermathecal gland; sp, spermatheca. Schematic drawing breaks spermathecal duct at &quot; A &quot; to accommodate drawing to page size. Estimated length of spermathecal duct 1.7 mm." figureDoi="10.3897/zookeys.1044.59674.figure10" httpUri="https://binary.pensoft.net/fig/553686" pageId="0" pageNumber="229">10</figureCitation>
); spermathecal duct very long, by measurement of drawing with mapping opisometer, 1.7 mm (!); spermathecal reservoir indistinctly annulated, at right angle to basal atrium and duct, spermathecal gland duct entering spermatheca near bend; spermathecal gland with narrow duct leading to broader reservoir, the reservoir bearing a small apical assemblage of ductule-bearing secretory cells on a narrow duct; gonocoxa bipartite (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paratachys spp., left female gonocoxa, ventral view A P. terryli; Kauai: Namolokama Mtn., 23 - v- 2005 (CUIC) B P. haleakalae; East Maui: Haleakala N. P., Kipahulu Vy., 30 - iv- 1991 (CUIC) C P. aaa; Hawai'i I.; Mountain View, Kazumura Cave, 21 - xi- 2018, lot HI 00713 (CUIC). Abbreviations: afs, apical fringe seta; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite; gc 2, apical gonocoxite; les, lateral ensiform setae." figureDoi="10.3897/zookeys.1044.59674.figure4" httpUri="https://binary.pensoft.net/fig/553680" pageId="0" pageNumber="229">4C</figureCitation>
); basal gonocoxite 1 narrow, elongate, a single apical fringe seta near apex of lateral apodeme; apical gonocoxite 2 broadened basally, with lateral extension directed dorsad to ventral surface of coxite bearing two lateral ensiform setae; one dorsal ensiform seta just apicad position of apical lateral ensiform seta; two apical nematiform setae situated in elongate fossa situated halfway between position of dorsal ensiform seta and tightly rounded apex of gonocoxite 2.
</paragraph>
<caption doi="10.3897/zookeys.1044.59674.figure10" httpUri="https://binary.pensoft.net/fig/553686" pageId="0" pageNumber="229" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" pageId="0" pageNumber="229">Figure 10.</emphasis>
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys aaa</emphasis>
</taxonomicName>
female reproductive tract, ventral view; Mountain View, Kazumura Cave, 21-xi-2018, lot HI00713 (CUIC). Abbreviations: bc, bursa copulatrix; co, common oviduct; sd, spermathecal duct; sg, spermathecal gland; sp, spermatheca. Schematic drawing breaks spermathecal duct at
<normalizedToken originalValue="“A”">&quot;A&quot;</normalizedToken>
to accommodate drawing to page size. Estimated length of spermathecal duct 1.7 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="etymology">
<paragraph pageId="0" pageNumber="229">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="229">
The species name
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys aaa</emphasis>
</taxonomicName>
incorporates the Hawaiian word '
<emphasis italics="true" pageId="0" pageNumber="229">
<normalizedToken originalValue="aa.‘ā">a'a.'a</normalizedToken>
</emphasis>
, meaning lava cave (
<bibRefCitation author="Na Puke Wehewehe ' Ōlelo Hawai'i" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="229" publicationUrl="wehewehe.org" refId="B74" refString="Na Puke Wehewehe 'Ōlelo Hawai'i, 2020. Ulukau Hawaiian Electronic Library. wehewehe.org" title="Ulukau Hawaiian Electronic Library." url="wehewehe.org" year="2020">
<normalizedToken originalValue="Nā">Na</normalizedToken>
Puke Wehewehe 'Ōlelo
<normalizedToken originalValue="Hawaii">Hawai'i</normalizedToken>
2020
</bibRefCitation>
). Being a Hawaiian word, the epithet is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="229" type="distribution">
<paragraph pageId="0" pageNumber="229">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="229">
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys aaa</emphasis>
</taxonomicName>
has attained a very broad subterranean distribution that includes lava tube caves within Mauna Loa and Kilauea volcanic flows. The species prefers the deep zone of lava tube caves (
<bibRefCitation author="Howarth, FG" journalOrPublisher="Entomologia Generalis" pageId="0" pageNumber="229" pagination="17 - 26" refId="B46" refString="Howarth, FG, 1982. Bioclimatic and geologic factors governing the evolution and distribution of Hawaiian cave insects. Entomologia Generalis 8: 17 - 26" title="Bioclimatic and geologic factors governing the evolution and distribution of Hawaiian cave insects." volume="8" year="1982">Howarth 1982</bibRefCitation>
), and can thus be considered troglobitic. The type locality, Kazumura Cave northeast of Kilauea (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributional records of Paratachys aaa. Abbreviations: H, Hilo; K, caldera of Kilauea Volcano; ML, summit caldera of Mauna Loa Volcano." figureDoi="10.3897/zookeys.1044.59674.figure11" httpUri="https://binary.pensoft.net/fig/553687" pageId="0" pageNumber="229">11</figureCitation>
), was formed approximately AD 1445 (
<bibRefCitation DOI="https://doi.org/10.1007/s004450050279" author="Clague, DA" journalOrPublisher="Bulletin of Volcanology" pageId="0" pageNumber="229" pagination="363 - 381" refId="B22" refString="Clague, DA, Hagstrum, JT, Champion, DE, Beeson, MH, 1999. Kilauea summit overflows: their ages and distribution in the Puna District, Hawai'i. Bulletin of Volcanology 61: 363 - 381, DOI: https://doi.org/10.1007/s004450050279" title="Kilauea summit overflows: their ages and distribution in the Puna District, Hawai'i." url="https://doi.org/10.1007/s004450050279" volume="61" year="1999">Clague et al. 1999</bibRefCitation>
), during an eruption of Mauna Loa that lasted approximately 50 years. Pauoa Cave and Burn Cave near the eastern tip of
<normalizedToken originalValue="Hawaii">Hawai'i</normalizedToken>
Island were formed from Puna volcanics derived from Kilauea eruptions that occurred less than 1000 years ago. Similarly, the caves in Kau accessed from sites in Ocean View Estates near South Point (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributional records of Paratachys aaa. Abbreviations: H, Hilo; K, caldera of Kilauea Volcano; ML, summit caldera of Mauna Loa Volcano." figureDoi="10.3897/zookeys.1044.59674.figure11" httpUri="https://binary.pensoft.net/fig/553687" pageId="0" pageNumber="229">11</figureCitation>
) were all formed during eruptive episodes of Mauna Loa involving k3 flows that range in age from 750-1500 years old (
<bibRefCitation DOI="https://doi.org/10.3133/ds144" author="Trusdell, FA" journalOrPublisher="Kontyu" pageId="0" pageNumber="229" refId="B91" refString="Trusdell, FA, Wolfe, EW, Morris, J, 2005. Digital database of the Geologic map of the island of Hawaii. DS 144. U.S. Geological Survey, Reston. https://doi.org/10.3133/ds144" title="Digital database of the Geologic map of the island of Hawaii. DS 144. U. S. Geological Survey, Reston." url="https://doi.org/10.3133/ds144" year="2005">Trusdell et al. 2005</bibRefCitation>
). The northwesterly Bobcat Cave north of the Mauna Loa caldera (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributional records of Paratachys aaa. Abbreviations: H, Hilo; K, caldera of Kilauea Volcano; ML, summit caldera of Mauna Loa Volcano." figureDoi="10.3897/zookeys.1044.59674.figure11" httpUri="https://binary.pensoft.net/fig/553687" pageId="0" pageNumber="229">11</figureCitation>
) is situated at the margin of a k3 flow overlying an older k2 flow, giving an age of origin for that cave of at most 1500 years ago. And most recently, Kaumana Cave just WSW of Hilo, formed during the 1880-1881 eruption of Mauna Loa. Thus
<taxonomicName lsidName="P. aaa" pageId="0" pageNumber="229" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">P. aaa</emphasis>
</taxonomicName>
occupies a disparate array of recently formed lava tube caves derived from several eruptive episodes. That these tiny beetles have attained such a broad distributional range among caves of different flows supports
<bibRefCitation DOI="https://doi.org/10.1126/science.175.4019.325" author="Howarth, FG" journalOrPublisher="Science" pageId="0" pageNumber="229" pagination="325 - 326" refId="B44" refString="Howarth, FG, 1972. Cavernicoles in lava tubes on the Island of Hawaii. Science 175: 325 - 326, DOI: https://doi.org/10.1126/science.175.4019.325" title="Cavernicoles in lava tubes on the Island of Hawaii." url="https://doi.org/10.1126/science.175.4019.325" volume="175" year="1972">
<normalizedToken originalValue="Howarths">Howarth's</normalizedToken>
(1972)
</bibRefCitation>
proposal that Hawaiian cave animals occupy both the larger lava tube caves accessible to humans (and collectors), but also the fractal network of mesocaverns ranging from 0.1-20 cm in diameter that connect those tubes. Add lignified organic layers sandwiched between older and newer flows (
<bibRefCitation DOI="https://doi.org/10.1029/JB094iB12p17465" author="Moore, JG" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="229" refId="B73" refString="Moore, JG, Clague, DA, Holcomb, RT, Lipman, PW, Normark, WR, Torresan, ME, 1989. Prodigious submarine landslides on the Hawaiian Ridge. Journal of Geophysical Research 94 (No. B12): 17465-17484. https://doi.org/10.1029/JB094iB12p17465" title="Prodigious submarine landslides on the Hawaiian Ridge. Journal of Geophysical Research 94 (No. B 12): 17465 - 17484." url="https://doi.org/10.1029/JB094iB12p17465" year="1989">Moore et al. 1989</bibRefCitation>
) to the mix, and one obtains a complex web of interconnected subterranean voids that supports a variety of microorganisms and the food web built upon them.
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys</emphasis>
</taxonomicName>
beetles serve as an apex predator in this system (
<bibRefCitation author="Howarth, FG" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="229" pagination="139 - 151" refId="B45" refString="Howarth, FG, 1973. The cavernicolous fauna of Hawaiian lava tubes, 1. Introduction. Pacific Insects 15: 139 - 151" title="The cavernicolous fauna of Hawaiian lava tubes, 1. Introduction." volume="15" year="1973">Howarth 1973</bibRefCitation>
), with their active foraging necessarily leading to dispersal among variously available and connected tubes and mesocaverns (
<bibRefCitation DOI="https://doi.org/10.1146/annurev.en.28.010183.002053" author="Howarth, FG" journalOrPublisher="Annual Review of Entomology" pageId="0" pageNumber="229" pagination="365 - 389" refId="B47" refString="Howarth, FG, 1983. Ecology of cave arthropods. Annual Review of Entomology 28: 365 - 389, DOI: https://doi.org/10.1146/annurev.en.28.010183.002053" title="Ecology of cave arthropods." url="https://doi.org/10.1146/annurev.en.28.010183.002053" volume="28" year="1983">Howarth 1983</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.5038/1827-806X.16.1.1" author="Howarth, FG" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="229" pagination="1 - 16" refId="B48" refString="Howarth, FG, 1987. The evolution of non-relictual tropical troglobites. International Journal of Speleology 16: 1 - 16, DOI: https://doi.org/10.5038/1827-806X.16.1.1" title="The evolution of non-relictual tropical troglobites." url="https://doi.org/10.5038/1827-806X.16.1.1" volume="16" year="1987">1987</bibRefCitation>
). Whereas above-ground elevational variation in habitats coupled with geographical variation in rainfall and temperature serve to define geographic distributions, the relatively more homogeneous subterranean voids would allow beetle dispersal among variously connected mesocaverns. Given that their bodies are only 2 mm long, requiring only occasional meals and encounters with conspecifics of the opposite sex to support persistent populations among the variously connected cave systems,
<taxonomicName lsidName="P. aaa" pageId="0" pageNumber="229" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">P. aaa</emphasis>
</taxonomicName>
have dispersed underground to colonize a broad swath of recently built
<normalizedToken originalValue="Hawaii">Hawai'i</normalizedToken>
Island.
</paragraph>
<caption doi="10.3897/zookeys.1044.59674.figure11" httpUri="https://binary.pensoft.net/fig/553687" pageId="0" pageNumber="229" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="229">
<emphasis bold="true" pageId="0" pageNumber="229">Figure 11.</emphasis>
Distributional records of
<taxonomicName authorityName="Liebherr" authorityYear="2021" class="Insecta" family="Carabidae" genus="Paratachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paratachys aaa" order="Coleoptera" pageId="0" pageNumber="229" phylum="Arthropoda" rank="species" species="aaa">
<emphasis italics="true" pageId="0" pageNumber="229">Paratachys aaa</emphasis>
</taxonomicName>
. Abbreviations: H, Hilo; K, caldera of Kilauea Volcano; ML, summit caldera of Mauna Loa Volcano.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>

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<paragraph id="9C3E76E0E30E4D3BB87600CB059BB531" pageId="15" pageNumber="142">
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<taxonomicNameLabel id="0381242C2BB647599EDE6EC5274961B8" pageId="15" pageNumber="142">sp. n.</taxonomicNameLabel>
Figure 7A E
</paragraph>
</subSubSection>
<subSubSection id="A9B5AF46CE512D04847DB21DDD2CEC04" pageId="15" pageNumber="142" type="diagnosis">
<paragraph id="62C9603AFDC7543D9BAF6D0E095759F2" pageId="15" pageNumber="142">Diagnosis.</paragraph>
<paragraph id="A876F5A11AFB94F97C1C0DEFED80A417" pageId="15" pageNumber="142">
<taxonomicName id="D790F4E34BB023F11FB48ABC1009A5ED" class="Hexapoda" family="Ismaridae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus laevigatus" order="Hymenoptera" pageId="15" pageNumber="142" phylum="Arthropoda" rank="species" species="laevigatus">Ismarus laevigatus</taxonomicName>
sp. n. is similar to
<taxonomicName id="EED6EFCA285186E26FA3ECC84447E193" lsidName="I. spinalis" pageId="15" pageNumber="142" rank="species" species="spinalis">I. spinalis</taxonomicName>
Kolyada &amp; Chemyreva, 2016 from the Palaearctic region. It differs mainly in the female antennal segments ratio and male antennal structure: in
<taxonomicName id="12EA219379527DA9753D95671CA2AEBF" lsidName="I. spinalis" pageId="15" pageNumber="142" rank="species" species="spinalis">I. spinalis</taxonomicName>
A7-A14 subquadrate, male A3 and A4 with sharp keels; in
<taxonomicName id="D104F27CEA429DD4C216B095C8D25FE4" lsidName="I. laevigatus" pageId="15" pageNumber="142" rank="species" species="laevigatus">I. laevigatus</taxonomicName>
sp. n. A7-A14 slightly longer than wide, male A4 with blade carina.
</paragraph>
</subSubSection>
<subSubSection id="3F9A49E47D04A093FA39C35611D43E84" pageId="15" pageNumber="142" type="type material">
<paragraph id="763D4178943D98EC4D0C17CB63AC3800" pageId="15" pageNumber="142">Type material</paragraph>
<paragraph id="D12D681218DDE4F1A8FE2CA62C87FDEE" pageId="15" pageNumber="142">(2♀♀9♂♂). Holotype, 1♀, SOUTH AFRICA: 75 km West South West of Natal, Estcourt, Cathedral Peaks Forest Station, 1500 m alt., 7-31.XII.1979 (MT), S. &amp; J. Peck leg., CJDAF010082 (deposited in CNCI). Allotype, 1♂, SOUTH AFRICA: 75 km West South West of Natal, Estcourt, Cathedral Peaks Forest Station, 1500 m alt., 7-31.XII.1979 (MT), S. &amp; J. Peck leg., CJDAF010084 (CNCI). Paratype, SOUTH AFRICA: 1♀5♂♂, 75 km West South West of Natal, Estcourt, Cathedral Peaks Forest Station, 1500 m alt., 7-31.XII.1979 (MT), S. &amp; J. Peck leg., CJDAF010083, 010085-010089 (CNCI); 1♂, ditto, 11-31.X.1979 (MT), S. &amp; J. Peck leg. (CNCI); 1♂, E. Cape Province, Katberg, 4000 ft alt., XII.1932, R. E. Turner leg., CJDAF010090 (NHMUK); 1♂, ditto, 14-26.XI.1932, R. E. Turner leg., CJDAF010091 (NHMUK).</paragraph>
</subSubSection>
<subSubSection id="56EBA52B6225A82D01C00AC09922ADC3" lastPageId="17" lastPageNumber="144" pageId="15" pageNumber="142" type="description">
<paragraph id="C8E52C1EF3D60453D1B7754CF921D0E1" pageId="15" pageNumber="142">Description.</paragraph>
<paragraph id="0438F1D2728A3C501ECD0B9A4E23935B" pageId="15" pageNumber="142">
Holotype (Female). Head. Head in dorsal view much wider than long (3:2), slightly wider than width of mesosoma (25:23) (Fig. 7
<normalizedToken id="4A3F415C5083DCF2F3E037B340AD0731" originalValue="AB">A-B</normalizedToken>
); POL: 11; LOL: 6; OOL: 9 (Fig. 7B); ocelli large, LOL slightly longer than diameter of lateral ocellus (12:11); vertex behind ocelli nearly flat in lateral view; eye large and without setae; inner orbits, frons and temple with few sparse setae; above antennal sockets, face and cheek with few long setae; antenna much shorter than body length (13:18); scape and pedicel with scattered setae; A3-A15 with dense and short setae; antennal segments in following proportions (length:width): 18:5; 7:4; 10:3; 10:2.5; 8:4; 6:4.5; 6:4.5; 6:4.5; 6:5; 6:5; 6:5; 6:5; 6:5; 6:6; 11:5 (Fig. 7A).
</paragraph>
<caption id="61CE23DCB8B4FC0A9514A6D4F9DC81F7" pageId="15" pageNumber="142">
<paragraph id="92CDE611592859E0D442D8A778530461" pageId="15" pageNumber="142">
Figure 7.
<taxonomicName id="B659DB8D1C3B66D04CF9AFEA5327088F" class="Hexapoda" family="Ismaridae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus laevigatus" order="Hymenoptera" pageId="15" pageNumber="142" phylum="Arthropoda" rank="species" species="laevigatus">Ismarus laevigatus</taxonomicName>
sp. n. (
<normalizedToken id="7D34B0007F46C41BE712FC0B30DFBB2E" originalValue="AB">A-B</normalizedToken>
, D Female C, E Male). A, C Habitus in lateral view B Head in dorsal view D Mesosoma in dorsal view E Antenna (A3-A5).
</paragraph>
</caption>
<paragraph id="5BAB3639EAF1A4F1EB333BC301366EC2" pageId="15" pageNumber="142">Mesosoma. Pronotum in dorsal view punctate-rugose with whitish long setae; pronotal shoulders angled; upper part of lateral pronotum smooth and concave in the middle, lower part of lateral pronotum punctate-rugose; mesoscutum smooth and convex with pairs of long setae in front of scutellar pit; notauli present anteriorly as large pits (Fig. 7D); humeral sulcus deep and short, as long as length of tegula; scutellum smooth and slightly convex, posterior rim rounded (Fig. 7D); anterior scutellar pit large and deep, shorter than remaining scutellar disc, slightly crenulate at bottom, median keel present (Fig. 7D); mesopleuron smooth with deep crenulate line along posterior margin; metapleuron punctate to rugose and covered with dense whitish long setae.</paragraph>
<paragraph id="A09B2ADA69B67E413EF8577E411CD9D1" pageId="15" pageNumber="142">
Wings. Radial cell completely closed, 2.1
<normalizedToken id="5C3637EBD4B6E482158360F01988C0A2" originalValue="×">x</normalizedToken>
as long as wide and as long as marginal vein (Fig. 7A).
</paragraph>
<paragraph id="12404B349D321C5D18EC70D5DDFE440A" pageId="15" pageNumber="142">Legs. Fore and mid legs slender; hind tibiae incrassate posteriorly, its maximum width slightly wider than hind femora (9:8).</paragraph>
<paragraph id="BD0C6B0808BF510E32F82A214A8129EA" pageId="15" pageNumber="142">
Metasoma. Petiole subquadrate, rugose dorsally; tergites smooth, with scattered setigerous punctures; base of second tergite with several short costae basally and very long median furrow, 0.90
<normalizedToken id="7769C2D30E82B95BEAFE9F315DD0FD40" originalValue="×">x</normalizedToken>
length of second; sutures between tergites complete and deeply impressed.
</paragraph>
<paragraph id="7CF00143FD4AD9A3E43E4DF7301583B3" pageId="15" pageNumber="142">Colour. Body black; antennae, tegulae and legs uniformly yellow except inner part of tibiae brown; wings hyaline, covered with brown setae.</paragraph>
<paragraph id="71C3C0A1BF0F58C29102F7707EE26E77" pageId="15" pageNumber="142">Measurements. Head length 0.47 mm, width 0.74 mm; mesosoma length 0.97 mm, width 0.69 mm; metasoma length 1.20 mm; fore wing length 2.52 mm; body length 2.63 mm.</paragraph>
<paragraph id="91F9E8CEDAEE9BE22F738D5A8BC7634D" lastPageId="17" lastPageNumber="144" pageId="16" pageNumber="143">
<pageBreakToken id="98A754F2B8538182BE363562E967F1C5" pageId="16" pageNumber="143" start="start">Male</pageBreakToken>
(Allotype). Body length 1.88 mm. Similar to female, POL: 8; LOL: 5; OOL: 7; body dark brown except metasoma brown, antennae brown except scape and pedicel yellow, legs yellow except hind tibiae and tarsi yellowish brown to brown; anterior margin of lateral pronotum rugose; blade-like carina on A4 percurrent (Fig. 7E); antennal
<pageBreakToken id="CBEC8CB3B4B21516C9D8A433E9563284" pageId="17" pageNumber="144" start="start">segments</pageBreakToken>
in following proportions: 13:4; 7:3.5; 7:3.5; 7:3.5; 5:4; 5:4; 5:4; 5:4; 5:4; 5:4; 5:4; 5:4; 5:4; 8:4 (Fig. 7C).
</paragraph>
</subSubSection>
<subSubSection id="589B5CB05594B5995E026F85F24B6722" pageId="17" pageNumber="144" type="variation">
<paragraph id="4047BB695E415BE9FF93472618747C2B" pageId="17" pageNumber="144">Variation.</paragraph>
<paragraph id="169CFFA2D5F055C28C7CB9EEA4D56266" pageId="17" pageNumber="144">
Body length 1.88-2.63 mm in both sexes. Median furrow very long extending 0.80.9
<normalizedToken id="53927D0A61EAEB4DF14AF3FEC76388B1" originalValue="×">x</normalizedToken>
length of second tergite in both sexes.
</paragraph>
</subSubSection>
<subSubSection id="0B0EAD314B9B0A7B1E7F6BB078BE7443" pageId="17" pageNumber="144" type="distribution">
<paragraph id="FBDB5D4AE4579E724DBAE11C7A555C80" pageId="17" pageNumber="144">Distribution.</paragraph>
<paragraph id="8F648738014E8A2B7007469177333308" pageId="17" pageNumber="144">South Africa.</paragraph>
</subSubSection>
<subSubSection id="90D62E8D8A437A24B230D6E82A18C4FD" pageId="17" pageNumber="144" type="etymology">
<paragraph id="0D28BBC729AF0C6027FCC73450AF3CFB" pageId="17" pageNumber="144">Etymology.</paragraph>
<paragraph id="ED708463077140A03006DC6EFE6AA091" pageId="17" pageNumber="144">
The specific name
<taxonomicName id="0B8AC76DCF12B056C874101E996AD7EA" genus="Chrysidoidea" lsidName="Chrysidoidea laevigatus" pageId="17" pageNumber="144" rank="species" species="laevigatus">laevigatus</taxonomicName>
is derived from the Latin adjective which means smooth.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,143 @@
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<paragraph id="7F67B74D839DD3C136204B65000F831D" pageNumber="421">
<taxonomicName id="2BD469A1F7AA2159CAE489382F6D0E61" ID-CoL="5Y55Y" authority="Peters 1860" authorityName="Peters" authorityYear="1860" class="Mammalia" family="Phyllostomidae" genus="Chiroderma" kingdom="Animalia" order="Chiroptera" pageNumber="421" phylum="Chordata" rank="species" species="villosum">
<emphasis id="75C0EE581F60507A6904A2267296313C" italics="true" pageNumber="421">Chiroderma villosum</emphasis>
Peters 1860
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CAD7611490558A66029D496AB905D165" pageNumber="421" type="reference_group">
<paragraph id="658D21759D883E545E5A4795B8685A10" pageNumber="421">
<treatmentCitation id="BCA4C680F343EA1E7B159D0915DD0CC9" author="Peters" authority="Peters 1860" authorityName="Peters" authorityYear="1860" class="Mammalia" family="Phyllostomidae" genus="Chiroderma" journal="Monatsb. K. Preuss. Akad. Wiss. Berlin" kingdom="Animalia" order="Chiroptera" page="748" pageNumber="421" phylum="Chordata" rank="species" species="villosum" title="Chiroderma villosum" volume="1860" year="1860">
<taxonomicName id="121EB32DB6B6AC444C6E56D583633A7B" ID-CoL="5Y55Y" authority="Peters 1860" authorityName="Peters" authorityYear="1860" class="Mammalia" family="Phyllostomidae" genus="Chiroderma" kingdom="Animalia" order="Chiroptera" pageNumber="421" phylum="Chordata" rank="species" species="villosum">
<emphasis id="2DD1E0FE2FE9A36477873B8B340231AD" italics="true" pageNumber="421">Chiroderma villosum</emphasis>
Peters 1860
</taxonomicName>
,
<bibCitation id="3CDDFB1EEBEB82966B144A12581F9E34" author="Peters" journal="Monatsb. K. Preuss. Akad. Wiss. Berlin" pageNumber="421" pagination="748" title="Chiroderma villosum" volume="1860" year="1860">Monatsb. K. Preuss. Akad. Wiss. Berlin, 1860: 748</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86D3F2AB28D0DF96CDFB1C87E9B4A86F" pageNumber="421" type="materials_examined">
<paragraph id="1B2BBD1855F9BB796EDB7BF558FDB2F1" pageNumber="421">
<emphasis id="6706D3355CC4FD25C27FB2EBEE51789D" inLineHeading="true" pageNumber="421">Type Locality:</emphasis>
<materialsCitation id="80EBF181BC71514837B3D73D3534D1AE" ID-GBIF-Occurrence="4867283445" country="Brazil" pageNumber="421">
<collectingCountry id="215BF6B813019AEDA60B19776B8744B8" name="Brazil">Brazil</collectingCountry>
; see
<bibRefCitation id="7D58D143B231B634AE1609F2175F8790" author="Carter, D. C. &amp; P. G. Dolan" refId="ref53891" refString="Carter, D. C., and P. G. Dolan. 1978. Catalog of type specimens of Neotropical bats in selected European museums. Special Publications, The Museum, Texas Tech University Press, 15: 1 - 136." year="1978">Carter and Dolan (1978)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8034DEEDBBFFB74FD77D511116782DD" pageNumber="421" type="vernacular_names">
<paragraph id="81A4AA91D46326C381896F35A246D2D0" pageNumber="421">
<emphasis id="E8416FD89ECAD9C8F97120159EEE4B2C" inLineHeading="true" pageNumber="421">Vernacular Names:</emphasis>
<vernacularName id="AF0FE4402EB8AEC19C8125D2E3F59BDF" language="eng" pageNumber="421">Hairy Big-eyed Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="55F1AAA2C3B1D64ED1800B4F81A51076" pageNumber="421" type="child_taxa">
<paragraph id="C97BEB9B6A43B81976EFAF2CB81668AB" pageNumber="421">
<emphasis id="B325F1986DA4BA359919210FF48E2E75" inLineHeading="true" pageNumber="421">Subspecies:</emphasis>
:
</paragraph>
<paragraph id="8DF58A92432F552A5D5FACB01200D537" pageNumber="421">
Subspecies
<taxonomicName id="280DBE3C3F21DC09E29C35EA06CAEE0D" authority="Peters 1860" authorityName="Peters" authorityYear="1860" class="Mammalia" family="Phyllostomidae" genus="Chiroderma" kingdom="Animalia" order="Chiroptera" pageNumber="421" phylum="Chordata" rank="subSpecies" species="villosum" subSpecies="villosum">
<emphasis id="9B2EFD36A0D5EBF1F98E3506D6A48580" italics="true" pageNumber="421">Chiroderma villosum</emphasis>
subsp.
<emphasis id="D980E49F065BE01B787C94E1E35FEF4F" italics="true" pageNumber="421">villosum</emphasis>
Peters 1860
</taxonomicName>
</paragraph>
<paragraph id="A62FF56D02F7BBB5597B5DA40A478815" pageNumber="421">
Subspecies
<taxonomicName id="10D422D601D3AAE7628831EF16014C86" authority="J. A. Allen 1900" authorityName="J. A. Allen" authorityYear="1900" class="Mammalia" family="Phyllostomidae" genus="Chiroderma" kingdom="Animalia" order="Chiroptera" pageNumber="421" phylum="Chordata" rank="subSpecies" species="villosum" subSpecies="jesupi">
<emphasis id="77C93217EF5B654A076C9B2220DDD085" italics="true" pageNumber="421">Chiroderma villosum</emphasis>
subsp.
<emphasis id="8E9C3280B9FF54E55D563C21FBCBDEEC" italics="true" pageNumber="421">jesupi</emphasis>
J. A.
<bibRefCitation id="7471B7BD78D45259E3C2A185967E3D21" author="Allen, J. A." refId="ref5416" refString="Allen, J. A. 1900. Description of new American marsupials. Bulletin of the American Museum of Natural History, 13: 191 - 199." year="1900">Allen 1900</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6BC96EAD40CB42DE7C000D32620E669E" pageNumber="421" type="distribution">
<paragraph id="A3A97A6B2F6F31B7C38A24459D266D9A" pageNumber="421">
<emphasis id="6AF9B04E787D9C6673440B5C91DF2783" inLineHeading="true" pageNumber="421">Distribution:</emphasis>
<collectingRegion id="9EA5E5E9852ECB4E0A63D947507D0433" country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
(
<collectingCountry id="94848032F606DA77609FC94E43E676B7" name="Mexico">Mexico</collectingCountry>
) south to S
<collectingCountry id="6081ABF597F935E1E2E743C60CE712DE" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="C79ED3C71711774FA37D2B2EACC11E5C" name="Bolivia">Bolivia</collectingCountry>
and
<collectingCountry id="0B1D3A26444CD43F697A9ECB43990581" name="Peru">Peru</collectingCountry>
;
<collectingCountry id="0136766C0247BC57DD7EF3A895197257" name="Trinidad and Tobago">Trinidad and Tobago</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="498E6B1C354AF5B8B18D667A4A39646F" pageNumber="421" type="conservation">
<paragraph id="98D9354372DB059BEDC7875F4EF33595" pageNumber="421">
<emphasis id="EDD28BA392B43E89E310F4CAC3C20BB2" inLineHeading="true" pageNumber="421">Conservation:</emphasis>
<collectionCode id="A4A7AB7A8EA49281D750F346426560C5">IUCN</collectionCode>
2003 and
<collectionCode id="C22CD194B8C752FF543C895C48396E82">IUCN</collectionCode>
/
<collectionCode id="CDC33D94E1171EA67C587F3BEC59CABA" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="2E228BF520AE7284E6B91A55D7837431" pageNumber="421" type="discussion">
<paragraph id="7890CA8C413C046DD9A52D3A3FD36385" pageNumber="421">
<emphasis id="3910965A25B3CF9F0809D17D25E9AC3D" inLineHeading="true" pageNumber="421">Discussion:</emphasis>
See
<bibRefCitation id="A8B92004616F947C6AF745C408F82652" author="Handley, C. O., Jr." refId="ref138975" refString="Handley, C. O., Jr. 1960. Descriptions of new bats from Panama. Proceedings of the United States National Museum, 112: 459 - 479." year="1960">Handley (1960)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.903.37775" ID-GBIF-Dataset="d1383bc2-3cb4-4e6e-a637-c30574637a45" ID-PMC="PMC6976704" ID-Pensoft-Pub="1313-2970-903-1" ID-Pensoft-UUID="DC1A8D36F34457478AD7EBDCD89B21CF" ID-PubMed="31997887" ID-ZooBank="60467E58F35B4D2DBC6182568F56AD89" ModsDocID="1313-2970-903-1" checkinTime="1579185758659" checkinUser="pensoft" docAuthor="Cerretti, Pierfilippo, Badano, Davide, Gisondi, Silvia, Giudice, Giuseppe Lo &amp; Pape, Thomas" docDate="2020" docId="0C026A8A9BF958C687B0D02322EF9DF6" docLanguage="en" docName="ZooKeys 903: 1-130" docOrigin="ZooKeys 903" docSource="http://dx.doi.org/10.3897/zookeys.903.37775" docTitle="Kinabalumyia Cerretti &amp; Pape 2020, gen. nov." docType="treatment" docUuid="5E01603E-D354-4B84-BA64-0064DD960B14" docUuidSource="ZooBank" docVersion="6" id="DC1A8D36F34457478AD7EBDCD89B21CF" lastPageNumber="1" masterDocId="DC1A8D36F34457478AD7EBDCD89B21CF" masterDocTitle="The world woodlouse flies (Diptera, Rhinophoridae)" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1668168302666" updateUser="ExternalLinkService">
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<mods:role>
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<mods:namePart>Giudice, Giuseppe Lo</mods:namePart>
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<mods:role>
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<treatment ID-GBIF-Taxon="161756140" LSID="urn:lsid:zoobank.org:act:5E01603E-D354-4B84-BA64-0064DD960B14" httpUri="http://treatment.plazi.org/id/0C026A8A9BF958C687B0D02322EF9DF6" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="0C026A8A-9BF9-58C6-87B0-D02322EF9DF6" authority="Cerretti &amp; Pape" authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">Kinabalumyia Cerretti &amp; Pape</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">gen. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Kinabalumyia gen. nov. A-C, F-I Kinabalumyia pinax sp. nov. (Malaysia, Sabah) D-E Kinabalumyia sp. ♀ (Philippines) A K. pinax sp. nov., habitus in lateral view B K. pinax sp. nov., abdomen in dorsal view C K. pinax sp. nov., head in lateral view D Kinabalumyia sp. ♀, head in frontal view, ♀ E Kinabalumyia sp. ♀, head in lateral view F K. pinax sp. nov., wing G-I K. pinax sp. nov.: epandrial complex in posterior (G) and lateral (H) view I hypandrial complex in lateral view [A-C holotype; F-I paratypes]." figureDoi="10.3897/zookeys.903.37775.figure18" httpUri="https://binary.pensoft.net/fig/372568" pageId="0" pageNumber="1">Fig. 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Unambiguous character state changes</paragraph>
<paragraph pageId="0" pageNumber="1">
(Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of characters and states." httpUri="http://table.plazi.org/id/A6B55C4624CF9A1BE4266A3B81981756" pageId="0" pageNumber="1" tableUuid="A6B55C4624CF9A1BE4266A3B81981756">1</tableCitation>
, Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phylogeny of Rhinophoridae based on morphological evidence. Single tree obtained under IW (total fit: 62.33), enforcing a k-value of 4, with unambiguous character state changes mapped above branches [black squares: global apomorphies (i. e., uncontradicted and unreversed apomorphic character states; white squares: local apomorphies (i. e., homoplasious character state due to convergence or reversal)] and new taxa highlighted in bold." figureDoi="10.3897/zookeys.903.37775.figure20" httpUri="https://binary.pensoft.net/fig/372570" pageId="0" pageNumber="1">20</figureCitation>
). Global apomorphies: 81:1; local apomorphies: 21:1, 53:1, 59:1.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: head higher than long in lateral view. Facial ridge approx. 1.3 times as long as frons. Ocellar setae present though small. Frons 0.9-1.1 times as wide as compound eye in dorsal view. Median vertical setae strong and slightly converging. Two or three slightly convergent and reclinate frontal setae, descending to approx. half level of pedicel. Fronto-orbital plate nearly bare with one proclinate orbital seta (no sexual dimorphism). One upper lateroclinate orbital seta. Parafacial bare, at its narrowest point approx. twice as wide as maximum diameter of arista. Vibrissal angle strongly receding. Vibrissa well developed, arising slightly below the level of lower facial margin. Lower facial margin not sunken and slightly visible in lateral view. Facial ridge strongly concave with two or three short setulae above vibrissa. Face concave but antennae not hidden and clearly visible in lateral view. Antenna long and wide (in lateral view), much longer than height of gena. Postpedicel 4.5-6.5 times as long as pedicel. Postpedicel axe-shaped in male, more or less stick-like in female. Arista bare (or apparently so). First and second aristomere slightly thickened and strongly elongated. Lunule bare. Gena, in profile, approx. 3/5 (male), 1/2 (female) as high as compound eye. Palpus pretty short, dark brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
: prosternum bare. Postpronotum with two setae. Two postsutural supra-alar setae (first postsutural supra-alar seta absent). Scutellum with one pair of strong lateral setae and one pair of short, crossed and horizontal apical setae. Anatergite with a tuft of short setulae below lower calypter. Subscutellum moderately swollen, mostly, though not entirely, sclerotised. Metathoracic spiracular lappets nearly undeveloped. Lower calypter distinctly tongue-shaped (ground-plan trait of
<taxonomicName authorityName="Robineau-Desvoidy" authorityYear="1863" class="Insecta" family="Rhinophoridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Rhinophoridae</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphological terminology of Rhinophoridae, head, thorax and abdomen. A Chaetotaxy of head, thorax and abdomen in dorsal view B, C arrangement of setae of postpronotum: B three setae arranged in shallow triangle C three setae standing in a nearly right-angled triangle D thorax, sclerites and chaetotaxy, in lateral view E wing veins and cells. Drawings by Giulia Bellanti." figureDoi="10.3897/zookeys.903.37775.figure2" httpUri="https://binary.pensoft.net/fig/372552" pageId="0" pageNumber="1">2E</figureCitation>
). Costal sector cs2 setose ventrally. Costal spine not differentiated from general costal setae. Costal sector cs5 clearly longer than costal sector cs2. Vein R1 entirely bare. Vein R4+5 bare dorsally. Bend of vein M1 absent. Crossvein dm-m forming a right angle with proximal section of M4. Vein CuA+CuP reaching wing margin. Preapical anterodorsal seta of fore tibia slightly longer than preapical dorsal seta. Fore tarsus not compressed. Tibiae of mid and hind leg normally developed. Hind tibia with three dorsal preapical setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
: tergites virtually without microtomentum and with relatively strong and recumbent general setulae; syntergite 1+2, tergites 3 and 4 with a pair of strong median marginal setae, without median discal setae, tergite 5 with strong marginal setae without discal setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male terminalia</emphasis>
: posterior margin of sternite 5 slightly concave, almost straight. Tergite 6 plate-like, with median marginal setae; tergite 6 and syntergosternite 7+8 fused. Connection between sternite 6 and syntergosternite 7+8 fused on right side. Cerci well developed and entirely fused medially into a syncercus. Surstylus long, narrow (almost stick-like), slightly enlarged distally and gently curved anteriorly. Surstylus not fused to epandrium. Bacilliform sclerite articulated (i.e., not fused) to laterobasal margin of surstylus. Hypandrial arms not fused medially. Connection between phallic guide and pregonite membranous. Pregonite well developed and lobe-like. Postgonite without anterior seta. Epiphallus well developed apically, pointed and attached dorsomedially to basiphallus. Extension of dorsal sclerite of distiphallus divided medially into two hemisclerites which are proximally fused to dorsal sclerite of distiphallus. Median process of ventral sclerotisation of distiphallus present, divided longitudinally and interrupted proximally, i.e., not connected to ventral plate. Acrophallus simple and scale-like spinules not differentiated.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Oriental - Indonesia (Bali), Malaysia (Sabah), Philippines (Palawan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia pinax" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia pinax</emphasis>
</taxonomicName>
Cerretti &amp; Pape, sp. nov., by present designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The generic name is a composite word formed from the name of the type locality of the type species, Mount Kinabalu, and from the Greek word
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="μύγα">μύγα</normalizedToken>
</emphasis>
(miga), meaning fly. The name should be treated as a feminine noun.
</paragraph>
<caption doi="10.3897/zookeys.903.37775.figure18" httpUri="https://binary.pensoft.net/fig/372568" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia</emphasis>
</taxonomicName>
gen. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A-C, F-I</emphasis>
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia pinax" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia pinax</emphasis>
</taxonomicName>
sp. nov. (Malaysia, Sabah)
<emphasis bold="true" pageId="0" pageNumber="1">D-E</emphasis>
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia</emphasis>
</taxonomicName>
sp. ♀ (Philippines)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName lsidName="K. pinax" pageId="0" pageNumber="1" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">K. pinax</emphasis>
</taxonomicName>
sp. nov., habitus in lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName lsidName="K. pinax" pageId="0" pageNumber="1" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">K. pinax</emphasis>
</taxonomicName>
sp. nov., abdomen in dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName lsidName="K. pinax" pageId="0" pageNumber="1" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">K. pinax</emphasis>
</taxonomicName>
sp. nov., head in lateral view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia</emphasis>
</taxonomicName>
sp. ♀, head in frontal view, ♀
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Cerretti &amp; Pape" authorityYear="2020" class="Insecta" family="Rhinophoridae" genus="Kinabalumyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kinabalumyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Kinabalumyia</emphasis>
</taxonomicName>
sp. ♀, head in lateral view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName lsidName="K. pinax" pageId="0" pageNumber="1" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">K. pinax</emphasis>
</taxonomicName>
sp. nov., wing
<emphasis bold="true" pageId="0" pageNumber="1">G-I</emphasis>
<taxonomicName lsidName="K. pinax" pageId="0" pageNumber="1" rank="species" species="pinax">
<emphasis italics="true" pageId="0" pageNumber="1">K. pinax</emphasis>
</taxonomicName>
sp. nov.: epandrial complex in posterior (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and lateral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) view
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
hypandrial complex in lateral view [
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
holotype;
<emphasis bold="true" pageId="0" pageNumber="1">F-I</emphasis>
paratypes].
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.834.28800" ID-GBIF-Dataset="0ab2498c-ae69-4bde-8fc2-271f6d40253c" ID-PMC="PMC6495053" ID-Pensoft-Pub="1313-2970-834-1" ID-PubMed="31105437" ID-ZBK="A9309D4615834D33A6B7F824BC3160FD" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-834-1" ModsDocOrigin="ZooKeys 834" ModsDocTitle="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)" checkinTime="1555318553194" checkinUser="pensoft" docAuthor="Inkhavilay, Khamla, Sutcharit, Chirasak, Bantaowong, Ueangfa, Chanabun, Ratmanee, Siriwut, Warut, Srisonchai, Ruttapon, Pholyotha, Arthit, Jirapatrasilp, Parin &amp; Panha, Somsak" docDate="2019" docId="0C02A1C12FC76AE1F026A263AC70ACCD" docLanguage="en" docName="ZooKeys 834: 1-166" docOrigin="ZooKeys 834" docSource="http://dx.doi.org/10.3897/zookeys.834.28800" docTitle="Cyclophorus franzhuberi Thach 2017" docType="treatment" docVersion="5" lastPageNumber="1" masterDocId="8E3DFF87FFA2874BEC66D510A643FFDE" masterDocTitle="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)" masterLastPageNumber="166" masterPageNumber="1" pageNumber="1" updateTime="1668167097005" updateUser="ExternalLinkService">
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<mods:title>Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)</mods:title>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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<mods:name type="personal">
<mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:namePart>Bantaowong, Ueangfa</mods:namePart>
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<mods:role>
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</mods:role>
<mods:namePart>Chanabun, Ratmanee</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siriwut, Warut</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pholyotha, Arthit</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:identifier type="ZooBank">A9309D4615834D33A6B7F824BC3160FD</mods:identifier>
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<treatment ID-GBIF-Taxon="156200583" LSID="urn:lsid:plazi:treatment:0C02A1C12FC76AE1F026A263AC70ACCD" httpUri="http://treatment.plazi.org/id/0C02A1C12FC76AE1F026A263AC70ACCD" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName authority="Thach, 2017" authorityName="Thach" authorityYear="2017" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus franzhuberi" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="franzhuberi">Cyclophorus franzhuberi Thach, 2017</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus franzhuberi" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="franzhuberi">Cyclophorus franzhuberi</taxonomicName>
Thach, 2017: 14, figs 54-56. Type locality: Thakhek, Khammouane Province, Central Laos.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype MNHN-IM-2000-33194 (Fig. 6F).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from the type locality in Laos (
<bibRefCitation author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio, USA" pageId="0" pageNumber="1" title="New shells of Southeast Asia with 2 new genera and 85 new species." year="2017">Thach 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">No material of this species was found, and only the type specimens were examined.</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 6. A, B
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx messageri" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="messageri">Dioryx messageri</taxonomicName>
A syntype MNHN-IM-2000-31785 and B CUMZ collection C
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus heudei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="heudei">Metalycaeus heudei</taxonomicName>
, syntype MNHN-IM-2012-27169 D
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus laosensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="laosensis">Metalycaeus laosensis</taxonomicName>
, holotype MNHN-IM-2012-27172 E
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus floridus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="floridus">Cyclophorus floridus</taxonomicName>
, CUMZ collection F
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus franzhuberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="franzhuberi">Cyclophorus franzhuberi</taxonomicName>
, holotype MNHN-IM-2000-33194. Photos: B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
(C, D).
</paragraph>
</caption>
</subSubSection>
</treatment>
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@ -0,0 +1,742 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.211.89453" ID-Pensoft-Pub="1314-2003-211-1" ID-Pensoft-UUID="20DF43DC40DB5AF9BC61DF4AD7C16EF7" ModsDocID="1314-2003-211-1" checkinTime="1665870600856" checkinUser="pensoft" docAuthor="Siti-Munirah, Mat Yunoh, Suhaida, Mustafa &amp; Eddie, Chan" docDate="2022" docId="0C02AA5AE1A1500DA6495FD2F2626DD2" docLanguage="en" docName="PhytoKeys 211: 1-11" docOrigin="PhytoKeys 211" docPubDate="2022-10-11" docSource="http://dx.doi.org/10.3897/phytokeys.211.89453" docTitle="Thismia limkokthayi Siti-Munirah &amp; E. Chan 2022, sp. nov." docType="treatment" docVersion="2" id="20DF43DC40DB5AF9BC61DF4AD7C16EF7" lastPageNumber="1" masterDocId="20DF43DC40DB5AF9BC61DF4AD7C16EF7" masterDocTitle="Thismia limkokthayi (Thismiaceae): A new mycoheterotrophic species from Genting Highlands in Pahang, Malaysia" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1665870901284" updateUser="ExternalLinkService">
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<mods:namePart>Suhaida, Mustafa</mods:namePart>
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<mods:namePart>Eddie, Chan</mods:namePart>
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<taxonomicName LSID="0C02AA5A-E1A1-500D-A649-5FD2F2626DD2" authority="Siti-Munirah &amp; E. Chan" authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi" status="sp. nov.">Thismia limkokthayi Siti-Munirah &amp; E.Chan</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F2" captionText="Figure 1. Thismia limkokthayi with scale (all in centimetres (cm )) A plants with mature (FRI 91138 c) and young flowers with long stems B mature flower with a very short stem (FRI 91138 b) C stamens (FRI FRI 91138 a). Photos by Siti-Munirah MY." figureDoi="10.3897/phytokeys.211.89453.figure1" httpUri="https://binary.pensoft.net/fig/754973" pageId="0" pageNumber="1">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F3" captionText="Figure 2. Thismia limkokthayi A coralliform roots B full flower C claviform tip of mitre appendage D longitudinal section of floral tube, showing inner surface E outer tepal and inner tepal F side view of mitre G mitre viewed from above H six stamens, viewed from below I outer view of a stamen J inner view of two pendulous stamens K detail of the side lobe on the supraconnective apex (side lobe with truncate tips that are recurved inwards) L ovary with style, stigma and flower bud surrounded by bracts M stigmas. All from FRI FRI 91138 a (A, D, I, J, K, L, M), FRI 91138 b (H) and FRI 91138 c (B, C, E, F, G). Photos by Siti-Munirah MY; images not to scale (see dimensions in description and Fig. 1)." figureDoi="10.3897/phytokeys.211.89453.figure2" httpUri="https://binary.pensoft.net/fig/754974" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Thismia limkokthayi Siti-Munirah &amp; Eddie Chan A group photo with the T. limkokthayi (white arrow) in its habitat (from left; Angan A, Siti-Munirah MY, Eddie C, Ahmad Norsidar AH and Suhaida M) B-C different stages of anthesis of T. limkokthayi in its habitat (in situ) B two mature flowers from one individual C a fresh flower (on the left) and a new young flower starting to bloom (on the right). Photos by Siti-Munirah MY." figureDoi="10.3897/phytokeys.211.89453.figure3" httpUri="https://binary.pensoft.net/fig/754975" pageId="0" pageNumber="1">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
is distinguished from closely-related species by the following combination of characteristics: black to dark brown floral tube and appendages, the presence of mitre fovea between the bases of claviform appendages, the presence of longitudinal ribs and absent transverse bars on the inner side of the floral tube, the apex of stamens with a central lobe (prolongation of the rib) and two lateral lobes (the tips of each are recurved) and dark purple stigma.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
. Peninsular
<materialsCitation accessionNumber="FRI91138" collectingDate="2022-04-28" collectorName="Siti-Munirah MY" country="Malaysia" elevation="1137" location="Genting Highlands" municipality="Bentong District" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion country="Malaysia" name="Pahang">Pahang</collectingRegion>
,
<collectingMunicipality>Bentong District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0C02AA5AE1A1500DA6495FD2F2626DD2:F6117D98C70B827651C4969BA76EA914" country="Malaysia" municipality="Bentong District" name="Genting Highlands" stateProvince="Pahang">Genting Highlands</location>
, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.137" unit="m" value="1137.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.137" unit="m" value="1137.0">1137 m</elevation>
</quantity>
elev.,
<collectingDate value="2022-04-28">28 April 2022</collectingDate>
,
<collectorName>Siti-Munirah MY</collectorName>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91138">FRI91138</accessionNumber>
c (
<typeStatus>holotype</typeStatus>
KEP!, spirit collection, barcode no. SC12022)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Achlorophyllous herbs up to ca.
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.4" unit="cm" value="14.0">14 cm</quantity>
.
<emphasis bold="true" pageId="0" pageNumber="1">Roots</emphasis>
coralliform, surface hairy, apices brownish-white.
<emphasis bold="true" pageId="0" pageNumber="1">Stems</emphasis>
up to ca.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.5" unit="cm" value="4.5">4.5 cm</quantity>
tall or very short (possibly for young plants), ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
in diameter, erect, ascending, white and becoming brownish with age, glabrous, terete.
<emphasis bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
glabrous, pale brown, dark brown in the upper part or towards the apex, scale-like, triangular-ovate to lanceolate, up to
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.2" unit="mm" value="12.0">12 mm</quantity>
long,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
wide at the base, apex acute to acuminate, spirally arranged, more crowded in the upper part of the stem.
<emphasis bold="true" pageId="0" pageNumber="1">Floral bracts</emphasis>
3, pale brown to dark brown towards the apex, similar to upper leaves, but slightly larger,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10-12 mm</quantity>
long, apex acute to acuminate,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
wide at the base.
<emphasis bold="true" pageId="0" pageNumber="1">Pedicels</emphasis>
lengthen up to
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
during flower growth,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
wide.
<emphasis bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
solitary or in clusters of 2, opening in succession in the latter case and forming loose monochasial inflorescences, lateral flower located in the axil of one of the floral bracts of the terminal flower and also bearing its own floral bracts; each flower up to
<quantity metricMagnitude="-2" metricUnit="m" metricValue="8.0" unit="cm" value="8.0">8 cm</quantity>
long (including ovary and appendages);
<emphasis bold="true" pageId="0" pageNumber="1">perianth</emphasis>
actinomorphic with 6 tepals fused to form a floral tube with a dome-shaped mitre with 3 slender, claviform appendages on its top;
<emphasis bold="true" pageId="0" pageNumber="1">floral tube</emphasis>
black brownish/brown blackish, urceolate, ca.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20-25 mm</quantity>
long, ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 mm</quantity>
wide, constricted just above the ovary, widest in the upper part;
<emphasis bold="true" pageId="0" pageNumber="1">outer surface</emphasis>
with 6 longitudinal ribs, glabrous, rough, black to dark brown;
<emphasis bold="true" pageId="0" pageNumber="1">inner surface</emphasis>
with 6 greenish longitudinal ribs, without transverse bars, black to dark brown;
<emphasis bold="true" pageId="0" pageNumber="1">outer tepal lobes</emphasis>
3, brownish orange, minute, ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
long,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
wide at base, broadly triangular, erect;
<emphasis bold="true" pageId="0" pageNumber="1">inner tepal lobes</emphasis>
3, black to dark brown, thick, cuneate, surface glabrous, apically adnate to form a dome-shaped mitre;
<emphasis bold="true" pageId="0" pageNumber="1">mitre</emphasis>
with 3 lateral, round-shaped, ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 mm</quantity>
wide apertures, 3 hood-like accessory lobes, more curved during the early stages of flowering and flattening when the flower is older and matured;
<emphasis bold="true" pageId="0" pageNumber="1">mitre appendages</emphasis>
each ca.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27-30 mm</quantity>
long, their base wide and flattened, forming a fovea in the centre of the mitre, becoming narrower above, claviform at apex, glabrous, dark brown to pale orange towards tip.
<emphasis bold="true" pageId="0" pageNumber="1">Stamens</emphasis>
6, pendulous from the apical margin of the floral tube;
<emphasis bold="true" pageId="0" pageNumber="1">annulus</emphasis>
absent;
<emphasis bold="true" pageId="0" pageNumber="1">filaments</emphasis>
orange and white, curved downwards, with bases slightly emerged above floral tube apex, not connate, forming 6 apertures apparent when viewed from above;
<emphasis bold="true" pageId="0" pageNumber="1">connectives</emphasis>
broad, orange-yellowish in lower half and black/dark brown in upper half, laterally connate to form a tube, ca.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.2" unit="mm" value="12.0">12 mm</quantity>
long, each with prominent longitudinal rib extending along the entire length of the inner surface of the connective; supraconnective apex with one central lobe (extension of the rib) and two smaller side lobes with tips recurved inwards/truncate, glabrous;
<emphasis bold="true" pageId="0" pageNumber="1">lateral appendage</emphasis>
skirt-like, black, protruding towards the floral tube, not reaching connective apex, glabrous including on margins, only sparsely hairy on each horn-like corner; individual
<emphasis bold="true" pageId="0" pageNumber="1">stamens</emphasis>
with 2 thecae (abaxial, dehiscing towards the inner surface of the floral tube), each
<emphasis bold="true" pageId="0" pageNumber="1">theca</emphasis>
oblong,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
long;
<emphasis bold="true" pageId="0" pageNumber="1">interstaminal glands</emphasis>
elliptic-oblong, translucent, inserted on the line of fusion between connectives.
<emphasis bold="true" pageId="0" pageNumber="1">Ovary</emphasis>
inferior, obconical, ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
long, brown to dark, with 6 longitudinal ribs, unilocular;
<emphasis bold="true" pageId="0" pageNumber="1">placentas</emphasis>
3;
<emphasis bold="true" pageId="0" pageNumber="1">style</emphasis>
short, ca.
<quantity metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
long, black/purplish;
<emphasis bold="true" pageId="0" pageNumber="1">stigma</emphasis>
3-lobed, black-purplish,
<emphasis bold="true" pageId="0" pageNumber="1">stigma lobe</emphasis>
oblong, ca.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
long, folded, bifid at apex, surface slightly papillose.
<emphasis bold="true" pageId="0" pageNumber="1">Fruit</emphasis>
not observed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus isOtherTypeStatus="true">Additional specimens examined</typeStatus>
(
<typeStatus>paratypes</typeStatus>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Peninsular
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion country="Malaysia" name="Pahang">Pahang</collectingRegion>
, Bentong, Genting Highlands, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.137" unit="m" value="1137.0">1137 m</quantity>
elev.,
<materialsCitation accessionNumber="FRI91138" collectingDate="2022-04-07" collectorName="Siti-Munirah MY, Eddie C, Suhaida M" specimenCount="1">
<collectingDate value="2022-04-07">7 April 2022</collectingDate>
,
<collectorName>Siti-Munirah MY</collectorName>
,
<collectorName>Eddie C</collectorName>
&amp;
<collectorName>Suhaida M</collectorName>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91138">FRI91138</accessionNumber>
a (KEP, spirit collection, barcode no. SC12021)
</materialsCitation>
;
<materialsCitation accessionNumber="FRI91138" collectingDate="2022-04-07" collectorName="Siti-Munirah MY, Eddie C, Suhaida M" specimenCount="1">
<collectingDate value="2022-04-07">7 April 2022</collectingDate>
,
<collectorName>Siti-Munirah MY</collectorName>
,
<collectorName>Eddie C</collectorName>
&amp;
<collectorName>Suhaida M</collectorName>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91138">FRI91138</accessionNumber>
b (KEP, spirit collection, barcode no. SC12020)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Endemic to Peninsular Malaysia, Pahang. Currently known only from the type locality.</paragraph>
<caption doi="10.3897/phytokeys.211.89453.figure1" httpUri="https://binary.pensoft.net/fig/754973" pageId="0" pageNumber="1" start="Figure 1" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
with scale (all in centimetres (cm))
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
plants with mature (FRI91138c) and young flowers with long stems
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
mature flower with a very short stem (FRI91138b)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
stamens (FRI FRI91138a). Photos by Siti-Munirah MY.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.211.89453.figure2" httpUri="https://binary.pensoft.net/fig/754974" pageId="0" pageNumber="1" start="Figure 2" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
coralliform roots
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
full flower
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
claviform tip of mitre appendage
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
longitudinal section of floral tube, showing inner surface
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
outer tepal and inner tepal
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
side view of mitre
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
mitre viewed from above
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
six stamens, viewed from below
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
outer view of a stamen
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
inner view of two pendulous stamens
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
detail of the side lobe on the supraconnective apex (side lobe with truncate tips that are recurved inwards)
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
ovary with style, stigma and flower bud surrounded by bracts
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
stigmas. All from FRI FRI91138a (
<emphasis bold="true" pageId="0" pageNumber="1">A, D, I, J, K, L, M</emphasis>
), FRI91138b (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and FRI91138c (
<emphasis bold="true" pageId="0" pageNumber="1">B, C, E, F, G</emphasis>
). Photos by Siti-Munirah MY; images not to scale (see dimensions in description and Fig.
<figureCitation captionStart="Figure 1" captionStartId="F2" captionText="Figure 1. Thismia limkokthayi with scale (all in centimetres (cm )) A plants with mature (FRI 91138 c) and young flowers with long stems B mature flower with a very short stem (FRI 91138 b) C stamens (FRI FRI 91138 a). Photos by Siti-Munirah MY." figureDoi="10.3897/phytokeys.211.89453.figure1" httpUri="https://binary.pensoft.net/fig/754973" pageId="0" pageNumber="1">1</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
In lower montane forest and upper dipterocarp forest, on moist soil in shade, near an open space (hiking trail; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Thismia limkokthayi Siti-Munirah &amp; Eddie Chan A group photo with the T. limkokthayi (white arrow) in its habitat (from left; Angan A, Siti-Munirah MY, Eddie C, Ahmad Norsidar AH and Suhaida M) B-C different stages of anthesis of T. limkokthayi in its habitat (in situ) B two mature flowers from one individual C a fresh flower (on the left) and a new young flower starting to bloom (on the right). Photos by Siti-Munirah MY." figureDoi="10.3897/phytokeys.211.89453.figure3" httpUri="https://binary.pensoft.net/fig/754975" pageId="0" pageNumber="1">3A</figureCitation>
) and sloping area. This species occurs in a healthy undisturbed forest at an altitude of about 1137 m. The site is in a private forest and within a watershed. The forest area remains intact, apart from where it was affected by recent flooding and water surge (19 December 2021). The impact of this natural disaster completely reshaped part of the river area. However, this area is currently recovering. Fortunately, the only known population of
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
is located away from the riverbank, on the slope of the main trail. As a result of this discovery, the trail was moved to another part of the forest. The flowering period is from March to April.
</paragraph>
<caption doi="10.3897/phytokeys.211.89453.figure3" httpUri="https://binary.pensoft.net/fig/754975" pageId="0" pageNumber="1" start="Figure 3" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
Siti-Munirah &amp; Eddie Chan
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
group photo with the
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
(white arrow) in its habitat (from left; Angan A, Siti-Munirah MY, Eddie C, Ahmad Norsidar AH and Suhaida M)
<emphasis bold="true" pageId="0" pageNumber="1">B-C</emphasis>
different stages of anthesis of
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
in its habitat (in situ)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
two mature flowers from one individual
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
a fresh flower (on the left) and a new young flower starting to bloom (on the right). Photos by Siti-Munirah MY.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
was named in honour of Tan Sri Lim Kok Thay, Chairman of the Genting Group, who is closely involved in efforts to develop eco-tourism facilities and amenities supporting the preservation and sustainability of important biodiversity assets and sites in Genting Highlands.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="1">Preliminary conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">
Following the IUCN Red List Categories and Criteria (
<bibRefCitation author="Imhof, S" journalOrPublisher="Proceedings of the Linnean Society of London" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B7" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
), this species has been assessed as Critically Endangered (CR B2ab (iii); D) due to its small population and threats to its microhabitat. It is currently known from only one locality (the type locality) and is certainly a very rare species since only three individuals have been observed. Although the type locality is in a private forest, it may be exposed to future tourist activities in the Resorts World Genting Awana area. The habitat of the species is located on the original trail leading from the entrance of Clearwater Way to Chin Swee Caves Temple. However, since this is private land, the protection of this area from disturbance remains possible. As such, efforts must be made to locate this species in the surrounding area.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
species are characterised by their peculiar appearance and flower morphology, which are distinctive and simultaneously very diverse (
<bibRefCitation DOI="https://doi.org/10.1093/botlinnean/boaa017" author="Shepeleva, EA" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="287 - 315" refId="B10" refString="Shepeleva, EA, Schelkunov, MI, Hrones, M, Sochor, M, Dancak, M, Merckx, VSFT, Kikuchi, IABS, Chantanaorrapint, S, Suetsugu, K, Tsukaya, H, Mar, SS, Luu, HT, Li, H-Q, Logacheva, MD, Nuraliev, MS, 2020. Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287 - 315, DOI: https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." url="https://doi.org/10.1093/botlinnean/boaa017" volume="193" year="2020">Shepeleva et al. 2020</bibRefCitation>
). Most
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
species have various unique structural combinations. Amongst them, there are species belonging to the
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
group of which inner tepals are fused together in their upper part and form a roof-like or hat-like structure called a mitre. In Peninsular Malaysia, there are two groups of mitriform
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
: first including
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
species with mitre appendages (e.g.
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia clavigera</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Siti-Munirah" authorityYear="2018" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia kelantanensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigeroides" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigeroides">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigeroides</emphasis>
</taxonomicName>
) and second including
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
species without appendages (e.g.
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. latiffiana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="latiffiana">
<emphasis italics="true" pageId="0" pageNumber="1">T. latiffiana</emphasis>
</taxonomicName>
Siti-Munirah &amp; Dome and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sitimeriamiae" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sitimeriamiae">
<emphasis italics="true" pageId="0" pageNumber="1">T. sitimeriamiae</emphasis>
</taxonomicName>
Siti-Munirah, Dome &amp; Thorogood).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
is easily recognised by a combination of the following characteristics: coralliform roots, erect outer perianth lobes, mitriform inner perianth lobes with three erect, slender, claviform appendages that form a mitre fovea with their flattened bases in the centre, absence of the annulus, presence of longitudinal ribs on the inner side of the floral tube (or absence of the reticulate inner side), the apex of stamens with a central lobe (prolongation of the rib) and two lateral lobes (the tips of each are recurved) and black-purple stigma.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on the infrageneric classification of
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.319.3.2" author="Kumar, P" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" pagination="225 - 240" refId="B9" refString="Kumar, P, Gale, SW, Li, JH, Bouamanivong, S, Fischer, GA, 2017. Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification. Phytotaxa 319 (3): 225 - 240, DOI: https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification." url="https://doi.org/10.11646/phytotaxa.319.3.2" volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia</emphasis>
</taxonomicName>
species in the
<taxonomicName authority="Geomitra" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia (Thismia) subsp. section" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Geomitra" subGenus="Thismia">Thismia subgenus Thismia section Geomitra</taxonomicName>
(Becc.) Kumar &amp; S.W. Gale, mainly based on the presence of inner tepals forming a mitre, each with filiform appendage and central mitral appendages that are free from each other, with the outer tepals always being short (less than 2 mm long). Two species were included in section
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Geomitra">Thismia Geomitra</taxonomicName>
:
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
(Becc.) F. Muell. and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. betung-kerihunensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="betung-kerihunensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. betung-kerihunensis</emphasis>
</taxonomicName>
Tsukaya &amp; H. Okada.
</paragraph>
<paragraph pageId="0" pageNumber="1">
To date,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
has not been included in any DNA-based analysis. According to the classification proposed by
<bibRefCitation DOI="https://doi.org/10.1093/botlinnean/boaa017" author="Shepeleva, EA" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="287 - 315" refId="B10" refString="Shepeleva, EA, Schelkunov, MI, Hrones, M, Sochor, M, Dancak, M, Merckx, VSFT, Kikuchi, IABS, Chantanaorrapint, S, Suetsugu, K, Tsukaya, H, Mar, SS, Luu, HT, Li, H-Q, Logacheva, MD, Nuraliev, MS, 2020. Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287 - 315, DOI: https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." url="https://doi.org/10.1093/botlinnean/boaa017" volume="193" year="2020">Shepeleva et al. (2020)</bibRefCitation>
,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
is possibly related to species in clade 3 because it is characterised by coralliform roots, inner tepals fused into a mitre and free mitre appendages extending from a central point. Within clade 3 (i.e. section
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Sarcosiphon">Thismia Sarcosiphon</taxonomicName>
sensu
<bibRefCitation author="Jonker, FP" editor="van Steenis, CGGJ" journalOrPublisher="Noordhoff-Kolf, Leiden, series 1 (4)" pageId="0" pageNumber="1" pagination="13 - 26" refId="B8" refString="Jonker, FP, 1948. Burmanniaceae. In: van Steenis, CGGJ, Ed., Flora Malesiana. Noordhoff-Kolf, Leiden, series 1(4): 13 - 26" title="Burmanniaceae." volumeTitle="Flora Malesiana." year="1948">Jonker 1948</bibRefCitation>
), there are two morphologically distinct groups characterised by the presence of a prominent central rib along the inner side of the connective and the absence of an annulus (
<bibRefCitation DOI="https://doi.org/10.3372/wi.50.50106" author="Dancak, M" journalOrPublisher="Willdenowia" pageId="0" pageNumber="1" pagination="65 - 76" refId="B4" refString="Dancak, M, Hrones, M, Sochor, M, 2020. Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo. Willdenowia 50 (1): 65 - 76, DOI: https://doi.org/10.3372/wi.50.50106" title="Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo." url="https://doi.org/10.3372/wi.50.50106" volume="50" year="2020">
<normalizedToken originalValue="Dančák">Dancak</normalizedToken>
et al. 2020
</bibRefCitation>
). They can be distinguished by the presence of distinct appendages at the top of the mitre (
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
group) or their absence (
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. goodii" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="goodii">
<emphasis italics="true" pageId="0" pageNumber="1">T. goodii</emphasis>
</taxonomicName>
group). However, their exact relationship cannot be clarified until more species in both groups are sequenced.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Currently, there are five species known in the
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
group:
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. betung-kerihunensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="betung-kerihunensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. betung-kerihunensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigeroides" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigeroides">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigeroides</emphasis>
</taxonomicName>
Chantanaorr &amp; Seelanan,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. kelantanensis</emphasis>
</taxonomicName>
Siti-Munirah and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sumatrana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sumatrana">
<emphasis italics="true" pageId="0" pageNumber="1">T. sumatrana</emphasis>
</taxonomicName>
Suetsugu &amp; Tsukaya (
<bibRefCitation DOI="https://doi.org/10.1600/036364412X616639" author="Tsukaya, H" journalOrPublisher="Systematic Botany" pageId="0" pageNumber="1" pagination="53 - 57" refId="B15" refString="Tsukaya, H, Okada, H, 2012. A new species of Thismia (Thismiaceae) from West Kalimantan, Borneo. Systematic Botany 37 (1): 53 - 57, DOI: https://doi.org/10.1600/036364412X616639" title="A new species of Thismia (Thismiaceae) from West Kalimantan, Borneo." url="https://doi.org/10.1600/036364412X616639" volume="37" year="2012">Tsukaya and Okada 2012</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/s12225-018-9767-0" author="Siti-Munirah, MY" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" refId="B11" refString="Siti-Munirah, MY, 2018. Thismia kelantanensis (Thismiaceae), a new species from Kelantan, Peninsular Malaysia. Kew Bulletin 73(3): e42. https://doi.org/10.1007/s12225-018-9767-0" title="Thismia kelantanensis (Thismiaceae), a new species from Kelantan, Peninsular Malaysia. Kew Bulletin 73 (3): e 42." url="https://doi.org/10.1007/s12225-018-9767-0" year="2018">Siti-Munirah 2018</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.113.29103" author="Suetsugu, K" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="59 - 67" refId="B14" refString="Suetsugu, K, Tsukaya, H, Nurainas, N, Okada, H, 2018. Thismia sumatrana (Thismiaceae), a new species from West Sumatra, Indonesia, with discussions on the taxonomic identity of Thismia clavigera. PhytoKeys 113: 59 - 67, DOI: https://doi.org/10.3897/phytokeys.113.29103" title="Thismia sumatrana (Thismiaceae), a new species from West Sumatra, Indonesia, with discussions on the taxonomic identity of Thismia clavigera." url="https://doi.org/10.3897/phytokeys.113.29103" volume="113" year="2018">Suetsugu et al. 2018</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1600/036364421X16128061189468" author="Chantanaorrapint, S" journalOrPublisher="Systematic Botany" pageId="0" pageNumber="1" pagination="18 - 23" refId="B2" refString="Chantanaorrapint, S, Seelanan, T, 2021. Thismia clavigeroides (Thismiaceae), a new mycoheterotrophic species from Thailand. Systematic Botany 46 (1): 18 - 23, DOI: https://doi.org/10.1600/036364421X16128061189468" title="Thismia clavigeroides (Thismiaceae), a new mycoheterotrophic species from Thailand." url="https://doi.org/10.1600/036364421X16128061189468" volume="46" year="2021">Chantanaorrapint and Seelanan 2021</bibRefCitation>
).
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
clearly differs from all these species by the absence of transverse bars on the inner surface of the floral tube. Notably,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
clearly resembles
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. kelantanensis</emphasis>
</taxonomicName>
, an endemic species from Kelantan (Malay Peninsula), which also has three slender, claviform appendages at the tip of the mitre and erect outer tepals. However,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. kelantanensis</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
by its bluish filaments and stamens, as well as the six-part cap on the mitre. Additionally, it also resembles
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigeroides" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigeroides">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigeroides</emphasis>
</taxonomicName>
Chantanaorr &amp; Seelanan from Thailand and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sumatrana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sumatrana">
<emphasis italics="true" pageId="0" pageNumber="1">T. sumatrana</emphasis>
</taxonomicName>
Suetsugu &amp; Tsukaya from Sumatra in its general appearance. However,
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
differs from both of these species by the curved outer lobes on the supraconnective apex. Moreover, the connective appendage of
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sumatrana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sumatrana">
<emphasis italics="true" pageId="0" pageNumber="1">T. sumatrana</emphasis>
</taxonomicName>
is hairy, while it is glabrous in
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
and the flower colour of
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigeroides" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigeroides">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigeroides</emphasis>
</taxonomicName>
(including the stamens) is almost whitish, while that of
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
is blackish-brown. A comparison of morphological characteristics between
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
and other related species is presented in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological differences between Thismia limkokthayi and related species. The characters of previously-described species are taken from the protologues and recent publications on T. betung-kerihunensis (Tsukaya and Okada 2012), T. clavigera (Chantanaorrapint and Chantanaorrapint 2009), T. kelantanensis (Siti-Munirah 2018) and T. sumatrana (Suetsugu et al. 2018)." httpUri="http://table.plazi.org/id/3A45A696C69340C8179B303244C62E77" pageId="0" pageNumber="1" tableUuid="3A45A696C69340C8179B303244C62E77">1</tableCitation>
.
</paragraph>
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<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
Morphological differences between
<taxonomicName authorityName="Siti-Munirah &amp; E. Chan" authorityYear="2022" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">Thismia limkokthayi</emphasis>
</taxonomicName>
and related species. The characters of previously-described species are taken from the protologues and recent publications on
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. betung-kerihunensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="betung-kerihunensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. betung-kerihunensis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1600/036364412X616639" author="Tsukaya, H" journalOrPublisher="Systematic Botany" pageId="0" pageNumber="1" pagination="53 - 57" refId="B15" refString="Tsukaya, H, Okada, H, 2012. A new species of Thismia (Thismiaceae) from West Kalimantan, Borneo. Systematic Botany 37 (1): 53 - 57, DOI: https://doi.org/10.1600/036364412X616639" title="A new species of Thismia (Thismiaceae) from West Kalimantan, Borneo." url="https://doi.org/10.1600/036364412X616639" volume="37" year="2012">Tsukaya and Okada 2012</bibRefCitation>
),
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
(
<bibRefCitation author="Chantanaorrapint, S" journalOrPublisher="Thai Forest Bulletin (Botany)" pageId="0" pageNumber="1" pagination="27 - 31" refId="B1" refString="Chantanaorrapint, S, Chantanaorrapint, A, 2009. Thismia clavigera (Thismiaceae), a new record for Thailand. Thai Forest Bulletin (Botany) 37: 27 - 31" title="Thismia clavigera (Thismiaceae), a new record for Thailand." volume="37" year="2009">Chantanaorrapint and Chantanaorrapint 2009</bibRefCitation>
),
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. kelantanensis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1007/s12225-018-9767-0" author="Siti-Munirah, MY" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" refId="B11" refString="Siti-Munirah, MY, 2018. Thismia kelantanensis (Thismiaceae), a new species from Kelantan, Peninsular Malaysia. Kew Bulletin 73(3): e42. https://doi.org/10.1007/s12225-018-9767-0" title="Thismia kelantanensis (Thismiaceae), a new species from Kelantan, Peninsular Malaysia. Kew Bulletin 73 (3): e 42." url="https://doi.org/10.1007/s12225-018-9767-0" year="2018">Siti-Munirah 2018</bibRefCitation>
) and
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sumatrana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sumatrana">
<emphasis italics="true" pageId="0" pageNumber="1">T. sumatrana</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.113.29103" author="Suetsugu, K" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="59 - 67" refId="B14" refString="Suetsugu, K, Tsukaya, H, Nurainas, N, Okada, H, 2018. Thismia sumatrana (Thismiaceae), a new species from West Sumatra, Indonesia, with discussions on the taxonomic identity of Thismia clavigera. PhytoKeys 113: 59 - 67, DOI: https://doi.org/10.3897/phytokeys.113.29103" title="Thismia sumatrana (Thismiaceae), a new species from West Sumatra, Indonesia, with discussions on the taxonomic identity of Thismia clavigera." url="https://doi.org/10.3897/phytokeys.113.29103" volume="113" year="2018">Suetsugu et al. 2018</bibRefCitation>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. limkokthayi" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="limkokthayi">
<emphasis italics="true" pageId="0" pageNumber="1">T. limkokthayi</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. betung-kerihunensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="betung-kerihunensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. betung-kerihunensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigera" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigera">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigera</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. clavigeroides" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="clavigeroides">
<emphasis italics="true" pageId="0" pageNumber="1">T. clavigeroides</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. kelantanensis" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kelantanensis">
<emphasis italics="true" pageId="0" pageNumber="1">T. kelantanensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Liliopsida" family="Burmanniaceae" kingdom="Plantae" lsidName="T. sumatrana" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sumatrana">
<emphasis italics="true" pageId="0" pageNumber="1">T. sumatrana</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Colour of floral tube</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Black-brownish/brown-blackish</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">White with indigo and brown to pale brown with purple-dark blue</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">White-orangish</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Whitish</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pale to bright and dark blue-purplish translucent,</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Colour of mitre</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Black-brownish/brown-blackish</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Blue-green</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Yellowish-orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pale brown or grey</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Yellow to bright orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Colour of inner tepal lobes</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Brown-blackish</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Blue-green</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pale brown or grey</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Bright orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Outer tepal</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Erect</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Erect</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Erect</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Reflexed</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Erect</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Reflexed</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Colour of filament</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Orange and white</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Blue-green</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">White</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Bright blue</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Colour of appendages</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Dark brown to pale orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pale blue tinged with orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pale brown or grey below, blue-green at tip</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Orange</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Unknown</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Fovea</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Apex of supraconnective</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">One central lobe (extension of the rib) and two smaller side lobes with tips recurved inwards/truncate, glabrous</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">One central lobe (extension of the rib) and two smaller side lobes, hairy</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Acute</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">One central lobe (extension of the rib) and two smaller side lobes</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Acuminate</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Acute, hairy</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Presence of transverse bars</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Present</td>
</tr>
</table>
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Type status:
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. Occurrence: sex:
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<paragraph id="662A6428A47C276F1C030860DC796267" pageId="0" pageNumber="5447">Type locality: [West Papua], Arfak Mts, Angi Lakes, 6000 ft.</paragraph>
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<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
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<mods:name type="personal">
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</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>1758</mods:dateIssued>
<mods:publisher>Laurentius Salvius</mods:publisher>
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<mods:placeTerm>Stockholm</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:location>
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</mods:location>
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<mods:identifier type="DOI">https://doi.org/10.5962/bhl.title.542</mods:identifier>
<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3917846" ID-GBIF-Taxon="164813250" ID-Zenodo-Dep="3917846" LSID="urn:lsid:zoobank.org:act:6DC219C0-C547-4613-BEF7-E1F08227D662" httpUri="http://treatment.plazi.org/id/0C02E8E243F9FF7811CB706DFCB04C7F" lastPageNumber="561" pageNumber="560" sourcePageUrl="https://www.biodiversitylibrary.org/page/727471">
<subSubSection type="nomenclature">
<paragraph pageNumber="560">
<taxonomicName LSID="urn:lsid:zoobank.org:act:6DC219C0-C547-4613-BEF7-E1F08227D662" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Eulophidae" genus="Ichneumon" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="camelus" status="spec. nov." zbkClass="Insecta" zbkKingdom="Animalia" zbkOrder="Hymenoptera">Ichneumon camelus</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="560">I. abdomine mucronato atro lateribus albo maculato, thorace laevi.</paragraph>
<paragraph pageNumber="560">
<emphasis italics="true">Uddm. diss.</emphasis>
82.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="560">
<emphasis italics="true">Habitat in</emphasis>
Ligno
<emphasis italics="true">putrido.</emphasis>
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="560">
<emphasis italics="true">Antennae nigrae</emphasis>
;
<emphasis italics="true">praecedentium</emphasis>
1, 2, 3
<emphasis italics="true">flavae.</emphasis>
</paragraph>
<footnote>
<paragraph pageNumber="560">
Ichneumonis
<emphasis italics="true">Larvae plerumque intra alia insecta,</emphasis>
&amp;
<emphasis italics="true">imprimis intra Larvas Lepidopterorum</emphasis>
aliorumque vivunt, quas intus consumunt.
</paragraph>
</footnote>
<paragraph pageNumber="561">
<pageBreakToken pageNumber="561">*</pageBreakToken>
*
<emphasis italics="true">Scutello albido</emphasis>
;
<emphasis italics="true">Antennis fascia albida annulatis.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,188 @@
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<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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</mods:role>
<mods:namePart>Beaver, Roger A.</mods:namePart>
<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>983</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
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<treatment ID-GBIF-Taxon="168754550" LSID="urn:lsid:zoobank.org:act:F8082757-4BED-4036-BE76-64A89A26FB2D" httpUri="http://treatment.plazi.org/id/0C02EAAB62BA53D7813F4B613913D1FB" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/F8082757-4BED-4036-BE76-64A89A26FB2D" authority="Smith &amp; Beaver &amp; Cognato, 2020" authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Microperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microperus sagmatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sagmatus" status="sp. nov.">Microperus sagmatus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 70" captionStartId="F70" captionText="Figure 70. Dorsal, lateral and declivital view of Microperus sagmatus holotype, 1.75 - 1.95 mm (A, B, E), and M. undulatus, 2.0 - 2.1 mm (C, D, F)." figureDoi="10.3897/zookeys.983.52630.figure70" httpUri="https://binary.pensoft.net/fig/470870" pageId="0" pageNumber="1">Fig. 70A, B, E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, female, Thailand: Suranthani [= Surat Thani], durian or[chard], 01.xii.[20]10, Wisut Sittichaya, EToH-trap (MSUC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
, female, Malaysia: Penang, B[atu] Ferringhi, 6.i-1.ii.1981, T. Palm (RABC, 1); Thailand: as holotype (MSUC, 1); Prachuab Khiri Khan: Kui Buri N.P., 27.iii.2008, S. Stevens et al., ex
<normalizedToken originalValue="krachid">'krachid'</normalizedToken>
(NHMUK, 2; MSUC, 2; QSBG, 1; RABC, 2); Songkhla, Ratthapum distr., ex durian branch, 4.xi.2008, W. Sittichaya (RABC, 1; QSBG 1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
1.75-1.95 mm long (mean = 1.83 mm; n = 5); 2.69-2.79
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. This species is distinguished by the elytral disc shallowly transversely impressed with a saddle-like depression; elytral interstriae costate with strong interstrial spines posterior to the saddle; and declivity steep, slightly flattened.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Microperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microperus sagmatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sagmatus">
<emphasis italics="true" pageId="0" pageNumber="1">Microperus sagmatus</emphasis>
</taxonomicName>
closely resembles
<taxonomicName lsidName="M. undulatus" pageId="0" pageNumber="1" rank="species" species="undulatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. undulatus</emphasis>
</taxonomicName>
but is distinguished by the discal impression deeper, antero-posteriorly narrower, with steeper anterior and posterior slopes, strial punctures on impression with rounded granules, interstrial spines on disc behind saddle stronger, and backwardly hooked, not pointing dorsally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" baseAuthorityName="Schedl" baseAuthorityYear="1975" class="Insecta" family="Curculionidae" genus="Microperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microperus cruralis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis italics="true" pageId="0" pageNumber="1">Microperus cruralis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="M. nugax" pageId="0" pageNumber="1" rank="species" species="nugax">
<emphasis italics="true" pageId="0" pageNumber="1">M. nugax</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="M. undulatus" pageId="0" pageNumber="1" rank="species" species="undulatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. undulatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(female).</emphasis>
1.75-1.95 mm long (mean = 1.83 mm; n = 5); 2.69-2.79
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Body ferruginous. Legs and antennae light brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, shagreened, punctate; punctures large, shallow, sparse, setose; punctures bearing a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, slightly impressed. Antennal scape short and thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 2; segment 1 corneous, feebly convex on anterior face, occupying basal 1/3, nearly covering posterior face; segment 2 narrow, soft; segment 1 present on posterior face.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
: 1.05
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. In dorsal view subquadrate and parallel-sided, type 3, sides parallel in basal 2/3, weakly rounded anteriorly with prominent anterolateral corners; anterior margin without a row of serrations. In lateral view tall, type 2, disc flat, summit at midpoint. Anterior slope with densely spaced, broad asperities, becoming lower and more strongly transverse towards summit. Disc shagreened, alutaceous, impunctate, glabrous, some moderately long hair-like setae at margins. Lateral margins obliquely costate. Base weakly bisinuate, posterior angles acutely rounded, almost subquadrate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytra</emphasis>
: 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum. Scutellum minute, convex, slightly raised above elytral surface. Elytral mycangium present as a dispersed median setal tuft of setae extending along elytral base. Elytral base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 4/5, then narrowly rounded to apex. Disc shiny, a moderately deep transverse saddle-like impression at midpoint, striae and interstriae flat, nearly glabrous anteriad of depression, strial punctures on impression with rounded granules, interstriae costate with strong backwardly hooked spines posteriad of depression, spines setose with long hair-like setae. Declivity occupying 1/3 of elytral length, shagreened, dull, steeply rounded, face slightly flattened; striae flat, parallel, punctate, punctures very large, shallow subcontiguous, setose, setae recumbent, as long a puncture; interstriae irregularly denticulate along their lengths, denticles small, irregularly spaced and sized, each bearing a long, erect hair-like seta, interstriae 1 and 3, weakly convex, 2 and 4 flat. Posterolateral margin carinate to interstriae 7.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
: procoxae contiguous; prosternal coxal piece tall, conical. Protibiae obliquely triangular, broadest at apical 1/3; posterior face smooth; apical 1/3 of outer margin with seven moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae flattened; outer margin evenly rounded with nine and eight moderate to large socketed denticles, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
G.
<emphasis italics="true" pageId="0" pageNumber="1">sagma</emphasis>
= pack-saddle. In reference to the shape of the elytra. An adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">West Malaysia, Thailand.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="host">
<paragraph pageId="0" pageNumber="1">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="1">
Recorded from
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Durio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Durio zibethinus" order="Malvales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="zibethinus">
<emphasis italics="true" pageId="0" pageNumber="1">Durio zibethinus</emphasis>
</taxonomicName>
(durian) (
<taxonomicName family="Malvaceae" lsidName="" pageId="0" pageNumber="1" rank="family">Malvaceae</taxonomicName>
), and an undetermined tree,
<normalizedToken originalValue="krachid">'krachid'</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specimens from Thailand were included under
<taxonomicName lsidName="M. undulatus" pageId="0" pageNumber="1" rank="species" species="undulatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. undulatus</emphasis>
</taxonomicName>
by
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3875.1.1" author="Beaver, RA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 82" refId="B26" refString="Beaver, RA, Sittichaya, W, Liu, L-Y, 2014. A synopsis of the scolytine ambrosia beetles of Thailand (Coleoptera: Curculionidae: Scolytinae). Zootaxa 3875: 1 - 82, DOI: https://doi.org/10.11646/zootaxa.3875.1.1" title="A synopsis of the scolytine ambrosia beetles of Thailand (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.11646/zootaxa.3875.1.1" volume="3875" year="2014">Beaver et al. (2014)</bibRefCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.983.52630.figure70" httpUri="https://binary.pensoft.net/fig/470870" pageId="0" pageNumber="1" start="Figure 70" startId="F70">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 70.</emphasis>
Dorsal, lateral and declivital view of
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Microperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microperus sagmatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sagmatus">
<emphasis italics="true" pageId="0" pageNumber="1">Microperus sagmatus</emphasis>
</taxonomicName>
holotype, 1.75-1.95 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, B, E</emphasis>
), and
<taxonomicName lsidName="M. undulatus" pageId="0" pageNumber="1" rank="species" species="undulatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. undulatus</emphasis>
</taxonomicName>
, 2.0-2.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">C, D, F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
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@ -0,0 +1,864 @@
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<mods:title id="539D2BC85B31DD3A7BC5EBF5D14811C6">A new species of the genus Amolops (Amphibia: Ranidae) and the first national record of Amolops vitreus from China</mods:title>
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<mods:namePart id="71072EFEB821DFDEF6BE91D9BDCE5FB5">Yu, Zhong-Bin</mods:namePart>
<mods:affiliation id="BD069D982A6841A95D0392BB7875E855">State Key Laboratory of Genetic Resources and Evolution &amp; Yunnan key laboratory of biodiversity and ecological conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
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</mods:role>
<mods:namePart id="204D4A9CCFAC2C7D14ABDBE48759C657">Lu, Chen-Qi</mods:namePart>
<mods:affiliation id="FE323B02B5D90BF7234FD0B64A5139B1">State Key Laboratory of Genetic Resources and Evolution &amp; Yunnan key laboratory of biodiversity and ecological conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Kunming College of Life Science, University of the Chinese Academy of Sciences, Kunming, Yunnan 650204, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="14D48CC2043A63356D102AD051E296B4">Zhang, Yin-Peng</mods:namePart>
<mods:nameIdentifier id="47EB6F9F7548F89ECFEB4D4CA931DC48" type="ORCID">https://orcid.org/0000-0001-6857-7445</mods:nameIdentifier>
<mods:affiliation id="203F05151C5D64881CB9D6790613A6AA">Department of Ecology and Evolutionary Biology, University of Arizona, Tucson 85721, USA</mods:affiliation>
</mods:name>
<mods:name id="46C0F15E75A791547362C3DDA2205958" type="personal">
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<mods:namePart id="ADCE34203FFD67678025D8A195A6C1A2">Dong, Wen-Jie</mods:namePart>
<mods:affiliation id="717A45E9AED377247454702CFFFAA02E">State Key Laboratory of Genetic Resources and Evolution &amp; Yunnan key laboratory of biodiversity and ecological conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Kunming College of Life Science, University of the Chinese Academy of Sciences, Kunming, Yunnan 650204, China</mods:affiliation>
</mods:name>
<mods:name id="494101B61B8F6DB4E6D1429657A1847F" type="personal">
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</mods:role>
<mods:namePart id="C628B8E9675ED2175B1FD3FFE01AA2EF">Liu, Xiao-Long</mods:namePart>
<mods:affiliation id="38C11EE7E6387AC830A7A66595CA649A">Key Laboratory of Freshwater Fish Reproduction and Development, Ministry of Education, Southwest University, Chongqing 400715, China</mods:affiliation>
</mods:name>
<mods:name id="FE599BBD433CF7CA04CB5BF8E4B6D1C5" type="personal">
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<mods:roleTerm id="FE537D2864089E1C630F2AA21B4463AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD8CCC916B9D3094A28492C813207848">Kilunda, Felista Kasyoka</mods:namePart>
<mods:affiliation id="5720C61253F8F9A839F4187DF88DC786">State Key Laboratory of Genetic Resources and Evolution &amp; Yunnan key laboratory of biodiversity and ecological conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Kunming College of Life Science, University of the Chinese Academy of Sciences, Kunming, Yunnan 650204, China</mods:affiliation>
</mods:name>
<mods:name id="9771273162431BA10D40CBE4FDEBED03" type="personal">
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</mods:role>
<mods:namePart id="03FCD9498F5F11CE74294C95FD975034">Xiong, Yun</mods:namePart>
<mods:affiliation id="555ACAF1B20F975AF53530273DF1E350">Gongshan Bureau of Gaoligongshan National Nature Reserve, Nujiang, Yunnan 673500, China</mods:affiliation>
</mods:name>
<mods:name id="DB5B9647E275CDB7A61BB9D6D147C884" type="personal">
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</mods:role>
<mods:namePart id="192EC7B6EA99574891C9063F1569318B">Jiang, Yun-Fang</mods:namePart>
<mods:affiliation id="A32531CD2BABD0D3C4827B85EEBFD6C7">Lushui Bureau of Gaoligongshan National Nature Reserve, Nujiang, Yunnan 673299, China</mods:affiliation>
</mods:name>
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<mods:roleTerm id="101A55DA58177554F941C42D8825477B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BCD6710B5095460F666FCB047E378EE">Ouyang, Hong</mods:namePart>
<mods:affiliation id="27B7291E0104D8284681C2907EB3FC02">Lushui Bureau of Gaoligongshan National Nature Reserve, Nujiang, Yunnan 673299, China</mods:affiliation>
</mods:name>
<mods:name id="962E65DD3F98BCB82C5BE0A135B6D947" type="personal">
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<mods:roleTerm id="65447B8AC8B06CF4670FBA8678BC1BCD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="55D0D39980EEADC09AFA49C391063258">Fu, Zhong-Xiong</mods:namePart>
<mods:affiliation id="A6F5227BEA6452A073E3A4562647E72C">Yunnan Senye Biotechnology Co., Ltd, Xishuangbanna, Yunnan 666100, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5E3AB5C106ECB88F8E79B08EDF41468A">He, Yun-Biao</mods:namePart>
<mods:affiliation id="24E9619C4E734B0F842425B50B2D7B48">Fugong Bureau of Gaoligongshan National Nature Reserve, Nujiang, Yunnan 673400, China</mods:affiliation>
</mods:name>
<mods:name id="39EB7D59EAD4ABD44CF2BF6D0511FF72" type="personal">
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</mods:role>
<mods:namePart id="EEF01D7DF45044536F5EBBE45D78F358">Yuan, Zhi-Yong</mods:namePart>
<mods:nameIdentifier id="BF4DE540F846FD9374FF408C8098AA28" type="ORCID">https://orcid.org/0000-0001-5991-3021</mods:nameIdentifier>
<mods:affiliation id="57E78724BCAF45687282BC2FAAF7F0CC">Key Laboratory of Freshwater Fish Reproduction and Development, Ministry of Education, Southwest University, Chongqing 400715, China</mods:affiliation>
<mods:nameIdentifier id="721F1548210D34B2DE7B0CE065AEA799" type="email">yuanzhiyongkiz@126.com</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="402049931C7C09CFB0531AC091B5C453">Che, Jing</mods:namePart>
<mods:affiliation id="D52BD47239161A099F0FC9891620E66D">State Key Laboratory of Genetic Resources and Evolution &amp; Yunnan key laboratory of biodiversity and ecological conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
<mods:nameIdentifier id="7CFBB6726B6729B51C801CB6235F0FF1" type="email">chej@mail.kiz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1B43CCD65323D2B5030F94E66002B673">text</mods:typeOfResource>
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<mods:titleInfo id="E6BA0DF465024A88B54D5E53105A0392">
<mods:title id="2603D4B390A30F7133B17B02669CC2F4">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="F41B9CB892AFA7598240102A373C2089">
<mods:date id="5A82F4BBCD6D078CA53DD30978783453">2024</mods:date>
<mods:detail id="5B4BEE4B2AABFE170668F13261BB9CB4" type="pubDate">
<mods:number id="73D42D631C250A987C25F155F49FFDBE">2024-03-14</mods:number>
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<mods:detail id="E61124F4B2E2E97FE5AA607D0689D285" type="volume">
<mods:number id="33E212B1F9BF5CB4BDD469B42A3B9FFD">74</mods:number>
</mods:detail>
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<mods:start id="706F152DFF8150F159068A7820074BD0">343</mods:start>
<mods:end id="3CBFCC17F70BDC16D08E5A3CD0389F81">357</mods:end>
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<mods:url id="81A21FFEBDB27151B3D1D2025D4FF1FB">http://dx.doi.org/10.3897/vz.74.e108013</mods:url>
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<mods:classification id="ACBE8B952F893AE97DB4662CB4A505CE">journal article</mods:classification>
<mods:identifier id="7FA5F45794A61A617269E5F1B0FBF613" type="DOI">http://dx.doi.org/10.3897/vz.74.e108013</mods:identifier>
<mods:identifier id="D8D6668F0FADF8C661F4E50AF6DC4DF0" type="Pensoft-Pub">2625-8498-74-343</mods:identifier>
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<treatment id="0C02F36B5F6E52C6A80A97144CF19DA1" LSID="urn:lsid:zoobank.org:act:5B4E8AC7-37AD-4192-83E3-A05E49393B7B" httpUri="http://treatment.plazi.org/id/0C02F36B5F6E52C6A80A97144CF19DA1" lastPageNumber="343" pageId="0" pageNumber="343">
<subSubSection id="8BC0B92CB3F5AE44C71AA69E87423DE6" pageId="0" pageNumber="343" type="nomenclature">
<paragraph id="B19D48049D7897AACA3286CD2B1A4424" pageId="0" pageNumber="343">
<taxonomicName id="BEF45E6C57C2A7B0F829699EE8E71AA2" LSID="https://zoobank.org/5B4E8AC7-37AD-4192-83E3-A05E49393B7B" authority="Wu, Yu, Lu, Yuan &amp; Che" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi" status="sp. nov.">Amolops yangi Wu, Yu, Lu, Yuan &amp; Che</taxonomicName>
<taxonomicNameLabel id="F3C177F77EC577C63C0EFDDD4E5C58BE" pageId="0" pageNumber="343">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="50C6E5B28AAE0FE3B0725DF08092F6D3" pageId="0" pageNumber="343" type="holotype">
<paragraph id="056BE9A341855D1F482775086CB39786" pageId="0" pageNumber="343">Holotype.</paragraph>
<paragraph id="AE70FEB1CDB04818437E0B3990E0A003" pageId="0" pageNumber="343">
Adult female (KIZ 038643), from Ega, Lushui, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (
<geoCoordinate id="59B211DB8FEE74E0F348E1016CBB5A13" degrees="26.43744" direction="north" orientation="latitude" precision="1" value="26.43744">26.43744°N</geoCoordinate>
,
<geoCoordinate id="FE92EA6D7676666AF1740C0327E5E090" degrees="98.75044" direction="east" orientation="longitude" precision="1" value="98.75044">98.75044°E</geoCoordinate>
; elevation 3496 m a.s.l.), collected by Zhong-Bin Yu, Dong An, Tian-En Chen on 07, August, 2021.
</paragraph>
</subSubSection>
<subSubSection id="1015E23535B19C2D20295B18910C05D1" pageId="0" pageNumber="343" type="paratypes">
<paragraph id="290521BA4572B12D939FB35D0550A540" pageId="0" pageNumber="343">Paratypes.</paragraph>
<paragraph id="3351CACC0BBC4262553DCE1E25F9978B" pageId="0" pageNumber="343">
One adult male KIZ 038645 from Ega, Lushui, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (
<geoCoordinate id="C66EE8554DE4989FDED1EB4C04465B3D" degrees="26.44924" direction="north" orientation="latitude" precision="1" value="26.44924">26.44924°N</geoCoordinate>
,
<geoCoordinate id="510294E549F738FC623A9085D90AEFAA" degrees="98.76762" direction="east" orientation="longitude" precision="1" value="98.76762">98.76762°E</geoCoordinate>
; elevation 2915 m a.s.l.), collected by Zhong-Bin Yu, Dong An, Tian-En Chen on 07, August, 2021; one adult male KIZ 050788 from Yaping road, Lumadeng Township, Fugong county, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (
<geoCoordinate id="315127331013A16E0C733B83C9E5407A" degrees="27.15890" direction="north" orientation="latitude" precision="1" value="27.1589">27.15890°N</geoCoordinate>
,
<geoCoordinate id="90D0AC336BCB451B6F333FE828C931EB" degrees="98.79712" direction="east" orientation="longitude" precision="1" value="98.79712">98.79712°E</geoCoordinate>
; elevation 2451 m a.s.l.), collected by Zhong-Bin Yu, Dong An on 05, August, 2022.
</paragraph>
</subSubSection>
<subSubSection id="347EAD9753FF0E7C6BF9B3BD8E7E3D3A" pageId="0" pageNumber="343" type="etymology">
<paragraph id="A51EF59FD03D4BC4D7D09F800B8B9DC8" pageId="0" pageNumber="343">Etymology.</paragraph>
<paragraph id="9D2E5FAF883E66D972AA00E1E8FD28B1" pageId="0" pageNumber="343">
The specific epithet &quot;
<emphasis id="DA0A2453B8B865EA2D3DE34E1A3501ED" italics="true" pageId="0" pageNumber="343">yangi</emphasis>
&quot; is a patronymic noun in the genitive singular; derived from the name of Prof. Da-Tong Yang of the Kunming Institute of Zoology, CAS, China. We acknowledge his great contributions to the herpetological research in southwestern China. We suggest the Chinese formal name as
<normalizedToken id="BEAF36F04D168980DAF5D161F66DAADF" originalValue="“杨氏湍蛙”">&quot;杨氏湍蛙&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="BB48DB97C71D66A2CF157BD590B6155C" pageId="0" pageNumber="343" type="diagnosis">
<paragraph id="033208311F35E1F88D9B16E5D5269048" pageId="0" pageNumber="343">Diagnosis.</paragraph>
<paragraph id="6E07542BDF5FABEAF6B26F20776B3507" pageId="0" pageNumber="343">
<taxonomicName id="9557CEA4A255C472934D23378C86A2DE" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="7D1DB28BDA70903CFC211E5F19F73041" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="3CEE230CF5688F5A92D72C3ABBDBAD03" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
is assigned to the genus
<taxonomicName id="4D5541BBDC0A39E81ABA7CCAFAA4D073" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="genus">
<emphasis id="14C4E029E6343F7F15351AF4D3217C7B" italics="true" pageId="0" pageNumber="343">Amolops</emphasis>
</taxonomicName>
based on molecular phylogenetic analyses and can be distinguished from its congeners by a combination of the following characters: (1) medium body size (SVL 46.3-51.8 mm in males and at least 51.5 mm in female); (2) vomerine teeth developed, on two short oblique between choanae, equal in distance from each other as to choanae; (3) supernumerary tubercles present at the base of each finger; (4) tympanum indistinct; (5) three metacarpal tubercles, inner metacarpal tubercle long, outer metacarpal tubercle relatively small, oval, median one rounded; (6) supratympanic fold indistinct; (7) discontinuous glandular dorsolateral fold from rear of eye to near vent; (8) circummarginal grooves present on tips of outer three fingers, absent on first finger; (9) iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths, black reticulations throughout; (10) rictal gland absent; (11) dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots; (12) dorsal parts of limbs, fingers and toes with black crossbars; (13) vocal sac absent in males; (14) male with orange nuptial pad at the base of first finger.
</paragraph>
</subSubSection>
<subSubSection id="F45A03E50A4101FD2441109D55DEA05A" pageId="0" pageNumber="343" type="description">
<paragraph id="B46D871384C07CA9EE5E4A330FF41993" pageId="0" pageNumber="343">Description of holotype</paragraph>
<paragraph id="347A8E81317002AECA30D8DBB1122EA1" pageId="0" pageNumber="343">
<emphasis id="0138DF3123F9BEB43EA5D7113B499C3F" bold="true" pageId="0" pageNumber="343">
(all measurements in mm; see Table
<tableCitation id="C668F628B90FEA1D64335F9938882878" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (mm) of Amolops yangi sp. nov. and A. vitreus. The asterisk (*) indicates the holotype." httpUri="http://table.plazi.org/id/94A921A1A38F6838C1FCF2ECC2413F8C" pageId="0" pageNumber="343" tableUuid="94A921A1A38F6838C1FCF2ECC2413F8C">1</tableCitation>
).
</emphasis>
KIZ 038643, sexually mature female, body size moderate, adult female (SVL 51.5 mm); head length larger than wide (HL/SVL 34.2%, HW/SVL 32.6%); top of head flat; snout short (SL/HL 46.0%), snout rounded in dorsal view (Fig.
<figureCitation id="BF9643C776E93BAA35137FCE03376E59" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4A</figureCitation>
), obtusely rounded in profile, projecting beyond margin of lower jaw; loreal region and concave and oblique; canthus rostralis distinct, slightly constricted behind nostrils; dorsal region of snout flattened; eyes relatively large (ED/HL 29.0%), slightly protuberant in dorsal view and notably protruding in profile (Fig.
<figureCitation id="58D8CC11C2C8A2E7202556C725418BEF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4A</figureCitation>
), eye diameter shorter than snout length (ED/SL 63.0%); nostrils oval, laterally orientated, slightly protuberant, closer to anterior corner of eye than to tip of snout; pupil oval, horizontal; tympanum indistinct, circular in shape, relatively small (TD/HL 10.2%), tympanum diameter about one third of eye diameter (TD/ED 35.3%); internarial distance (INS/SVL 13.4%) larger than width of upper eyelid (INS/UEW 168.3%) and interorbital distance (INS/IOS 160.5%); tongue cordiform, deeply notched posteriorly; vomerine teeth developed, on two short oblique between choanae, equal in distance from each other as to choanae; choanae oval; maxillary teeth developed; a small tooth-like projection on anteromedial edge of mandible.
</paragraph>
<caption id="94A921A1A38F6838C1FCF2ECC2413F8C" ID-Table-UUID="94A921A1A38F6838C1FCF2ECC2413F8C" httpUri="http://table.plazi.org/id/94A921A1A38F6838C1FCF2ECC2413F8C" pageId="0" pageNumber="343" start="Table 1" startId="T1">
<paragraph id="067A26061250362E3180852FED4BEAE8" pageId="0" pageNumber="343">
<emphasis id="728F111C72FB7198C524B8ECED7DE4FC" bold="true" pageId="0" pageNumber="343">Table 1.</emphasis>
Measurements (mm) of
<taxonomicName id="6ADF24008342317F845FB48E183FB38F" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="C678CF8D1AC459A07B61AEB58C17F0CF" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="385EA7CAE2EB91BD9ACA69E20D3874ED" bold="true" pageId="0" pageNumber="343">sp. nov</emphasis>
. and
<taxonomicName id="99F75E768F081FADFBA65D14F4D3DDAF" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. vitreus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="vitreus">
<emphasis id="75261166026C8306F26B0D166D2746FD" italics="true" pageId="0" pageNumber="343">A. vitreus</emphasis>
</taxonomicName>
. The asterisk (*) indicates the holotype.
</paragraph>
</caption>
<paragraph id="4809C945C944E5B88192215F78AC79BF" pageId="0" pageNumber="343">
<table id="5443F8A55D84BE3E2C0989F43BC01034" pageId="0" pageNumber="343">
<tr id="B64C98B5411AA08CD14234671E76192A" pageId="0" pageNumber="343">
<td id="9A66BBE7990C35315AC9CEC4EA3312D3" colspan="1" isEmpty="true" pageId="0" pageNumber="343" rowspan="1">-</td>
<td id="6D0308234E1AAEC8F74984F783871F71" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="4E859DF5A4E372FEEDEB319570D2848B" bold="true" pageId="0" pageNumber="343">
<taxonomicName id="285CA64F75C072B9EB4DB285CBEFD74E" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. yangi" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="4C3B4A98838EE3484CAF2D49ECE2C774" italics="true" pageId="0" pageNumber="343">A. yangi</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
<td id="81067D177D68D80D0408792E42D97F2A" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="5D6F8EB0C20F1751CEE833DB2BF3B309" bold="true" pageId="0" pageNumber="343">
<taxonomicName id="B9FC9E1465DDEFAF50D468C44DEDDBC9" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. yangi" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="5F9E0EACB12BDED71AE619D0DED0EB9F" italics="true" pageId="0" pageNumber="343">A. yangi</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
<td id="FE518F7BD437615675CFA32F62352664" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="D65A3026F174F3D8A7F3EBEFD38ECDDA" bold="true" pageId="0" pageNumber="343">
<taxonomicName id="63BDAE025C4CD788E3C4EDAFBFE1E072" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. yangi" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="9737E4E9B99438D05DC00AD7EC6E0382" italics="true" pageId="0" pageNumber="343">A. yangi</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
*
</td>
<td id="A3B6F819B0B8E513D39F9AD8730316A5" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<taxonomicName id="1EF146380457590122AF0EF1FD85A8E9" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. vitreus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="vitreus">
<emphasis id="922A059E928454196F9F677142F1CF9B" bold="true" italics="true" pageId="0" pageNumber="343">A. vitreus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="A19FC936355DED1DF35A8990DEEF5873" pageId="0" pageNumber="343">
<td id="5FC7FD156B9D054B9B2C01E2DFEA5964" colspan="1" pageId="0" pageNumber="343" rowspan="1">Sex</td>
<td id="7DCA5039FF8366F1C264D5B7E89F0304" colspan="1" pageId="0" pageNumber="343" rowspan="1"></td>
<td id="58C3B4F17E702A8A541B809056A80034" colspan="1" pageId="0" pageNumber="343" rowspan="1"></td>
<td id="326AFCAE6D0DCAE4B3F7DBACE444E3FF" colspan="1" pageId="0" pageNumber="343" rowspan="1"></td>
<td id="1A4CE1BC6039EEBEB38C05A35D9FA8B7" colspan="1" pageId="0" pageNumber="343" rowspan="1"></td>
</tr>
<tr id="7B6842C96663471024CE31BC3DD67734" pageId="0" pageNumber="343">
<td id="8113465D740F83BA488CFED21F96CAA1" colspan="1" pageId="0" pageNumber="343" rowspan="1">Catalog No.</td>
<td id="4042E9D76A7C0FB8764256ECD56C6879" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="549DB4947B889D0B00B821D9AE0C2FAD" bold="true" pageId="0" pageNumber="343">KIZ</emphasis>
050788
</td>
<td id="6B00E544226187FA893C466CE126750C" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="054574A8A49CC065A9688F3961A57E60" bold="true" pageId="0" pageNumber="343">KIZ</emphasis>
038645
</td>
<td id="9D83B62E13B0373619CE08DD5A56B0ED" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="311F3427130BC54341F259436A5B3335" bold="true" pageId="0" pageNumber="343">KIZ</emphasis>
038643
</td>
<td id="2645A2943B2220216C6A60674EF4152C" colspan="1" pageId="0" pageNumber="343" rowspan="1">
<emphasis id="70C968BDCE45B8B5B0172C7F086C8D4B" bold="true" pageId="0" pageNumber="343">KIZ</emphasis>
050452
</td>
</tr>
<tr id="D56F4F1598164E92FDC2ECA7B04E7C2A" pageId="0" pageNumber="343">
<td id="2D17541E3CDC9A68A797165C27143380" colspan="1" pageId="0" pageNumber="343" rowspan="1">SVL</td>
<td id="93608846F225B10926496371780670CE" colspan="1" pageId="0" pageNumber="343" rowspan="1">51.8</td>
<td id="7EA71602966DA7E2D5DEA743520396FC" colspan="1" pageId="0" pageNumber="343" rowspan="1">46.3</td>
<td id="D98E489461AF92DAE25ABB633C4B75B0" colspan="1" pageId="0" pageNumber="343" rowspan="1">51.5</td>
<td id="5857793BF51F2AEFA2E3ECEF3EA35837" colspan="1" pageId="0" pageNumber="343" rowspan="1">38.9</td>
</tr>
<tr id="6ED147C864784C1C300265508C5CA932" pageId="0" pageNumber="343">
<td id="7DA068D4EF960044181F022462153370" colspan="1" pageId="0" pageNumber="343" rowspan="1">HL</td>
<td id="0811A10E8742FC42B36AF17AAE48EB98" colspan="1" pageId="0" pageNumber="343" rowspan="1">17.0</td>
<td id="05D73A53E4754B52BEB230B34CF7399B" colspan="1" pageId="0" pageNumber="343" rowspan="1">16.8</td>
<td id="AE545C9FED6C646D92F3BDA596FDEC07" colspan="1" pageId="0" pageNumber="343" rowspan="1">17.6</td>
<td id="6D4E39FE605B0C5685A0E2D9C5F25AB9" colspan="1" pageId="0" pageNumber="343" rowspan="1">15.4</td>
</tr>
<tr id="0B36E9FA3F94366FBBDF88FC358FC181" pageId="0" pageNumber="343">
<td id="755D2647476C8D930141E0E7DC0886B4" colspan="1" pageId="0" pageNumber="343" rowspan="1">HL/SVL</td>
<td id="FCC0D7A7CA426D8500B70AC66ACDF4D3" colspan="1" pageId="0" pageNumber="343" rowspan="1">32.8%</td>
<td id="544A8737AB5BDCC594C7984B69A3E9A2" colspan="1" pageId="0" pageNumber="343" rowspan="1">36.3%</td>
<td id="11A755A2D59285C2C241E8F946102074" colspan="1" pageId="0" pageNumber="343" rowspan="1">34.2%</td>
<td id="F61B0F0076F5C5C956F268E7C172087D" colspan="1" pageId="0" pageNumber="343" rowspan="1">39.6%</td>
</tr>
<tr id="CEEB4F477EE2A72E1425AB96605B8B5F" pageId="0" pageNumber="343">
<td id="4F48709758330376C2E135B47DFBC35B" colspan="1" pageId="0" pageNumber="343" rowspan="1">HW</td>
<td id="DFDD29DB9D88199D8E3589DF6EB063D5" colspan="1" pageId="0" pageNumber="343" rowspan="1">16.7</td>
<td id="0253596891D09BAC860CB5CBAD028208" colspan="1" pageId="0" pageNumber="343" rowspan="1">15.4</td>
<td id="918974AC7B8996383B522E6164B57F22" colspan="1" pageId="0" pageNumber="343" rowspan="1">16.8</td>
<td id="D9417B179D659B95FC6142B0CFA61025" colspan="1" pageId="0" pageNumber="343" rowspan="1">13.9</td>
</tr>
<tr id="3A8217BC6B524BB55701CECFC09BC348" pageId="0" pageNumber="343">
<td id="D3FAD261C703A91B2B7F20FF684BEDAD" colspan="1" pageId="0" pageNumber="343" rowspan="1">HW/SVL</td>
<td id="7012CE07837DAA4BCA9434B6D93D9FA9" colspan="1" pageId="0" pageNumber="343" rowspan="1">32.2%</td>
<td id="CC8D98052B561E984E2280610F4018F9" colspan="1" pageId="0" pageNumber="343" rowspan="1">33.3%</td>
<td id="29D3F85B098F4C549B0B93FD4799399C" colspan="1" pageId="0" pageNumber="343" rowspan="1">32.6%</td>
<td id="F25688C7C14AA5AA40A4AFDE47349731" colspan="1" pageId="0" pageNumber="343" rowspan="1">35.7%</td>
</tr>
<tr id="34A315C254C190A8B8834C41C4753201" pageId="0" pageNumber="343">
<td id="9D0A71835837864F5FAC43808A634033" colspan="1" pageId="0" pageNumber="343" rowspan="1">SL</td>
<td id="96046E8731CFBBA7E05DF202135D28B9" colspan="1" pageId="0" pageNumber="343" rowspan="1">7.2</td>
<td id="38A62C5FB3891F27A9FF2848498F92F1" colspan="1" pageId="0" pageNumber="343" rowspan="1">7.7</td>
<td id="C0DCBEC010C2B3FB82FA3CA3D3D46323" colspan="1" pageId="0" pageNumber="343" rowspan="1">8.1</td>
<td id="4B3E7BFE2CFA3BEE3672E37CD5BB973D" colspan="1" pageId="0" pageNumber="343" rowspan="1">6.5</td>
</tr>
<tr id="BC9DED8185CAB009DE26B63F51D83A82" pageId="0" pageNumber="343">
<td id="D5BD71DB863F3F2A4F037A436DA8120E" colspan="1" pageId="0" pageNumber="343" rowspan="1">SL/SVL</td>
<td id="9FC91D8FE60D691FBD1D4A50AE9B375C" colspan="1" pageId="0" pageNumber="343" rowspan="1">13.9%</td>
<td id="90764CA9D3475411C217BE96E8189137" colspan="1" pageId="0" pageNumber="343" rowspan="1">16.6%</td>
<td id="FEE3A4B6098EA94A8F3D1030FA8CFEC8" colspan="1" pageId="0" pageNumber="343" rowspan="1">15.7%</td>
<td id="C4AAFBCEA5F61D9A3AD4FE8EB3D51394" colspan="1" pageId="0" pageNumber="343" rowspan="1">16.7%</td>
</tr>
<tr id="30BDB6A87B0048EBC657EE65199F76B7" pageId="0" pageNumber="343">
<td id="C3E9B4F5E46897CCB1AE52F5267E52A5" colspan="1" pageId="0" pageNumber="343" rowspan="1">SL/HL</td>
<td id="8932D07A9EB4F0FAADA2F3C98ADBC62C" colspan="1" pageId="0" pageNumber="343" rowspan="1">42.4%</td>
<td id="3E5848A9F2B7203946060A3F530FF2A5" colspan="1" pageId="0" pageNumber="343" rowspan="1">45.8%</td>
<td id="EB3931A9997B36FD2A5C83826C261669" colspan="1" pageId="0" pageNumber="343" rowspan="1">46.0%</td>
<td id="F8DD23101942DAB311FEECC4E13A1E12" colspan="1" pageId="0" pageNumber="343" rowspan="1">42.2%</td>
</tr>
<tr id="5366EFC10092E7B51701B0D0535E3A11" pageId="0" pageNumber="343">
<td id="A42E25B2507CE72FC962575D9E7B51A4" colspan="1" pageId="0" pageNumber="343" rowspan="1">INS</td>
<td id="3BCA718B9B9395802A3067AAD5AE9F3A" colspan="1" pageId="0" pageNumber="343" rowspan="1">6.7</td>
<td id="D5C5C75ADB3F1DE434C2694BC1F25DC1" colspan="1" pageId="0" pageNumber="343" rowspan="1">6.5</td>
<td id="FA395ED9C168BF6B9A323F36D630317D" colspan="1" pageId="0" pageNumber="343" rowspan="1">6.9</td>
<td id="63B4270258A04693D9A6C2781C04E25D" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.6</td>
</tr>
<tr id="79F151DF194425FBEE221314F4C710ED" pageId="0" pageNumber="343">
<td id="A464FEE03648001060EC8F08AAB9CA54" colspan="1" pageId="0" pageNumber="343" rowspan="1">INS/SVL</td>
<td id="F1A7D725C89A22AAADBDC073B648430B" colspan="1" pageId="0" pageNumber="343" rowspan="1">12.9%</td>
<td id="5332A47BAC53CCF38C48DD444BE78B32" colspan="1" pageId="0" pageNumber="343" rowspan="1">14.0%</td>
<td id="76FC890857221717E0D3A40C3EE1C2AD" colspan="1" pageId="0" pageNumber="343" rowspan="1">13.4%</td>
<td id="1AF182DAABB0FA87CBE77ACE861C2E76" colspan="1" pageId="0" pageNumber="343" rowspan="1">11.8%</td>
</tr>
<tr id="904E53D6D1DB48E98F5CB5DDB7C593D1" pageId="0" pageNumber="343">
<td id="0755BBC5680A56E3C539D634A1496942" colspan="1" pageId="0" pageNumber="343" rowspan="1">IOS</td>
<td id="25F22F74F60EBF69B3365FCAB91EC074" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.1</td>
<td id="5E38DD1E662DEB415BDAA1D5883A4784" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.8</td>
<td id="73D2DBA46D62F7D04C7D6C055142634A" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.3</td>
<td id="A1D0D74637F52F810DA535132BC82F93" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.1</td>
</tr>
<tr id="DC6D6BD563E385418992A84E82282579" pageId="0" pageNumber="343">
<td id="BCBBA1A04EC26BE7B2E7E8E060D4433B" colspan="1" pageId="0" pageNumber="343" rowspan="1">INS/IOS</td>
<td id="BE3EAF8C011DF18633C01B0D4E28D441" colspan="1" pageId="0" pageNumber="343" rowspan="1">163.4%</td>
<td id="98FC3A1E8D6C3E04F3B6EC9E720A6340" colspan="1" pageId="0" pageNumber="343" rowspan="1">135.4%</td>
<td id="AC81C45F8D2C736ED186E735ACFCF397" colspan="1" pageId="0" pageNumber="343" rowspan="1">160.5%</td>
<td id="A2D3ECE982BA75DE036F504EDE5E5A5B" colspan="1" pageId="0" pageNumber="343" rowspan="1">112.2%</td>
</tr>
<tr id="AC655E864AE14FF52F813C07020172BB" pageId="0" pageNumber="343">
<td id="F87D111E7563C0437356411CFD55F49F" colspan="1" pageId="0" pageNumber="343" rowspan="1">NED</td>
<td id="579B94FD4FD6FB0B4107ED9BECC3E7B8" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.3</td>
<td id="47D6A794AA90D3522D56B5368210085A" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.2</td>
<td id="05483E3FE39C4D1C8952934C8CC8F6E0" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.3</td>
<td id="9AE288C58FDE123FE7F003FD161AB56B" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.4</td>
</tr>
<tr id="5A632A8E2EDE62497E6648ED91A0CB3C" pageId="0" pageNumber="343">
<td id="94370EC2B1EBDD3FA441278584FB6006" colspan="1" pageId="0" pageNumber="343" rowspan="1">UEW</td>
<td id="E271C8D6438AF85196826FD7B03CC4E7" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.1</td>
<td id="DC49683AA86002B10735A470318D2BF0" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.6</td>
<td id="91A3031FC7076036F1500CC85684C269" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.1</td>
<td id="C0A4019B6088067A9EDA5514CE19FD32" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.8</td>
</tr>
<tr id="515D5F95C763588FCD9D9F9DB633A9FD" pageId="0" pageNumber="343">
<td id="45FDB1436F5F348E64DA94FD90334336" colspan="1" pageId="0" pageNumber="343" rowspan="1">INS/UEW</td>
<td id="F821B324848D658D3CD091D37FFD17D0" colspan="1" pageId="0" pageNumber="343" rowspan="1">163.4%</td>
<td id="2E3093C45431B3BB0F1389C159601362" colspan="1" pageId="0" pageNumber="343" rowspan="1">180.6</td>
<td id="4DD9F2A8368F7BBFC4EE98EAE7EB5632" colspan="1" pageId="0" pageNumber="343" rowspan="1">168.3%</td>
<td id="B31DC540D4B50D96FF5C86238D785825" colspan="1" pageId="0" pageNumber="343" rowspan="1">121.1%</td>
</tr>
<tr id="FD88EFB219D1127BE253A994CC0B04BA" pageId="0" pageNumber="343">
<td id="F76E64D8DBAEE0ABE366C48F8FD31973" colspan="1" pageId="0" pageNumber="343" rowspan="1">UEW/IOS</td>
<td id="95630DDC4A58F3059A0258B9165A1362" colspan="1" pageId="0" pageNumber="343" rowspan="1">100.0%</td>
<td id="6E3B8F298312B1DA55C3F5EB203A23B8" colspan="1" pageId="0" pageNumber="343" rowspan="1">75.0%</td>
<td id="F6453DB9371159698DA6C751B2F2F690" colspan="1" pageId="0" pageNumber="343" rowspan="1">95.3%</td>
<td id="BA963CE732B311E8111761B810959244" colspan="1" pageId="0" pageNumber="343" rowspan="1">92.7%</td>
</tr>
<tr id="D390CA53B61B3C0542297610B383391C" pageId="0" pageNumber="343">
<td id="C7FD3282FC6E8C488663948A0C650220" colspan="1" pageId="0" pageNumber="343" rowspan="1">ED</td>
<td id="246CC63F0318CECA89FC9DE6790AD144" colspan="1" pageId="0" pageNumber="343" rowspan="1">5.0</td>
<td id="1CA86DEA9ED81414E77AD9CF6AFB7262" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.9</td>
<td id="A76D2544307D973043F9EA56FCCADD93" colspan="1" pageId="0" pageNumber="343" rowspan="1">5.1</td>
<td id="A6C4BF5AD77CE438F69CFDF3457B16EF" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.4</td>
</tr>
<tr id="D99BCF99D4EEA5E0DFEEB29D9192AEC0" pageId="0" pageNumber="343">
<td id="858B7A42ED8907DB2C2E15D2499BA2B1" colspan="1" pageId="0" pageNumber="343" rowspan="1">ED/HL</td>
<td id="01DF3BB2FD97CB0E59CD0EB1B63E4381" colspan="1" pageId="0" pageNumber="343" rowspan="1">29.4%</td>
<td id="8F476B409E1AFFF0039F487A022CCE44" colspan="1" pageId="0" pageNumber="343" rowspan="1">29.2%</td>
<td id="31D75BA6AE8DDA8B9C7D650ED2829B4B" colspan="1" pageId="0" pageNumber="343" rowspan="1">29.0%</td>
<td id="9532802ED2F43B018066D84C8E642016" colspan="1" pageId="0" pageNumber="343" rowspan="1">28.6%</td>
</tr>
<tr id="5D06EB257A3054877D4E4E5FF6F48512" pageId="0" pageNumber="343">
<td id="A63CCF43CCC854313C2FE86500E93AF7" colspan="1" pageId="0" pageNumber="343" rowspan="1">ED/SL</td>
<td id="E7736B07F893080EC115EBF4F95DCEF3" colspan="1" pageId="0" pageNumber="343" rowspan="1">69.4%</td>
<td id="D1BAEA49AA014E698F26BB68DFA93822" colspan="1" pageId="0" pageNumber="343" rowspan="1">63.6%</td>
<td id="4C9F576827E8842EDBEC0E0F8652FBD5" colspan="1" pageId="0" pageNumber="343" rowspan="1">63.0%</td>
<td id="2AF3AAE6731140C2A364688C7D6FD1BB" colspan="1" pageId="0" pageNumber="343" rowspan="1">67.7%</td>
</tr>
<tr id="BFEF655CFEF06CDD5C8FC7070105E3FD" pageId="0" pageNumber="343">
<td id="0E93E4FF154029208487440F263A62BA" colspan="1" pageId="0" pageNumber="343" rowspan="1">TD</td>
<td id="E768D53CBA6D08893466483B4A67E59B" colspan="1" pageId="0" pageNumber="343" rowspan="1">2.6</td>
<td id="EB00AC20F505D9FA7C6977F057600152" colspan="1" pageId="0" pageNumber="343" rowspan="1">2.4</td>
<td id="11DD267BAE9C9B644D2F25DBC8878A45" colspan="1" pageId="0" pageNumber="343" rowspan="1">1.8</td>
<td id="0C99190C00DD38664DFB8A037C7038C4" colspan="1" pageId="0" pageNumber="343" rowspan="1">2.7</td>
</tr>
<tr id="897CBA1BA5AE655E1D4B5E854738C66A" pageId="0" pageNumber="343">
<td id="2ED4BC4F6DA93F461F44CC57E9D427AF" colspan="1" pageId="0" pageNumber="343" rowspan="1">TD/ED</td>
<td id="E3BF30AC144957D52BAFC3B48AF537B4" colspan="1" pageId="0" pageNumber="343" rowspan="1">52.0%</td>
<td id="D33F2AF385917CBA238AF4F40E4207CD" colspan="1" pageId="0" pageNumber="343" rowspan="1">49.0%</td>
<td id="6FBE90EAAD0BD7FBB47BA00AAF140764" colspan="1" pageId="0" pageNumber="343" rowspan="1">35.3%</td>
<td id="9F68523A3267FEB9569459E613E76425" colspan="1" pageId="0" pageNumber="343" rowspan="1">61.4%</td>
</tr>
<tr id="23ED83E21C1222EDDF36B864A831559E" pageId="0" pageNumber="343">
<td id="42A97F792C94E3E4DA198EA06A5882B6" colspan="1" pageId="0" pageNumber="343" rowspan="1">LAHL</td>
<td id="AC3433B7F388288037A904C683E752AE" colspan="1" pageId="0" pageNumber="343" rowspan="1">26.9</td>
<td id="A4D74902F7810590C6515FB7F24EC68F" colspan="1" pageId="0" pageNumber="343" rowspan="1">25.0</td>
<td id="797499F51230F1C25A63BE213EC46B36" colspan="1" pageId="0" pageNumber="343" rowspan="1">29.1</td>
<td id="5ED3C51F148F6FFC7EA869E7C41A0F5F" colspan="1" pageId="0" pageNumber="343" rowspan="1">19.6</td>
</tr>
<tr id="F36A4A750CFEE82B05DE9DE4498A2C5F" pageId="0" pageNumber="343">
<td id="BE05BC2615859DC2DB95C978C9E85B79" colspan="1" pageId="0" pageNumber="343" rowspan="1">LAHL/SVL</td>
<td id="2DF907DD64AA928A01496C6616ADC4F1" colspan="1" pageId="0" pageNumber="343" rowspan="1">51.9%</td>
<td id="D3B3C29C778FAB77BA989EEBA6C3314C" colspan="1" pageId="0" pageNumber="343" rowspan="1">54.0%</td>
<td id="C2D6AEAF904E9C9E89F93B2049184867" colspan="1" pageId="0" pageNumber="343" rowspan="1">56.5%</td>
<td id="514168746FBBF884C5596604A354954C" colspan="1" pageId="0" pageNumber="343" rowspan="1">50.4%</td>
</tr>
<tr id="B09C43EC26073E673875E7C45D3518F3" pageId="0" pageNumber="343">
<td id="3257EA5611F7EA2F80EA5C2599759155" colspan="1" pageId="0" pageNumber="343" rowspan="1">HND</td>
<td id="603B344AAA94582C8E34C51CE53B1D7A" colspan="1" pageId="0" pageNumber="343" rowspan="1">17.1</td>
<td id="75E412087B9AC5D9F63572FB177B9278" colspan="1" pageId="0" pageNumber="343" rowspan="1">15.4</td>
<td id="31D81105D103D8329CEF86C5B1BFA87B" colspan="1" pageId="0" pageNumber="343" rowspan="1">18.4</td>
<td id="670C32DEEA6FD77084818C823449224B" colspan="1" pageId="0" pageNumber="343" rowspan="1">12.2</td>
</tr>
<tr id="3206573933573F2678A4CCBB7F8434F9" pageId="0" pageNumber="343">
<td id="36F66F46E93B5916AF28FD5DCD28C09C" colspan="1" pageId="0" pageNumber="343" rowspan="1">HND/SVL</td>
<td id="D8680EDACD3D58416538EBBD039B5E48" colspan="1" pageId="0" pageNumber="343" rowspan="1">33.0%</td>
<td id="E87DB467FAD9D0FDF766927D2661E2A6" colspan="1" pageId="0" pageNumber="343" rowspan="1">33.3%</td>
<td id="F0B64E3AC1BB7A1CF0BBCF7C6EB8B7DD" colspan="1" pageId="0" pageNumber="343" rowspan="1">35.7%</td>
<td id="CECC18D16CC8DA7113569724279B5CB2" colspan="1" pageId="0" pageNumber="343" rowspan="1">31.4%</td>
</tr>
<tr id="0F25EF2534EBE5356B5313ABC0D86A07" pageId="0" pageNumber="343">
<td id="1FD48B433A458F1F13099835ACFCBB2D" colspan="1" pageId="0" pageNumber="343" rowspan="1">LAD</td>
<td id="3138029142B415306DCBB0D58624B8A2" colspan="1" pageId="0" pageNumber="343" rowspan="1">5.0</td>
<td id="5BAFFE8728ED058844CAE1DEA6145037" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.8</td>
<td id="6BECCFE05B8C19C35FC2DA12D618D8BC" colspan="1" pageId="0" pageNumber="343" rowspan="1">4.4</td>
<td id="89E07F5D87DC99ADE981F48EFB85FA9F" colspan="1" pageId="0" pageNumber="343" rowspan="1">3.8</td>
</tr>
<tr id="C8506CBC903A4C5C2E86E88AADF1BA2E" pageId="0" pageNumber="343">
<td id="8D7CE17D3A51F4EC0A3B6E48746EEB58" colspan="1" pageId="0" pageNumber="343" rowspan="1">LAD/SVL</td>
<td id="AF7D3A80959BF77BA58A71CFFF5DAA2A" colspan="1" pageId="0" pageNumber="343" rowspan="1">9.7%</td>
<td id="1133F0B0F6D90A15FFD3254414A6EE5F" colspan="1" pageId="0" pageNumber="343" rowspan="1">10.4%</td>
<td id="86F3D6378975C4BB1983E6080C71F3BC" colspan="1" pageId="0" pageNumber="343" rowspan="1">8.5%</td>
<td id="CC4452F5E3FF7AAC7DC970E6ACAAFC18" colspan="1" pageId="0" pageNumber="343" rowspan="1">9.8%</td>
</tr>
<tr id="5E5D4831C13D0696573252F6E9E0E340" pageId="0" pageNumber="343">
<td id="214788C3FB781A68312FDE0A945D181D" colspan="1" pageId="0" pageNumber="343" rowspan="1">FEM</td>
<td id="CFB1802C2BF532D606BF28C24077D1CA" colspan="1" pageId="0" pageNumber="343" rowspan="1">26.3</td>
<td id="57F0B859DCDE85DD44C4CB9FC983C27F" colspan="1" pageId="0" pageNumber="343" rowspan="1">22.7</td>
<td id="72BD6B0D0FF922C0F4CA1DADA6E01D83" colspan="1" pageId="0" pageNumber="343" rowspan="1">25.3</td>
<td id="F29E3124F8E701615B33ACA1EB088081" colspan="1" pageId="0" pageNumber="343" rowspan="1">22.2</td>
</tr>
<tr id="85112F332DE2EDECF21A8D872CA533FC" pageId="0" pageNumber="343">
<td id="E0E3D4383E221524D06CEC56CA0341BF" colspan="1" pageId="0" pageNumber="343" rowspan="1">TIB</td>
<td id="A7535C58916474DD3AEBF203EF771373" colspan="1" pageId="0" pageNumber="343" rowspan="1">27.1</td>
<td id="8FE3CEE0B03B9B745330B1F3897FA6D2" colspan="1" pageId="0" pageNumber="343" rowspan="1">24.3</td>
<td id="D0F41C0A992EF03D96E450DDD3BDE80C" colspan="1" pageId="0" pageNumber="343" rowspan="1">27.2</td>
<td id="199C543C92BD364327B2008386106E60" colspan="1" pageId="0" pageNumber="343" rowspan="1">23.5</td>
</tr>
<tr id="DB46AFB8FED619DAA2B5BCCDB966A9FD" pageId="0" pageNumber="343">
<td id="141142FD105F60D2A468F02E7E1E8DA5" colspan="1" pageId="0" pageNumber="343" rowspan="1">FEM/TIB</td>
<td id="44FCC950E7C4C451F452465CC5F9E1B1" colspan="1" pageId="0" pageNumber="343" rowspan="1">97.0%</td>
<td id="C157E42787CF929FB9FAFDCEB401B96C" colspan="1" pageId="0" pageNumber="343" rowspan="1">93.4%</td>
<td id="0F098B73481FF6710AD7F97E9CEFF6DE" colspan="1" pageId="0" pageNumber="343" rowspan="1">93.0%</td>
<td id="A18E2FF7E70728789754ADDC02DB6482" colspan="1" pageId="0" pageNumber="343" rowspan="1">94.5%</td>
</tr>
<tr id="D3E63E3D20C1DF483B0CC5099F54AF8B" pageId="0" pageNumber="343">
<td id="A7B145CCBE1D08CEBAE0FFAC1F327897" colspan="1" pageId="0" pageNumber="343" rowspan="1">FTL</td>
<td id="7A3D05A3613137F6098F69018BA2C410" colspan="1" pageId="0" pageNumber="343" rowspan="1">27.7</td>
<td id="92540436F2A4383601E16E007C584E93" colspan="1" pageId="0" pageNumber="343" rowspan="1">26.8</td>
<td id="23F3265F769EE81EA3A4C8AE16E6AD08" colspan="1" pageId="0" pageNumber="343" rowspan="1">28.0</td>
<td id="2946EE073C80D31E86F679F28F90AB66" colspan="1" pageId="0" pageNumber="343" rowspan="1">20.3</td>
</tr>
<tr id="3A99A676D0DB022167A83D93582E1034" pageId="0" pageNumber="343">
<td id="610EC98AD55221BA60B9FD2B69F6C647" colspan="1" pageId="0" pageNumber="343" rowspan="1">FEM/FTL</td>
<td id="BA455651E6676BF8AE22C8321F2F7156" colspan="1" pageId="0" pageNumber="343" rowspan="1">94.9%</td>
<td id="A557A6BB0355BB1F918AC076E2319D61" colspan="1" pageId="0" pageNumber="343" rowspan="1">84.7%</td>
<td id="1BC68F658073C9D744833BA08F380BFD" colspan="1" pageId="0" pageNumber="343" rowspan="1">90.4%</td>
<td id="CB3B39A668CEEC9AB97C139CC6E31E03" colspan="1" pageId="0" pageNumber="343" rowspan="1">109.4%</td>
</tr>
</table>
</paragraph>
<caption id="6C6CB5C940F4DA305B6363A4D2E1E535" doi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343" start="Figure 4" startId="F4">
<paragraph id="7F45357657EE7F6E4A80C0F99769FB46" pageId="0" pageNumber="343">
<emphasis id="F843C76C4C90DCF463144F9921271496" bold="true" pageId="0" pageNumber="343">Figure 4.</emphasis>
<taxonomicName id="1C028CB6BEF7CCD46A2AF7390DDE1A3B" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="AC519798F3660D5DC60B0C565C2F9DCA" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="11E0CE43BDEBA989E8C3313A83E0BDE4" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
(Holotype KIZ 038643).
<emphasis id="B0943A31BD708F847A8AEB56133A940E" bold="true" pageId="0" pageNumber="343">A</emphasis>
Lateral view,
<emphasis id="BAE4E74B617010327B032F4B67B1CEE4" bold="true" pageId="0" pageNumber="343">B</emphasis>
dorsal view,
<emphasis id="CBA888B21804C36A9018E950BC2068CD" bold="true" pageId="0" pageNumber="343">C</emphasis>
ventral view of thighs,
<emphasis id="08B253DC8AA3A8A769C41C24DCFEFE8B" bold="true" pageId="0" pageNumber="343">D</emphasis>
ventral view of hand,
<emphasis id="63804A0246A1FC2A487C043B9ED78800" bold="true" pageId="0" pageNumber="343">E</emphasis>
foot,
<emphasis id="9B63A6462CD563BA40978F00A127245F" bold="true" pageId="0" pageNumber="343">F</emphasis>
habitat. Photos by Zhong-Bin Yu.
</paragraph>
</caption>
<paragraph id="87AF4F9234F6AB6B496BBABCFF35F0E4" pageId="0" pageNumber="343">
Forelimbs moderately long and robust, forelimb and hand length (29.1 mm) longer than half body size (LAHL/SVL 56.5%); relative length of fingers: I&lt;II&lt;IV&lt;III; circummarginal grooves present on tips of outer three fingers, absent on first finger; subarticular tubercles prominent and oval, formula 1, 1, 2, 2; supernumerary tubercles present at the base of each finger; webbing between fingers absent; narrow lateral fringes of fingers III and IV; three metacarpal tubercles, inner metacarpal tubercle long, outer metacarpal tubercle relatively small, oval, median one rounded (Fig.
<figureCitation id="A2EC52606D3B12517D30AD0170ABDC94" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4D</figureCitation>
).
</paragraph>
<paragraph id="4A16F3F1C35C960BCCDE6EB504148E9D" pageId="0" pageNumber="343">
Hindlimbs long and robust, femoral length shorter than the tibia length (FEM/TIB 93.0%) and the foot length (FEM/FTL 90.4%); tibiotarsal articulation of adpressed limb reaching between nostrils and eyes when hindlimb stretched alongside of body; the heels overlapping when the tibias are perpendicular to the body axis; relative toes lengths: I&lt;II&lt;III&lt;V&lt;IV; narrow lateral fringes of preaxial side of toe I and postaxial side of toe V; tips of all toes expanded into discs with circummarginal grooves; toes fully webbed except for fourth toe, in which web reaches beyond distal subarticular tubercle; subarticular tubercles oval and distinct, formula 1, 1, 2, 3, 2; supernumerary tubercles absent; inner metatarsal tubercle long, outer metatarsal tubercle absent (Fig.
<figureCitation id="A561E59A0FFBC392D193460C0D81C044" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4E</figureCitation>
).
</paragraph>
<paragraph id="BA0A7C3E120652B9533798211BECDE90" pageId="0" pageNumber="343">
Dorsal surface of head, body, limbs, fingers, toes and flank of body relatively smooth; loreal region densely scattered with raised tubercles; temporal region and posterior angle of the jaw with dense tubercles; skin ventrally smooth, including throat, chest, abdomen, and ventral surface of limbs; discontinuous glandular dorsolateral fold from rear of eye to near vent; supratympanic fold indistinct; rictal gland absent (Fig.
<figureCitation id="B733951183BB393A926D14EDC832631B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4A-E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="29D6FE313733F1506207B1CCF8AA16A9" pageId="0" pageNumber="343" type="color of holotype in life">
<paragraph id="663FCEC834BD672EA2889FDCC4B55B80" pageId="0" pageNumber="343">Color of holotype in life.</paragraph>
<paragraph id="1FA617B529675FDBC075FCD97BD5B09A" pageId="0" pageNumber="343">
For coloration of the holotype in life see Figure
<figureCitation id="D8CEBAED1074333B6C6A764B7418DA28" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4A-E</figureCitation>
. Dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots; throat, ventral surface of the head, chest and anterior abdomen mostly yellow, with scattered grayish spots; upper part of flanks green, lower part of flanks green-yellow; a black stripe below edge of the canthus rostralis extending from the snout tip across the eyes, to the anterior edge of supratympanic fold; upper lips with three dark bars; ventral surface of thighs orangish, densely scattered with small yellow spots; dorsal parts of limbs, fingers and toes with black crossbars; ventral surface of fingers and toes orange; toes webbing yellowish-gray; the inside of lower arm with black stripe; ventral surface of all fingers discs, subarticular tubercles, supernumerary tubercles, and metacarpal tubercles orange, ventral surface of outer three toes discs, subarticular tubercles, and metatarsal tubercle grey; ventral surface of inner two toes discs orange; iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths, black reticulations throughout.
</paragraph>
</subSubSection>
<subSubSection id="92D1051AE6A76252E24F78202BF47F0B" pageId="0" pageNumber="343" type="color of holotype in preservative">
<paragraph id="E170DA2E62A0EA0CBB11C90FFBDE9625" pageId="0" pageNumber="343">Color of holotype in preservative.</paragraph>
<paragraph id="5690E2D821B5320C72BBA85E0C31B1C0" pageId="0" pageNumber="343">
For coloration of the holotype in preservative see Fig.
<figureCitation id="08C42E2C74F51E5982DD99DC8431125F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Dorsal view, B ventral view, C ventral view of hand, D ventral view of foot, (D) Amolops yangi sp. nov. (Holotype KIZ 038643) in preservative. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure5" httpUri="https://binary.pensoft.net/fig/1019898" pageId="0" pageNumber="343">5A-D</figureCitation>
. After two years of storage in ethanol, the dorsal surface fading to metallic blue with irregular black spots; black crossbars present on dorsal surfaces of limbs, fingers and toes still clear; throat, chest, belly, and ventral surface of limbs fading to cream-yellow, with irregular gray pigmentations; ventral surface of the hands and toes cream-yellow, digit tips and subarticular tubercles fading to cream-yellow or grayish-white.
</paragraph>
<caption id="CE1A98FA9A3CE73DEB92FF13A5CD8191" doi="10.3897/vz.74.e108013.figure5" httpUri="https://binary.pensoft.net/fig/1019898" pageId="0" pageNumber="343" start="Figure 5" startId="F5">
<paragraph id="D30B27E3FE5919E22F8F0E3CEF4B6DB3" pageId="0" pageNumber="343">
<emphasis id="6DDBA5A0A5A4A407864667279EC3DE21" bold="true" pageId="0" pageNumber="343">Figure 5.</emphasis>
A Dorsal view,
<emphasis id="A00260B68DFDB2D12C8838E170B1CDA7" bold="true" pageId="0" pageNumber="343">B</emphasis>
ventral view,
<emphasis id="42DE9A5ABB402E94A3A415FA6F53E38B" bold="true" pageId="0" pageNumber="343">C</emphasis>
ventral view of hand,
<emphasis id="4DB9042D51B121EE0BFA94BDC49B879D" bold="true" pageId="0" pageNumber="343">D</emphasis>
ventral view of foot, (
<emphasis id="0E7FA35EFCB9FBCAE6F450821BA4C5A9" bold="true" pageId="0" pageNumber="343">D</emphasis>
)
<taxonomicName id="6A599B9F93D71306E9F1EED2FE4D8D58" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="F27CCDA0F06DA22512795C2CF43DC4D1" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="2B4E3FA58736431E6C4D504056FCCA49" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
(Holotype KIZ 038643) in preservative. Photos by Zhong-Bin Yu.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7FA851DFE253F92FC1DD533FEF6DD1AC" pageId="0" pageNumber="343" type="description">
<paragraph id="363E47AD3602F351598583FF76153B46" pageId="0" pageNumber="343">Male secondary sexual characteristics.</paragraph>
<paragraph id="B58382052D8866CCEA781C72C64A2243" pageId="0" pageNumber="343">Adult males possess orange nuptial pads covering the base of first finger; absence of vocal sacs in males.</paragraph>
</subSubSection>
<subSubSection id="E1DEDCA012520ADF8574AE69AAAD89BB" pageId="0" pageNumber="343" type="distribution">
<paragraph id="BE6D0B0E8FD2A2A52A5DAB1083251BCC" pageId="0" pageNumber="343">Distribution and ecology.</paragraph>
<paragraph id="FF7A25CC88CBFBC90F16BDB6E580AD64" pageId="0" pageNumber="343">
<taxonomicName id="E801760306B052EA0B6DFAD577B464BD" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="DD456E42928A028BEE54367E624C65D9" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="51E8979559A4AFA74A6398BCDFBAB171" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
is currently known from two localities in the Gaoligong Mountains. These are Fugong and Lushui County, both in Yunnan Province, China. These two localities are separated by a straight-line distance of approximately 80 km. The new species inhabits the banks of rocky, fast-flowing streams or perches on shrubs (ca. 0.5 m above the ground) along the swift flowing streams (Fig.
<figureCitation id="D1189E584F28EC29A50F57B05B31EC7C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Amolops yangi sp. nov. (Holotype KIZ 038643). A Lateral view, B dorsal view, C ventral view of thighs, D ventral view of hand, E foot, F habitat. Photos by Zhong-Bin Yu." figureDoi="10.3897/vz.74.e108013.figure4" httpUri="https://binary.pensoft.net/fig/1019897" pageId="0" pageNumber="343">4F</figureCitation>
). The new species primarily inhabits high altitude mountainous areas with elevations ranging from 2500 to 3500 m. The breeding season is currently uncertain. Other sympatric amphibian species found in the same habitat included
<taxonomicName id="35479CBD809E26EB9811474906F4EF2E" baseAuthorityName="Ye in Sichuan Institute of Biology Herpetology Department" baseAuthorityYear="1977" class="Amphibia" family="Ranidae" genus="Nanorana" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanorana chayuensis" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="chayuensis">
<emphasis id="BDA5CC438107F0B4CA9B17EF3765F27F" italics="true" pageId="0" pageNumber="343">Nanorana chayuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2847FF99D9CC5B9574A584F1C8D7F09A" baseAuthorityName="Jiang" baseAuthorityYear="1983" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. viridimaculatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="viridimaculatus">
<emphasis id="013A0398744C2F3583812A7DB8016558" italics="true" pageId="0" pageNumber="343">A. viridimaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="69243C81B00A7E71F5A4FA03B023C278" authorityName="Yang &amp; Su In Yang, Su &amp; Li" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Scutiger" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scutiger gongshanensis" order="Lepidoptera" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="gongshanensis">
<emphasis id="1252E0F18313006B054BC323EB02EB5B" italics="true" pageId="0" pageNumber="343">Scutiger gongshanensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="D2D2FE111DC35EFA3AA4223BA9C0AC49" baseAuthorityName="Fei, Ye &amp; Huang" baseAuthorityYear="1990" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xenophrys glandulosa" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="11B513078BF70F6474393EACDEF4C207" italics="true" pageId="0" pageNumber="343">Xenophrys glandulosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEF63DB414AC698CD6206E597E91EE08" pageId="0" pageNumber="343" type="comparisons">
<paragraph id="DB47FE819FECAAA4B49BA92B2CFD477F" pageId="0" pageNumber="343">Comparisons.</paragraph>
<paragraph id="20819FA99310E2C3F1F1A4621776E7CA" pageId="0" pageNumber="343">
Phylogenetic analyses indicated that the new species belongs to the
<taxonomicName id="FED0FDB2FC2A49F41CECDD35F4133124" baseAuthorityName="Jiang" baseAuthorityYear="1983" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. viridimaculatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="viridimaculatus">
<emphasis id="02FBC83589A26C8F1D175E7AC25239C5" italics="true" pageId="0" pageNumber="343">A. viridimaculatus</emphasis>
</taxonomicName>
group with strong support. Geographically, the new species is found in Fugong and Lushui, Yunnan Province, China that belong to Gaoligong Mountains, and close to northern Myanmar. Therefore, we compared
<taxonomicName id="7A88514D2DE651A09A24CA761D863D26" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="4AC609BC4CAA5A74B54C4DB1A1888CDF" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="C1F5A02196A6C8DDB9F582C0B73D6F04" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
with morphologically, geographically, and molecularly similar species, which include
<taxonomicName id="6ACC60AC371230BED53F64DCEFD358A9" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. chayuensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="chayuensis">
<emphasis id="9852E624D5FEACCA5B6958D29219A3C6" italics="true" pageId="0" pageNumber="343">A. chayuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="79D8E5E0B5F5A8A0077AA6C53CFF00A2" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. bellulus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="bellulus">
<emphasis id="8CB0CBDF9671A535EFF972DBB8750392" italics="true" pageId="0" pageNumber="343">A. bellulus</emphasis>
</taxonomicName>
,
<taxonomicName id="FC21882964C2083D057BE4955C911E30" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. putaoensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="putaoensis">
<emphasis id="44D40BED3D4A21701733E4586E05F1DE" italics="true" pageId="0" pageNumber="343">A. putaoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5216A2D69A07F5266B51A26A9A52ADE7" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. binchachaensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="binchachaensis">
<emphasis id="EA06A6377BDA9123D815289FBF5421A5" italics="true" pageId="0" pageNumber="343">A. binchachaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F89E90A270F9ADA790BDEB6C7276DF70" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. deng" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="deng">
<emphasis id="BE0072E21EE1C65981C46DF33154477A" italics="true" pageId="0" pageNumber="343">A. deng</emphasis>
</taxonomicName>
,
<taxonomicName id="C8B74FB436E80E54E0E9FB632427A9B9" authorityName="Su, Yang &amp; Li" authorityYear="1986" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. jinjiangensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="jinjiangensis">
<emphasis id="D056690347C83BC915074DD51BB83A71" italics="true" pageId="0" pageNumber="343">A. jinjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B019AA9479AE24913B32BD55F520BC1B" baseAuthorityName="Jiang" baseAuthorityYear="1983" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. viridimaculatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="viridimaculatus">
<emphasis id="0CD4CA5D26CF481F96CEBA699442D204" italics="true" pageId="0" pageNumber="343">A. viridimaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="62D566273727CE1EDCE59F1DA6374C85" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. kaulbacki" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="kaulbacki">
<emphasis id="22E1B419B641BE4BCBB1A0C76B170105" italics="true" pageId="0" pageNumber="343">A. kaulbacki</emphasis>
</taxonomicName>
,
<taxonomicName id="85FEF4AAAD9CE5A054043A0698F542A8" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. marmoratus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="marmoratus">
<emphasis id="C5E0947FF6A27AC3DC76C05BB80EAD7F" italics="true" pageId="0" pageNumber="343">A. marmoratus</emphasis>
</taxonomicName>
,
<taxonomicName id="D3432DA94D3A7C29FB0650652625401B" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. afghanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="afghanus">
<emphasis id="C69505B32BE55DD5638E373A7906C5AA" italics="true" pageId="0" pageNumber="343">A. afghanus</emphasis>
</taxonomicName>
, and
<taxonomicName id="B5978AFC4A819FC77D9AA035A565058E" authorityName="Liu &amp; Yang" authorityYear="2000" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. tuberodepressus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="tuberodepressus">
<emphasis id="85848D6D9B9E32ED639A4381C13511CB" italics="true" pageId="0" pageNumber="343">A. tuberodepressus</emphasis>
</taxonomicName>
,
<taxonomicName id="DECF6AC051BA2F08E66B0AACA5BBB4ED" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. beibengensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="beibengensis">
<emphasis id="3E55C1B14C3204DDA6EDB9ACF97C4326" italics="true" pageId="0" pageNumber="343">A. beibengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5647D650DB623989B5DD90DAD725708C" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. wangyufani" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="wangyufani">
<emphasis id="AC35528F45095F2C080DAFEEB1B6BFE2" italics="true" pageId="0" pageNumber="343">A. wangyufani</emphasis>
</taxonomicName>
,
<taxonomicName id="80814E40A1A46D7D81D2A541252CD752" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. formosus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="873878D8CFC1FC5D2336DE92830FABF9" italics="true" pageId="0" pageNumber="343">A. formosus</emphasis>
</taxonomicName>
,
<taxonomicName id="9FCA1B58B6C512399E6F526E8275DDE0" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. medogensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="medogensis">
<emphasis id="5ABA49D73A616138F0D9354D8350AF1B" italics="true" pageId="0" pageNumber="343">A. medogensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDAA48BBC9A0FBB4FD7B696E55384C2" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pallasitatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="pallasitatus">
<emphasis id="34C470119A88E517C923259DC935F79E" italics="true" pageId="0" pageNumber="343">A. pallasitatus</emphasis>
</taxonomicName>
,
<taxonomicName id="6E7849F4BCF7833704F4542BF55CEB8F" authorityName="Biju, Mahony &amp; Kamei" authorityYear="2010" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. nidorbellus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="nidorbellus">
<emphasis id="AE6AC8F811A2A72F507D222281C0FA65" italics="true" pageId="0" pageNumber="343">A. nidorbellus</emphasis>
</taxonomicName>
,
<taxonomicName id="1B1844CF232360A11B34B77DE9F5411A" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. himalayanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis id="CD3267AC1218F0D6CD0AD2F716866FCC" italics="true" pageId="0" pageNumber="343">A. himalayanus</emphasis>
</taxonomicName>
,
<taxonomicName id="07820C715648A135AB4EF14E19E9109D" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. wangyali" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="wangyali">
<emphasis id="3D717CAD258484D5652F7F0CE5060D8E" italics="true" pageId="0" pageNumber="343">A. wangyali</emphasis>
</taxonomicName>
,
<taxonomicName id="DF70C04774E87F02DF02666CBBFB6BDB" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. longimanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="65AFA29DB3E15BAAC7122A8206F87DB5" italics="true" pageId="0" pageNumber="343">A. longimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="1D3416CAB14C3D734FDC745258757254" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ailao" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="ailao">
<emphasis id="D30B0FF54BBC4CD7B91BEA01ABA1127E" italics="true" pageId="0" pageNumber="343">A. ailao</emphasis>
</taxonomicName>
,
<taxonomicName id="D954517DCEF8FEDBEEF76AB86BBBB568" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. chanakya" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="chanakya">
<emphasis id="7E992DBFEB6C252A137AEB198223B746" italics="true" pageId="0" pageNumber="343">A. chanakya</emphasis>
</taxonomicName>
, and
<taxonomicName id="1333B5E5411B26FFF5BC5A35B9DFF982" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. tawang" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="tawang">
<emphasis id="6BC71354CB3BAA09A2A5B193B4D19E76" italics="true" pageId="0" pageNumber="343">A. tawang</emphasis>
</taxonomicName>
(
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<bibRefCitation id="A4487623E2A6B8CBB9A191238CFAC360" DOI="https://doi.org/10.26515/rzsi/v122/i3/2022/169417" author="Saikia, B" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="343" pagination="247 - 266" refId="B40" refString="Saikia, B, Laskar, MA, Dinesh, KP, Shabnam, A, Sinha, B, 2022. Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological ' Viridimaculatus species group'. Records of the Zoological Survey of India 122: 247 - 266, DOI: https://doi.org/10.26515/rzsi/v122/i3/2022/169417" title="Description of two new species of Amolops (Anura: Ranidae) from Arunachal Pradesh, Northeast India under the morphological ' Viridimaculatus species group'." url="https://doi.org/10.26515/rzsi/v122/i3/2022/169417" volume="122" year="2022">Saikia et al. 2022</bibRefCitation>
;
<bibRefCitation id="28C1DD78CEEA5CDAE402689B3587FA68" DOI="https://doi.org/10.1186/s40851-023-00214-9" author="Tang, S" journalOrPublisher="Zoological Letters" pageId="0" pageNumber="343" pagination="1 - 19" refId="B48" refString="Tang, S, Sun, T, Liu, S, Luo, S, Yu, G, Du, LN, 2023. A new species of cascade frog (Anura: Ranidae: Amolops) from central Yunnan, China. Zoological Letters 9: 1 - 19, DOI: https://doi.org/10.1186/s40851-023-00214-9" title="A new species of cascade frog (Anura: Ranidae: Amolops) from central Yunnan, China." url="https://doi.org/10.1186/s40851-023-00214-9" volume="9" year="2023">Tang et al. 2023</bibRefCitation>
).
</paragraph>
<paragraph id="9E595CB04599A73FAB670A8F6EF48A42" pageId="0" pageNumber="343">
<taxonomicName id="9CFE8575A5C396EE8D15A730682BF3E7" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="CE2A43A9E070169EF12D9F7FCE83AB8B" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="658D479BDB4F62E38FDF40AF67F92C43" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
is significantly different from
<taxonomicName id="C3B7AD8675DC37528EE5B73306C40C40" baseAuthorityName="Jiang" baseAuthorityYear="1983" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. viridimaculatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="viridimaculatus">
<emphasis id="FEF07BB695FD458F5F0CBC04E0EF3A91" italics="true" pageId="0" pageNumber="343">A. viridimaculatus</emphasis>
</taxonomicName>
by discontinuous glandular dorsolateral fold (vs. absent), supratympanic fold indistinct (vs. distinct), SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. 72.7-82.3 in males and 83.0-94.3 in females), dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots (vs. dorsum and flank with nearly round green or yellowish green spots, scattered with small green spots); from
<taxonomicName id="9582C34829F9DE9ADA08AC2C5C9D5165" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. kaulbacki" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="kaulbacki">
<emphasis id="C87E8C52D474F76048E04AC529A5E9F1" italics="true" pageId="0" pageNumber="343">A. kaulbacki</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 72.6-82.6 mm in males and 82.7-87.2 mm in females), discontinuous glandular dorsolateral fold (vs. absent), supratympanic fold indistinct (vs. distinct), iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths, black reticulations throughout (vs. eyes brownish black with scattered yellow spotting and yellow ring around iris); from
<taxonomicName id="45A220358AACB71C7F73B77E0DD2261A" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. beibengensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="beibengensis">
<emphasis id="459F14E5E1188AF81E0BCB71950E046C" italics="true" pageId="0" pageNumber="343">A. beibengensis</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 75.8 mm males and 90.2-93.2 mm in females), supratympanic fold indistinct (vs. distinct, wide and thick), discontinuous glandular dorsolateral fold from rear of eye to near vent (vs. absent); from
<taxonomicName id="420F8B96E0AC1D033C53F5B32F34CC6E" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. wangyufani" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="wangyufani">
<emphasis id="C608B5B3A2A558E0226898FAEB65A0E0" italics="true" pageId="0" pageNumber="343">A. wangyufani</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 68.3-69.0 mm males and 83.4 mm in female), vomerine teeth developed, on two short oblique between choanae, equal in distance from each other as to choanae (vs. vomerine teeth developed, the two rows are almost in touch), discontinuous glandular dorsolateral fold from rear of eye to near vent (vs. absent); from
<taxonomicName id="C34ED5FDA0CEA33D8FB2266F7F4A0088" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. medogensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="medogensis">
<emphasis id="0C4D2F264F8FCE980979EC5FB06CEC51" italics="true" pageId="0" pageNumber="343">A. medogensis</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 95.0 mm male and 72.4-96.9 mm in females), tibiotarsal articulation of adpressed limb reaching between nostrils and eyes (vs. beyond tip of snout), supratympanic fold indistinct (vs. distinct, wide and thick); from
<taxonomicName id="96D81103423932CA88FC65CB2DA2828F" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. himalayanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis id="F6028445B7F356A6BEA86CC6DA834894" italics="true" pageId="0" pageNumber="343">A. himalayanus</emphasis>
</taxonomicName>
by dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots (vs. dark brown, interspersed with irregular yellow spots), vocal sac absent in males (vs. externally visible vocal sacs present); from
<taxonomicName id="C2782BFE159396FFFF7C9AB1B325DD96" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. pallasitatus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="pallasitatus">
<emphasis id="DCDEE22F708F0D9D7686D4967AB707F4" italics="true" pageId="0" pageNumber="343">A. pallasitatus</emphasis>
</taxonomicName>
by SVL 51.5 mm in female (vs. SVL 70.6-72.3 mm in females), discontinuous glandular dorsolateral fold from rear of eye to near vent (vs. absent), dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots (vs. dorsum yellow-green, with irregular dark brown blotches without margins); from
<taxonomicName id="80AE75BA9D37F6D6B2E31A353B68EB87" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. wangyali" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="wangyali">
<emphasis id="30DDCC945EADC42A890E2B92D5D245E1" italics="true" pageId="0" pageNumber="343">A. wangyali</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 71.4-76.7 mm males and 80.5-89.6 mm in females), dorsal surface of the head, back, limbs, fingers, and toes green, interspersed with irregular black spots (vs. large brown irregularly shaped blotches on dorsum of head and body), rictal gland absent (vs. a distinct patch of rictal glands at rear of jaw on either side); from
<taxonomicName id="1611833AE667BB35D4C3FFEC1EFD960B" authorityName="Biju, Mahony &amp; Kamei" authorityYear="2010" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. nidorbellus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="nidorbellus">
<emphasis id="CB620F312519CE1994EB15F06E735C99" italics="true" pageId="0" pageNumber="343">A. nidorbellus</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 76.4-82.3 mm males and 85.4-98.0 mm in females), dorsum green, interspersed with irregular black spots (vs. dorsally brown with small irregularly arranged cobalt green spots), discontinuous glandular dorsolateral fold (vs. absent); from
<taxonomicName id="FFB1FB69637962367B1BBB4BEEC75474" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. longimanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="7B56F447F2151E646BB4B878801B77D1" italics="true" pageId="0" pageNumber="343">A. longimanus</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males and 51.5 mm in female (vs. SVL 30 mm), nostrils closer to anterior corner of eye than to tip of snout (vs. nostrils a litter nearer to tip of snout than the eye), above tympanum to the forelimb a thick glandular parotoid-like swelling absent (vs. present), eye diameter (ED/SL 63.0%) shorter than snout length (vs. snout about as long as the eye diameter, ED/SL 97.7%); from
<taxonomicName id="FA7AC7DBC712BF777BCF51FD904BE6A9" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. formosus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="4A50401A2E44303A4732B1976B41C45C" italics="true" pageId="0" pageNumber="343">A. formosus</emphasis>
</taxonomicName>
by dorsal parts of limbs, fingers and toes with black crossbars (vs. legs and toes with black white-dotted crossbars), inner metatarsal tubercle long, outer metatarsal tubercle absent (vs. metatarsal tubercle indistinct), dorsum green, interspersed with irregular black spots (vs. upper parts green, marbled with black, the black spots enclosing a number of small whitish dots); from
<taxonomicName id="63952A84604E295E3130D740EB01CD0E" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. chanakya" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="chanakya">
<emphasis id="FAB171658CA43D0BC7A2A7829261BCDD" italics="true" pageId="0" pageNumber="343">A. chanakya</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males (vs. SVL 76.4 mm), tympanum indistinct (vs. tympanum distinct, about 40% of eye length), absence of vocal sacs in males (vs. vocal sac externally visible), dorsal color green, interspersed with irregular black spots (vs. dorsal color dull brick-red, spotted with irregular cocoa-brown spots, these cocoa-brown spots enclosing a number of smaller dull brick-red spots); from
<taxonomicName id="71C9387077FA1366A77D983EA6A9FD6F" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. tawang" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="tawang">
<emphasis id="3ECBC6D8FB793F8F3AF3EB4728151369" italics="true" pageId="0" pageNumber="343">A. tawang</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males (vs. SVL 82.5 mm), tibiotarsal articulation of adpressed limb reaching between nostrils and eyes (vs. reaching up to snout), dorsal color green, interspersed with irregular black spots (vs. dorsal color olivegreen, spotted with large, irregular shaped dark-brown spots, brown spots enclosing a number of small olivegreen dots).
</paragraph>
<paragraph id="28FF8990F64543D4AD0FB0ED6E1EB835" pageId="0" pageNumber="343">
<taxonomicName id="D62A9B0E0DFB3BC1AA7A37011B49DBBE" authorityName="Wu, Yu, Lu, Yuan &amp; Che" authorityYear="2024" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops yangi" order="Anura" pageId="0" pageNumber="343" phylum="Chordata" rank="species" species="yangi">
<emphasis id="C7DF39822EC0956AB3627D637A9854E9" italics="true" pageId="0" pageNumber="343">Amolops yangi</emphasis>
</taxonomicName>
<emphasis id="E0046D2C5C5F5FEB718120BF8E1146D5" bold="true" pageId="0" pageNumber="343">sp. nov.</emphasis>
is significantly different from
<taxonomicName id="511F16BFCEF1D2C654A38B9C63D81FF4" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. chayuensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="chayuensis">
<emphasis id="E3C111882CDBB571702125CB3BCC8BFA" italics="true" pageId="0" pageNumber="343">A. chayuensis</emphasis>
</taxonomicName>
by absence of vocal sacs in males (vs. pair of external subgular vocal sacs); discontinuous glandular dorsolateral fold from rear of eye to near vent (vs. dorsolateral fold prominent); upper part of flanks green, lower part of flanks green-yellow (vs. upper part of flanks brown, lower part of flanks light green or white with dark brown blotches); from
<taxonomicName id="C5A5122FB12859072F87FB8BB276ADEA" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. bellulus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="bellulus">
<emphasis id="E1632C8F82E24A613B55B7A84A7647AC" italics="true" pageId="0" pageNumber="343">A. bellulus</emphasis>
</taxonomicName>
by tympanum indistinct (vs. tympanum distinct), iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths, black reticulations throughout (vs. upper half of iris golden yellow with some irregular brown spots, lower half dark brown); from
<taxonomicName id="5C430BC0DFBD0D1F26AD5D5B275E43C5" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. putaoensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="putaoensis">
<emphasis id="F6A8E7749AA1484431D4BAFCD14236A3" italics="true" pageId="0" pageNumber="343">A. putaoensis</emphasis>
</taxonomicName>
by SVL of adult male 46.3-51.8 mm (vs. 37.6-40.2 mm), rictal gland absent (vs. two rictal glands present), iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths, black reticulations throughout (vs. upper one-fourth of iris bronze with black reticulations, lower three-fourths dark), absence of vocal sacs in males (vs. pair of internal subgular vocal sacs present); from
<taxonomicName id="269263BDE244E656258F3CFCA07BABD0" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. binchachaensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="binchachaensis">
<emphasis id="CD3174F83507B72CDFDEB978E385CBA7" italics="true" pageId="0" pageNumber="343">A. binchachaensis</emphasis>
</taxonomicName>
by SVL of adult female 51.5 mm (vs. 65.0 mm), tympanum indistinct, relatively small, about one third of eye diameter (vs. tympanum big), supratympanic fold indistinct (vs. supratympanic fold absent), dorsal surface of the head, back, limbs, fingers, and toes green (vs. light yellow); from
<taxonomicName id="162A91257826ACC6B8C5A4749770B491" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. deng" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="deng">
<emphasis id="2A4B3549A0D1011689F513B33A831AA8" italics="true" pageId="0" pageNumber="343">A. deng</emphasis>
</taxonomicName>
by SVL of adult female 51.5 mm (vs. 68.5-72.0 mm), tympanum indistinct, relatively small, about one third of eye diameter (vs. tympanum distinct, slightly less than half of eye diameter), tibiotarsal articulation of adpressed limb reaching between nostrils and eyes (vs. tibiotarsal articulation of adpressed limb reaching beyond tip of snout); from
<taxonomicName id="018141E46D4F752B3F00C5B94F705F32" authorityName="Su, Yang &amp; Li" authorityYear="1986" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. jinjiangensis" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="jinjiangensis">
<emphasis id="019030FDB2FB2676879DF1C103018BE6" italics="true" pageId="0" pageNumber="343">A. jinjiangensis</emphasis>
</taxonomicName>
by supratympanic fold indistinct (vs. distinct), dorsal surface of head, body, limbs, fingers, toes and flank of body relatively smooth (vs. rough with tubercles), the absence of a pair of large tubercles on sides of cloaca (vs. present); from
<taxonomicName id="C1CB200EAFE3C9AAC3119987540C8AE6" authorityName="Liu &amp; Yang" authorityYear="2000" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. tuberodepressus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="tuberodepressus">
<emphasis id="808B9909E035FFBD272288D5DBD66C1A" italics="true" pageId="0" pageNumber="343">A. tuberodepressus</emphasis>
</taxonomicName>
by supratympanic fold indistinct (vs. present, wide),
<normalizedToken id="1A313CF8D4B099EE19BF5D790FD4F428" originalValue="flanks">flanks</normalizedToken>
smooth (vs. with
<normalizedToken id="67D45DFAC5B5D1326EFABCF5AB6F24C5" originalValue="flatter">flatter</normalizedToken>
tubercles), relative length of fingers I&lt;II&lt;IV&lt;III (vs. II&lt;IV&lt;I&lt;III), vomerine teeth developed (vs. weak); from
<taxonomicName id="E9E0BC1626EFFF9CFEEBF33DC9049D5A" authorityName="Duméril In C. Duméril &amp; A. Duméril" authorityYear="1851" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ailao" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="ailao">
<emphasis id="9356CBBEF7510BE59A5635D2857DD9A2" italics="true" pageId="0" pageNumber="343">A. ailao</emphasis>
</taxonomicName>
by SVL 46.3-51.8 mm in males (vs. SVL 33.0-35.1 mm in males), vomerine teeth developed (vs. absent), tibiotarsal articulation of adpressed limb reaching between nostrils and eyes (vs. beyond anterior corner of eye), iris distinctly bicolored, golden-yellow in upper one-fourth and reddish brown in lower three-fourths (vs. iris light brown with dark wash); from
<taxonomicName id="961C1C0ECA183A5607A089B80B1DA81A" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. marmoratus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="marmoratus">
<emphasis id="62415EC3FCBA4BFC6C58ECECC2AA8EAB" italics="true" pageId="0" pageNumber="343">A. marmoratus</emphasis>
</taxonomicName>
by tibiotarsal articulation of adpressed limb reaching between nostrils and eyes (vs. beyond tip of snout), discontinuous glandular dorsolateral fold from rear of eye to near vent (vs. distinct dorsolateral fold absent), rictal gland absent (vs. multiple small globular rictal glands on right side, single rictal gland on left side, just posterior to jaw); from
<taxonomicName id="D53F6FD97A549AF52268752B21F18AFD" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. afghanus" pageId="0" pageNumber="343" phylum="Arthropoda" rank="species" species="afghanus">
<emphasis id="4871AE0BA6E2ABAC004E7E3CE9D6085A" italics="true" pageId="0" pageNumber="343">A. afghanus</emphasis>
</taxonomicName>
by SVL 51.5 mm in female (vs. SVL 67.7-94.1 mm in females), rictal gland absent (vs. indistinct rictal glands present on one side of head at posterior end of jaw), absence of vocal sacs in males (vs. males with dual gular pouches).
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="4AF5647AC2D6F0E821F8214609874A2B" ID-CoL="38478" ID-ENA="2923130" authority="Foerster, 1868" authorityName="Foerster" authorityYear="1868" class="Insecta" family="Ichneumonidae" genus="Dusona" kingdom="Animalia" lsidName="Dusona blanda" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="blanda">
Dusona blanda (
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<taxonomicName id="39DE4B2777856837BAC6AA063D465809" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" lsidName="Campoplex blandus" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="blandus">Campoplex blandus</taxonomicName>
<normalizedToken id="901BEB7676F96AC311E20B8163ECC285" originalValue="Förster">Foerster</normalizedToken>
, 1868
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<taxonomicName id="FCDD7010352662F253988353F753C867" class="Insecta" family="Ichneumonidae" genus="Ichneumonoidea" kingdom="Animalia" lsidName="Ichneumonoidea remota" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="remota">remota</taxonomicName>
(
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, 1868,
<taxonomicName id="D0110616F3191648C468D049FC6DCA7A" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" lsidName="Campoplex" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">Campoplex</taxonomicName>
)
</paragraph>
<paragraph id="6C3EEB9F8D8663FAD1FDDE4005D5130D" pageId="0" pageNumber="9042">
<taxonomicName id="F53B0CA4AABAD18621F803FA7A21EE99" class="Insecta" family="Ichneumonidae" genus="Ichneumonoidea" kingdom="Animalia" lsidName="Ichneumonoidea forsselli" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="forsselli">forsselli</taxonomicName>
(Holmgren, 1872,
<taxonomicName id="6C7B081AAD6AEDC3093C2EFF78394954" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" lsidName="Campoplex" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">Campoplex</taxonomicName>
)
</paragraph>
<paragraph id="FDA243DB7A3BE6BD61CE0C49E0CEC1D0" pageId="0" pageNumber="9042">
<taxonomicName id="0196020EBAECD83566203820BF27ACF4" class="Insecta" family="Ichneumonidae" genus="Ichneumonoidea" kingdom="Animalia" lsidName="Ichneumonoidea punctiventris" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="punctiventris">punctiventris</taxonomicName>
(Woldstedt, 1877,
<taxonomicName id="7BB59F5E19C95E037ED45662836C26E7" class="Insecta" family="Ichneumonidae" genus="Casinaria" kingdom="Animalia" lsidName="Casinaria" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">Casinaria</taxonomicName>
)
</paragraph>
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<paragraph id="83F146AC0DDC2F85E37865D3DC39CD4E" pageId="0" pageNumber="9042">Distribution</paragraph>
<paragraph id="C61337BAA95F92F91325A0EF33181577" pageId="0" pageNumber="9042">England, Scotland, Isle of Man</paragraph>
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Genus
<taxonomicName id="4634330DD165F3C50B10D967C25FA464" ID-CoL="89M8" ID-ENA="58758" LSID="0C032E95-3A93-5168-AEE3-4785094E5A8C" authority="Foerster, 1863" authorityName="Foerster" authorityYear="1863" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Xynobius Foerster, 1863</taxonomicName>
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<taxonomicName id="8F5A7E951568F673E5E88DF01A878DEA" authorityName="Foerster" authorityYear="1863" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Xynobius</taxonomicName>
Foerster, 1863: 235. Type species (by original designation):
<taxonomicName id="32171E3979080A7DD013BDFB8C6E8B9D" authorityName="Forster" authorityYear="1862" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius pallipes" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="pallipes">Xynobius pallipes</taxonomicName>
Foerster, 1863 (=
<taxonomicName id="DA4D25E5A25A837FA948C04DD7D7B67E" authorityName="Haliday" authorityYear="1837" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius caelatus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="caelatus">Opius caelatus</taxonomicName>
Haliday, 1837).
</paragraph>
<paragraph id="9DA3D6471219A77F5242DB8EAD2CC1B4" pageId="0" pageNumber="219">
<taxonomicName id="1FB7ED8E475D9D5552A96094DED5A7DC" authorityName="Foerster" authorityYear="1862" class="Insecta" family="Braconidae" genus="Aclisis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aclisis" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Aclisis</taxonomicName>
Foerster, 1863: 267. Type species (by original designation):
<taxonomicName id="2628A68A8DC7A8F1E695234E78305FC4" authorityName="Foerster" authorityYear="1862" class="Insecta" family="Braconidae" genus="Aclisis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aclisis isomera" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="isomera">Aclisis isomera</taxonomicName>
Foerster, 1863 (=
<taxonomicName id="8B17F366323D87843054A9E8BA7E8320" authorityName="Haliday" authorityYear="1837" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius caelatus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="caelatus">Opius caelatus</taxonomicName>
Haliday, 1837). Synonymised by
<bibRefCitation id="74F75632B63228FFD1438E8BEB29132D" author="Fischer, M" journalOrPublisher="fuer Zoologie und Botanik am Landesmuseum Joanneum" pageId="0" pageNumber="219" refId="B4" refString="Fischer, M, 1972. HymenopteraBraconidae (Opiinae I). (Palaearktische Region). Das Tierreich.91(1973): 1-620." title="HymenopteraBraconidae (Opiinae I). (Palaearktische Region). Das Tierreich. 91 (1973): 1 - 620." year="1972">Fischer (1972)</bibRefCitation>
.
</paragraph>
<paragraph id="3182A1B1FC7D58E32EE44B61C8D915F1" pageId="0" pageNumber="219">
<taxonomicName id="01246398FC2A6857A4D7D49944B3D58B" class="Insecta" family="Braconidae" genus="Holconotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holconotus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Holconotus</taxonomicName>
Foerster, 1863: 259 (not
<normalizedToken id="1D38255ACD7EABC87FA3264E705B0821" originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
, 1846). Type species (by original designation):
<taxonomicName id="C78BFC1BB7E12AA8A3F6DDC18C0251EA" authorityName="Wesmael" authorityYear="1835" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius comatus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="comatus">Opius comatus</taxonomicName>
Wesmael, 1835). Synonymised by
<bibRefCitation id="162CAEB6B84AA9B9729896AC864BF249" author="van Achterberg, C" journalOrPublisher="Zooelogische Mededeelingen" pageId="0" pageNumber="219" pagination="313 - 329" refId="B25" refString="van Achterberg, C, 2004. New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae). Zooelogische Mededeelingen 78: 313 - 329" title="New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae)." volume="78" year="2004">van Achterberg (2004)</bibRefCitation>
.
</paragraph>
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<taxonomicName id="9E1AAAB203A27B4E9062E7BD388BF866" authorityName="Ashmead" authorityYear="1900" class="Insecta" family="Braconidae" genus="Aulonotus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aulonotus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Aulonotus</taxonomicName>
Ashmead, 1900: 368 (new name for
<taxonomicName id="6DFC12F9E0F90C3963F7C6E10FC834D1" class="Insecta" family="Braconidae" genus="Holconotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holconotus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Holconotus</taxonomicName>
Foerster). Type species (by original designation):
<taxonomicName id="567BC56ADC25AB820EEA232682F0CCD0" authorityName="Wesmael" authorityYear="1835" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius comatus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="comatus">Opius comatus</taxonomicName>
Wesmael, 1835). Synonymised by
<bibRefCitation id="BE0A73B0BBD27B7B0101EEE5D9C89C78" author="Tobias, VI" editor="Medvedev, GS" journalOrPublisher="Opr. Faune SSSR 147 (3) 5" pageId="0" pageNumber="219" pagination="7 - 231" refId="B20" refString="Tobias, VI, Jakimavicius, A, 1986. Alysiinae and Opiinae. In: Medvedev, GS, Ed., Opredelitel Nasekomych Evrospeiskoi Tsasti SSSR 3, Peredpontdatokrylye 4. Opr. Faune SSSR 147(3)5: 7 - 231" title="Alysiinae and Opiinae." volumeTitle="Opredelitel Nasekomych Evrospeiskoi Tsasti SSSR 3, Peredpontdatokrylye 4." year="1986">
Tobias and
<normalizedToken id="B451E7019AD1862C78748E21AAF80F8E" originalValue="Jakimavičius">Jakimavicius</normalizedToken>
(1986)
</bibRefCitation>
.
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<paragraph id="90A0396CE92AB89CD55F158EB07CEC27" pageId="0" pageNumber="219">
<taxonomicName id="AD514CBD0BCBBA7F299C45F93B67F745" authorityName="Viereck" authorityYear="1913" class="Insecta" family="Braconidae" genus="Eristernaulax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eristernaulax" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Eristernaulax</taxonomicName>
Viereck, 1913: 362. Type species (by original designation):
<taxonomicName id="CE79DB065EF24AE344B60888D9C52658" class="Insecta" family="Braconidae" genus="Eristernaulax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eristernaulax leucotaenia" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="leucotaenia">Eristernaulax leucotaenia</taxonomicName>
Viereck, 1913). Synonymised by
<bibRefCitation id="A3EE3221FC07ABF06A5B3D3A9628114B" author="van Achterberg, C" journalOrPublisher="Zooelogische Mededeelingen" pageId="0" pageNumber="219" pagination="313 - 329" refId="B25" refString="van Achterberg, C, 2004. New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae). Zooelogische Mededeelingen 78: 313 - 329" title="New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae)." volume="78" year="2004">van Achterberg (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="0B70550792EA5D1C37B69DA9DC768364" pageId="0" pageNumber="219">
<taxonomicName id="37DEC220A0DA36F05180930E351BB8E6" authorityName="Fischer" authorityYear="1987" class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
Fischer, 1984: 610, 611 (as subgenus of
<taxonomicName id="DAFFCCDEF79969A45C3D8200C78CFF7A" authorityName="Wesmael" authorityYear="1835" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Opius</taxonomicName>
Wesmael), 1998: 25 (key to species); Wharton, 1988: 356; 2006: 338 (as subgenus of
<taxonomicName id="97479AB5F71ADA4A56601CE4B98530F8" class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
Foerster, 1863; possible paraphyly in
<taxonomicName id="8DED329719B2079D423C1D900A6B615D" authorityName="Foerster" authorityYear="1863" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Xynobius</taxonomicName>
). Type species (by original designation):
<taxonomicName id="ADF49D6DEE887FFF3CBEA18BE588A9BB" authorityName="Thomson" authorityYear="1895" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius macrocerus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="macrocerus">Opius macrocerus</taxonomicName>
Thomson, 1895. Synonymised by
<bibRefCitation id="475BFE3B3E94E2CD21CA9897CD0C9C46" author="van Achterberg, C" journalOrPublisher="Zooelogische Mededeelingen" pageId="0" pageNumber="219" pagination="313 - 329" refId="B25" refString="van Achterberg, C, 2004. New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae). Zooelogische Mededeelingen 78: 313 - 329" title="New Indo-Australian subgenera and species of the genera Xynobius Foerster and Ademoneuron Fischer (Hymenoptera: Braconidae: Opiinae)." volume="78" year="2004">van Achterberg (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="904570DFF89C23FB8A851642E6D8F64B" pageId="0" pageNumber="219">
<taxonomicName id="2246917A620A48FF5E6C4E235A044326" authorityName="Fischer" authorityYear="1998" class="Insecta" family="Braconidae" genus="Xynobiotenes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xynobiotenes" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Xynobiotenes</taxonomicName>
Fischer, 1998: 23 (as subgenus of
<taxonomicName id="F7E8159F7F7408BB488F965D149978B4" class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
Foerster, 1863). Type species (by original designation):
<taxonomicName id="4B580F6B9E07722D97AFC0B7AC602F2C" authorityName="Fischer" authorityYear="1962" class="Insecta" family="Braconidae" genus="Opius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opius scutellatus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="scutellatus">Opius scutellatus</taxonomicName>
Fischer, 1962. Synonymised by
<bibRefCitation id="4932EC00AA809B41DD52D1512C071E11" DOI="https://doi.org/10.3897/zookeys.326.5911" author="Li, X-Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="219" pagination="1 - 186" refId="B9" refString="Li, X-Y, van, Achterberg C, Tan, J-C, 2013. Revision of the subfamily Opiinae (Hymenoptera, Braconidae) from Hunan (China), including thirty-six new species and two new genera. ZooKeys 268: 1 - 186, DOI: https://doi.org/10.3897/zookeys.326.5911" title="Revision of the subfamily Opiinae (Hymenoptera, Braconidae) from Hunan (China), including thirty-six new species and two new genera." url="https://doi.org/10.3897/zookeys.326.5911" volume="268" year="2013">Li et al. (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A9C6A895C1C39283EECE60E7419BEE89" pageId="0" pageNumber="219" type="diagnosis">
<paragraph id="88C3D6E2984B64F07BD28864EAD1365D" pageId="0" pageNumber="219">Diagnosis.</paragraph>
<paragraph id="D798096AF7198D3F866609F1D52E03BA" pageId="0" pageNumber="219">
Hypoclypeal depression distinct and ventral margin of clypeus above upper level of mandibular condyles (Figs
<figureCitation id="9D49CBAABCF53E7423BCA592B554207C" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">6</figureCitation>
,
<figureCitation id="1088F2074F1984376CE18562A9BBF0FB" captionStart="Figures 1424" captionStartId="F4" captionText="Figures 14 - 24. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 14 wings 15 mesosoma, lateral view 16 mesosoma, dorsal view 17 metasoma, dorsal view 18 propodeum, dorsal view 19 hypopygium, ventral view 20 head, anterior view 21 head, dorsal view 22 1 st- 3 rd metasomal tergites, dorsal view 23 antenna 24 ovipositor and its sheath, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures14-24" httpUri="https://binary.pensoft.net/fig/997929" pageId="0" pageNumber="219">20</figureCitation>
,
<figureCitation id="7BB7863BC1045DB53989613052ADF3FC" captionStart="Figures 3041" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">34</figureCitation>
,
<figureCitation id="1D3B36C1D834C5994C7BDE980E682609" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">48</figureCitation>
,
<figureCitation id="52D9B71C26D992F539786E08028AD795" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">58</figureCitation>
,
<figureCitation id="C310ADE716AC003CB4AFC860DE1D3ABF" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">70</figureCitation>
); mandible without acute basal lamella; notauli complete (Fig.
<figureCitation id="E396A515E3886E9F2C0F3C0789FD0A95" captionStart="Figures 3041" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
) or largely absent except a pair of short anterior impressions (Figs
<figureCitation id="307EEFD63368AD6F83002BB84F603326" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">4</figureCitation>
,
<figureCitation id="A3AAA0F62FE4A65B896A7C5856B04ED2" captionStart="Figures 1424" captionStartId="F4" captionText="Figures 14 - 24. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 14 wings 15 mesosoma, lateral view 16 mesosoma, dorsal view 17 metasoma, dorsal view 18 propodeum, dorsal view 19 hypopygium, ventral view 20 head, anterior view 21 head, dorsal view 22 1 st- 3 rd metasomal tergites, dorsal view 23 antenna 24 ovipositor and its sheath, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures14-24" httpUri="https://binary.pensoft.net/fig/997929" pageId="0" pageNumber="219">16</figureCitation>
,
<figureCitation id="80797BBBC369D09E4F03328DC31E2292" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">57</figureCitation>
); medio-posterior depression variable, often present (Figs
<figureCitation id="53F34092E2A13683C4482BFC197C0038" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">4</figureCitation>
,
<figureCitation id="1DD6D60B98E529CB32B0B36113591ADB" captionStart="Figures 3041" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
,
<figureCitation id="F33895F9E4A2FCE0F5DEA9D0796966EA" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">45</figureCitation>
,
<figureCitation id="0FBA75B34F0637DB87F922F322B494A6" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">57</figureCitation>
,
<figureCitation id="3673D7D6906BC9C6636B9D04C00F10F3" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">67</figureCitation>
); precoxal sulcus distinct and no sternaulus; vein r of fore wing more or less angled with vein 3-SR and distinctly shorter than vein 2-SR (Figs
<figureCitation id="780894F2814AA7B574A070D1ADE7BDC3" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">2</figureCitation>
,
<figureCitation id="EFA9300F0DB2247CD697B113A058408F" captionStart="Figures 1424" captionStartId="F4" captionText="Figures 14 - 24. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 14 wings 15 mesosoma, lateral view 16 mesosoma, dorsal view 17 metasoma, dorsal view 18 propodeum, dorsal view 19 hypopygium, ventral view 20 head, anterior view 21 head, dorsal view 22 1 st- 3 rd metasomal tergites, dorsal view 23 antenna 24 ovipositor and its sheath, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures14-24" httpUri="https://binary.pensoft.net/fig/997929" pageId="0" pageNumber="219">14</figureCitation>
,
<figureCitation id="CCD73AF4DF8A0D3A5D5573E3A0175A9E" captionStart="Figures 3041" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">30</figureCitation>
,
<figureCitation id="0C3AA593FA193FF9018E6C1D78AEBC09" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">43</figureCitation>
,
<figureCitation id="95E413168FA3030B9915CBE76B0E27C6" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">55</figureCitation>
,
<figureCitation id="13BF5659F00ADCFC67E5A4C5B1818C26" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">65</figureCitation>
); pterostigma either narrowed apically or parallel-sided to slightly widened apically (subgenus
<taxonomicName id="B715E6E38DC80E2AFDD25CE12460E887" class="Insecta" family="Braconidae" genus="Xynobiotenes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xynobiotenes (Stigmatopoea)" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="subGenus" subGenus="Stigmatopoea">Xynobiotenes Stigmatopoea</taxonomicName>
: Figs
<figureCitation id="09CACA33563A55202AB503305EAA3E85" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">55</figureCitation>
,
<figureCitation id="8029756A94E6E4985976918C442D2ED6" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">65</figureCitation>
); dorsope distinct (Figs
<figureCitation id="7B6F8D54E2E6E7D84C0E69A60A8500C9" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">5</figureCitation>
,
<figureCitation id="851FEDDBA99807DAFE7DB902A73A9B27" captionStart="Figures 212" captionStartId="F2" captionText="Figures 2 - 12. Xynobius azonius Han &amp; van Achterberg sp. nov., holotype, ♀ 2 wings 3 mesosoma, lateral view 4 mesosoma, dorsal view 5 metasoma, dorsal view 6 head, anterior view 7 head, dorsal view 8 propodeum, dorsal view 9 1 st metasomal tergite, dorsal view 10 ovipositor and sheath, latero-ventral view 11 hind leg 12 antenna." figureDoi="10.3897/zookeys.1193.115831.figures2-12" httpUri="https://binary.pensoft.net/fig/997927" pageId="0" pageNumber="219">9</figureCitation>
,
<figureCitation id="6F97014162ABFCB4B5A5E6F699BF2AA6" captionStart="Figures 1424" captionStartId="F4" captionText="Figures 14 - 24. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 14 wings 15 mesosoma, lateral view 16 mesosoma, dorsal view 17 metasoma, dorsal view 18 propodeum, dorsal view 19 hypopygium, ventral view 20 head, anterior view 21 head, dorsal view 22 1 st- 3 rd metasomal tergites, dorsal view 23 antenna 24 ovipositor and its sheath, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures14-24" httpUri="https://binary.pensoft.net/fig/997929" pageId="0" pageNumber="219">22</figureCitation>
,
<figureCitation id="1C6E14486ADE624D42E31D881E40F15A" captionStart="Figures 2528" captionStartId="F5" captionText="Figures 25 - 28. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 25 1 st metasomal tergite, dorsal view 26 1 st metasomal tergite, dorso-posterior view 27 mandible, lateral view 28 hind leg, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures25-28" httpUri="https://binary.pensoft.net/fig/997930" pageId="0" pageNumber="219">25</figureCitation>
,
<figureCitation id="A7B51263CBAAF8E1642976C098EEE61C" captionStart="Figures 2528" captionStartId="F5" captionText="Figures 25 - 28. Xynobius brevifemora Han &amp; van Achterberg, sp. nov., holotype, ♀ 25 1 st metasomal tergite, dorsal view 26 1 st metasomal tergite, dorso-posterior view 27 mandible, lateral view 28 hind leg, lateral view." figureDoi="10.3897/zookeys.1193.115831.figures25-28" httpUri="https://binary.pensoft.net/fig/997930" pageId="0" pageNumber="219">26</figureCitation>
,
<figureCitation id="917E6B8B5F8816F6BF47DE8BD7AC6927" captionStart="Figures 3041" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">37</figureCitation>
,
<figureCitation id="7B65CC7BAAE66BA67713D4B2E234006A" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">45</figureCitation>
,
<figureCitation id="5413E15841FF7144A1A87D6A33232340" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">46</figureCitation>
,
<figureCitation id="56D992189A1F19C4F6E6DA905E889A8D" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">50</figureCitation>
,
<figureCitation id="19EA072AB3154F7FE6E99A965CD6DF7A" captionStart="Figures 4353" captionStartId="F9" captionText="Figures 43 - 53. Xynobius stipitoides Han &amp; van Achterberg, sp. nov., holotype, ♀ 43 wings 44 mesosoma, lateral view 45 mesosoma, dorsal view 46 metasoma, dorsal view 47 head, anterior view 48 head, dorsal view 49 head, lateral view 50 propodeum, dorsal view 51 1 st metasomal tergite, dorsal view 52 antenna 53 hind leg and ovipositor, lateral view. The red arrow points to the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures43-53" httpUri="https://binary.pensoft.net/fig/997934" pageId="0" pageNumber="219">51</figureCitation>
,
<figureCitation id="01D84C331BCFDE25E831A5A7EB64304B" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">57</figureCitation>
,
<figureCitation id="D54F95DC736E3903359474A0B5C9C804" captionStart="Figures 5564" captionStartId="F11" captionText="Figures 55 - 64. Xynobius geniculatus (Thomson), ♀, South Korea 55 wings 56 mesosoma, lateral view 57 mesosoma, dorsal view 58 1 st metasomal tergite, dorsal view 59 metasoma, dorsal view 60 head, anterior view 61 head and pronotum, dorsal view 62 propodeum, dorsal view 63 ovipositor and sheath, latero-ventral view 64 antenna. The red arrow indicates the dorsope." figureDoi="10.3897/zookeys.1193.115831.figures55-64" httpUri="https://binary.pensoft.net/fig/997936" pageId="0" pageNumber="219">58</figureCitation>
,
<figureCitation id="D83A0708AE455CE9645648BD45D7C2DC" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">68</figureCitation>
,
<figureCitation id="C9C250F91493BD5432E38E4309EFBEB6" captionStart="Figures 6575" captionStartId="F12" captionText="Figures 65 - 75. Xynobius stipitatus (Tobias), holotype, ♀, Russia 65 wings 66 mesosoma lateral 67 mesosoma dorsal 68 metasoma dorsal 69 propodeum and 1 st metasomal tergite latero-dorsal 70 head anterior 71 head and pronotum dorsal 72 head latero-anterior 73 ovipositor sheath lateral 74 hind leg 75 antenna. Photographs: Konstantin Samartsev." figureDoi="10.3897/zookeys.1193.115831.figures65-75" httpUri="https://binary.pensoft.net/fig/997937" pageId="0" pageNumber="219">69</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5CE7B12226EEC5BAB1150F6C1BF7341B" pageId="0" pageNumber="219" type="distribution">
<paragraph id="47949EC452AAF8FD6B0A205BD534D970" pageId="0" pageNumber="219">Distribution.</paragraph>
<paragraph id="D064F194A6BB227F8FDA97F6A5BD1F5A" pageId="0" pageNumber="219">Cosmopolitan.</paragraph>
</subSubSection>
<subSubSection id="D79669ED87CF5D4D25CD3CD01868FA97" pageId="0" pageNumber="219" type="biology_ecology">
<paragraph id="B3D34A70B8ACEF5E3DD0488F6642BCBC" pageId="0" pageNumber="219">Biology.</paragraph>
<paragraph id="DFBE21FEEA0F17E2C04FC77A7F0FB296" pageId="0" pageNumber="219">
Koinobionts endoparasitoids of mining dipterous larvae (species of the genus
<taxonomicName id="09DCEF7B8511B70C652DA9F9D711D782" class="Insecta" family="Agromyzidae" genus="Agromyza" kingdom="Animalia" lsidName="Agromyza" order="Diptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="6186FC6D6B85F913DDB5E516F65B9C29" italics="true" pageId="0" pageNumber="219">Agromyza</emphasis>
</taxonomicName>
Fallen, 1810;
<taxonomicName id="F54870B92055E8003BDEE7761B577FC7" class="Insecta" family="Agromyzidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="219" rank="family">Agromyzidae</taxonomicName>
), or of fruit-infesting larvae (species of the genera
<taxonomicName id="4B495E39CCAAFD9DBC442F164CFD88A4" genus="Euliea" lsidName="Euliea" pageId="0" pageNumber="219" rank="genus">
<emphasis id="FCA0946283226BE0684A756A923D4CF4" italics="true" pageId="0" pageNumber="219">Euliea</emphasis>
</taxonomicName>
Walker, 1835, and
<taxonomicName id="2837E675AA493A97F7FAF32334DF2D07" class="Insecta" family="Tephritidae" genus="Trypeta" kingdom="Animalia" lsidName="Trypeta" order="Diptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="A535A7C4E8A411ECBE3FA9725CBFE694" italics="true" pageId="0" pageNumber="219">Trypeta</emphasis>
</taxonomicName>
Meigen, 1803;
<taxonomicName id="29A8368A12420C41A5C49151637C80B2" class="Hexapoda" family="Tephritidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
).
</paragraph>
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</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:namePart>Hazir, Canan</mods:namePart>
<mods:affiliation>Adnan Menderes University, Health Services Vocational College, 09100 Aydin, Turkey</mods:affiliation>
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<mods:namePart>Keskin, Nevin</mods:namePart>
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<mods:namePart>Scheuchl, Erwin</mods:namePart>
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<taxonomicName LSID="http://species-id.net/wiki/Andrena_sericata" authority="Imhoff, 1866" authorityName="Imhoff" authorityYear="1866" class="Hexapoda" family="Andrenidae" genus="Andrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Andrena sericata" order="Hymenoptera" pageId="23" pageNumber="82" phylum="Arthropoda" rank="species" species="sericata">Andrena sericata Imhoff, 1866</taxonomicName>
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<paragraph pageId="23" pageNumber="82">Distribution in Turkey.</paragraph>
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, Antalya (
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).
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<paragraph pageId="23" pageNumber="82">Material examined.</paragraph>
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:
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,
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,
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, 38 m, 23.II.2006, 1 ♀, 5 ♂♂,
<normalizedToken originalValue="Gölhisar-Mesutlu">Goelhisar-Mesutlu</normalizedToken>
<normalizedToken originalValue="arası">arasi</normalizedToken>
,
<geoCoordinate degrees="37" direction="north" minutes="47" orientation="latitude" precision="15" seconds="57" value="37.799168">37°47'57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="27" direction="east" minutes="55" orientation="longitude" precision="15" seconds="42" value="27.928333">27°55'42&quot;E</geoCoordinate>
, 48 m, 23.II.2006, 7 ♀♀, 2 ♂♂,
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<normalizedToken originalValue="köyü">koeyue</normalizedToken>
,
<geoCoordinate degrees="37" direction="north" minutes="48" orientation="latitude" precision="15" seconds="06" value="37.801666">37°48'06&quot;N</geoCoordinate>
,
<geoCoordinate degrees="28" direction="east" minutes="00" orientation="longitude" precision="15" seconds="55" value="28.015278">28°00'55&quot;E</geoCoordinate>
, 46 m, 23.II.2006, 6 ♀♀, 6 ♂♂,
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,
<geoCoordinate degrees="37" direction="north" minutes="47" orientation="latitude" precision="15" seconds="24" value="37.79">37°47'24&quot;N</geoCoordinate>
,
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, 35 m, 23.II.2006, 1♀, 3 ♂♂,
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,
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,
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, 61 m, 23.II.2006, 7 ♂♂, leg. B.
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, S.
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, Umurlu,
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<normalizedToken originalValue="köyü">koeyue</normalizedToken>
,
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,
<geoCoordinate degrees="27" direction="east" minutes="55" orientation="longitude" precision="15" seconds="50" value="27.930555">27°55'50&quot;E</geoCoordinate>
, 30 m, 13.III.2010, 8 ♀♀, leg. E. Scheuchl.
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<bibRefCitation id="10E045477DE42F68658B41B46B1104D0" author="Jaech, M A" journalOrPublisher="Water beetles of China" pageId="0" pageNumber="118110" refId="B10960370" refString="Jaech, M A, Easton, E R, 1998. . Water beetles of China Vol. 2" volume="Vol. 2" year="1998">
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Girault, 1933
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Soyka, 1946
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Soyka, 1946
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Soyka, 1946
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Soyka, 1946
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Soyka, 1946
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Soyka, 1946
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Soyka, 1950
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Soyka, 1950
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Soyka, 1950
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Ogloblin, 1962
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<taxonomicName LSID="0C043511-C290-5F70-AF6F-4E2C5549A367" authority="Burukovsky, 1972" authorityName="Burukovsky" authorityYear="1972" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus)" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) Burukovsky, 1972</taxonomicName>
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<paragraph pageId="0" pageNumber="29">Type species.</paragraph>
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<taxonomicName authorityName="Ives" authorityYear="1891" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus brasiliensis subsp. var. var. aztecus" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="variety" species="brasiliensis" subSpecies="var." variety="aztecus">Penaeus brasiliensis var. aztecus</taxonomicName>
Ives, 1891.
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<subSubSection pageId="0" pageNumber="29" type="gender of subgenus">
<paragraph pageId="0" pageNumber="29">Gender of subgenus.</paragraph>
<paragraph pageId="0" pageNumber="29">Masculine.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="diagnosis">
<paragraph pageId="0" pageNumber="29">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="29">Rostrum usually bearing 2 ventral teeth. Median sulcus at postrostral carina generally distinct, long. Adrostral sulcus extending to near posterior margin of carapace. Gastrofrontal carina distinct, with posterior end not turning anteriorly. Cervical carina with dorsal end a distance from dorsal carapace. Hepatic carina distinct. First pereiopod with strong ischial spine. Fifth pereiopod bearing exopod. Sixth abdominal somite with distinct dorsolateral sulcus. Telson without lateral spines. Thelycum closed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="species included">
<paragraph pageId="0" pageNumber="29">Species included.</paragraph>
<paragraph pageId="0" pageNumber="29">
<taxonomicName authorityName="Ives" authorityYear="1891" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) aztecus" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="aztecus" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) aztecus</taxonomicName>
Ives, 1891,
<taxonomicName class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) brasiensis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="brasiensis" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) brasiensis</taxonomicName>
Latreille, 1817,
<taxonomicName authorityName="Kingsley" authorityYear="1878" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) brevirostris" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="brevirostris" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) brevirostris</taxonomicName>
Kingsley, 1878,
<taxonomicName authorityName="Holmes" authorityYear="1900" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) californiensis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="californiensis" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) californiensis</taxonomicName>
Holmes, 1900,
<taxonomicName authorityName="Burkenroad" authorityYear="1939" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) duorarum" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="duorarum" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) duorarum</taxonomicName>
Burkenroad, 1939,
<taxonomicName baseAuthorityName="Tavares &amp; Guzmao" baseAuthorityYear="2016" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) isabelae" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="isabelae" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) isabelae</taxonomicName>
Tavares &amp;
<normalizedToken originalValue="Gusmão">Gusmao</normalizedToken>
, 2016,
<taxonomicName authorityName="Perez Farfante" authorityYear="1967" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) notialis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="notialis" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) notialis</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Farfante, 1967,
<taxonomicName authorityName="Perez Farfante" authorityYear="1967" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) paulensis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="paulensis" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) paulensis</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Farfante, 1967,
<taxonomicName class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Farfantepenaeus) subtilis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="subtilis" subGenus="Farfantepenaeus">Penaeus (Farfantepenaeus) subtilis</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Farfante, 1967.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="remarks">
<paragraph pageId="0" pageNumber="29">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="29">
This subgenus together with
<taxonomicName authority="" authorityName="Perez Farfante" authorityYear="1969" class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus (Litopenaeus)" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Litopenaeus">Penaeus (Litopenaeus)</taxonomicName>
are often called the American
<taxonomicName class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="29">Penaeus</emphasis>
</taxonomicName>
. Morphologically these two subgenera are markedly different from each other and had long been thought to be evolutionary far apart (see
<bibRefCitation author="Burkenroad, MD" journalOrPublisher="Bulletin of the Bingham Oceanographic Collection" pageId="0" pageNumber="29" pagination="1 - 109" refId="B4" refString="Burkenroad, MD, 1934. Littoral Penaeidea chiefly from the Bingham Oceanographic Collection, with a revision of Penaeopsis and descriptions of two new genera and eleven new American species. Bulletin of the Bingham Oceanographic Collection 4: 1 - 109" title="Littoral Penaeidea chiefly from the Bingham Oceanographic Collection, with a revision of Penaeopsis and descriptions of two new genera and eleven new American species." volume="4" year="1934">Burkenroad 1934</bibRefCitation>
;
<bibRefCitation author="Kubo, I" journalOrPublisher="Journal of the Tokyo College of Fisheries" pageId="0" pageNumber="29" pagination="1 - 467" refId="B26" refString="Kubo, I, 1949. Studies on penaeids of Japanese and its adjacent waters. Journal of the Tokyo College of Fisheries 36: 1 - 467" title="Studies on penaeids of Japanese and its adjacent waters." volume="36" year="1949">Kubo 1949</bibRefCitation>
;
<bibRefCitation author="Perez Farfante, I" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="29" refId="B36" refString="Perez Farfante, I, 1969. Western Atlantic shrimps of the genus P.naeus. Fish Bulletin 67: 461-591." title="Western Atlantic shrimps of the genus P. naeus. Fish Bulletin 67: 461 - 591." year="1969">
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Farfante 1969
</bibRefCitation>
;
<bibRefCitation author="Dall, W" journalOrPublisher="Advances in Marine Biology" pageId="0" pageNumber="29" pagination="1 - 484" refId="B13" refString="Dall, W, Hill, BJ, Rothlisberg, NW, Staples, DJ, 1990. The biology of Penaeidae. Advances in Marine Biology 27: 1 - 484" title="The biology of Penaeidae." volume="27" year="1990">Dall et al. 1990</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.18353/crustacea.24.0_146" author="von Sternberg, R" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="29" pagination="146 - 156" refId="B42" refString="von Sternberg, R, Motoh, H, 1995. Notes on the phylogeny of the American Penaeus shrimps (Decapoda: Dendrobranchiata: Penaeidae). Crustacean Research 24 (0): 146 - 156, DOI: https://doi.org/10.18353/crustacea.24.0_146" title="Notes on the phylogeny of the American Penaeus shrimps (Decapoda: Dendrobranchiata: Penaeidae)." url="https://doi.org/10.18353/crustacea.24.0_146" volume="24" year="1995">von Sternberg and Motoh 1995</bibRefCitation>
;
<bibRefCitation author="Perez Farfante, I" journalOrPublisher="Memoires du Museum National d'Histoire naturelle, Paris" pageId="0" pageNumber="29" pagination="1 - 233" refId="B37" refString="Perez Farfante, I, Kensley, B, 1997. Penaeoid and sergestoid shrimps and prawns of the world. Keys and diagnoses for the families and genera. Memoires du Museum National d'Histoire naturelle, Paris 175: 1 - 233" title="Penaeoid and sergestoid shrimps and prawns of the world. Keys and diagnoses for the families and genera." volume="175" year="1997">
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Farfante and Kensley 1997
</bibRefCitation>
;
<bibRefCitation author="von Sternberg, R" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="29" refId="B41" refString="von Sternberg, R, 1997. Phylogenetic and systematic position of the Penaeus subgenus Litopenaeus (Decapoda: Penaeidae). Revistra de Biologia Tropical 44(3)-45(1): 441-451." title="Phylogenetic and systematic position of the Penaeus subgenus Litopenaeus (Decapoda: Penaeidae). Revistra de Biologia Tropical 44 (3) - 45 (1): 441 - 451." year="1997">von Sternberg 1997</bibRefCitation>
). They are, however, very closely related genetically (see
<bibRefCitation DOI="https://doi.org/10.1016/j.aquaculture.2022.738955" author="Yang, CH" journalOrPublisher="Aquaculture (Amsterdam, Netherlands)" pageId="0" pageNumber="29" pagination="1 - 10" refId="B49" refString="Yang, CH, Ma, KY, Chu, KH, Chan, TY, 2023. Making sense of the taxonomy of the most commercially important shrimps Penaeus Fabricius, 1798 s. l. (Crustacea: Decapoda: Penaeidae), a way forward. Aquaculture (Amsterdam, Netherlands) 563: 1 - 10, DOI: https://doi.org/10.1016/j.aquaculture.2022.738955" title="Making sense of the taxonomy of the most commercially important shrimps Penaeus Fabricius, 1798 s. l. (Crustacea: Decapoda: Penaeidae), a way forward." url="https://doi.org/10.1016/j.aquaculture.2022.738955" volume="563" year="2023">Yang et al. 2023</bibRefCitation>
). At present only one morphological character, the sixth abdominal somite with dorsolateral sulcus, is found to separate the American
<taxonomicName class="Malacostraca" family="Penaeidae" genus="Penaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Penaeus" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="29">Penaeus</emphasis>
</taxonomicName>
from other congeneric species. Recent molecular analysis has suggested that
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="P. (Farfantepenaeus) notialis" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="notialis" subGenus="Farfantepenaeus">P. (Farfantepenaeus) notialis</taxonomicName>
, originally described as a subspecies of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="P. (Farfantepenaeus) duorarum" order="Decapoda" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="duorarum" subGenus="Farfantepenaeus">P. (Farfantepenaeus) duorarum</taxonomicName>
, may not be distinct at the species level (
<bibRefCitation DOI="https://doi.org/10.1071/IS18044" author="Timm, L" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="29" pagination="488 - 500" refId="B46" refString="Timm, L, Browder, JA, Simon, S, Jackson, TL, Zink, IC, Bracken-Grissom, HD, 2019. A tree money grows on: the first inclusive molecular phylogeny of the economically important pink shrimp (Decapoda: Farfantepenaeus) reveals cryptic diversity. Invertebrate Systematics 33: 488 - 500, DOI: https://doi.org/10.1071/IS18044" title="A tree money grows on: the first inclusive molecular phylogeny of the economically important pink shrimp (Decapoda: Farfantepenaeus) reveals cryptic diversity." url="https://doi.org/10.1071/IS18044" volume="33" year="2019">Timm et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="45A20E40939169971D5DAD656305269B" pageId="0" pageNumber="79255">Mediterranean-Asiatic</paragraph>
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<paragraph id="C37F53B79632DDB02B5BD553196335BD" pageId="0" pageNumber="79255">
References:
<bibRefCitation id="B8E44BFDE48AAB698615F7BB2504F845" author="Gaston, J" journalOrPublisher="SHILAP Revista de Lepidopterologia" pageId="0" pageNumber="79255" pagination="505 - 517" refId="B7551676" refString="Gaston, J, Vives, A, 2018. Revision del genero Gymnancyla Zeller, 1848 en Espana continental y descripcion de una nueva especie (Lepidoptera: Pyralidae, Phycitinae). SHILAP Revista de Lepidopterologia 46: 505 - 517" title="Revision del genero Gymnancyla Zeller, 1848 en Espana continental y descripcion de una nueva especie (Lepidoptera: Pyralidae, Phycitinae)" volume="46" year="2018">
<normalizedToken id="8836FFA55C3D19FEFB5026189A59E724" originalValue="Gastón">Gaston</normalizedToken>
and Vives (2018)
</bibRefCitation>
. Biological data: Univoltine. Flight period: VIII.
</paragraph>
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<mods:title id="2C9565E90B3B099890E9C4742D6A74FF">The genus Litophyton Forskal, 1775 (Octocorallia, Alcyonacea, Nephtheidae) in the Red Sea and the western Indian Ocean</mods:title>
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<paragraph id="878133B781B2B563EF1D6A258BBE97CA" pageId="11" pageNumber="12">Taxon classification Animalia Alcyonacea Nephtheidae</paragraph>
</subSubSection>
<subSubSection id="085708B8298F8E0463681621BB7EAEEA" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="17182944EEAFF4F7ADABF4A36CBE8875" pageId="11" pageNumber="12">
<taxonomicName id="76D34554CD507A428DC7B5496D652067" ID-CoL="3VMCN" authority="Kuekenthal, 1913" authorityName="Kuekenthal" authorityYear="1913" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">
Litophyton laevis (
<normalizedToken id="5DAADEA7C89A96892827317989F82A16" originalValue="Kükenthal">Kuekenthal</normalizedToken>
, 1913)
</taxonomicName>
Figures 2A, 36, 37, 38, 39, 40, 41
</paragraph>
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<paragraph id="405A0B61ED24B27AD35944F72FE229D3" pageId="11" pageNumber="12">
<taxonomicName id="8DF860FD4034974630ED0090C2DB74BC" class="Anthozoa" family="Nephtheidae" genus="Nephthya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephthya laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">Nephthya laevis</taxonomicName>
<normalizedToken id="87BB20698537E9159B7B230D23483594" originalValue="Kükenthal">Kuekenthal</normalizedToken>
, 1913: 20, figs 9-13, pl. 2 fig. 5 (Red Sea, Jeddah).
</paragraph>
<paragraph id="E0CB6DCB5708E8325749FAEBAD897DBF" pageId="11" pageNumber="12">
<taxonomicName id="4BA967B48996BAE5759179D9BAD8FBAC" class="Anthozoa" family="Nephtheidae" genus="Nephthea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephthea laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">Nephthea laevis</taxonomicName>
;
<bibRefCitation id="E6A20B7BC672826B045DE991D1D91DDC" author="Roxas, HA" journalOrPublisher="Philippine Journal of Science" pageId="29" pageNumber="30" pagination="345 - 470" title="Philippine Alcyonaria, II. The Families Alcyoniidae and Nephthyidae." volume="50" year="1933">Roxas 1933</bibRefCitation>
: 415;
<bibRefCitation id="4013B221088FECEB50FD2F35A1CA7654" author="Verseveldt, J" journalOrPublisher="Israel Journal of Zoology" pageId="30" pageNumber="31" pagination="209 - 229" title="Report on some Octocorallia (Alcyonacea) from the northern Red Sea." volume="19" year="1970">Verseveldt 1970</bibRefCitation>
: 219, figs 5-6, pl. 3 fig. 1 (Gulf of Suez, Et Tur).
</paragraph>
<paragraph id="628033DED4B54FC5B4489B19FC7351A1" pageId="11" pageNumber="12">
<taxonomicName id="5ABD15773D9866D43A9A531D71FA8D56" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">Litophyton laevis</taxonomicName>
Not
<taxonomicName id="2771D6005FE01E17D058904D16A39B15" class="Anthozoa" family="Nephtheidae" genus="Nephthea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephthea laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">Nephthea laevis</taxonomicName>
;
<bibRefCitation id="CC72F3CB46BACEB53EE4FAA117CC0EE5" author="Verseveldt, J" journalOrPublisher="Israel Journal of Zoology" pageId="30" pageNumber="31" pagination="1 - 37" title="Alcyonacea (Octocorallia) from the Red Sea, with a discussion of a new Sinularia species from Ceylon." volume="23" year="1974 b">Verseveldt 1974b</bibRefCitation>
: 2 (Red Sea, Gulf of Aqaba, El Hamira; listed only; =
<taxonomicName id="BB68D7EE0FB6C6949B1B15D998248D0B" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton arboreum" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="arboreum">Litophyton arboreum</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="52B3CDF2EE5D627A952B0AF6B24628CB" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="14D536CF859223C0B4250864F296093A" pageId="11" pageNumber="12">Material examined.</paragraph>
<paragraph id="5F74EF47F0BFB565189D06FCEB3ED716" pageId="11" pageNumber="12">
ZMB 6818, holotype,
<normalizedToken id="A38F305991869C22F3B0E62092E69DD2" originalValue="Kükth">Kuekth</normalizedToken>
det., Rotes Meer, Djidda, Pola Exp.; RMNH Coel. 6821, Red Sea, Gulf of Suez, Et Tur, 6. July 1969, coll. L. Fishelson; ZMTAU NS 8306, Red Sea, Gulf of Suez, El-Bilaiyim lagoon, 24 August 1971, coll. D. Popper; ZMTAU Co 25971, Red Sea, Gulf of Suez, Jubal Is., Bluf point, depth 4 m, 24 March 1988, coll. Y. Benayahu; ZMTAU Co 26126 3211, Red Sea, Gulf of Suez, between Shaduan and Tawilla Is., 25 September 1989, coll. Y. Benayahu; ZMTAU Co 26231, Red Sea, Gulf of Suez, Ras Gahra, 26 September 1974, coll. Y. Benayahu; ZMTAU Co 28550 (E258); Red Sea, Dahlak Archipelago, Daliacus; depth 3 m; 18 October 1993, coll. Y. Benayahu; ZMTAU Co 28585 (E121), Red Sea, Dahlak Archipelago, Dur Gam, depth 3 m, 14 October 1993, coll. Y. Benayahu.
</paragraph>
</subSubSection>
<subSubSection id="19E2AB103B5C9787956AE8236DD9E89D" pageId="11" pageNumber="12" type="removed from the species">
<paragraph id="3CC5B62374F6992AEA536069BB080AE4" pageId="11" pageNumber="12">Removed from the species.</paragraph>
<paragraph id="7C75C061E3C369FA30E1F7F8EC91927F" pageId="11" pageNumber="12">
RMNH 8941, Red Sea, Gulf of Aqaba, Ophir Bay, 30 August 1967, coll. Hebrew Univ.- Smiths. Red Sea project (identified by Verseveldt as
<taxonomicName id="94882903469C64166ADE3F42777120A4" class="Anthozoa" family="Nephtheidae" genus="Nephthea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephthea laevis" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="laevis">Nephthea laevis</taxonomicName>
=
<taxonomicName id="25930538A9C8FDC1FCD2223DC5E7AB1A" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton arboreum" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="arboreum">Litophyton arboreum</taxonomicName>
).
</paragraph>
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<subSubSection id="FE9F69D74BA58D94BF604B258045700D" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="06488B03D09489EB79CEACBF9BE672F0" pageId="11" pageNumber="12">Diagnosis.</paragraph>
<paragraph id="7FBB135212D31B0B9D4CD666653A431D" pageId="11" pageNumber="12">
<taxonomicName id="E9CF1EDDE22A92B14EFF6CFEF6F97C05" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">Litophyton</taxonomicName>
with the internal spindles of the base of the stalk short and slender, up to 0.15 mm wide and 0.5 mm long.
</paragraph>
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<subSubSection id="EFBC3EC926CC55EF60431F260D9685CA" pageId="11" pageNumber="12" type="description">
<paragraph id="A925366B5E60CE4E4E0FC02A7B44E88B" pageId="11" pageNumber="12">Description.</paragraph>
<paragraph id="1AE7E9CB755603B3A57ADE0FC5579409" pageId="11" pageNumber="12">The holotype is 8 cm high and 5 cm wide; the short colony stalk divides in several main stems shortly above its base (Figure 36A). Polyps are crowded at the end of the lobes arranged in globular to oval-shaped structures.</paragraph>
<paragraph id="EB78A816494E9A01AE85DA7442F3455D" pageId="11" pageNumber="12">
The polyps are up to about 0.7 mm high and 0.6 mm wide (Figure 37A). Supporting bundle not projecting, composed of spindles with simple tubercles, outer side and distal end with larger tubercles. Length of these spindles is up to 0.8 mm (Figure 37B). Polyp body sclerites irregularly arranged, the smallest are present adaxially; abaxially they merge into the smaller spindles of the supporting bundle (Figure 37C). The tentacle sclerites resemble the smallest adaxial polyp sclerites (Figures 37D, 39
<normalizedToken id="445521E325CFC880AE00A1A987DA223F" originalValue="BC">B-C</normalizedToken>
).
</paragraph>
<paragraph id="E6642727ED06DC1DDF751D7C1FF71995" pageId="11" pageNumber="12">Surface layer top of stalk. Rods and spindles, up to 0.45 mm long, with simple tubercles (Figure 37E).</paragraph>
<paragraph id="6215133F688BCDF68D91D6BBE547631F" pageId="11" pageNumber="12">Surface layer base of stalk. Radiates and derivatives of these, up to 0.15 mm long, with simple tubercles; a few are unilaterally spinose (Figure 38A). A few spindles and unilaterally spinose spindles are also present, up to 0.45 mm long, with simple tubercles.</paragraph>
<paragraph id="E7BF276F39DE0F8F80791B6DE9F9ECB6" pageId="11" pageNumber="12">Interior base of stalk. Spindles, up to 0.5 mm long, with simple sparse tubercles (Figure 38B). Several spindles have one or more side branches.</paragraph>
<paragraph id="EE7187D676F1DA1359313AEBAE6F12D3" pageId="11" pageNumber="12">Colour. The colony is whitish.</paragraph>
</subSubSection>
<subSubSection id="BA445696B20B9FE3FDC0626D62465CA2" pageId="11" pageNumber="12" type="distribution">
<paragraph id="B0BCBC96019318B07A7CE90616377D7D" pageId="11" pageNumber="12">Distribution.</paragraph>
<paragraph id="B33CDF03CC3EC29809FADCEA5AD1E7E9" pageId="11" pageNumber="12">Red Sea: Gulf of Suez, Dahlak Archipelago.</paragraph>
</subSubSection>
<subSubSection id="50A97A963A9E2144349EAAF74D235B16" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="remarks">
<paragraph id="DF301DA26F17440A38D3F982523913E8" pageId="11" pageNumber="12">Remarks.</paragraph>
<paragraph id="CF2B39DEE53A5C192CE10F6602938A7E" pageId="11" pageNumber="12">
<bibRefCitation id="C82E86FA99816B9E8B706058066AE540" author="Kuekenthal, W" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften Wien, Mathematisch-Naturwissenschaftliche Klasse" pageId="28" pageNumber="29" pagination="1 - 33" title="Alcyonaria des Roten Meeres. Expeditionen S. M. Schiff &quot; Pola &quot; in das Rote Meer. Zoologische Ergebnisse 29." volume="89" year="1913">
<normalizedToken id="E6885367A2EF8376443AF422F3D1E9A6" originalValue="Kükenthal">Kuekenthal</normalizedToken>
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mentioned four specimens with his description, in Berlin I found only one specimen, labelled holotype, which is the same one that
<normalizedToken id="10DF9FD58AB39942A97011CC6CADF553" originalValue="Kükenthal">Kuekenthal</normalizedToken>
used in his description. He mentioned longer interior spindles, up to 1 mm long. I assume I missed the longer ones as only few interior spindles are present in the microscope slide made of the stalk of ZMB 6818.
</paragraph>
<paragraph id="61386EF4DA6F56FD1BA71785E3E98A70" pageId="12" pageNumber="13">
<pageBreakToken id="85077D35E417E0F965F5B96600417941" pageId="12" pageNumber="13" start="start">The</pageBreakToken>
species can be confused with
<taxonomicName id="D38F0970A7045AF6267535E854A45370" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton simulatum" order="Alcyonacea" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="simulatum">Litophyton simulatum</taxonomicName>
, but the latter has wider, more branched internal spindles.
</paragraph>
<paragraph id="EFF9D8CE33EB9AF1D243FE25F3C759AA" pageId="12" pageNumber="13">ZMTAU Co 26126 3211 has been used for SEM images of sclerites (Figures 39-41).</paragraph>
<paragraph id="9874E2CBDE77937B02D7E68AE0BE4652" pageId="12" pageNumber="13">
RMNH Coel. 8941 has spindles up to 1.3 mm long in the interior of the stalk, quite some of them with blunt ends and therefore I re-identified it as
<taxonomicName id="12DF5B78D53D2CCE7AD81FD87C8F8F32" class="Anthozoa" family="Nephtheidae" genus="Litophyton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litophyton arboreum" order="Alcyonacea" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="arboreum">Litophyton arboreum</taxonomicName>
.
</paragraph>
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<bibCitation id="FD3CE6D73C0901E28A6DDE6DCD9C10B9" author="Linnaeus" pagination="829" title="Atractylis gummifera" volumeTitle="Species Plantarum 2" year="1753">
<taxonomicName id="21B43530895AE891552CE416A28864A8" ID-CoL="5VWBW" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Atractylis" higherTaxonomySource="GBIF,CoL,IPNI" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="gummifera">
<emphasis id="858C89847A07DBCE739B274C8D2CBABC" bold="true" pageNumber="338">Atractylis gummifera</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="9922988E7EFA97637985F00763B148C4">
<emphasis id="0FBB1CD00BAC96619B6771D591778E9C" italics="true" pageNumber="338">Species Plantarum</emphasis>
2
</volumeTitle>
: 829. 1753
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52F16A0560BB0EE08F5FA3E49D456F98" pageNumber="338" type="distribution">
<paragraph id="08CF037457EE62EB387AB02021A15F1D" pageNumber="338">&quot;Habitat in Creta, Italia.&quot; RCN: 5999.</paragraph>
</subSubSection>
<subSubSection id="5582D97B928E83633F2EF173981E8268" pageNumber="338" type="materials_examined">
<paragraph id="8E7ED42EE1D06243B843E1087C40A08A" pageNumber="338">
<materialsCitation id="E3C388A992CDB75DEF49639C7E0ADF78" ID-GBIF-Occurrence="2612071187" collectionCode="LINN" specimenCode="971.1" typeStatus="lectotype">
<typeStatus id="7A3C9D6E93DAAEAD7000141CBCE70573">Lectotype</typeStatus>
(Petit in
<emphasis id="9C853C6488A5DC5697EE6A5C202E3CB4" italics="true" pageNumber="338">
Bull. Mus. Natl. Hist. Nat., B,
<taxonomicName id="FE32C52DDE92D02B58DD3E63E383013A" class="Magnoliopsida" family="Malvaceae" genus="Adansonia" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">Adansonia</taxonomicName>
</emphasis>
9: 412. 1988 [1987]): Herb. Linn. No. 971.1 (
<collectionCode id="DA8FB43232ADDE4551EBF73E01003C9B">LINN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6CF7BE58544CA0DA6C39FE0D01165678" pageNumber="338" type="nomenclature">
<paragraph id="B28B4701D412C28230142F53A9E896FB" pageNumber="338">
Current name:
<taxonomicName id="21B5C2836E0CDCF57CC2E2D4D8F09A40" ID-CoL="69C42" ID-ENA="143183" authority="(L.) Less." authorityName="Less." baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Carlina" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="gummifera">
<emphasis id="49F91CD2C71CEE50209F7B44A543D75F" italics="true" pageNumber="338">
<emphasis id="F3385B872D247E3F5E76D46079BA79CA" bold="true" italics="true" pageNumber="338">Carlina gummifera</emphasis>
</emphasis>
(L.) Less.
</taxonomicName>
(
<taxonomicName id="68F10DF4B0AA6CF74BCE287531C3A482" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,116 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.878.38408" ID-GBIF-Dataset="61d76a4c-8496-40dd-843e-3dd87a54ecd1" ID-PMC="PMC6791897" ID-Pensoft-Pub="1313-2970-878-1" ID-Pensoft-UUID="77A4C4CD21F25D92B2AFB96AFDA8DFC0" ID-PubMed="31632174" ID-ZooBank="9B51B440ACFC4E1A91EA32B28554AF56" ModsDocID="1313-2970-878-1" checkinTime="1570497189296" checkinUser="pensoft" docAuthor="Kocic, Korana, Petrovic, Andjeljko, Ckrkic, Jelisaveta, Mitrovic, Milana &amp; Zeljko Tomanovic," docDate="2019" docId="0C04A02A6E1156A4BCE7B535FC511E64" docLanguage="en" docName="ZooKeys 878: 1-22" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.38408" docTitle="(Fovephedrus) Chen 1986" docType="treatment" docVersion="5" id="77A4C4CD21F25D92B2AFB96AFDA8DFC0" lastPageNumber="1" masterDocId="77A4C4CD21F25D92B2AFB96AFDA8DFC0" masterDocTitle="Phylogenetic relationships and subgeneric classification of European Ephedrus species (Hymenoptera, Braconidae, Aphidiinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="1" updateTime="1668167844277" updateUser="ExternalLinkService">
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<mods:title>Phylogenetic relationships and subgeneric classification of European Ephedrus species (Hymenoptera, Braconidae, Aphidiinae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kocic, Korana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petrovic, Andjeljko</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ckrkic, Jelisaveta</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mitrovic, Milana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zeljko Tomanovic,</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2019</mods:date>
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<mods:number>878</mods:number>
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<mods:start>1</mods:start>
<mods:end>22</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.878.38408</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.878.38408</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-878-1</mods:identifier>
<mods:identifier type="ZooBank">9B51B440ACFC4E1A91EA32B28554AF56</mods:identifier>
<mods:identifier type="Pensoft-UUID">77A4C4CD21F25D92B2AFB96AFDA8DFC0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159654813" LSID="urn:lsid:plazi:treatment:0C04A02A6E1156A4BCE7B535FC511E64" httpUri="http://treatment.plazi.org/id/0C04A02A6E1156A4BCE7B535FC511E64" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
Subgenus
<taxonomicName LSID="0C04A02A-6E11-56A4-BCE7-B535FC511E64" authority="Chen, 1986" authorityName="Chen" authorityYear="1986" family="Braconidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="(Fovephedrus)" order="Hymenoptera" pageId="0" pageNumber="1" rank="subGenus" subGenus="Fovephedrus">Fovephedrus Chen, 1986</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" genus="Fovephedrus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Fovephedrus radiatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="radiatus">
<emphasis italics="true" pageId="0" pageNumber="1">Fovephedrus radiatus</emphasis>
</taxonomicName>
Chen, 1986.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Name derived from presence of fovea on mesoscutum of type species</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">3Rsa vein shorter than 2Rsa (3Rsa / 2Rsa less than 1), petiole subquadrate.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">F1 with smaller number of longitudinal placodes (0-3, rarely 4). Mesoscutum usually with one or two mesoscuteal foveae, sometimes lacking both. Scutellar sulcus always undivided. 3RSa shorter than 2Rsa (0.55-0.95). 3Rsb/3Rsa 2.65-3.4. Petiole short and broad, 1.3-1.5 times as long as wide, lacking post lateral excavations. Ovipositor sheaths varying from stout to slender.</paragraph>
<paragraph pageId="0" pageNumber="1">
Species in Europe:
<taxonomicName class="Insecta" family="Braconidae" genus="Ephedrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephedrus persicae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="persicae">
<emphasis italics="true" pageId="0" pageNumber="1">Ephedrus persicae</emphasis>
</taxonomicName>
Froggatt, 1904,
<taxonomicName class="Insecta" family="Braconidae" genus="Ephedrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephedrus chaitophori" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chaitophori">
<emphasis italics="true" pageId="0" pageNumber="1">Ephedrus chaitophori</emphasis>
</taxonomicName>
<normalizedToken originalValue="Gärdenfors">Gaerdenfors</normalizedToken>
, 1986,
<taxonomicName class="Insecta" family="Braconidae" genus="Ephedrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephedrus lonicerae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lonicerae">
<emphasis italics="true" pageId="0" pageNumber="1">Ephedrus lonicerae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Tomanović">Tomanovic</normalizedToken>
, Kavallieratos &amp;
<normalizedToken originalValue="Starý">Stary</normalizedToken>
, 2009,
<taxonomicName class="Insecta" family="Braconidae" genus="Ephedrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephedrus tamaricis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tamaricis">
<emphasis italics="true" pageId="0" pageNumber="1">Ephedrus tamaricis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Tomanović">Tomanovic</normalizedToken>
&amp;
<normalizedToken originalValue="Petrović">Petrovic</normalizedToken>
, 2016
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,198 @@
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.51.33529" ID-GBIF-Dataset="2dcbe70b-c6ad-4dbc-b8ad-c047a5322057" ID-PMC="PMC6520330" ID-Pensoft-Pub="1314-4049-51-107" ID-PubMed="31139006" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-51-107" ModsDocOrigin="MycoKeys 51" ModsDocTitle="Two new species of Phylloporus (Fungi, Boletales) from tropical Quercus forests in eastern Mexico" checkinTime="1557379565242" checkinUser="pensoft" docAuthor="Montoya, Leticia, Garay-Serrano, Edith &amp; Bandala, Victor M." docDate="2019" docId="0C04E2411FB2BBADDEACCFD182F872F0" docLanguage="en" docName="MycoKeys 51: 107-123" docOrigin="MycoKeys 51" docSource="http://dx.doi.org/10.3897/mycokeys.51.33529" docTitle="Phylloporus rimosus Bandala, Montoya &amp; Garay, sp. nov." docType="treatment" docVersion="4" lastPageNumber="107" masterDocId="FFC9314EFFE0FF8BFFFEFF88FF970A3E" masterDocTitle="Two new species of Phylloporus (Fungi, Boletales) from tropical Quercus forests in eastern Mexico" masterLastPageNumber="123" masterPageNumber="107" pageNumber="107" updateTime="1668136210528" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of Phylloporus (Fungi, Boletales) from tropical Quercus forests in eastern Mexico</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Montoya, Leticia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garay-Serrano, Edith</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bandala, Victor M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>MycoKeys</mods:title>
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<mods:date>2019</mods:date>
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<mods:number>51</mods:number>
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<mods:start>107</mods:start>
<mods:end>123</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.51.33529</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-51-107</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="157078302" LSID="urn:lsid:plazi:treatment:0C04E2411FB2BBADDEACCFD182F872F0" httpUri="http://treatment.plazi.org/id/0C04E2411FB2BBADDEACCFD182F872F0" lastPageNumber="107" pageId="0" pageNumber="107">
<subSubSection pageId="0" pageNumber="107" type="nomenclature">
<paragraph pageId="0" pageNumber="107">
<taxonomicName LSID="829439" authority="Bandala, Montoya &amp; Garay" class="Agaricomycetes" family="Boletaceae" genus="Phylloporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phylloporus rimosus" order="Boletales" pageId="0" pageNumber="107" phylum="Basidiomycota" rank="species" species="rimosus">Phylloporus rimosus Bandala, Montoya &amp; Garay</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="107">sp. nov.</taxonomicNameLabel>
Figs 2a, b, 3, 4
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="holotype">
<paragraph pageId="0" pageNumber="107">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="107">
MEXICO. Veracruz: Municipality of Coatepec,
<normalizedToken originalValue="Vaquería">Vaqueria</normalizedToken>
, gregarious in soil, under
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus oleoides" order="Fagales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="oleoides">Quercus oleoides</taxonomicName>
Schltdl. &amp; Cham., 27 June 2012, Montoya 4834 (XAL).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="diagnosis">
<paragraph pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="107">
Recognized by the combination of pileus vinaceous to grayish-vinaceous, surface becoming rimose-areolate with development, the stipe apex with ribbed appearance and scabrous or even with tiny rigid scales and gills staining blue. Its stature (pileus 27-80 mm diam., stipe 27-80
<normalizedToken originalValue="×">x</normalizedToken>
7-12 mm), basidiospores and pleurocystidia size and shape, prevents confusion with
<taxonomicName lsidName="P. purpurellus" pageId="0" pageNumber="107" rank="species" species="purpurellus">P. purpurellus</taxonomicName>
Singer or with
<taxonomicName lsidName="P. scabripes" pageId="0" pageNumber="107" rank="species" species="scabripes">P. scabripes</taxonomicName>
B. Ortiz &amp; M.A. Neves.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="gene sequences ex-holotype">
<paragraph pageId="0" pageNumber="107">Gene sequences ex-holotype.</paragraph>
<paragraph pageId="0" pageNumber="107">MK226546 (ITS), MK226554 (LSU), MK314102 (tef-1α).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="etymology">
<paragraph pageId="0" pageNumber="107">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="107">Referring to the rimose pileus surface.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Description.</paragraph>
<paragraph pageId="0" pageNumber="107">
Pileus 27-80 mm diam, convex to plane-convex, at times faintly depressed at center or even infundibuliform; surface velvety, uniform but frequently rimose-areolate, or fracturing and forming rivulose patches, cracked when seen under lens, vinaceous to grayish-vinaceous (7D4-D5, 7C4; 5YR 3/4, 4/3, 4/4-25Y 6/6), darker in some areas especially towards the margin, or yellowish, reddish-yellow, reddish-brown or even yellowish-beige (10YR 5/4, 6/6) in other parts especially towards the center, some specimens even reddish-vinaceous (7E8-E7) with brownish tinges (7D6-6E8), mature specimens fading to brownish when exposed to the sun; margin slightly incurved, edge entire, at times undulate. Lamellae subdecurrent to decurrent, 9-15 mm broad, close, bright yellow (3A7, 5A6-A7; 5Y8/8; 4A16), mustard yellow with age (4A6-A7; 4B7-B8), staining blue or greenish-blue when handled, stains becoming reddish or brownish-vinaceous after several minutes, old specimens or specimens long exposed to the sun developing reddish spots at lamellae sides or even dark brownish red or brown at edge; somewhat sinuous when the hymenophore is seen frontally, veined or anastomosed mostly in the area below the pileus and intervenose or even somewhat labyrinthiform, especially when young; lamellullae of different sizes, edge entire. Stipe 27-80
<normalizedToken originalValue="×">x</normalizedToken>
7-12 mm, cylindrical, curved, somewhat sinuous, compact, apex with ribbed appearance by decurrent lines of the lamellae, surface pruinose, scabrous or even with tiny rigid scales, cracked after long exposure to the sun, beige (10YR 6/6-8) or pale yellow (4A/2), or whitish at the bottom of the surface and covered with a reddish or oxide-red (25YR 4/6) pruina, at the middle area reddish-beige (8D16), at times caespitose. Basal mycelium whitish-cream with some yellow spots or even mustard yellow (5Y8/6). Context yellow, staining pinkish or pinkish-brown. KOH 3% reddish (10YR 3/6 to 2.5YR 3/4) on pileus, stipe surface and context; NH4OH 10% greenish-blue (5Y 2.5/1) on pileus surface, the center of the stain becoming reddish (2.5YR 3/6), brownish at the hymenium, negative in the context and faintly green or negative on stipe surface. Odor mild to slightly citric. Taste mild.
</paragraph>
<paragraph pageId="0" pageNumber="107">
Basidiospores (9-) 9.5-14 (-15)
<normalizedToken originalValue="×">x</normalizedToken>
3.5-5
<normalizedToken originalValue="µm">µm</normalizedToken>
, X = 11-12.3
<normalizedToken originalValue="×">x</normalizedToken>
4.3-4.6
<normalizedToken originalValue="µm">µm</normalizedToken>
, Q = 2.5 2.8
<normalizedToken originalValue="µm">µm</normalizedToken>
, subfusiform, with suprahilar depression, somewhat ventricose, apex attenuated, yellow to amber yellow in KOH, wall slightly thickened (up to 0.5
<normalizedToken originalValue="µm">µm</normalizedToken>
thick). Basidia 29-50 (-55)
<normalizedToken originalValue="×">x</normalizedToken>
7-10 (-11)
<normalizedToken originalValue="µm">µm</normalizedToken>
, clavate, tetrasporic, rarely trisporic, hyaline, thin walled, unclamped. Pleurocystidia 42-105 (-120)
<normalizedToken originalValue="×">x</normalizedToken>
9-27
<normalizedToken originalValue="µm">µm</normalizedToken>
, narrowly to broadly utriform, at times cylindrical or subclavate, rarely sphaeropedunculate (52-58
<normalizedToken originalValue="×">x</normalizedToken>
20-23
<normalizedToken originalValue="μm">μm</normalizedToken>
), thin-walled, at times thickened in some areas, some with incrustations, hyaline, abundant, unclamped. Cheilocystidia (33-) 34-70 (-75)
<normalizedToken originalValue="×">x</normalizedToken>
8-17 (-19)
<normalizedToken originalValue="µm">µm</normalizedToken>
, narrowly utriform, hyaline, thin-walled, at times thickened towards the apex, unclamped. Pileipellis a trichodermis, with anticlinally oriented hyphae, tightly interwoven, frequently disposed in mounds, hyphae 8-16
<normalizedToken originalValue="µm">µm</normalizedToken>
broad, wall slightly thickened (up to 1
<normalizedToken originalValue="µm">µm</normalizedToken>
), hyaline yellowish-brown; terminal elements 23-64
<normalizedToken originalValue="×">x</normalizedToken>
8-14
<normalizedToken originalValue="µm">µm</normalizedToken>
, cylindrical, slightly inflated, other or clavate, pale yellowish-brown. Pileus trama hyphae 5-16
<normalizedToken originalValue="µm">µm</normalizedToken>
broad, in a lax interwoven arrangement, hyaline, thin walled. Hymenophoral trama arranged in a more or less regular central strand and somewhat divergent on both sides of the strand, with cylindrical hyphae 7-19
<normalizedToken originalValue="µm">µm</normalizedToken>
broad; some slightly inflated, hyaline, thin-walled, unclamped.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="habitat">
<paragraph pageId="0" pageNumber="107">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="107">
In soil, solitary or gregarious, in tropical oak forest, under
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus oleoides" order="Fagales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="oleoides">Quercus oleoides</taxonomicName>
and
<taxonomicName lsidName="Q. sapotifolia" pageId="0" pageNumber="107" rank="species" species="sapotifolia">Q. sapotifolia</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="materials_examined">
<paragraph pageId="0" pageNumber="107">Additional studied material.</paragraph>
<paragraph pageId="0" pageNumber="107">
MEXICO. Veracruz: Alto Lucero Co., NE Mesa de Venticuatro, 4 Oct 2016, Garrido14; 19 Sep 2017,
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
37. Zentla Co. Road Puentecilla-La
<normalizedToken originalValue="Piña">Pina</normalizedToken>
, 2 July 2009, Ramos 195. Around town of Zentla, 15 June 2016, Montoya 5232a; Montoya 5238; 23 June 2016,
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
5, Hervert 84; 30 June 2016, Cesar 61, Hervert 93; 6 July 2016, Caro 69; 30 Aug 2016, Garrido 3; 24 Aug 2017, Garay 368; 7 Sep 2017,
<normalizedToken originalValue="César">Cesar</normalizedToken>
84 (all at XAL).
</paragraph>
<caption pageId="0" pageNumber="107">
<paragraph pageId="0" pageNumber="107">
Figure 2. Basidiomes of
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Phylloporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phylloporus" order="Boletales" pageId="0" pageNumber="107" phylum="Basidiomycota" rank="genus">Phylloporus</taxonomicName>
species. a, b
<taxonomicName lsidName="P. rimosus" pageId="0" pageNumber="107" rank="species" species="rimosus">P. rimosus</taxonomicName>
(a Garrido 3, b Montoya 5232a) c
<taxonomicName lsidName="P. quercophilus" pageId="0" pageNumber="107" rank="species" species="quercophilus">P. quercophilus</taxonomicName>
(LM5239 holotype). Scale bars: 10 mm.
</paragraph>
</caption>
<caption pageId="0" pageNumber="107">
<paragraph pageId="0" pageNumber="107">
Figure 3.
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Phylloporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phylloporus rimosus" order="Boletales" pageId="0" pageNumber="107" phylum="Basidiomycota" rank="species" species="rimosus">Phylloporus rimosus</taxonomicName>
(Montoya 4834, holotype). a Basidiospores b hymenophoral trama c longitudinal section of pileipellis. Scale bars: 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(a), 25
<normalizedToken originalValue="μm">μm</normalizedToken>
(b), 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(c).
</paragraph>
</caption>
<caption pageId="0" pageNumber="107">
<paragraph pageId="0" pageNumber="107">
Figure 4.
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Phylloporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phylloporus rimosus" order="Boletales" pageId="0" pageNumber="107" phylum="Basidiomycota" rank="species" species="rimosus">Phylloporus rimosus</taxonomicName>
(Montoya 4834, holotype). a Basidiospores b basidia c cheilocystidia d pleurocystidia. Scale bars: 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(a), 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<normalizedToken originalValue="bd">b-d</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,76 @@
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<taxonomicName id="8DA1C3061164583143182E1272528FCF" ID-CoL="6TFGW" ID-ENA="115846" authority="Delle Chiaje, 1827" authorityName="Delle Chiaje" authorityYear="1827" class="Polychaeta" family="Hesionidae" genus="Oxydromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxydromus flexuosus" order="Phyllodocida" pageId="0" pageNumber="20997" phylum="Annelida" rank="species" species="flexuosus">Oxydromus flexuosus (Delle Chiaje, 1827)</taxonomicName>
</paragraph>
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<paragraph id="CF86BE89C44020A90A0CDAA8829DE7EA" pageId="0" pageNumber="20997">
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(Delle Chiaje, 1827)
</paragraph>
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<paragraph id="28010E760C44020F4FBE5A2BF7BA792F" pageId="0" pageNumber="20997">Type locality: Mediterranean (Gulf of Naples).</paragraph>
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</treatment>
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View file

@ -0,0 +1,101 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.102.24380" ID-PMC="PMC6033956" ID-Pensoft-Pub="1314-2003-102-1" ID-Pensoft-UUID="AA226A35FFF8FFBC37621A40C2518C67" ID-PubMed="30002597" ID-Zenodo-Dep="1306325" ModsDocID="1314-2003-102-1" checkinTime="1530324112772" checkinUser="pensoft" docAuthor="DeFilipps, Robert A. &amp; Krupnick, Gary A." docDate="2018" docId="0C06097211D35DCF96C11C65641E9426" docLanguage="en" docName="PhytoKeys 102: 1-341" docOrigin="PhytoKeys 102" docPubDate="2018-06-28" docSource="http://dx.doi.org/10.3897/phytokeys.102.24380" docTitle="Acacia nilotica Delile (= A. arabica (Lam.) Willd." docType="treatment" docVersion="3" id="AA226A35FFF8FFBC37621A40C2518C67" lastPageNumber="77" masterDocId="AA226A35FFF8FFBC37621A40C2518C67" masterDocTitle="The medicinal plants of Myanmar" masterLastPageNumber="341" masterPageNumber="1" pageNumber="77" updateTime="1668138950833" updateUser="ExternalLinkService">
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</mods:role>
<mods:namePart>DeFilipps, Robert A.</mods:namePart>
<mods:affiliation>Deceased</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krupnick, Gary A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1357-4826</mods:nameIdentifier>
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, PO Box 37012, MRC- 166, Washington, DC, 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">krupnick@si.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:number>2018-06-28</mods:number>
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<mods:number>102</mods:number>
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<paragraph pageId="76" pageNumber="77">
<taxonomicName LSID="0C060972-11D3-5DCF-96C1-1C65641E9426" authority="(L.) Delile (= A. arabica (Lam.) Willd.)" baseAuthorityName="L.) Delile (= A. arabica (Lam.) Willd." class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia nilotica" order="Fabales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="species" species="nilotica">
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nilotica (L.) Delile (= A. arabica (Lam.) Willd.)
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<subSubSection pageId="76" pageNumber="77" type="names">
<paragraph pageId="76" pageNumber="77">Names.</paragraph>
<paragraph pageId="76" pageNumber="77">
<emphasis bold="true" pageId="76" pageNumber="77">Myanmar</emphasis>
:
<emphasis italics="true" pageId="76" pageNumber="77">babu</emphasis>
,
<emphasis italics="true" pageId="76" pageNumber="77">babul</emphasis>
,
<emphasis italics="true" pageId="76" pageNumber="77">subyu</emphasis>
.
<emphasis bold="true" pageId="76" pageNumber="77">English</emphasis>
: babul, gum-arabic, Indian gum tree, suntwood.
</paragraph>
</subSubSection>
<subSubSection pageId="76" pageNumber="77" type="range">
<paragraph pageId="76" pageNumber="77">Range.</paragraph>
<paragraph pageId="76" pageNumber="77">Tropical Africa; widely naturalized in India. Naturalized in Myanmar.</paragraph>
</subSubSection>
<subSubSection pageId="76" pageNumber="77" type="use">
<paragraph pageId="76" pageNumber="77">Use.</paragraph>
<paragraph pageId="76" pageNumber="77">
<emphasis italics="true" pageId="76" pageNumber="77">Bark</emphasis>
: Used as an astringent.
</paragraph>
</subSubSection>
<subSubSection pageId="76" pageNumber="77" type="notes">
<paragraph pageId="76" pageNumber="77">Note.</paragraph>
<paragraph pageId="76" pageNumber="77">
In India the bark is employed as an astringent (
<bibRefCitation author="Jain, SK" journalOrPublisher="Phytotherapy Research" pageId="173" pageNumber="174" refId="B50" refString="Jain, SK, DeFilipps, RA, 1991. Medicinal Plants of India. 2 Vols. Reference Publications, Inc, Algonac." title="Medicinal Plants of India. 2 Vols. Reference Publications, Inc, Algonac." year="1991">Jain and DeFilipps 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="76" pageNumber="77" type="reference">
<paragraph pageId="76" pageNumber="77">Reference.</paragraph>
<paragraph pageId="76" pageNumber="77">
<bibRefCitation author="Nordal, A" journalOrPublisher="Taxon" pageId="174" pageNumber="175" refId="B85" refString="Nordal, A, 1963. The Medicinal Plants and Crude Drugs of Burma. Hellstrom &amp; Nordahls Boktrykkeri, Oslo." title="The Medicinal Plants and Crude Drugs of Burma. Hellstrom &amp; Nordahls Boktrykkeri, Oslo." year="1963">Nordal (1963)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,329 @@
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Budaeva, Nataliya</mods:namePart>
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</mods:role>
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</mods:role>
<mods:namePart>Capa, Maria</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glasby, Christopher J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9464-1938</mods:nameIdentifier>
<mods:affiliation>Museum and Art Gallery of the Northern Territory, Darwin, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hsueh, Pan-Wen</mods:namePart>
<mods:affiliation>Department of Life Sciences, National Chung Hsing University, Taichung City, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hutchings, Pat</mods:namePart>
<mods:affiliation>Australian Museum Research Institute, Sydney, Australia &amp; Macquarie University, Sydney, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jimi, Naoto</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8586-3320</mods:nameIdentifier>
<mods:affiliation>National Institute of Polar Research, Tachikawa, Tokyo, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kongsrud, Jon A.</mods:namePart>
<mods:affiliation>Department of Natural History, University Museum of Bergen, University of Bergen, Bergen, Norway</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Langeneck, Joachim</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3665-8683</mods:nameIdentifier>
<mods:affiliation>Department of Biology, University of Pisa, Pisa, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meissner, Karin</mods:namePart>
<mods:affiliation>Forschungsinstitut Senckenberg, DZMB, Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Murray, Anna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1765-1286</mods:nameIdentifier>
<mods:affiliation>Australian Museum Research Institute, Sydney, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nikolic, Mark</mods:namePart>
<mods:affiliation>Museums Victoria, Melbourne, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paxton, Hannelore</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7086-5219</mods:nameIdentifier>
<mods:affiliation>Australian Museum Research Institute, Sydney, Australia &amp; Macquarie University, Sydney, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ramos, Dino</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4069-5383</mods:nameIdentifier>
<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schulze, Anja</mods:namePart>
<mods:affiliation>Texas A &amp; M University at Galveston, Galveston, TX, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sobczyk, Robert</mods:namePart>
<mods:affiliation>Department of Zoology of Invertebrates and Hydrobiology, University of Lodz, Lodz, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watson, Charlotte</mods:namePart>
<mods:affiliation>Museum and Art Gallery of the Northern Territory, Darwin, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiklund, Helena</mods:namePart>
<mods:affiliation>Natural History Museum, London, UK &amp; Gothenburg Global Biodiversity Centre and University of Gothenburg, Gothenburg, Sweden</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Robin S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9441-2131</mods:nameIdentifier>
<mods:affiliation>Museums Victoria, Melbourne, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhadan, Anna</mods:namePart>
<mods:affiliation>Biological Faculty, Lomonosov Moscow State University, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Jinghuai</mods:namePart>
<mods:affiliation>South China Sea Environmental Monitoring Centre, State Oceanic Administration, Guangzhou, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1020</mods:number>
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<mods:start>1</mods:start>
<mods:end>198</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1020.57921</mods:url>
</mods:location>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1020.57921</mods:identifier>
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<treatment ID-GBIF-Taxon="179910900" LSID="urn:lsid:plazi:treatment:0C067329177817E32432F12206DC9883" httpUri="http://treatment.plazi.org/id/0C067329177817E32432F12206DC9883" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Maciolek, 1981" authorityName="Maciolek" authorityYear="1981" class="Polychaeta" family="Spionidae" genus="Aurospio" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Aurospio dibranchiata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="dibranchiata">Aurospio cf. dibranchiata Maciolek, 1981</taxonomicName>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Spionidae A Aurospio cf. dibranchiata: anterior end, dorsal view (AM W. 52242) B Aurospio sp. nov. 1, anterior end, dorsal view (AM W. 52240) C Prionospio cf. amarsupiata, anterior end, dorsal view (AM W. 52221) D Dipolydora notialis, heavy spine with crest of bristles on convex side from chaetiger 5 (AM W. 52245) E Dipolydora notialis, bidentate hooded hooks with smooth, curved shafts without constriction from posterior chaetiger (AM W. 52245) F Laonice cf. blakei: anterior end, dorsal-oblique view (AM W. 52226) G Spiophanes anoculata, posteriorly incomplete specimen in dorsal-oblique view, metameric dorsal ciliated organ bordered by pigment of dark orange or ochre colour (AM W. 52222). Scale bars: 100 µm (A, B, C); 1 µm (D, E); 200 µm (F); 500 µm (G)." figureDoi="10.3897/zookeys.1020.57921.figure31" httpUri="https://binary.pensoft.net/fig/513299" pageId="0" pageNumber="1">Fig. 31A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Prostomium round anteriorly, elongated posteriorly (keel), extending to middle or the end of chaetiger 1, without appendages. Prostomial peaks and eyes absent. Peristomium fused to first chaetiger, with golden pigments dorsally along posterior margin of the prostomium. Dorsal crests and interparapodial pouches absent. Cirriform branchiae on chaetigers 3 and 4, small, often partially hidden by parapodial dorsal lamellae to which they are fused basally. Dorsal lamellae large and foliaceous from chaetigers 2-6, smaller and round thereafter. Particularly long capillaries present in anterior chaetigers. Multidentate long-shafted hooded hooks present in noto- and neuropodia, in neuropodia starting on chaetigers 10, much later in notopodia according to original description. Sabre chaetae from chaetiger 10.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Diagnostic characters are not consistently observable in all specimens due to their poor condition (all anterior fragments, longest anterior fragment with 19 chaetigers). Notopodial hooks were not present in examined specimens (all short anterior fragments). In some specimens a few branchiae were still present and the start of sabre chaetae and neuropodial hooks on chaetigers 10 could be observed. We here tentatively identify the examined specimens as
<taxonomicName authorityName="Maciolek" authorityYear="1981" class="Polychaeta" family="Spionidae" genus="Aurospio" kingdom="Animalia" lsidName="Aurospio dibranchiata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="dibranchiata">Aurospio cf. dibranchiata</taxonomicName>
Maciolek, 1981. The species is known to occur in deep waters of the Atlantic and central Pacific Oceans, but has not been reported yet from near Australia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="records">
<paragraph pageId="0" pageNumber="1">Records.</paragraph>
<paragraph pageId="0" pageNumber="1">8 specimens. Suppl. material 1: ops. 16, 27, 31, 42, 79 (AM). 10 specimens. Suppl. material 1: ops. 23, 27, 31, 76 (NHMUK).</paragraph>
<caption doi="10.3897/zookeys.1020.57921.figure31" httpUri="https://binary.pensoft.net/fig/513299" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
<taxonomicName authorityName="Gunton &amp; Kupriyanova &amp; Alvestad &amp; Avery &amp; Blake &amp; Biriukova &amp; Böggemann &amp; Borisova &amp; Budaeva &amp; Burghardt &amp; Capa &amp; Georgieva &amp; Glasby &amp; Hsueh &amp; Hutchings &amp; Jimi &amp; Kongsrud &amp; Langeneck &amp; Meißner &amp; Murray &amp; Nikolic &amp; Paxton &amp; Ramos &amp; Schulze &amp; Sobczyk &amp; Watson &amp; Wiklund &amp; Wilson &amp; Zhadan &amp; Zhang" authorityYear="2021" family="Spionidae" kingdom="Animalia" lsidName="" order="Phyllodocida" pageId="0" pageNumber="1" rank="family">Spionidae</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Maciolek" authorityYear="1981" class="Polychaeta" family="Spionidae" genus="Aurospio" kingdom="Animalia" lsidName="Aurospio dibranchiata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="dibranchiata">Aurospio cf. dibranchiata</taxonomicName>
: anterior end, dorsal view (AM W.52242)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName class="Polychaeta" family="Spionidae" genus="Aurospio" kingdom="Animalia" lsidName="Aurospio" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Aurospio</emphasis>
</taxonomicName>
sp. nov. 1, anterior end, dorsal view (AM W.52240)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName authorityName="Neal &amp; Altamira in Paterson, Neal, Altamira, Soto, Smith, Menot, Billett, Cunha, Marchais-Laguionie and Glover" authorityYear="2016" class="Polychaeta" family="Spionidae" genus="Prionospio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prionospio amarsupiata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="amarsupiata">Prionospio cf. amarsupiata</taxonomicName>
, anterior end, dorsal view (AM W.52221)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName baseAuthorityName="Blake &amp; Kudenov" baseAuthorityYear="1978" class="Polychaeta" family="Spionidae" genus="Dipolydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipolydora notialis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="notialis">
<emphasis italics="true" pageId="0" pageNumber="1">Dipolydora notialis</emphasis>
</taxonomicName>
, heavy spine with crest of bristles on convex side from chaetiger 5 (AM W.52245)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName baseAuthorityName="Blake &amp; Kudenov" baseAuthorityYear="1978" class="Polychaeta" family="Spionidae" genus="Dipolydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipolydora notialis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="notialis">
<emphasis italics="true" pageId="0" pageNumber="1">Dipolydora notialis</emphasis>
</taxonomicName>
, bidentate hooded hooks with smooth, curved shafts without constriction from posterior chaetiger (AM W.52245)
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Sikorski &amp; Jirkov in Sikorski, Jirkov and Tzetlin" authorityYear="1988" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" lsidName="Laonice blakei" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="blakei">Laonice cf. blakei</taxonomicName>
: anterior end, dorsal-oblique view (AM W.52226)
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName authorityName="Hartman" authorityYear="1960" class="Polychaeta" family="Spionidae" genus="Spiophanes" kingdom="Animalia" lsidName="Spiophanes anoculata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="anoculata">
<emphasis italics="true" pageId="0" pageNumber="1">Spiophanes anoculata</emphasis>
</taxonomicName>
, posteriorly incomplete specimen in dorsal-oblique view, metameric dorsal ciliated organ bordered by pigment of dark orange or ochre colour (AM W.52222). Scale bars: 100
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A, B, C</emphasis>
); 1
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">D, E</emphasis>
); 200
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); 500
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,100 @@
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<bibRefCitation author="Kocak, AOe" pageId="453" pageNumber="454" refId="B407" refString="Kocak, AOe, Kemal, M, 2008. New synonyms and replacement names in the genus group taxa of Araneida. Centre for Entomological Studies Ankara, Miscellaneous Papers 139-140: 1-4." title="New synonyms and replacement names in the genus group taxa of Araneida. Centre for Entomological Studies Ankara, Miscellaneous Papers 139 - 140: 1 - 4." year="2008">
<normalizedToken originalValue="Koçak">Kocak</normalizedToken>
and Kemal 2008
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: 3 [T]
</paragraph>
<paragraph pageId="384" pageNumber="385">
<taxonomicName class="Arachnida" family="Theridiidae" genus="Theridium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theridium antonii" order="Araneae" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="antonii">Theridium antonii</taxonomicName>
Keyserling, 1884;
<bibRefCitation author="Banks, N" pageId="433" pageNumber="434" publicationUrl="10.5479/si.03629236.72.i" refId="B26" refString="Banks, N, 1910. Catalogue of nearctic spiders. Bulletin of the United States National Museum 72: 1-80. doi: 10.5479/si.03629236.72.i" title="Catalogue of nearctic spiders. Bulletin of the United States National Museum 72: 1 - 80. doi:" url="10.5479/si.03629236.72.i" year="1910">Banks 1910</bibRefCitation>
: 19;
<bibRefCitation author="Marx, G" pageId="459" pageNumber="460" publicationUrl="10.5479/si.00963801.782.497" refId="B506" refString="Marx, G, 1890. Catalogue of the described Araneae of temperate North America. Proceedings of the United States National Museum 12: 497-594. doi: 10.5479/si.00963801.782.497" title="Catalogue of the described Araneae of temperate North America. Proceedings of the United States National Museum 12: 497 - 594. doi:" url="10.5479/si.00963801.782.497" year="1890">Marx 1890</bibRefCitation>
: 519
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<paragraph pageId="384" pageNumber="385">
<taxonomicName class="Arachnida" family="Theridiidae" genus="Theridion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridion antoni" order="Araneae" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="antoni">Theridion antoni</taxonomicName>
Keyserling, 1884;
<bibRefCitation author="Petrunkevitch, A" pageId="461" pageNumber="462" publicationUrl="10.5962/bhl.title.23819" refId="B567" refString="Petrunkevitch, A, 1911. A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History 29: 1-791. doi: 10.5962/bhl.title.23819" title="A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History 29: 1 - 791. doi:" url="10.5962/bhl.title.23819" year="1911">Petrunkevitch 1911</bibRefCitation>
: 191
</paragraph>
<paragraph pageId="384" pageNumber="385">
<taxonomicName class="Arachnida" family="Theridiidae" genus="Theridion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridion antonii" order="Araneae" pageId="384" pageNumber="385" phylum="Arthropoda" rank="species" species="antonii">Theridion antonii</taxonomicName>
Keyserling, 1884;
<bibRefCitation author="Fox, I" pageId="445" pageNumber="446" refId="B264" refString="Fox, I, 1940. Notes on nearctic spiders chiefly of the family Theridiidae. Proceedings of the Biological Society of Washington 53: 39-45." title="Notes on nearctic spiders chiefly of the family Theridiidae. Proceedings of the Biological Society of Washington 53: 39 - 45." year="1940">Fox 1940</bibRefCitation>
: 42;
<bibRefCitation author="Jackman, JA" pageId="451" pageNumber="452" refId="B378" refString="Jackman, JA, 1997. A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." title="A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." year="1997">Jackman 1997</bibRefCitation>
: 169;
<bibRefCitation author="Levi, HW" pageId="454" pageNumber="455" refId="B427" refString="Levi, HW, 1957a. The spider genera Enoplognatha, Theridion, and Paidisca in America north of Mexico (Araneae, Theridiidae). Bulletin of the American Museum of Natural History 112: 1-123." title="The spider genera Enoplognatha, Theridion, and Paidisca in America north of Mexico (Araneae, Theridiidae). Bulletin of the American Museum of Natural History 112: 1 - 123." year="1957 a">Levi 1957a</bibRefCitation>
: 60, mf, desc. (figs 196-197, 205, 215-216, 219-220);
<bibRefCitation author="Levi, HW" pageId="457" pageNumber="458" refId="B474" refString="Levi, HW, Randolph, DE, 1975. A key and checklist of American spiders of the family Theridiidae north of Mexico (Araneae). Journal of Arachnology 3: 31-51." title="A key and checklist of American spiders of the family Theridiidae north of Mexico (Araneae). Journal of Arachnology 3: 31 - 51." year="1975">Levi and Randolph 1975</bibRefCitation>
: 43;
<bibRefCitation author="Roewer, CF" pageId="467" pageNumber="468" refId="B679" refString="Roewer, CF, 1942. Katalog der Araneae von 1758 bis 1940. Bremen, Germany, volume 1." title="Katalog der Araneae von 1758 bis 1940. Bremen, Germany, volume 1." year="1942">Roewer 1942</bibRefCitation>
: 501
</paragraph>
</subSubSection>
<subSubSection pageId="384" pageNumber="385" type="distribution">
<paragraph pageId="384" pageNumber="385">Distribution.</paragraph>
<paragraph pageId="384" pageNumber="385">Bexar</paragraph>
</subSubSection>
<subSubSection pageId="384" pageNumber="385" type="type">
<paragraph pageId="384" pageNumber="385">Type.</paragraph>
<paragraph pageId="384" pageNumber="385">Texas (male, Bexar Co., San Antonio, no date, no collector, holotype, USNM)</paragraph>
</subSubSection>
<subSubSection pageId="384" pageNumber="385" type="etymology">
<paragraph pageId="384" pageNumber="385">Etymology.</paragraph>
<paragraph pageId="384" pageNumber="385">locality (city)</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,209 @@
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<mods:namePart>Szumik, Claudia</mods:namePart>
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</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pereyra, Veronica</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Szumik, Victoria E. Goloboff</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina &amp; Facultad de Ciencias Naturales e Instituto Miguel Lillo, Universidad Nacional de Tucuman, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Costa-Pinto, Paula Jessica</mods:namePart>
<mods:affiliation>Instituto Nacional de Pesquisas da Amazonia, INPA, Manaus, Amazonas, Brazil &amp; Programa de Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Juarez, Maria Laura</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina</mods:affiliation>
<mods:nameIdentifier type="email">lau_zoo@yahoo.com.ar</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:identifier type="ZooBank">015929874A644BA89D830B4A2F162E27</mods:identifier>
<mods:identifier type="Pensoft-UUID">A9678B3356615CE9BE576ADD2001E4A0</mods:identifier>
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<treatment ID-GBIF-Taxon="193943948" LSID="urn:lsid:plazi:treatment:0C06867EE88657649F89BB53222A868F" httpUri="http://treatment.plazi.org/id/0C06867EE88657649F89BB53222A868F" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
Genus
<taxonomicName LSID="0C06867E-E886-5764-9F89-BB53222A868F" authority="Davis, 1940" authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embolyntha Davis, 1940</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="reference_group">
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Gray" authorityYear="1832" class="Insecta" family="Archembiidae" genus="Embius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Embius" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embius</taxonomicName>
? Griffith &amp; Pidgeon, 1832: 786, name and illustration, type species
<taxonomicName class="Insecta" family="Archembiidae" genus="Embius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Embius brasilinesis" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="brasilinesis">Embius brasilinesis</taxonomicName>
Gray.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Gray" authorityYear="1832" class="Insecta" family="Archembiidae" genus="Olyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Olyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Olyntha</taxonomicName>
Griffith &amp; Pidgeon, 1832: 347, as subgenus of
<taxonomicName class="Insecta" family="Archembiidae" genus="Embia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embia</taxonomicName>
Latreille, type species
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1832" class="Insecta" family="Archembiidae" genus="Olyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Olyntha brasiliensis" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="brasiliensis">Olyntha brasiliensis</taxonomicName>
Griffith and Pidgeon;
<bibRefCitation DOI="https://doi.org/10.1111/j.1095-8339.1834.tb00029.x" author="Westwood, JO" journalOrPublisher="Transactions of the Linnean Society of London (Zoology)" pageId="0" pageNumber="129" pagination="369 - 374" refId="B58" refString="Westwood, JO, 1837. Characters of Embia, a genus of Insects allied to the white ants (Termites); with descriptions of the species of which it is composed. Transactions of the Linnean Society of London (Zoology) 17: 369 - 374, DOI: https://doi.org/10.1111/j.1095-8339.1834.tb00029.x" title="Characters of Embia, a genus of Insects allied to the white ants (Termites); with descriptions of the species of which it is composed." url="https://doi.org/10.1111/j.1095-8339.1834.tb00029.x" volume="17" year="1837">Westwood 1837</bibRefCitation>
: 373, redescription;
<bibRefCitation author="Burmeister, H" journalOrPublisher="Berlin" pageId="0" pageNumber="129" pagination="757 - 1050" refId="B1" refString="Burmeister, H, 1839. Handbuch der Entomologie. Berlin 2: 757 - 1050" title="Handbuch der Entomologie." volume="2" year="1839">Burmeister 1839</bibRefCitation>
: 770, distinctive characters;
<bibRefCitation author="Walker, F" journalOrPublisher="London, Part 3" pageId="0" pageNumber="129" refId="B57" refString="Walker, F, 1853. Catalogue of the specimens of Neuropterous Insects in the collection of the British Museum. London, Part 3" title="Catalogue of the specimens of Neuropterous Insects in the collection of the British Museum." year="1853">Walker 1853</bibRefCitation>
: 532, catalog, diagnosis, species included;
<bibRefCitation author="Krauss, HA" journalOrPublisher="Zoologica (Stuttgart)" pageId="0" pageNumber="129" pagination="1 - 78" refId="B20" refString="Krauss, HA, 1911. Monographie der Embien. Zoologica (Stuttgart) 23: 1 - 78" title="Monographie der Embien." volume="23" year="1911">Krauss 1911</bibRefCitation>
: 27, erected to genus;
<bibRefCitation author="Enderlein, G" journalOrPublisher="Hayez, imprimeur de l'Academie Royale de Belgique" pageId="0" pageNumber="129" pagination="1 - 121" refId="B14" refString="Enderlein, G, 1912. Embiidinen. Collections zoologiques deu Baron E. de S. Longchamps. Catalogue Systematique et Descriptif. Bruxelles, M. Hayez, imprimeur de l'Academie Royale de Belgique 3: 1 - 121" title="Embiidinen. Collections zoologiques deu Baron E. de S. Longchamps. Catalogue Systematique et Descriptif. Bruxelles, M." volume="3" year="1912">Enderlein 1912</bibRefCitation>
: 29, as junior synonym of
<taxonomicName class="Insecta" family="Archembiidae" genus="Embia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embia</taxonomicName>
;
<bibRefCitation author="Davis, C" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="129" pagination="323 - 344" refId="B8" refString="Davis, C, 1940b. Taxonomic notes on the Order Embioptera. Part XVI: The genus Embia Latreille. Proceedings of the Linnean Society of New South Wales 65: 323 - 344" title="Taxonomic notes on the Order Embioptera. Part XVI: The genus Embia Latreille." volume="65" year="1940 b">Davis 1940b</bibRefCitation>
: 323, 324, invalid name preoccupied by a genus of
<taxonomicName higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="129" rank="order">Coleoptera</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embolyntha</taxonomicName>
Davis, 1940c: 344, description and diagnosis, key to the species, type species
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1832" class="Insecta" family="Archembiidae" genus="Olyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Olyntha brasiliensis" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="brasiliensis">Olyntha brasiliensis</taxonomicName>
,
<taxonomicName authorityName="Gray" authorityYear="1832" class="Insecta" family="Archembiidae" genus="Embius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Embius" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embius</taxonomicName>
and
<taxonomicName authorityName="Gray" authorityYear="1832" class="Insecta" family="Archembiidae" genus="Olyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Olyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Olyntha</taxonomicName>
preoccupied names;
<bibRefCitation DOI="https://doi.org/10.5479/si.00963801.94-3175.401" author="Ross, ES" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="129" pagination="401 - 504" refId="B31" refString="Ross, ES, 1944. A revision of the Embioptera or Web-Spinners, of the New World. Proceedings of the Entomological Society of Washington 94: 401 - 504, DOI: https://doi.org/10.5479/si.00963801.94-3175.401" title="A revision of the Embioptera or Web-Spinners, of the New World." url="https://doi.org/10.5479/si.00963801.94-3175.401" volume="94" year="1944">Ross 1944</bibRefCitation>
: 412, discussion;
<bibRefCitation author="Ross, ES" journalOrPublisher="The Wasmann Journal of Biology" pageId="0" pageNumber="129" pagination="29 - 36" refId="B34" refString="Ross, ES, 1971. A new neotropical genus and species of Embioptera. The Wasmann Journal of Biology 29: 29 - 36" title="A new neotropical genus and species of Embioptera." volume="29" year="1971">Ross 1971</bibRefCitation>
: 29, genus restricted only to type species;
<bibRefCitation author="Szumik, C" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="129" pagination="140 - 153" refId="B45" refString="Szumik, C, 1997. Two new neotropical genera of Embiidae (Embioptera, Insecta). Journal of the New York Entomological Society 105: 140 - 153" title="Two new neotropical genera of Embiidae (Embioptera, Insecta)." volume="105" year="1997">Szumik 1997</bibRefCitation>
: 141, phylogeny;
<bibRefCitation author="Szumik, C" editor="Coscaron, S" journalOrPublisher="Ediciones del Sur, Argentina" pageId="0" pageNumber="129" pagination="32 - 37" refId="B46" refString="Szumik, C, 1998a. Capitulo 4: Embioptera. In: Coscaron, S, Morrone, J, Eds., Biodiversidad de artropodos argentinos. Ediciones del Sur, Argentina: 32 - 37" title="Capitulo 4: Embioptera." volumeTitle="Biodiversidad de artropodos argentinos." year="1998 a">Szumik 1998a</bibRefCitation>
: 34, genera key;
<bibRefCitation author="Ross, ES" journalOrPublisher="Occasional Papers of the California Academy of Sciences" pageId="0" pageNumber="129" pagination="1 - 86" refId="B39" refString="Ross, ES, 2001. Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina). Occasional Papers of the California Academy of Sciences 150: 1 - 86" title="Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina)." volume="150" year="2001">Ross 2001</bibRefCitation>
: 26, diagnosis;
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
: 226, diagnosis, composition, phylogeny;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-0031.2008.00228.x" author="Szumik, C" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="993 - 1005" refId="B52" refString="Szumik, C, Edgerly, J, Hayashi, C, 2008. Phylogeny of Embiopterans (Insecta). Cladistics 24: 993 - 1005, DOI: https://doi.org/10.1111/j.1096-0031.2008.00228.x" title="Phylogeny of Embiopterans (Insecta)." url="https://doi.org/10.1111/j.1096-0031.2008.00228.x" volume="24" year="2008">Szumik et al. 2008</bibRefCitation>
: 997, phylogeny;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2055.1.1" author="Miller, KB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 34" refId="B23" refString="Miller, KB, 2009. Genus -and family-group names in the order Embioptera (Insecta). Zootaxa 2055: 1 - 34, DOI: https://doi.org/10.11646/zootaxa.2055.1.1" title="Genus - and family-group names in the order Embioptera (Insecta)." url="https://doi.org/10.11646/zootaxa.2055.1.1" volume="2055" year="2009">Miller 2009</bibRefCitation>
: 10, catalog;
<bibRefCitation author="Szumik, C" editor="Rafael, JA" journalOrPublisher="Holos Editora, Ribeirao Preto" pageId="0" pageNumber="129" pagination="348 - 354" refId="B50" refString="Szumik, C, 2012. Capitulo 20. Embioptera. In: Rafael, JA, Melo, GAR, Carvalho, CJB, Casari, SAC, Constatino, R, Eds., Insetos do Brasil, Diversidade e Taxonomia. Holos Editora, Ribeirao Preto: 348 - 354" title="Capitulo 20. Embioptera." volumeTitle="Insetos do Brasil, Diversidade e Taxonomia." year="2012">Szumik 2012</bibRefCitation>
: 352, list of genera from Brazil;
<bibRefCitation DOI="https://doi.org/10.1206/3933.1" author="Szumik, C" journalOrPublisher="Novitates" pageId="0" pageNumber="129" pagination="1 - 31" refId="B54" refString="Szumik, C, Juarez, ML, Ramirez, MJ, Goloboff, P, Pereyra, V, 2019. Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera. Novitates 3933: 1 - 31, DOI: https://doi.org/10.1206/3933.1" title="Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera." url="https://doi.org/10.1206/3933.1" volume="3933" year="2019">Szumik et al. 2019</bibRefCitation>
: 9, tympanal hearing, silk ejectors, leg chaetotaxy, phylogeny.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Argocercembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argocercembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Argocercembia</taxonomicName>
Ross, 2001: 63, type species
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Argocercembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argocercembia guyana" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="guyana">Argocercembia guyana</taxonomicName>
Ross;
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
: 226, junior synonym of
<taxonomicName authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Embolyntha</taxonomicName>
Davis.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Embolyntha</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" kingdom="Animalia" lsidName="" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="family">Archembiidae</taxonomicName>
by Sm with anterior margin membranous, 10Lp1 simple and starting at inner caudal angle of 10L, 10Lp1 leaf-like with many longitudinal carinae (
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Composition and distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Embolyntha</emphasis>
</taxonomicName>
has two species (
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
):
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1832" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha brasiliensis" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis italics="true" pageId="0" pageNumber="129">Embolyntha brasiliensis</emphasis>
</taxonomicName>
(Gray, 1832) from Brazil (without any specific location) and
<taxonomicName baseAuthorityName="Ross" baseAuthorityYear="2001" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha guyana" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="guyana">
<emphasis italics="true" pageId="0" pageNumber="129">Embolyntha guyana</emphasis>
</taxonomicName>
(Ross, 2001) from Guyana and Brazil (see Catalog for details). In addition, one new species from Brazil is described below.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">South America.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="relationships">
<paragraph pageId="0" pageNumber="129">Relationships.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Argocercembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argocercembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Argocercembia</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName authorityName="Davis" authorityYear="1940" class="Insecta" family="Archembiidae" genus="Embolyntha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolyntha" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Embolyntha</emphasis>
</taxonomicName>
by
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik (2004)</bibRefCitation>
because of the minimal differences between them. Both taxa appeared grouped together by sharing 10Lp1 shape, a small medial bladder in males, and LC1dp in a medial position, both forming a well-supported group. In the higher classification of the Order (
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-0031.2008.00228.x" author="Szumik, C" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="993 - 1005" refId="B52" refString="Szumik, C, Edgerly, J, Hayashi, C, 2008. Phylogeny of Embiopterans (Insecta). Cladistics 24: 993 - 1005, DOI: https://doi.org/10.1111/j.1096-0031.2008.00228.x" title="Phylogeny of Embiopterans (Insecta)." url="https://doi.org/10.1111/j.1096-0031.2008.00228.x" volume="24" year="2008">Szumik et al. 2008</bibRefCitation>
: 927), the genus remains monophyletic and as sister group of the monotypic genus
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Scelembiidae" genus="Ochrembia" kingdom="Animalia" lsidName="Ochrembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Ochrembia</emphasis>
</taxonomicName>
. Finally, these groups remain supported in a more recent study (
<bibRefCitation DOI="https://doi.org/10.1206/3933.1" author="Szumik, C" journalOrPublisher="Novitates" pageId="0" pageNumber="129" pagination="1 - 31" refId="B54" refString="Szumik, C, Juarez, ML, Ramirez, MJ, Goloboff, P, Pereyra, V, 2019. Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera. Novitates 3933: 1 - 31, DOI: https://doi.org/10.1206/3933.1" title="Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera." url="https://doi.org/10.1206/3933.1" volume="3933" year="2019">Szumik et al. 2019</bibRefCitation>
: 9) where new evidence from ultrastructure traits on leg chaetotaxy as well as the chordotonal organ were included.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,370 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.979.52059" ID-GBIF-Dataset="85bb4820-5880-44f0-82d3-163f1a684cb0" ID-PMC="PMC7609529" ID-Pensoft-Pub="1313-2970-979-35" ID-Pensoft-UUID="2F4B35A2CA7C5E0E9B1AB671EE37390C" ID-PubMed="33192131" ID-ZooBank="2B5B9363B75C4392A23A78186989B7F3" ModsDocID="1313-2970-979-35" checkinTime="1603838524667" checkinUser="pensoft" docAuthor="Gadallah, Neveen S., Soliman, Ahmed M. &amp; Dhafer, Hathal M. Al" docDate="2020" docId="0C0687C8CEE8524E810C85BE7763F7B1" docLanguage="en" docName="ZooKeys 979: 35-86" docOrigin="ZooKeys 979" docSource="http://dx.doi.org/10.3897/zookeys.979.52059" docTitle="Epitranus similis Gadallah &amp; Soliman 2020, sp. nov." docType="treatment" docUuid="A190D08C-92B7-4662-9660-B1C642740266" docUuidSource="ZooBank" docVersion="4" id="2F4B35A2CA7C5E0E9B1AB671EE37390C" lastPageNumber="35" masterDocId="2F4B35A2CA7C5E0E9B1AB671EE37390C" masterDocTitle="First record of the subfamily Epitraninae from Saudi Arabia (Hymenoptera, Chalcidoidea, Chalcididae), with the description of three new species" masterLastPageNumber="86" masterPageNumber="35" pageNumber="35" updateTime="1668169650305" updateUser="ExternalLinkService">
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<mods:title>First record of the subfamily Epitraninae from Saudi Arabia (Hymenoptera, Chalcidoidea, Chalcididae), with the description of three new species</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gadallah, Neveen S.</mods:namePart>
<mods:affiliation>Entomology Department, Faculty of Science, Cairo University, Giza, Egypt</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soliman, Ahmed M.</mods:namePart>
<mods:affiliation>Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. BOX 2460, Riyadh 11451, Saudi Arabia &amp; Zoology Department, Faculty of Science (Boys), Al-Azhar University, P. O. Box 11884, Nasr City, Cairo, Egypt</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5284-713X</mods:nameIdentifier>
<mods:nameIdentifier type="email">ammsoliman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dhafer, Hathal M. Al</mods:namePart>
<mods:affiliation>Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. BOX 2460, Riyadh 11451, Saudi Arabia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4911-2332</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>979</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>35</mods:start>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="168650525" LSID="urn:lsid:zoobank.org:act:A190D08C-92B7-4662-9660-B1C642740266" httpUri="http://treatment.plazi.org/id/0C0687C8CEE8524E810C85BE7763F7B1" lastPageNumber="35" pageId="0" pageNumber="35">
<subSubSection pageId="0" pageNumber="35" type="nomenclature">
<paragraph pageId="0" pageNumber="35">
<taxonomicName LSID="0C0687C8-CEE8-524E-810C-85BE7763F7B1" authority="Gadallah &amp; Soliman" authorityName="Gadallah &amp; Soliman" authorityYear="2020" class="Insecta" family="Chalcididae" genus="Epitranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epitranus similis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">Epitranus similis Gadallah &amp; Soliman</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">Figures 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B metasoma (dorsal and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure9" httpUri="https://binary.pensoft.net/fig/467086" pageId="0" pageNumber="35">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="type material">
<paragraph pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
♂.
<emphasis bold="true" pageId="0" pageNumber="35">Kingdom of Saudi Arabia</emphasis>
, Asir, Abha, Garf Raydah Natural Reserve [
<geoCoordinate degrees="18" direction="north" minutes="11" orientation="latitude" precision="1" seconds="35.74" value="18.19326">18°11'35.74&quot;N</geoCoordinate>
,
<geoCoordinate degrees="42" direction="east" minutes="23" orientation="longitude" precision="1" seconds="30.24" value="42.391735">42°23'30.24&quot;E</geoCoordinate>
, Alt. 1805 m], sweeping net, 5.IX.2015, leg. Ahmed M. Soliman [KSMA];
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Paratypes</emphasis>
: 2♂,
<emphasis bold="true" pageId="0" pageNumber="35">Kingdom of Saudi Arabia</emphasis>
, same data as for holotype [KSMA].
</paragraph>
<caption doi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figure 6.</emphasis>
<taxonomicName authorityName="Gadallah &amp; Soliman" authorityYear="2020" class="Insecta" family="Chalcididae" genus="Epitranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epitranus similis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="35">Epitranus similis</emphasis>
</taxonomicName>
Gadallah &amp; Soliman, sp. nov. (holotype male)
<emphasis bold="true" pageId="0" pageNumber="35">A, B, C</emphasis>
habitus (lateral, dorsal, and ventral views respectively).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="diagnosis">
<paragraph pageId="0" pageNumber="35">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="35">
Frontal lobe distinctly long, sub-trapezoidal in shape as its sides are straight and regularly converging ventrally, with a longitudinal median carina extending on its whole length, and free margin with three lobes (outer lobes notched subapically) (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7B</figureCitation>
); OOL 0.80-0.85
<normalizedToken originalValue="×">x</normalizedToken>
as long as OD, and ca. 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as AOL (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7C</figureCitation>
); POL ca. 4.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as OOL (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7C</figureCitation>
); F1 moderately long, ca. 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7D</figureCitation>
); pronotal collar laterally, scapula anterolaterally, propodeum on prestigmatic areola, mesepisternum, metepimeron dorsally and basoventral surface of metacoxa densely clothed with whitish setae masking integument beneath (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">6A, C</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8B</figureCitation>
); pronotal collum with dense, long golden setae on two submedian patches (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7C</figureCitation>
); propodeal median areola deep, with lateral ridges converging posteriorly to meet before the adpetiolar areola (Y-like raised carina) (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8A</figureCitation>
); metafemur toothed ventrally, with nine or ten spaced teeth following the stout sub-basal tooth (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8D, E</figureCitation>
); tarsal groove of metatibia fully occupying the completely delimited smooth and deep metatibial process, reaching the sub-basal prominence anteriorly (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8E</figureCitation>
); petiole very long, 7.12-7.65
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B metasoma (dorsal and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure9" httpUri="https://binary.pensoft.net/fig/467086" pageId="0" pageNumber="35">9A</figureCitation>
); gaster relatively short (1.25-1.40
<normalizedToken originalValue="×">x</normalizedToken>
as long as height in profile) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">6A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figure 7.</emphasis>
<taxonomicName authorityName="Gadallah &amp; Soliman" authorityYear="2020" class="Insecta" family="Chalcididae" genus="Epitranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epitranus similis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="35">Epitranus similis</emphasis>
</taxonomicName>
Gadallah &amp; Soliman, sp. nov. (holotype male)
<emphasis bold="true" pageId="0" pageNumber="35">A</emphasis>
head (frontal view)
<emphasis bold="true" pageId="0" pageNumber="35">B</emphasis>
lower part of face showing frontal lobe (frontal view)
<emphasis bold="true" pageId="0" pageNumber="35">C</emphasis>
head &amp; pronotum (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="35">D</emphasis>
antennal pedicel and flagellum
<emphasis bold="true" pageId="0" pageNumber="35">E</emphasis>
mesosoma and part of gastral petiole (dorsal view).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="description">
<paragraph pageId="0" pageNumber="35">Description.</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Male</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="35">holotype</emphasis>
). Body length 5.5 mm. fore wing length 3.3 mm.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Head</emphasis>
(Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7A-D</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8B</figureCitation>
). Triangular in frontal view, 1.25
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high, wider than mesoscutum in dorsal view (1.2
<normalizedToken originalValue="×">x</normalizedToken>
), 2.6
<normalizedToken originalValue="×">x</normalizedToken>
as wide as its length in profile. Frontovertex 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as eye height; AOL 2.0
<normalizedToken originalValue="×">x</normalizedToken>
OOL; OOL 0.85
<normalizedToken originalValue="×">x</normalizedToken>
OD; POL 4.3
<normalizedToken originalValue="×">x</normalizedToken>
OOL; supra antennal surface absent; frons transversely finely strigulate medially beneath antennal scape (at scrobe), laterally with sparse setiferous punctures, the setae lanceolate and long; preorbital carina absent; malar area mostly polished, with scattered superficial setiferous punctures; malar space ca. 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long eye height in profile; malar carina absent; gena broad, nearly smooth, with a row of setae directed inwards along post-orbital carina that is well-developed, lamellate and joining genal carina at a level of ventral edge of eye; post-orbital carina hardly converging to the higher edge of the eye (nearly parallel); suborbital carina weak. Occiput alutaceous, with sparse setiferous punctures (setae long and dispersed, pale yellow). Interantennal projection absent; interantennal distance ca. 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as torulus diameter; frontal lobe quite long, with subantennal distance 4.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as interantennal distance, ventral margin with two pairs of indentations delimiting three lobes, the outer ones are narrowly rounded; projection with a sharp longitudinal median carina extended throughout its length; surface of projection with long, lanceolate and whitish setae.
</paragraph>
<caption doi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figure 8.</emphasis>
<taxonomicName authorityName="Gadallah &amp; Soliman" authorityYear="2020" class="Insecta" family="Chalcididae" genus="Epitranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epitranus similis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="35">Epitranus similis</emphasis>
</taxonomicName>
Gadallah &amp; Soliman, sp. nov. (holotype male)
<emphasis bold="true" pageId="0" pageNumber="35">A</emphasis>
mesoscutellum, metanotum &amp; propodeum (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="35">B</emphasis>
head &amp; mesosoma (lateral view)
<emphasis bold="true" pageId="0" pageNumber="35">C</emphasis>
fore wing (parts of wing membrane and MV and STV magnified)
<emphasis bold="true" pageId="0" pageNumber="35">D, E</emphasis>
hind leg, excluding coxa (outer and ventral views respectively).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7D</figureCitation>
). Scape 1.35
<normalizedToken originalValue="×">x</normalizedToken>
as long as eye height, ending very closely to median ocellus, its ventral face strongly excavated, the excavation densely and finely pubescent; pedicel hardly longer than wide (1.15
<normalizedToken originalValue="×">x</normalizedToken>
); anellus transverse (0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide); F1 moderate, 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as F2 and F7 as well; clava 2.35
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Mesosoma</emphasis>
(Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A head (frontal view) B lower part of face showing frontal lobe (frontal view) C head &amp; pronotum (dorsal view) D antennal pedicel and flagellum E mesosoma and part of gastral petiole (dorsal view)." figureDoi="10.3897/zookeys.979.52059.figure7" httpUri="https://binary.pensoft.net/fig/467084" pageId="0" pageNumber="35">7C, E</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8A, B</figureCitation>
). 1.65
<normalizedToken originalValue="×">x</normalizedToken>
as long as mesoscutum width. Pronotal collar 2.45
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long, mostly smooth on disc, with scattered punctures bearing fine setae; collar laterally and collum densely setiferous punctulate, the setae forming tufts that are inwardly oriented, white on the former and golden yellow on the latter; pronotal sides slightly convex; lateral carinae sharp and extending dorsally, not meeting medially; humeral angle rounded; pronotal lateral panel finely alutaceous, shiny. Mesoscutum 2.25
<normalizedToken originalValue="×">x</normalizedToken>
as long as median length of pronotum, middle lobe finely alutaceous anteriorly, followed by deep and sparse setiferous punctures bearing fine setae; scapula nearly smooth, with few scattered punctures, antero-laterally clothed with dense lanceolate whitish setae; notauli deep, foveolate; axilla with dense, upwardly directed whitish setae, integument smooth beneath; tegula large, depressed near posterior margin, smooth anteriorly, finely alutaceous posteriorly and laterally, with broad angulate posterior margin that distinctly overlap base of hind wing. Mesoscutellum convex, hardly longer than wide (1.1
<normalizedToken originalValue="×">x</normalizedToken>
), irregularly deeply foveate, the foveae large, widely separated medially and closer laterally, posterior margin broadly rounded, strigose. Propodeum deeply areolate; median areola 1.37
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with lateral ridges converging posteriorly and meeting slightly before adpetiolar areola; lateral areola transversely carinate; prestigmatic areola densely setose, setae oriented inwards. Adscrobal area of mesopleuron densely clothed with long, suberect and whitish setae, femoral scrobe finely transversely strigose, ventral shelf of mesepisternum sparsely punctate, with adpressed long setae, interspaces between punctures smooth; epicnemial carina lamellate. Metepimeron closely foveolate throughout (bottom of foveae smooth), with dense, long adpressed and whitish setae; metepisternum densely reticulate, with two median carinae diverging posteriorly followed by two large and sharp submedian teeth; subcoxal teeth small; adpetiolar area concave, longitudinally striated, with a median longitudinal carina ends posteriorly with a strong subpentagonal subpetiolar areola.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Wings</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8C</figureCitation>
). Fore wing 3.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, bare on upper side, sparsely setose subapically on underside; MV ca. 0.72
<normalizedToken originalValue="×">x</normalizedToken>
as long as costal cell; STV rudimentary (1.25
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide), strongly diverging from anterior margin of wing at an angle of ca. 80°. Hind wing hyaline and asetose, with three hamuli.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Hind leg</emphasis>
(Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">6C</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8D, E</figureCitation>
). Metacoxa widened basally, 2.45
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as metafemur, densely setose on ventral side, punctured but with transverse ridges near apex on outer dorsal side. Metafemur 1.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with dense setiferous punctures on outer face, its ventral margin with a triangular sub-basal tooth followed by nine teeth that are equally separated, progressively smaller towards apex. Tarsal scrobe of metatibia fully occupying the completely delimited, smooth and deep metatibial process, reaching sub-basal prominence anteriorly (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Epitranus subinops Soliman &amp; Gadallah, sp. nov. (holotype female) A propodeum and metasoma (dorsal view) B head including antennal scape and mesosoma (lateral view) D fore wing (parts of wing membrane and MV and STV magnified)." figureDoi="10.3897/zookeys.979.52059.figure12" httpUri="https://binary.pensoft.net/fig/467089" pageId="0" pageNumber="35">12E</figureCitation>
), the edge of the later with four denticles concealed by pubescence (could be seen when examined from dorsal view).
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Metasoma</emphasis>
(Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">6A</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B metasoma (dorsal and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure9" httpUri="https://binary.pensoft.net/fig/467086" pageId="0" pageNumber="35">9A, B</figureCitation>
). Petiole quite long, 7.12
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, 1.58
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of Gt1 in dorsal view, and 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of gaster in dorsal view, dorsally with two, lateral and sublateral, longitudinal ridges that is vague along the apical two-thirds, the area between sublateral ridges transversely wrinkled. Gaster ovoid in dorsal view, 1.25
<normalizedToken originalValue="×">x</normalizedToken>
as long as its height in lateral view. Gt1 long (0.75
<normalizedToken originalValue="×">x</normalizedToken>
as long as gaster in dorsal view), deeply concave posteriorly, sparsely finely setiferous punctate (setae fine and short, punctures dense postero-laterally); remaining tergites short, sparsely finely punctate and finely setose.
</paragraph>
<caption doi="10.3897/zookeys.979.52059.figure9" httpUri="https://binary.pensoft.net/fig/467086" pageId="0" pageNumber="35" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figure 9.</emphasis>
<taxonomicName authorityName="Gadallah &amp; Soliman" authorityYear="2020" class="Insecta" family="Chalcididae" genus="Epitranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epitranus similis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="35">Epitranus similis</emphasis>
</taxonomicName>
Gadallah &amp; Soliman, sp. nov. (holotype male)
<emphasis bold="true" pageId="0" pageNumber="35">A, B</emphasis>
metasoma (dorsal and ventral views respectively).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Color</emphasis>
(Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A, B, C habitus (lateral, dorsal, and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure6" httpUri="https://binary.pensoft.net/fig/467083" pageId="0" pageNumber="35">6A-C</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Epitranus similis Gadallah &amp; Soliman, sp. nov. (holotype male) A mesoscutellum, metanotum &amp; propodeum (dorsal view) B head &amp; mesosoma (lateral view) C fore wing (parts of wing membrane and MV and STV magnified) D, E hind leg, excluding coxa (outer and ventral views respectively)." figureDoi="10.3897/zookeys.979.52059.figure8" httpUri="https://binary.pensoft.net/fig/467085" pageId="0" pageNumber="35">8C</figureCitation>
). Body black, except the following parts, bright reddish brown: frontal lobe, malar area, a relatively broad strip around eye, gena, pronotum, a lateral longitudinal strip on middle lobe of mesoscutum, lateral and posterior borders and postero-inner corner of scapula, inner part of axilla, area around epicnemial carina on mesopleuron, upper part of metepimeron, and posterior part of adpetiolar area on metepisternum. Gaster reddish brown with black tint dorsally. Antennal scape and pedicel reddish brown, flagellum brown. Legs reddish brown, with black tint on mesofemur, ventral face of metacoxa and outer face of metafemur. Tegula glassy golden yellow. Wings hyaline with brown veins that are paler on hind wing. Genitalia pale yellow.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Female</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="variation">
<paragraph pageId="0" pageNumber="35">Variation.</paragraph>
<paragraph pageId="0" pageNumber="35">The paratype specimens differ from the holotype specimen in the predominance of red brown color on: head (except post-orbital and occipital carinae in one of the paratype specimens or a band along occipital carina, post-orbital carina and a narrow longitudinal median strip on the frons in the other paratype specimen); middle lobe of mesoscutum (except a triangular area on disc); lateral lobe of mesoscutum (except an oval area on disc); the whole axilla, mesoscutellum (except longitudinal median strip); the whole metapleuron; metacoxa and metafemur (except black tint on the former).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="remarks">
<paragraph pageId="0" pageNumber="35">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="35">
The new species is morphologically similar to
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
(Schmitz) (Democratic Republic of Congo) especially the identical frontal projection; the absence of interantennal projection; similar flagellum; the presence of outstanding setae on mesosoma; similar STV, and similar petiole. But differs from it by the partly reddish head and mesosoma (entirely black in
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
); the presence of distinctive setation on different parts of mesosoma as reported above (mesosoma with regular setation in
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
); propodeum with petiolate median areola (complete in
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
); shorter and relatively stouter metacoxa (quite slender in
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="etymology">
<paragraph pageId="0" pageNumber="35">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="35">
The word
<taxonomicName lsidName="similis" pageId="0" pageNumber="35" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="35">similis</emphasis>
</taxonomicName>
is an adjective in Latin and means similar or resembling, referring to the similarity of this species to
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="35" rank="species" species="nitidus">
<emphasis italics="true" pageId="0" pageNumber="35">E. nitidus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="host">
<paragraph pageId="0" pageNumber="35">Hosts.</paragraph>
<paragraph pageId="0" pageNumber="35">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="distribution">
<paragraph pageId="0" pageNumber="35">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="35">Saudi Arabia (Asir region).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,71 @@
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<taxonomicName id="937D7CFC2FACA3FD926134B6BC0D8EF0" ID-CoL="7C2WZ" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Dilleniaceae" genus="Tetracera" kingdom="Plantae" order="Dilleniales" phylum="Tracheophyta" rank="species" species="volubilis" status="spec. nov.">Tetracera volubilis</taxonomicName>
,
<taxonomicNameLabel id="111E99F5603DE43DEB8A96B4380205F7" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
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<paragraph id="B867EBEF5CFC39565991F81D5A91A100" pageNumber="533">
1. TETRACERA.
<emphasis id="424BE99C8968DFC2AA36A6B658F570D7" italics="true">Hort. cliff. 214.</emphasis>
*
</paragraph>
<paragraph id="E41FB461AE932E3F49820122EC6BE9DB" pageNumber="533">
Petraea floribus spicatis, scabris lauri foliis.
<emphasis id="63AED811E004B1E9D20B5626B3872D64" italics="true">Amm. herb. 581.</emphasis>
</paragraph>
<paragraph id="EB7404202CA058B37DDF41DAE98DC385" pageNumber="533">
Fagus americanus, ulmi amplissimis foliis, capsulis bigemellis.
<emphasis id="2AA64E22009318E720190CF2C3F6F9A5" italics="true">Pluk. amalt.87.</emphasis>
</paragraph>
<paragraph id="B17C771C55120269071D66BFC851A9BC" pageNumber="533">
Arbor americana convolvulacea platyphyllos barbadensibus dicta, foliis serratis.
<emphasis id="AF165CC78A099973BEE1DAA565B45AA4" italics="true">Pluk. alm. 48. t.146. f.1.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="DC6E090F7608E5AB74DBFEF69127A8ED" type="distribution">
<paragraph id="8E220381B0FA53D09181607FE0C08F08" pageNumber="533">
<emphasis id="9D0EA16D5E82634BF5769B26A56072D9" italics="true">Habitat in</emphasis>
Barbados
<emphasis id="140712DDC5071C692ACEE574CF47D5EF" italics="true">,</emphasis>
Vera Cruce
<emphasis id="572CD3BCCECB771BC4FF91E3FDEE24F0" italics="true">,</emphasis>
Marilandia
<emphasis id="C794F9CC2A8F2D601C6AAE44F7D36A06" italics="true">. ♄</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BC5A913321ACEFD58C9DF6ADEB379558" type="description">
<paragraph id="0849DAD361A096085EACC6A244F3A5CB" pageNumber="533">
<heading id="29B2C8B0E767AF76934A90869EDE3BB0" level="4">PENTAGYNIA.</heading>
</paragraph>
</subSubSection>
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View file

@ -0,0 +1,130 @@
<document id="FD7EC799E1AA0AAFCE8EAFBEB341C3BE" ID-DOI="10.5281/zenodo.292249" ID-ISBN="3-7643-0556-8" ID-Zenodo-Dep="292249" checkinTime="1646762436682" checkinUser="donat" docAuthor="Hess, Hans Ernst, Landolt, Elias &amp; Hirzel, Rosmarie" docDate="1976" docId="0C072F136A98432037F0D153DDB68EC7" docLanguage="de" docName="gentianaceae.xml" docOrigin="Birkhaeuser Verlag" docSource="https://doi.org/10.5281/zenodo.292249" docTitle="Gentiana insubrica H. Kunz" docType="treatment" docVersion="3" lastPageNumber="36" masterDocId="EE3340D459253352DBE5606A2DCD8699" masterDocTitle="Flora der Schweiz und angrenzender Gebiete. Band 3. Plumbaginaceae bis Compositae (2 nd edition): Gentianaceae" pageId="null" pageNumber="36" sec="gentianaceae.s47" updateTime="1649917633437" updateUser="donat">
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<mods:title id="B6C9F6842F819D723A6E675664ADAC05">Flora der Schweiz und angrenzender Gebiete. Band 3. Plumbaginaceae bis Compositae (2 nd edition): Gentianaceae</mods:title>
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<mods:namePart id="5EB5E934AECF6E70A84EDF549FE7F5CB">Hess, Hans Ernst</mods:namePart>
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</mods:role>
<mods:namePart id="60CC38E3E6F7AB72EDACC5F11FE081AC">Landolt, Elias</mods:namePart>
</mods:name>
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<mods:roleTerm id="122013707890961196BCAAFCF9F25B76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3831C24E558E369AFF1A312D0A71B3E4">Hirzel, Rosmarie</mods:namePart>
</mods:name>
<mods:typeOfResource id="12151222471335B7769B58225F8DA2FA">text</mods:typeOfResource>
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<mods:dateIssued id="055FD5464B9FC9A63F343F80607B8A9C">1976</mods:dateIssued>
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<taxonomicName id="2C873D2CC084DACC01AC5CA34EEBC217" authority="H. Kunz" authorityName="H. Kunz" class="Magnoliopsida" family="Gentianaceae" genus="Gentiana" kingdom="Plantae" order="Gentianales" pageId="null" pageNumber="36" phylum="Tracheophyta" rank="species" species="insubrica">
Gentiana
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H. Kunz
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</subSubSection>
<subSubSection id="215138B239D1FDEBA00293505A7BB843" pageId="null" pageNumber="36" type="reference_group">
<paragraph id="8D16C1D466A6E8DD4CB48ADA1F219785" pageId="null" pageNumber="36">
(
<taxonomicName id="5F159658D367FDCD022494DCC7B4089A" class="Magnoliopsida" family="Gentianaceae" genus="Gentianella" kingdom="Plantae" order="Gentianales" pageId="null" pageNumber="36" phylum="Tracheophyta" rank="species" species="insubrica">
<emphasis id="10E21F506AC8A43C071A99CCB7B4434E" italics="true" pageId="null" pageNumber="36">Gentianella insubrica</emphasis>
</taxonomicName>
[H. Kunz] Holub)
</paragraph>
</subSubSection>
<subSubSection id="F52E38B8D572443B8AA48D08A4744276" pageId="null" pageNumber="36" type="vernacular_names">
<paragraph id="1A7A5D246F3B532FF8DA1A25EF4F0266" pageId="null" pageNumber="36">Insubrischer Enzian</paragraph>
</subSubSection>
</title>
<subSubSection id="E23EA6CCB132384C4761FD5811432090" pageId="null" pageNumber="36" type="description">
<paragraph id="9AEA45045DF0A1C4C4B83644830CD263" pageId="null" pageNumber="36">
10-25 cm hoch.
<emphasis id="0E70079C32E94FC98C922AA5160D19B4" italics="true" pageId="null" pageNumber="36">Kelch bis auf etwa</emphasis>
<normalizedToken id="C235ABAEBEC20EDC02D53CA423B7D544" originalValue="½" pageId="null" pageNumber="36">1/2</normalizedToken>
-⅓
<emphasis id="98ACA7D241D397269DFC4ECF522DCC3D" italics="true" pageId="null" pageNumber="36">
der
<normalizedToken id="9FE24FD02B624AB7D8E42DFCEE5D1BAD" originalValue="Länge" pageId="null" pageNumber="36">Laenge</normalizedToken>
5
</emphasis>
(
<emphasis id="1BC44A31DB3F58F35B62DF9604CA7D56" italics="true" pageId="null" pageNumber="36">selten 4</emphasis>
)
<emphasis id="45FE7C54CCCFA190480E611173B7B769" italics="true" pageId="null" pageNumber="36">teilig</emphasis>
, mit spitzen Buchten; Zipfel lanzettlich, zugespitzt,
<emphasis id="D08EC3EE35ADCDDA25297B972A8824F8" italics="true" pageId="null" pageNumber="36">ungleich</emphasis>
(
<emphasis id="F3960F096F687B380593ED1CAB67AF2F" italics="true" pageId="null" pageNumber="36">1</emphasis>
-
<emphasis id="B1F5A5F3390FFE166E2B4E6AAD97BFB4" italics="true" pageId="null" pageNumber="36">3 Zipfel bedeutend breiter als die andern</emphasis>
),
<emphasis id="0BA4C4B5FA9038B4F317CBE07007ACA9" italics="true" pageId="null" pageNumber="36">
meist
<normalizedToken id="0801F3FDB9133D7CAFC5FCE8E502DF3C" originalValue="länger" pageId="null" pageNumber="36">laenger</normalizedToken>
als die Kronzipfel
</emphasis>
, am Rande meist von etwa 0,1 mm langen Papillen rauh (10fache
<normalizedToken id="DD68EB8E58E3F8E48BA9E46261107DC0" originalValue="Vergrößerung" pageId="null" pageNumber="36">Vergroesserung</normalizedToken>
!) und nach
<normalizedToken id="FECE2AAA1E83EAFCB5106A91E06AB2B2" originalValue="außen" pageId="null" pageNumber="36">aussen</normalizedToken>
umgerollt.
<emphasis id="8341DD0C1E050D26DA796F3E4793C230" italics="true" pageId="null" pageNumber="36">Krone blauviolett</emphasis>
(selten
<normalizedToken id="2E7B155B6CC15CA97259E91B3CFDFFBC" originalValue="weiß" pageId="null" pageNumber="36">weiss</normalizedToken>
), mit 5 (selten 4)
<emphasis id="5CF1E02D77022705D04565AA5AC25E5A" italics="true" pageId="null" pageNumber="36">7</emphasis>
-
<emphasis id="3909350BD04A214DDDC6A32080D7C558" italics="true" pageId="null" pageNumber="36">
10 mm langen Zipfeln. Fruchtknoten und Frucht
<normalizedToken id="EA5B5510323D87E80C7AEC1501558C5A" originalValue="über" pageId="null" pageNumber="36">ueber</normalizedToken>
dem Kelch deutlich gestielt
</emphasis>
(Stiel 2-4 mm lang). -
<normalizedToken id="E333B900235900757E2C52F139A68176" originalValue="Blüte" pageId="null" pageNumber="36">Bluete</normalizedToken>
: Sommer und Herbst.
</paragraph>
<paragraph id="0BB14B01EC00BBC36F09E96DECBEA46C" pageId="null" pageNumber="36">
<emphasis id="4D374A5D15946208A52496915BAD525C" italics="true" pageId="null" pageNumber="36">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="EC16E1DBAC8EA7D207EC3C0CAB31F581" italics="true" pageId="null" pageNumber="36">36:</emphasis>
Material aus dem
<normalizedToken id="30912BA5E3D80661894771D09E146CC2" originalValue="südlichen" pageId="null" pageNumber="36">suedlichen</normalizedToken>
Tessin (Favarger 1952).
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<paragraph id="1BB717D984DCF65439673D430D7F472F" pageId="null" pageNumber="36">
<emphasis id="47AD6D16CD0C37ECB5A8CF87D1909FBD" italics="true" pageId="null" pageNumber="36">Standort</emphasis>
(wenig bekannt). Montan und subalpin. Kalkhaltige
<normalizedToken id="9E57E4C6F6CE284721F4848DFDCCDDD0" originalValue="Böden" pageId="null" pageNumber="36">Boeden</normalizedToken>
. Trockene Wiesen.
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<paragraph id="448E36E9B115DF49837F13056E5DDF66" pageId="null" pageNumber="36">
<emphasis id="F34BC3933614D630A2E91B65A40E19F5" italics="true" pageId="null" pageNumber="36">Verbreitung. Insubrische Pflanze</emphasis>
(genaue Verbreitung nicht bekannt):
<normalizedToken id="FB686A8086D222F193050A8ED68F0F4D" originalValue="Südliches" pageId="null" pageNumber="36">Suedliches</normalizedToken>
Tessin (vgl. Kunz 1940; dort auch Beschreibung und Abgrenzung der Art) und angrenzende italienische Gebiete.
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,449 @@
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e71492" ID-GBIF-Dataset="2b24f5fb-ba29-46a0-8de0-2d0f2ac78020" ID-GBIF-Taxon="193856274" ID-Pensoft-Pub="1864-8312-80-75" ID-Pensoft-UUID="372E1AF5D7675176AD2805DFE2F0E6DA" ID-ZooBank="F85F11D5313D4A75B3907FDCCCA9376F" ModsDocID="1864-8312-80-75" checkinTime="1646282020555" checkinUser="pensoft" docAuthor="Barroso, Paulo Cezar Salgado, Menezes, Rodolpho Santos Telles, de Oliveira, Marcio Luiz &amp; Somavilla, Alexandre" docDate="2022" docId="0C07695ACEDD5A5F92DEBAAADAAFA62A" docLanguage="en" docName="ArthropodSystPhyl 80: 75-97" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-03-02" docSource="http://dx.doi.org/10.3897/asp.80.e71492" docTitle="Angiopolybia pallens" docType="treatment" docVersion="3" id="372E1AF5D7675176AD2805DFE2F0E6DA" lastPageNumber="75" masterDocId="372E1AF5D7675176AD2805DFE2F0E6DA" masterDocTitle="A systematic review of the Neotropical social wasp genus Angiopolybia Araujo, 1946 (Hymenoptera: Vespidae): species delimitation, morphological diagnosis, and geographical distribution" masterLastPageNumber="97" masterPageNumber="75" pageId="0" pageNumber="75" updateTime="1646286233131" updateUser="ExternalLinkService">
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<taxonomicName LSID="0C07695A-CEDD-5A5F-92DE-BAAADAAFA62A" authority="(Lepeletier, 1836)" baseAuthorityName="Lepeletier" baseAuthorityYear="1836" class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">Angiopolybia pallens (Lepeletier, 1836)</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lectotype, female of Polybia infernalis de Saussure, 1856 [currently Angiopolybia pallens (Lepeletier, 1836)]: a. lateral view, b. dorsal view, c. head in frontal view. Male: d. head in frontal view, e. lateral view, f. dorsal view. Scale: 1 mm. Source: Agniele Touret-Alby MNHN for images a, b, and c." figureDoi="10.3897/asp.80.e71492.figure1" httpUri="https://binary.pensoft.net/fig/652691" pageId="0" pageNumber="75">Figs 1a-f</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Structure variation of the nest of Angiopolybia pallens: a. typical form in external view, b. and c. variation found in external and internal view, respectively." figureDoi="10.3897/asp.80.e71492.figure2" httpUri="https://binary.pensoft.net/fig/652692" pageId="0" pageNumber="75">, 2a-c</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of three Angiopolybia species. Angiopolybia pallens: a. paramere, b. aedeagus in left lateral view, c. aedeagus in ventral view, d. digitus, e. cuspis (image mirrored). Angiopolybia obidensis: f. paramere, g. aedeagus in left lateral view, h. aedeagus in ventral view, i. digitus, j. cuspis. Angiopolybia paraensis: k. paramere, l. aedeagus in left lateral view, m. aedeagus in ventral view, n. digitus, o. cuspis. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e71492.figure6" httpUri="https://binary.pensoft.net/fig/652696" pageId="0" pageNumber="75">, 6a-e</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Morphological variation of Angiopolybia species. Angiopolybia pallens: a. Cayenne, French Guiana; b. Amazonas, Brazil; c. Bahia, Brazil; d. Sao Paulo, Brazil; e. Loreto, Peru; f. Napo, Ecuador; g. Chapare, Bolivia. Angiopolybia zischkai: h. Pipeline, Panama. Angiopolybia obidensis: i. Roraima, Brazil. Angiopolybia paraensis: j. Roraima (brown and yellow specimen), Brazil; k. Napo, Ecuador (brown specimen); l. Crique Alama, French Guiana (dark brown specimen). Scale: 1 mm." figureDoi="10.3897/asp.80.e71492.figure7" httpUri="https://binary.pensoft.net/fig/652697" pageId="0" pageNumber="75">, 7a-g</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="reference_group">
<paragraph pageId="0" pageNumber="75">
<taxonomicName authority="Guérin-Méneville, 1838" authorityName="Lepeletier" authorityYear="1836" class="Insecta" family="Vespidae" genus="Rhopalidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhopalidia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">Rhopalidia pallens</taxonomicName>
Lepeletier, 1836: 539; Spinola, 1851: 63 (nest); de Saussure, 1854: 189 (synonym of
<taxonomicName class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia pallipes" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallipes">Polybia pallipes</taxonomicName>
(Olivier, 1792)); du Buysson, 1906: 342 (synonym of
<taxonomicName class="Insecta" family="Vespidae" genus="Apoica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apoica pallida subsp. var. var. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="variety" species="pallida" subSpecies="var." variety="pallens">Apoica pallida var. pallens</taxonomicName>
(Fabricius, 1804)); Ducke, 1910: 542 (specimen of the collection of Spinola =
<taxonomicName family="Vespidae" genus="S." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. infernalis" order="Hymenoptera" pageId="0" pageNumber="75" rank="species" species="infernalis">S. infernalis</taxonomicName>
(de Saussure,1854)); Schulz, 1912: 60 (synonym:
<taxonomicName family="Vespidae" genus="S." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. infernalis" order="Hymenoptera" pageId="0" pageNumber="75" rank="species" species="infernalis">S. infernalis</taxonomicName>
(de Saussure, 1854)).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authorityName="Erichson" authorityYear="1848" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes rufina" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="rufina">Polistes rufina</taxonomicName>
Erichson, 1848: 590; Spinola, 1851: 79 (synonym of
<taxonomicName authority="Guérin-Méneville, 1838" authorityName="Lepeletier" authorityYear="1836" class="Insecta" family="Vespidae" genus="Rhopalidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhopalidia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">Rhopalidia pallens</taxonomicName>
Lepeletier, 1836).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">Polybia infernalis</taxonomicName>
de Saussure, 1854: 195, plate XXV: fig. 3 (in division
<emphasis italics="true" pageId="0" pageNumber="75">My</emphasis>
); Ducke, 1905a: 662 (synonym:
<taxonomicName family="Vespidae" genus="P." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. ampullaria" order="Hymenoptera" pageId="0" pageNumber="75" rank="species" species="ampullaria">P. ampullaria</taxonomicName>
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
, 1856); Richards, 1943: 45 (invalid designation of the type species of
<taxonomicName authority="Ducke, 1910" authorityName="Ducke" authorityYear="1910" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stelopolybia" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="genus">Stelopolybia</taxonomicName>
); Richards, 1978: 232 (lectotype designation). Type locality: &quot;Le Para&quot;, female (MNHN, Paris) [examined by images].
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authorityName="Moebius" authorityYear="1856" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia ampullaria" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="ampullaria">Polybia ampullaria</taxonomicName>
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
, 1856: 133, 155 (key identification of nests), 165 (plate VII), VII (figs. 1-8 - female, nest).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes flavopectus" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="flavopectus">Eumenes flavopectus</taxonomicName>
Provancher, 1888: 422.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authority="Ducke, 1910" authorityName="Ducke" authorityYear="1910" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stelopolybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">Stelopolybia infernalis</taxonomicName>
; Ducke, 1910: 519 (key), 524 (synonym:
<taxonomicName authorityName="Moebius" authorityYear="1856" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia ampullaria" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="ampullaria">Polybia ampullaria</taxonomicName>
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
, 1856), 525 (fig. 12 - nest); Ducke, 1913: 331 (synonym:
<taxonomicName family="Vespidae" genus="R." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="R. pallens" order="Hymenoptera" pageId="0" pageNumber="75" rank="species" species="pallens">R. pallens</taxonomicName>
Lepeletier, 1836).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authority="Ducke, 1910" authorityName="Ducke" authorityYear="1910" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stelopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">Stelopolybia pallens</taxonomicName>
; Bequaert, 1944: 293 (key), 294 (synonym:
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">Polybia infernalis</taxonomicName>
Saussure, 1854,
<taxonomicName authorityName="Moebius" authorityYear="1856" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia ampullaria" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="ampullaria">Polybia ampullaria</taxonomicName>
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
, 1856,
<taxonomicName authorityName="Erichson" authorityYear="1848" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes rufina" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="rufina">Polistes rufina</taxonomicName>
Erichson, 1884,
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes flavopectus" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="flavopectus">Eumenes flavopectus</taxonomicName>
<bibRefCitation author="Provancher, L 1889" journalOrPublisher="Darveau, Quebec" pageId="0" pageNumber="75" refId="B54" refString="Provancher, L 1889, 1888. Faune Entomologique du Canada: Traitant des Hymenopteres. Additiones et corrections au volume II. Typographie de C. Darveau, Quebec" title="Faune Entomologique du Canada: Traitant des Hymenopteres. Additiones et corrections au volume II. Typographie de C." year="1888">Provancher 1888</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Lepeletier" baseAuthorityYear="1836" class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">Angiopolybia pallens</taxonomicName>
; Araujo, 1946: 169; Richards, 1978: 231 (key), 232 (description of male, and diagnose of female and male);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1427.1.4" author="Andena, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="57 - 64" refId="B1" refString="Andena, SR, Noll, FB, Carpenter, JM, 2007. Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini). Zootaxa 1427: 57 - 64, DOI: https://doi.org/10.11646/zootaxa.1427.1.4" title="Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini)." url="https://doi.org/10.11646/zootaxa.1427.1.4" volume="1427" year="2007">Andena et al., 2007</bibRefCitation>
: 59 (phylogenetic discussion), 60 (table 2 - characters matrix), 61 (figs. 1A, 2A), 62 (figs. 3B, 5 - cladogram), 63 (key), 64 (locality of examined material);
<bibRefCitation DOI="https://doi.org/10.4238/2014.January.8.7" author="Carvalho, AF" journalOrPublisher="Genetics and Molecular Research" pageId="0" pageNumber="75" pagination="89 - 94" refId="B14" refString="Carvalho, AF, Santos, GMM, Menezes, RST, Costa, MA, 2014. Genetic diversity of Angiopolybia pallens (Lepeletier) (Hymenoptera, Vespidae, Polistinae) explained by the disjunction of South American rainforests. Genetics and Molecular Research 13 (1): 89 - 94, DOI: https://doi.org/10.4238/2014.January.8.7" title="Genetic diversity of Angiopolybia pallens (Lepeletier) (Hymenoptera, Vespidae, Polistinae) explained by the disjunction of South American rainforests." url="https://doi.org/10.4238/2014.January.8.7" volume="13" year="2014">Carvalho et al., 2014</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1093/biolinnean/blaa215" author="Carvalho, AF" journalOrPublisher="Biological Journal of the Linnean Society" pageId="0" pageNumber="75" pagination="955 - 969" refId="B12" refString="Carvalho, AF, Menezes, RST, Miranda, EA, Costa, MA, Del Lama, MA, 2021. Comparative phylogeography and palaeomodelling reveal idiosyncratic responses to climate changes in Neotropical paper wasps. Biological Journal of the Linnean Society 132 (4): 955 - 969, DOI: https://doi.org/10.1093/biolinnean/blaa215" title="Comparative phylogeography and palaeomodelling reveal idiosyncratic responses to climate changes in Neotropical paper wasps." url="https://doi.org/10.1093/biolinnean/blaa215" volume="132" year="2021">2021</bibRefCitation>
(evolutionary hypothesis of species distribution).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName authority="Ducke, 1910" authorityName="Ducke" authorityYear="1910" class="Insecta" family="Vespidae" genus="Stelopolybia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stelopolybia (Angiopolybia) pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens" subGenus="Angiopolybia">Stelopolybia (Angiopolybia) pallens</taxonomicName>
; Richards and Richards, 1951: 77 (list and notes about the nests).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">Angiopolybia infernalis</taxonomicName>
; Overal, 1978: 9 (list of species).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="type locality">
<paragraph pageId="0" pageNumber="75">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="75">Cayenne, French Guiana.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="diagnosis">
<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="75">Anterior wing of 7-8.5 mm; eyes with medium-sized and sparse bristles; rounded gena; pronotal lamella low on the anterior margin, one fifth of the height of antennal socket; pronotal lobe developed in the anterior lateral region, below the pronotal fovea; defined and deep pronotal fovea; axillary fossa with anterior margin directed to the posterior region; posterior submedian translucent mark of the propodeum inserted in a round depression; basal metapleural area with parallel upper and lower margins.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="redescription">
<paragraph pageId="0" pageNumber="75">Redescription of female (Fig. 1a, b, c).</paragraph>
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">Size.</emphasis>
(1) Head 1.27 mm long, 2 mm high, and 2.27 mm wide; (2) mesosoma 3.50 mm long, anterior wing 8.11 mm long, and posterior wing 5.27 mm long; (3) metasoma 5.53 mm long.
<emphasis bold="true" pageId="0" pageNumber="75">Head.</emphasis>
(1) Lateral ocelli with 0.15 mm and median ocellus with 0.17 mm of diameter, not inserted in a declivity of the vertex, and the lateral ocelli separated from the eyes for twice its diameter. (2) Compound eyes with medium-sized and sparse bristles. (3) Frons with interantennal space about twice the height of antennal socket. Anterior tentorial fovea closer to the antennal socket than to the eye. The central region of the frons with long bristles. (4) Antennal socket 0.22 mm high. (5) Clypeus as high as wide, contact with eyes for a distance greater than twice the height of antennal socket, and lateral lobe touching the eye. Medium bristles in the basal half and long bristles in the apical half. (6) Gena with half of the width of the eye at the level of the ocular sinus.
<emphasis bold="true" pageId="0" pageNumber="75">Mesosoma.</emphasis>
(1) Anterior lamella of pronotum with a height of one fifth of the height of antennal socket. Pronotal fovea with a circular shape, deep and with slight anterior prominence. (2) Mesoscutum convex and as long as wide. (3) Tegula 1.7 times longer than wide. (4) Axillary fossa with anterior margin directing to the posterior region. (5) Propodeum with translucent posterior submedian mark, anterior to the propodeal valve, inserted in a round depression. Propodeal valve complete and expanded, median region with half of the height of antennal socket. (6) Anterior wing with prestigma 1.4 times longer than wide. (7) Posterior wing with eight hamuli.
<emphasis bold="true" pageId="0" pageNumber="75">Metasoma.</emphasis>
(1) Metasomal tergum I two times longer than broad. Tergum with angulation in the posterior third, in lateral view. (2) Metasomal tergum II 0.82 times longer than broad.
<emphasis bold="true" pageId="0" pageNumber="75">Color.</emphasis>
Brown in general. Yellow: lateral of the apex, lateral of the front, interantennal region, disc of the clypeus, mandibles, gena, antennal segments: scape, dorsal of FL6 (flagellomere 6) and 7, FL8-10, band contouring the posterior margin of the pronotum, tegula, thin bands in the posterior margins of the metasomal terga I, II and III, and metasomal sterna II and III. Yellowish brown: central longitudinal and lateral region of the mesoscutum, tibiae and tarsi, femora median and posterior, metasomal tergum I, anterior half of metasomal tergum II, metasomal sterna I-IV. Black: FL1-5, ventral of FL 6 and 7, metasomal terga 4-6 and metasomal sterna 5 and 6. Wings with hyaline cells, pterostigma and venation in general yellowish-brown, except brown in the veins C, Sc+R, M+Cu and M.
</paragraph>
<caption doi="10.3897/asp.80.e71492.figure1" httpUri="https://binary.pensoft.net/fig/652691" pageId="0" pageNumber="75" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">Figure 1.</emphasis>
Lectotype, female of
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia infernalis</emphasis>
</taxonomicName>
de Saussure, 1856 [currently
<taxonomicName baseAuthorityName="Lepeletier" baseAuthorityYear="1836" class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">Angiopolybia pallens</emphasis>
</taxonomicName>
(Lepeletier, 1836)]:
<emphasis bold="true" pageId="0" pageNumber="75">a.</emphasis>
lateral view,
<emphasis bold="true" pageId="0" pageNumber="75">b.</emphasis>
dorsal view,
<emphasis bold="true" pageId="0" pageNumber="75">c.</emphasis>
head in frontal view. Male:
<emphasis bold="true" pageId="0" pageNumber="75">d.</emphasis>
head in frontal view,
<emphasis bold="true" pageId="0" pageNumber="75">e.</emphasis>
lateral view,
<emphasis bold="true" pageId="0" pageNumber="75">f.</emphasis>
dorsal view. Scale: 1 mm. Source:
<normalizedToken originalValue="Agnièle">Agniele</normalizedToken>
Touret-Alby MNHN for images
<emphasis bold="true" pageId="0" pageNumber="75">a</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="75">b</emphasis>
, and
<emphasis bold="true" pageId="0" pageNumber="75">c</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="redescription">
<paragraph pageId="0" pageNumber="75">Redescription of male (Fig. 1d, e, f).</paragraph>
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">Size.</emphasis>
(1) Head 1.01 mm long, 1.83 mm high, and 2.09 mm wide; (2) mesosoma 2.87 mm long, anterior wing 7.1 mm long, and posterior wing 4.5 mm long; (3) metasoma 6.08 mm long.
<emphasis bold="true" pageId="0" pageNumber="75">Head.</emphasis>
(1) Lateral ocelli with 0.17 mm and median ocellus with 0.18 mm of diameter. (2) Compound eyes with small-sized and sparse bristles. (3) Frons with interantennal space with 1.85 times the height of antennal socket. Anterior tentorial fovea closer to the eye that to the antennal socket. (4) Antennal socket with 0.21 mm high. (5) Clypeus as high as wide and apex less acute than in the female. Pubescence stronger than in the female. (6) Gena with half of the width of the eye at the level of the ocular sinus.
<emphasis bold="true" pageId="0" pageNumber="75">Mesosoma.</emphasis>
(1) Mesoscutum 0.9 times longer than wide. (2) Propodeum with translucent posterior submedian mark, anterior to the propodeal valve, inserted in a not round depression. (3) Anterior wing with prestigma 1.6 times longer than wide.
<emphasis bold="true" pageId="0" pageNumber="75">Metasoma.</emphasis>
(1) Metasomal tergum I 1.9 times longer than broad. (2) Metasomal tergum II 0.8 times longer than wide.
<emphasis bold="true" pageId="0" pageNumber="75">
Genitalia (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of three Angiopolybia species. Angiopolybia pallens: a. paramere, b. aedeagus in left lateral view, c. aedeagus in ventral view, d. digitus, e. cuspis (image mirrored). Angiopolybia obidensis: f. paramere, g. aedeagus in left lateral view, h. aedeagus in ventral view, i. digitus, j. cuspis. Angiopolybia paraensis: k. paramere, l. aedeagus in left lateral view, m. aedeagus in ventral view, n. digitus, o. cuspis. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e71492.figure6" httpUri="https://binary.pensoft.net/fig/652696" pageId="0" pageNumber="75">6a</figureCitation>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="75">e). Paramere</emphasis>
1.6 mm long and 0.6 mm wide; parameral spine with one fifth of the paramere, curved upwards and with small-sized and sparse bristles; lobe with a rounded apex and slightly curved downwards.
<emphasis bold="true" pageId="0" pageNumber="75">Aedeagus</emphasis>
1.2 mm long; enlarged valve with a small emargination in the tip; apical portion 0.42 mm long and straight, ventral margin with denticles directed for the anterior region; denticulation decreasing in size from the apex to the base and more sclerotized than the rest of the apical portion; small-sized bristles with alveolar base, closer in the lower half and sparse in the upper half; median expansion without denticles and with acute apex; lateral apodeme not flattened dorsoventrally at the apex; basal apodeme arched to the venter.
<emphasis bold="true" pageId="0" pageNumber="75">Digitus</emphasis>
2.5 times longer than wide; apical process not curved in the region of the upper half and with bristles of alveolar base small and sparse; rounded anteroventral lobe with a strip of black scale-like bristles crossing it obliquely at the base of the digitus; bristles absent in the basal articulation.
<emphasis bold="true" pageId="0" pageNumber="75">Cuspis</emphasis>
approximately 0.48 mm long, with five black scale-like bristles on the lateral lobe, and small bristles with alveolar base and close throughout the area of the cuspis, except sparse in the central region and on the ventral margin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="morphological variation (fig. 7ag)">
<paragraph pageId="0" pageNumber="75">Morphological variation (Fig. 7a-g).</paragraph>
<paragraph pageId="0" pageNumber="75">
Anterior wing between 7-8.5 mm in length. Anterior margin of the pronotum (below the fovea) more curved. Pronotum, in lateral view, with frontal region more projected forward. Coloration varied between populations from black with yellow marks [like
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
dark morph (sensu Richards, 1978)] to yellowish brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="nest (fig. 2ac)">
<paragraph pageId="0" pageNumber="75">Nest (Fig. 2a-c).</paragraph>
<paragraph pageId="0" pageNumber="75">
The nest of
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
was initially described by
<bibRefCitation author="Moebius, K" journalOrPublisher="Abhandlungen aus dem Gebiete der Naturwissenschaften" pageId="0" pageNumber="75" pagination="117 - 171" refId="B45" refString="Moebius, K, 1856. Die Nester der geselligen Wespen. Abhandlungen aus dem Gebiete der Naturwissenschaften 3: 117 - 171" title="Die Nester der geselligen Wespen." volume="3" year="1856">
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
(1856)
</bibRefCitation>
for
<taxonomicName authorityName="Moebius" authorityYear="1856" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia ampullaria" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="ampullaria">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia ampullaria</emphasis>
</taxonomicName>
(junior synonym of the taxon), and later described by
<bibRefCitation author="Wenzel, JW" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="75" pagination="1 - 39" refId="B72" refString="Wenzel, JW, 1998. A generic key to the nests of Hornets, Yellowjackets, and paper wasps worldwide (Vespidae: Vespinae, Polistinae). American Museum Novitates 3224: 1 - 39" title="A generic key to the nests of Hornets, Yellowjackets, and paper wasps worldwide (Vespidae: Vespinae, Polistinae)." volume="3224" year="1998">Wenzel (1998)</bibRefCitation>
as a nest with a flask-shaped envelope and with a long downward entrance with its hole horizontally; pedicel initially single and later being able to be multiple; flexible card with long fibers, usually yellow or amber; and adjacent combs, suspended or fused from each other and without contact with the envelope. Additionally, the following nest variation was found: built on an irregular substrate (thin branch with several leaves), connected to the substrate by a central pedicel (thick) and several support pedicels (fines); 11 combs overlapping and also connected by a central pedicel and multiple support, with the third and fourth combs with the largest circumferences and combs decreasing in circumference towards the ends, and without contact with the envelope; hexagonal cells of the combs with diameter of 2.5 mm; single envelope with long fibers arranged longitudinally, circumference gradually decreasing towards the entrance and without entrance of tubular shape.
</paragraph>
<caption doi="10.3897/asp.80.e71492.figure2" httpUri="https://binary.pensoft.net/fig/652692" pageId="0" pageNumber="75" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">Figure 2.</emphasis>
Structure variation of the nest of
<taxonomicName baseAuthorityName="Lepeletier" baseAuthorityYear="1836" class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">Angiopolybia pallens</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="75">a.</emphasis>
typical form in external view,
<emphasis bold="true" pageId="0" pageNumber="75">b.</emphasis>
and
<emphasis bold="true" pageId="0" pageNumber="75">c.</emphasis>
variation found in external and internal view, respectively.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="comparative comments">
<paragraph pageId="0" pageNumber="75">Comparative comments.</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Lepeletier" baseAuthorityYear="1836" class="Insecta" family="Vespidae" genus="Angiopolybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angiopolybia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">Angiopolybia pallens</emphasis>
</taxonomicName>
is distinguished by the presence of a lobe in the lateroanterior region of the pronotum, absent in
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. zischkai" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="75">A. zischkai</emphasis>
</taxonomicName>
, and metapleural basalar area with parallel upper and lower margins, diverging in
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. zischkai" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="75">A. zischkai</emphasis>
</taxonomicName>
. Some
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
specimens can be confused with
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. zischkai" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="75">A. zischkai</emphasis>
</taxonomicName>
because it can resemble the typical form of
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. zischkai" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis italics="true" pageId="0" pageNumber="75">A. zischkai</emphasis>
</taxonomicName>
, of color darker.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="additional comments">
<paragraph pageId="0" pageNumber="75">Additional comments.</paragraph>
<paragraph pageId="0" pageNumber="75">
Despite the geographic disjunction by thousand kilometers in the distribution of
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
between the Amazon and Atlantic Forest (
<bibRefCitation DOI="https://doi.org/10.4238/2014.January.8.7" author="Carvalho, AF" journalOrPublisher="Genetics and Molecular Research" pageId="0" pageNumber="75" pagination="89 - 94" refId="B14" refString="Carvalho, AF, Santos, GMM, Menezes, RST, Costa, MA, 2014. Genetic diversity of Angiopolybia pallens (Lepeletier) (Hymenoptera, Vespidae, Polistinae) explained by the disjunction of South American rainforests. Genetics and Molecular Research 13 (1): 89 - 94, DOI: https://doi.org/10.4238/2014.January.8.7" title="Genetic diversity of Angiopolybia pallens (Lepeletier) (Hymenoptera, Vespidae, Polistinae) explained by the disjunction of South American rainforests." url="https://doi.org/10.4238/2014.January.8.7" volume="13" year="2014">Carvalho et al. 2014</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1093/biolinnean/blaa215" author="Carvalho, AF" journalOrPublisher="Biological Journal of the Linnean Society" pageId="0" pageNumber="75" pagination="955 - 969" refId="B12" refString="Carvalho, AF, Menezes, RST, Miranda, EA, Costa, MA, Del Lama, MA, 2021. Comparative phylogeography and palaeomodelling reveal idiosyncratic responses to climate changes in Neotropical paper wasps. Biological Journal of the Linnean Society 132 (4): 955 - 969, DOI: https://doi.org/10.1093/biolinnean/blaa215" title="Comparative phylogeography and palaeomodelling reveal idiosyncratic responses to climate changes in Neotropical paper wasps." url="https://doi.org/10.1093/biolinnean/blaa215" volume="132" year="2021">2021</bibRefCitation>
), considering the morphological analysis of specimens collected in both biomes, we verified that the differences, when found, are very subtle and follow the variation found between different populations. Considering the populations from the Atlantic Forest, the color varied from yellow with brown marks to completely brown, while among the populations from the Amazon Forest it varied from black with yellow marks to yellowish brown, with some forms existing in both biomes. The occipital carina present in
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
, despite weak or not, found in this study not reported by
<bibRefCitation author="Richards, OW" journalOrPublisher="The Bulletin of Zoological Nomenclature" pageId="0" pageNumber="75" refId="B60" refString="Richards, OW, 1978. The social wasps of the Americas: excluding the Vespinae. British Museum (Natural History) London, 580 pp." title="The social wasps of the Americas: excluding the Vespinae. British Museum (Natural History) London, 580 pp." year="1978">Richards (1978)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1427.1.4" author="Andena, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="57 - 64" refId="B1" refString="Andena, SR, Noll, FB, Carpenter, JM, 2007. Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini). Zootaxa 1427: 57 - 64, DOI: https://doi.org/10.11646/zootaxa.1427.1.4" title="Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini)." url="https://doi.org/10.11646/zootaxa.1427.1.4" volume="1427" year="2007">Andena et al. (2007)</bibRefCitation>
. They treated the occipital carina absent for the taxon. The prestigma longer than wide, also, was not cited by
<bibRefCitation author="Richards, OW" journalOrPublisher="The Bulletin of Zoological Nomenclature" pageId="0" pageNumber="75" refId="B60" refString="Richards, OW, 1978. The social wasps of the Americas: excluding the Vespinae. British Museum (Natural History) London, 580 pp." title="The social wasps of the Americas: excluding the Vespinae. British Museum (Natural History) London, 580 pp." year="1978">Richards (1978)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1427.1.4" author="Andena, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="57 - 64" refId="B1" refString="Andena, SR, Noll, FB, Carpenter, JM, 2007. Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini). Zootaxa 1427: 57 - 64, DOI: https://doi.org/10.11646/zootaxa.1427.1.4" title="Phylogenetic analysis of the Neotropical social wasps of the genus Angiopolybia Araujo, 1946 (Hymenoptera, Vespidae, Epiponini)." url="https://doi.org/10.11646/zootaxa.1427.1.4" volume="1427" year="2007">Andena et al. (2007)</bibRefCitation>
. However, this character was found by
<bibRefCitation author="Silveira, OT" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="75" pagination="69 - 72" refId="B65" refString="Silveira, OT, Carpenter, JM, 1995. A new species of Agelaia Lepeletier from Brazilian Amazonia (Hymenoptera: Vespidae; Polistinae). Journal of the New York Entomological Society 103 (1): 69 - 72" title="A new species of Agelaia Lepeletier from Brazilian Amazonia (Hymenoptera: Vespidae; Polistinae)." volume="103" year="1995">Silveira and Carpenter (1995)</bibRefCitation>
for some specimens, and here we also found the two forms, prestigma about as long as wide and longer than wide. The redescription of
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
was made based on the lectotype of
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia infernalis</emphasis>
</taxonomicName>
de Saussure, 1854 (junior synonym of
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
). The information about the male specimen described is: BRA, Roraima,
<normalizedToken originalValue="Amajarí">Amajari</normalizedToken>
, Serra do
<normalizedToken originalValue="Tepequém">Tepequem</normalizedToken>
, SESC
<normalizedToken originalValue="Tepequém">Tepequem</normalizedToken>
. 1-16.i.2016 /
<taxonomicName genus="Malaise" lsidName="Malaise" pageId="0" pageNumber="75" rank="genus">Malaise</taxonomicName>
grande, J.A. Rafael, F.F. Xavier Filho col.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="lectotype">
<paragraph pageId="0" pageNumber="75">Lectotype.</paragraph>
<paragraph pageId="0" pageNumber="75">
♀, TYPE / MUSEUM PARIS,
<normalizedToken originalValue="Amérique">Amerique</normalizedToken>
, Leprieur 1834 /
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia infernalis</emphasis>
</taxonomicName>
Sauss., Type. / 289694 /
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia infernalis</emphasis>
</taxonomicName>
Sauss., Lectotype ♀, Richards, 1971 (MNHN, Paris), record MNHN, Paris EY25588 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lectotype, female of Polybia infernalis de Saussure, 1856 [currently Angiopolybia pallens (Lepeletier, 1836)]: a. lateral view, b. dorsal view, c. head in frontal view. Male: d. head in frontal view, e. lateral view, f. dorsal view. Scale: 1 mm. Source: Agniele Touret-Alby MNHN for images a, b, and c." figureDoi="10.3897/asp.80.e71492.figure1" httpUri="https://binary.pensoft.net/fig/652691" pageId="0" pageNumber="75">1a</figureCitation>
,
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lectotype, female of Polybia infernalis de Saussure, 1856 [currently Angiopolybia pallens (Lepeletier, 1836)]: a. lateral view, b. dorsal view, c. head in frontal view. Male: d. head in frontal view, e. lateral view, f. dorsal view. Scale: 1 mm. Source: Agniele Touret-Alby MNHN for images a, b, and c." figureDoi="10.3897/asp.80.e71492.figure1" httpUri="https://binary.pensoft.net/fig/652691" pageId="0" pageNumber="75">1b</figureCitation>
,
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lectotype, female of Polybia infernalis de Saussure, 1856 [currently Angiopolybia pallens (Lepeletier, 1836)]: a. lateral view, b. dorsal view, c. head in frontal view. Male: d. head in frontal view, e. lateral view, f. dorsal view. Scale: 1 mm. Source: Agniele Touret-Alby MNHN for images a, b, and c." figureDoi="10.3897/asp.80.e71492.figure1" httpUri="https://binary.pensoft.net/fig/652691" pageId="0" pageNumber="75">1c</figureCitation>
).
<taxonomicName authorityName="de Saussure" authorityYear="1854" class="Insecta" family="Vespidae" genus="Polybia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polybia infernalis" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis italics="true" pageId="0" pageNumber="75">Polybia infernalis</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName authority="Guérin-Méneville, 1838" authorityName="Lepeletier" authorityYear="1836" class="Insecta" family="Vespidae" genus="Rhopalidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhopalidia pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">Rhopalidia pallens</emphasis>
</taxonomicName>
Lepeletier, 1836, but here we used it to the redescription of the species. Type specimen analyzed by images.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="materials_examined">
<paragraph pageId="0" pageNumber="75">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="75">
We examined
<specimenCount type="female">469 females</specimenCount>
and
<specimenCount type="male">32 males</specimenCount>
for
<taxonomicName class="Insecta" genus="A." kingdom="Animalia" lsidName="A. pallens" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="pallens">
<emphasis italics="true" pageId="0" pageNumber="75">A. pallens</emphasis>
</taxonomicName>
; see supplementary material S1.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="distribution">
<paragraph pageId="0" pageNumber="75">Geographic distribution.</paragraph>
<paragraph pageId="0" pageNumber="75">
Bolivia: Beni, Cochabamba; Brazil: Acre, Alagoas (
<emphasis bold="true" pageId="0" pageNumber="75">new record</emphasis>
),
<normalizedToken originalValue="Amapá">Amapa</normalizedToken>
, Amazonas, Bahia,
<normalizedToken originalValue="Ceará">Ceara</normalizedToken>
,
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo,
<normalizedToken originalValue="Maranhão">Maranhao</normalizedToken>
, Mato Grosso,
<normalizedToken originalValue="Pará">Para</normalizedToken>
, Pernambuco,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Rio de Janeiro,
<normalizedToken originalValue="Rondônia">Rondonia</normalizedToken>
, Roraima, Santa Catarina,
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo Sergipe; Colombia: Meta,
<normalizedToken originalValue="Caquetá">Caqueta</normalizedToken>
,
<normalizedToken originalValue="Nariño">Narino</normalizedToken>
,
<normalizedToken originalValue="Vaupés">Vaupes</normalizedToken>
, Putumayo, Amazonas; Ecuador: Esmereldas, Napo, Pichincha; French Guiana; Guyana; Panama; Peru: Loreto, San Martin,
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
, Pasco,
<normalizedToken originalValue="Junín">Junin</normalizedToken>
, Cuzco; Suriname; Trinidad and Tobago; Venezuela: Monagas (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Map of South and Central America showing the geographic distribution of Angiopolybia species: a. Angiopolybia pallens, b. Angiopolybia zischkai, c. Angiopolybia obidensis, and d. Angiopolybia paraensis." figureDoi="10.3897/asp.80.e71492.figure10" httpUri="https://binary.pensoft.net/fig/652700" pageId="0" pageNumber="75">10a</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="6ACF59C2B62C6F83B1842296DF1D38FA" type="GBIF-Dataset">60ca36d6-4c1f-4050-88e2-38e8b28f3f5c</mods:identifier>
<mods:identifier id="F17CBA4CB80A16E1D313471AA5A98194" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="DDCA51797F41ABF17EB0B971FF8BFF74" type="Zenodo-Dep">3743146</mods:identifier>
<mods:identifier id="57F475AC668F575F850CD328000D376C" type="ZooBank">urn:lsid:zoobank.org:pub:21B9187A-D055-4946-8922-53FD10A159CE</mods:identifier>
</mods:mods>
<treatment id="0C0787D1FF90837DFC06F96CFB35EA3E" ID-DOI="http://doi.org/10.5281/zenodo.3809870" ID-GBIF-Taxon="163536913" ID-Zenodo-Dep="3809870" LSID="urn:lsid:zoobank.org:act:E67019D9-A3A3-4A9C-9CC4-CE6279752520" httpUri="http://treatment.plazi.org/id/0C0787D1FF90837DFC06F96CFB35EA3E" lastPageId="7" lastPageNumber="144" pageId="4" pageNumber="141">
<subSubSection id="CCB4654CFF90837EFC06F96CFA4AE8C0" box="[944,1324,1715,1743]" pageId="4" pageNumber="141" type="nomenclature">
<paragraph id="841136C7FF90837EFC06F96CFA4AE8C0" blockId="4.[944,1324,1715,1775]" box="[944,1324,1715,1743]" pageId="4" pageNumber="141">
<heading id="DF5981ABFF90837EFC06F96CFA4AE8C0" box="[944,1324,1715,1743]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="141" reason="2">
<taxonomicName id="43AE4D44FF90837EFC06F96CFBF4E8C1" ID-CoL="8H2QC" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[944,1170,1715,1742]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF90837EFC06F96CFBF4E8C1" bold="true" box="[944,1170,1715,1742]" italics="true" pageId="4" pageNumber="141">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB2FF96AFA4AE8C0" box="[1177,1324,1717,1743]" pageId="4" pageNumber="141" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFB96F90BFAF3E923" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFB96F90BFBDDE8E1" blockId="4.[944,1324,1715,1775]" box="[1056,1211,1748,1775]" pageId="4" pageNumber="141">
(
<figureCitation id="1C952A42FF90837EFB91F90BFB0FE8E1" box="[1063,1129,1748,1774]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Figs 1</figureCitation>
;
<figureCitation id="1C952A42FF90837EFBC1F90BFBE3E8E1" box="[1143,1157,1748,1774]" captionStart="FIG" captionStartId="6.[132,143,1987,2005]" captionTargetBox="[163,1420,231,1895]" captionTargetId="figure@6.[132,1455,216,1902]" captionTargetPageId="6" captionText="FIG. 2. — Polletomyia subulata n. gen., n. sp.: A, male terminalia, lateral view; B, male epandrium, proctiger and epandrial claspers, dorsal view; C, female terminalia, external structure, ventral view; D, female terminalia, internal structure, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743150" httpUri="https://zenodo.org/record/3743150/files/figure.png" pageId="4" pageNumber="141">2</figureCitation>
;
<figureCitation id="1C952A42FF90837EFB25F90BFBD5E8E0" box="[1171,1203,1748,1775]" captionStart="FIG" captionStartId="10.[207,218,1142,1160]" captionTargetBox="[132,1456,215,1098]" captionTargetId="figure@10.[127,1461,209,1106]" captionTargetPageId="10" captionText="FIG. 5. — Photomicrographs of male wings: A, Polletomyia subulata n. gen., n. sp.; B, Myiomystax trilineata n. gen., n. sp. Scale bars: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3743156" httpUri="https://zenodo.org/record/3743156/files/figure.png" pageId="4" pageNumber="141">5A</figureCitation>
)
</paragraph>
<paragraph id="841136C7FF90837EFCF1F8C5FAF3E923" blockId="4.[839,1429,1818,1836]" box="[839,1429,1818,1836]" pageId="4" pageNumber="141">
<uri id="F03F3AC5FF90837EFCF1F8C5FAF3E923" box="[839,1429,1818,1836]" pageId="4" pageNumber="141">
urn:lsid:zoobank.org:act:
<uuid id="F0080C12FF90837EFBBAF8C5FAF3E923" box="[1036,1429,1818,1836]" pageId="4" pageNumber="141">E67019D9-A3A3-4A9C-9CC4-CE6279752520</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFC9BF88FFBC2E966" box="[813,1188,1872,1897]" pageId="4" pageNumber="141" type="distribution">
<paragraph id="841136C7FF90837EFC9BF88FFBC2E966" blockId="4.[813,1188,1872,1897]" box="[813,1188,1872,1897]" pageId="4" pageNumber="141">
<typeStatus id="5B158865FF90837EFC9BF88FFC3BE96A" box="[813,861,1872,1896]" pageId="4" pageNumber="141">TYPE</typeStatus>
LOCALITY. —
<collectingCountry id="FCB97657FF90837EFC5DF88EFB5CE966" box="[1003,1082,1873,1897]" name="French Guiana" pageId="4" pageNumber="141">Guyane</collectingCountry>
, Mitaraka.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837DFC9BF85BFE3BED0D" lastPageId="7" lastPageNumber="144" pageId="4" pageNumber="141" type="materials_examined">
<paragraph id="841136C7FF90837DFC9BF85BFE3BED0D" blockId="4.[813,1456,1924,2027]" lastBlockId="7.[132,776,216,770]" lastPageId="7" lastPageNumber="144" pageId="4" pageNumber="141">
TYPE MATERIAL. —
<materialsCitation id="34C63C9AFF90837EFC59F85BFAD6E9E4" ID-GBIF-Occurrence="2595987901" collectingDate="2015-02-27" collectingDateMax="2015-03-06" collectingDateMin="2015-02-27" collectionCode="YPT" collectorName="Mitaraka &amp; Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" location="French Guiana" longLatPrecision="1" longitude="-54.448055" pageId="4" pageNumber="141" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B6DAEAD5FF90837EFC59F85BFB28E993" bold="true" box="[1007,1102,1924,1948]" pageId="4" pageNumber="141">
<typeStatus id="5B158865FF90837EFC59F85BFB28E993" box="[1007,1102,1924,1948]" pageId="4" pageNumber="141" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B6DAEAD5FF90837EFBEDF85BFB89E993" bold="true" box="[1115,1263,1924,1948]" pageId="4" pageNumber="141">
<collectingCountry id="FCB97657FF90837EFBEDF85BFB89E993" box="[1115,1263,1924,1948]" name="French Guiana" pageId="4" pageNumber="141">French Guiana</collectingCountry>
</emphasis>
<specimenCount id="92A8FD4EFF90837EFAB1F858FA7DE991" box="[1287,1307,1927,1950]" pageId="4" pageNumber="141" type="male"></specimenCount>
;
<collectorName id="295B5311FF90837EFA9FF85BFAE2E993" box="[1321,1412,1924,1948]" pageId="4" pageNumber="141">Mitaraka</collectorName>
, site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF90837EFC74F840FB29E9B8" box="[962,1103,1951,1975]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="4" pageNumber="141" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF90837EFBEEF841FB8EE9B9" box="[1112,1256,1950,1974]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="4" pageNumber="141" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<elevation id="0F83D1F4FF90837EFB44F840FA31E9B9" box="[1266,1367,1950,1975]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="141" unit="m" value="29.0">
<quantity id="43569B22FF90837EFB44F840FA43E9B9" box="[1266,1317,1951,1975]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="141" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF90837EFC23F866FB50E9DE" box="[917,1078,1977,2001]" pageId="4" pageNumber="141" value="2015-02-27" valueMax="2015-03-06" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF90837EFC23F866FB50E9DE" box="[917,1078,1977,2001]" pageId="4" pageNumber="141" value="2015-02-27" valueMax="2015-03-06" valueMin="2015-02-27">27.II-6.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF90837EFBF6F866FBD6E9DE" box="[1088,1200,1977,2001]" pageId="4" pageNumber="141">Marc Pollet</collectorName>
leg. (sample code: MITA- RAKA/125),
<collectionCode id="E2BFAE02FF90837EFC18F80BFCBAE9E4" box="[942,988,2004,2027]" pageId="4" pageNumber="141">YPT</collectionCode>
; specimen dissected, mounted on slide; MNHN.
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFF32FF07FE4FEF2A" ID-GBIF-Occurrence="2595987914" collectingDate="2015-02-27" collectingDateMax="2015-03-06" collectingDateMin="2015-02-27" collectionCode="YPT, MNHN, LACM" collectorName="Mitaraka &amp; Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" location="French Guiana" longLatPrecision="1" longitude="-54.448055" pageId="7" pageNumber="144" specimenCode="LACM 340591-340594, LACM 340595" specimenCount="20" specimenCount-female="4" specimenCount-male="16" typeStatus="paratype">
<emphasis id="B6DAEAD5FF93837DFF32FF07FEE4EEFF" bold="true" box="[132,386,216,240]" pageId="7" pageNumber="144">
<typeStatus id="5B158865FF93837DFF32FF07FF80EEFF" box="[132,230,216,240]" pageId="7" pageNumber="144" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="FCB97657FF93837DFF46FF07FEE4EEFF" box="[240,386,216,240]" name="French Guiana" pageId="7" pageNumber="144">French Guiana</collectingCountry>
</emphasis>
<specimenCount id="92A8FD4EFF93837DFE21FF06FEDBEEFD" box="[407,445,217,242]" pageId="7" pageNumber="144" type="male">8 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF93837DFE7FFF06FE8AEEFD" box="[457,492,217,242]" pageId="7" pageNumber="144" type="female">2 ♀</specimenCount>
; same data as
<typeStatus id="5B158865FF93837DFDC2FF06FDAAEEFE" box="[628,716,217,241]" pageId="7" pageNumber="144" type="holotype">holotype</typeStatus>
;
<collectionCode id="E2BFAE02FF93837DFD63FF06FC65EEFF" box="[725,771,217,240]" pageId="7" pageNumber="144">YPT</collectionCode>
;
<collectionCode id="E2BFAE02FF93837DFF32FF2BFFB1EF04" box="[132,215,244,267]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="7" pageNumber="144">MNHN</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFF51FF2CFE76EF02" box="[231,272,243,269]" pageId="7" pageNumber="144" type="male">4 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF93837DFEA9FF2CFE23EF02" box="[287,325,243,269]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
),
<collectionCode id="E2BFAE02FF93837DFEEDFF2CFEC4EF04" box="[347,418,243,267]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFE04FF2CFEBDEF02" box="[434,475,243,269]" pageId="7" pageNumber="144" type="male">4 ♂</specimenCount>
<specimenCode id="D4089EBCFF93837DFE55FF2CFDB3EF04" box="[483,725,243,267]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340591-340594</specimenCode>
,
<specimenCount id="92A8FD4EFF93837DFD57FF2CFC60EF02" box="[737,774,243,269]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
<specimenCode id="D4089EBCFF93837DFF32FED1FE44EF2A" box="[132,290,269,294]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340595</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFEF5FED1FD04EF7A" ID-GBIF-Occurrence="2595987908" collectingDate="2015-02-27" collectingDateMax="2015-03-05" collectingDateMin="2015-02-27" collectionCode="BPT, LACM" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" longLatPrecision="1" longitude="-54.448055" pageId="7" pageNumber="144" specimenCode="LACM 340600-340601, LACM 340602" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFEF5FED1FE0AEF28" box="[323,364,270,295]" pageId="7" pageNumber="144" type="male">2 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF93837DFECDFED1FEC4EF28" box="[379,418,270,295]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
; site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF93837DFDC2FED1FC65EF29" box="[628,771,270,294]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFF32FEF7FE71EF4F" box="[132,279,296,320]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<elevation id="0F83D1F4FF93837DFE95FEF7FEE9EF4F" box="[291,399,296,320]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="144" unit="m" value="29.0">
<quantity id="43569B22FF93837DFE95FEF7FE3EEF4F" box="[291,344,296,320]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="144" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF93837DFDE9FEF7FC65EF4F" box="[607,771,296,320]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-05" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF93837DFDE9FEF7FC65EF4F" box="[607,771,296,320]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-05" valueMin="2015-02-27">27.II-5.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFF32FE9CFF9FEF55" box="[132,249,322,346]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/126);
<collectionCode id="E2BFAE02FF93837DFD32FE9CFDD5EF55" box="[644,691,323,346]" pageId="7" pageNumber="144">BPT</collectionCode>
;
<collectionCode id="E2BFAE02FF93837DFD09FE9CFC60EF54" box="[703,774,323,347]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFF3AFE80FFC6EF79" box="[140,160,351,374]" pageId="7" pageNumber="144" type="male"></specimenCount>
<specimenCode id="D4089EBCFF93837DFF1EFE82FEFCEF7A" box="[168,410,349,373]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340600-340601</specimenCode>
,
<specimenCount id="92A8FD4EFF93837DFE13FE80FED0EF79" box="[421,438,351,374]" pageId="7" pageNumber="144" type="female"></specimenCount>
<specimenCode id="D4089EBCFF93837DFE09FE82FD3CEF7A" box="[447,602,349,373]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340602</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFDCDFE82FE3DEFD0" ID-GBIF-Occurrence="2595987906" collectingDate="2015-02-27" collectingDateMax="2015-03-08" collectingDateMin="2015-02-27" collectionCode="YPT, LACM" collectorName="Marc Pollet" country="French Guiana" elevation="25" latitude="2.2363334" longLatPrecision="1" longitude="-54.447083" pageId="7" pageNumber="144" specimenCode="LACM 340596, LACM 340597" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFDCDFE82FDC2EF79" box="[635,676,349,374]" pageId="7" pageNumber="144" type="male">3 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF93837DFD04FE82FDBEEF79" box="[690,728,349,374]" pageId="7" pageNumber="144" type="female">2 ♀</specimenCount>
; site MIT-C-RBF1;
<geoCoordinate id="E19A5000FF93837DFEACFEA7FEC1EF9F" box="[282,423,376,400]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="10.8" value="2.2363334">02°1410.8”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFE06FEA8FD26EF80" box="[432,576,375,399]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="49.5" value="-54.447083">54°2649.5”W</geoCoordinate>
;
<elevation id="0F83D1F4FF93837DFDFCFEA8FDC9EF80" box="[586,687,375,399]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">
<quantity id="43569B22FF93837DFDFCFEA8FD1AEF80" box="[586,636,375,399]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">25 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF93837DFF5BFE4DFEE8EFA5" box="[237,398,402,426]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF93837DFF5BFE4DFEE8EFA5" box="[237,398,402,426]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">27.II-8.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFE21FE4DFD6EEFA5" box="[407,520,402,426]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITA- RAKA/133);
<collectionCode id="E2BFAE02FF93837DFEB4FE72FE48EFCB" box="[258,302,429,452]" pageId="7" pageNumber="144">YPT</collectionCode>
; MNHN (
<specimenCount id="92A8FD4EFF93837DFE25FE73FEDEEFC9" box="[403,440,428,454]" pageId="7" pageNumber="144" type="male">2 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF93837DFE77FE73FE85EFC9" box="[449,483,428,454]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
),
<collectionCode id="E2BFAE02FF93837DFE43FE73FD5CEFCB" box="[501,570,428,452]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFDFEFE73FD0CEFC9" box="[584,618,428,454]" pageId="7" pageNumber="144" type="male">1 ♂</specimenCount>
<specimenCode id="D4089EBCFF93837DFDD8FE73FC65EFCB" box="[622,771,428,452]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340596</specimenCode>
,
<specimenCount id="92A8FD4EFF93837DFF32FE18FFCAEFEF" box="[132,172,455,480]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
<specimenCode id="D4089EBCFF93837DFF02FE18FE32EFD0" box="[180,340,455,479]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340597</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFEC3FE18FC61EC1C" ID-GBIF-Occurrence="2595987910" collectingDate="2015-02-27" collectingDateMax="2015-03-08" collectingDateMin="2015-02-27" collectionCode="WPT" collectorName="Marc Pollet" country="French Guiana" elevation="25" latitude="2.2363334" longLatPrecision="1" longitude="-54.447083" pageId="7" pageNumber="144" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFEC3FE18FEFBEFEF" box="[373,413,455,480]" pageId="7" pageNumber="144" type="female">2 ♀</specimenCount>
; site MIT-C-RBF1;
<geoCoordinate id="E19A5000FF93837DFDC7FE18FC65EFD0" box="[625,771,455,479]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="10.8" value="2.2363334">02°1410.8”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFF32FE3EFE71EFF6" box="[132,279,481,505]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="49.5" value="-54.447083">54°2649.5”W</geoCoordinate>
;
<elevation id="0F83D1F4FF93837DFE95FE3EFEE9EFF6" box="[291,399,481,505]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">
<quantity id="43569B22FF93837DFE95FE3EFE3EEFF6" box="[291,344,481,505]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">25 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF93837DFDE9FE3EFC65EFF6" box="[607,771,481,505]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF93837DFDE9FE3EFC65EFF6" box="[607,771,481,505]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">27.II-8.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFF32FE23FF93EC1B" box="[132,245,508,532]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/134);
<collectionCode id="E2BFAE02FF93837DFDC4FE23FDCCEC1C" box="[626,682,508,531]" pageId="7" pageNumber="144">WPT</collectionCode>
; MNHN
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFF21FDC9FD6CEC6C" ID-GBIF-Occurrence="2595987903" collectingDate="2015-02-27" collectingDateMax="2015-03-08" collectingDateMin="2015-02-27" collectionCode="BPT" collectorName="Marc Pollet" country="French Guiana" elevation="25" latitude="2.2363334" longLatPrecision="1" longitude="-54.447083" pageId="7" pageNumber="144" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFF21FDC9FFA6EC3F" box="[151,192,534,560]" pageId="7" pageNumber="144" type="male">1 ♂</specimenCount>
; site MIT-C-RBF1;
<geoCoordinate id="E19A5000FF93837DFE22FDC9FD43EC21" box="[404,549,534,558]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="10.8" value="2.2363334">02°1410.8”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFD87FDC9FDA3EC21" box="[561,709,534,558]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="49.5" value="-54.447083">54°2649.5”W</geoCoordinate>
;
<elevation id="0F83D1F4FF93837DFD67FDC9FFD7EC46" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">
<quantity id="43569B22FF93837DFD67FDC9FC61EC21" box="[721,775,534,558]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="144" unit="m" value="25.0">25 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF93837DFEC0FDEFFD71EC47" box="[374,535,560,584]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF93837DFEC0FDEFFD71EC47" box="[374,535,560,584]" pageId="7" pageNumber="144" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">27.II-8.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFD96FDEEFDE9EC46" box="[544,655,561,585]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/135);
<collectionCode id="E2BFAE02FF93837DFEC9FD93FEC8EC6C" box="[383,430,588,611]" pageId="7" pageNumber="144">BPT</collectionCode>
; MNHN
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFDA9FD94FD2EECBD" ID-GBIF-Occurrence="2595987909" collectingDate="2015-03-05" collectingDateMax="2015-03-10" collectingDateMin="2015-03-05" collectionCode="BPT, MNHN, LACM" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" longLatPrecision="1" longitude="-54.448055" pageId="7" pageNumber="144" specimenCode="LACM 340598" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFDA9FD94FD23EC6A" box="[543,581,587,613]" pageId="7" pageNumber="144" type="male">3 ♂</specimenCount>
; site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF93837DFF32FDB9FE69EC71" box="[132,271,614,638]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFEAEFDBAFEC1EC72" box="[280,423,613,637]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<quantity id="43569B22FF93837DFE06FDB9FE87EC72" box="[432,481,614,638]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="144" unit="m" value="29.0">
<elevation id="0F83D1F4FF93837DFE06FDB9FE87EC72" box="[432,481,614,638]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="144" unit="m" value="29.0">29 m</elevation>
</quantity>
a.s.l.; lowland rainforest;
<date id="F0101007FF93837DFD65FDBAFFBCEC97" pageId="7" pageNumber="144" value="2015-03-05" valueMax="2015-03-10" valueMin="2015-03-05">
<collectingDate id="E054E9EFFF93837DFD65FDBAFFBCEC97" pageId="7" pageNumber="144" value="2015-03-05" valueMax="2015-03-10" valueMin="2015-03-05">5-10. III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFF52FD5FFE33EC97" box="[228,341,640,664]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/137);
<collectionCode id="E2BFAE02FF93837DFD63FD5FFC62EC98" box="[725,772,640,663]" pageId="7" pageNumber="144">BPT</collectionCode>
;
<collectionCode id="E2BFAE02FF93837DFF32FD44FFB1ECBD" box="[132,215,667,690]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="7" pageNumber="144">MNHN</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFF50FD44FE68ECBB" box="[230,270,667,692]" pageId="7" pageNumber="144" type="male">2 ♂</specimenCount>
),
<collectionCode id="E2BFAE02FF93837DFE95FD44FE0CECBC" box="[291,362,667,691]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF93837DFECCFD44FEC6ECBB" box="[378,416,667,692]" pageId="7" pageNumber="144" type="male">1 ♂</specimenCount>
<specimenCode id="D4089EBCFF93837DFE11FD44FD27ECBD" box="[423,577,666,691]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340598</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF93837DFDD6FD44FE3BED0D" ID-GBIF-Occurrence="2595987907" collectingDate="2015-03-04" collectingDateMax="2015-03-10" collectingDateMin="2015-03-04" collectionCode="BPT" collectorName="Marc Pollet" country="French Guiana" elevation="28" latitude="2.2368057" longLatPrecision="1" longitude="-54.45225" pageId="7" pageNumber="144" specimenCode="RBF2, LACM 340599" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="92A8FD4EFF93837DFDD6FD44FDE2ECBB" box="[608,644,667,692]" pageId="7" pageNumber="144" type="female">1 ♀</specimenCount>
; site MIT-A-
<specimenCode id="D4089EBCFF93837DFF32FD6AFFA6ECC2" box="[132,192,693,717]" pageId="7" pageNumber="144">RBF2</specimenCode>
;
<geoCoordinate id="E19A5000FF93837DFF71FD6AFE37ECC2" box="[199,337,693,717]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="7" pageNumber="144" precision="1" seconds="12.5" value="2.2368057">02°1412.5”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF93837DFEEFFD6AFE83ECC2" box="[345,485,693,717]" degrees="54" direction="west" minutes="27" orientation="longitude" pageId="7" pageNumber="144" precision="1" seconds="08.1" value="-54.45225">54°2708.1”W</geoCoordinate>
;
<quantity id="43569B22FF93837DFE5BFD6AFD79ECC3" box="[493,543,693,717]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="144" unit="m" value="28.0">
<elevation id="0F83D1F4FF93837DFE5BFD6AFD79ECC3" box="[493,543,693,717]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="144" unit="m" value="28.0">28 m</elevation>
</quantity>
a.s.l.; lowland rainforest;
<date id="F0101007FF93837DFF32FD10FE6BECE8" box="[132,269,719,743]" pageId="7" pageNumber="144" value="2015-03-04" valueMax="2015-03-10" valueMin="2015-03-04">
<collectingDate id="E054E9EFFF93837DFF32FD10FE6BECE8" box="[132,269,719,743]" pageId="7" pageNumber="144" value="2015-03-04" valueMax="2015-03-10" valueMin="2015-03-04">4-10.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF93837DFEAFFD0FFEEDECE8" box="[281,395,719,743]" pageId="7" pageNumber="144">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/149);
<collectionCode id="E2BFAE02FF93837DFF32FD35FFD5ED0E" box="[132,179,746,769]" pageId="7" pageNumber="144">BPT</collectionCode>
;
<specimenCode id="D4089EBCFF93837DFF0BFD35FE3FED0D" box="[189,345,746,770]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="144" type="Museum">LACM 340599</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="D0D1664FFF91837FFF32F84BFE3EE9E5" ID-DOI="http://doi.org/10.5281/zenodo.3743148" ID-Zenodo-Dep="3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="5" pageNumber="142" startId="5.[132,143,1940,1958]" targetBox="[160,1421,243,1885]" targetPageId="5">
<paragraph id="841136C7FF91837FFF32F84BFE3EE9E5" blockId="5.[132,1457,1940,2026]" pageId="5" pageNumber="142">
FIG. 1. —
<taxonomicName id="43AE4D44FF91837FFF63F84BFEE7E9A9" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[213,385,1940,1958]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF91837FFF63F84BFEE7E9A9" box="[213,385,1940,1958]" italics="true" pageId="5" pageNumber="142">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF91837FFE31F84BFE9BE9A9" box="[391,509,1940,1958]" pageId="5" pageNumber="142" rank="species">n. gen., n. sp.</taxonomicNameLabel>
:
<emphasis id="B6DAEAD5FF91837FFDB2F84AFD77E9A8" bold="true" box="[516,529,1941,1959]" pageId="5" pageNumber="142">A</emphasis>
, male head, frontal view;
<emphasis id="B6DAEAD5FF91837FFD5EF84AFD93E9A8" bold="true" box="[744,757,1941,1959]" pageId="5" pageNumber="142">B</emphasis>
, wing;
<emphasis id="B6DAEAD5FF91837FFC84F84AFC26E9A8" bold="true" box="[818,832,1941,1959]" pageId="5" pageNumber="142">C</emphasis>
, male antennal flagellomeres 3-4, dorsal view;
<emphasis id="B6DAEAD5FF91837FFB7FF84AFBB1E9A8" bold="true" box="[1225,1239,1941,1959]" pageId="5" pageNumber="142">D</emphasis>
, male antennal flagellomeres 12-14, dorsal view;
<emphasis id="B6DAEAD5FF91837FFEF8F874FE3CE9B2" bold="true" box="[334,346,1963,1981]" pageId="5" pageNumber="142">E</emphasis>
, male terminalia,epandrium removed, dorsal view;
<emphasis id="B6DAEAD5FF91837FFCB5F874FC68E9B2" bold="true" box="[771,782,1963,1981]" pageId="5" pageNumber="142">F</emphasis>
, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view;
<emphasis id="B6DAEAD5FF91837FFF32F81DFFF4E9DB" bold="true" box="[132,146,1986,2004]" pageId="5" pageNumber="142">G</emphasis>
, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm.
</paragraph>
</caption>
<caption id="D0D1664FFF92837CFF32F81CFAE8E9E4" ID-DOI="http://doi.org/10.5281/zenodo.3743150" ID-Zenodo-Dep="3743150" httpUri="https://zenodo.org/record/3743150/files/figure.png" pageId="6" pageNumber="143" startId="6.[132,143,1987,2005]" targetBox="[163,1420,231,1895]" targetPageId="6">
<paragraph id="841136C7FF92837CFF32F81CFAE8E9E4" blockId="6.[132,1455,1987,2028]" pageId="6" pageNumber="143">
FIG. 2. —
<taxonomicName id="43AE4D44FF92837CFF63F81CFEE7E9DA" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[213,385,1987,2005]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF92837CFF63F81CFEE7E9DA" box="[213,385,1987,2005]" italics="true" pageId="6" pageNumber="143">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF92837CFE31F81CFE99E9DA" box="[391,511,1987,2005]" pageId="6" pageNumber="143" rank="species">
<taxonomicNameLabel id="ADE957AEFF92837CFE31F81CFEA3E9DA" box="[391,453,1987,2005]" pageId="6" pageNumber="143" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</taxonomicNameLabel>
:
<emphasis id="B6DAEAD5FF92837CFDB0F81CFD75E9DA" bold="true" box="[518,531,1987,2005]" pageId="6" pageNumber="143">A</emphasis>
, male terminalia, lateral view;
<emphasis id="B6DAEAD5FF92837CFCA6F81CFC7BE9DA" bold="true" box="[784,797,1987,2005]" pageId="6" pageNumber="143">B</emphasis>
, male epandrium, proctiger and epandrial claspers, dorsal view;
<emphasis id="B6DAEAD5FF92837CFA8BF81CFA2DE9DA" bold="true" box="[1341,1355,1987,2005]" pageId="6" pageNumber="143">C</emphasis>
, female terminalia, external structure, ventral view;
<emphasis id="B6DAEAD5FF92837CFE65F805FE87E9E3" bold="true" box="[467,481,2010,2028]" pageId="6" pageNumber="143">D</emphasis>
, female terminalia, internal structure, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="CCB4654CFF93837DFF32FCFDFDF5ED60" pageId="7" pageNumber="144" type="etymology">
<paragraph id="841136C7FF93837DFF32FCFDFDF5ED60" blockId="7.[132,775,802,879]" pageId="7" pageNumber="144">ETYMOLOGY. — The specific epithet is derived form the Latin word “subulatus”, meaning awl-shaped, in reference to the prominent, spiniform seta inserted at the apex of each gonostylus.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFF32FC4FFD5BEDA6" box="[132,573,912,937]" pageId="7" pageNumber="144" type="diagnosis">
<paragraph id="841136C7FF93837DFF32FC4FFD5BEDA6" blockId="7.[132,573,912,937]" box="[132,573,912,937]" pageId="7" pageNumber="144">
DIAGNOSIS. — See above,
<taxonomicName id="43AE4D44FF93837DFE30FC4FFE94EDA7" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[390,498,912,936]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF93837DFE30FC4FFE94EDA7" box="[390,498,912,936]" italics="true" pageId="7" pageNumber="144">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF93837DFE4FFC4EFD5BEDA6" box="[505,573,913,937]" pageId="7" pageNumber="144" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFF32FC17FEB4EDF4" pageId="7" pageNumber="144" type="distribution">
<paragraph id="841136C7FF93837DFF32FC17FEB4EDF4" blockId="7.[132,775,968,1020]" pageId="7" pageNumber="144">
DISTRIBUTION. — Currently known from multiple localities in the Mitaraka range of
<collectingCountry id="FCB97657FF93837DFE8FFC3CFEABEDF4" box="[313,461,995,1019]" name="French Guiana" pageId="7" pageNumber="144">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFF32FBC3FD9AEA41" pageId="7" pageNumber="144" type="biology_ecology">
<paragraph id="841136C7FF93837DFF32FBC3FD9AEA41" blockId="7.[132,775,1052,1102]" pageId="7" pageNumber="144">HABITAT. — Little information regarding specific habitat is available; however, all specimens were collected in lowland rainforest.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFF32FBAAFB35EA3E" pageId="7" pageNumber="144" type="description">
<paragraph id="841136C7FF93837DFF32FBAAFE73EA81" blockId="7.[132,776,1141,2030]" box="[132,277,1141,1167]" pageId="7" pageNumber="144">DESCRIPTION</paragraph>
<paragraph id="841136C7FF93837DFF32FB49FFDFEABF" blockId="7.[132,776,1141,2030]" box="[132,185,1174,1200]" pageId="7" pageNumber="144">
<emphasis id="B6DAEAD5FF93837DFF32FB49FFDFEABF" box="[132,185,1174,1200]" italics="true" pageId="7" pageNumber="144">Male</emphasis>
</paragraph>
<paragraph id="841136C7FF93837DFF32FB69FB78ECFE" blockId="7.[132,776,1141,2030]" lastBlockId="7.[813,1456,217,753]" pageId="7" pageNumber="144">
Measurements, (N = 5) head width
<quantity id="43569B22FF93837DFDA5FB69FD1CEADF" box="[531,634,1206,1232]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="144" unit="mm" value="0.37">0.37 mm</quantity>
(0.34-0.40), head height
<quantity id="43569B22FF93837DFEA6FB09FE1EEAFF" box="[272,376,1238,1264]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="144" unit="mm" value="0.33">0.33 mm</quantity>
(0.30-0.34), wing length
<quantity id="43569B22FF93837DFD29FB09FC61EAFF" box="[671,775,1238,1264]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="144" unit="mm" value="1.7">1.70 mm</quantity>
(1.64-1.76), wing width
<quantity id="43569B22FF93837DFE33FB29FE8FEB1F" box="[389,489,1270,1296]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="144" unit="mm" value="0.44">0.44 mm</quantity>
. Head with vertex extended slightly posteriorly, eyebridge comprised of 3 facet rows, contiguous but slightly constricted at median. Antennae: scape cylindrical, about 1.5 times longer than wide; pedicel globular; flagellomeres fusiform, with paired ascoids on f4 f7, single ascoids on f8 f11. Frons and clypeus distinctly divided; frontal scar patch bilobed posteriorly, contiguous medially. Mouthparts inconspicuous, obscured by clypeus, extending slightly beyond apex of palp segment 1; palpi typical of
<taxonomicName id="43AE4D44FF93837DFF32F9CBFE68E821" authorityName="Newman" authorityYear="1834" box="[132,270,1556,1582]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
, with apical segment annulated; palp segment proportions: 1, 2, 2.18, 2.18; labellum about as wide as anterior margin of clypeus. Wing: strongly lanceolate; radial and medial fork positioned at a level basal to the apex of CuA; R
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terminating in C posterior to wing apex; M
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incomplete at base; CuA conspicuously widened at base. Male terminalia: bilaterally symmetrical; hypandrium flat, band-like, arched posteriorly, fused with gonocoxites laterally; hypandrium in some specimens appearing folded along median (
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); intersegmental membrane connecting between anterior margin of hypandrium and posterior margin of abdominal segment 8, covering base of ejaculatory apodeme; ejaculatory apodeme dorsoventrally compressed at base, dome-like at center, articulating posteriorly with aedeagal sclerites; aedeagus comprised of four blade-like sclerites (
<figureCitation id="1C952A42FF93837DFD27F80CFDADE9E2" box="[657,715,2003,2029]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="7" pageNumber="144">Fig. 1</figureCitation>
E-G) surrounded by a tubular membrane; innermost aedeagal sclerites spathiform, expanding dorsally in extended aedeagus; outer sclerites sinuous, longer than inner sclerites, expanding lateroventrally in extended aedeagus (
<figureCitation id="1C952A42FF93837DFB55FEE6FA4BEF5C" box="[1251,1325,313,339]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="7" pageNumber="144">Fig. 1F</figureCitation>
); parameres dorsoventrally compressed, flat, with pointed, hook-like projection posteromedially; gonostyli tapered gradually from base to apex, curved ventrally along apical half, appearing pitted ventrally; gonocoxites reniform, about two times as long as wide, setose, with setae alveoli laterally and dorsally; gonocoxal apodemes inflated, triangular in dorsoventral view, bulbous in lateral view, with dorsal paramere extending from posterior margin; subepandrial sclerite narrow, band-like, with lateral arms claw-shaped in dorsoventral view, linked to base of epandrial claspers; epandrium rectangular, narrowing posteriorly; epandrial claspers elongate, tapered, curved dorsally from base to apex, with single, setiform tenaculum inserted dorsoapically.
</paragraph>
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<emphasis id="B6DAEAD5FF93837DFC9BFCC8FC10ED3E" box="[813,886,791,817]" italics="true" pageId="7" pageNumber="144">Female</emphasis>
</paragraph>
<paragraph id="841136C7FF93837DFC9BFCE7FB35EA3E" blockId="7.[813,1456,791,1073]" pageId="7" pageNumber="144">
Measurements, (N = 3) head width
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(0.32-0.36), head height
<quantity id="43569B22FF93837DFC0FFC88FB47ED7D" box="[953,1057,855,882]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="144" unit="mm" value="0.33">0.33 mm</quantity>
(0.32-0.34), wing length
<quantity id="43569B22FF93837DFAFEFC88FAD6ED7D" box="[1352,1456,855,882]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="7" pageNumber="144" unit="mm" value="1.84">1.84 mm</quantity>
(1.76-1.88), wing width
<quantity id="43569B22FF93837DFBF5FCA8FBCDED9E" box="[1091,1195,887,913]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="144" unit="mm" value="0.48">0.48 mm</quantity>
(0.44-0.52). Head and wing generally as in male; palp segment proportions: 1, 2.50, 3, 3. Terminalia: cerci elongate, heavily sclerotized, spatulate in lateral view, with numerous setae inserted on medial surface; posterior margin of sternum 8 bilobed, oviduct complex, ornate internally (
<figureCitation id="1C952A42FF93837DFC59FBC9FB25EA3F" box="[1007,1091,1046,1072]" captionStart="FIG" captionStartId="6.[132,143,1987,2005]" captionTargetBox="[163,1420,231,1895]" captionTargetId="figure@6.[132,1455,216,1902]" captionTargetPageId="6" captionText="FIG. 2. — Polletomyia subulata n. gen., n. sp.: A, male terminalia, lateral view; B, male epandrium, proctiger and epandrial claspers, dorsal view; C, female terminalia, external structure, ventral view; D, female terminalia, internal structure, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743150" httpUri="https://zenodo.org/record/3743150/files/figure.png" pageId="7" pageNumber="144">Fig. 2D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,218 @@
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<mods:title id="915870D91E9316CEB331792EE71F3D1D">Descriptions of two new genera of Maruinini (Diptera, Psychodidae, Psychodinae) from the Mitaraka range of French Guiana</mods:title>
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<paragraph id="841136C7FF90837EFE8DFAC9FD37EB3F" blockId="4.[315,593,1301,1328]" box="[315,593,1301,1328]" pageId="4" pageNumber="141">
<heading id="DF5981ABFF90837EFE8DFAC9FD37EB3F" box="[315,593,1301,1328]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="141" reason="2">
Genus
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<emphasis id="B6DAEAD5FF90837EFE3EFACAFE99EB20" box="[392,511,1301,1327]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFDB0FAC9FD37EB3F" box="[518,593,1302,1328]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="CCB4654CFF90837EFF32FA53FD1AEBAB" box="[132,636,1420,1445]" pageId="4" pageNumber="141" type="type_taxon">
<paragraph id="841136C7FF90837EFF32FA53FD1AEBAB" blockId="4.[132,636,1420,1445]" box="[132,636,1420,1445]" pageId="4" pageNumber="141">
<typeStatus id="5B158865FF90837EFF32FA53FFD3EBAE" box="[132,181,1420,1444]" pageId="4" pageNumber="141">TYPE</typeStatus>
SPECIES. —
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<emphasis id="B6DAEAD5FF90837EFE98FA53FE96EBAB" box="[302,496,1420,1444]" italics="true" pageId="4" pageNumber="141">Polletomyia subulata</emphasis>
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<taxonomicNameLabel id="ADE957AEFF90837EFE41FA52FD1AEBAB" box="[503,636,1421,1445]" pageId="4" pageNumber="141" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="CCB4654CFF90837EFF32FA64FF8AEBE1" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFF32FA64FF8AEBE1" blockId="4.[132,776,1467,1518]" pageId="4" pageNumber="141">
ILLUSTRATIONS. — Head:
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; Wing:
<figureCitation id="1C952A42FF90837EFD97FA63FD0FEBDB" box="[545,617,1468,1492]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1B</figureCitation>
; Male genitalia:
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E-G.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F9DAFD9BE85C" pageId="4" pageNumber="141" type="etymology">
<paragraph id="841136C7FF90837EFF32F9DAFD9BE85C" blockId="4.[132,775,1541,1619]" pageId="4" pageNumber="141">
ETYMOLOGY. — This genus is named in honor of Dr Marc Pollet, who collected the
<typeStatus id="5B158865FF90837EFE98F9FEFE31E836" box="[302,343,1569,1593]" pageId="4" pageNumber="141">type</typeStatus>
specimens, forwarded them to the author and provided supporting information during the course of this study.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F9B5FDE6E959" pageId="4" pageNumber="141" type="diagnosis">
<paragraph id="841136C7FF90837EFF32F9B5FDE6E959" blockId="4.[132,776,1642,1878]" pageId="4" pageNumber="141">
DIAGNOSIS. — The following combination of characters is diagnostic for the genus: male head: antennal flagellomeres fusiform, with paired, unbranched ascoids on f4 f7 (
<figureCitation id="1C952A42FF90837EFE4BF940FD2FE8B8" box="[509,585,1695,1719]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1C</figureCitation>
), single ascoids on f8 f11; male terminalia: bilaterally symmetrical; aedeagus comprised of four movable, blade-like sclerites surrounded by membranous sheath; outer aedeagal sclerites sinuous, spreading laterally when extended posteriorly (
<figureCitation id="1C952A42FF90837EFEEAF8D6FEC4E92E" box="[348,418,1801,1825]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1F</figureCitation>
); gonostyli with stout, conical setae inserted apically (
<figureCitation id="1C952A42FF90837EFE85F8FBFE0EE933" box="[307,360,1828,1852]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1</figureCitation>
E-G); epandrial claspers each with single, setiform tenaculum inserted dorsoapically (
<figureCitation id="1C952A42FF90837EFD9DF8E1FD14E959" box="[555,626,1854,1878]" captionStart="FIG" captionStartId="6.[132,143,1987,2005]" captionTargetBox="[163,1420,231,1895]" captionTargetId="figure@6.[132,1455,216,1902]" captionTargetPageId="6" captionText="FIG. 2. — Polletomyia subulata n. gen., n. sp.: A, male terminalia, lateral view; B, male epandrium, proctiger and epandrial claspers, dorsal view; C, female terminalia, external structure, ventral view; D, female terminalia, internal structure, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743150" httpUri="https://zenodo.org/record/3743150/files/figure.png" pageId="4" pageNumber="141">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F8B2FE91E9AF" pageId="4" pageNumber="141" type="distribution">
<paragraph id="841136C7FF90837EFF32F8B2FE91E9AF" blockId="4.[132,775,1901,1953]" pageId="4" pageNumber="141">
DISTRIBUTION. —
<taxonomicName id="43AE4D44FF90837EFE8CF8B1FEC0E989" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[314,422,1902,1926]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFE8CF8B1FEC0E989" box="[314,422,1902,1926]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFE1CF8B1FE8AE989" box="[426,492,1902,1926]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
is currently known only from the Mitaraka range of
<collectingCountry id="FCB97657FF90837EFEE8F857FE95E9AF" box="[350,499,1928,1952]" name="French Guiana" pageId="4" pageNumber="141">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F868FD9AE9E5" pageId="4" pageNumber="141" type="biology_ecology">
<paragraph id="841136C7FF90837EFF32F868FD9AE9E5" blockId="4.[132,775,1975,2026]" pageId="4" pageNumber="141">HABITAT. — Little information regarding specific habitat is available; however, all specimens were collected in lowland rainforest.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFC9BFF06FB6FE800" pageId="4" pageNumber="141" type="discussion">
<paragraph id="841136C7FF90837EFC9BFF06FCF7EEFE" blockId="4.[813,1456,217,1551]" box="[813,913,217,243]" pageId="4" pageNumber="141">REMARKS</paragraph>
<paragraph id="841136C7FF90837EFC9BFF26FBFAEC7D" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
The relatively small size of
<taxonomicName id="43AE4D44FF90837EFBF0FF26FBDBEF1C" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1094,1213,249,275]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFBF0FF26FBDBEF1C" box="[1094,1213,249,275]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB75FF25FA68EF1B" box="[1219,1294,250,276]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
, as well as their lanceolate wings and dark coloration make it possible to differentiate them in a sample comprised of many psychodid genera and species. These superficial characters also make it possible to associate males and females, as the latter are otherwise lacking in characters that readily differentiate them from other psychodine females. Indeed, if further species of
<taxonomicName id="43AE4D44FF90837EFC9BFE07FCC2EFFD" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[813,932,472,498]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFC9BFE07FCC2EFFD" box="[813,932,472,498]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFC1CFE06FC95EFFC" box="[938,1011,473,499]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
are discovered, it may become difficult or impossible to differentiate females within the genus. Despite the relative ease of differentiating them in a bulk sample, accurate diagnosis of this genus is dependent on examination of slide-mounted male terminalia.
</paragraph>
<paragraph id="841136C7FF90837EFCF5FDA7FCB2EA3F" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
Currently, the following character states are sufficient for distinguishing this genus from all other
<taxonomicName id="43AE4D44FF90837EFB41FD47FA08ECBD" authorityName="Enderlein" authorityYear="1937" box="[1271,1390,664,690]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
: male terminalia bilaterally symmetrical; gonostyli each with a conical, spiniform seta inserted apically; tenacula setiform. Nonetheless, additional characters are included above in the diagnosis because they are also unique to the genus, thereby making the diagnosis more robust. Some other genera of
<taxonomicName id="43AE4D44FF90837EFC9BFC88FCF9ED7E" authorityName="Enderlein" authorityYear="1937" box="[813,927,855,881]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
also have bilaterally symmetrical male terminalia in some or all constituent species (e.g.
<taxonomicName id="43AE4D44FF90837EFB7BFCA8FA76ED9E" authorityName="Enderlein" authorityYear="1937" box="[1229,1296,887,913]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFB7BFCA8FA76ED9E" box="[1229,1296,887,913]" italics="true" pageId="4" pageNumber="141">Alepia</emphasis>
</taxonomicName>
,
<taxonomicName id="43AE4D44FF90837EFAABFCA8FC2CEDBF" authorityName="Wagner" authorityYear="1981" class="Insecta" family="Psychodidae" genus="Caenobrunettia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAABFCA8FC2CEDBF" italics="true" pageId="4" pageNumber="141">Caenobrunettia</emphasis>
</taxonomicName>
,
<taxonomicName id="43AE4D44FF90837EFCECFC48FB3FEDBD" authority="Muller, 1895" authorityName="Muller" authorityYear="1895" box="[858,1113,919,946]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFCECFC48FCDDEDBF" box="[858,955,919,944]" italics="true" pageId="4" pageNumber="141">Maruina</emphasis>
Müller, 1895
</taxonomicName>
, and
<taxonomicName id="43AE4D44FF90837EFB2EFC48FA70EDBF" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1176,1302,919,944]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFB2EFC48FA70EDBF" box="[1176,1302,919,944]" italics="true" pageId="4" pageNumber="141">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFA96FC48FA17EDBE" box="[1312,1393,919,945]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
). Yet, each of these groups has a unique build to their aedeagus that is distinct from
<taxonomicName id="43AE4D44FF90837EFBA7FC09FBEDEDFF" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1041,1163,982,1008]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFBA7FC09FBEDEDFF" box="[1041,1163,982,1008]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB25FC08FB86EDFE" box="[1171,1248,983,1009]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
and none of them have conspicuous, spiniform setae inserted at the apices of their gonostyli.
</paragraph>
<paragraph id="841136C7FF90837EFCF5FBE9FB6FE800" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
Aside from the male terminalia, some character states present in
<taxonomicName id="43AE4D44FF90837EFCCEFB89FC89EA7F" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[888,1007,1110,1136]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFCCEFB89FC89EA7F" box="[888,1007,1110,1136]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFC42FB88FB5BEA7E" box="[1012,1085,1111,1137]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
are found as well in other genera of
<taxonomicName id="43AE4D44FF90837EFC9BFBA9FCFBEA9F" authorityName="Enderlein" authorityYear="1937" box="[813,925,1142,1168]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
, such as a contiguous eye bridge comprised of three rows of ommatidia. These states are also present in
<taxonomicName id="43AE4D44FF90837EFAE6FB48FAC9EABF" authorityName="Muller" authorityYear="1895" box="[1360,1455,1175,1200]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAE6FB48FAC9EABF" box="[1360,1455,1175,1200]" italics="true" pageId="4" pageNumber="141">Maruina</emphasis>
</taxonomicName>
(so far only in subgenus
<taxonomicName id="43AE4D44FF90837EFB9DFB69FA7FEADF" authority="Hogue, 1973" authorityName="Hogue" authorityYear="1973" box="[1067,1305,1206,1232]" class="Insecta" family="Psychodidae" genus="Aculcina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFB9DFB69FBE3EADF" box="[1067,1157,1206,1232]" italics="true" pageId="4" pageNumber="141">Aculcina</emphasis>
Hogue, 1973
</taxonomicName>
) and
<taxonomicName id="43AE4D44FF90837EFAE5FB69FCAEEAFF" authority="Satchell, 1955" authorityName="Satchell" authorityYear="1955" class="Insecta" family="Psychodidae" genus="Arisemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAE5FB69FAC9EADF" box="[1363,1455,1206,1232]" italics="true" pageId="4" pageNumber="141">Arisemus</emphasis>
<bibRefCitation id="E03F4B36FF90837EFC9BFB09FCAEEAFF" author="SATCHELL G. H." box="[813,968,1237,1264]" pageId="4" pageNumber="141" pagination="85 - 93" refId="ref7264" refString="SATCHELL G. H. 1955. - Two new subgenera of Psychodidae (Diptera) from Jamaica, with descriptions of five new species. Annals and Magazine of Natural History 12: 85 - 93. https: // doi. org / 10.1080 / 00222935508651831" type="journal article" year="1955">Satchell, 1955</bibRefCitation>
</taxonomicName>
and have been indicated by some authors as diagnostic for those genera (e.g.
<bibRefCitation id="E03F4B36FF90837EFB3DFB2AFAF4EB1F" author="CAMICO J. &amp; CORDEIRO D. &amp; CHAGAS C." box="[1163,1426,1269,1296]" pageId="4" pageNumber="141" pagination="296 - 316" refId="ref6793" refString="CAMICO J., CORDEIRO D. &amp; CHAGAS C. 2019. - Maruina Muller, 1895 (Diptera: Psychodidae): description of four new species from Brazil, with revised diagnoses for the genus and subgenera. Zootaxa 4652 (2): 296 - 316. https: // doi. org / 10.11646 / zootaxa. 4652.2.4" type="journal article" year="2019">
Camico
<emphasis id="B6DAEAD5FF90837EFB51FB29FA7DEB00" box="[1255,1307,1269,1295]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2019:
<quantity id="43569B22FF90837EFADEFB29FAF4EB1F" box="[1384,1426,1270,1296]" metricMagnitude="0" metricUnit="m" metricValue="7.5691999999999995" pageId="4" pageNumber="141" unit="in" value="298.0">298</quantity>
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in their diagnosis of
<emphasis id="B6DAEAD5FF90837EFC59FAC9FB69EB20" box="[1007,1039,1302,1327]" italics="true" pageId="4" pageNumber="141">M.</emphasis>
(
<taxonomicName id="43AE4D44FF90837EFBA9FACAFB1FEB20" authorityName="Hogue" authorityYear="1973" box="[1055,1145,1301,1327]" class="Insecta" family="Psychodidae" genus="Aculcina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFBA9FACAFB1FEB20" box="[1055,1145,1301,1327]" italics="true" pageId="4" pageNumber="141">Aculcina</emphasis>
</taxonomicName>
)). Despite this, the presence of these character states in multiple genera indicates that they are not diagnostic in and of themselves. Rather, it is necessary to determine whether there are other nuances that distinguish the eye bridge of each genus from others that have the same general appearance. In the case of
<taxonomicName id="43AE4D44FF90837EFB1BFA6AFA40EBC0" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1197,1318,1461,1487]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFB1BFA6AFA40EBC0" box="[1197,1318,1461,1487]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFA86FA6AFAE6EBC0" box="[1328,1408,1461,1487]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
, the eye bridge is not particularly distinct, so it is omitted as a diagnostic character.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFC9BF9EBFA75E860" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFC9BF9EBFCDBE843" blockId="4.[813,1299,1588,1647]" box="[813,957,1588,1614]" pageId="4" pageNumber="141">DESCRIPTION</paragraph>
<paragraph id="841136C7FF90837EFC9BF98BFA75E860" blockId="4.[813,1299,1588,1647]" box="[813,1299,1620,1647]" pageId="4" pageNumber="141">
See below,
<taxonomicName id="43AE4D44FF90837EFC15F98BFB1FE861" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[931,1145,1620,1646]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF90837EFC15F98BFB1FE861" box="[931,1145,1620,1646]" italics="true" pageId="4" pageNumber="141">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB36F98AFA75E860" box="[1152,1299,1621,1647]" pageId="4" pageNumber="141" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="CC714562822B35F3356BC4BC9F8849C4">Descriptions of two new genera of Maruinini (Diptera, Psychodidae, Psychodinae) from the Mitaraka range of French Guiana</mods:title>
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<mods:detail id="437E029A2E07C74828B146DD1C30CEC4" type="volume">
<mods:number id="D422A7D79076A5836900045B6FA7655A">42</mods:number>
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<mods:detail id="0AF11C04B891C0EC5FCCD6186B3BC068" type="issue">
<mods:number id="4F0A73380103566D482A72AFD3BF3BC8">10</mods:number>
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<mods:start id="1E56B6B7D51A53B004DFC2EF84880CB4">139</mods:start>
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<mods:classification id="FD5E3BAD3E51709E920B372E3ADC8AF8">journal article</mods:classification>
<mods:identifier id="C0517CE4BC0ADEB75AEF411FC24D1E1E" type="DOI">10.5252/zoosystema2020v42a10</mods:identifier>
<mods:identifier id="A77AB62192875C8F9D9626BBF6C4D42E" type="GBIF-Dataset">60ca36d6-4c1f-4050-88e2-38e8b28f3f5c</mods:identifier>
<mods:identifier id="700634763F90D3F414A956C61F250BCB" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="2CBCAF4372AC9DFD31A9FB3FBACC59CD" type="Zenodo-Dep">3743146</mods:identifier>
<mods:identifier id="201157A1F128EE5771838EEDB55466D3" type="ZooBank">urn:lsid:zoobank.org:pub:21B9187A-D055-4946-8922-53FD10A159CE</mods:identifier>
</mods:mods>
<treatment id="0C0787D1FF938370FC57FBA9FD11E9E1" ID-DOI="http://doi.org/10.5281/zenodo.3809874" ID-GBIF-Taxon="163536909" ID-Zenodo-Dep="3809874" LSID="urn:lsid:zoobank.org:act:510873DE-6E58-4A95-9A5C-1D311DC38A70" httpUri="http://treatment.plazi.org/id/0C0787D1FF938370FC57FBA9FD11E9E1" lastPageId="10" lastPageNumber="147" pageId="7" pageNumber="144">
<subSubSection id="CCB4654CFF93837DFC57FBA9FB9DEA9E" box="[993,1275,1142,1169]" pageId="7" pageNumber="144" type="nomenclature">
<paragraph id="841136C7FF93837DFC57FBA9FB9DEA9E" blockId="7.[993,1275,1142,1169]" box="[993,1275,1142,1169]" pageId="7" pageNumber="144">
<heading id="DF5981ABFF93837DFC57FBA9FB9DEA9E" box="[993,1275,1142,1169]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="144" reason="2">
Genus
<taxonomicName id="43AE4D44FF93837DFB98FBA8FBCFEA9F" ID-CoL="8H3JS" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1070,1193,1143,1168]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF93837DFB98FBA8FBCFEA9F" box="[1070,1193,1143,1168]" italics="true" pageId="7" pageNumber="144">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF93837DFB06FBA8FB9DEA9E" box="[1200,1275,1143,1169]" pageId="7" pageNumber="144" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFCF1FB69FAF3EAC7" box="[839,1429,1206,1224]" pageId="7" pageNumber="144" type="description">
<paragraph id="841136C7FF93837DFCF1FB69FAF3EAC7" blockId="7.[839,1429,1206,1224]" box="[839,1429,1206,1224]" pageId="7" pageNumber="144">
<uri id="F03F3AC5FF93837DFCF1FB69FAF3EAC7" box="[839,1429,1206,1224]" pageId="7" pageNumber="144">
urn:lsid:zoobank.org:act:
<uuid id="F0080C12FF93837DFBA6FB69FAF3EAC7" box="[1040,1429,1206,1224]" pageId="7" pageNumber="144">510873DE-6E58-4A95-9A5C-1D311DC38A70</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BFB31FA56EB09" box="[813,1328,1262,1287]" pageId="7" pageNumber="144" type="type_taxon">
<paragraph id="841136C7FF93837DFC9BFB31FA56EB09" blockId="7.[813,1328,1262,1287]" box="[813,1328,1262,1287]" pageId="7" pageNumber="144">
<typeStatus id="5B158865FF93837DFC9BFB31FC3BEB0C" box="[813,861,1262,1286]" pageId="7" pageNumber="144">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="43AE4D44FF93837DFC61FB30FBC2EB09" ID-CoL="8H4CK" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[983,1188,1262,1286]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF93837DFC61FB30FBC2EB09" box="[983,1188,1262,1286]" italics="true" pageId="7" pageNumber="144">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF93837DFB1CFB30FA56EB09" box="[1194,1328,1263,1287]" pageId="7" pageNumber="144" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BFAFCFCFEEB5A" pageId="7" pageNumber="144" type="description">
<paragraph id="841136C7FF93837DFC9BFAFCFCFEEB5A" blockId="7.[813,1456,1315,1366]" pageId="7" pageNumber="144">
ILLUSTRATIONS. — Wing:
<figureCitation id="1C952A42FF93837DFB87FAFCFB1FEB34" box="[1073,1145,1315,1339]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="7" pageNumber="144">Fig. 3A</figureCitation>
; Head:
<figureCitation id="1C952A42FF93837DFB7FFAFCFA77EB34" box="[1225,1297,1315,1339]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="7" pageNumber="144">Fig. 3B</figureCitation>
; Male genitalia:
<figureCitation id="1C952A42FF93837DFC9BFAE1FC05EB59" box="[813,867,1342,1366]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="7" pageNumber="144">Fig. 3</figureCitation>
C-D.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BFAADFA4EEBFB" pageId="7" pageNumber="144" type="etymology">
<paragraph id="841136C7FF93837DFC9BFAADFA4EEBFB" blockId="7.[813,1456,1394,1524]" pageId="7" pageNumber="144">
ETYMOLOGY. — This genus name is derived from the Latin “myia”, meaning fly and “mystax”, meaning mustache, in reference to the maxillary palps of the male. Before treating the specimens with KOH, the setose palps of male
<taxonomicName id="43AE4D44FF93837DFBEFFA1CFBB4EBD5" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1113,1234,1474,1498]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF93837DFBEFFA1CFBB4EBD5" box="[1113,1234,1474,1498]" italics="true" pageId="7" pageNumber="144">M. trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF93837DFB6EFA1DFA3DEBD6" box="[1240,1371,1474,1498]" pageId="7" pageNumber="144" rank="species">n. gen., n. sp.</taxonomicNameLabel>
gave the appearance of the fly having a prominent mustache.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BF9CEFB46E943" pageId="7" pageNumber="144" type="diagnosis">
<paragraph id="841136C7FF93837DFC9BF9CEFB46E943" blockId="7.[813,1456,1553,1868]" pageId="7" pageNumber="144">DIAGNOSIS. — Male head: antennal flagellomeres 1-5 elongate, cylindrical; all flagellomeres lacking ascoids; clypeus elongate, expanded laterally, with lateral margins overlapping medial margins of eyes; palpi conspicuously elongate, with segment 1 hook-like, segment 4 inflated apically, capitate; palp segments with clusters of spatulate setae inserted dorsally and ventrally; palps folded behind head at rest. Male terminalia: bilaterally symmetrical; gonocoxites glabrous, without setae alveoli; aedeagal sclerites paired, laterally compressed, spatulate. Female head: maxillary palpi not elongated. Female terminalia: posterior margin of sternum 8 trilobed, with median lobe wider than flanking lobes; oviduct with microtrichia internally on membrane.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BF8B6FBF9E993" pageId="7" pageNumber="144" type="distribution">
<paragraph id="841136C7FF93837DFC9BF8B6FBF9E993" blockId="7.[813,1455,1897,1948]" pageId="7" pageNumber="144">
DISTRIBUTION. —
<taxonomicName id="43AE4D44FF93837DFC57F8B5FB36E98E" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[993,1104,1898,1921]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF93837DFC57F8B5FB36E98E" box="[993,1104,1898,1921]" italics="true" pageId="7" pageNumber="144">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF93837DFBE2F8B5FBF0E98D" box="[1108,1174,1898,1922]" pageId="7" pageNumber="144" rank="genus">n. gen.</taxonomicNameLabel>
is currently known only from the Mitaraka range of
<collectingCountry id="FCB97657FF93837DFBB0F85BFBFDE993" box="[1030,1179,1924,1948]" name="French Guiana" pageId="7" pageNumber="144">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF93837DFC9BF867FAC2E9E4" pageId="7" pageNumber="144" type="biology_ecology">
<paragraph id="841136C7FF93837DFC9BF867FAC2E9E4" blockId="7.[813,1455,1976,2027]" pageId="7" pageNumber="144">HABITAT. — Little information regarding specific habitat is available; however, all specimens were collected in lowland rainforest.</paragraph>
</subSubSection>
<caption id="D0D1664FFF9C8372FF32F81CFB9EE9E4" ID-DOI="http://doi.org/10.5281/zenodo.3743152" ID-Zenodo-Dep="3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="8" pageNumber="145" startId="8.[132,143,1987,2005]" targetBox="[151,1438,231,1842]" targetPageId="8">
<paragraph id="841136C7FF9C8372FF32F81CFB9EE9E4" blockId="8.[132,1455,1987,2027]" pageId="8" pageNumber="145">
FIG. 3. —
<taxonomicName id="43AE4D44FF9C8372FF60F81CFEE3E9DA" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[214,389,1987,2005]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="145" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9C8372FF60F81CFEE3E9DA" box="[214,389,1987,2005]" italics="true" pageId="8" pageNumber="145">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9C8372FE3DF81CFD65E9DA" box="[395,515,1987,2005]" pageId="8" pageNumber="145" rank="species">n. gen., n. sp.</taxonomicNameLabel>
:
<emphasis id="B6DAEAD5FF9C8372FDBDF81CFD7EE9DA" bold="true" box="[523,536,1987,2005]" pageId="8" pageNumber="145">A</emphasis>
, wing;
<emphasis id="B6DAEAD5FF9C8372FDE3F81CFD04E9DA" bold="true" box="[597,610,1987,2005]" pageId="8" pageNumber="145">B</emphasis>
, male head, frontal view;
<emphasis id="B6DAEAD5FF9C8372FC8BF81CFC2DE9DA" bold="true" box="[829,843,1987,2005]" pageId="8" pageNumber="145">C</emphasis>
, male epandrium, proctiger and epandrial claspers, dorsal view;
<emphasis id="B6DAEAD5FF9C8372FAD8F81CFA1AE9DA" bold="true" box="[1390,1404,1987,2005]" pageId="8" pageNumber="145">D</emphasis>
, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm.
</paragraph>
</caption>
<caption id="D0D1664FFF9D8373FF32F81CFA2CE9E4" ID-DOI="http://doi.org/10.5281/zenodo.3743154" ID-Zenodo-Dep="3743154" httpUri="https://zenodo.org/record/3743154/files/figure.png" pageId="9" pageNumber="146" startId="9.[132,143,1987,2005]" targetBox="[302,1290,256,1858]" targetPageId="9">
<paragraph id="841136C7FF9D8373FF32F81CFA2CE9E4" blockId="9.[132,1455,1987,2028]" pageId="9" pageNumber="146">
FIG. 4. —
<taxonomicName id="43AE4D44FF9D8373FF64F81CFE19E9DA" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[210,383,1987,2005]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="146" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9D8373FF64F81CFE19E9DA" box="[210,383,1987,2005]" italics="true" pageId="9" pageNumber="146">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9D8373FE35F81CFE90E9DA" box="[387,502,1987,2005]" pageId="9" pageNumber="146" rank="species">n. gen., n. sp.</taxonomicNameLabel>
:
<emphasis id="B6DAEAD5FF9D8373FE4DF81CFD6EE9DA" bold="true" box="[507,520,1987,2005]" pageId="9" pageNumber="146">A</emphasis>
, male terminalia,lateral view;
<emphasis id="B6DAEAD5FF9D8373FD48F81CFC6DE9DA" bold="true" box="[766,779,1987,2005]" pageId="9" pageNumber="146">B</emphasis>
, female mouthparts and maxillary palpi,frontal view;
<emphasis id="B6DAEAD5FF9D8373FB09F81CFBABE9DA" bold="true" box="[1215,1229,1987,2005]" pageId="9" pageNumber="146">C</emphasis>
, female terminalia,external structure, ventral view;
<emphasis id="B6DAEAD5FF9D8373FEF0F805FE32E9E3" bold="true" box="[326,340,2010,2028]" pageId="9" pageNumber="146">D</emphasis>
, female terminalia, internal structure and egg, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="D0D1664FFF9E8370FF79FBA9FA02EA87" ID-DOI="http://doi.org/10.5281/zenodo.3743156" ID-Zenodo-Dep="3743156" box="[207,1380,1142,1161]" httpUri="https://zenodo.org/record/3743156/files/figure.png" pageId="10" pageNumber="147" startId="10.[207,218,1142,1160]" targetBox="[132,1456,215,1098]" targetPageId="10">
<paragraph id="841136C7FF9E8370FF79FBA9FA02EA87" blockId="10.[207,1380,1142,1161]" box="[207,1380,1142,1161]" pageId="10" pageNumber="147">
FIG. 5. — Photomicrographs of male wings:
<emphasis id="B6DAEAD5FF9E8370FD8AFBA8FD2FEA86" bold="true" box="[572,585,1143,1161]" pageId="10" pageNumber="147">A</emphasis>
,
<taxonomicName id="43AE4D44FF9E8370FDE2FBA9FD99EA87" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[596,767,1142,1160]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9E8370FDE2FBA9FD99EA87" box="[596,767,1142,1160]" italics="true" pageId="10" pageNumber="147">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FCB3FBA9FC1CEA87" box="[773,890,1142,1160]" pageId="10" pageNumber="147" rank="species">n. gen., n. sp.</taxonomicNameLabel>
;
<emphasis id="B6DAEAD5FF9E8370FC37FBA8FCE8EA86" bold="true" box="[897,910,1143,1161]" pageId="10" pageNumber="147">B</emphasis>
,
<taxonomicName id="43AE4D44FF9E8370FC2EFBA9FB20EA87" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[920,1094,1142,1160]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9E8370FC2EFBA9FB20EA87" box="[920,1094,1142,1160]" italics="true" pageId="10" pageNumber="147">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FBFDFBA9FBD9EA87" box="[1099,1215,1142,1160]" pageId="10" pageNumber="147" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Scale bars: 0.5 mm
</paragraph>
</caption>
<subSubSection id="CCB4654CFF9E8370FF32FB0AFE85E981" pageId="10" pageNumber="147" type="discussion">
<paragraph id="841136C7FF9E8370FF32FB0AFF8EEAE1" blockId="10.[132,775,1237,1934]" box="[132,232,1237,1263]" pageId="10" pageNumber="147">REMARKS</paragraph>
<paragraph id="841136C7FF9E8370FF32FB29FEC5E881" blockId="10.[132,775,1237,1934]" pageId="10" pageNumber="147">
In contrast to
<taxonomicName id="43AE4D44FF9E8370FE9EFB2AFEC5EB00" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[296,419,1269,1295]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FE9EFB2AFEC5EB00" box="[296,419,1269,1295]" italics="true" pageId="10" pageNumber="147">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FE1AFB29FE9BEB1F" box="[428,509,1270,1296]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
, the relatively large size and light coloration of
<taxonomicName id="43AE4D44FF9E8370FE37FAC9FE99EB20" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[385,511,1302,1327]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FE37FAC9FE99EB20" box="[385,511,1302,1327]" italics="true" pageId="10" pageNumber="147">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FDB0FAC9FD37EB3F" box="[518,593,1302,1328]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
make it possible to easily differentiate them in samples containing many psychodid genera. Moreover, while both genera have lanceolate wings, the latter is more distinctive due to several veins being darkly pigmented (
<figureCitation id="1C952A42FF9E8370FE6FFA4AFD71EBBF" box="[473,535,1429,1456]" captionStart="FIG" captionStartId="10.[207,218,1142,1160]" captionTargetBox="[132,1456,215,1098]" captionTargetId="figure@10.[127,1461,209,1106]" captionTargetPageId="10" captionText="FIG. 5. — Photomicrographs of male wings: A, Polletomyia subulata n. gen., n. sp.; B, Myiomystax trilineata n. gen., n. sp. Scale bars: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3743156" httpUri="https://zenodo.org/record/3743156/files/figure.png" pageId="10" pageNumber="147">Fig. 5</figureCitation>
). The unique wing of
<taxonomicName id="43AE4D44FF9E8370FF32FA69FE64EBC0" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[132,258,1462,1487]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FF32FA69FE64EBC0" box="[132,258,1462,1487]" italics="true" pageId="10" pageNumber="147">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FEBEFA6AFE35EBC0" box="[264,339,1461,1487]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
also makes it easy to associate males and females of this genus. It remains to be determined whether these characteristics of the wing will be present in the wings of congeners, if any are discovered. Although less important compared to those of
<taxonomicName id="43AE4D44FF9E8370FEDBF9EBFE81E841" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[365,487,1588,1614]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FEDBF9EBFE81E841" box="[365,487,1588,1614]" italics="true" pageId="10" pageNumber="147">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FE5AF9EAFD5FE840" box="[492,569,1589,1615]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
, accurate diagnosis of this genus is still dependent on examination of slidemounted male terminalia.
</paragraph>
<paragraph id="841136C7FF9E8370FF2DF94AFE85E981" blockId="10.[132,775,1237,1934]" pageId="10" pageNumber="147">
<taxonomicName id="43AE4D44FF9E8370FF2DF94AFE7EE8A1" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[155,280,1685,1710]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FF2DF94AFE7EE8A1" box="[155,280,1685,1710]" italics="true" pageId="10" pageNumber="147">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FE97F94AFE09E8A0" box="[289,367,1685,1711]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
males have a bilaterally symmetrical terminalia, with an aedeagus that extends as an open-close mechanism, much like
<taxonomicName id="43AE4D44FF9E8370FE38F90BFD6CE8E1" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[398,522,1748,1774]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FE38F90BFD6CE8E1" box="[398,522,1748,1774]" italics="true" pageId="10" pageNumber="147">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FDA5F90BFD05E8E1" box="[531,611,1748,1774]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="43AE4D44FF9E8370FD28F90AFC66E8E1" authorityName="Muller" authorityYear="1895" box="[670,768,1749,1774]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF9E8370FD28F90AFC66E8E1" box="[670,768,1749,1774]" italics="true" pageId="10" pageNumber="147">Maruina</emphasis>
</taxonomicName>
. However, they differ from the other genera by having paired aedeagal sclerites as opposed to two pairs (as in
<taxonomicName id="43AE4D44FF9E8370FD39F8CBFC60E921" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[655,774,1812,1838]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FD39F8CBFC60E921" box="[655,774,1812,1838]" italics="true" pageId="10" pageNumber="147">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FF32F8EBFFB4E941" box="[132,210,1844,1870]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
) or multiple aedeagal spines (as in
<taxonomicName id="43AE4D44FF9E8370FDEDF8EBFDDCE942" authorityName="Muller" authorityYear="1895" box="[603,698,1844,1869]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF9E8370FDEDF8EBFDDCE942" box="[603,698,1844,1869]" italics="true" pageId="10" pageNumber="147">Maruina</emphasis>
</taxonomicName>
).
<taxonomicName id="43AE4D44FF9E8370FD62F8EBFFB0E962" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9E8370FD62F8EBFFB0E962" italics="true" pageId="10" pageNumber="147">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FF6AF88BFE40E961" box="[220,294,1876,1902]" pageId="10" pageNumber="147" rank="genus">n. gen.</taxonomicNameLabel>
males are also unique among these genera in that their gonocoxites lack setae.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9E8370FF32F86CFD11E9E1" pageId="10" pageNumber="147" type="description">
<paragraph id="841136C7FF9E8370FF32F86CFE73E9C4" blockId="10.[132,631,1971,2030]" box="[132,277,1971,1997]" pageId="10" pageNumber="147">DESCRIPTION</paragraph>
<paragraph id="841136C7FF9E8370FF32F80CFD11E9E1" blockId="10.[132,631,1971,2030]" box="[132,631,2003,2030]" pageId="10" pageNumber="147">
See below,
<taxonomicName id="43AE4D44FF9E8370FF4CF80BFEBBE9E2" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[250,477,2003,2029]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9E8370FF4CF80BFEBBE9E2" box="[250,477,2003,2029]" italics="true" pageId="10" pageNumber="147">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FE52F80BFD11E9E1" box="[484,631,2004,2030]" pageId="10" pageNumber="147" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,358 @@
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<mods:title id="B55193C1DCD3184C63BB1093BF402938">Descriptions of two new genera of Maruinini (Diptera, Psychodidae, Psychodinae) from the Mitaraka range of French Guiana</mods:title>
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<mods:namePart id="D689970813AA9E51E543A8C1597B7668">Curler, Gregory</mods:namePart>
<mods:affiliation id="B97B385436CED480AB1DD3C1D782D220">Mississippi Entomological Museum, Mississippi State University, Old Highway 12, P. O. Drawer 9775, Mississippi State, MS 39762 - 9775 (USA) Collaborator, Smithsonian Institution, National Museum of Natural History; 10 &amp; Constitution NW, Washington, DC 20560 - 0169 (USA) gcurler @ gmail. com</mods:affiliation>
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<mods:title id="2F9C3BD73CAD99D8B432499CEBC02DE2">Zoosystema</mods:title>
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<mods:part id="2509667C3B459E3AB43D1BEDE364551C">
<mods:date id="0934C3F7CD5FF61A91270BAF7533005B">2020</mods:date>
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<mods:number id="A6106C3DFAC0DBEEE9FEB7AF1BF0B89F">2020-04-07</mods:number>
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<treatment id="0C0787D1FF9E8371FC1FFB0AFBE3ED3E" ID-DOI="http://doi.org/10.5281/zenodo.3809876" ID-GBIF-Taxon="163536910" ID-Zenodo-Dep="3809876" LSID="urn:lsid:zoobank.org:act:4BF6B1A7-8DED-4AFA-9184-A51145B25143" httpUri="http://treatment.plazi.org/id/0C0787D1FF9E8371FC1FFB0AFBE3ED3E" lastPageId="11" lastPageNumber="148" pageId="10" pageNumber="147">
<subSubSection id="CCB4654CFF9E8370FC1FFB0AFA55EAFF" box="[937,1331,1237,1264]" pageId="10" pageNumber="147" type="nomenclature">
<paragraph id="841136C7FF9E8370FC1FFB0AFA55EAFF" blockId="10.[937,1331,1237,1296]" box="[937,1331,1237,1264]" pageId="10" pageNumber="147">
<heading id="DF5981ABFF9E8370FC1FFB0AFA55EAFF" box="[937,1331,1237,1264]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="147" reason="2">
<taxonomicName id="43AE4D44FF9E8370FC1FFB0AFBFFEAFF" ID-CoL="8H4CK" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[937,1177,1237,1264]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="species" species="trilineata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF9E8370FC1FFB0AFBFFEAFF" bold="true" box="[937,1177,1237,1264]" italics="true" pageId="10" pageNumber="147">Myiomystax trilineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9E8370FB16FB09FA55EAFF" box="[1184,1331,1238,1264]" pageId="10" pageNumber="147" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9E8370FB97FB2AFAF5EB4B" pageId="10" pageNumber="147" type="description">
<paragraph id="841136C7FF9E8370FB97FB2AFBDDEB1F" blockId="10.[937,1331,1237,1296]" box="[1057,1211,1269,1296]" pageId="10" pageNumber="147">
(
<figureCitation id="1C952A42FF9E8370FB9EFB2AFB0FEB1F" box="[1064,1129,1269,1296]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="10" pageNumber="147">Figs 3</figureCitation>
;
<figureCitation id="1C952A42FF9E8370FBC1FB29FBE3EB1F" box="[1143,1157,1270,1296]" captionStart="FIG" captionStartId="9.[132,143,1987,2005]" captionTargetBox="[302,1290,256,1858]" captionTargetId="figure@9.[260,1328,215,1893]" captionTargetPageId="9" captionText="FIG. 4. — Myiomystax trilineata n. gen., n. sp.:A, male terminalia,lateral view; B, female mouthparts and maxillary palpi,frontal view;C, female terminalia,external structure, ventral view; D, female terminalia, internal structure and egg, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743154" httpUri="https://zenodo.org/record/3743154/files/figure.png" pageId="10" pageNumber="147">4</figureCitation>
;
<figureCitation id="1C952A42FF9E8370FB25FB2AFBD4EB1F" box="[1171,1202,1269,1296]" captionStart="FIG" captionStartId="10.[207,218,1142,1160]" captionTargetBox="[132,1456,215,1098]" captionTargetId="figure@10.[127,1461,209,1106]" captionTargetPageId="10" captionText="FIG. 5. — Photomicrographs of male wings: A, Polletomyia subulata n. gen., n. sp.; B, Myiomystax trilineata n. gen., n. sp. Scale bars: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3743156" httpUri="https://zenodo.org/record/3743156/files/figure.png" pageId="10" pageNumber="147">5B</figureCitation>
)
</paragraph>
<paragraph id="841136C7FF9E8370FCFEFAEDFAF5EB4B" blockId="10.[840,1427,1330,1348]" box="[840,1427,1330,1348]" pageId="10" pageNumber="147">
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</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9E8370FC9BFAB2FBC2EB89" box="[813,1188,1389,1414]" pageId="10" pageNumber="147" type="distribution">
<paragraph id="841136C7FF9E8370FC9BFAB2FBC2EB89" blockId="10.[813,1188,1389,1414]" box="[813,1188,1389,1414]" pageId="10" pageNumber="147">
<typeStatus id="5B158865FF9E8370FC9BFAB2FC3BEB8C" box="[813,861,1389,1413]" pageId="10" pageNumber="147">TYPE</typeStatus>
LOCALITY. —
<collectingCountry id="FCB97657FF9E8370FC5DFAB1FB5CEB89" box="[1003,1082,1390,1414]" name="French Guiana" pageId="10" pageNumber="147">Guyane</collectingCountry>
, Mitaraka.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9E8370FC9BFA79FBC5E9E5" pageId="10" pageNumber="147" type="materials_examined">
<paragraph id="841136C7FF9E8370FC9BFA79FBC5E9E5" blockId="10.[813,1456,1446,2026]" pageId="10" pageNumber="147">
TYPE MATERIAL. —
<materialsCitation id="34C63C9AFF9E8370FC58FA79FAD6E802" ID-GBIF-Occurrence="2595987905" collectingDate="2015-02-27" collectingDateMax="2015-03-06" collectingDateMin="2015-02-27" collectionCode="YPT, MNHN" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" location="Mitaraka" longLatPrecision="1" longitude="-54.448055" pageId="10" pageNumber="147" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B6DAEAD5FF9E8370FC58FA79FB2BEBB1" bold="true" box="[1006,1101,1446,1470]" pageId="10" pageNumber="147">
<typeStatus id="5B158865FF9E8370FC58FA79FB2BEBB1" box="[1006,1101,1446,1470]" pageId="10" pageNumber="147" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B6DAEAD5FF9E8370FBECFA79FB88EBB1" bold="true" box="[1114,1262,1446,1470]" pageId="10" pageNumber="147">
<collectingCountry id="FCB97657FF9E8370FBECFA79FB88EBB1" box="[1114,1262,1446,1470]" name="French Guiana" pageId="10" pageNumber="147">French Guiana</collectingCountry>
</emphasis>
<specimenCount id="92A8FD4EFF9E8370FAB0FA76FA7CEBCF" box="[1286,1306,1449,1472]" pageId="10" pageNumber="147" type="male"></specimenCount>
;
<location id="8171601CFF9E8370FA9FFA79FAE2EBB1" LSID="urn:lsid:plazi:treatment:0C0787D1FF9E8371FC1FFB0AFBE3ED3E:8171601CFF9E8370FA9FFA79FAE2EBB1" box="[1321,1412,1446,1470]" country="French Guiana" latitude="2.2342777" longLatPrecision="1" longitude="-54.448055" name="Mitaraka" pageId="10" pageNumber="147">Mitaraka</location>
, site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF9E8370FC74FA1EFB29EBD6" box="[962,1103,1473,1497]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="147" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF9E8370FBEEFA1EFB8EEBD6" box="[1112,1256,1473,1497]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="10" pageNumber="147" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<elevation id="0F83D1F4FF9E8370FB44FA1EFA31EBD6" box="[1266,1367,1473,1497]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">
<quantity id="43569B22FF9E8370FB44FA1EFA43EBD7" box="[1266,1317,1473,1497]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF9E8370FC2EFA04FB5DEBFC" box="[920,1083,1499,1523]" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-06" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF9E8370FC2EFA04FB5DEBFC" box="[920,1083,1499,1523]" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-06" valueMin="2015-02-27">27.II-6.III.2015</collectingDate>
</date>
;
<collectionCode id="E2BFAE02FF9E8370FBF1FA03FB13EBFC" box="[1095,1141,1500,1523]" pageId="10" pageNumber="147">YPT</collectionCode>
;
<collectorName id="295B5311FF9E8370FB37FA03FB92EBFC" box="[1153,1268,1499,1523]" pageId="10" pageNumber="147">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/125); specimen dissected, mounted on slide;
<collectionCode id="E2BFAE02FF9E8370FAE1FA29FACAE802" box="[1367,1452,1526,1549]" pageId="10" pageNumber="147">MNHN</collectionCode>
.
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FC9BF9CFFAC9E852" ID-GBIF-Occurrence="2595987913" collectingDate="2015-02-27" collectingDateMax="2015-03-06" collectingDateMin="2015-02-27" collectionCode="SW" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" location="French Guiana" longLatPrecision="1" longitude="-54.448055" pageId="10" pageNumber="147" specimenCode="LACM 340595" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B6DAEAD5FF9E8370FC9BF9CFFB41E827" bold="true" box="[813,1063,1552,1576]" pageId="10" pageNumber="147">
<typeStatus id="5B158865FF9E8370FC9BF9CFFCE8E827" box="[813,910,1552,1576]" pageId="10" pageNumber="147" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="FCB97657FF9E8370FC20F9CFFB41E827" box="[918,1063,1552,1576]" name="French Guiana" pageId="10" pageNumber="147">French Guiana</collectingCountry>
</emphasis>
<specimenCount id="92A8FD4EFF9E8370FB8FF9CFFB3BE825" box="[1081,1117,1552,1578]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
; site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF9E8370FAA9F9CFFACAE827" box="[1311,1452,1552,1576]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="147" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF9E8370FC9BF9F4FCA7E84C" box="[813,961,1579,1603]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="10" pageNumber="147" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<elevation id="0F83D1F4FF9E8370FC7DF9F4FB52E84C" box="[971,1076,1579,1603]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">
<quantity id="43569B22FF9E8370FC7DF9F4FB66E84D" box="[971,1024,1579,1603]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF9E8370FAB7F9F4FA16E84C" box="[1281,1392,1579,1603]" pageId="10" pageNumber="147" value="2015-02-27">
<collectingDate id="E054E9EFFF9E8370FAB7F9F4FA16E84C" box="[1281,1392,1579,1603]" pageId="10" pageNumber="147" value="2015-02-27">27.II.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF9E8370FACCF9F4FC00E852" pageId="10" pageNumber="147">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/018);
<collectionCode id="E2BFAE02FF9E8370FB52F99AFA6DE852" box="[1252,1291,1605,1629]" pageId="10" pageNumber="147">SW</collectionCode>
;
<specimenCode id="D4089EBCFF9E8370FAA2F99AFAC9E852" box="[1300,1455,1605,1629]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340595</specimenCode>
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FC89F9BFFAFBE89D" ID-GBIF-Occurrence="2595987915" collectionCode="YPT, LACM" country="French Guiana" location="French Guiana" pageId="10" pageNumber="147" specimenCode="LACM 340603-340604, LACM 340605" specimenCount="12" specimenCount-female="4" specimenCount-male="8" typeStatus="paratype">
<specimenCount id="92A8FD4EFF9E8370FC89F9BFFC0FE876" box="[831,873,1632,1657]" pageId="10" pageNumber="147" type="male">4 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FCCEF9BFFCF9E876" box="[888,927,1632,1657]" pageId="10" pageNumber="147" type="female">2 ♀</specimenCount>
; same data as holotype;
<collectionCode id="E2BFAE02FF9E8370FB28F9BFFBA8E878" box="[1182,1230,1632,1655]" pageId="10" pageNumber="147">YPT</collectionCode>
; MNHN (
<specimenCount id="92A8FD4EFF9E8370FA89F9BFFA0CE876" box="[1343,1386,1632,1657]" pageId="10" pageNumber="147" type="male">2 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FACFF9BFFAC6E876" box="[1401,1440,1632,1657]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
),
<collectionCode id="E2BFAE02FF9E8370FC9BF9A5FC11E89D" box="[813,887,1658,1682]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF9E8370FC31F9A5FCD4E89B" box="[903,946,1658,1684]" pageId="10" pageNumber="147" type="male">2 ♂</specimenCount>
<specimenCode id="D4089EBCFF9E8370FC0CF9A5FBD0E89D" box="[954,1206,1658,1684]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340603-340604</specimenCode>
,
<specimenCount id="92A8FD4EFF9E8370FB74F9A5FB8CE89B" box="[1218,1258,1658,1684]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
<specimenCode id="D4089EBCFF9E8370FB44F9A5FAF0E89D" box="[1266,1430,1658,1682]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340605</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FC9BF94AFB04E8F3" ID-GBIF-Occurrence="2595987911" collectingDate="2015-02-27" collectingDateMax="2015-03-08" collectingDateMin="2015-02-27" collectionCode="YPT, LACM" collectorName="Marc Pollet" country="French Guiana" elevation="25" latitude="2.2363334" longLatPrecision="1" longitude="-54.447083" pageId="10" pageNumber="147" specimenCode="LACM 340606" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="92A8FD4EFF9E8370FC9BF94AFC3EE8A1" box="[813,856,1685,1710]" pageId="10" pageNumber="147" type="male">1 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FCD1F94AFCE9E8A1" box="[871,911,1685,1710]" pageId="10" pageNumber="147" type="female">2 ♀</specimenCount>
; site MIT-C-RBF1;
<geoCoordinate id="E19A5000FF9E8370FBDBF94AFA63E8A2" box="[1133,1285,1685,1709]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="147" precision="1" seconds="10.8" value="2.2363334">02°1410.8”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF9E8370FAA7F94BFACDE8A3" box="[1297,1451,1684,1708]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="10" pageNumber="147" precision="1" seconds="49.5" value="-54.447083">54°2649.5”W</geoCoordinate>
;
<elevation id="0F83D1F4FF9E8370FC9BF970FCF9E8C8" box="[813,927,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="147" unit="m" value="25.0">
<quantity id="43569B22FF9E8370FC9BF970FC03E8C8" box="[813,869,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="147" unit="m" value="25.0">25 m</quantity>
a.s.l.
</elevation>
; lowland rainforest;
<date id="F0101007FF9E8370FBCDF970FA41E8C8" box="[1147,1319,1711,1737]" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF9E8370FBCDF970FA41E8C8" box="[1147,1319,1711,1737]" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-08" valueMin="2015-02-27">27.II-8.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF9E8370FA82F96FFAC9E8C8" box="[1332,1455,1711,1735]" pageId="10" pageNumber="147">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/133);
<collectionCode id="E2BFAE02FF9E8370FB03F915FB82E8EE" box="[1205,1252,1738,1761]" pageId="10" pageNumber="147">YPT</collectionCode>
; MNHN (
<specimenCount id="92A8FD4EFF9E8370FAE0F913FA0DE8EC" box="[1366,1387,1740,1763]" pageId="10" pageNumber="147" type="male"></specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FACFF915FAC6E8EC" box="[1401,1440,1738,1763]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
),
<collectionCode id="E2BFAE02FF9E8370FC9BF93BFC11E8F3" box="[813,887,1764,1788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF9E8370FC3FF93BFCC9E8F2" box="[905,943,1764,1789]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
<specimenCode id="D4089EBCFF9E8370FC0EF93BFB3DE8F3" box="[952,1115,1764,1788]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340606</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FBCBF93BFB47E969" ID-GBIF-Occurrence="2595987912" collectingDate="2015-03-06" collectingDateMax="2015-03-10" collectingDateMin="2015-03-06" collectionCode="YPT, MNHN, LACM" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" longLatPrecision="1" longitude="-54.448055" pageId="10" pageNumber="147" specimenCode="LACM 340607-340611, LACM 340612-340613" specimenCount="28" specimenCount-female="10" specimenCount-male="18" typeStatus="paratype">
<specimenCount id="92A8FD4EFF9E8370FBCBF93BFBCEE8F2" box="[1149,1192,1764,1789]" pageId="10" pageNumber="147" type="male">9 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FB01F93BFBB8E8F2" box="[1207,1246,1764,1789]" pageId="10" pageNumber="147" type="female">5 ♀</specimenCount>
; site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF9E8370FC9BF920FCDFE918" box="[813,953,1791,1815]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="147" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF9E8370FC77F921FB36E919" box="[961,1104,1790,1814]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="10" pageNumber="147" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<quantity id="43569B22FF9E8370FBEFF920FBEDE919" box="[1113,1163,1791,1815]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">
<elevation id="0F83D1F4FF9E8370FBEFF920FBEDE919" box="[1113,1163,1791,1815]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">29 m</elevation>
</quantity>
a.s.l.; lowland rainforest;
<date id="F0101007FF9E8370FACAF921FCE2E93E" pageId="10" pageNumber="147" value="2015-03-06" valueMax="2015-03-10" valueMin="2015-03-06">
<collectingDate id="E054E9EFFF9E8370FACAF921FCE2E93E" pageId="10" pageNumber="147" value="2015-03-06" valueMax="2015-03-10" valueMin="2015-03-06">6-10. III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF9E8370FC3BF8C6FC98E93E" box="[909,1022,1817,1841]" pageId="10" pageNumber="147">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/136);
<collectionCode id="E2BFAE02FF9E8370FACAF8C6FACDE93F" box="[1404,1451,1817,1840]" pageId="10" pageNumber="147">YPT</collectionCode>
;
<collectionCode id="E2BFAE02FF9E8370FC9BF8EBFCE5E944" box="[813,899,1844,1867]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="10" pageNumber="147">MNHN</collectionCode>
(
<specimenCount id="92A8FD4EFF9E8370FC27F8ECFCDCE942" box="[913,954,1843,1869]" pageId="10" pageNumber="147" type="male">4 ♂</specimenCount>
,
<specimenCount id="92A8FD4EFF9E8370FC71F8ECFC8BE942" box="[967,1005,1843,1869]" pageId="10" pageNumber="147" type="female">3 ♀</specimenCount>
),
<collectionCode id="E2BFAE02FF9E8370FBB4F8ECFB2DE944" box="[1026,1099,1843,1867]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM</collectionCode>
(
<specimenCount id="92A8FD4EFF9E8370FBECF8ECFBE5E942" box="[1114,1155,1843,1869]" pageId="10" pageNumber="147" type="male">5 ♂</specimenCount>
<specimenCode id="D4089EBCFF9E8370FB3FF8ECFA19E944" box="[1161,1407,1843,1869]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340607-340611</specimenCode>
,
<specimenCount id="92A8FD4EFF9E8370FA3FF8ECFAC9E942" box="[1417,1455,1843,1869]" pageId="10" pageNumber="147" type="female">2 ♀</specimenCount>
<specimenCode id="D4089EBCFF9E8370FC9BF891FB7CE969" box="[813,1050,1870,1895]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="147" type="Museum">LACM 340612-340613</specimenCode>
)
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FB82F891FCE5E9BA" ID-GBIF-Occurrence="2595987902" collectingDate="2015-02-27" collectingDateMax="2015-03-02" collectingDateMin="2015-02-27" collectionCode="SLAM" collectorName="Marc Pollet" country="French Guiana" elevation="29" latitude="2.2342777" longLatPrecision="1" longitude="-54.448055" pageId="10" pageNumber="147" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="92A8FD4EFF9E8370FB82F891FB31E968" box="[1076,1111,1870,1895]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
; site MIT-C-RBF2;
<geoCoordinate id="E19A5000FF9E8370FAABF891FACDE969" box="[1309,1451,1870,1894]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="147" precision="1" seconds="03.4" value="2.2342777">02°1403.4”N</geoCoordinate>
,
<geoCoordinate id="E19A5000FF9E8370FC9BF8B7FCA7E98F" box="[813,961,1896,1920]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="10" pageNumber="147" precision="1" seconds="53.0" value="-54.448055">54°2653.0”W</geoCoordinate>
;
<elevation id="0F83D1F4FF9E8370FC7CF8B7FB57E98F" box="[970,1073,1896,1920]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">
<quantity id="43569B22FF9E8370FC7CF8B7FC98E98F" box="[970,1022,1896,1920]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="147" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
; tropical wet forest (bas fond);
<date id="F0101007FF9E8370FAD4F8B7FCE7E994" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-02" valueMin="2015-02-27">
<collectingDate id="E054E9EFFF9E8370FAD4F8B7FCE7E994" pageId="10" pageNumber="147" value="2015-02-27" valueMax="2015-03-02" valueMin="2015-02-27">27.II-2. III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF9E8370FC3FF85CFC90E994" box="[905,1014,1923,1947]" pageId="10" pageNumber="147">Marc Pollet</collectorName>
leg. (sample code: MITARAKA/184);
<collectionCode id="E2BFAE02FF9E8370FADEF85CFACAE994" box="[1384,1452,1923,1947]" pageId="10" pageNumber="147">SLAM</collectionCode>
; MNHN
</materialsCitation>
<materialsCitation id="34C63C9AFF9E8370FC20F842FBF9E9E5" ID-GBIF-Occurrence="2595987904" collectingDate="2015-03-14" collectionCode="SLAM" collectorName="Julien Touroult &amp; Eddy Poirier" country="French Guiana" pageId="10" pageNumber="147" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="92A8FD4EFF9E8370FC20F842FCDCE9B8" box="[918,954,1949,1975]" pageId="10" pageNumber="147" type="female">1 ♀</specimenCount>
; site different sites near base camp and along trails;
<date id="F0101007FF9E8370FC9BF868FCC3E9C0" box="[813,933,1975,1999]" pageId="10" pageNumber="147" value="2015-03-14">
<collectingDate id="E054E9EFFF9E8370FC9BF868FCC3E9C0" box="[813,933,1975,1999]" pageId="10" pageNumber="147" value="2015-03-14">14.III.2015</collectingDate>
</date>
;
<collectorName id="295B5311FF9E8370FC07F867FB2DE9DF" box="[945,1099,1976,2000]" pageId="10" pageNumber="147">Julien Touroult</collectorName>
&amp;
<collectorName id="295B5311FF9E8370FBC7F867FB97E9DF" box="[1137,1265,1976,2000]" pageId="10" pageNumber="147">Eddy Poirier</collectorName>
leg. (sample code: MITARAKA/191);
<collectionCode id="E2BFAE02FF9E8370FC43F80DFB5DE9E5" box="[1013,1083,2002,2026]" pageId="10" pageNumber="147">SLAM</collectionCode>
; MNHN
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9F8371FF32FF07FEAAEF29" pageId="11" pageNumber="148" type="etymology">
<paragraph id="841136C7FF9F8371FF32FF07FEAAEF29" blockId="11.[132,775,216,294]" pageId="11" pageNumber="148">ETYMOLOGY. — The specific epithet is derived form the Latin word “linea”, meaning line, in reference to the three darkly pigmented wing veins present in this species.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9F8371FF32FEE2FD26EF59" box="[132,576,317,342]" pageId="11" pageNumber="148" type="diagnosis">
<paragraph id="841136C7FF9F8371FF32FEE2FD26EF59" blockId="11.[132,576,317,342]" box="[132,576,317,342]" pageId="11" pageNumber="148">
DIAGNOSIS. — See above,
<taxonomicName id="43AE4D44FF9F8371FE30FEE1FE90EF5A" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[390,502,318,341]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="148" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF9F8371FE30FEE1FE90EF5A" box="[390,502,318,341]" italics="true" pageId="11" pageNumber="148">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF9F8371FE4AFEE1FD26EF59" box="[508,576,318,342]" pageId="11" pageNumber="148" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9F8371FF32FEB2FEB4EF90" pageId="11" pageNumber="148" type="distribution">
<paragraph id="841136C7FF9F8371FF32FEB2FEB4EF90" blockId="11.[132,775,365,416]" pageId="11" pageNumber="148">
DISTRIBUTION. — Currently known from multiple localities in the Mitaraka range of
<collectingCountry id="FCB97657FF9F8371FE8FFE58FEABEF90" box="[313,461,391,415]" name="French Guiana" pageId="11" pageNumber="148">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9F8371FF32FE68FD9AEFE6" pageId="11" pageNumber="148" type="biology_ecology">
<paragraph id="841136C7FF9F8371FF32FE68FD9AEFE6" blockId="11.[132,775,439,489]" pageId="11" pageNumber="148">HABITAT. — Little information regarding specific habitat is available; however, all specimens were collected in lowland rainforest.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF9F8371FF32FDC8FBE3ED3E" pageId="11" pageNumber="148" type="description">
<paragraph id="841136C7FF9F8371FF32FDC8FE73EC3F" blockId="11.[132,776,535,2030]" box="[132,277,535,561]" pageId="11" pageNumber="148">DESCRIPTION</paragraph>
<paragraph id="841136C7FF9F8371FF32FDE7FFDFEC5D" blockId="11.[132,776,535,2030]" box="[132,185,568,594]" pageId="11" pageNumber="148">
<emphasis id="B6DAEAD5FF9F8371FF32FDE7FFDFEC5D" box="[132,185,568,594]" italics="true" pageId="11" pageNumber="148">Male</emphasis>
</paragraph>
<paragraph id="841136C7FF9F8371FF32FD87FAD7EF5C" blockId="11.[132,776,535,2030]" lastBlockId="11.[813,1457,217,339]" pageId="11" pageNumber="148">
Measurements, (N = 4) head width
<quantity id="43569B22FF9F8371FE58FD87FD2AEC7D" box="[494,588,600,626]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="148" unit="mm" value="0.41">0.41 mm</quantity>
(0.38-0.44), head height
<quantity id="43569B22FF9F8371FF67FDA7FE51EC9D" box="[209,311,632,658]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="11" pageNumber="148" unit="mm" value="0.47">0.47 mm</quantity>
(0.44-0.48), wing length
<quantity id="43569B22FF9F8371FDE0FDA7FDDAEC9D" box="[598,700,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="11" pageNumber="148" unit="mm" value="2.17">2.17 mm</quantity>
(2.08- 2.24), wing width
<quantity id="43569B22FF9F8371FEE4FD48FED0ECBD" box="[338,438,663,690]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="11" pageNumber="148" unit="mm" value="0.54">0.54 mm</quantity>
(0.52-0.56). Head with vertex extended slightly posteriorly, eyebridge comprised of 3 facet rows, contiguous but strongly constricted at median. Frons and clypeus distinctly divided; frontal scar patch bilobed posteriorly, contiguous medially; clypeus elongated laterally, with lateral margins overlapping medial margins of eyes. Antennae: scape cylindrical, about 2.5 times longer than wide; pedicel globular; flagellomeres 1-5 elongate, cylindrical; flagellomeres 6-11 fusiform; flagellomeres 12-14 reduced, bead-like; flagellomere 14 with apical process digitiform, setose, approximately as long as node; all flagellomeres lacking ascoids. Mouthparts extending slightly beyond base of palp segment 1; labellum about as wide as anterior margin of clypeus; palpi conspicuously elongate, with segment 1 hook-like; segment 2 clavate, inflating from base to apex; segment 3 elongate, inflated at midlength; segment 4 inflated apically, capitate; palp segments with clusters of spatulate setae inserted dorsally and ventrally; palps folded behind head at rest. Wing: strongly lanceolate; radial fork positioned slightly distal to apex of CuA; medial fork positioned at the level of apex of CuA; R
<subScript id="182A3482FF9F8371FE76FADDFEACEB1C" attach="left" box="[448,458,1282,1299]" fontSize="7" pageId="11" pageNumber="148">5</subScript>
terminating in C posterior to wing apex; wing veins R
<subScript id="182A3482FF9F8371FEC3FAFDFE18EB3C" attach="left" box="[373,382,1314,1331]" fontSize="7" pageId="11" pageNumber="148">1</subScript>
, R
<subScript id="182A3482FF9F8371FE2DFAFDFEC2EB3C" attach="left" box="[411,420,1314,1331]" fontSize="7" pageId="11" pageNumber="148">5</subScript>
and M
<subScript id="182A3482FF9F8371FE5AFAFDFE93EB3C" attach="left" box="[492,501,1314,1331]" fontSize="7" pageId="11" pageNumber="148">4</subScript>
pigmented brown, appearing more heavily sclerotized than other wing veins (
<figureCitation id="1C952A42FF9F8371FD1EFAEAFD91EB5F" box="[680,759,1333,1360]" captionStart="FIG" captionStartId="10.[207,218,1142,1160]" captionTargetBox="[132,1456,215,1098]" captionTargetId="figure@10.[127,1461,209,1106]" captionTargetPageId="10" captionText="FIG. 5. — Photomicrographs of male wings: A, Polletomyia subulata n. gen., n. sp.; B, Myiomystax trilineata n. gen., n. sp. Scale bars: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3743156" httpUri="https://zenodo.org/record/3743156/files/figure.png" pageId="11" pageNumber="148">Fig. 5B</figureCitation>
); costal vein with light pigment between apices of R
<subScript id="182A3482FF9F8371FD15FABDFDCBEB7C" attach="left" box="[675,685,1378,1395]" fontSize="7" pageId="11" pageNumber="148">1</subScript>
and R
<subScript id="182A3482FF9F8371FD40FABDFC66EB7C" attach="left" box="[758,768,1378,1395]" fontSize="7" pageId="11" pageNumber="148">3</subScript>
, M
<subScript id="182A3482FF9F8371FF2BFA5EFFC1EB9D" attach="left" box="[157,167,1409,1426]" fontSize="7" pageId="11" pageNumber="148">1</subScript>
and M
<subScript id="182A3482FF9F8371FF45FA5EFF9BEB9D" attach="left" box="[243,253,1409,1426]" fontSize="7" pageId="11" pageNumber="148">4</subScript>
; pigmented veins contrasting against other lightly colored wing veins. Male terminalia: bilaterally symmetrical; hypandrium flat, band-like, arched posteriorly, fused with gonocoxites laterally; intersegmental membrane connecting between anterior margin of hypandrium and posterior margin of abdominal segment 8, covering base of ejaculatory apodeme; ejaculatory apodeme dorsoventrally compressed anterolaterally, dome-like at center, articulating ventrally with gonocoxal apodemes, posteriorly with aedeagal sclerites; aedeagus comprised of paired, laterally compressed sclerites, spatulate in lateral view; spatulate sclerites enveloped by tubular membrane; parameres not differentiated from aedeagus; gonostyli tapered, sinuous from base to apex (
<figureCitation id="1C952A42FF9F8371FEE7F92BFEC1E901" box="[337,423,1780,1806]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="11" pageNumber="148">Fig. 3D</figureCitation>
), digitiform apically, setose, with paired, elongate setae inserted medially near apex (
<figureCitation id="1C952A42FF9F8371FD11F8CBFC66E921" box="[679,768,1812,1838]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="11" pageNumber="148">Figs 3D</figureCitation>
;
<figureCitation id="1C952A42FF9F8371FF32F8EBFFC2E941" box="[132,164,1844,1870]" captionStart="FIG" captionStartId="9.[132,143,1987,2005]" captionTargetBox="[302,1290,256,1858]" captionTargetId="figure@9.[260,1328,215,1893]" captionTargetPageId="9" captionText="FIG. 4. — Myiomystax trilineata n. gen., n. sp.:A, male terminalia,lateral view; B, female mouthparts and maxillary palpi,frontal view;C, female terminalia,external structure, ventral view; D, female terminalia, internal structure and egg, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743154" httpUri="https://zenodo.org/record/3743154/files/figure.png" pageId="11" pageNumber="148">4A</figureCitation>
); gonocoxites glabrous, without setae alveoli (
<figureCitation id="1C952A42FF9F8371FD13F8ECFD99E941" box="[677,767,1843,1870]" captionStart="FIG" captionStartId="8.[132,143,1987,2005]" captionTargetBox="[151,1438,231,1842]" captionTargetId="figure@8.[132,1455,216,1862]" captionTargetPageId="8" captionText="FIG. 3. — Myiomystax trilineata n. gen., n. sp.: A, wing; B, male head, frontal view; C, male epandrium, proctiger and epandrial claspers, dorsal view; D, male terminalia, epandrium removed, dorsal view. Abbreviations: see Material and methods. Scale bars: A, 0.5 mm; B, 0.25 mm; C-D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743152" httpUri="https://zenodo.org/record/3743152/files/figure.png" pageId="11" pageNumber="148">Figs 3D</figureCitation>
;
<figureCitation id="1C952A42FF9F8371FF32F88BFFC2E961" box="[132,164,1876,1902]" captionStart="FIG" captionStartId="9.[132,143,1987,2005]" captionTargetBox="[302,1290,256,1858]" captionTargetId="figure@9.[260,1328,215,1893]" captionTargetPageId="9" captionText="FIG. 4. — Myiomystax trilineata n. gen., n. sp.:A, male terminalia,lateral view; B, female mouthparts and maxillary palpi,frontal view;C, female terminalia,external structure, ventral view; D, female terminalia, internal structure and egg, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743154" httpUri="https://zenodo.org/record/3743154/files/figure.png" pageId="11" pageNumber="148">4A</figureCitation>
), cylindrical, about two times as long as wide; gonocoxal apodemes inflated, rounded in dorsoventral view, bulbous in lateral view, linked to subepandrial sclerite anteromedially; subepandrial sclerite narrow, divided medially, with puckered seam at median, with posterolateral arms ribbon-like, linked to base of epandrial claspers; epandrium rectangular along posterior half in dorsoventral view, anterior half narrowing abruptly; epandrial claspers elongate, tapered, curved dorsally from base to apex, with single, spatulate tenaculum inserted dorsoapically.
</paragraph>
<paragraph id="841136C7FF9F8371FC9BFEA7FC10EF9D" blockId="11.[813,1456,376,818]" box="[813,886,376,402]" pageId="11" pageNumber="148">
<emphasis id="B6DAEAD5FF9F8371FC9BFEA7FC10EF9D" box="[813,886,376,402]" italics="true" pageId="11" pageNumber="148">Female</emphasis>
</paragraph>
<paragraph id="841136C7FF9F8371FC9BFE46FBE3ED3E" blockId="11.[813,1456,376,818]" pageId="11" pageNumber="148">
Measurements, (N = 4) head width
<quantity id="43569B22FF9F8371FB0AFE46FA44EFBC" box="[1212,1314,409,435]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="148" unit="mm" value="0.44">0.44 mm</quantity>
(0.42-0.46), head height
<quantity id="43569B22FF9F8371FC0FFE66FB47EFDC" box="[953,1057,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="148" unit="mm" value="0.41">0.41 mm</quantity>
(0.40-0.42), wing length
<quantity id="43569B22FF9F8371FAFEFE67FAD6EFDC" box="[1352,1456,440,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" pageId="11" pageNumber="148" unit="mm" value="2.16">2.16 mm</quantity>
(2.04-2.36), wing width
<quantity id="43569B22FF9F8371FB83FE07FBF1EFFC" box="[1077,1175,472,499]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="148" unit="mm" value="0.53">0.53 mm</quantity>
(0.52-0.56). Head mostly as in male. Antennae with flagellomeres 1 and 2 elongate, cylindrical, shorter than in male; apical flagellomeres missing in specimens observed. Mouthparts as in male; maxillary palpi more typical of
<taxonomicName id="43AE4D44FF9F8371FC6AFD87FB02EC7D" authorityName="Newman" authorityYear="1834" box="[988,1124,600,626]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="148" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
, segments not elongated; palp segment proportions: 1, 1.30, 1, 2.30. Wing venation as in male. Terminalia: cerci conical in ventral view, with ventral surface covered by setulae, numerous setae inserted laterally and ventrally; posterior margin of sternum 8 trilobed, with median lobe wider than flanking lobes; oviduct with microtrichia internally on membrane.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,127 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.470.8548" ID-PMC="PMC4304036" ID-Pensoft-Pub="1313-2970-470-17" ID-Pensoft-UUID="FFDAFF85127CFFB3AA5915611C3A767A" ID-PubMed="25632243" ID-Zenodo-Dep="578680" ID-ZooBank="0E78A6A90B82401199EED5895E7F8A9E" ModsDocID="1313-2970-470-17" checkinTime="1451244867417" checkinUser="pensoft" docAuthor="Breure, Abraham S. H. &amp; Ablett, Jonathan D." docDate="2015" docId="0C07A67419072513AD1ACB6B841E29E4" docLanguage="en" docName="ZooKeys 470: 17-143" docOrigin="ZooKeys 470" docPubDate="2015-01-12" docSource="http://dx.doi.org/10.3897/zookeys.470.8548" docTitle="Bulimus iris Pfeiffer 1853" docType="treatment" docVersion="5" id="FFDAFF85127CFFB3AA5915611C3A767A" lastPageNumber="32" masterDocId="FFDAFF85127CFFB3AA5915611C3A767A" masterDocTitle="Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London" masterLastPageNumber="143" masterPageNumber="17" pageNumber="32" updateTime="1668159795410" updateUser="ExternalLinkService">
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<mods:title>Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London</mods:title>
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<mods:namePart>Breure, Abraham S. H.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK</mods:affiliation>
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<mods:number>2015-01-12</mods:number>
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<treatment ID-GBIF-Taxon="152057468" LSID="urn:lsid:plazi:treatment:0C07A67419072513AD1ACB6B841E29E4" httpUri="http://treatment.plazi.org/id/0C07A67419072513AD1ACB6B841E29E4" lastPageNumber="32" pageId="15" pageNumber="32">
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<paragraph pageId="15" pageNumber="32">
<taxonomicName LSID="0C07A674-1907-2513-AD1A-CB6B841E29E4" authority="Pfeiffer, 1853" authorityName="Pfeiffer" authorityYear="1853" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus iris" order="Stylommatophora" pageId="15" pageNumber="32" phylum="Mollusca" rank="species" species="iris">Bulimus iris Pfeiffer, 1853</taxonomicName>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Porphyrobaphe species. i-ii Porphyrobaphe (Porphyrobaphe) approximata Fulton, 1896, syntype NHMUK 1895.12.19.44 (H = 65.7) iii-v Porphyrobaphe (Porphyrobaphe) iris (Pfeiffer, 1853), lectotype NHMUK 20100506 (H = 72.6)." pageId="15" pageNumber="32">Figs 14iii-v</figureCitation>
<figureCitation captionStart="Figure L10" captionStartId="F40" captionText="Figure L 10. i Bulimus integer Pfeiffer, 185 5 ii Bulimus iris Pfeiffer, 1853 iii Bulimus irroratus Reeve, 1849." pageId="15" pageNumber="32">, L10ii</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="reference_group">
<paragraph pageId="15" pageNumber="32">
<taxonomicName authorityName="Pfeiffer" authorityYear="1853" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus iris" order="Stylommatophora" pageId="15" pageNumber="32" phylum="Mollusca" rank="species" species="iris">Bulimus iris</taxonomicName>
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Monographia heliceorum viventium: sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum" pageId="49" pageNumber="66" refId="B72" refString="Pfeiffer, L, 1853b. . Monographia heliceorum viventium: sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum 3" volume="3" year="1853 b">Pfeiffer 1853b</bibRefCitation>
: 313;
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Monographia heliceorum viventium: sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum" pageId="49" pageNumber="66" pagination="135 - 138" refId="B74" refString="Pfeiffer, L, 1854b. Descriptions of fourteen new species of land shells, from the collection of Hugh Cuming Esq. In: Proceedings of the Zoological Society of London.: 135 - 138" title="Descriptions of fourteen new species of land shells, from the collection of Hugh Cuming Esq." volumeTitle="Proceedings of the Zoological Society of London." year="1854 b">Pfeiffer 1854b</bibRefCitation>
: 136; Pfeiffer 1855 in
<bibRefCitation author="Kuester, HC" journalOrPublisher="Systematisches Conchylien-Cabinet von Martini und Chemnitz" pageId="48" pageNumber="65" refId="B52" refString="Kuester, HC, Pfeiffer, L, 1840-1855 1840-1865. Die Gattungen Bulimus, Partula, Achatinella, Achatina und Azeca. Systematisches Conchylien-Cabinet von Martini und Chemnitz 13 (1)" title="Die Gattungen Bulimus, Partula, Achatinella, Achatina und Azeca." volume="13" year="1840 - 1855">
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
and Pfeiffer 1840-1865
</bibRefCitation>
: 244, pl. 65 figs 4-5.
</paragraph>
<paragraph pageId="15" pageNumber="32">
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Bulimulidae" genus="Porphyrobaphe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porphyrobaphe iris" order="Stylommatophora" pageId="15" pageNumber="32" phylum="Mollusca" rank="species" species="iris">Porphyrobaphe iris</taxonomicName>
;
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Manual of Conchology" pageId="50" pageNumber="67" pagination="1 - 258" refId="B101" refString="Pilsbry, HA, 1899. American Bulimulidae: North American and Antillean Drymaeus, Leiostracus, Orthalicinae and Amphibuliminae. Manual of Conchology 12: 1 - 258" title="American Bulimulidae: North American and Antillean Drymaeus, Leiostracus, Orthalicinae and Amphibuliminae." volume="12" year="1899">Pilsbry 1899</bibRefCitation>
: 157, pl. 51 figs 28-29.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="type locality">
<paragraph pageId="15" pageNumber="32">Type locality.</paragraph>
<paragraph pageId="15" pageNumber="32">&quot;Le Ceja, Rio Negro Novae Granadae (Bland)&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="label">
<paragraph pageId="15" pageNumber="32">Label.</paragraph>
<paragraph pageId="15" pageNumber="32">
&quot;La Ceja. Rio Negro / New Grenada&quot;, taxon label in
<normalizedToken originalValue="Pfeiffers">Pfeiffer's</normalizedToken>
handwriting. M.C. label style I.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="dimensions">
<paragraph pageId="15" pageNumber="32">Dimensions.</paragraph>
<paragraph pageId="15" pageNumber="32">&quot;Long. 64, diam. 32 mill.&quot;. Figured specimen H 72.6, D 41.1, W 5.8.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="type material">
<paragraph pageId="15" pageNumber="32">Type material.</paragraph>
<paragraph pageId="15" pageNumber="32">NHMUK 20100506, lectotype (Cuming coll.).</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="remarks">
<paragraph pageId="15" pageNumber="32">Remarks.</paragraph>
<paragraph pageId="15" pageNumber="32">
Pfeiffer did not state on how many specimens his description was based, but described the species from &quot;Mus. Cuming&quot;. Although the specimen is larger than the measurements given by Pfeiffer, there is little doubt it is from the original series as the label confirms the original locality; it also bears the text &quot;please to name this / '
<emphasis italics="true" pageId="15" pageNumber="32">Blandi</emphasis>
' after the collector&quot;. The specimen is now designated lectotype (
<emphasis bold="true" pageId="15" pageNumber="32">design. n.</emphasis>
) to fixate this taxon, which needs further study to clarify its status. The mentioning in
<bibRefCitation author="Linares, EL" journalOrPublisher="Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota" pageId="48" pageNumber="65" refId="B53" refString="Linares, EL, Vera, ML, 2012. Catalogo de los moluscos continentales de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota" title="Catalogo de los moluscos continentales de Colombia." year="2012">Linares and Vera (2012</bibRefCitation>
: 157) of &quot;ZMUZ 511864&quot; as lectotype for this taxon is erroneous, as this refers to the type specimen of the junior subjective synonym
<taxonomicName authorityName="Mousson" authorityYear="1873" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus wallisianus" order="Pulmonata" pageId="15" pageNumber="32" phylum="Mollusca" rank="species" species="wallisianus">
<emphasis italics="true" pageId="15" pageNumber="32">Bulimus wallisianus</emphasis>
</taxonomicName>
Mousson, 1873 (see
<bibRefCitation author="Breure, ASH" journalOrPublisher="Backhuys, Rotterdam" pageId="45" pageNumber="62" pagination="1 - 4" refId="B8" refString="Breure, ASH, 1976. Types of Bulimulidae (Gastropoda, Euthyneura) in the Zoologisches Museum, Universitaet Zuerich. In: Malacologische opstellen ter gelegenheid van het tienjarig bestaan van de malacologische contactgroep 'Amsterdam en omstreken De Kreukel'. Backhuys, Rotterdam: 1 - 4" title="Types of Bulimulidae (Gastropoda, Euthyneura) in the Zoologisches Museum, Universitaet Zuerich. In: Malacologische opstellen ter gelegenheid van het tienjarig bestaan van de malacologische contactgroep ' Amsterdam en omstreken De Kreukel'" year="1976">Breure 1976</bibRefCitation>
: 3). The current systematic position follows
<bibRefCitation author="Richardson, CL" journalOrPublisher="Tryonia" pageId="51" pageNumber="68" pagination="1 - 164" refId="B113" refString="Richardson, CL, 1993. Bulimulacea: catalog of species. Amphibulimidae, Anadromidae, Grangerellidae, Odontostomidae, Orthalicidae. Tryonia 27: 1 - 164" title="Bulimulacea: catalog of species. Amphibulimidae, Anadromidae, Grangerellidae, Odontostomidae, Orthalicidae." volume="27" year="1993">Richardson (1993</bibRefCitation>
: 119).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="32" type="current systematic position">
<paragraph pageId="15" pageNumber="32">Current systematic position.</paragraph>
<paragraph pageId="15" pageNumber="32">
<taxonomicName class="Gastropoda" family="Orthalicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="15" pageNumber="32" phylum="Mollusca" rank="family">Orthalicidae</taxonomicName>
,
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Orthalicidae" genus="Porphyrobaphe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porphyrobaphe (Porphyrobaphe) iris" order="Stylommatophora" pageId="15" pageNumber="32" phylum="Mollusca" rank="species" species="iris" subGenus="Porphyrobaphe">Porphyrobaphe (Porphyrobaphe) iris</taxonomicName>
(Pfeiffer, 1853).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,104 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.711.14753" ID-PMC="PMC5674181" ID-Pensoft-Pub="1313-2970-711-1" ID-Pensoft-UUID="FFC4FE3CFFAAFF87F42FFF91FFACFFC3" ID-PubMed="29134026" ID-Zenodo-Dep="1149948" ID-ZooBank="D94E8B43C7A7447386D4FFBFAD6852DC" ModsDocID="1313-2970-711-1" checkinTime="1508771341586" checkinUser="pensoft" docAuthor="Lopes, Daniela A., Gomes, Delir Correa &amp; Knoff, Marcelo" docDate="2017" docId="0C080EB78D658AF3AEFFC6E5A8603D89" docLanguage="en" docName="ZooKeys 711: 1-52" docOrigin="ZooKeys 711" docPubDate="2017-10-23" docSource="http://dx.doi.org/10.3897/zookeys.711.14753" docTitle="Aprocta brevipenis Rodrigues &amp; Rodrigues 1980" docType="treatment" docVersion="6" id="FFC4FE3CFFAAFF87F42FFF91FFACFFC3" lastPageNumber="7" masterDocId="FFC4FE3CFFAAFF87F42FFF91FFACFFC3" masterDocTitle="Type material of Acanthocephala, Nematoda and other non-helminths phyla (Cnidaria, Annelida, and Arthropoda) housed in the Helminthological Collection of the Oswaldo Cruz Institute / FIOCRUZ (CHIOC), Rio de Janeiro, Brazil, from 1979 to 2016" masterLastPageNumber="52" masterPageNumber="1" pageNumber="7" updateTime="1668165017486" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type material of Acanthocephala, Nematoda and other non-helminths phyla (Cnidaria, Annelida, and Arthropoda) housed in the Helminthological Collection of the Oswaldo Cruz Institute / FIOCRUZ (CHIOC), Rio de Janeiro, Brazil, from 1979 to 2016</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lopes, Daniela A.</mods:namePart>
<mods:affiliation>Laboratorio de Helmintos Parasitos de Vertebrados, Instituto Oswaldo Cruz, FIOCRUZ, Av. Brasil, 4365 Rio de Janeiro, RJ, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gomes, Delir Correa</mods:namePart>
<mods:affiliation>Laboratorio de Helmintos Parasitos de Vertebrados, Instituto Oswaldo Cruz, FIOCRUZ, Av. Brasil, 4365 Rio de Janeiro, RJ, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knoff, Marcelo</mods:namePart>
<mods:affiliation>Laboratorio de Helmintos Parasitos de Vertebrados, Instituto Oswaldo Cruz, FIOCRUZ, Av. Brasil, 4365 Rio de Janeiro, RJ, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">knoffm@ioc.fiocruz.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>711</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>52</mods:end>
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<mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="Pensoft-Pub">1313-2970-711-1</mods:identifier>
<mods:identifier type="ZooBank">D94E8B43C7A7447386D4FFBFAD6852DC</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFC4FE3CFFAAFF87F42FFF91FFACFFC3</mods:identifier>
<mods:identifier type="Zenodo-Dep">1149948</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="135452890" LSID="urn:lsid:plazi:treatment:0C080EB78D658AF3AEFFC6E5A8603D89" httpUri="http://treatment.plazi.org/id/0C080EB78D658AF3AEFFC6E5A8603D89" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection pageId="6" pageNumber="7" type="nomenclature">
<paragraph pageId="6" pageNumber="7">
<taxonomicName LSID="0C080EB7-8D65-8AF3-AEFF-C6E5A8603D89" authority="Rodrigues &amp; Rodrigues, 1980" authorityName="Rodrigues &amp; Rodrigues" authorityYear="1980" class="Chromadorea" family="Aproctidae" genus="Aprocta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aprocta brevipenis" order="Rhabditida" pageId="6" pageNumber="7" phylum="Nematoda" rank="species" species="brevipenis">Aprocta brevipenis Rodrigues &amp; Rodrigues, 1980</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="type host">
<paragraph pageId="6" pageNumber="7">Type host.</paragraph>
<paragraph pageId="6" pageNumber="7">
<taxonomicName authority="guira" class="Aves" family="Cuculidae" genus="Guira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Guira guira" order="Cuculiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="guira">
<emphasis italics="true" pageId="6" pageNumber="7">Guira guira</emphasis>
</taxonomicName>
(Gmelin, 1788) (
<taxonomicName authorityName="Lesson" authorityYear="1830" class="Aves" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" pageId="6" pageNumber="7" phylum="Chordata" rank="class">Aves</taxonomicName>
:
<taxonomicName class="Aves" family="Cuculidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Columbiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="family">Cuculidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="infection site">
<paragraph pageId="6" pageNumber="7">Infection site.</paragraph>
<paragraph pageId="6" pageNumber="7">Ocular cavity.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="type locality">
<paragraph pageId="6" pageNumber="7">Type locality.</paragraph>
<paragraph pageId="6" pageNumber="7">
Brazil,
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo State, Ilha Seca.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="holotype">
<paragraph pageId="6" pageNumber="7">Holotype.</paragraph>
<paragraph pageId="6" pageNumber="7">♂ CHIOC 31808 a.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="paratypes">
<paragraph pageId="6" pageNumber="7">Paratypes.</paragraph>
<paragraph pageId="6" pageNumber="7">CHIOC 31808 b (allotype ♀), 31808 c-d (♀♀).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="reference">
<paragraph pageId="6" pageNumber="7">Reference.</paragraph>
<paragraph pageId="6" pageNumber="7">
<bibRefCitation author="Rodrigues, HO" journalOrPublisher="Atas da Sociedade de Biologia do Rio de Janeiro" pageId="38" pageNumber="39" pagination="12 - 14" refId="B95" refString="Rodrigues, HO, Rodrigues, SS, 1980. Nova especie do genero Aprocta Linstow, 1883 (Nematoda, Aproctoidea). Atas da Sociedade de Biologia do Rio de Janeiro 21: 12 - 14" title="Nova especie do genero Aprocta Linstow, 1883 (Nematoda, Aproctoidea)." volume="21" year="1980">Rodrigues and Rodrigues (1980)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

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<mods:title id="121DCA66074010E5887B4ED06DE1D750">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9007859C5B8400DFEED2207" ID-DOI="http://doi.org/10.5281/zenodo.7300269" ID-GBIF-Taxon="204205234" ID-Zenodo-Dep="7300269" LSID="urn:lsid:plazi:treatment:0C081779B9007859C5B8400DFEED2207" httpUri="http://treatment.plazi.org/id/0C081779B9007859C5B8400DFEED2207" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="CCBBF5E4B9007859C5B8400DFDBC273E" box="[151,597,546,573]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="841EA66FB9007859C5B8400DFDBC273E" blockId="19.[151,662,546,609]" box="[151,597,546,573]" pageId="19" pageNumber="20">
<heading id="DF561103B9007859C5B8400DFDBC273E" bold="true" box="[151,597,546,573]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
<emphasis id="B6D57A7DB9007859C5B8400DFDBC273E" bold="true" box="[151,597,546,573]" pageId="19" pageNumber="20">
<taxonomicName id="43A1DDECB9007859C5B8400DFE5E273F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,439,546,573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="camerolatula" status="sp. nov.">
<emphasis id="B6D57A7DB9007859C5B8400DFE5E273F" bold="true" box="[151,439,546,573]" italics="true" pageId="19" pageNumber="20">Hydraena camerolatula</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9007859C4EA400CFDBC273E" box="[453,597,546,573]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5B84068FD7F2763" box="[151,662,583,609]" pageId="19" pageNumber="20" type="description">
<paragraph id="841EA66FB9007859C5B84068FD7F2763" blockId="19.[151,662,546,609]" box="[151,662,583,609]" pageId="19" pageNumber="20">
<figureCitation id="1C9ABAEAB9007859C5B84068FF0C2763" box="[151,229,583,609]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="19" pageNumber="20">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB9007859C5DE4068FEE42763" box="[241,269,583,609]" captionStart="FIGURE 23" captionStartId="48.[151,250,1985,2009]" captionTargetBox="[221,1361,195,1940]" captionTargetId="figure-25@48.[180,1407,181,1961]" captionTargetPageId="48" captionText="FIGURE 23. Hydraena camerolatula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284913" httpUri="https://zenodo.org/record/7284913/files/figure.png" pageId="19" pageNumber="20">23</figureCitation>
(habitus), 23 (aedeagus), 43 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5B840A0FEFB27F3" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="841EA66FB9007859C5B840A0FEFB27F3" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5B840A0FEAF27AA" bold="true" box="[151,326,654,681]" pageId="19" pageNumber="20">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9007859C46440A0FC1227CF" ID-GBIF-Occurrence="3962178311" collectingDate="1949-01-07" collectingDateMax="1949-01-11" collectingDateMin="1949-01-07" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9007859C46440A0FE5A27AB" box="[331,435,655,681]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
(male):
<location id="817EF0B4B9007859C72140A0FD6E27AB" LSID="urn:lsid:plazi:treatment:0C081779B9007859C5B8400DFEED2207:817EF0B4B9007859C72140A0FD6E27AB" box="[526,647,655,681]" country="Cameroon" municipality="Northwest Province" name="Cameroon" pageId="19" pageNumber="20">Cameroon</location>
:
<collectingMunicipality id="647A3C15B9007859C7A040A0FC9927AB" box="[655,880,655,681]" pageId="19" pageNumber="20">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB9007859C6AD40A0FB4227AB" box="[898,1195,655,681]" name="Cameroon" pageId="19" pageNumber="20">BRITISH CAMEROONS</collectingCountry>
. / Mamfe,
<date id="F01F80AFB9007859C03340A0FA7527AB" box="[1308,1436,655,681]" pageId="19" pageNumber="20" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">
<collectingDate id="E05B7947B9007859C03340A0FA7527AB" box="[1308,1436,655,681]" pageId="19" pageNumber="20" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">7-11.i.1949</collectingDate>
</date>
/
<collectorName id="2954C3B9B9007859C587409CFEF527CF" box="[168,284,691,717]" pageId="19" pageNumber="20">B. Malkin</collectorName>
coll. // Rain forest; clear stream / Gravel and sand” (
<collectionCode id="E2B03EAAB9007859C6AD409CFC1A27CF" box="[898,1011,691,717]" pageId="19" pageNumber="20">UKNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9007859C128409CFEE727F3" ID-GBIF-Occurrence="3962178333" collectingDate="1949-01-07" collectingDateMax="1949-01-11" collectingDateMin="1949-01-07" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="19" pageNumber="20" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9007859C128409CFB8427CF" box="[1031,1133,691,717]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
: Same data as holotype (1
<collectionCode id="E2B03EAAB9007859C5B840F8FEEF27F3" box="[151,262,727,753]" pageId="19" pageNumber="20">UKNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E840D5FBCA26A7" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="841EA66FB9007859C5E840D5FBCA26A7" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E840D5FE382616" bold="true" box="[199,465,762,788]" pageId="19" pageNumber="20">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9007859C12740D4FB942617" box="[1032,1149,763,789]" name="Cameroon" pageId="19" pageNumber="20">Cameroon</collectingCountry>
by the combination of the moderately densely punctate dorsum, the relatively wide mesoventral intercoxal process (P2, l/w ca. 2/1), and the aedeagus (
<figureCitation id="1C9ABAEAB9007859C423416CFE89265F" box="[268,352,835,861]" captionStart="FIGURE 23" captionStartId="48.[151,250,1985,2009]" captionTargetBox="[221,1361,195,1940]" captionTargetId="figure-25@48.[180,1407,181,1961]" captionTargetPageId="48" captionText="FIGURE 23. Hydraena camerolatula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284913" httpUri="https://zenodo.org/record/7284913/files/figure.png" pageId="19" pageNumber="20">Fig. 23</figureCitation>
).
<taxonomicName id="43A1DDECB9007859C45C416CFD95265F" authorityName="Perkins" authorityYear="2022" box="[371,636,835,861]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB9007859C45C416CFD95265F" box="[371,636,835,861]" italics="true" pageId="19" pageNumber="20">Hydraena cameropetila</emphasis>
</taxonomicName>
is also densely punctate, but much smaller (1.20), and lacks plaques.
<taxonomicName id="43A1DDECB9007859C0AE416CFEA02683" authorityName="Perkins" authorityYear="2022" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB9007859C0AE416CFEA02683" italics="true" pageId="19" pageNumber="20">H. camerocrebrata</emphasis>
</taxonomicName>
is also densely punctate, but much smaller (ca. 1.33 vs. 1.50), and the plaques are larger (P2/w/l/s ratios ca. 1/1/6/6 vs. 1/0.2/2/2). Also refer to the diagnosis of
<taxonomicName id="43A1DDECB9007859C66841A4FBF526A7" authorityName="Perkins" authorityYear="2022" box="[839,1052,907,933]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB9007859C66841A4FBF526A7" box="[839,1052,907,933]" italics="true" pageId="19" pageNumber="20">H. camerocrebrata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E84181FCCE239B" pageId="19" pageNumber="20" type="description">
<paragraph id="841EA66FB9007859C5E84181FF1D21C7" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E84181FEB826CA" bold="true" box="[199,337,942,968]" pageId="19" pageNumber="20">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9007859C48B4180FDEF26CB" box="[420,518,943,969]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.49/0.62; head width 0.36; pronotum 0.37/0.50, PA 0.42, PB 0.44; elytra 0.88/0.62. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9007859C12E41FCFBBC26EF" box="[1025,1109,979,1005]" captionStart="FIGURE 23" captionStartId="48.[151,250,1985,2009]" captionTargetBox="[221,1361,195,1940]" captionTargetId="figure-25@48.[180,1407,181,1961]" captionTargetPageId="48" captionText="FIGURE 23. Hydraena camerolatula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284913" httpUri="https://zenodo.org/record/7284913/files/figure.png" pageId="19" pageNumber="20">Fig. 23</figureCitation>
). Dorsum light brown to dark brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head moderately finely sparsely punctulate, pronotum moderately coarsely, moderately densely punctate, punctures much larger than those of frons, interstices shining, ca. 12xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae moderately deep, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB9007859C5E846E0FEB3207B" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">Elytra with summit of posterior declivity slightly beyond midlength, where elytra transversely quite convex; lateral explanate margins moderately wide; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB9007859C5B847ACFCCE239B" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">Venter: Mentum and postmentum very finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina sinuate in profile. P2 relatively wide, l/w ca. 2/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, very indistinct, straight, parallel, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1x P2, or slightly narrower than P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with slightly off center shallow notch in tergite, into which minute apical process of ventrite fits.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E8448DFC2123BF" box="[199,968,1698,1725]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="841EA66FB9007859C5E8448DFC2123BF" blockId="19.[151,1437,654,1797]" box="[199,968,1698,1725]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E8448DFE8823BE" bold="true" box="[199,353,1698,1724]" pageId="19" pageNumber="20">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9007859C7E3448CFD1523BF" box="[716,764,1699,1725]" pageId="19" pageNumber="20">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9007859C648448CFC5223BF" box="[871,955,1699,1725]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="19" pageNumber="20">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E844E8FEED2207" pageId="19" pageNumber="20" type="etymology">
<paragraph id="841EA66FB9007859C5E844E8FEED2207" blockId="19.[151,1437,654,1797]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E844E8FEA723E3" bold="true" box="[199,334,1735,1761]" pageId="19" pageNumber="20">Etymology.</emphasis>
Named in reference to the known distribution and the relatively wide mesoventral intercoxal process (P2).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,167 @@
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<mods:title id="0CD8AAC31E13516BD27455E488A9F52A">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9017859C5B846A9FC6C24D3" ID-DOI="http://doi.org/10.5281/zenodo.7300200" ID-GBIF-Taxon="204205233" ID-Zenodo-Dep="7300200" LSID="urn:lsid:plazi:treatment:0C081779B9017859C5B846A9FC6C24D3" httpUri="http://treatment.plazi.org/id/0C081779B9017859C5B846A9FC6C24D3" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="CCBBF5E4B9017858C5B846A9FD9821A2" box="[151,625,1158,1185]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="841EA66FB9017858C5B846A9FD9821A2" blockId="18.[151,662,1158,1220]" box="[151,625,1158,1185]" pageId="18" pageNumber="19">
<heading id="DF561103B9017858C5B846A9FD9821A2" bold="true" box="[151,625,1158,1185]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="3">
<emphasis id="B6D57A7DB9017858C5B846A9FD9821A2" bold="true" box="[151,625,1158,1185]" pageId="18" pageNumber="19">
<taxonomicName id="43A1DDECB9017858C5B846A9FE3B21A2" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,466,1158,1185]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cameroconfinis" status="sp. nov.">
<emphasis id="B6D57A7DB9017858C5B846A9FE3B21A2" bold="true" box="[151,466,1158,1185]" italics="true" pageId="18" pageNumber="19">Hydraena cameroconfinis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9017858C4CE46A9FD9821A2" box="[481,625,1158,1184]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5B84685FD7F21C6" box="[151,662,1194,1220]" pageId="18" pageNumber="19" type="description">
<paragraph id="841EA66FB9017858C5B84685FD7F21C6" blockId="18.[151,662,1158,1220]" box="[151,662,1194,1220]" pageId="18" pageNumber="19">
<figureCitation id="1C9ABAEAB9017858C5B84685FF0C21C6" box="[151,229,1194,1220]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="18" pageNumber="19">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB9017858C5DE4685FEE421C6" box="[241,269,1194,1220]" captionStart="FIGURE 22" captionStartId="47.[151,250,1991,2015]" captionTargetBox="[316,1272,181,1967]" captionTargetId="figure-15@47.[316,1272,181,1967]" captionTargetPageId="47" captionText="FIGURE 22. Hydraena cameroconfinis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284909" httpUri="https://zenodo.org/record/7284909/files/figure.png" pageId="18" pageNumber="19">22</figureCitation>
(habitus), 22 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5B846DDFEB72056" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="841EA66FB9017858C5B846DDFEB72056" blockId="18.[151,1437,1266,2013]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5B846DDFEA3200E" bold="true" box="[151,330,1266,1292]" pageId="18" pageNumber="19">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9017858C47C46DDFF032032" ID-GBIF-Occurrence="3962178349" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9017858C47C46DDFE52200E" box="[339,443,1266,1292]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB9017858C73046DDFDC4200E" bold="true" box="[543,557,1266,1292]" pageId="18" pageNumber="19"></emphasis>
<collectingCountry id="FCB6E6FFB9017858C70246DDFD4B200E" box="[557,674,1266,1292]" name="Cameroon" pageId="18" pageNumber="19">Cameroon</collectingCountry>
:
<quantity id="43590B8AB9017858C79B46DDFD14200E" box="[692,765,1266,1293]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B9017858C62946DCFC5E200E" box="[774,951,1266,1293]" pageId="18" pageNumber="19">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB9017858C13046DDFB68200E" box="[1055,1153,1266,1292]" pageId="18" pageNumber="19" value="1982-05-01">
<collectingDate id="E05B7947B9017858C13046DDFB68200E" box="[1055,1153,1266,1292]" pageId="18" pageNumber="19" value="1982-05-01">1.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B9017858C1A046DCFACC200E" box="[1167,1317,1266,1293]" pageId="18" pageNumber="19">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB9017858C5B04739FF0B2032" box="[159,226,1302,1328]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="18" pageNumber="19" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9017858C5D84738FEB32056" ID-GBIF-Occurrence="3962178320" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="18" pageNumber="19" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9017858C5D84738FE812033" box="[247,360,1303,1329]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<collectorName id="2954C3B9B9017858C45A4739FE5A2032" box="[373,435,1302,1328]" pageId="18" pageNumber="19">Same</collectorName>
data as holotype (122
<collectionCode id="E2B03EAAB9017858C7934739FD172032" box="[700,766,1302,1328]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="18" pageNumber="19" type="Museum">MCZ</collectionCode>
);
<quantity id="43590B8AB9017858C63C4739FC892032" box="[787,864,1302,1328]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="km" value="12.0">12 km</quantity>
.
<collectorName id="2954C3B9B9017858C6424738FBF42032" box="[877,1053,1302,1329]" pageId="18" pageNumber="19">N. Ngaoundere</collectorName>
, lake margin,
<date id="F01F80AFB9017858C1EE4739FACD2032" box="[1217,1316,1302,1328]" pageId="18" pageNumber="19" value="1982-05-01">
<collectingDate id="E05B7947B9017858C1EE4739FACD2032" box="[1217,1316,1302,1328]" pageId="18" pageNumber="19" value="1982-05-01">1.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9017858C04B4738FF022056" pageId="18" pageNumber="19">P. D. Perkins</collectorName>
(1
<collectionCode id="E2B03EAAB9017858C43E4715FEBA2056" box="[273,339,1338,1364]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="18" pageNumber="19" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5E84771FC9B230A" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="841EA66FB9017858C5E84771FC9B230A" blockId="18.[151,1437,1266,2013]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E84771FE3D207A" bold="true" box="[199,468,1374,1400]" pageId="18" pageNumber="19">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9017858C10B4771FB70207A" box="[1060,1177,1374,1400]" name="Cameroon" pageId="18" pageNumber="19">Cameroon</collectingCountry>
by the combination of the comparatively large size (ca.
<quantity id="43590B8AB9017858C72647ADFD84209F" box="[521,621,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="18" pageNumber="19" unit="mm" value="1.62">1.62 mm</quantity>
vs. 1.101.55 for other known species), the narrow metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB9017858C4474789FE4420C2" box="[360,429,1446,1472]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
). The dorsal habitus is similar to that of
<taxonomicName id="43A1DDECB9017858C6574788FBDD20C2" authorityName="Perkins" authorityYear="2022" box="[888,1076,1446,1472]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="camerocubita">
<emphasis id="B6D57A7DB9017858C6574788FBDD20C2" box="[888,1076,1446,1472]" italics="true" pageId="18" pageNumber="19">H. camerocubita</emphasis>
</taxonomicName>
, but
<taxonomicName id="43A1DDECB9017858C1414788FAD520C2" authorityName="Perkins" authorityYear="2022" box="[1134,1340,1446,1472]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cameroconfinis">
<emphasis id="B6D57A7DB9017858C1414788FAD520C2" box="[1134,1340,1446,1472]" italics="true" pageId="18" pageNumber="19">H. cameroconfinis</emphasis>
</taxonomicName>
is much larger (ca. 1.62 vs.
<quantity id="43590B8AB9017858C44447E5FE3B20E6" box="[363,466,1482,1508]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="18" pageNumber="19" unit="mm" value="1.26">1.26 mm</quantity>
), and has different ventral proportions (P2/w/l/s ca. 1/0.25/1.5/1 vs. 1/0.15/0.15/2). The aedeagi of the two species are markedly dissimilar (
<figureCitation id="1C9ABAEAB9017858C7F347C1FCD0230B" box="[732,825,1518,1545]" captionStart="FIGURE 15" captionStartId="40.[151,250,1979,2003]" captionTargetBox="[256,1332,181,1956]" captionTargetId="figure-25@40.[256,1332,181,1956]" captionTargetPageId="40" captionText="FIGURE 15. Hydraena camerocubita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284886" httpUri="https://zenodo.org/record/7284886/files/figure.png" pageId="18" pageNumber="19">Figs. 15</figureCitation>
,
<figureCitation id="1C9ABAEAB9017858C66A47C1FC8C230A" box="[837,869,1518,1544]" captionStart="FIGURE 22" captionStartId="47.[151,250,1991,2015]" captionTargetBox="[316,1272,181,1967]" captionTargetId="figure-15@47.[316,1272,181,1967]" captionTargetPageId="47" captionText="FIGURE 22. Hydraena cameroconfinis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284909" httpUri="https://zenodo.org/record/7284909/files/figure.png" pageId="18" pageNumber="19">22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017859C5E8443DFBED2467" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="841EA66FB9017858C5E8443DFF1D222A" blockId="18.[151,1437,1266,2013]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E8443DFEB8232E" bold="true" box="[199,337,1554,1580]" pageId="18" pageNumber="19">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9017858C48B443DFDEF232E" box="[420,518,1554,1580]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.62/0.68; head width 0.38; pronotum 0.39/0.52, PA 0.44, PB 0.45; elytra 0.94/0.68. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9017858C1074419FB962352" box="[1064,1151,1590,1616]" captionStart="FIGURE 22" captionStartId="47.[151,250,1991,2015]" captionTargetBox="[316,1272,181,1967]" captionTargetId="figure-15@47.[316,1272,181,1967]" captionTargetPageId="47" captionText="FIGURE 22. Hydraena cameroconfinis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284909" httpUri="https://zenodo.org/record/7284909/files/figure.png" pageId="18" pageNumber="19">Fig. 22</figureCitation>
). Dorsum brown to light brown, head and pronotal macula darker, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 23xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB9017858C5E8451DFBA922BA" blockId="18.[151,1437,1266,2013]" pageId="18" pageNumber="19">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row.Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB9017859C5E845ECFBED2467" blockId="18.[151,1437,1266,2013]" lastBlockId="19.[151,1437,150,465]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.25/1.5/1. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques narrow, straight, slightly converging toward one another anteriorly, weakly raised, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS concave, width at straight posterior margin ca. 1x P2. Pro- and mesotibiae slender, straight. Metatibia slender, straight, inner margin with row of short setae. Abdominal apex with deep asymmetrical slightly off-center notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E84341FC21248B" box="[199,968,366,393]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="841EA66FB9007859C5E84341FC21248B" blockId="19.[151,1437,150,465]" box="[199,968,366,393]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E84341FE88248A" bold="true" box="[199,353,366,392]" pageId="19" pageNumber="20">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9007859C7E34340FD15248B" box="[716,764,367,393]" pageId="19" pageNumber="20">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9007859C6484340FC52248B" box="[871,955,367,393]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="19" pageNumber="20">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9007859C5E843BCFC6C24D3" pageId="19" pageNumber="20" type="etymology">
<paragraph id="841EA66FB9007859C5E843BCFC6C24D3" blockId="19.[151,1437,150,465]" pageId="19" pageNumber="20">
<emphasis id="B6D57A7DB9007859C5E843BCFEA724AF" bold="true" box="[199,334,403,429]" pageId="19" pageNumber="20">Etymology.</emphasis>
Named in reference to the known distribution, near the northern limit of the Sudano-Guinean zone (zone 2), just south of the very dry Sudano-Sahelian zone (zone 1).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,148 @@
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<mods:title id="F695CDF5BB1FB6C04EB34C282ABF918E">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<subSubSection id="CCBBF5E4B902785BC5B8451DFD67224E" box="[151,654,1842,1869]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="841EA66FB902785BC5B8451DFD67224E" blockId="17.[151,662,1842,1905]" box="[151,654,1842,1869]" pageId="17" pageNumber="18">
<heading id="DF561103B902785BC5B8451DFD67224E" bold="true" box="[151,654,1842,1869]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<emphasis id="B6D57A7DB902785BC5B8451DFD67224E" bold="true" box="[151,654,1842,1869]" pageId="17" pageNumber="18">
<taxonomicName id="43A1DDECB902785BC5B8451DFE19224F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,496,1842,1869]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="camerocompressa" status="sp. nov.">
<emphasis id="B6D57A7DB902785BC5B8451DFE19224F" bold="true" box="[151,496,1842,1869]" italics="true" pageId="17" pageNumber="18">Hydraena camerocompressa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B902785BC4D1451CFD67224E" box="[510,654,1842,1869]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5B84578FD7F2273" box="[151,662,1879,1905]" pageId="17" pageNumber="18" type="description">
<paragraph id="841EA66FB902785BC5B84578FD7F2273" blockId="17.[151,662,1842,1905]" box="[151,662,1879,1905]" pageId="17" pageNumber="18">
<figureCitation id="1C9ABAEAB902785BC5B84578FF0C2273" box="[151,229,1879,1905]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="17" pageNumber="18">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB902785BC5DE4578FEE42273" box="[241,269,1879,1905]" captionStart="FIGURE 21" captionStartId="46.[151,250,1986,2010]" captionTargetBox="[253,1333,181,1963]" captionTargetId="figure-25@46.[253,1333,181,1963]" captionTargetPageId="46" captionText="FIGURE 21. Hydraena camerocompressa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284905" httpUri="https://zenodo.org/record/7284905/files/figure.png" pageId="17" pageNumber="18">21</figureCitation>
(habitus), 21 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5B845B0FD3722DF" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="841EA66FB902785BC5B845B0FD3722DF" blockId="17.[151,1436,1950,2013]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5B845B0FEA322BA" bold="true" box="[151,330,1950,1977]" pageId="17" pageNumber="18">Type Material.</emphasis>
<materialsCitation id="34C9AC32B902785BC47C45B0FF0322DF" ID-GBIF-Occurrence="3962178301" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB902785BC47C45B0FE5222BB" box="[339,443,1951,1977]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB902785BC73045B1FDC422BA" bold="true" box="[543,557,1950,1976]" pageId="17" pageNumber="18"></emphasis>
<collectingCountry id="FCB6E6FFB902785BC70245B0FD4B22BB" box="[557,674,1951,1977]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
:
<quantity id="43590B8AB902785BC79B45B1FD1422BB" box="[692,765,1950,1977]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B902785BC62945B0FC5E22BB" box="[774,951,1951,1977]" pageId="17" pageNumber="18">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB902785BC13045B0FB6822BB" box="[1055,1153,1951,1977]" pageId="17" pageNumber="18" value="1982-05-01">
<collectingDate id="E05B7947B902785BC13045B0FB6822BB" box="[1055,1153,1951,1977]" pageId="17" pageNumber="18" value="1982-05-01">1.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B902785BC1A045B0FACC22BB" box="[1167,1317,1951,1977]" pageId="17" pageNumber="18">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB902785BC5B045ECFF0B22DF" box="[159,226,1987,2013]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B902785BC5DA45ECFD3322DF" ID-GBIF-Occurrence="3962178334" box="[245,730,1987,2013]" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="17" pageNumber="18" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB902785BC5DA45ECFE8F22DF" box="[245,358,1987,2013]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
: Same data as holotype (6
<collectionCode id="E2B03EAAB902785BC7BE45ECFD3A22DF" box="[657,723,1987,2013]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5E842B9FAB4241E" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="841EA66FB9017858C5E842B9FAB4241E" blockId="18.[151,1437,150,1076]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E842B9FE3825B2" bold="true" box="[199,465,150,176]" pageId="18" pageNumber="19">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9017858C12742B9FB9425B2" box="[1032,1149,150,176]" name="Cameroon" pageId="18" pageNumber="19">Cameroon</collectingCountry>
by the combination of the very narrow P2 (l/w ca. 4/1), the unusual ratios of P2 width and plaque shape (P2/w/l/s ca. 1/1/6/5), and the aedeagus (
<figureCitation id="1C9ABAEAB9017858C5B042F1FF1A25FA" box="[159,243,222,248]" captionStart="FIGURE 21" captionStartId="46.[151,250,1986,2010]" captionTargetBox="[253,1333,181,1963]" captionTargetId="figure-25@46.[253,1333,181,1963]" captionTargetPageId="46" captionText="FIGURE 21. Hydraena camerocompressa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284905" httpUri="https://zenodo.org/record/7284905/files/figure.png" pageId="18" pageNumber="19">Fig. 21</figureCitation>
). Although the P2 is very narrow, as is that of
<taxonomicName id="43A1DDECB9017858C7D842F0FC2225FA" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[759,971,222,248]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="camerocrebrata" status="sp. nov.">
<emphasis id="B6D57A7DB9017858C7D842F0FC2225FA" box="[759,971,222,248]" italics="true" pageId="18" pageNumber="19">H. camerocrebrata</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB9017858C6FD42F1FBF825FA" bold="true" box="[978,1041,222,248]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="ADE6C706B9017858C6FD42F1FBF825FA" box="[978,1041,222,248]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the habitus and aedeagi of the two species markedly differ.
<taxonomicName id="43A1DDECB9017858C485432CFDB6241E" authorityName="Perkins" authorityYear="2022" box="[426,607,258,284]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB9017858C485432CFDB6241E" box="[426,607,258,284]" italics="true" pageId="18" pageNumber="19">H. cameropetila</emphasis>
</taxonomicName>
(CAM13) also has a very narrow P2, but lacks metaventral plaques.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5E84309FBD726CA" pageId="18" pageNumber="19" type="description">
<paragraph id="841EA66FB9017858C5E84309FEC4273E" blockId="18.[151,1437,150,1076]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E84309FEB82442" bold="true" box="[199,337,294,320]" pageId="18" pageNumber="19">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9017858C48B4309FDEF2442" box="[420,518,294,320]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.40/0.61; head width 0.33; pronotum 0.33/0.48, PA 0.37, PB 0.39; elytra 0.83/0.61. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9017858C1074365FB972466" box="[1064,1150,330,356]" captionStart="FIGURE 21" captionStartId="46.[151,250,1986,2010]" captionTargetBox="[253,1333,181,1963]" captionTargetId="figure-25@46.[253,1333,181,1963]" captionTargetPageId="46" captionText="FIGURE 21. Hydraena camerocompressa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284905" httpUri="https://zenodo.org/record/7284905/files/figure.png" pageId="18" pageNumber="19">Fig. 21</figureCitation>
). Dorsum brown to light brown, head and pronotal macula darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae shallow, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB9017858C5E84069FE7427F2" blockId="18.[151,1437,150,1076]" pageId="18" pageNumber="19">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB9017858C5B840D4FBD726CA" blockId="18.[151,1437,150,1076]" pageId="18" pageNumber="19">Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/6/5. P1 laminate; median carina weakly sinuate in profile. P2 narrow, l/ w ca. 4/1, sides parallel, apex blunt. Plaques narrow, straight, parallel, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 1.5x P2. Pro- meso- and metatibiae slender, straight. Abdominal apex with deep asymmetrical slightly off-center notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5E841FDFC2126EE" box="[199,968,978,1005]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="841EA66FB9017858C5E841FDFC2126EE" blockId="18.[151,1437,150,1076]" box="[199,968,978,1005]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E841FDFE8826EE" bold="true" box="[199,353,978,1004]" pageId="18" pageNumber="19">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9017858C7E341FCFD1526EF" box="[716,764,979,1005]" pageId="18" pageNumber="19">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9017858C64841FDFC5226EE" box="[871,955,978,1004]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="18" pageNumber="19">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9017858C5E841D9FF262136" pageId="18" pageNumber="19" type="etymology">
<paragraph id="841EA66FB9017858C5E841D9FF262136" blockId="18.[151,1437,150,1076]" pageId="18" pageNumber="19">
<emphasis id="B6D57A7DB9017858C5E841D9FEA72112" bold="true" box="[199,334,1014,1040]" pageId="18" pageNumber="19">Etymology.</emphasis>
Named in reference to the known distribution and the very narrow mesoventral intercoxal process (P2).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,145 @@
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<mods:title id="8F0648E034E1F114AEA039A9790A47E8">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<subSubSection id="CCBBF5E4B902785BC5B840A1FD6A27AA" box="[151,643,654,681]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="841EA66FB902785BC5B840A1FD6A27AA" blockId="17.[151,662,654,717]" box="[151,643,654,681]" pageId="17" pageNumber="18">
<heading id="DF561103B902785BC5B840A1FD6A27AA" bold="true" box="[151,643,654,681]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<emphasis id="B6D57A7DB902785BC5B840A1FD6A27AA" bold="true" box="[151,643,654,681]" pageId="17" pageNumber="18">
<taxonomicName id="43A1DDECB902785BC5B840A1FE0D27AB" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,484,654,681]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="camerocompacta" status="sp. nov.">
<emphasis id="B6D57A7DB902785BC5B840A1FE0D27AB" bold="true" box="[151,484,654,681]" italics="true" pageId="17" pageNumber="18">Hydraena camerocompacta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B902785BC4DC40A0FD6A27AA" box="[499,643,654,681]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5B8409CFD7F27CF" box="[151,662,691,717]" pageId="17" pageNumber="18" type="description">
<paragraph id="841EA66FB902785BC5B8409CFD7F27CF" blockId="17.[151,662,654,717]" box="[151,662,691,717]" pageId="17" pageNumber="18">
<figureCitation id="1C9ABAEAB902785BC5B8409CFF0C27CF" box="[151,229,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="17" pageNumber="18">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB902785BC5DE409CFEE427CF" box="[241,269,691,717]" captionStart="FIGURE 20" captionStartId="45.[151,250,1993,2017]" captionTargetBox="[278,1316,193,1954]" captionTargetId="figure-15@45.[217,1370,181,1969]" captionTargetPageId="45" captionText="FIGURE 20. Hydraena camerocompacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284901" httpUri="https://zenodo.org/record/7284901/files/figure.png" pageId="17" pageNumber="18">20</figureCitation>
(habitus), 20 (aedeagus), 40 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5B840D4FD05263B" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="841EA66FB902785BC5B840D4FD05263B" blockId="17.[151,1437,762,1761]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5B840D4FEAE2616" bold="true" box="[151,327,762,789]" pageId="17" pageNumber="18">Type Material.</emphasis>
<materialsCitation id="34C9AC32B902785BC46240D4FF03263B" ID-GBIF-Occurrence="3962178312" collectingDate="1982-01-30" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB902785BC46240D4FE5C2617" box="[333,437,763,789]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB902785BC73440D4FD7F2617" box="[539,662,763,789]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B902785BC78F40D4FCCB2617" box="[672,802,763,789]" pageId="17" pageNumber="18">N. W. Prov.</collectorName>
,
<quantity id="43590B8AB902785BC60340D4FC8E2617" box="[812,871,763,789]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7 km</quantity>
.
<collectorName id="2954C3B9B902785BC65F40D4FC122617" box="[880,1019,763,789]" pageId="17" pageNumber="18">E. Bamenda</collectorName>
, stream,
<date id="F01F80AFB902785BC17340D4FB212617" box="[1116,1224,763,789]" pageId="17" pageNumber="18" value="1982-01-30">
<collectingDate id="E05B7947B902785BC17340D4FB212617" box="[1116,1224,763,789]" pageId="17" pageNumber="18" value="1982-01-30">30.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B902785BC02E40D4FA7A2617" box="[1281,1427,763,789]" pageId="17" pageNumber="18">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB902785BC5B04130FF0B263B" box="[159,226,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B902785BC5DA4130FD01263B" ID-GBIF-Occurrence="3962178305" box="[245,744,799,825]" collectingDate="1982-01-30" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="17" pageNumber="18" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB902785BC5DA4130FE8F263B" box="[245,358,799,825]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
: Same data as holotype (12
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E8416DFDF626EF" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="841EA66FB902785BC5E8416DFDF626EF" blockId="17.[151,1437,762,1761]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E8416DFE3D265E" bold="true" box="[199,468,834,860]" pageId="17" pageNumber="18">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB902785BC10B416CFB70265F" box="[1060,1177,835,861]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
by the combination of the relatively large body size (ca.
<quantity id="43590B8AB902785BC7334148FD6F2683" box="[540,646,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="18" unit="mm" value="1.6">1.60 mm</quantity>
), the broad body form, the antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, the wide lateral explanate margins of the elytra, and the aedeagus (
<figureCitation id="1C9ABAEAB902785BC01441A4FA7926A7" box="[1339,1424,907,933]" captionStart="FIGURE 20" captionStartId="45.[151,250,1993,2017]" captionTargetBox="[278,1316,193,1954]" captionTargetId="figure-15@45.[217,1370,181,1969]" captionTargetPageId="45" captionText="FIGURE 20. Hydraena camerocompacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284901" httpUri="https://zenodo.org/record/7284901/files/figure.png" pageId="17" pageNumber="18">Fig. 20</figureCitation>
). Somewhat similar to
<taxonomicName id="43A1DDECB902785BC4AB4180FDB226CA" authorityName="Perkins" authorityYear="2022" box="[388,603,943,968]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="camerocontinua">
<emphasis id="B6D57A7DB902785BC4AB4180FDB226CA" box="[388,603,943,968]" italics="true" pageId="17" pageNumber="18">H. camerocontinua</emphasis>
</taxonomicName>
in body size and form, but differing therefrom in denser dorsal punctation, ventral morphology, and aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E841D9FE67239B" pageId="17" pageNumber="18" type="description">
<paragraph id="841EA66FB902785BC5E841D9FB5D200F" blockId="17.[151,1437,762,1761]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E841D9FEB82112" bold="true" box="[199,337,1014,1040]" pageId="17" pageNumber="18">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB902785BC48B41D8FDEF2113" box="[420,518,1015,1041]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.59/0.71; head width 0.40; pronotum 0.40/0.55, PA 0.45, PB 0.48; elytra 0.93/0.71. Habitus as illustrated (
<figureCitation id="1C9ABAEAB902785BC1064634FB692137" box="[1065,1152,1051,1077]" captionStart="FIGURE 20" captionStartId="45.[151,250,1993,2017]" captionTargetBox="[278,1316,193,1954]" captionTargetId="figure-15@45.[217,1370,181,1969]" captionTargetPageId="45" captionText="FIGURE 20. Hydraena camerocompacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284901" httpUri="https://zenodo.org/record/7284901/files/figure.png" pageId="17" pageNumber="18">Fig. 20</figureCitation>
). Dorsum brown to dark brown or black, frons and diffusely margined pronotal disc darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately sparcely moderately coarsely punctulate, more sparsely punctulate on pronotal disc than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 12xef, interstices ca. 23xpd or slightly larger; punctures of clypeus very fine. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB902785BC5E84738FB64209F" blockId="17.[151,1437,762,1761]" pageId="17" pageNumber="18">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures moderately coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB902785BC5B84788FE67239B" blockId="17.[151,1437,762,1761]" pageId="17" pageNumber="18">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/3/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, very slightly arcuate, converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia slender, very slightly arcuate. Meso- and metatibiae slender, straight. Abdominal apex asymmetrical, with off-center deep notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E8448DFC2123BF" box="[199,968,1698,1725]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="841EA66FB902785BC5E8448DFC2123BF" blockId="17.[151,1437,762,1761]" box="[199,968,1698,1725]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E8448DFE8823BE" bold="true" box="[199,353,1698,1724]" pageId="17" pageNumber="18">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB902785BC7E3448CFD1523BF" box="[716,764,1699,1725]" pageId="17" pageNumber="18">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB902785BC648448CFC5223BF" box="[871,955,1699,1725]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E844E8FB5023E3" box="[199,1209,1735,1761]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="841EA66FB902785BC5E844E8FB5023E3" blockId="17.[151,1437,762,1761]" box="[199,1209,1735,1761]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E844E8FEA723E3" bold="true" box="[199,334,1735,1761]" pageId="17" pageNumber="18">Etymology.</emphasis>
Named in reference to the known distribution, and to the compact body form.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,169 @@
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<mods:title id="6B4F6ADF13265C2C10758FFAC82F6198">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B903785AC5B842B8FA6121EB" ID-DOI="http://doi.org/10.5281/zenodo.7300196" ID-GBIF-Taxon="204205230" ID-Zenodo-Dep="7300196" LSID="urn:lsid:plazi:treatment:0C081779B903785AC5B842B8FA6121EB" httpUri="http://treatment.plazi.org/id/0C081779B903785AC5B842B8FA6121EB" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="CCBBF5E4B903785AC5B842B8FDBB25B3" box="[151,594,151,178]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="841EA66FB903785AC5B842B8FDBB25B3" blockId="16.[151,662,151,213]" box="[151,594,151,178]" pageId="16" pageNumber="17">
<heading id="DF561103B903785AC5B842B8FDBB25B3" bold="true" box="[151,594,151,178]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<emphasis id="B6D57A7DB903785AC5B842B8FDBB25B3" bold="true" box="[151,594,151,178]" pageId="16" pageNumber="17">
<taxonomicName id="43A1DDECB903785AC5B842B8FE5A25B0" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,435,151,178]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cameropetila" status="sp. nov.">
<emphasis id="B6D57A7DB903785AC5B842B8FE5A25B0" bold="true" box="[151,435,151,178]" italics="true" pageId="16" pageNumber="17">Hydraena cameropetila</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B903785AC4ED42B8FDBB25B3" box="[450,594,151,177]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5B84294FD7F25D7" box="[151,662,187,213]" pageId="16" pageNumber="17" type="description">
<paragraph id="841EA66FB903785AC5B84294FD7F25D7" blockId="16.[151,662,151,213]" box="[151,662,187,213]" pageId="16" pageNumber="17">
<figureCitation id="1C9ABAEAB903785AC5B84294FF0D25D7" box="[151,228,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="16" pageNumber="17">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB903785AC5DE4294FEE425D7" box="[241,269,187,213]" captionStart="FIGURE 18" captionStartId="43.[151,250,1945,1969]" captionTargetBox="[206,1394,202,1889]" captionTargetId="figure-15@43.[156,1431,181,1921]" captionTargetPageId="43" captionText="FIGURE 18. Hydraena cameropetila, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284894" httpUri="https://zenodo.org/record/7284894/files/figure.png" pageId="16" pageNumber="17">18</figureCitation>
(habitus), 18 (aedeagus), 43 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5B8432CFEFB2467" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="841EA66FB903785AC5B8432CFEFB2467" blockId="16.[151,1437,259,1258]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5B8432CFEAF241F" bold="true" box="[151,326,259,285]" pageId="16" pageNumber="17">Type Material.</emphasis>
<materialsCitation id="34C9AC32B903785AC464432CFC1A2443" ID-GBIF-Occurrence="3962178341" collectingDate="1949-01-07" collectingDateMax="1949-01-11" collectingDateMin="1949-01-07" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB903785AC464432CFE5A241F" box="[331,435,259,285]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(male):
<location id="817EF0B4B903785AC721432CFD6E241F" LSID="urn:lsid:plazi:treatment:0C081779B903785AC5B842B8FA6121EB:817EF0B4B903785AC721432CFD6E241F" box="[526,647,259,285]" country="Cameroon" municipality="Northwest Province" name="Cameroon" pageId="16" pageNumber="17">Cameroon</location>
:
<collectingMunicipality id="647A3C15B903785AC7A0432CFC99241F" box="[655,880,259,285]" pageId="16" pageNumber="17">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB903785AC6AD432CFB42241F" box="[898,1195,259,285]" name="Cameroon" pageId="16" pageNumber="17">BRITISH CAMEROONS</collectingCountry>
. / Mamfe,
<date id="F01F80AFB903785AC033432CFA75241F" box="[1308,1436,259,285]" pageId="16" pageNumber="17" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">
<collectingDate id="E05B7947B903785AC033432CFA75241F" box="[1308,1436,259,285]" pageId="16" pageNumber="17" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">7-11.i.1949</collectingDate>
</date>
/
<collectorName id="2954C3B9B903785AC5874307FEF22443" box="[168,283,295,322]" pageId="16" pageNumber="17">B. Malkin</collectorName>
coll. // Rain forest; clear stream / Gravel and sand” (
<collectionCode id="E2B03EAAB903785AC6544308FC052443" box="[891,1004,295,321]" pageId="16" pageNumber="17">UKNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B903785AC6D04307FEE72467" ID-GBIF-Occurrence="3962178346" collectingDate="1949-01-07" collectingDateMax="1949-01-11" collectingDateMin="1949-01-07" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="16" pageNumber="17" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB903785AC6D04307FB992440" box="[1023,1136,296,322]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
: Same data as holotype (2
<collectionCode id="E2B03EAAB903785AC5B84364FEEF2467" box="[151,262,331,357]" pageId="16" pageNumber="17">UKNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5E84340FD3D273C" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="841EA66FB903785AC5E84340FD3D273C" blockId="16.[151,1437,259,1258]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E84340FE3D248B" bold="true" box="[199,468,367,393]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB903785AC10B4340FB70248B" box="[1060,1177,367,393]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
by the combination of absence of metaventral plaques, the very narrow P2, and the aedeagus (
<figureCitation id="1C9ABAEAB903785AC6EE43BCFBFC24AF" box="[961,1045,403,429]" captionStart="FIGURE 18" captionStartId="43.[151,250,1945,1969]" captionTargetBox="[206,1394,202,1889]" captionTargetId="figure-15@43.[156,1431,181,1921]" captionTargetPageId="43" captionText="FIGURE 18. Hydraena cameropetila, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284894" httpUri="https://zenodo.org/record/7284894/files/figure.png" pageId="16" pageNumber="17">Fig. 18</figureCitation>
). For characters differentiating the two species refer to the diagnosis of
<emphasis id="B6D57A7DB903785AC7164397FD1524D3" box="[569,764,440,465]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="43A1DDECB903785AC7164397FD1024D3" authorityName="Perkins" authorityYear="2022" box="[569,761,440,465]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="camerosetosa">H. camerosetosa</taxonomicName>
.
</emphasis>
Although
<taxonomicName id="43A1DDECB903785AC65B4397FBC224D3" authorityName="Perkins" authorityYear="2022" box="[884,1067,439,465]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB903785AC65B4397FBC224D3" box="[884,1067,439,465]" italics="true" pageId="16" pageNumber="17">H. cameropetila</emphasis>
</taxonomicName>
is smaller than
<taxonomicName id="43A1DDECB903785AC1CF4397FA7524D3" authorityName="Perkins" authorityYear="2022" box="[1248,1436,440,465]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB903785AC1CF4397FA7524D3" box="[1248,1436,440,465]" italics="true" pageId="16" pageNumber="17">H. camerosetosa</emphasis>
</taxonomicName>
in body length (ca. 1.20 vs.
<quantity id="43590B8AB903785AC4E243F4FDDD24F7" box="[461,564,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="16" pageNumber="17" unit="mm" value="1.33">1.33 mm</quantity>
), the width of the mesocoxae are the same in the two species, making the ratio of P2 width/mesocoxa width approximately 0.12 for
<taxonomicName id="43A1DDECB903785AC7DF402FFC4F271B" authorityName="Perkins" authorityYear="2022" box="[752,934,511,537]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB903785AC7DF402FFC4F271B" box="[752,934,511,537]" italics="true" pageId="16" pageNumber="17">H. cameropetila</emphasis>
</taxonomicName>
and 0.40
<taxonomicName id="43A1DDECB903785AC137402FFB3D271B" authorityName="Perkins" authorityYear="2022" box="[1048,1236,512,537]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB903785AC137402FFB3D271B" box="[1048,1236,512,537]" italics="true" pageId="16" pageNumber="17">H. camerosetosa</emphasis>
</taxonomicName>
. Also refer to the diagnoses of
<taxonomicName id="43A1DDECB903785AC404400BFE2B273F" authorityName="Perkins" authorityYear="2022" box="[299,450,547,573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="millenaria">
<emphasis id="B6D57A7DB903785AC404400BFE2B273F" box="[299,450,547,573]" italics="true" pageId="16" pageNumber="17">H. millenaria</emphasis>
</taxonomicName>
and
<taxonomicName id="43A1DDECB903785AC4D7400BFD25273F" authorityName="Perkins" authorityYear="2022" box="[504,716,547,573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB903785AC4D7400BFD25273F" box="[504,716,547,573]" italics="true" pageId="16" pageNumber="17">H. camerocrebrata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5E84068FE0B21A3" pageId="16" pageNumber="17" type="description">
<paragraph id="841EA66FB903785AC5E84068FC85263B" blockId="16.[151,1437,259,1258]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E84068FEB82763" bold="true" box="[199,337,583,609]" pageId="16" pageNumber="17">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB903785AC48B4068FDEF2763" box="[420,518,583,609]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.20/0.50; head width 0.30; pronotum 0.29/0.42, PA 0.32, PB 0.36; elytra 0.74/0.50. Habitus as illustrated (
<figureCitation id="1C9ABAEAB903785AC1064044FB962787" box="[1065,1151,619,645]" captionStart="FIGURE 18" captionStartId="43.[151,250,1945,1969]" captionTargetBox="[206,1394,202,1889]" captionTargetId="figure-15@43.[156,1431,181,1921]" captionTargetPageId="43" captionText="FIGURE 18. Hydraena cameropetila, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284894" httpUri="https://zenodo.org/record/7284894/files/figure.png" pageId="16" pageNumber="17">Fig. 18</figureCitation>
). Dorsum dark brown to black, pronotum without macula, palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 23xef, interstices ca. 1xpd or less; punctures of clypeus moderately fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep, anteromedian fovea shallow.
</paragraph>
<paragraph id="841EA66FB903785AC5E8416CFB5A26CB" blockId="16.[151,1437,259,1258]" pageId="16" pageNumber="17">Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 0.5xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming weak angle with one another.</paragraph>
<paragraph id="841EA66FB903785AC5B841FBFAB126EF" blockId="16.[151,1437,259,1258]" box="[151,1368,979,1006]" pageId="16" pageNumber="17">Venter: Mentum and postmentum densely finely punctulate, dull. Genae raised, dull, without posterior ridge.</paragraph>
<paragraph id="841EA66FB903785AC5B841D8FE0B21A3" blockId="16.[151,1437,259,1258]" pageId="16" pageNumber="17">Metaventral plaques absent. P1 laminate; median carina sinuate in profile. P2 very narrow, l/w ca. 4/1, sides parallel, blunt apex raised slightly above mesoventral intercoxal process.. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 4x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with very small indistinct apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5E84684FC2121C7" box="[199,968,1195,1222]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="841EA66FB903785AC5E84684FC2121C7" blockId="16.[151,1437,259,1258]" box="[199,968,1195,1222]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E84684FE8821C7" bold="true" box="[199,353,1195,1221]" pageId="16" pageNumber="17">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB903785AC7E34683FD1521C4" box="[716,764,1196,1222]" pageId="16" pageNumber="17">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB903785AC6484684FC5221C7" box="[871,955,1195,1221]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="16" pageNumber="17">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B903785AC5E846E0FA6121EB" box="[199,1416,1231,1258]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="841EA66FB903785AC5E846E0FA6121EB" blockId="16.[151,1437,259,1258]" box="[199,1416,1231,1258]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E846E0FEA721EB" bold="true" box="[199,334,1231,1257]" pageId="16" pageNumber="17">Etymology.</emphasis>
Named in reference to the distribution and the very narrow mesoventral intercoxal process (P2).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="6564CDC32780EADABB70B81C9F7A70DD">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<emphasis id="B6D57A7DB903785AC5B847ACFE2C209C" bold="true" box="[151,453,1411,1438]" italics="true" pageId="16" pageNumber="17">Hydraena cameroancylis</emphasis>
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,
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<paragraph id="841EA66FB903785AC5B84788FD7F20C3" blockId="16.[151,662,1411,1473]" box="[151,662,1447,1473]" pageId="16" pageNumber="17">
<figureCitation id="1C9ABAEAB903785AC5B84788FF0C20C3" box="[151,229,1447,1473]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="16" pageNumber="17">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB903785AC5DE4788FEE420C3" box="[241,269,1447,1473]" captionStart="FIGURE 19" captionStartId="44.[151,250,1981,2005]" captionTargetBox="[244,1274,202,1932]" captionTargetId="figure-25@44.[216,1372,181,1958]" captionTargetPageId="44" captionText="FIGURE 19. Hydraena cameroancylis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284898" httpUri="https://zenodo.org/record/7284898/files/figure.png" pageId="16" pageNumber="17">19</figureCitation>
(habitus), 19 (aedeagus), 41 (map)
</paragraph>
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<paragraph id="841EA66FB903785AC5B847C0FC41232F" blockId="16.[151,1437,1519,2014]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5B847C0FEA0230B" bold="true" box="[151,329,1519,1545]" pageId="16" pageNumber="17">Type Material.</emphasis>
<materialsCitation id="34C9AC32B903785AC47E47C0FE57232F" ID-GBIF-Occurrence="3962178347" collectingDate="1983-01-02" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Isali River" municipality="Nouma" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB903785AC47E47C0FE50230B" box="[337,441,1519,1545]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB903785AC70A47C0FD49230B" box="[549,672,1519,1545]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
:
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,
<quantity id="43590B8AB903785AC61947C0FC68230B" box="[822,897,1519,1546]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="km" value="15.0">15 km</quantity>
. NW Yaounde,
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nr.
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,
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<collectingDate id="E05B7947B903785AC01647C0FA71230B" box="[1337,1432,1519,1545]" pageId="16" pageNumber="17" value="1983-01-02">2.i.1983</collectingDate>
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, leg.
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” (
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)
</materialsCitation>
.
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<typeStatus id="5B1A18CDB903785AC4E6443BFDC6232C" box="[457,559,1556,1582]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
: Same data as holotype (1
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)
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.
</paragraph>
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<paragraph id="841EA66FB903785AC5E84418FD1323BF" blockId="16.[151,1437,1519,2014]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E84418FE3D2353" bold="true" box="[199,468,1591,1617]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
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by the combination of the anterolateral pronotal foveae large and deep, posterolateral pronotal foveae absent, the brown to dark brown dorsum, and the aedeagus (
<figureCitation id="1C9ABAEAB903785AC4E54450FDF7239B" box="[458,542,1663,1689]" captionStart="FIGURE 19" captionStartId="44.[151,250,1981,2005]" captionTargetBox="[244,1274,202,1932]" captionTargetId="figure-25@44.[216,1372,181,1958]" captionTargetPageId="44" captionText="FIGURE 19. Hydraena cameroancylis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284898" httpUri="https://zenodo.org/record/7284898/files/figure.png" pageId="16" pageNumber="17">Fig. 19</figureCitation>
). Similar in dorsal habitus to
<taxonomicName id="43A1DDECB903785AC64244AFFBD6239B" authorityName="Perkins" authorityYear="2022" box="[877,1087,1663,1689]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cameropalmula">
<emphasis id="B6D57A7DB903785AC64244AFFBD6239B" box="[877,1087,1663,1689]" italics="true" pageId="16" pageNumber="17">H. cameropalmula</emphasis>
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and
<taxonomicName id="43A1DDECB903785AC15944AFFAE7239B" authorityName="Perkins" authorityYear="2022" box="[1142,1294,1663,1689]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="millenaria">
<emphasis id="B6D57A7DB903785AC15944AFFAE7239B" box="[1142,1294,1663,1689]" italics="true" pageId="16" pageNumber="17">H. millenaria</emphasis>
</taxonomicName>
; reliable determinations will require examination of the aedeagus.
</paragraph>
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<subSubSection id="CCBBF5E4B903785BC5E844E8FBE024F7" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="841EA66FB903785AC5E844E8FB1122DF" blockId="16.[151,1437,1519,2014]" pageId="16" pageNumber="17">
<emphasis id="B6D57A7DB903785AC5E844E8FEB823E3" bold="true" box="[199,337,1735,1761]" pageId="16" pageNumber="17">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB903785AC48B44E8FDEF23E3" box="[420,518,1735,1761]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.35/0.60; head width 0.36; pronotum 0.33/0.48, PA 0.40, PB 0.43; elytra 0.85/0.60. Habitus as illustrated (
<figureCitation id="1C9ABAEAB903785AC10644C4FB962207" box="[1065,1151,1771,1797]" captionStart="FIGURE 19" captionStartId="44.[151,250,1981,2005]" captionTargetBox="[244,1274,202,1932]" captionTargetId="figure-25@44.[216,1372,181,1958]" captionTargetPageId="44" captionText="FIGURE 19. Hydraena cameroancylis, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284898" httpUri="https://zenodo.org/record/7284898/files/figure.png" pageId="16" pageNumber="17">Fig. 19</figureCitation>
). Dorsum brown to dark brown, frons and diffusely margined pronotal disc darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately sparcely moderately coarsely punctulate, more sparsely punctulate on pronotal disc than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 12xef, interstices ca. 12xpd or slightly larger; punctures of clypeus very fine, very dense, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae large and deep, posterolateral pronotal foveae absent, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB902785BC5E842B8FB3B241F" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="18">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures moderately coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderate angle with one another.</paragraph>
<paragraph id="841EA66FB902785BC5B84308FBE024F7" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="18">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2.5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, short, slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. All tibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E843D1FC21271B" box="[199,968,510,537]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="841EA66FB902785BC5E843D1FC21271B" blockId="17.[151,1436,151,573]" box="[199,968,510,537]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E843D1FE88271A" bold="true" box="[199,353,510,536]" pageId="17" pageNumber="18">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB902785BC7E343D0FD15271B" box="[716,764,511,537]" pageId="17" pageNumber="18">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB902785BC64843D0FC53271B" box="[871,954,511,537]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B902785BC5E8400CFA7E273F" box="[199,1431,547,573]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="841EA66FB902785BC5E8400CFA7E273F" blockId="17.[151,1436,151,573]" box="[199,1431,547,573]" pageId="17" pageNumber="18">
<emphasis id="B6D57A7DB902785BC5E8400CFEA0273F" bold="true" box="[199,329,547,573]" pageId="17" pageNumber="18">Etymology.</emphasis>
Named in reference to the known distribution, and to the strong hook-shaped process of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="1E66AC42D32DCE9FA92CA05B05CE5427">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<heading id="DF561103B904785DC5B8448DFD9D23BE" bold="true" box="[151,628,1698,1725]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="B6D57A7DB904785DC5B8448DFD9D23BE" bold="true" box="[151,628,1698,1725]" pageId="23" pageNumber="24">
<taxonomicName id="43A1DDECB904785DC5B8448DFE3F23BF" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,470,1698,1725]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="camerocameola" status="sp. nov.">
<emphasis id="B6D57A7DB904785DC5B8448DFE3F23BF" bold="true" box="[151,470,1698,1725]" italics="true" pageId="23" pageNumber="24">Hydraena camerocameola</emphasis>
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,
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<paragraph id="841EA66FB904785DC5B844E8FD7F23E3" blockId="23.[151,662,1698,1761]" box="[151,662,1734,1761]" pageId="23" pageNumber="24">
<figureCitation id="1C9ABAEAB904785DC5B844E8FF0C23E3" box="[151,229,1734,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="23" pageNumber="24">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB904785DC5DE44E8FEE423E3" box="[241,269,1735,1761]" captionStart="FIGURE 30" captionStartId="55.[151,250,2006,2030]" captionTargetBox="[287,1377,191,1959]" captionTargetId="figure-15@55.[198,1390,181,1982]" captionTargetPageId="55" captionText="FIGURE 30. Hydraena camerocameola, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284935" httpUri="https://zenodo.org/record/7284935/files/figure.png" pageId="23" pageNumber="24">30</figureCitation>
(habitus), 30 (aedeagus), 39 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5B84520FA9C224F" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="841EA66FB904785DC5B84520FA9C224F" blockId="23.[151,1437,1806,2013]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5B84520FEA1222A" bold="true" box="[151,328,1806,1833]" pageId="23" pageNumber="24">Type Material.</emphasis>
<materialsCitation id="34C9AC32B904785DC4614520FEC5224F" ID-GBIF-Occurrence="3962178308" collectingDate="1982-01-14" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB904785DC4614520FE5F222B" box="[334,438,1807,1833]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB904785DC73B4521FDCB222A" bold="true" box="[532,546,1806,1832]" pageId="23" pageNumber="24"></emphasis>
<collectingCountry id="FCB6E6FFB904785DC70D4520FD72222B" box="[546,667,1807,1833]" name="Cameroon" pageId="23" pageNumber="24">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B904785DC78A4520FCEC222B" box="[677,773,1807,1833]" pageId="23" pageNumber="24">N. Prov.</collectorName>
,
<quantity id="43590B8AB904785DC6204520FCB0222B" box="[783,857,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="km" value="22.0">22 km</quantity>
.
<collectorName id="2954C3B9B904785DC64C4520FC36222B" box="[867,991,1807,1833]" pageId="23" pageNumber="24">N. Maroua</collectorName>
, desert stream,
<date id="F01F80AFB904785DC1A44520FB1E222B" box="[1163,1271,1807,1833]" pageId="23" pageNumber="24" value="1982-01-14">
<collectingDate id="E05B7947B904785DC1A44520FB1E222B" box="[1163,1271,1807,1833]" pageId="23" pageNumber="24" value="1982-01-14">14.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B904785DC01D4520FF21224F" pageId="23" pageNumber="24">P. D. Perkins</collectorName>
” (
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)
</materialsCitation>
.
<materialsCitation id="34C9AC32B904785DC418451CFA9C224F" ID-GBIF-Occurrence="3962178348" box="[311,1397,1842,1869]" collectingDate="1982-05-27" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="23" pageNumber="24" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB904785DC418451CFE41224F" box="[311,424,1843,1869]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
: Same data as holotype (9
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); Same locality,
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<collectingDate id="E05B7947B904785DC6E2451CFBD7224F" box="[973,1086,1843,1869]" pageId="23" pageNumber="24" value="1982-05-27">27.v.1982</collectingDate>
</date>
, leg.
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(5
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)
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</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E84579FEE922DF" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="841EA66FB904785DC5E84579FEE922DF" blockId="23.[151,1437,1806,2013]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E84579FE382272" bold="true" box="[199,465,1878,1904]" pageId="23" pageNumber="24">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB904785DC1274578FB942273" box="[1032,1149,1879,1905]" name="Cameroon" pageId="23" pageNumber="24">Cameroon</collectingCountry>
by the combination of the rather elongate body form, the very coarse and very dense pronotal and elytral punctures, interstices narrow walls, shining, the narrow, long plaques, located at sides of deep longitudinal groove-like median sulcus, and the aedeagus (
<figureCitation id="1C9ABAEAB904785DC5B045ECFF1D22DF" box="[159,244,1987,2013]" captionStart="FIGURE 30" captionStartId="55.[151,250,2006,2030]" captionTargetBox="[287,1377,191,1959]" captionTargetId="figure-15@55.[198,1390,181,1982]" captionTargetPageId="55" captionText="FIGURE 30. Hydraena camerocameola, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284935" httpUri="https://zenodo.org/record/7284935/files/figure.png" pageId="23" pageNumber="24">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90B7852C5E842B9FC2A265E" pageId="24" pageNumber="25" type="description">
<paragraph id="841EA66FB90B7852C5E842B9FA8924D3" blockId="24.[151,1437,150,933]" pageId="24" pageNumber="25">
<emphasis id="B6D57A7DB90B7852C5E842B9FEB825B2" bold="true" box="[199,337,150,176]" pageId="24" pageNumber="25">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB90B7852C4B342B9FE1725B2" box="[412,510,150,176]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.48/0.60; head width 0.36; pronotum 0.34/0.44, PA 0.40, PB 0.42; elytra 0.92/0.60. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90B7852C6C54295FBD625D6" box="[1002,1087,186,212]" captionStart="FIGURE 30" captionStartId="55.[151,250,2006,2030]" captionTargetBox="[287,1377,191,1959]" captionTargetId="figure-15@55.[198,1390,181,1982]" captionTargetPageId="55" captionText="FIGURE 30. Hydraena camerocameola, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284935" httpUri="https://zenodo.org/record/7284935/files/figure.png" pageId="24" pageNumber="25">Fig. 30</figureCitation>
). Body form elongate. Dorsum generally brown to dark brown, maxillary palpi, testaceous, pronotum with diffusely margined weakly defined dark brown macula. Dorsum of head and pronotum coarsely, very densely punctulate. Frons punctures distinctly smaller and less dense than pronotal punctures, interstices shining; punctures of frons disc and ca. 23xef, interstices ca. 12xpd. Pronotal punctures very coarse and very dense, interstices narrow walls, shining. Clypeus very finely sparsely punctulate in median 1/3, shining; lateral areas densely micropunctulate, dull. Labrum apicomedially excised, free margins rounded, very weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral foveae deep, confluent, posterosubmedial foveae deep, very coarsely punctate.
</paragraph>
<paragraph id="841EA66FB90B7852C5E843F5FDC02762" blockId="24.[151,1437,150,933]" pageId="24" pageNumber="25">Elytra with summit of posterior declivity at about midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures. Intervals not raised, width ca. 0.5xpd or less, sometimes narrow walls, as are interstices between punctures of a row. Apices in dorsal aspect jointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB90B7852C5B84044FC2A265E" blockId="24.[151,1437,150,933]" pageId="24" pageNumber="25">Venter: Mentum and postmentum rather coarsely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2.3/1.3. P1 laminate; median carina sinuate in profile. P2 narrow, l/w ca. 3/1, sides slightly converging toward blunt apex raised slightly above mesoventral intercoxal process. Plaques narrow, long compared to P2, straight, slightly converging toward one another anteriorly, weakly raised, located at sides of deep longitudinal groove-like median sulcus. AIS width at straight posterior margin ca. 2x P2. Protibia moderately arcuate, slightly widened on inner margin over distal 1/3. Meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90B7852C5E84149FC212682" box="[199,968,870,897]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="841EA66FB90B7852C5E84149FC212682" blockId="24.[151,1437,150,933]" box="[199,968,870,897]" pageId="24" pageNumber="25">
<emphasis id="B6D57A7DB90B7852C5E84149FE882682" bold="true" box="[199,353,870,896]" pageId="24" pageNumber="25">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB90B7852C7E34148FD152683" box="[716,764,871,897]" pageId="24" pageNumber="25">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90B7852C6484149FC522682" box="[871,955,870,896]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="24" pageNumber="25">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90B7852C5E841A5FB2126A6" box="[199,1224,906,933]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="841EA66FB90B7852C5E841A5FB2126A6" blockId="24.[151,1437,150,933]" box="[199,1224,906,933]" pageId="24" pageNumber="25">
<emphasis id="B6D57A7DB90B7852C5E841A5FEA726A6" bold="true" box="[199,334,906,932]" pageId="24" pageNumber="25">Etymology.</emphasis>
Named in reference to the known distribution, and to the cameo-like aedeagus.
</paragraph>
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</treatment>
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@ -0,0 +1,137 @@
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<subSubSection id="CCBBF5E4B904785DC5B84045FD8B2786" box="[151,610,618,645]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="841EA66FB904785DC5B84045FD8B2786" blockId="23.[151,662,618,681]" box="[151,610,618,645]" pageId="23" pageNumber="24">
<heading id="DF561103B904785DC5B84045FD8B2786" bold="true" box="[151,610,618,645]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="B6D57A7DB904785DC5B84045FD8B2786" bold="true" box="[151,610,618,645]" pageId="23" pageNumber="24">
<taxonomicName id="43A1DDECB904785DC5B84045FE2D2787" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,452,618,645]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="camerobuloba" status="sp. nov.">
<emphasis id="B6D57A7DB904785DC5B84045FE2D2787" bold="true" box="[151,452,618,645]" italics="true" pageId="23" pageNumber="24">Hydraena camerobuloba</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B904785DC4FC4044FD8B2786" box="[467,610,618,645]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5B840A0FD7F27AB" box="[151,662,654,681]" pageId="23" pageNumber="24" type="description">
<paragraph id="841EA66FB904785DC5B840A0FD7F27AB" blockId="23.[151,662,618,681]" box="[151,662,654,681]" pageId="23" pageNumber="24">
<figureCitation id="1C9ABAEAB904785DC5B840A0FF0C27AB" box="[151,229,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="23" pageNumber="24">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB904785DC5DE40A0FEE427AB" box="[241,269,655,681]" captionStart="FIGURE 29" captionStartId="54.[151,250,1990,2014]" captionTargetBox="[230,1365,191,1947]" captionTargetId="figure-25@54.[169,1418,181,1966]" captionTargetPageId="54" captionText="FIGURE 29. Hydraena camerobuloba, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284931" httpUri="https://zenodo.org/record/7284931/files/figure.png" pageId="23" pageNumber="24">29</figureCitation>
(habitus), 29 (aedeagus), 43 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5B840F8FB662617" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="841EA66FB904785DC5B840F8FB662617" blockId="23.[151,1437,726,1617]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5B840F8FEAE27F2" bold="true" box="[151,327,726,753]" pageId="23" pageNumber="24">Type Material.</emphasis>
<materialsCitation id="34C9AC32B904785DC46340F8FD892617" ID-GBIF-Occurrence="3962178306" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB904785DC46340F8FE5D27F3" box="[332,436,727,753]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
(male):
<location id="817EF0B4B904785DC73E40F8FD6327F3" LSID="urn:lsid:plazi:treatment:0C081779B904785DC5B84045FAD62353:817EF0B4B904785DC73E40F8FD6327F3" box="[529,650,727,753]" country="Cameroon" municipality="Northwest Province" name="Cameroon" pageId="23" pageNumber="24">Cameroon</location>
:
<collectingMunicipality id="647A3C15B904785DC7BC40F8FC9D27F3" box="[659,884,727,753]" pageId="23" pageNumber="24">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB904785DC6A840F8FB4527F3" box="[903,1196,727,753]" name="Cameroon" pageId="23" pageNumber="24">BRITISH CAMEROONS</collectingCountry>
/ Matute, / Tiko Plantation /
<date id="F01F80AFB904785DC5C240D5FEA62617" box="[237,335,762,789]" pageId="23" pageNumber="24" value="1949-05-05">
<collectingDate id="E05B7947B904785DC5C240D5FEA62617" box="[237,335,762,789]" pageId="23" pageNumber="24" value="1949-05-05">5.v.1949</collectingDate>
</date>
,
<collectorName id="2954C3B9B904785DC47440D4FE262617" box="[347,463,763,789]" pageId="23" pageNumber="24">B. Malkin</collectorName>
” (
<collectionCode id="E2B03EAAB904785DC4C740D4FDB02617" box="[488,601,763,789]" pageId="23" pageNumber="24">UKNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B904785DC74540D4FB652617" ID-GBIF-Occurrence="3962178357" box="[618,1164,763,789]" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="23" pageNumber="24" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB904785DC74540D4FD322617" box="[618,731,763,789]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
: Same data as holotype (48
<collectionCode id="E2B03EAAB904785DC13B40D4FB6A2617" box="[1044,1155,763,789]" pageId="23" pageNumber="24">UKNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E84131FA702683" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="841EA66FB904785DC5E84131FA702683" blockId="23.[151,1437,726,1617]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E84131FE23263A" bold="true" box="[199,458,798,824]" pageId="23" pageNumber="24">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB904785DC1284130FB9E263B" box="[1031,1143,799,825]" name="Cameroon" pageId="23" pageNumber="24">Cameroon</collectingCountry>
by the combination of the pronotum color, mostly black, contrasting with light brown to testaceous color narrowly along anterior and posterior margins, the elytral margins in posterior aspect forming very strong angle with one another, and the aedeagus (
<figureCitation id="1C9ABAEAB904785DC0134148FA672683" box="[1340,1422,871,897]" captionStart="FIGURE 29" captionStartId="54.[151,250,1990,2014]" captionTargetBox="[230,1365,191,1947]" captionTargetId="figure-25@54.[169,1418,181,1966]" captionTargetPageId="54" captionText="FIGURE 29. Hydraena camerobuloba, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284931" httpUri="https://zenodo.org/record/7284931/files/figure.png" pageId="23" pageNumber="24">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E841A5FE86230B" pageId="23" pageNumber="24" type="description">
<paragraph id="841EA66FB904785DC5E841A5FB49217F" blockId="23.[151,1437,726,1617]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E841A5FEB826A6" bold="true" box="[199,337,906,932]" pageId="23" pageNumber="24">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB904785DC48B41A4FDEF26A7" box="[420,518,907,933]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.25/0.54; head width 0.33; pronotum 0.30/0.43, PA 0.38, PB 0.39; elytra 0.75/0.54. Habitus as illustrated (
<figureCitation id="1C9ABAEAB904785DC6D04180FBBA26CB" box="[1023,1107,943,969]" captionStart="FIGURE 29" captionStartId="54.[151,250,1990,2014]" captionTargetBox="[230,1365,191,1947]" captionTargetId="figure-25@54.[169,1418,181,1966]" captionTargetPageId="54" captionText="FIGURE 29. Hydraena camerobuloba, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284931" httpUri="https://zenodo.org/record/7284931/files/figure.png" pageId="23" pageNumber="24">Fig. 29</figureCitation>
). Dorsum with head and most of pronotum black, pronotum narrowly along anterior and posterior margins light brown to testaceous, palpi testaceous. Frons and clypeus finely, moderately densely punctulate, interstices shining. Pronotum moderately coarsely, densely punctulate, interstices shining; punctures of pronotal disc ca. 23xef, interstices ca. 1xpd or less. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep.
</paragraph>
<paragraph id="841EA66FB904785DC5E846A8FAAA200F" blockId="23.[151,1437,726,1617]" pageId="23" pageNumber="24">Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect slightly separately rounded, in posterior aspect margins forming very strong angle with one another.</paragraph>
<paragraph id="841EA66FB904785DC5B84738FE86230B" blockId="23.[151,1437,726,1617]" pageId="23" pageNumber="24">Venter: Mentum and postmentum moderately finely punctulate, shining. Genae raised, dull, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, straight, parallel or nearly so, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E8443DFC21232F" box="[199,968,1554,1581]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="841EA66FB904785DC5E8443DFC21232F" blockId="23.[151,1437,726,1617]" box="[199,968,1554,1581]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E8443DFE88232E" bold="true" box="[199,353,1554,1580]" pageId="23" pageNumber="24">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB904785DC7E3443CFD15232F" box="[716,764,1555,1581]" pageId="23" pageNumber="24">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB904785DC648443CFC52232F" box="[871,955,1555,1581]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="23" pageNumber="24">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E84418FAD62353" box="[199,1343,1591,1617]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="841EA66FB904785DC5E84418FAD62353" blockId="23.[151,1437,726,1617]" box="[199,1343,1591,1617]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E84418FEA72353" bold="true" box="[199,334,1591,1617]" pageId="23" pageNumber="24">Etymology.</emphasis>
Named in reference to the distribution and the relatively large distal lobe of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,135 @@
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<mods:title id="F0B686ECFF32E6E83013941B77BE8EF0">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B905785DC5B84715FBD6271B" ID-DOI="http://doi.org/10.5281/zenodo.7300206" ID-GBIF-Taxon="204205226" ID-Zenodo-Dep="7300206" LSID="urn:lsid:plazi:treatment:0C081779B905785DC5B84715FBD6271B" httpUri="http://treatment.plazi.org/id/0C081779B905785DC5B84715FBD6271B" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="CCBBF5E4B905785CC5B84715FDA62056" box="[151,591,1338,1365]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="841EA66FB905785CC5B84715FDA62056" blockId="22.[151,662,1338,1401]" box="[151,591,1338,1365]" pageId="22" pageNumber="23">
<heading id="DF561103B905785CC5B84715FDA62056" bold="true" box="[151,591,1338,1365]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
<emphasis id="B6D57A7DB905785CC5B84715FDA62056" bold="true" box="[151,591,1338,1365]" pageId="22" pageNumber="23">
<taxonomicName id="43A1DDECB905785CC5B84715FE592057" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,432,1338,1365]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cameroarida" status="sp. nov.">
<emphasis id="B6D57A7DB905785CC5B84715FE592057" bold="true" box="[151,432,1338,1365]" italics="true" pageId="22" pageNumber="23">Hydraena cameroarida</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B905785CC4904715FDA62056" box="[447,591,1338,1364]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5B84771FD7F207A" box="[151,662,1374,1401]" pageId="22" pageNumber="23" type="description">
<paragraph id="841EA66FB905785CC5B84771FD7F207A" blockId="22.[151,662,1338,1401]" box="[151,662,1374,1401]" pageId="22" pageNumber="23">
<figureCitation id="1C9ABAEAB905785CC5B84771FF0C207B" box="[151,229,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="22" pageNumber="23">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB905785CC5DE4771FEE4207A" box="[241,269,1374,1400]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">28</figureCitation>
(habitus), 28 (aedeagus), 39 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5B84789FEBC20E6" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="841EA66FB905785CC5B84789FEBC20E6" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5B84789FEA220C2" bold="true" box="[151,331,1446,1472]" pageId="22" pageNumber="23">Type Material.</emphasis>
<materialsCitation id="34C9AC32B905785CC47A4789FEBC20E6" ID-GBIF-Occurrence="3962178313" collectingDate="1982-01-14" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="22" pageNumber="23" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB905785CC47A4789FE5420C2" box="[341,445,1446,1472]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB905785CC7034789FD4E20C2" box="[556,679,1446,1472]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B905785CC79A4788FCFE20C3" box="[693,791,1447,1473]" pageId="22" pageNumber="23">N. Prov.</collectorName>
,
<quantity id="43590B8AB905785CC60A4789FC9B20C2" box="[805,882,1446,1472]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="km" value="22.0">22 km</quantity>
.
<collectorName id="2954C3B9B905785CC6AF4788FC1620C3" box="[896,1023,1447,1473]" pageId="22" pageNumber="23">N. Maroua</collectorName>
, desert stream,
<date id="F01F80AFB905785CC1994789FAC820C2" box="[1206,1313,1446,1472]" pageId="22" pageNumber="23" value="1982-01-14">
<collectingDate id="E05B7947B905785CC1994789FAC820C2" box="[1206,1313,1446,1472]" pageId="22" pageNumber="23" value="1982-01-14">14.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B905785CC04B4788FF0720E6" pageId="22" pageNumber="23">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB905785CC42847E5FEA320E6" box="[263,330,1482,1508]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5E847C1FE402376" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="841EA66FB905785CC5E847C1FE402376" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E847C1FE38230A" bold="true" box="[199,465,1518,1544]" pageId="22" pageNumber="23">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB905785CC12747C1FB94230A" box="[1032,1149,1518,1544]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
by the combination of the brown to dark brown or black dorsum, the deep anterolateral pronotal foveae, and the aedeagus (
<figureCitation id="1C9ABAEAB905785CC1CF443DFADC232E" box="[1248,1333,1554,1580]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">Fig. 28</figureCitation>
). Similar in habitus and size to
<taxonomicName id="43A1DDECB905785CC4A34418FD8D2352" authorityName="Perkins" authorityYear="2022" box="[396,612,1591,1616]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="camerocontinua">
<emphasis id="B6D57A7DB905785CC4A34418FD8D2352" box="[396,612,1591,1616]" italics="true" pageId="22" pageNumber="23">H. camerocontinua</emphasis>
</taxonomicName>
, but differing therefrom in denser dorsal punctation, ventral morphology, and aedeagal characters.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785DC5E84451FC1624D3" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="841EA66FB905785CC5E84451FB0F2296" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E84451FEB8239A" bold="true" box="[199,337,1662,1688]" pageId="22" pageNumber="23">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB905785CC48B4451FDEF239A" box="[420,518,1662,1688]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.37/0.57; head width 0.34; pronotum 0.32/0.45, PA 0.38, PB 0.40; elytra 0.80/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB905785CC106448DFB6923BE" box="[1065,1152,1698,1724]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">Fig. 28</figureCitation>
). Dorsum brown to dark brown or black, frons and pronotal disc darker, maxillary palpi light brown, tip of last palpomere not darker. Frons finely moderately densely punctulate, ca. 1xef, interstices ca. 12xpd, interstices shining. Pronotum moderately coarsely moderately densely punctate, ca. 2xpd of frons punctures; interstices ca. 1xpd or slightly greater, shining. Clypeus medially very finely sparsely punctulate, shining; laterally very finely densely punctulate, dull. Labrum apicomedially excised, free margins not upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae deep, posterolateral shallow, posterosubmedial moderately deep.
</paragraph>
<paragraph id="841EA66FB905785DC5E845B1FEB325FB" blockId="22.[151,1437,1446,2013]" lastBlockId="23.[151,1437,151,537]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">Elytra with summit of posterior declivity at or very near posterior 1/3; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB904785DC5B8432CFC1624D3" blockId="23.[151,1437,151,537]" pageId="23" pageNumber="24">Venter: Mentum sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight or very slightly arcuate, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slender, straight. All tibiae slender, straight. Abdominal apex symmetrical, with deep apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E843F5FC2124F7" box="[199,968,474,501]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="841EA66FB904785DC5E843F5FC2124F7" blockId="23.[151,1437,151,537]" box="[199,968,474,501]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E843F5FE8824F6" bold="true" box="[199,353,474,500]" pageId="23" pageNumber="24">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB904785DC7E343F4FD1524F7" box="[716,764,475,501]" pageId="23" pageNumber="24">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB904785DC64843F4FC5224F7" box="[871,955,475,501]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="23" pageNumber="24">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E843D0FBD6271B" box="[199,1087,511,537]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="841EA66FB904785DC5E843D0FBD6271B" blockId="23.[151,1437,151,537]" box="[199,1087,511,537]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E843D0FEA7271B" bold="true" box="[199,334,511,537]" pageId="23" pageNumber="24">Etymology.</emphasis>
Named in reference to the known distribution, and to the arid area.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,144 @@
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<mods:title id="C709053F567F33D342D647BF3F410D41">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B906785CC5B84521FB2421A3" ID-DOI="http://doi.org/10.5281/zenodo.7300204" ID-GBIF-Taxon="204205225" ID-Zenodo-Dep="7300204" LSID="urn:lsid:plazi:treatment:0C081779B906785CC5B84521FB2421A3" httpUri="http://treatment.plazi.org/id/0C081779B906785CC5B84521FB2421A3" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="CCBBF5E4B906785FC5B84521FD4B222A" box="[151,674,1806,1833]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="841EA66FB906785FC5B84521FD4B222A" blockId="21.[151,674,1806,1869]" box="[151,674,1806,1833]" pageId="21" pageNumber="22">
<heading id="DF561103B906785FC5B84521FD4B222A" bold="true" box="[151,674,1806,1833]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="B6D57A7DB906785FC5B84521FD4B222A" bold="true" box="[151,674,1806,1833]" pageId="21" pageNumber="22">
<taxonomicName id="43A1DDECB906785FC5B84521FDEA222B" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,515,1806,1833]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="camerosemicordata" status="sp. nov.">
<emphasis id="B6D57A7DB906785FC5B84521FDEA222B" bold="true" box="[151,515,1806,1833]" italics="true" pageId="21" pageNumber="22">Hydraena camerosemicordata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B906785FC73D4520FD4B222A" box="[530,674,1806,1833]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5B8451CFD7F224F" box="[151,662,1843,1869]" pageId="21" pageNumber="22" type="description">
<paragraph id="841EA66FB906785FC5B8451CFD7F224F" blockId="21.[151,674,1806,1869]" box="[151,662,1843,1869]" pageId="21" pageNumber="22">
<figureCitation id="1C9ABAEAB906785FC5B8451CFF0C224F" box="[151,229,1843,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="21" pageNumber="22">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB906785FC5DE451CFEE4224F" box="[241,269,1843,1869]" captionStart="FIGURE 27" captionStartId="52.[151,250,1988,2012]" captionTargetBox="[296,1303,213,1939]" captionTargetId="figure-25@52.[213,1374,181,1964]" captionTargetPageId="52" captionText="FIGURE 27. Hydraena camerosemicordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284925" httpUri="https://zenodo.org/record/7284925/files/figure.png" pageId="21" pageNumber="22">27</figureCitation>
(habitus), 27 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5B84554FD3722BB" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="841EA66FB906785FC5B84554FD3722BB" blockId="21.[151,1436,1914,2013]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5B84554FEA32296" bold="true" box="[151,330,1914,1941]" pageId="21" pageNumber="22">Type Material.</emphasis>
<materialsCitation id="34C9AC32B906785FC47C4554FF0322BB" ID-GBIF-Occurrence="3962178327" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB906785FC47C4554FE522297" box="[339,443,1915,1941]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB906785FC7304555FDC42296" bold="true" box="[543,557,1914,1940]" pageId="21" pageNumber="22"></emphasis>
<collectingCountry id="FCB6E6FFB906785FC7024554FD4B2297" box="[557,674,1915,1941]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
:
<quantity id="43590B8AB906785FC79B4555FD142297" box="[692,765,1914,1941]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B906785FC6294554FC5E2297" box="[774,951,1915,1941]" pageId="21" pageNumber="22">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB906785FC1304554FB682297" box="[1055,1153,1915,1941]" pageId="21" pageNumber="22" value="1982-05-01">
<collectingDate id="E05B7947B906785FC1304554FB682297" box="[1055,1153,1915,1941]" pageId="21" pageNumber="22" value="1982-05-01">1.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B906785FC1A04554FACC2297" box="[1167,1317,1915,1941]" pageId="21" pageNumber="22">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB906785FC5B045B0FF0B22BB" box="[159,226,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B906785FC5DA45B0FD3322BB" ID-GBIF-Occurrence="3962178352" box="[245,730,1951,1977]" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="21" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB906785FC5DA45B0FE8F22BB" box="[245,358,1951,1977]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
: Same data as holotype (6
<collectionCode id="E2B03EAAB906785FC7BE45B0FD3A22BB" box="[657,723,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785CC5E845EDFE1E2443" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="841EA66FB906785CC5E845EDFE1E2443" blockId="21.[151,1436,1914,2013]" lastBlockId="22.[151,1437,150,1185]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E845EDFE3D22DE" bold="true" box="[199,468,1986,2012]" pageId="21" pageNumber="22">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB906785FC10B45ECFB7022DF" box="[1060,1177,1987,2013]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
by the combination of the chordate and convex pronotum, the very narrow metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB905785CC1B642B9FB0425B2" box="[1177,1261,150,176]" captionStart="FIGURE 27" captionStartId="52.[151,250,1988,2012]" captionTargetBox="[296,1303,213,1939]" captionTargetId="figure-25@52.[213,1374,181,1964]" captionTargetPageId="52" captionText="FIGURE 27. Hydraena camerosemicordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284925" httpUri="https://zenodo.org/record/7284925/files/figure.png" pageId="22" pageNumber="23">Fig. 27</figureCitation>
). Differentiated from
<taxonomicName id="43A1DDECB905785CC5F94294FE4225D6" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[214,427,187,212]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="camerocarinata" status="sp. nov.">
<emphasis id="B6D57A7DB905785CC5F94294FE4225D6" box="[214,427,187,212]" italics="true" pageId="22" pageNumber="23">H. camerocarinata</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB905785CC49D4295FE1B25D6" bold="true" box="[434,498,186,212]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="ADE6C706B905785CC49D4295FE1B25D6" box="[434,498,186,212]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, a species which also has a semichordate pronotum, by the more convex and more chordate pronotum, the deeper pronotal foveae, the area between the antero- and posterolateral pronotal foveae more strongly convex, the more acuminate elytra, and the different proportions of the ventral characters (P2/w/l/s ca. 1/0.2/2/2 vs. 1/0.15/0.15/2).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5E84365FDAD215A" pageId="22" pageNumber="23" type="description">
<paragraph id="841EA66FB905785CC5E84365FCD327AB" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E84365FEB82466" bold="true" box="[199,337,330,356]" pageId="22" pageNumber="23">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB905785CC48B4365FDEF2466" box="[420,518,330,356]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.37/0.54; head width 0.31; pronotum 0.37/0.44, PA 0.35, PB 0.31; elytra 0.81/0.54. Habitus as illustrated (
<figureCitation id="1C9ABAEAB905785CC13D4341FB8E248A" box="[1042,1127,366,392]" captionStart="FIGURE 27" captionStartId="52.[151,250,1988,2012]" captionTargetBox="[296,1303,213,1939]" captionTargetId="figure-25@52.[213,1374,181,1964]" captionTargetPageId="52" captionText="FIGURE 27. Hydraena camerosemicordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284925" httpUri="https://zenodo.org/record/7284925/files/figure.png" pageId="22" pageNumber="23">Fig. 27</figureCitation>
). Pronotum semicordiform, anterior margin much wider than posterior margin. Dorsum brown except head and pronotal fascia black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately punctulate, more sparsely punctulate on fascia than on surrounding areas, interstices effacedly microreticulate, weakly shining; punctures of frons disc fine and sparse, 1xef or less; punctures of pronotal disc ca. 12xef, interstices ca. 13xpd; punctures of clypeus fine, moderately dense. Labrum apicomedial excision deep and wide, free margins weakly upturned, angulate laterally and medially. Pronotum distinctly convex, especially transversely, anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae deep, posterosubmedial foveae deep, area between antero- and posterolateral foveae distinctly convex.
</paragraph>
<paragraph id="841EA66FB905785CC5E8409DFCF3265E" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="23">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures 12xpd and deeper than largest pronotal punctures, punctures becoming gradually slightly smaller toward posterior. Intervals not raised, width ca. 1xpd, interstices between punctures of a row ca. 23xpd, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rather sharply rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB905785CC5B84148FDAD215A" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="23">Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, lines, straight, convergent anteriorly, very weakly if at all raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Pro-, meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5E8464DFC21217E" box="[199,968,1122,1149]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="841EA66FB905785CC5E8464DFC21217E" blockId="22.[151,1437,150,1185]" box="[199,968,1122,1149]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E8464DFE88217E" bold="true" box="[199,353,1122,1148]" pageId="22" pageNumber="23">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB905785CC7E3464CFD15217F" box="[716,764,1123,1149]" pageId="22" pageNumber="23">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB905785CC648464DFC52217E" box="[871,955,1122,1148]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="22" pageNumber="23">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5E846A9FB2421A3" box="[199,1229,1158,1185]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="841EA66FB905785CC5E846A9FB2421A3" blockId="22.[151,1437,150,1185]" box="[199,1229,1158,1185]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E846A9FEA721A2" bold="true" box="[199,334,1158,1184]" pageId="22" pageNumber="23">Etymology.</emphasis>
Named in reference to the known distribution and the semi-chordate pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,135 @@
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<mods:title id="5C1AE4524F1A1907A6F394F3B4F8EC4B">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B906785FC5B84069FB1223BF" ID-DOI="http://doi.org/10.5281/zenodo.7300273" ID-GBIF-Taxon="204205224" ID-Zenodo-Dep="7300273" LSID="urn:lsid:plazi:treatment:0C081779B906785FC5B84069FB1223BF" httpUri="http://treatment.plazi.org/id/0C081779B906785FC5B84069FB1223BF" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="CCBBF5E4B906785FC5B84069FD9D2762" box="[151,628,582,609]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="841EA66FB906785FC5B84069FD9D2762" blockId="21.[151,662,582,645]" box="[151,628,582,609]" pageId="21" pageNumber="22">
<heading id="DF561103B906785FC5B84069FD9D2762" bold="true" box="[151,628,582,609]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="B6D57A7DB906785FC5B84069FD9D2762" bold="true" box="[151,628,582,609]" pageId="21" pageNumber="22">
<taxonomicName id="43A1DDECB906785FC5B84069FE3F2763" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,470,582,609]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="camerocarinata" status="sp. nov.">
<emphasis id="B6D57A7DB906785FC5B84069FE3F2763" bold="true" box="[151,470,582,609]" italics="true" pageId="21" pageNumber="22">Hydraena camerocarinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B906785FC4CB4068FD9D2762" box="[484,628,582,609]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5B84044FD7F2787" box="[151,662,619,645]" pageId="21" pageNumber="22" type="description">
<paragraph id="841EA66FB906785FC5B84044FD7F2787" blockId="21.[151,662,582,645]" box="[151,662,619,645]" pageId="21" pageNumber="22">
<figureCitation id="1C9ABAEAB906785FC5B84044FF0C2787" box="[151,229,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="21" pageNumber="22">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB906785FC5DE4044FEE42787" box="[241,269,619,645]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">26</figureCitation>
(habitus), 26 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5B8409CFF0727F3" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="841EA66FB906785FC5B8409CFF0727F3" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5B8409CFEA327CE" bold="true" box="[151,330,690,717]" pageId="21" pageNumber="22">Type Material.</emphasis>
<materialsCitation id="34C9AC32B906785FC47C409CFF0727F3" ID-GBIF-Occurrence="3962178307" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB906785FC47C409CFE5227CF" box="[339,443,691,717]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB906785FC730409DFDC427CE" bold="true" box="[543,557,690,716]" pageId="21" pageNumber="22"></emphasis>
<collectingCountry id="FCB6E6FFB906785FC702409CFD4B27CF" box="[557,674,691,717]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
:
<quantity id="43590B8AB906785FC79B409DFD1427CF" box="[692,765,690,717]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B906785FC629409CFC5E27CF" box="[774,951,691,717]" pageId="21" pageNumber="22">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB906785FC130409CFB6827CF" box="[1055,1153,691,717]" pageId="21" pageNumber="22" value="1982-05-01">
<collectingDate id="E05B7947B906785FC130409CFB6827CF" box="[1055,1153,691,717]" pageId="21" pageNumber="22" value="1982-05-01">1.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B906785FC1A0409CFACC27CF" box="[1167,1317,691,717]" pageId="21" pageNumber="22">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB906785FC5B040F8FF0B27F3" box="[159,226,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E840D5FCC3265F" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="841EA66FB906785FC5E840D5FCC3265F" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E840D5FE382616" bold="true" box="[199,465,762,788]" pageId="21" pageNumber="22">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB906785FC12740D4FB942617" box="[1032,1149,763,789]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
by the combination of the nearly semicordate pronotum, the widely separated and carinate mesoventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB906785FC0154130FA79263B" box="[1338,1424,799,825]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
). Refer also to the diagnosis of
<taxonomicName id="43A1DDECB906785FC4C9416CFD0D265F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[486,740,835,861]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="camerosemicordata" status="sp. nov.">
<emphasis id="B6D57A7DB906785FC4C9416CFD0D265F" box="[486,740,835,861]" italics="true" pageId="21" pageNumber="22">H. camerosemicordata</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB906785FC7C4416CFCC3265F" bold="true" box="[747,810,835,861]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="ADE6C706B906785FC7C4416CFCC3265F" box="[747,810,835,861]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84149FE452377" pageId="21" pageNumber="22" type="description">
<paragraph id="841EA66FB906785FC5E84149FDA221A3" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84149FEB82682" bold="true" box="[199,337,870,896]" pageId="21" pageNumber="22">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB906785FC48B4148FDEF2683" box="[420,518,871,897]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.35/0.57; head width 0.33; pronotum 0.35/0.44, PA 0.37, PB 0.33; elytra 0.81/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB906785FC12341A4FB8926A7" box="[1036,1120,907,933]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
). Pronotum almost semicordiform, anterior margin much wider than posterior margin. Dorsum brown except head and pronotal fascia black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately punctulate, more sparsely punctulate on fascia than on surrounding areas, interstices shining; punctures of frons disc fine and sparse, less than 1xef; punctures of pronotal disc ca. 2xef, interstices ca. 13xpd or slightly larger; punctures of clypeus fine, moderately dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB906785FC5E84684FD3E2057" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures 23xpd and deeper than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, very narrow, width ca. 1/2xpd or less, as are interstices between punctures of a row, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB906785FC5B84770FE452377" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.15/0.15/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, carinate lines, straight, convergent anteriorly, very weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Protibiae slender, very weakly arcuate. Mesotibiae slender, straight. Metatibiae slender, straight, very slightly wider at midlength than apically. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84451FC21239B" box="[199,968,1662,1689]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="841EA66FB906785FC5E84451FC21239B" blockId="21.[151,1437,690,1725]" box="[199,968,1662,1689]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84451FE88239A" bold="true" box="[199,353,1662,1688]" pageId="21" pageNumber="22">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB906785FC7E34450FD15239B" box="[716,764,1663,1689]" pageId="21" pageNumber="22">type</typeStatus>
locality (
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).
</paragraph>
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<emphasis id="B6D57A7DB906785FC5E8448CFEA723BF" bold="true" box="[199,334,1699,1725]" pageId="21" pageNumber="22">Etymology.</emphasis>
Named in reference to the known distribution and the carinate metaventral plaques.
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<mods:title id="0BDDB595AC1DFBAC4D4F6AD55FAE7A72">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<emphasis id="B6D57A7DB907785EC5B842B8FE3D25B0" bold="true" box="[151,468,151,178]" italics="true" pageId="20" pageNumber="21">Hydraena cameropalmula</emphasis>
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,
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<paragraph id="841EA66FB907785EC5B84294FD7F25D7" blockId="20.[151,662,151,213]" box="[151,662,187,213]" pageId="20" pageNumber="21">
<figureCitation id="1C9ABAEAB907785EC5B84294FF0C25D7" box="[151,229,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="20" pageNumber="21">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB907785EC5DE4294FEE425D7" box="[241,269,187,213]" captionStart="FIGURE 24" captionStartId="49.[151,250,1989,2013]" captionTargetBox="[251,1350,189,1935]" captionTargetId="figure-15@49.[219,1369,181,1966]" captionTargetPageId="49" captionText="FIGURE 24. Hydraena cameropalmula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284915" httpUri="https://zenodo.org/record/7284915/files/figure.png" pageId="20" pageNumber="21">24</figureCitation>
(habitus), 24 (aedeagus), 43 (map)
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<paragraph id="841EA66FB907785EC5B8432CFD8D2442" blockId="20.[151,1437,258,1178]" pageId="20" pageNumber="21">
<emphasis id="B6D57A7DB907785EC5B8432CFEAE241E" bold="true" box="[151,327,258,285]" pageId="20" pageNumber="21">Type Material.</emphasis>
<materialsCitation id="34C9AC32B907785EC463432DFD8D2442" ID-GBIF-Occurrence="3962178310" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB907785EC463432DFE5D241E" box="[332,436,258,284]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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/ Matute, / Tiko Plantation /
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<emphasis id="B6D57A7DB907785EC5E84365FE3D2466" bold="true" box="[199,468,330,356]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
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by the combination of the pronotum with a macula, the absence of posterosubmedial pronotal foveae, and the aedeagus (
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). Reliable determinations to distinguish this species from other similar members in Aedeagal Species Group 3 will require dissection of males.
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<emphasis id="B6D57A7DB907785EC5E843F6FEB824F1" bold="true" box="[199,337,473,499]" pageId="20" pageNumber="21">Description</emphasis>
. Size:
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(length/width, mm): body (length to elytral apices) 1.33/0.58; head width 0.34; pronotum 0.34/0.47, PA 0.38, PB 0.38; elytra 0.82/0.58. Habitus as illustrated (
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). Dorsum testaceous to dark brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely sparsely punctulate, pronotum moderately coarsely, densely punctate, punctures much larger than those of frons, interstices shining, ca. 12xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral foveae very shallow, posterosubmedial foveae absent.
</paragraph>
<paragraph id="841EA66FB907785EC5E840FCFB78265A" blockId="20.[151,1437,258,1178]" pageId="20" pageNumber="21">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB907785EC5B8414CFAB32150" blockId="20.[151,1437,258,1178]" pageId="20" pageNumber="21">Venter: Mentum and postmentum finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/4/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, parallel or nearly so, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2.5x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with apicomedian shallow notch in tergite, into which minute apical process of ventrite fits.</paragraph>
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<emphasis id="B6D57A7DB907785EC5E84673FE882174" bold="true" box="[199,353,1116,1142]" pageId="20" pageNumber="21">Distribution.</emphasis>
Currently known only from the
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</paragraph>
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<emphasis id="B6D57A7DB907785EC5E846AFFEA72198" bold="true" box="[199,334,1152,1178]" pageId="20" pageNumber="21">Etymology.</emphasis>
Named in reference to the known distribution and the palmate shape of the aedeagus.
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<paragraph id="841EA66FB907785EC5B846C4FDDA2007" blockId="20.[151,662,1259,1322]" box="[151,563,1259,1286]" pageId="20" pageNumber="21">
<heading id="DF561103B907785EC5B846C4FDDA2007" bold="true" box="[151,563,1259,1286]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
<emphasis id="B6D57A7DB907785EC5B846C4FDDA2007" bold="true" box="[151,563,1259,1286]" pageId="20" pageNumber="21">
<taxonomicName id="43A1DDECB907785EC5B846C4FE7D2004" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,404,1259,1286]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millenaria" status="sp. nov.">
<emphasis id="B6D57A7DB907785EC5B846C4FE7D2004" bold="true" box="[151,404,1259,1286]" italics="true" pageId="20" pageNumber="21">Hydraena millenaria</emphasis>
</taxonomicName>
,
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</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B907785EC5B84720FD7F202B" box="[151,662,1295,1322]" pageId="20" pageNumber="21" type="description">
<paragraph id="841EA66FB907785EC5B84720FD7F202B" blockId="20.[151,662,1259,1322]" box="[151,662,1295,1322]" pageId="20" pageNumber="21">
<figureCitation id="1C9ABAEAB907785EC5B84720FF0C202B" box="[151,229,1295,1321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="20" pageNumber="21">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB907785EC5DE4720FEE42028" box="[241,269,1295,1322]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">25</figureCitation>
(habitus), 25 (aedeagus), 40 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B907785EC5B84778FCD22097" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="841EA66FB907785EC5B84778FCD22097" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
<emphasis id="B6D57A7DB907785EC5B84778FEA12073" bold="true" box="[151,328,1367,1393]" pageId="20" pageNumber="21">Type Material.</emphasis>
<materialsCitation id="34C9AC32B907785EC4614778FEB82097" ID-GBIF-Occurrence="3962178350" collectingDate="1982-02-01" collectionCode="MCZ" collectorName="N. W. Prov. &amp; P. D. Perkins" country="Cameroon" location="Bamenda" municipality="Skyline Hotel" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB907785EC4614778FE5F2073" box="[334,438,1367,1393]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB907785EC7314778FD702073" box="[542,665,1367,1393]" name="Cameroon" pageId="20" pageNumber="21">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B907785EC78B4777FCC12070" box="[676,808,1368,1394]" pageId="20" pageNumber="21">N. W. Prov.</collectorName>
,
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, stream nr.
<collectingMunicipality id="647A3C15B907785EC1314778FB522073" box="[1054,1211,1367,1393]" pageId="20" pageNumber="21">Skyline Hotel</collectingMunicipality>
,
<date id="F01F80AFB907785EC1E94778FAC52073" box="[1222,1324,1367,1393]" pageId="20" pageNumber="21" value="1982-02-01">
<collectingDate id="E05B7947B907785EC1E94778FAC52073" box="[1222,1324,1367,1393]" pageId="20" pageNumber="21" value="1982-02-01">1.ii.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B907785EC0484777FF072097" pageId="20" pageNumber="21">P. D. Perkins</collectorName>
” (
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)
</materialsCitation>
.
<materialsCitation id="34C9AC32B907785EC4734753FCDE2097" ID-GBIF-Occurrence="3962178302" box="[348,823,1403,1430]" collectingDate="1982-02-01" collectionCode="MCZ" collectorName="N. W. Prov. &amp; P. D. Perkins" country="Cameroon" location="Bamenda" municipality="Skyline Hotel" pageId="20" pageNumber="21" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB907785EC4734753FE2B2094" box="[348,450,1404,1430]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
: Same data as holotype (1
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B907785EC5E847B0FDA8234B" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="841EA66FB907785EC5E847B0FDA8234B" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
<emphasis id="B6D57A7DB907785EC5E847B0FE3820BB" bold="true" box="[199,465,1439,1465]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB907785EC12747B0FB9420BB" box="[1032,1149,1439,1465]" name="Cameroon" pageId="20" pageNumber="21">Cameroon</collectingCountry>
by the combination of the absence of metaventral plaques, the frons and pronotum finely sparsely punctulate, the punctures of the pronotum very consistent in size throughout, and the aedeagus (
<figureCitation id="1C9ABAEAB907785EC7C347C8FCA92300" box="[748,832,1511,1538]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">Fig. 25</figureCitation>
). The dorsal punctation is much finer and sparser than that of
<taxonomicName id="43A1DDECB907785EC5CA4423FE702327" authorityName="Perkins" authorityYear="2022" box="[229,409,1547,1573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB907785EC5CA4423FE702327" box="[229,409,1547,1573]" italics="true" pageId="20" pageNumber="21">H. cameropetila</emphasis>
</taxonomicName>
and
<taxonomicName id="43A1DDECB907785EC4E14423FD612327" authorityName="Perkins" authorityYear="2022" box="[462,648,1548,1573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB907785EC4E14423FD612327" box="[462,648,1548,1573]" italics="true" pageId="20" pageNumber="21">H. camerosetosa</emphasis>
</taxonomicName>
, two species which also lack metaventral plaques. All three species are in different Aedeagal Species Groups.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B907785FC5E8447CFBFC2443" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="841EA66FB907785EC5E8447CFAF3226B" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
<emphasis id="B6D57A7DB907785EC5E8447CFEB8236F" bold="true" box="[199,337,1619,1645]" pageId="20" pageNumber="21">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB907785EC4B3447CFE17236F" box="[412,510,1619,1645]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.20/0.53; head width 0.33; pronotum 0.31/0.44, PA 0.34, PB 0.39; elytra 0.75/0.53. Habitus as illustrated (
<figureCitation id="1C9ABAEAB907785EC6CC4458FBDE2390" box="[995,1079,1655,1682]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">Fig. 25</figureCitation>
). Dorsum brown to dark brown, head, pronotal fascia and elytra dark brown, pronotum rather widely bordered anteriorly and posteriorly slightly lighter brown, maxillary palpi testaceous, tip of last palpomere not darker. Frons and pronotum finely sparsely punctulate, punctures of pronotum slightly larger than those of frons, very consistent in size throughout, interstices shining, ca. 13xpd. Clypeus medially very finely sparsely punctulate, shining; laterally densely micropunctulate, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral and posterosubmedian foveae shallow.
</paragraph>
<paragraph id="841EA66FB907785EC5E8455CFBA922FB" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals shining, not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB906785FC5B842B8FBFC2443" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="22">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex slightly asymmetrical, with deep and wide apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84365FC212467" box="[199,968,330,357]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="841EA66FB906785FC5E84365FC212467" blockId="21.[151,1437,151,429]" box="[199,968,330,357]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84365FE882466" bold="true" box="[199,353,330,356]" pageId="21" pageNumber="22">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB906785FC7E34364FD152467" box="[716,764,331,357]" pageId="21" pageNumber="22">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB906785FC6484364FC522467" box="[871,955,331,357]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="21" pageNumber="22">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84340FDD424AF" pageId="21" pageNumber="22" type="etymology">
<paragraph id="841EA66FB906785FC5E84340FDD424AF" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84340FEA7248B" bold="true" box="[199,334,367,393]" pageId="21" pageNumber="22">Etymology.</emphasis>
Named in reference to the fact that the authors goal of describing more than 1000 new species of water beetles is now happily attained.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,160 @@
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<mods:title id="F45BE12C5720648E140D15E43F4ADDF9">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<heading id="DF561103B9087851C5B84309FDBD2442" bold="true" box="[151,596,294,321]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
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<emphasis id="B6D57A7DB9087851C5B84309FE5C2443" bold="true" box="[151,437,294,321]" italics="true" pageId="27" pageNumber="28">Hydraena camerodigita</emphasis>
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,
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</subSubSection>
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<paragraph id="841EA66FB9087851C5B84364FD222467" blockId="27.[151,715,294,357]" box="[151,715,330,357]" pageId="27" pageNumber="28">
<figureCitation id="1C9ABAEAB9087851C5B84364FF0C2467" box="[151,229,330,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="27" pageNumber="28">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB9087851C5DE4365FEE42467" box="[241,269,330,357]" captionStart="FIGURE 35" captionStartId="60.[151,250,1971,1995]" captionTargetBox="[204,1375,195,1913]" captionTargetId="figure-25@60.[178,1410,181,1947]" captionTargetPageId="60" captionText="FIGURE 35. Hydraena camerodigita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284953" httpUri="https://zenodo.org/record/7284953/files/figure.png" pageId="27" pageNumber="28">35</figureCitation>
(habitus), 35 (aedeagus), 40, 42 (maps)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5B843BCFE7224F7" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="841EA66FB9087851C5B843BCFE7224F7" blockId="27.[151,1437,402,1329]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5B843BCFEAE24AE" bold="true" box="[151,327,402,429]" pageId="27" pageNumber="28">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9087851C46243BCFF0324D3" ID-GBIF-Occurrence="3962178335" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9087851C46243BCFE5C24AF" box="[333,437,403,429]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9087851C73343BCFD7E24AF" box="[540,663,403,429]" name="Cameroon" pageId="27" pageNumber="28">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9087851C78E43BCFCE924AF" box="[673,768,403,429]" pageId="27" pageNumber="28">N. Prov.</collectorName>
,
<quantity id="43590B8AB9087851C62543BDFCB924AF" box="[778,848,402,429]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="27" pageNumber="28" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B9087851C67943BCFBEA24AF" box="[854,1027,403,429]" pageId="27" pageNumber="28">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB9087851C14B43BCFB2E24AF" box="[1124,1223,403,429]" pageId="27" pageNumber="28" value="1982-05-01">
<collectingDate id="E05B7947B9087851C14B43BCFB2E24AF" box="[1124,1223,403,429]" pageId="27" pageNumber="28" value="1982-05-01">1.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9087851C02E43BCFA7A24AF" box="[1281,1427,403,429]" pageId="27" pageNumber="28">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9087851C5B04398FF0B24D3" box="[159,226,439,465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9087851C5DB4398FE7E24F7" ID-GBIF-Occurrence="3962178351" collectingDate="1982-02-01" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="27" pageNumber="28" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9087851C5DB4398FE8C24D3" box="[244,357,439,465]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
<collectorName id="2954C3B9B9087851C4404398FE4424D3" box="[367,429,439,465]" pageId="27" pageNumber="28">Same</collectorName>
data as holotype (2
<collectionCode id="E2B03EAAB9087851C7A24398FD2624D3" box="[653,719,439,465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
);
<collectorName id="2954C3B9B9087851C7CE4398FC8C24D3" box="[737,869,439,465]" pageId="27" pageNumber="28">N. W. Prov.</collectorName>
,
<collectorName id="2954C3B9B9087851C6404398FC3524D3" box="[879,988,439,465]" pageId="27" pageNumber="28">Bamenda</collectorName>
, stream nr.
<collectorName id="2954C3B9B9087851C1754398FB1E24D3" box="[1114,1271,439,465]" pageId="27" pageNumber="28">Skyline Hotel</collectorName>
,
<date id="F01F80AFB9087851C02D4398FA8124D3" box="[1282,1384,439,465]" pageId="27" pageNumber="28" value="1982-02-01">
<collectingDate id="E05B7947B9087851C02D4398FA8124D3" box="[1282,1384,439,465]" pageId="27" pageNumber="28" value="1982-02-01">1.ii.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9087851C5B843F4FEC124F7" box="[151,296,475,501]" pageId="27" pageNumber="28">P. D. Perkins</collectorName>
(4
<collectionCode id="E2B03EAAB9087851C46243F4FE6624F7" box="[333,399,475,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E843D1FA752763" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="841EA66FB9087851C5E843D1FA752763" blockId="27.[151,1437,402,1329]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E843D1FE38271A" bold="true" box="[199,465,510,536]" pageId="27" pageNumber="28">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9087851C12743D0FB94271B" box="[1032,1149,511,537]" name="Cameroon" pageId="27" pageNumber="28">Cameroon</collectingCountry>
by the combination of the narrow P2 (l/w ca. 4/1), the very small, short and narrow plaques, the deep anterolateral pronotal foveae, and the aedeagus (
<figureCitation id="1C9ABAEAB9087851C5DB4068FEA12763" box="[244,328,582,609]" captionStart="FIGURE 35" captionStartId="60.[151,250,1971,1995]" captionTargetBox="[204,1375,195,1913]" captionTargetId="figure-25@60.[178,1410,181,1947]" captionTargetPageId="60" captionText="FIGURE 35. Hydraena camerodigita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284953" httpUri="https://zenodo.org/record/7284953/files/figure.png" pageId="27" pageNumber="28">Fig. 35</figureCitation>
). Similar in habitus to
<taxonomicName id="43A1DDECB9087851C7684068FCEB2762" authorityName="Perkins" authorityYear="2022" box="[583,770,583,608]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB9087851C7684068FCEB2762" box="[583,770,583,608]" italics="true" pageId="27" pageNumber="28">H. camerosetosa</emphasis>
</taxonomicName>
, but differing therefrom in ventral and aedeagal characters.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E84045FDAD21EB" pageId="27" pageNumber="28" type="description">
<paragraph id="841EA66FB9087851C5E84045FB01265F" blockId="27.[151,1437,402,1329]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E84045FEB82786" bold="true" box="[199,337,618,644]" pageId="27" pageNumber="28">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9087851C48B4044FDEF2787" box="[420,518,619,645]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.25/0.56; head width 0.34; pronotum 0.32/0.43, PA 0.37, PB 0.38; elytra 0.76/0.56. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9087851C10640A0FB9627AB" box="[1065,1151,654,681]" captionStart="FIGURE 35" captionStartId="60.[151,250,1971,1995]" captionTargetBox="[204,1375,195,1913]" captionTargetId="figure-25@60.[178,1410,181,1947]" captionTargetPageId="60" captionText="FIGURE 35. Hydraena camerodigita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284953" httpUri="https://zenodo.org/record/7284953/files/figure.png" pageId="27" pageNumber="28">Fig. 35</figureCitation>
). Dorsum brown to dark brown, frons and pronotal disc often darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately finely and moderately sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 12xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae deep, posterolateral and posterosubmedial pronotal foveae moderately deep
</paragraph>
<paragraph id="841EA66FB9087851C5E84148FCA82113" blockId="27.[151,1437,402,1329]" pageId="27" pageNumber="28">Elytra with posterior declivity very gradual, summit of at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB9087851C5E84634FDAD21EB" blockId="27.[151,1437,402,1329]" pageId="27" pageNumber="28">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/ w/l/s) ca. 1/0.5/1/1.5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides parallel, blunt apex raised slightly above mesoventral intercoxal process. Plaques very small, short, narrow, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. All tibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E846DDFB13200F" box="[199,1274,1266,1293]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="841EA66FB9087851C5E846DDFB13200F" blockId="27.[151,1437,402,1329]" box="[199,1274,1266,1293]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E846DDFE88200E" bold="true" box="[199,353,1266,1292]" pageId="27" pageNumber="28">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9087851C7E346DCFD15200F" box="[716,764,1267,1293]" pageId="27" pageNumber="28">type</typeStatus>
locality and a site at Bamenda (
<figureCitation id="1C9ABAEAB9087851C14B46DCFB2B200F" box="[1124,1218,1267,1293]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="27" pageNumber="28">Figs. 40</figureCitation>
,
<figureCitation id="1C9ABAEAB9087851C1E246DCFB04200F" box="[1229,1261,1267,1293]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="27" pageNumber="28">42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E84738FA652033" box="[199,1420,1303,1329]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="841EA66FB9087851C5E84738FA652033" blockId="27.[151,1437,402,1329]" box="[199,1420,1303,1329]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E84738FEA72033" bold="true" box="[199,334,1303,1329]" pageId="27" pageNumber="28">Etymology.</emphasis>
Named in reference to the known distribution, and to the finger-like main-piece of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,155 @@
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<mods:title id="B8F3F38BDB1A929E0FB0C5EBCB48238C">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<subSubSection id="CCBBF5E4B9087851C5B847ADFD68209E" box="[151,641,1410,1437]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="841EA66FB9087851C5B847ADFD68209E" blockId="27.[151,662,1410,1473]" box="[151,641,1410,1437]" pageId="27" pageNumber="28">
<heading id="DF561103B9087851C5B847ADFD68209E" bold="true" box="[151,641,1410,1437]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B6D57A7DB9087851C5B847ADFD68209E" bold="true" box="[151,641,1410,1437]" pageId="27" pageNumber="28">
<taxonomicName id="43A1DDECB9087851C5B847ADFE0A209F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,483,1410,1437]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cameroflagellata" status="sp. nov.">
<emphasis id="B6D57A7DB9087851C5B847ADFE0A209F" bold="true" box="[151,483,1410,1437]" italics="true" pageId="27" pageNumber="28">Hydraena cameroflagellata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9087851C4DE47ACFD68209E" box="[497,641,1410,1437]" pageId="27" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5B84788FD7F20C3" box="[151,662,1446,1473]" pageId="27" pageNumber="28" type="description">
<paragraph id="841EA66FB9087851C5B84788FD7F20C3" blockId="27.[151,662,1410,1473]" box="[151,662,1446,1473]" pageId="27" pageNumber="28">
<figureCitation id="1C9ABAEAB9087851C5B84788FF0C20C3" box="[151,229,1446,1473]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="27" pageNumber="28">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB9087851C5DE4788FEE420C3" box="[241,269,1447,1473]" captionStart="FIGURE 36" captionStartId="61.[151,250,1867,1891]" captionTargetBox="[189,1419,207,1820]" captionTargetId="figure-15@61.[163,1423,181,1844]" captionTargetPageId="61" captionText="FIGURE 36. Hydraena cameroflagellata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284955" httpUri="https://zenodo.org/record/7284955/files/figure.png" pageId="27" pageNumber="28">36</figureCitation>
(habitus), 36 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5B847C0FBD42353" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="841EA66FB9087851C5B847C0FBD42353" blockId="27.[151,1437,1518,2013]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5B847C0FEAE230A" bold="true" box="[151,327,1518,1545]" pageId="27" pageNumber="28">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9087851C46247C0FDAC232F" ID-GBIF-Occurrence="3962178340" collectingDate="1982-05-22" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Nyong River" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9087851C46247C0FE5C230B" box="[333,437,1519,1545]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9087851C73247C0FD71230B" box="[541,664,1519,1545]" name="Cameroon" pageId="27" pageNumber="28">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9087851C78D47C0FCF5230B" box="[674,796,1519,1545]" pageId="27" pageNumber="28">C. S. Prov.</collectorName>
,
<quantity id="43590B8AB9087851C60A47C1FC86230B" box="[805,879,1518,1545]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="km" value="15.0">15 km</quantity>
.
<collectorName id="2954C3B9B9087851C65647C0FBF8230B" box="[889,1041,1519,1545]" pageId="27" pageNumber="28">S. Mbalmayo</collectorName>
,
<location id="817EF0B4B9087851C13347C0FB47230B" LSID="urn:lsid:plazi:treatment:0C081779B9087856C5B847ADFACD2763:817EF0B4B9087851C13347C0FB47230B" box="[1052,1198,1519,1545]" country="Cameroon" name="Nyong River" pageId="27" pageNumber="28">Nyong River</location>
, ex. leaves at margin,
<date id="F01F80AFB9087851C5B8443CFEE1232F" box="[151,264,1555,1581]" pageId="27" pageNumber="28" value="1982-05-22">
<collectingDate id="E05B7947B9087851C5B8443CFEE1232F" box="[151,264,1555,1581]" pageId="27" pageNumber="28" value="1982-05-22">22.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9087851C467443CFE09232F" box="[328,480,1555,1581]" pageId="27" pageNumber="28">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9087851C4D5443CFDD4232F" box="[506,573,1555,1581]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9087851C77E443CFBD02353" ID-GBIF-Occurrence="3962178323" collectingDate="1978-01-07" collectionCode="MCZ, LUM" collectorName="Gardensforhall &amp; Samuelson" country="Cameroon" location="Nyong River" pageId="27" pageNumber="28" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9087851C77E443CFD2B232F" box="[593,706,1555,1581]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
: Same data as holotype (2
<collectionCode id="E2B03EAAB9087851C6D5443CFBD5232F" box="[1018,1084,1555,1581]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
); Akonolinga, moist secondary forest and plantation, at light,
<date id="F01F80AFB9087851C4C74418FDAF2353" box="[488,582,1591,1617]" pageId="27" pageNumber="28" value="1978-01-07">
<collectingDate id="E05B7947B9087851C4C74418FDAF2353" box="[488,582,1591,1617]" pageId="27" pageNumber="28" value="1978-01-07">7.i.1978</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9087851C7AD4418FCC32353" box="[642,810,1591,1617]" pageId="27" pageNumber="28">Gardensforhall</collectorName>
&amp;
<collectorName id="2954C3B9B9087851C6614418FC202353" box="[846,969,1591,1617]" pageId="27" pageNumber="28">Samuelson</collectorName>
(1
<collectionCode id="E2B03EAAB9087851C6C14418FBD82353" box="[1006,1073,1591,1617]" pageId="27" pageNumber="28">LUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E84475FBC423BF" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="841EA66FB9087851C5E84475FBC423BF" blockId="27.[151,1437,1518,2013]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E84475FE3D2376" bold="true" box="[199,468,1626,1652]" pageId="27" pageNumber="28">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9087851C10B4474FB702377" box="[1060,1177,1627,1653]" name="Cameroon" pageId="27" pageNumber="28">Cameroon</collectingCountry>
by the combination of the small size (ca.
<quantity id="43590B8AB9087851C44C4451FE21239B" box="[355,456,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="28" unit="mm" value="1.05">1.05 mm</quantity>
, smallest species in
<collectingCountry id="FCB6E6FFB9087851C7874450FCC8239B" box="[680,801,1663,1689]" name="Cameroon" pageId="27" pageNumber="28">Cameroon</collectingCountry>
), the very narrow, very short, carinate or subcarinate, and widely separated plaques (P2/w/l/s ca. 1/0.2/2/2), and the male genitalia (
<figureCitation id="1C9ABAEAB9087851C6E3448CFBC823BF" box="[972,1057,1699,1725]" captionStart="FIGURE 36" captionStartId="61.[151,250,1867,1891]" captionTargetBox="[189,1419,207,1820]" captionTargetId="figure-15@61.[163,1423,181,1844]" captionTargetPageId="61" captionText="FIGURE 36. Hydraena cameroflagellata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284955" httpUri="https://zenodo.org/record/7284955/files/figure.png" pageId="27" pageNumber="28">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087856C5E844E9FD7B271B" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="841EA66FB9087851C5E844E9FC5A22DF" blockId="27.[151,1437,1518,2013]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E844E9FEB823E2" bold="true" box="[199,337,1734,1760]" pageId="27" pageNumber="28">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9087851C48B44E8FDEF23E3" box="[420,518,1735,1761]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.05/0.45; head width 0.27; pronotum 0.27/0.37, PA 0.31, PB 0.33; elytra 0.77/0.45. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9087851C10744C4FB962207" box="[1064,1151,1771,1797]" captionStart="FIGURE 36" captionStartId="61.[151,250,1867,1891]" captionTargetBox="[189,1419,207,1820]" captionTargetId="figure-15@61.[163,1423,181,1844]" captionTargetPageId="61" captionText="FIGURE 36. Hydraena cameroflagellata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284955" httpUri="https://zenodo.org/record/7284955/files/figure.png" pageId="27" pageNumber="28">Fig. 36</figureCitation>
). Dorsum light brown to brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely moderately sparsely punctulate, pronotum moderately finely, moderately densely punctate, punctures ca. 1xpd of largest punctures of frons, interstices shining, ca. 13xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral shallow or absent, posterosubmedian foveae shallow, medially confluent forming U-shaped impression.
</paragraph>
<paragraph id="841EA66FB90F7856C5E842B8FBB1241F" blockId="28.[151,1437,151,609]" pageId="28" pageNumber="29">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row.Apices in dorsal aspect separately rounded, in posterior aspect margins forming weak angle with one another.</paragraph>
<paragraph id="841EA66FB90F7856C5B84308FD7B271B" blockId="28.[151,1437,151,609]" pageId="28" pageNumber="29">Venter: Mentum moderately sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, slightly arcuate, slightly converging toward one another anteriorly, raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia slender, very weakly arcuate. Meso- metatibiae slender, straight. Abdominal apex symmetrical, with very small apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E8400DFC21273F" box="[199,968,546,573]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="841EA66FB90F7856C5E8400DFC21273F" blockId="28.[151,1437,151,609]" box="[199,968,546,573]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E8400DFE88273E" bold="true" box="[199,353,546,572]" pageId="28" pageNumber="29">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB90F7856C7E3400CFD15273F" box="[716,764,547,573]" pageId="28" pageNumber="29">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90F7856C648400CFC52273F" box="[871,955,547,573]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="28" pageNumber="29">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E84068FACD2763" box="[199,1316,583,609]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="841EA66FB90F7856C5E84068FACD2763" blockId="28.[151,1437,151,609]" box="[199,1316,583,609]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E84068FEA72763" bold="true" box="[199,334,583,609]" pageId="28" pageNumber="29">Etymology.</emphasis>
Named in reference to the known distribution, and to the very long aedeagal flagellum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,133 @@
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<mods:title id="E0F49B7DD7061932BB02F2BA6F860589">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9097851C5B841A5FF0425D7" ID-DOI="http://doi.org/10.5281/zenodo.7300212" ID-GBIF-Taxon="204205219" ID-Zenodo-Dep="7300212" LSID="urn:lsid:plazi:treatment:0C081779B9097851C5B841A5FF0425D7" httpUri="http://treatment.plazi.org/id/0C081779B9097851C5B841A5FF0425D7" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="CCBBF5E4B9097850C5B841A5FD8E26A6" box="[151,615,906,933]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="841EA66FB9097850C5B841A5FD8E26A6" blockId="26.[151,662,906,969]" box="[151,615,906,933]" pageId="26" pageNumber="27">
<heading id="DF561103B9097850C5B841A5FD8E26A6" bold="true" box="[151,615,906,933]" fontSize="11" level="2" pageId="26" pageNumber="27" reason="4">
<emphasis id="B6D57A7DB9097850C5B841A5FD8E26A6" bold="true" box="[151,615,906,933]" pageId="26" pageNumber="27">
<taxonomicName id="43A1DDECB9097850C5B841A5FE2026A7" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,457,906,933]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="camerodidacta" status="sp. nov.">
<emphasis id="B6D57A7DB9097850C5B841A5FE2026A7" bold="true" box="[151,457,906,933]" italics="true" pageId="26" pageNumber="27">Hydraena camerodidacta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9097850C4F841A4FD8E26A6" box="[471,615,906,933]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5B84180FD7F26CB" box="[151,662,942,969]" pageId="26" pageNumber="27" type="description">
<paragraph id="841EA66FB9097850C5B84180FD7F26CB" blockId="26.[151,662,906,969]" box="[151,662,942,969]" pageId="26" pageNumber="27">
<figureCitation id="1C9ABAEAB9097850C5B84180FF0C26CB" box="[151,229,942,969]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="26" pageNumber="27">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB9097850C5DE4180FEE426CB" box="[241,269,943,969]" captionStart="FIGURE 34" captionStartId="59.[151,250,1985,2009]" captionTargetBox="[253,1333,181,1961]" captionTargetId="figure-15@59.[253,1333,181,1961]" captionTargetPageId="59" captionText="FIGURE 34. Hydraena camerodidacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284949" httpUri="https://zenodo.org/record/7284949/files/figure.png" pageId="26" pageNumber="27">34</figureCitation>
(habitus), 34 (aedeagus), 40 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5B841D8FF072137" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="841EA66FB9097850C5B841D8FF072137" blockId="26.[151,1437,1014,2013]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5B841D8FEAE2112" bold="true" box="[151,327,1014,1041]" pageId="26" pageNumber="27">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9097850C46241D8FF072137" ID-GBIF-Occurrence="3962178319" collectingDate="1982-01-30" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="26" pageNumber="27" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9097850C46241D8FE5C2113" box="[333,437,1015,1041]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9097850C73441D8FD7F2113" box="[539,662,1015,1041]" name="Cameroon" pageId="26" pageNumber="27">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9097850C78F41D8FCCB2113" box="[672,802,1015,1041]" pageId="26" pageNumber="27">N. W. Prov.</collectorName>
,
<quantity id="43590B8AB9097850C60341D8FC8E2113" box="[812,871,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="27" unit="km" value="7.0">7 km</quantity>
.
<collectorName id="2954C3B9B9097850C65F41D8FC122113" box="[880,1019,1015,1041]" pageId="26" pageNumber="27">E. Bamenda</collectorName>
, stream,
<date id="F01F80AFB9097850C17341D8FB212113" box="[1116,1224,1015,1041]" pageId="26" pageNumber="27" value="1982-01-30">
<collectingDate id="E05B7947B9097850C17341D8FB212113" box="[1116,1224,1015,1041]" pageId="26" pageNumber="27" value="1982-01-30">30.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9097850C02D41D8FA7A2113" box="[1282,1427,1015,1041]" pageId="26" pageNumber="27">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9097850C5B04634FF0B2137" box="[159,226,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="26" pageNumber="27" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E84611FE8721EB" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="841EA66FB9097850C5E84611FE8721EB" blockId="26.[151,1437,1014,2013]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E84611FE3D215A" bold="true" box="[199,468,1086,1112]" pageId="26" pageNumber="27">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9097850C10B4610FB70215B" box="[1060,1177,1087,1113]" name="Cameroon" pageId="26" pageNumber="27">Cameroon</collectingCountry>
by the combination of the comparatively large body size (ca.
<quantity id="43590B8AB9097850C773464DFD2E217F" box="[604,711,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="26" pageNumber="27" unit="mm" value="1.65">1.65 mm</quantity>
), the finely sparsely punctulate frons and pronotum, punctures very consistent in size throughout, interstices shining, the anterolateral foveae moderately deep, posterolateral and posterosubmedian foveae very shallow or absent, the very narrow plaques, with margins very ill-defined, and the aedeagus (
<figureCitation id="1C9ABAEAB9097850C42246E0FE8821EB" box="[269,353,1231,1257]" captionStart="FIGURE 34" captionStartId="59.[151,250,1985,2009]" captionTargetBox="[253,1333,181,1961]" captionTargetId="figure-15@59.[253,1333,181,1961]" captionTargetPageId="59" captionText="FIGURE 34. Hydraena camerodidacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284949" httpUri="https://zenodo.org/record/7284949/files/figure.png" pageId="26" pageNumber="27">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E846DDFC1622BB" pageId="26" pageNumber="27" type="description">
<paragraph id="841EA66FB9097850C5E846DDFF0F232F" blockId="26.[151,1437,1014,2013]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E846DDFEB8200E" bold="true" box="[199,337,1266,1292]" pageId="26" pageNumber="27">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9097850C48B46DCFDEF200F" box="[420,518,1267,1293]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.65/0.69; head width 0.38; pronotum 0.38/0.53, PA 0.42, PB 0.48; elytra 0.98/0.69. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9097850C1064738FB692033" box="[1065,1152,1303,1329]" captionStart="FIGURE 34" captionStartId="59.[151,250,1985,2009]" captionTargetBox="[253,1333,181,1961]" captionTargetId="figure-15@59.[253,1333,181,1961]" captionTargetPageId="59" captionText="FIGURE 34. Hydraena camerodidacta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284949" httpUri="https://zenodo.org/record/7284949/files/figure.png" pageId="26" pageNumber="27">Fig. 34</figureCitation>
). Dorsum brown to dark brown, head, pronotal fascia and elytra dark brown, pronotum rather widely bordered anteriorly and posteriorly in light brown, maxillary palpi testaceous, tip of last palpomere not darker. Frons and pronotum finely sparsely punctulate, punctures of pronotum slightly larger than those of frons, very consistent in size throughout, interstices shining, ca. 13xpd. Clypeus medially very finely sparsely punctulate, shining; laterally densely micropunctulate, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral and posterosubmedian foveae very shallow or absent.
</paragraph>
<paragraph id="841EA66FB9097850C5E84418FB0B23BF" blockId="26.[151,1437,1014,2013]" pageId="26" pageNumber="27">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals shining, not raised, width ca. 23xpd, as are interstices between punctures of a row. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB9097850C5B844E8FC1622BB" blockId="26.[151,1437,1014,2013]" pageId="26" pageNumber="27">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely very densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.3/1.3/1.3. P1 laminate; median carina sinuate in profile. P2 moderately wide, l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, margins very ill-defined, slightly arcuate, slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1x P2.All tibiae slender. Protibia and mesotibia very weakly arcuate. Metatibia straight. Abdominal apex symmetrical, with deep, narrow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E845EDFC2122DF" box="[199,968,1986,2013]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="841EA66FB9097850C5E845EDFC2122DF" blockId="26.[151,1437,1014,2013]" box="[199,968,1986,2013]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E845EDFE8822DE" bold="true" box="[199,353,1986,2012]" pageId="26" pageNumber="27">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9097850C7E345ECFD1522DF" box="[716,764,1987,2013]" pageId="26" pageNumber="27">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9097850C64845ECFC5222DF" box="[871,955,1987,2013]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="26" pageNumber="27">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9087851C5E842B8FF0425D7" pageId="27" pageNumber="28" type="etymology">
<paragraph id="841EA66FB9087851C5E842B8FF0425D7" blockId="27.[151,1436,151,213]" pageId="27" pageNumber="28">
<emphasis id="B6D57A7DB9087851C5E842B8FEA725B3" bold="true" box="[199,334,151,177]" pageId="27" pageNumber="28">Etymology.</emphasis>
Named in reference to the known distribution, and to the two finger-like processes of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,133 @@
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<mods:title id="FA6DFFC9C987DAAFBF16C0D1F0BC99CE">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<heading id="DF561103B90A7853C5B847C1FD83230A" bold="true" box="[151,618,1518,1545]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<emphasis id="B6D57A7DB90A7853C5B847C1FD83230A" bold="true" box="[151,618,1518,1545]" pageId="25" pageNumber="26">
<taxonomicName id="43A1DDECB90A7853C5B847C1FE25230B" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,460,1518,1545]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="camerocordata" status="sp. nov.">
<emphasis id="B6D57A7DB90A7853C5B847C1FE25230B" bold="true" box="[151,460,1518,1545]" italics="true" pageId="25" pageNumber="26">Hydraena camerocordata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90A7853C4F447C0FD83230A" box="[475,618,1518,1545]" pageId="25" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5B8443CFD7F232F" box="[151,662,1554,1581]" pageId="25" pageNumber="26" type="description">
<paragraph id="841EA66FB90A7853C5B8443CFD7F232F" blockId="25.[151,662,1518,1581]" box="[151,662,1554,1581]" pageId="25" pageNumber="26">
<figureCitation id="1C9ABAEAB90A7853C5B8443CFF0C232F" box="[151,229,1554,1581]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="25" pageNumber="26">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90A7853C5DE443CFEE4232F" box="[241,269,1555,1581]" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">33</figureCitation>
(habitus), 33 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7850C5B84474FA8427F3" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="841EA66FB90A7853C5B84474FC81239B" blockId="25.[151,1437,1626,2013]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5B84474FEA02376" bold="true" box="[151,329,1626,1653]" pageId="25" pageNumber="26">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90A7853C47F4474FE9D239B" ID-GBIF-Occurrence="3962178317" collectingDate="1982-04-22" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Etoug Ebe" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90A7853C47F4474FE512377" box="[336,440,1627,1653]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB90A7853C70C4474FD772377" box="[547,670,1627,1653]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B90A7853C7864474FCCF2377" box="[681,806,1627,1653]" pageId="25" pageNumber="26">C. S. Prov.</collectorName>
,
<location id="817EF0B4B90A7853C61D4474FC432377" LSID="urn:lsid:plazi:treatment:0C081779B90A7850C5B847C1FADD263B:817EF0B4B90A7853C61D4474FC432377" box="[818,938,1627,1653]" country="Cameroon" name="Etoug Ebe" pageId="25" pageNumber="26">Etoug Ebe</location>
, stream with dead leaves,
<date id="F01F80AFB90A7853C1FC4474FAA52377" box="[1235,1356,1627,1653]" pageId="25" pageNumber="26" value="1982-04-22">
<collectingDate id="E05B7947B90A7853C1FC4474FAA52377" box="[1235,1356,1627,1653]" pageId="25" pageNumber="26" value="1982-04-22">22.iv.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B90A7853C0A64474FEF9239B" pageId="25" pageNumber="26">P. D. Perkins</collectorName>
” (
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)
</materialsCitation>
.
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<typeStatus id="5B1A18CDB90A7853C4514450FE06239B" box="[382,495,1663,1689]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
: Same data as holotype (2
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)
</materialsCitation>
.
</paragraph>
<paragraph id="841EA66FB90A7853C5E8448CFF2D2207" blockId="25.[151,1437,1626,2013]" pageId="25" pageNumber="26">
Differential Diagnosis. Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90A7853C12B448CFB9023BF" box="[1028,1145,1699,1725]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
by the combination of the large body size (ca.
<quantity id="43590B8AB90A7853C45744E9FE0923E3" box="[376,480,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="25" pageNumber="26" unit="mm" value="1.53">1.53 mm</quantity>
), the markedly cordiform pronotum, the very wide P1 and P2, and the aedeagus (
<figureCitation id="1C9ABAEAB90A7853C04044E8FF5F2207" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="841EA66FB90A7850C5E84520FE582443" blockId="25.[151,1437,1626,2013]" lastBlockId="26.[151,1437,151,825]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Description. Size:
<typeStatus id="5B1A18CDB90A7853C4B24520FE16222B" box="[413,511,1807,1833]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.53/0.69; head width 0.36; pronotum 0.38/0.52, PA 0.41, PB 0.38; elytra 0.90/0.69. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90A7853C133451CFB98224F" box="[1052,1137,1843,1869]" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">Fig. 33</figureCitation>
). Pronotum markedly cordiform, width at midlength much wider than anterior and posterior margins. Dorsum dark brown to black, except labrum, clypeus, and maxillary palpi light brown to testaceous, tip of last palpomere not darker. Frons moderately finely moderately sparsely punctate, punctures ca. 1xef. Pronotum coarsely densely punctate, especially anterior and posterior to moderately convex disc, interstices in densely punctate areas narrow walls; interstices on disc ca. 1xpd, shining; punctures laterally much smaller and sparser. Punctures of clypeus very fine, moderately dense on disc, very dense laterally. Labrum apicomedian excision deep and wide, free margins weakly upturned, angulate laterally and medially. Pronotum anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae deep, shallowly confluent; posterosubmedial foveae deep, area between antero- and posterolateral foveae distinctly convex.
</paragraph>
<paragraph id="841EA66FB9097850C5E84364FE74271B" blockId="26.[151,1437,151,825]" pageId="26" pageNumber="27">Elytra with posterior declivity rather abrupt, summit at or very near midlength; lateral explanate margins wide; on basal 1/3 punctures 1xpd or slightly larger and deeper than largest pronotal punctures, punctures becoming gradually slightly smaller laterally and toward posterior. Intervals not raised, width on disc ca. 1xpd, much wider laterally and posteriorly, interstices between punctures of a row ca. 16xpd, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rather sharply rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="841EA66FB9097850C5B8400CFA8427F3" blockId="26.[151,1437,151,825]" pageId="26" pageNumber="27">Venter: Mentum finely sparsely punctulate, shining. Postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.1/0.2/1.5. P1 and P2 very wide. P2 l/w ca. 1/1, sides parallel, apex blunt. Plaques very small, very short, widely separated carinate lines, located at sides of deep median depression. AIS very wide, weakly concave, width at straight posterior margin ca. 1.2x P2. Tibiae slender, straight. Protibia very slightly widened and bearing short spine at ca. distal 2/3. Mesotibia slightly widened at ca. distal 2/3. Abdominal apex symmetrical, with rather wide and deep apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E840D5FC212617" box="[199,968,762,789]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="841EA66FB9097850C5E840D5FC212617" blockId="26.[151,1437,151,825]" box="[199,968,762,789]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E840D5FEB02616" bold="true" box="[199,345,762,788]" pageId="26" pageNumber="27">Distribution</emphasis>
. Currently known only from the
<typeStatus id="5B1A18CDB9097850C7E340D4FD152617" box="[716,764,763,789]" pageId="26" pageNumber="27">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9097850C64840D4FC532617" box="[871,954,763,789]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="26" pageNumber="27">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E84130FADD263B" box="[199,1332,799,825]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="841EA66FB9097850C5E84130FADD263B" blockId="26.[151,1437,151,825]" box="[199,1332,799,825]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E84130FEA1263B" bold="true" box="[199,328,799,825]" pageId="26" pageNumber="27">Etymology</emphasis>
. Named in reference to the known distribution, and to the markedly cordiform pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="036F12D130701EBA77A61544ED5F3B01">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B90A7853C5B843BDFB62209F" ID-DOI="http://doi.org/10.5281/zenodo.7300278" ID-GBIF-Taxon="204205217" ID-Zenodo-Dep="7300278" LSID="urn:lsid:plazi:treatment:0C081779B90A7853C5B843BDFB62209F" httpUri="http://treatment.plazi.org/id/0C081779B90A7853C5B843BDFB62209F" lastPageNumber="26" pageId="25" pageNumber="26">
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<paragraph id="841EA66FB90A7853C5B843BDFD9A24AE" blockId="25.[151,662,402,465]" box="[151,627,402,429]" pageId="25" pageNumber="26">
<heading id="DF561103B90A7853C5B843BDFD9A24AE" bold="true" box="[151,627,402,429]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<emphasis id="B6D57A7DB90A7853C5B843BDFD9A24AE" bold="true" box="[151,627,402,429]" pageId="25" pageNumber="26">
<taxonomicName id="43A1DDECB90A7853C5B843BDFE3D24AF" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,468,402,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cameroconvexa" status="sp. nov.">
<emphasis id="B6D57A7DB90A7853C5B843BDFE3D24AF" bold="true" box="[151,468,402,429]" italics="true" pageId="25" pageNumber="26">Hydraena cameroconvexa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90A7853C4CC43BCFD9A24AE" box="[483,627,402,429]" pageId="25" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5B84398FD7F24D3" box="[151,662,438,465]" pageId="25" pageNumber="26" type="description">
<paragraph id="841EA66FB90A7853C5B84398FD7F24D3" blockId="25.[151,662,402,465]" box="[151,662,438,465]" pageId="25" pageNumber="26">
<figureCitation id="1C9ABAEAB90A7853C5B84398FF0C24D3" box="[151,229,438,465]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="25" pageNumber="26">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90A7853C5DE4398FEE424D3" box="[241,269,439,465]" captionStart="FIGURE 32" captionStartId="57.[151,250,1990,2014]" captionTargetBox="[260,1332,197,1943]" captionTargetId="figure-15@57.[192,1395,181,1966]" captionTargetPageId="57" captionText="FIGURE 32. Hydraena cameroconvexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284943" httpUri="https://zenodo.org/record/7284943/files/figure.png" pageId="25" pageNumber="26">32</figureCitation>
(habitus), 32 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5B843D0FE2B273F" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="841EA66FB90A7853C5B843D0FE2B273F" blockId="25.[151,1437,510,1437]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5B843D0FEA0271A" bold="true" box="[151,329,510,537]" pageId="25" pageNumber="26">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90A7853C47E43D0FE2B273F" ID-GBIF-Occurrence="3962178358" collectingDate="1983-01-02" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Isali River" municipality="Nouma" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90A7853C47E43D0FE50271B" box="[337,441,511,537]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB90A7853C70A43D0FD49271B" box="[549,672,511,537]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B90A7853C78343D0FCC3271B" box="[684,810,511,537]" pageId="25" pageNumber="26">C. S. Prov.</collectorName>
,
<quantity id="43590B8AB90A7853C61943D1FC68271B" box="[822,897,510,537]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="km" value="15.0">15 km</quantity>
. NW Yaounde,
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nr.
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,
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<collectingDate id="E05B7947B90A7853C01643D0FA71271B" box="[1337,1432,511,537]" pageId="25" pageNumber="26" value="1983-01-02">2.i.1983</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B90A7853C5E8400CFEB2273F" box="[199,347,547,573]" pageId="25" pageNumber="26">P. D. Perkins</collectorName>
” (
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).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5E84069FE2E27CF" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="841EA66FB90A7853C5E84069FE2E27CF" blockId="25.[151,1437,510,1437]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5E84069FE382762" bold="true" box="[199,465,582,608]" pageId="25" pageNumber="26">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90A7853C1274068FB942763" box="[1032,1149,583,609]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
by the combination of the large body size (ca.
<quantity id="43590B8AB90A7853C4574044FE372787" box="[376,478,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="25" pageNumber="26" unit="mm" value="1.74">1.74 mm</quantity>
, largest known in
<collectingCountry id="FCB6E6FFB90A7853C7844044FCCD2787" box="[683,804,619,645]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
), the pronotum with sides markedly arcuate over middle 1/3, the oval, transversely quite convex elytra, the very wide P1 and P2, the plaques widely separated carinate lines, and the aedeagus (
<figureCitation id="1C9ABAEAB90A7853C449409CFE5327CF" box="[358,442,691,717]" captionStart="FIGURE 32" captionStartId="57.[151,250,1990,2014]" captionTargetBox="[260,1332,197,1943]" captionTargetId="figure-15@57.[192,1395,181,1966]" captionTargetPageId="57" captionText="FIGURE 32. Hydraena cameroconvexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284943" httpUri="https://zenodo.org/record/7284943/files/figure.png" pageId="25" pageNumber="26">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5E840F9FA882057" pageId="25" pageNumber="26" type="description">
<paragraph id="841EA66FB90A7853C5E840F9FEC42113" blockId="25.[151,1437,510,1437]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5E840F9FEB027F2" bold="true" box="[199,345,726,752]" pageId="25" pageNumber="26">Description.</emphasis>
Size:
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(length/width, mm): body (length to elytral apices) 1.74/0.79; head width 0.42; pronotum 0.42/0.58, PA 0.47, PB 0.48; elytra 1.05/0.79. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90A7853C12840D4FBB22617" box="[1031,1115,763,789]" captionStart="FIGURE 32" captionStartId="57.[151,250,1990,2014]" captionTargetBox="[260,1332,197,1943]" captionTargetId="figure-15@57.[192,1395,181,1966]" captionTargetPageId="57" captionText="FIGURE 32. Hydraena cameroconvexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284943" httpUri="https://zenodo.org/record/7284943/files/figure.png" pageId="25" pageNumber="26">Fig. 32</figureCitation>
). Pronotum with sides markedly arcuate over middle 1/3, sides more strongly sinuate posterior to middle than anterior to middle. Frons finely and moderately sparsely punctulate, punctures ca. 1xef, interstices ca. 13xpd, shining. Pronotum coarsely densely punctate in anterior and posterior 1/3, interstices ca. 1xpd, shining; disc convex, very finely very sparsely punctulate, interstices ca. 36xpd, shining. Punctures of clypeus very fine, sparse on disc, interstices shining, very dense, dull laterally. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin emarginate over middle 1/2; anterolateral pronotal foveae deep; posterolateral and posterosubmedial pronotal foveae very shallow.
</paragraph>
<paragraph id="841EA66FB90A7853C5E84634FA7D21A3" blockId="25.[151,1437,510,1437]" pageId="25" pageNumber="26">Elytra oval, transversely quite convex, posterior declivity abrupt, summit of at or very near midlength; lateral explanate margins wide; punctures coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 23xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB90A7853C5B84684FA882057" blockId="25.[151,1437,510,1437]" pageId="25" pageNumber="26">Venter: Mentum finely sparsely punctulate, shining. Postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/1/1. P1 and P2 very wide. P2 l/w ca. 1/1, sides parallel, apex blunt. Plaques widely separated carinate lines, located at sides of deep median depression. AIS very wide, width at straight posterior margin ca. 1.2x P2. All tibiae slender. Protibia very slightly arcuate. Meso- and metatibiae straight. Abdominal apex asymmetrical, with deep apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5E84771FC21207B" box="[199,968,1374,1401]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="841EA66FB90A7853C5E84771FC21207B" blockId="25.[151,1437,510,1437]" box="[199,968,1374,1401]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5E84771FEB0207A" bold="true" box="[199,345,1374,1400]" pageId="25" pageNumber="26">Distribution</emphasis>
. Currently known only from the
<typeStatus id="5B1A18CDB90A7853C7E34770FD15207B" box="[716,764,1375,1401]" pageId="25" pageNumber="26">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90A7853C6484770FC53207B" box="[871,954,1375,1401]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="25" pageNumber="26">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5E847ACFB62209F" box="[199,1163,1411,1437]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="841EA66FB90A7853C5E847ACFB62209F" blockId="25.[151,1437,510,1437]" box="[199,1163,1411,1437]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5E847ACFEA1209F" bold="true" box="[199,328,1411,1437]" pageId="25" pageNumber="26">Etymology</emphasis>
. Named in reference to the known distribution, and to the convex dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="962263BA60A319E9C0E0F79D6DB345EB">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<taxonomicName id="43A1DDECB90B7852C5B841D9FE352112" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,476,1014,1041]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="camerocontinua" status="sp. nov.">
<emphasis id="B6D57A7DB90B7852C5B841D9FE352112" bold="true" box="[151,476,1014,1041]" italics="true" pageId="24" pageNumber="25">Hydraena camerocontinua</emphasis>
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,
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<figureCitation id="1C9ABAEAB90B7852C5B84635FF0C2137" box="[151,229,1050,1077]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="24" pageNumber="25">Figs. 5</figureCitation>
,
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(habitus), 31 (aedeagus), 42 (map)
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<emphasis id="B6D57A7DB90B7852C5B8464DFEAE217E" bold="true" box="[151,327,1122,1148]" pageId="24" pageNumber="25">Type Material.</emphasis>
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(male): “
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,
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, leg.
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Differential Diagnosis. Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90B7852C12B4685FB9021C6" box="[1028,1145,1194,1220]" name="Cameroon" pageId="24" pageNumber="25">Cameroon</collectingCountry>
by the combination of the relatively wide and short P2, l/w ca. 1/1, sides slightly diverging toward apex, apex very indistinct, fused with and at same level as mesoventral intercoxal process., the anterolateral pronotal foveae moderately deep, posterolateral pronotal foveae very shallow or absent, and the aedeagus (
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). Somewhat similar in habitus to
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<emphasis id="B6D57A7DB90B7852C1CB4738FA7F2032" box="[1252,1430,1302,1328]" italics="true" pageId="24" pageNumber="25">H. cameroarida</emphasis>
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, but differing in finer, sparser dorsal punctation, ventral characters, and aedeagal morphology.
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Description. Size:
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(length/width, mm): body (length to elytral apices) 1.47/0.67; head width 0.39; pronotum 0.37/0.52, PA 0.42, PB 0.47; elytra 0.91/0.67. Habitus as illustrated (
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). Dorsum brown to dark brown or black, frons and diffusely margined pronotal macula darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely moderately finely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae moderately deep, posterolateral pronotal foveae very shallow or absent, posterosubmedial foveae very shallow.
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<paragraph id="841EA66FB90B7852C5E84451FE36222A" blockId="24.[151,1437,1122,2013]" pageId="24" pageNumber="25">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 23xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB90B7853C5B8451CFC8F25D7" blockId="24.[151,1437,1122,2013]" lastBlockId="25.[151,1436,151,321]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.3/1/1.3. P1 laminate; median carina sinuate in profile. P2 relatively wide and short, l/w ca. 1/1, sides slightly diverging toward apex, apex very indistinct, fused with and at same level as mesoventral intercoxal process. Plaques narrow, slightly converging toward one another anteriorly, very weakly raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.3x P2. Protibia slender, arcuate. Meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
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<emphasis id="B6D57A7DB90A7853C5E842F1FEB025FA" bold="true" box="[199,345,222,248]" pageId="25" pageNumber="26">Distribution</emphasis>
. Currently known only from the
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locality (
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</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7853C5E8432CFE8C2443" pageId="25" pageNumber="26" type="etymology">
<paragraph id="841EA66FB90A7853C5E8432CFE8C2443" blockId="25.[151,1436,151,321]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5E8432CFEA1241F" bold="true" box="[199,328,259,285]" pageId="25" pageNumber="26">Etymology</emphasis>
. Named in reference to the known distribution, and to the fused intercoxal processes of the meso- and metaventrites.
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<paragraph id="841EA66FB90F7856C5B8409DFD8F27CE" blockId="28.[151,662,690,753]" box="[151,614,690,717]" pageId="28" pageNumber="29">
<heading id="DF561103B90F7856C5B8409DFD8F27CE" bold="true" box="[151,614,690,717]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B6D57A7DB90F7856C5B8409DFD8F27CE" bold="true" box="[151,614,690,717]" pageId="28" pageNumber="29">
<taxonomicName id="43A1DDECB90F7856C5B8409DFE2E27CF" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,455,690,717]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cameroradiata" status="sp. nov.">
<emphasis id="B6D57A7DB90F7856C5B8409DFE2E27CF" bold="true" box="[151,455,690,717]" italics="true" pageId="28" pageNumber="29">Hydraena cameroradiata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90F7856C4F9409CFD8F27CE" box="[470,614,690,717]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B90F7856C5B840F8FD7F27F3" box="[151,662,726,753]" pageId="28" pageNumber="29" type="description">
<paragraph id="841EA66FB90F7856C5B840F8FD7F27F3" blockId="28.[151,662,690,753]" box="[151,662,726,753]" pageId="28" pageNumber="29">
<figureCitation id="1C9ABAEAB90F7856C5B840F8FF0C27F3" box="[151,229,726,753]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="28" pageNumber="29">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90F7856C5DE40F8FEE427F3" box="[241,269,727,753]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">37</figureCitation>
(habitus), 37 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5B84130FD05265F" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="841EA66FB90F7856C5B84130FD05265F" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5B84130FEAE263A" bold="true" box="[151,327,798,825]" pageId="28" pageNumber="29">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90F7856C4624130FF03265F" ID-GBIF-Occurrence="3962178355" collectingDate="1981-12-12" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Yaounde" municipality="Bastos" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90F7856C4624130FE5C263B" box="[333,437,799,825]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB90F7856C7334130FD7E263B" box="[540,663,799,825]" name="Cameroon" pageId="28" pageNumber="29">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B90F7856C78E4130FCF2263B" box="[673,795,799,825]" pageId="28" pageNumber="29">C. S. Prov.</collectorName>
,
<location id="817EF0B4B90F7856C60B4130FC62263B" LSID="urn:lsid:plazi:treatment:0C081779B90F7856C5B8409DFB1823E3:817EF0B4B90F7856C60B4130FC62263B" box="[804,907,799,825]" country="Cameroon" municipality="Bastos" name="Yaounde" pageId="28" pageNumber="29">Yaounde</location>
:
<collectingMunicipality id="647A3C15B90F7856C6BB4130FC08263B" box="[916,993,799,825]" pageId="28" pageNumber="29">Bastos</collectingMunicipality>
, puddle,
<date id="F01F80AFB90F7856C16A4130FB2E263B" box="[1093,1223,799,825]" pageId="28" pageNumber="29" value="1981-12-12">
<collectingDate id="E05B7947B90F7856C16A4130FB2E263B" box="[1093,1223,799,825]" pageId="28" pageNumber="29" value="1981-12-12">12.xii.1981</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B90F7856C02E4130FA7A263B" box="[1281,1427,799,825]" pageId="28" pageNumber="29">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB90F7856C5B0416CFF0B265F" box="[159,226,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B90F7856C5DA416CFD01265F" ID-GBIF-Occurrence="3962178354" box="[245,744,835,861]" collectingDate="1981-12-12" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Yaounde" municipality="Bastos" pageId="28" pageNumber="29" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB90F7856C5DA416CFE8F265F" box="[245,358,835,861]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
: Same data as holotype (10
<collectionCode id="E2B03EAAB90F7856C7B0416CFD08265F" box="[671,737,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E84149FE2E26EF" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="841EA66FB90F7856C5E84149FE2E26EF" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E84149FE382682" bold="true" box="[199,465,870,896]" pageId="28" pageNumber="29">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90F7856C1274148FB942683" box="[1032,1149,871,897]" name="Cameroon" pageId="28" pageNumber="29">Cameroon</collectingCountry>
by the combination of the very sparcely and very finely punctulate head and pronotum, interstices strongly shining, the anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent, the absence of metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB90F7856C44941FCFE5326EF" box="[358,442,979,1005]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E841D9FDE3239B" pageId="28" pageNumber="29" type="description">
<paragraph id="841EA66FB90F7856C5E841D9FDDD2033" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E841D9FEB82112" bold="true" box="[199,337,1014,1040]" pageId="28" pageNumber="29">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB90F7856C48B41D8FDEF2113" box="[420,518,1015,1041]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.33/0.59; head width 0.34; pronotum 0.35/0.47, PA 0.38, PB 0.43; elytra 0.80/0.59. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90F7856C1074634FB962137" box="[1064,1151,1051,1077]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">Fig. 37</figureCitation>
). Dorsum brown to light brown, frons and diffusely margined pronotal macula slightly darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely finely punctulate, pronotum more finely sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of pronotal disc ca. 1xef, interstices ca. 35xpd or slightly larger; punctures of frons disc slightly larger and more closely spaced than those of pronotal disc; punctures of clypeus very fine, nearly obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent.
</paragraph>
<paragraph id="841EA66FB90F7856C5E84714FDD420E7" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins narrow; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 24xpd, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming moderately strong angle with one another.</paragraph>
<paragraph id="841EA66FB90F7856C5B847C0FDE3239B" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">Venter: Mentum and postmentum sparsely finely punctulate, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques absent, median depression deep. AIS width at straight posterior margin ca. 2x P2. All tibiae slender. Pro- and mesotibiae very slightly arcuate, metatibia straight. Abdominal apex symmetrical, with shallow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E8448DFC2123BF" box="[199,968,1698,1725]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="841EA66FB90F7856C5E8448DFC2123BF" blockId="28.[151,1437,798,1761]" box="[199,968,1698,1725]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E8448DFE8823BE" bold="true" box="[199,353,1698,1724]" pageId="28" pageNumber="29">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB90F7856C7E3448CFD1523BF" box="[716,764,1699,1725]" pageId="28" pageNumber="29">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90F7856C648448CFC5323BF" box="[871,954,1699,1725]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="28" pageNumber="29">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E844E8FB1823E3" box="[199,1265,1735,1761]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="841EA66FB90F7856C5E844E8FB1823E3" blockId="28.[151,1437,798,1761]" box="[199,1265,1735,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E844E8FEA723E3" bold="true" box="[199,334,1735,1761]" pageId="28" pageNumber="29">Etymology.</emphasis>
Named in reference to the known distribution, and to the highly reflective dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,167 @@
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<mods:title id="A52C712C29B3FF1797B3EEEA68A404A4">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<paragraph id="841EA66FB90F7856C5B84555FDB02296" blockId="28.[151,662,1914,1977]" box="[151,601,1914,1941]" pageId="28" pageNumber="29">
<heading id="DF561103B90F7856C5B84555FDB02296" bold="true" box="[151,601,1914,1941]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B6D57A7DB90F7856C5B84555FDB02296" bold="true" box="[151,601,1914,1941]" pageId="28" pageNumber="29">
<taxonomicName id="43A1DDECB90F7856C5B84555FE532297" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,442,1914,1941]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="camerosetosa" status="sp. nov.">
<emphasis id="B6D57A7DB90F7856C5B84555FE532297" bold="true" box="[151,442,1914,1941]" italics="true" pageId="28" pageNumber="29">Hydraena camerosetosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90F7856C4E64554FDB02296" box="[457,601,1914,1941]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="CCBBF5E4B90F7856C5B845B0FD7F22BB" box="[151,662,1950,1977]" pageId="28" pageNumber="29" type="description">
<paragraph id="841EA66FB90F7856C5B845B0FD7F22BB" blockId="28.[151,662,1914,1977]" box="[151,662,1950,1977]" pageId="28" pageNumber="29">
<figureCitation id="1C9ABAEAB90F7856C5B845B0FF0C22BB" box="[151,229,1950,1977]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="28" pageNumber="29">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90F7856C5DE45B0FEE422BB" box="[241,269,1951,1977]" captionStart="FIGURE 38" captionStartId="63.[151,250,1962,1986]" captionTargetBox="[296,1277,188,1925]" captionTargetId="figure-15@63.[226,1361,181,1939]" captionTargetPageId="63" captionText="FIGURE 38. Hydraena camerosetosa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284961" httpUri="https://zenodo.org/record/7284961/files/figure.png" pageId="28" pageNumber="29">38</figureCitation>
(habitus), 38 (aedeagus), 43 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90E7857C5B842B9FB6625D6" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="841EA66FB90E7857C5B842B9FB6625D6" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="30">
<emphasis id="B6D57A7DB90E7857C5B842B9FEAE25B2" bold="true" box="[151,327,150,176]" pageId="29" pageNumber="30">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90E7857C46342B9FD8925D6" ID-GBIF-Occurrence="3962178359" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90E7857C46342B9FE5D25B2" box="[332,436,150,176]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
(male):
<location id="817EF0B4B90E7857C73E42B9FD6325B2" LSID="urn:lsid:plazi:treatment:0C081779B90F7857C5B84555FB43217E:817EF0B4B90E7857C73E42B9FD6325B2" box="[529,650,150,176]" country="Cameroon" municipality="Northwest Province" name="Cameroon" pageId="29" pageNumber="30">Cameroon</location>
:
<collectingMunicipality id="647A3C15B90E7857C7BC42B9FC9D25B2" box="[659,884,150,176]" pageId="29" pageNumber="30">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB90E7857C6A842B9FB4525B2" box="[903,1196,150,176]" name="Cameroon" pageId="29" pageNumber="30">BRITISH CAMEROONS</collectingCountry>
/ Matute, / Tiko Plantation /
<date id="F01F80AFB90E7857C5C24295FEA625D7" box="[237,335,186,213]" pageId="29" pageNumber="30" value="1949-05-05">
<collectingDate id="E05B7947B90E7857C5C24295FEA625D7" box="[237,335,186,213]" pageId="29" pageNumber="30" value="1949-05-05">5.v.1949</collectingDate>
</date>
,
<collectorName id="2954C3B9B90E7857C4744294FE2625D6" box="[347,463,186,213]" pageId="29" pageNumber="30">B. Malkin</collectorName>
” (
<collectionCode id="E2B03EAAB90E7857C4C74295FDB025D6" box="[488,601,186,212]" pageId="29" pageNumber="30">UKNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B90E7857C7454294FB6525D6" ID-GBIF-Occurrence="3962178304" box="[618,1164,186,213]" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="29" pageNumber="30" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB90E7857C7454294FD3225D7" box="[618,731,187,213]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
: Same data as holotype (10
<collectionCode id="E2B03EAAB90E7857C13B4295FB6A25D6" box="[1044,1155,186,212]" pageId="29" pageNumber="30">UKNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90E7857C5E842F1FD9424D3" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="841EA66FB90E7857C5E842F1FD9424D3" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="30">
<emphasis id="B6D57A7DB90E7857C5E842F1FE3D25FA" bold="true" box="[199,468,222,248]" pageId="29" pageNumber="30">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90E7857C10B42F1FB7025FA" box="[1060,1177,222,248]" name="Cameroon" pageId="29" pageNumber="30">Cameroon</collectingCountry>
by the combination of the densely punctate dorsum, the absence of metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB90E7857C165432DFB77241E" box="[1098,1182,258,284]" captionStart="FIGURE 38" captionStartId="63.[151,250,1962,1986]" captionTargetBox="[296,1277,188,1925]" captionTargetId="figure-15@63.[226,1361,181,1939]" captionTargetPageId="63" captionText="FIGURE 38. Hydraena camerosetosa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284961" httpUri="https://zenodo.org/record/7284961/files/figure.png" pageId="29" pageNumber="30">Fig. 38</figureCitation>
).
<taxonomicName id="43A1DDECB90E7857C19D432CFA84241E" authorityName="Perkins" authorityYear="2022" box="[1202,1389,259,284]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB90E7857C19D432CFA84241E" box="[1202,1389,259,284]" italics="true" pageId="29" pageNumber="30">H. camerosetosa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="43A1DDECB90E7857C4284308FE572442" authorityName="Perkins" authorityYear="2022" box="[263,446,294,320]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB90E7857C4284308FE572442" box="[263,446,294,320]" italics="true" pageId="29" pageNumber="30">H. cameropetila</emphasis>
</taxonomicName>
, another species that lacks metaventral plaques, by a larger body size (ca. 1.33 vs.
<quantity id="43590B8AB90E7857C0444309FF2E2466" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="mm" value="1.2">1.20 mm</quantity>
), a markedly different aedeagus, and a wider mesoventral intercoxal process (P2), which is twice as wide as the P2 of
<taxonomicName id="43A1DDECB90E7857C5F44340FE78248A" authorityName="Perkins" authorityYear="2022" box="[219,401,366,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB90E7857C5F44340FE78248A" box="[219,401,366,392]" italics="true" pageId="29" pageNumber="30">H. cameropetila</emphasis>
</taxonomicName>
. Although
<taxonomicName id="43A1DDECB90E7857C7214340FD2D248A" authorityName="Perkins" authorityYear="2022" box="[526,708,366,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB90E7857C7214340FD2D248A" box="[526,708,366,392]" italics="true" pageId="29" pageNumber="30">H. cameropetila</emphasis>
</taxonomicName>
is smaller than
<taxonomicName id="43A1DDECB90E7857C6584340FBDB248A" authorityName="Perkins" authorityYear="2022" box="[887,1074,367,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB90E7857C6584340FBDB248A" box="[887,1074,367,392]" italics="true" pageId="29" pageNumber="30">H. camerosetosa</emphasis>
</taxonomicName>
in body length, the width of the mesocoxae are the same in the two species, making the ratio of P2 width/mesocoxa width approximately 0.12 for
<taxonomicName id="43A1DDECB90E7857C5B84398FEA524D2" authorityName="Perkins" authorityYear="2022" box="[151,332,438,464]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB90E7857C5B84398FEA524D2" box="[151,332,438,464]" italics="true" pageId="29" pageNumber="30">H. cameropetila</emphasis>
</taxonomicName>
and 0.40
<taxonomicName id="43A1DDECB90E7857C4944398FD9F24D2" authorityName="Perkins" authorityYear="2022" box="[443,630,439,464]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="camerosetosa">
<emphasis id="B6D57A7DB90E7857C4944398FD9F24D2" box="[443,630,439,464]" italics="true" pageId="29" pageNumber="30">H. camerosetosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90E7857C5E843F5FD442136" pageId="29" pageNumber="30" type="description">
<paragraph id="841EA66FB90E7857C5E843F5FB9227CE" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="30">
<emphasis id="B6D57A7DB90E7857C5E843F5FEB824F6" bold="true" box="[199,337,474,500]" pageId="29" pageNumber="30">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB90E7857C48B43F5FDEF24F6" box="[420,518,474,500]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.33/0.55; head width 0.34; pronotum 0.34/0.46, PA 0.39, PB 0.42; elytra 0.78/0.55. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90E7857C6D043D1FBBB271A" box="[1023,1106,510,536]" captionStart="FIGURE 38" captionStartId="63.[151,250,1962,1986]" captionTargetBox="[296,1277,188,1925]" captionTargetId="figure-15@63.[226,1361,181,1939]" captionTargetPageId="63" captionText="FIGURE 38. Hydraena camerosetosa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284961" httpUri="https://zenodo.org/record/7284961/files/figure.png" pageId="29" pageNumber="30">Fig. 38</figureCitation>
). Dorsum testaceous to black, head and pronotal macula dark brown to black, pronotum rather widely margined in light brown to testaceous, elytra brown, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely sparsely punctulate, pronotum moderately coarsely, densely punctate, punctures much larger than those of frons, interstices shining, ca. 1xpd or less. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB90E7857C5E840F9FB86265E" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="30">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming distinct angle with one another.</paragraph>
<paragraph id="841EA66FB90E7857C5B84148FD442136" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="30">Venter: Mentum finely punctulate, shining. Postmentum densely micropunctulate, dull. Genae raised, dull, without posterior ridge. Metaventral plaques absent. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with apicomedian indentation.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90E7857C5E84611FC21215A" box="[199,968,1086,1113]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="841EA66FB90E7857C5E84611FC21215A" blockId="29.[151,1437,150,1149]" box="[199,968,1086,1113]" pageId="29" pageNumber="30">
<emphasis id="B6D57A7DB90E7857C5E84611FE88215A" bold="true" box="[199,353,1086,1112]" pageId="29" pageNumber="30">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB90E7857C7E34610FD15215B" box="[716,764,1087,1113]" pageId="29" pageNumber="30">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90E7857C6484611FC52215A" box="[871,955,1086,1112]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="29" pageNumber="30">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90E7857C5E8464DFB43217E" box="[199,1194,1122,1149]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="841EA66FB90E7857C5E8464DFB43217E" blockId="29.[151,1437,150,1149]" box="[199,1194,1122,1149]" pageId="29" pageNumber="30">
<emphasis id="B6D57A7DB90E7857C5E8464DFEA7217E" bold="true" box="[199,334,1122,1148]" pageId="29" pageNumber="30">Etymology.</emphasis>
Named in reference to the entirely setose metaventrite, which lacks plaques.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,140 @@
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<mods:title id="8322A6EA690F38C2B7BC919535ADD029">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9187846C5B8464DFA78241F" ID-DOI="http://doi.org/10.5281/zenodo.7300190" ID-GBIF-Taxon="204205213" ID-Zenodo-Dep="7300190" LSID="urn:lsid:plazi:treatment:0C081779B9187846C5B8464DFA78241F" httpUri="http://treatment.plazi.org/id/0C081779B9187846C5B8464DFA78241F" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="CCBBF5E4B9187841C5B8464DFDBE217E" box="[151,599,1122,1149]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="841EA66FB9187841C5B8464DFDBE217E" blockId="11.[151,660,1122,1185]" box="[151,599,1122,1149]" pageId="11" pageNumber="12">
<heading id="DF561103B9187841C5B8464DFDBE217E" bold="true" box="[151,599,1122,1149]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="B6D57A7DB9187841C5B8464DFDBE217E" bold="true" box="[151,599,1122,1149]" pageId="11" pageNumber="12">
<taxonomicName id="43A1DDECB9187841C5B8464DFE51217F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,440,1122,1149]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="camerosicula" status="sp. nov.">
<emphasis id="B6D57A7DB9187841C5B8464DFE51217F" bold="true" box="[151,440,1122,1149]" italics="true" pageId="11" pageNumber="12">Hydraena camerosicula</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9187841C4E8464CFDBE217E" box="[455,599,1122,1149]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9187841C5B846A8FD7D21A3" box="[151,660,1159,1185]" pageId="11" pageNumber="12" type="description">
<paragraph id="841EA66FB9187841C5B846A8FD7D21A3" blockId="11.[151,660,1122,1185]" box="[151,660,1159,1185]" pageId="11" pageNumber="12">
<figureCitation id="1C9ABAEAB9187841C5B846A8FF0A21A3" box="[151,227,1159,1185]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="11" pageNumber="12">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB9187841C5DE46A8FEE521A3" box="[241,268,1159,1185]" captionStart="FIGURE 11" captionStartId="36.[151,250,1995,2019]" captionTargetBox="[277,1310,181,1971]" captionTargetId="figure-25@36.[277,1310,181,1971]" captionTargetPageId="36" captionText="FIGURE 11. Hydraena camerosicula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284872" httpUri="https://zenodo.org/record/7284872/files/figure.png" pageId="11" pageNumber="12">11</figureCitation>
(habitus), 11 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9187841C5B846E0FD37200F" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="841EA66FB9187841C5B846E0FD37200F" blockId="11.[151,1437,1230,2049]" pageId="11" pageNumber="12">
<emphasis id="B6D57A7DB9187841C5B846E0FEAE21EA" bold="true" box="[151,327,1230,1257]" pageId="11" pageNumber="12">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9187841C46246E0FF03200F" ID-GBIF-Occurrence="3962178328" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9187841C46246E0FE5C21EB" box="[333,437,1231,1257]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9187841C73346E0FD7E21EB" box="[540,663,1231,1257]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9187841C78E46E0FCE921EB" box="[673,768,1231,1257]" pageId="11" pageNumber="12">N. Prov.</collectorName>
,
<quantity id="43590B8AB9187841C62546E1FCB921EB" box="[778,848,1230,1257]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B9187841C67946E0FBEA21EB" box="[854,1027,1231,1257]" pageId="11" pageNumber="12">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB9187841C14B46E0FB2E21EB" box="[1124,1223,1231,1257]" pageId="11" pageNumber="12" value="1982-05-01">
<collectingDate id="E05B7947B9187841C14B46E0FB2E21EB" box="[1124,1223,1231,1257]" pageId="11" pageNumber="12" value="1982-05-01">1.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9187841C02E46E0FA7A21EB" box="[1281,1427,1231,1257]" pageId="11" pageNumber="12">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9187841C5B046DCFF0B200F" box="[159,226,1267,1293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="12" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9187841C5DA46DCFD33200F" ID-GBIF-Occurrence="3962178324" box="[245,730,1267,1293]" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="11" pageNumber="12" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9187841C5DA46DCFE8F200F" box="[245,358,1267,1293]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
: Same data as holotype (3
<collectionCode id="E2B03EAAB9187841C7BE46DCFD3A200F" box="[657,723,1267,1293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="12" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9187841C5E84739FBF3207B" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="841EA66FB9187841C5E84739FBF3207B" blockId="11.[151,1437,1230,2049]" pageId="11" pageNumber="12">
<emphasis id="B6D57A7DB9187841C5E84739FE382032" bold="true" box="[199,465,1302,1328]" pageId="11" pageNumber="12">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9187841C1274738FB942033" box="[1032,1149,1303,1329]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
by the combination of the comparatively large body size (ca.
<quantity id="43590B8AB9187841C7344714FD6B2057" box="[539,642,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="11" pageNumber="12" unit="mm" value="1.49">1.49 mm</quantity>
), the deep pronotal impressions, the comparatively wide P2, the narrow median depression between the metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB9187841C6954770FBE5207B" box="[954,1036,1375,1401]" captionStart="FIGURE 11" captionStartId="36.[151,250,1995,2019]" captionTargetBox="[277,1310,181,1971]" captionTargetId="figure-25@36.[277,1310,181,1971]" captionTargetPageId="36" captionText="FIGURE 11. Hydraena camerosicula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284872" httpUri="https://zenodo.org/record/7284872/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9187846C5E847ADFE2E25D7" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="841EA66FB9187841C5E847ADFE86239B" blockId="11.[151,1437,1230,2049]" pageId="11" pageNumber="12">
<emphasis id="B6D57A7DB9187841C5E847ADFEB8209E" bold="true" box="[199,337,1410,1436]" pageId="11" pageNumber="12">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9187841C48B47ACFDEF209F" box="[420,518,1411,1437]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.49/0.69; head width 0.39; pronotum 0.37/0.51, PA 0.41, PB 0.46; elytra 0.93/0.69. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9187841C1054788FB9720C3" box="[1066,1150,1447,1473]" captionStart="FIGURE 11" captionStartId="36.[151,250,1995,2019]" captionTargetBox="[277,1310,181,1971]" captionTargetId="figure-25@36.[277,1310,181,1971]" captionTargetPageId="36" captionText="FIGURE 11. Hydraena camerosicula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284872" httpUri="https://zenodo.org/record/7284872/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
). Dorsum brown to dark brown, frons and pronotal macula darker, maxillary palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately finely moderately sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, confluent with shallow posterolateral; posterosubmedial pronotal foveae moderately deep, markedly punctate.
</paragraph>
<paragraph id="841EA66FB9187841C5E8448CFEC8224F" blockId="11.[151,1437,1230,2049]" pageId="11" pageNumber="12">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB9187846C5B84578FE2E25D7" blockId="11.[151,1437,1230,2049]" lastBlockId="12.[151,1436,150,285]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/1. P1 laminate; median carina sinuate in profile. P2 comparatively wide, l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, slightly arcuate, parallel, tapering anteriorly, weakly raised, located at sides of moderately deep, narrow median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slender, straight. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E842F1FC2125FB" box="[199,968,222,249]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="841EA66FB91F7846C5E842F1FC2125FB" blockId="12.[151,1436,150,285]" box="[199,968,222,249]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E842F1FE8825FA" bold="true" box="[199,353,222,248]" pageId="12" pageNumber="13">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91F7846C7E342F0FD1525FB" box="[716,764,223,249]" pageId="12" pageNumber="13">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91F7846C64842F0FC5225FB" box="[871,955,223,249]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="12" pageNumber="13">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E8432CFA78241F" box="[199,1425,259,285]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="841EA66FB91F7846C5E8432CFA78241F" blockId="12.[151,1436,150,285]" box="[199,1425,259,285]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E8432CFEA7241F" bold="true" box="[199,334,259,285]" pageId="12" pageNumber="13">Etymology.</emphasis>
Named in reference to the known distribution, and to the dagger-shaped process of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,132 @@
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<mods:title id="F8595ECB856970F7F1C988FF5422E535">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9197840C5B84069FAF02377" ID-DOI="http://doi.org/10.5281/zenodo.7300186" ID-GBIF-Taxon="204205212" ID-Zenodo-Dep="7300186" LSID="urn:lsid:plazi:treatment:0C081779B9197840C5B84069FAF02377" httpUri="http://treatment.plazi.org/id/0C081779B9197840C5B84069FAF02377" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="CCBBF5E4B9197840C5B84069FDB32762" box="[151,602,582,609]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="841EA66FB9197840C5B84069FDB32762" blockId="10.[151,634,582,645]" box="[151,602,582,609]" pageId="10" pageNumber="11">
<heading id="DF561103B9197840C5B84069FDB32762" bold="true" box="[151,602,582,609]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="B6D57A7DB9197840C5B84069FDB32762" bold="true" box="[151,602,582,609]" pageId="10" pageNumber="11">
<taxonomicName id="43A1DDECB9197840C5B84069FE552763" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,444,582,609]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="camerojocula" status="sp. nov.">
<emphasis id="B6D57A7DB9197840C5B84069FE552763" bold="true" box="[151,444,582,609]" italics="true" pageId="10" pageNumber="11">Hydraena camerojocula</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9197840C4E54068FDB32762" box="[458,602,582,609]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5B84044FD932787" box="[151,634,619,645]" pageId="10" pageNumber="11" type="description">
<paragraph id="841EA66FB9197840C5B84044FD932787" blockId="10.[151,634,582,645]" box="[151,634,619,645]" pageId="10" pageNumber="11">
<figureCitation id="1C9ABAEAB9197840C5B84044FF0A2787" box="[151,227,619,645]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB9197840C5DE4044FF162787" box="[241,255,619,645]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">9</figureCitation>
(habitus), 9 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5B8409CFDD627F3" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="841EA66FB9197840C5B8409CFDD627F3" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5B8409CFEAE27CE" bold="true" box="[151,327,690,717]" pageId="10" pageNumber="11">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9197840C462409CFDD627F3" ID-GBIF-Occurrence="3962178356" collectingDate="1982-05-22" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Nyong River" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9197840C462409CFE5C27CF" box="[333,437,691,717]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9197840C732409CFD7127CF" box="[541,664,691,717]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9197840C78D409CFCF527CF" box="[674,796,691,717]" pageId="10" pageNumber="11">C. S. Prov.</collectorName>
,
<quantity id="43590B8AB9197840C60A409DFC8627CF" box="[805,879,690,717]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="km" value="15.0">15 km</quantity>
.
<collectorName id="2954C3B9B9197840C656409CFBF827CF" box="[889,1041,691,717]" pageId="10" pageNumber="11">S. Mbalmayo</collectorName>
,
<location id="817EF0B4B9197840C133409CFB4727CF" LSID="urn:lsid:plazi:treatment:0C081779B9197840C5B84069FAF02377:817EF0B4B9197840C133409CFB4727CF" box="[1052,1198,691,717]" country="Cameroon" name="Nyong River" pageId="10" pageNumber="11">Nyong River</location>
, ex. leaves at margin,
<date id="F01F80AFB9197840C5B840F8FEE127F3" box="[151,264,727,753]" pageId="10" pageNumber="11" value="1982-05-22">
<collectingDate id="E05B7947B9197840C5B840F8FEE127F3" box="[151,264,727,753]" pageId="10" pageNumber="11" value="1982-05-22">22.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9197840C46B40F8FE3E27F3" box="[324,471,727,753]" pageId="10" pageNumber="11">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9197840C4DF40F8FDDA27F3" box="[496,563,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="11" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5E840D5FABA2683" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="841EA66FB9197840C5E840D5FABA2683" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E840D5FE3D2616" bold="true" box="[199,468,762,788]" pageId="10" pageNumber="11">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9197840C10B40D4FB702617" box="[1060,1177,763,789]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
by the combination of the posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression, the metaventral plaques moderately narrow posteriorly, tapering to point anteriorly, and the aedeagus (
<figureCitation id="1C9ABAEAB9197840C17F416CFB7A265F" box="[1104,1171,835,861]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
). Dorsal habitus similar to other species in Aedeagal Group 1; reliable determinations will require examination of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5E841A5FD4E232F" pageId="10" pageNumber="11" type="description">
<paragraph id="841EA66FB9197840C5E841A5FDB221A3" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E841A5FEB826A6" bold="true" box="[199,337,906,932]" pageId="10" pageNumber="11">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9197840C48B41A4FDEF26A7" box="[420,518,907,933]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.35/0.60; head width 0.36; pronotum 0.32/0.46, PA 0.37, PB 0.41; elytra 0.82/0.60. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9197840C11D4180FB9326CB" box="[1074,1146,943,969]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
). Dorsum generally dark brown, except labrum, clypeus and anterolateral angles of pronotum lighter, palpi testaceous, tip of last palpomere not darker. Frons moderately finely and moderately densely punctulate, pronotum moderately coarsely, moderately densely punctate, punctures much larger than those of frons, interstices shining, ca. 12xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral foveae very shallow, posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression.
</paragraph>
<paragraph id="841EA66FB9197840C5E84684FD4E232F" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderately strong angle with one another. Venter: Mentum sparsely finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow posteriorly, tapering to point anteriorly, slightly converging toward one another anteriorly, weakly raised, located at sides of shallow median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5E84419FC212353" box="[199,968,1590,1617]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="841EA66FB9197840C5E84419FC212353" blockId="10.[151,1437,690,1653]" box="[199,968,1590,1617]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E84419FE882352" bold="true" box="[199,353,1590,1616]" pageId="10" pageNumber="11">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9197840C7E34418FD152353" box="[716,764,1591,1617]" pageId="10" pageNumber="11">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9197840C6484418FC532353" box="[871,954,1591,1617]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="10" pageNumber="11">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5E84474FAF02377" box="[199,1305,1627,1653]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="841EA66FB9197840C5E84474FAF02377" blockId="10.[151,1437,690,1653]" box="[199,1305,1627,1653]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E84474FEA72377" bold="true" box="[199,334,1627,1653]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named in reference to the known distribution, and to the smile shaped pronotal fovea.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,143 @@
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<mods:title id="94AF714C75DCEAD61C25EBE0C0E06710">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B9197841C5B844E9FB622113" ID-DOI="http://doi.org/10.5281/zenodo.7300188" ID-GBIF-Taxon="204205211" ID-Zenodo-Dep="7300188" LSID="urn:lsid:plazi:treatment:0C081779B9197841C5B844E9FB622113" httpUri="http://treatment.plazi.org/id/0C081779B9197841C5B844E9FB622113" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="CCBBF5E4B9197840C5B844E9FDB923E2" box="[151,592,1734,1761]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="841EA66FB9197840C5B844E9FDB923E2" blockId="10.[151,662,1734,1797]" box="[151,592,1734,1761]" pageId="10" pageNumber="11">
<heading id="DF561103B9197840C5B844E9FDB923E2" bold="true" box="[151,592,1734,1761]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="B6D57A7DB9197840C5B844E9FDB923E2" bold="true" box="[151,592,1734,1761]" pageId="10" pageNumber="11">
<taxonomicName id="43A1DDECB9197840C5B844E9FE5B23E3" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,434,1734,1761]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cameroovala" status="sp. nov.">
<emphasis id="B6D57A7DB9197840C5B844E9FE5B23E3" bold="true" box="[151,434,1734,1761]" italics="true" pageId="10" pageNumber="11">Hydraena cameroovala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B9197840C4EE44E8FDB923E2" box="[449,592,1734,1761]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5B844C4FD7F2207" box="[151,662,1771,1797]" pageId="10" pageNumber="11" type="description">
<paragraph id="841EA66FB9197840C5B844C4FD7F2207" blockId="10.[151,662,1734,1797]" box="[151,662,1771,1797]" pageId="10" pageNumber="11">
<figureCitation id="1C9ABAEAB9197840C5B844C4FF0A2207" box="[151,227,1771,1797]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB9197840C5DE44C4FEE42207" box="[241,269,1771,1797]" captionStart="FIGURE 10" captionStartId="35.[151,250,1994,2018]" captionTargetBox="[260,1301,189,1949]" captionTargetId="figure-15@35.[210,1377,181,1970]" captionTargetPageId="35" captionText="FIGURE 10. Hydraena cameroovala, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284870" httpUri="https://zenodo.org/record/7284870/files/figure.png" pageId="10" pageNumber="11">10</figureCitation>
(habitus), 10 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197840C5B8451CFBC62273" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="841EA66FB9197840C5B8451CFBC62273" blockId="10.[151,1437,1842,1977]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5B8451CFEAE224E" bold="true" box="[151,327,1842,1869]" pageId="10" pageNumber="11">Type Material.</emphasis>
<materialsCitation id="34C9AC32B9197840C462451CFDD22273" ID-GBIF-Occurrence="3962178315" collectingDate="1982-05-22" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Nyong River" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB9197840C462451CFE5C224F" box="[333,437,1843,1869]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB9197840C732451CFD71224F" box="[541,664,1843,1869]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B9197840C78D451CFCF5224F" box="[674,796,1843,1869]" pageId="10" pageNumber="11">C. S. Prov.</collectorName>
,
<quantity id="43590B8AB9197840C60A451DFC86224F" box="[805,879,1842,1869]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="km" value="15.0">15 km</quantity>
.
<collectorName id="2954C3B9B9197840C656451CFBF8224F" box="[889,1041,1843,1869]" pageId="10" pageNumber="11">S. Mbalmayo</collectorName>
,
<location id="817EF0B4B9197840C133451CFB47224F" LSID="urn:lsid:plazi:treatment:0C081779B9197841C5B844E9FB622113:817EF0B4B9197840C133451CFB47224F" box="[1052,1198,1843,1869]" country="Cameroon" name="Nyong River" pageId="10" pageNumber="11">Nyong River</location>
, ex. leaves at margin,
<date id="F01F80AFB9197840C5B84578FEE12273" box="[151,264,1879,1905]" pageId="10" pageNumber="11" value="1982-05-22">
<collectingDate id="E05B7947B9197840C5B84578FEE12273" box="[151,264,1879,1905]" pageId="10" pageNumber="11" value="1982-05-22">22.v.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B9197840C46B4578FE3E2273" box="[324,471,1879,1905]" pageId="10" pageNumber="11">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB9197840C4DF4578FDDA2273" box="[496,563,1879,1905]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="11" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B9197840C76A4578FBC22273" ID-GBIF-Occurrence="3962178332" box="[581,1067,1879,1905]" collectingDate="1982-05-22" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Nyong River" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB9197840C76A4578FD5F2273" box="[581,694,1879,1905]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
: Same data as holotype (4
<collectionCode id="E2B03EAAB9197840C6CE4578FBCA2273" box="[993,1059,1879,1905]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="11" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9197841C5E84555FC0A25D7" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="841EA66FB9197841C5E84555FC0A25D7" blockId="10.[151,1437,1842,1977]" lastBlockId="11.[151,1437,151,1041]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E84555FE3D2296" bold="true" box="[199,468,1914,1940]" pageId="10" pageNumber="11">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB9197840C10B4554FB702297" box="[1060,1177,1915,1941]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
by the combination of the oval body form, the small body size (ca.
<quantity id="43590B8AB9197840C7B445B0FCEC22BB" box="[667,773,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="11" unit="mm" value="1.18">1.18 mm</quantity>
), the posterosubmedial foveae moderately deep, confluent medially forming U-shaped impression, and the aedeagus (
<figureCitation id="1C9ABAEAB9187841C61A42B8FC6325B3" box="[821,906,151,177]" captionStart="FIGURE 10" captionStartId="35.[151,250,1994,2018]" captionTargetBox="[260,1301,189,1949]" captionTargetId="figure-15@35.[210,1377,181,1970]" captionTargetPageId="35" captionText="FIGURE 10. Hydraena cameroovala, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284870" httpUri="https://zenodo.org/record/7284870/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
). Similar in habitus to some other members of Aedeagal Group 1; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9187841C5E842F1FD4E26CB" pageId="11" pageNumber="12" type="description">
<paragraph id="841EA66FB9187841C5E842F1FD69271B" blockId="11.[151,1437,151,1041]" pageId="11" pageNumber="12">
<emphasis id="B6D57A7DB9187841C5E842F1FEB825FA" bold="true" box="[199,337,222,248]" pageId="11" pageNumber="12">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB9187841C48B42F0FDEF25FB" box="[420,518,223,249]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.18/0.56; head width 0.34; pronotum 0.31/0.44, PA 0.36, PB 0.39; elytra 0.70/0.56. Habitus as illustrated (
<figureCitation id="1C9ABAEAB9187841C12A432CFBB1241F" box="[1029,1112,259,285]" captionStart="FIGURE 10" captionStartId="35.[151,250,1994,2018]" captionTargetBox="[260,1301,189,1949]" captionTargetId="figure-15@35.[210,1377,181,1970]" captionTargetPageId="35" captionText="FIGURE 10. Hydraena cameroovala, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284870" httpUri="https://zenodo.org/record/7284870/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
). Body form quite oval. Dorsum generally brown to dark brown, maxillary palpi, labrum and clypeus testaceous, pronotum with diffusely margined weakly defined macula. Dorsum of head and pronotum moderately coarsely, often very densely punctulate, frons punctures distinctly finer and less dense than pronotal punctures, interstices shining; punctures of frons disc and pronotal disc ca. 23xef, interstices ca. 1xpd or less; punctures of clypeus very fine, very dense, dull. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral absent, posterosubmedial foveae moderately deep, confluent medially forming U-shaped impression.
</paragraph>
<paragraph id="841EA66FB9187841C5E8400CFE7427CF" blockId="11.[151,1437,151,1041]" pageId="11" pageNumber="12">Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming strong angle with one another.</paragraph>
<paragraph id="841EA66FB9187841C5B840F8FD4E26CB" blockId="11.[151,1437,151,1041]" pageId="11" pageNumber="12">Venter: Mentum sparsely finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow posteriorly, tapering to point anteriorly, slightly converging toward one another anteriorly, weakly raised, located at sides of shallow median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</paragraph>
</subSubSection>
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<emphasis id="B6D57A7DB9187841C5E841FDFE8826EE" bold="true" box="[199,353,978,1004]" pageId="11" pageNumber="12">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9187841C7E341FCFD1526EF" box="[716,764,979,1005]" pageId="11" pageNumber="12">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9187841C64841FCFC5326EF" box="[871,954,979,1005]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="11" pageNumber="12">Fig. 41</figureCitation>
).
</paragraph>
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<subSubSection id="CCBBF5E4B9187841C5E841D8FB622113" box="[199,1163,1015,1041]" pageId="11" pageNumber="12" type="etymology">
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<emphasis id="B6D57A7DB9187841C5E841D8FEA72113" bold="true" box="[199,334,1015,1041]" pageId="11" pageNumber="12">Etymology.</emphasis>
Named in reference to the known distribution, and to the oval body form.
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<mods:title id="89FF273F5F37427CE76D4311D565C99F">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<heading id="DF561103B91A7843C5B846E1FD8D21EA" bold="true" box="[151,612,1230,1257]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
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<taxonomicName id="43A1DDECB91A7843C5B846E1FE2C21EB" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,453,1230,1257]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="camerocorona" status="sp. nov.">
<emphasis id="B6D57A7DB91A7843C5B846E1FE2C21EB" bold="true" box="[151,453,1230,1257]" italics="true" pageId="9" pageNumber="10">Hydraena camerocorona</emphasis>
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,
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<paragraph id="841EA66FB91A7843C5B846DDFD93200E" blockId="9.[151,634,1230,1292]" box="[151,634,1266,1292]" pageId="9" pageNumber="10">
<figureCitation id="1C9ABAEAB91A7843C5B846DDFF0A200E" box="[151,227,1266,1292]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB91A7843C5DE46DDFF16200E" box="[241,255,1266,1292]" captionStart="FIGURE 8" captionStartId="33.[151,250,1993,2017]" captionTargetBox="[227,1359,181,1969]" captionTargetId="figure-15@33.[227,1359,181,1969]" captionTargetPageId="33" captionText="FIGURE 8. Hydraena camerocorona, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284864" httpUri="https://zenodo.org/record/7284864/files/figure.png" pageId="9" pageNumber="10">8</figureCitation>
(habitus), 8 (aedeagus), 42 (map)
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<paragraph id="841EA66FB91A7843C5B84715FD51207A" blockId="9.[151,1437,1338,2013]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5B84715FEAE2056" bold="true" box="[151,327,1338,1364]" pageId="9" pageNumber="10">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91A7843C4624715FD51207A" ID-GBIF-Occurrence="3962178322" collectingDate="1978-01-07" collectionCode="LUM" country="Cameroon" location="Akonolinga" municipality="Gardenforhall - Samuelson" pageId="9" pageNumber="10" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91A7843C4624715FE5C2056" box="[333,437,1338,1364]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
(male): “
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, moist secondary forest and plantation, at light,
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, Loc. #17 //
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” (
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<emphasis id="B6D57A7DB91A7843C5E847ADFE3D209E" bold="true" box="[199,468,1410,1436]" pageId="9" pageNumber="10">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91A7843C10B47ADFB70209E" box="[1060,1177,1410,1436]" name="Cameroon" pageId="9" pageNumber="10">Cameroon</collectingCountry>
by the combination of the very fine and widely spaced elytral punctures, the comparatively wide lateral explanate margins of the elytra, the pronotal macula, and the aedeagus (
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).
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<paragraph id="841EA66FB91A7843C5E847C1FA6A2206" blockId="9.[151,1437,1338,2013]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5E847C1FEB8230A" bold="true" box="[199,337,1518,1544]" pageId="9" pageNumber="10">Description</emphasis>
. Size:
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(length/width, mm): body (length to elytral apices) 1.26/0.56; head width 0.34; pronotum 0.31/0.4, PA 0.37, PB 0.38; elytra 0.75/0.56. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91A7843C6D6443DFBD5232E" box="[1017,1084,1554,1580]" captionStart="FIGURE 8" captionStartId="33.[151,250,1993,2017]" captionTargetBox="[227,1359,181,1969]" captionTargetId="figure-15@33.[227,1359,181,1969]" captionTargetPageId="33" captionText="FIGURE 8. Hydraena camerocorona, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284864" httpUri="https://zenodo.org/record/7284864/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
). Dorsum brown to dark brown or black, frons and diffusely margined pronotal macula darker, frons almost black, contrasting with light brown clypeus, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely finely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB91A7843C5E84521FE3622BA" blockId="9.[151,1437,1338,2013]" pageId="9" pageNumber="10">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 34xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB91A7840C5B845ECFCAC248B" blockId="9.[151,1437,1338,2013]" lastBlockId="10.[151,1437,151,501]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.7/4/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately wide, slightly converging toward one another anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. Tibiae slender, protibial very slightly arcuate, meso- and metatibiae straight. Abdominal apex symmetrical; last tergite with slightly off-center apical notch.</paragraph>
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<emphasis id="B6D57A7DB9197840C5E843BDFE8824AE" bold="true" box="[199,353,402,428]" pageId="10" pageNumber="11">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB9197840C7E343BCFD1524AF" box="[716,764,403,429]" pageId="10" pageNumber="11">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9197840C64843BCFC5224AF" box="[871,955,403,429]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="10" pageNumber="11">Fig. 42</figureCitation>
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</paragraph>
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<paragraph id="841EA66FB9197840C5E84398FF5524F7" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="11">
<emphasis id="B6D57A7DB9197840C5E84398FEA724D3" bold="true" box="[199,334,439,465]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named in reference to the known distribution, and the wide lateral explanate margins of the elytra.
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<paragraph id="841EA66FB91B7842C5B84045FD9A2786" blockId="8.[151,634,618,681]" box="[151,627,618,645]" pageId="8" pageNumber="9">
<heading id="DF561103B91B7842C5B84045FD9A2786" bold="true" box="[151,627,618,645]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
<emphasis id="B6D57A7DB91B7842C5B84045FD9A2786" bold="true" box="[151,627,618,645]" pageId="8" pageNumber="9">
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<emphasis id="B6D57A7DB91B7842C5B84045FE3D2787" bold="true" box="[151,468,618,645]" italics="true" pageId="8" pageNumber="9">Hydraena cameroconcava</emphasis>
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,
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<paragraph id="841EA66FB91B7842C5B840A0FD9327AB" blockId="8.[151,634,618,681]" box="[151,634,655,681]" pageId="8" pageNumber="9">
<figureCitation id="1C9ABAEAB91B7842C5B840A0FF0A27AB" box="[151,227,655,681]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB91B7842C5DE40A0FF1627AB" box="[241,255,655,681]" captionStart="FIGURE 6" captionStartId="31.[151,250,1996,2020]" captionTargetBox="[315,1272,181,1973]" captionTargetId="figure-15@31.[315,1272,181,1973]" captionTargetPageId="31" captionText="FIGURE 6. Hydraena cameroconcava, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284857" httpUri="https://zenodo.org/record/7284857/files/figure.png" pageId="8" pageNumber="9">6</figureCitation>
(habitus), 6 (aedeagus), 43 (map)
</paragraph>
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<subSubSection id="CCBBF5E4B91B7842C5B840F8FD8D2617" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="841EA66FB91B7842C5B840F8FD8D2617" blockId="8.[151,1437,726,1761]" pageId="8" pageNumber="9">
<emphasis id="B6D57A7DB91B7842C5B840F8FEAE27F2" bold="true" box="[151,327,726,753]" pageId="8" pageNumber="9">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91B7842C46340F8FD8D2617" ID-GBIF-Occurrence="3962178329" collectingDate="1949-05-05" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91B7842C46340F8FE5D27F3" box="[332,436,727,753]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
(male):
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:
<collectingMunicipality id="647A3C15B91B7842C7BC40F8FC9D27F3" box="[659,884,727,753]" pageId="8" pageNumber="9">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB91B7842C6A840F8FB4527F3" box="[903,1196,727,753]" name="Cameroon" pageId="8" pageNumber="9">BRITISH CAMEROONS</collectingCountry>
/ Matute, / Tiko Plantation /
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<collectingDate id="E05B7947B91B7842C5C240D5FEA62617" box="[237,335,762,789]" pageId="8" pageNumber="9" value="1949-05-05">5.v.1949</collectingDate>
</date>
,
<collectorName id="2954C3B9B91B7842C47440D4FE262617" box="[347,463,763,789]" pageId="8" pageNumber="9">B. Malkin</collectorName>
” (
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).
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</paragraph>
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<emphasis id="B6D57A7DB91B7842C5E84131FE38263A" bold="true" box="[199,465,798,824]" pageId="8" pageNumber="9">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
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by the combination of the absence of a distinct pronotal macula, the frons distinctly concave between median area and eye, the widely separated metaventral plaques that are markedly converging anteriorly (ratios P2/w/l/s ca. 1/1/4/5), and the aedeagus (
<figureCitation id="1C9ABAEAB91B7842C5B041A4FF0D26A7" box="[159,228,907,933]" captionStart="FIGURE 6" captionStartId="31.[151,250,1996,2020]" captionTargetBox="[315,1272,181,1973]" captionTargetId="figure-15@31.[315,1272,181,1973]" captionTargetPageId="31" captionText="FIGURE 6. Hydraena cameroconcava, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284857" httpUri="https://zenodo.org/record/7284857/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). Also refer to the diagnosis of
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<emphasis id="B6D57A7DB91B7842C76B41A4FCE126A6" box="[580,776,907,932]" italics="true" pageId="8" pageNumber="9">H. camerocorona</emphasis>
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for comments on the male genitalia.
</paragraph>
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<subSubSection id="CCBBF5E4B91B7842C5E84181FD4A239B" pageId="8" pageNumber="9" type="description">
<paragraph id="841EA66FB91B7842C5E84181FE2321C7" blockId="8.[151,1437,726,1761]" pageId="8" pageNumber="9">
<emphasis id="B6D57A7DB91B7842C5E84181FEB826CA" bold="true" box="[199,337,942,968]" pageId="8" pageNumber="9">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91B7842C48B4180FDEF26CB" box="[420,518,943,969]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.20/0.50; head width 0.32; pronotum 0.30/0.42, PA 0.36, PB 0.37; elytra 0.67/0.50. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91B7842C11C41FCFB9326EF" box="[1075,1146,979,1005]" captionStart="FIGURE 6" captionStartId="31.[151,250,1996,2020]" captionTargetBox="[315,1272,181,1973]" captionTargetId="figure-15@31.[315,1272,181,1973]" captionTargetPageId="31" captionText="FIGURE 6. Hydraena cameroconcava, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284857" httpUri="https://zenodo.org/record/7284857/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). Dorsum dark brown to black, frons and pronotal disc darker, light brown clypeus contrasting with black frons, palpi testaceous, tip of last palpomere not darker. Frons distinctly concave between median area and eye. Dorsum of head and pronotum finely sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial pronotal foveae rather deep
</paragraph>
<paragraph id="841EA66FB91B7842C5E846E0FEEC209F" blockId="8.[151,1437,726,1761]" pageId="8" pageNumber="9">Elytra with summit of posterior declivity at or very near posterior 1/3, where elytra transversely quite convex; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Abdominal apex with slightly off-center apical notch.</paragraph>
<paragraph id="841EA66FB91B7842C5B84788FD4A239B" blockId="8.[151,1437,726,1761]" pageId="8" pageNumber="9">Venter: Mentum and postmentum densely finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/4/5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides slightly converging toward rather narrow blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, markedly converging anteriorly, posteriorly separated by ca. 4x anterior separation, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS concave. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight.</paragraph>
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<emphasis id="B6D57A7DB91B7842C5E8448DFE8823BE" bold="true" box="[199,353,1698,1724]" pageId="8" pageNumber="9">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91B7842C7E3448CFD1523BF" box="[716,764,1699,1725]" pageId="8" pageNumber="9">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91B7842C648448CFC5223BF" box="[871,955,1699,1725]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="8" pageNumber="9">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91B7842C5E844E8FC6523E3" box="[199,908,1735,1761]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="841EA66FB91B7842C5E844E8FC6523E3" blockId="8.[151,1437,726,1761]" box="[199,908,1735,1761]" pageId="8" pageNumber="9">
<emphasis id="B6D57A7DB91B7842C5E844E8FEA723E3" bold="true" box="[199,334,1735,1761]" pageId="8" pageNumber="9">Etymology.</emphasis>
Named in reference to the concavities of the frons.
</paragraph>
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</treatment>
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<heading id="DF561103B91B7842C5B8451DFD69224E" bold="true" box="[151,640,1842,1869]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
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<emphasis id="B6D57A7DB91B7842C5B8451DFE08224F" bold="true" box="[151,481,1842,1869]" italics="true" pageId="8" pageNumber="9">Hydraena camerocontrasta</emphasis>
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,
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</paragraph>
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<subSubSection id="CCBBF5E4B91B7842C5B84578FD932273" box="[151,634,1879,1905]" pageId="8" pageNumber="9" type="description">
<paragraph id="841EA66FB91B7842C5B84578FD932273" blockId="8.[151,640,1842,1905]" box="[151,634,1879,1905]" pageId="8" pageNumber="9">
<figureCitation id="1C9ABAEAB91B7842C5B84578FF0A2273" box="[151,227,1879,1905]" captionStart="FIGURES 12" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 12. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 14." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
,
<figureCitation id="1C9ABAEAB91B7842C5DE4578FF162273" box="[241,255,1879,1905]" captionStart="FIGURE 7" captionStartId="32.[151,250,1989,2013]" captionTargetBox="[243,1353,199,1946]" captionTargetId="figure-25@32.[201,1386,181,1965]" captionTargetPageId="32" captionText="FIGURE 7. Hydraena camerocontrasta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284860" httpUri="https://zenodo.org/record/7284860/files/figure.png" pageId="8" pageNumber="9">7</figureCitation>
(habitus), 7 (aedeagus), 43 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91B7842C5B845B0FC0A22DF" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="841EA66FB91B7842C5B845B0FC0A22DF" blockId="8.[151,1436,1950,2013]" pageId="8" pageNumber="9">
<emphasis id="B6D57A7DB91B7842C5B845B0FEAF22BA" bold="true" box="[151,326,1950,1977]" pageId="8" pageNumber="9">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91B7842C46445B0FC0A22DF" ID-GBIF-Occurrence="3962178314" collectingDate="1949-01-07" collectingDateMax="1949-01-11" collectingDateMin="1949-01-07" collectionCode="UKNHM" collectorName="B. Malkin" country="Cameroon" location="Cameroon" municipality="Northwest Province" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91B7842C46445B0FE5A22BB" box="[331,435,1951,1977]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
(male):
<location id="817EF0B4B91B7842C72145B0FD6E22BB" LSID="urn:lsid:plazi:treatment:0C081779B91B7843C5B8451DFE4C217E:817EF0B4B91B7842C72145B0FD6E22BB" box="[526,647,1951,1977]" country="Cameroon" municipality="Northwest Province" name="Cameroon" pageId="8" pageNumber="9">Cameroon</location>
:
<collectingMunicipality id="647A3C15B91B7842C7A045B0FC9922BB" box="[655,880,1951,1977]" pageId="8" pageNumber="9">Northwest Province</collectingMunicipality>
. “
<collectingCountry id="FCB6E6FFB91B7842C6AD45B0FB4222BB" box="[898,1195,1951,1977]" name="Cameroon" pageId="8" pageNumber="9">BRITISH CAMEROONS</collectingCountry>
. / Mamfe,
<date id="F01F80AFB91B7842C03345B0FA7522BB" box="[1308,1436,1951,1977]" pageId="8" pageNumber="9" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">
<collectingDate id="E05B7947B91B7842C03345B0FA7522BB" box="[1308,1436,1951,1977]" pageId="8" pageNumber="9" value="1949-01-07" valueMax="1949-01-11" valueMin="1949-01-07">7-11.i.1949</collectingDate>
</date>
/
<collectorName id="2954C3B9B91B7842C58945ECFEF122DF" box="[166,280,1987,2013]" pageId="8" pageNumber="9">B. Malkin</collectorName>
coll. // Rain forest; clear stream / Gravel and sand” (
<collectionCode id="E2B03EAAB91B7842C64945ECFC3E22DF" box="[870,983,1987,2013]" pageId="8" pageNumber="9">UKNHM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91A7843C5E842B9FF26241E" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="841EA66FB91A7843C5E842B9FF26241E" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5E842B9FE3D25B2" bold="true" box="[199,468,150,176]" pageId="9" pageNumber="10">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91A7843C10B42B9FB7025B2" box="[1060,1177,150,176]" name="Cameroon" pageId="9" pageNumber="10">Cameroon</collectingCountry>
by the combination of the light brown clypeus contrasting with dark brown frons, the frons shallowly concave between median area and eye, and the male genitalia (
<figureCitation id="1C9ABAEAB91A7843C4E042F1FDFA25FA" box="[463,531,222,248]" captionStart="FIGURE 7" captionStartId="32.[151,250,1989,2013]" captionTargetBox="[243,1353,199,1946]" captionTargetId="figure-25@32.[201,1386,181,1965]" captionTargetPageId="32" captionText="FIGURE 7. Hydraena camerocontrasta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284860" httpUri="https://zenodo.org/record/7284860/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
). Also refer to the diagnosis of
<taxonomicName id="43A1DDECB91A7843C65F42F0FBDA25FA" authorityName="Perkins" authorityYear="2022" box="[880,1075,223,248]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="camerocorona">
<emphasis id="B6D57A7DB91A7843C65F42F0FBDA25FA" box="[880,1075,223,248]" italics="true" pageId="9" pageNumber="10">H. camerocorona</emphasis>
</taxonomicName>
for comments on the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91A7843C5E84309FD1D2112" pageId="9" pageNumber="10" type="description">
<paragraph id="841EA66FB91A7843C5E84309FDD1273E" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5E84309FEB82442" bold="true" box="[199,337,294,320]" pageId="9" pageNumber="10">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91A7843C48B4309FDEF2442" box="[420,518,294,320]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.22/0.54; head width 0.30; pronotum 0.29/0.44, PA 0.36, PB 0.39; elytra 0.76/0.54. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91A7843C11C4365FB932466" box="[1075,1146,330,356]" captionStart="FIGURE 7" captionStartId="32.[151,250,1989,2013]" captionTargetBox="[243,1353,199,1946]" captionTargetId="figure-25@32.[201,1386,181,1965]" captionTargetPageId="32" captionText="FIGURE 7. Hydraena camerocontrasta, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284860" httpUri="https://zenodo.org/record/7284860/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
). Dorsum brown to dark brown, frons and pronotal disc darker, light brown clypeus contrasting with dark brown frons, palpi testaceous, tip of last palpomere not darker. Frons shallowly concave between median area and eye. Dorsum of head and pronotum finely sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 13xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial pronotal foveae moderately deep
</paragraph>
<paragraph id="841EA66FB91A7843C5E84069FEEC2616" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">Elytra with summit of posterior declivity at or very near posterior 1/3, where elytra transversely quite convex; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Abdominal apex with slightly off-center apical notch.</paragraph>
<paragraph id="841EA66FB91A7843C5B84130FD1D2112" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">Venter: Mentum and postmentum moderately finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, parallel, raised very slightly, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia slightly arcuate, meso- and metatibiae straight. Abdominal apex with off center apical notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91A7843C5E84635FC212136" box="[199,968,1050,1077]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="841EA66FB91A7843C5E84635FC212136" blockId="9.[151,1437,150,1149]" box="[199,968,1050,1077]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5E84635FE882136" bold="true" box="[199,353,1050,1076]" pageId="9" pageNumber="10">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91A7843C7E34634FD152137" box="[716,764,1051,1077]" pageId="9" pageNumber="10">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91A7843C6484635FC522136" box="[871,955,1050,1076]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="9" pageNumber="10">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91A7843C5E84611FE4C217E" pageId="9" pageNumber="10" type="etymology">
<paragraph id="841EA66FB91A7843C5E84611FE4C217E" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">
<emphasis id="B6D57A7DB91A7843C5E84611FEA7215A" bold="true" box="[199,334,1086,1112]" pageId="9" pageNumber="10">Etymology.</emphasis>
Named in reference to the known distribution and the contrasting sizes of the parameres: right very elongate, left just a nub.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,139 @@
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<mods:title id="9117BE0006F039A2B7B8DB84662783E2">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B91C7845C5B841A5FB2022DF" ID-DOI="http://doi.org/10.5281/zenodo.7300261" ID-GBIF-Taxon="204205207" ID-Zenodo-Dep="7300261" LSID="urn:lsid:plazi:treatment:0C081779B91C7845C5B841A5FB2022DF" httpUri="http://treatment.plazi.org/id/0C081779B91C7845C5B841A5FB2022DF" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="CCBBF5E4B91C7845C5B841A5FD8E26A6" box="[151,615,906,933]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="841EA66FB91C7845C5B841A5FD8E26A6" blockId="15.[151,662,906,969]" box="[151,615,906,933]" pageId="15" pageNumber="16">
<heading id="DF561103B91C7845C5B841A5FD8E26A6" bold="true" box="[151,615,906,933]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="4">
<emphasis id="B6D57A7DB91C7845C5B841A5FD8E26A6" bold="true" box="[151,615,906,933]" pageId="15" pageNumber="16">
<taxonomicName id="43A1DDECB91C7845C5B841A5FE2026A6" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,457,906,933]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="camerofoveata" status="sp. nov.">
<emphasis id="B6D57A7DB91C7845C5B841A5FE2026A6" bold="true" box="[151,457,906,933]" italics="true" pageId="15" pageNumber="16">Hydraena camerofoveata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91C7845C4F841A4FD8E26A6" box="[471,615,906,933]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5B84180FD7F26CB" box="[151,662,943,969]" pageId="15" pageNumber="16" type="description">
<paragraph id="841EA66FB91C7845C5B84180FD7F26CB" blockId="15.[151,662,906,969]" box="[151,662,943,969]" pageId="15" pageNumber="16">
<figureCitation id="1C9ABAEAB91C7845C5B84180FF0D26CB" box="[151,228,943,969]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="15" pageNumber="16">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91C7845C5DE4180FEE426CB" box="[241,269,943,969]" captionStart="FIGURE 17" captionStartId="42.[151,250,1976,2000]" captionTargetBox="[275,1313,181,1952]" captionTargetId="figure-25@42.[275,1313,181,1952]" captionTargetPageId="42" captionText="FIGURE 17. Hydraena camerofoveata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284892" httpUri="https://zenodo.org/record/7284892/files/figure.png" pageId="15" pageNumber="16">17</figureCitation>
(habitus), 17 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5B841D8FE2B2137" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="841EA66FB91C7845C5B841D8FE2B2137" blockId="15.[151,1437,1014,2013]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5B841D8FEA02112" bold="true" box="[151,329,1014,1041]" pageId="15" pageNumber="16">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91C7845C47E41D8FE2B2137" ID-GBIF-Occurrence="3962178331" collectingDate="1983-01-02" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Isali River" municipality="Nouma" pageId="15" pageNumber="16" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91C7845C47E41D8FE502113" box="[337,441,1015,1041]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB91C7845C70A41D8FD492113" box="[549,672,1015,1041]" name="Cameroon" pageId="15" pageNumber="16">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B91C7845C78341D8FCC32113" box="[684,810,1015,1041]" pageId="15" pageNumber="16">C. S. Prov.</collectorName>
,
<quantity id="43590B8AB91C7845C61941D9FC682113" box="[822,897,1014,1041]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="km" value="15.0">15 km</quantity>
. NW Yaounde,
<location id="817EF0B4B91C7845C11B41D8FB432113" LSID="urn:lsid:plazi:treatment:0C081779B91C7845C5B841A5FB2022DF:817EF0B4B91C7845C11B41D8FB432113" box="[1076,1194,1015,1041]" country="Cameroon" municipality="Nouma" name="Isali River" pageId="15" pageNumber="16">Isali River</location>
nr.
<collectingMunicipality id="647A3C15B91C7845C1F741D8FAC42113" box="[1240,1325,1015,1041]" pageId="15" pageNumber="16">Nouma</collectingMunicipality>
,
<date id="F01F80AFB91C7845C01641D8FA712113" box="[1337,1432,1015,1041]" pageId="15" pageNumber="16" value="1983-01-02">
<collectingDate id="E05B7947B91C7845C01641D8FA712113" box="[1337,1432,1015,1041]" pageId="15" pageNumber="16" value="1983-01-02">2.i.1983</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B91C7845C5E84634FEB22137" box="[199,347,1051,1077]" pageId="15" pageNumber="16">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB91C7845C45B4634FE5E2137" box="[372,439,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="15" pageNumber="16" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E84611FDFA200F" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="841EA66FB91C7845C5E84611FDFA200F" blockId="15.[151,1437,1014,2013]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E84611FE3D215A" bold="true" box="[199,468,1086,1112]" pageId="15" pageNumber="16">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91C7845C10B4610FB70215B" box="[1060,1177,1087,1113]" name="Cameroon" pageId="15" pageNumber="16">Cameroon</collectingCountry>
by the combination of the small body size (ca.
<quantity id="43590B8AB91C7845C48D464CFDE0217F" box="[418,521,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="15" pageNumber="16" unit="mm" value="1.24">1.24 mm</quantity>
), the densely coarsely punctate dorsum, the anterolateral and posterolateral foveae deep, shallowly confluent, the posterosubmedian foveae deep, the plaques absent or very small and very faintly indicated by less dense hydrofuge pubescence, located at sides of very shallow median depression, and the aedeagus (
<figureCitation id="1C9ABAEAB91C7845C5B046E0FF1B21EB" box="[159,242,1231,1257]" captionStart="FIGURE 17" captionStartId="42.[151,250,1976,2000]" captionTargetBox="[275,1313,181,1952]" captionTargetId="figure-25@42.[275,1313,181,1952]" captionTargetPageId="42" captionText="FIGURE 17. Hydraena camerofoveata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284892" httpUri="https://zenodo.org/record/7284892/files/figure.png" pageId="15" pageNumber="16">Fig. 17</figureCitation>
). The habitus is quite similar to that of
<taxonomicName id="43A1DDECB91C7845C78846E0FC9221EB" authorityName="Perkins" authorityYear="2022" box="[679,891,1231,1257]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB91C7845C78846E0FC9221EB" box="[679,891,1231,1257]" italics="true" pageId="15" pageNumber="16">H. camerocrebrata</emphasis>
</taxonomicName>
, but that species has well developed metaventral plaques, and a different aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E84739FF342297" pageId="15" pageNumber="16" type="description">
<paragraph id="841EA66FB91C7845C5E84739FE98230B" blockId="15.[151,1437,1014,2013]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E84739FEB82032" bold="true" box="[199,337,1302,1328]" pageId="15" pageNumber="16">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91C7845C48B4738FDEF2033" box="[420,518,1303,1329]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.24/0.54; head width 0.31; pronotum 0.31/0.44, PA 0.37, PB 0.38; elytra 0.73/0.54. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91C7845C1064714FB962057" box="[1065,1151,1339,1365]" captionStart="FIGURE 17" captionStartId="42.[151,250,1976,2000]" captionTargetBox="[275,1313,181,1952]" captionTargetId="figure-25@42.[275,1313,181,1952]" captionTargetPageId="42" captionText="FIGURE 17. Hydraena camerofoveata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284892" httpUri="https://zenodo.org/record/7284892/files/figure.png" pageId="15" pageNumber="16">Fig. 17</figureCitation>
). Dorsum dark brown to black, maxillary palpi brown, tip of last palpomere not darker. Frons densely moderately coarsely, pronotum coarsely densely punctate, punctures much larger than those of frons, interstices shining, ca. 0.51xpd. Labrum apicomedially excised, free margins very weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae deep, shallowly confluent; posterosubmedian foveae deep; anteromedian fovea shallow.
</paragraph>
<paragraph id="841EA66FB91C7845C5E8443CFE0323BF" blockId="15.[151,1437,1014,2013]" pageId="15" pageNumber="16">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 12xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with short and fine seta. Intervals not raised, width ca. 0.51xpd, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming moderately strong angle with one another.</paragraph>
<paragraph id="841EA66FB91C7845C5B844E8FF342297" blockId="15.[151,1437,1014,2013]" pageId="15" pageNumber="16">Venter: Mentum and postmentum sparsely finely punctulate, shining. Genae raised, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent or very small and very faintly indicated by less dense hydrofuge pubescence, located at sides of very shallow median depression. AIS width at straight posterior margin ca. 2.5x P2. All tibiae straight, moderately stout. Abdominal apex symmetrical, with very small apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E845B1FC2122BB" box="[199,968,1950,1977]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="841EA66FB91C7845C5E845B1FC2122BB" blockId="15.[151,1437,1014,2013]" box="[199,968,1950,1977]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E845B1FE8822BA" bold="true" box="[199,353,1950,1976]" pageId="15" pageNumber="16">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91C7845C7E345B0FD1522BB" box="[716,764,1951,1977]" pageId="15" pageNumber="16">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91C7845C64845B0FC5322BB" box="[871,954,1951,1977]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="15" pageNumber="16">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E845ECFB2022DF" box="[199,1225,1987,2013]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="841EA66FB91C7845C5E845ECFB2022DF" blockId="15.[151,1437,1014,2013]" box="[199,1225,1987,2013]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E845ECFEA722DF" bold="true" box="[199,334,1987,2013]" pageId="15" pageNumber="16">Etymology.</emphasis>
Named in reference to the known distribution, and to the deep pronotal foveae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,158 @@
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<mods:title id="427677C8EFAD0028E3C9F04BA250C709">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<paragraph id="841EA66FB91D7844C5B84309FDB32442" blockId="14.[151,662,294,357]" box="[151,602,294,321]" pageId="14" pageNumber="15">
<heading id="DF561103B91D7844C5B84309FDB32442" bold="true" box="[151,602,294,321]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
<emphasis id="B6D57A7DB91D7844C5B84309FDB32442" bold="true" box="[151,602,294,321]" pageId="14" pageNumber="15">
<taxonomicName id="43A1DDECB91D7844C5B84309FE552443" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,444,294,321]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camerocubita" status="sp. nov.">
<emphasis id="B6D57A7DB91D7844C5B84309FE552443" bold="true" box="[151,444,294,321]" italics="true" pageId="14" pageNumber="15">Hydraena camerocubita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91D7844C4E54309FDB32442" box="[458,602,294,320]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="841EA66FB91D7844C5B84365FD7F2466" blockId="14.[151,662,294,357]" box="[151,662,330,357]" pageId="14" pageNumber="15">
<figureCitation id="1C9ABAEAB91D7844C5B84365FF0D2466" box="[151,228,330,356]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="14" pageNumber="15">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91D7844C5DE4365FEE42467" box="[241,269,330,357]" captionStart="FIGURE 15" captionStartId="40.[151,250,1979,2003]" captionTargetBox="[256,1332,181,1956]" captionTargetId="figure-25@40.[256,1332,181,1956]" captionTargetPageId="40" captionText="FIGURE 15. Hydraena camerocubita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284886" httpUri="https://zenodo.org/record/7284886/files/figure.png" pageId="14" pageNumber="15">15</figureCitation>
(habitus), 15 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5B843BDFB0224D2" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="841EA66FB91D7844C5B843BDFB0224D2" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5B843BDFEA324AE" bold="true" box="[151,330,402,428]" pageId="14" pageNumber="15">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91D7844C47C43BDFF0324D2" ID-GBIF-Occurrence="3962178343" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91D7844C47C43BDFE5224AE" box="[339,443,402,428]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB91D7844C73043BDFDC424AE" bold="true" box="[543,557,402,428]" pageId="14" pageNumber="15"></emphasis>
<collectingCountry id="FCB6E6FFB91D7844C70243BDFD4B24AE" box="[557,674,402,428]" name="Cameroon" pageId="14" pageNumber="15">Cameroon</collectingCountry>
:
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<collectorName id="2954C3B9B91D7844C62943BCFC5E24AE" box="[774,951,402,429]" pageId="14" pageNumber="15">N. Ngaoundere</collectorName>
, stream,
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<collectingDate id="E05B7947B91D7844C13043BDFB6824AE" box="[1055,1153,402,428]" pageId="14" pageNumber="15" value="1982-05-01">1.v.1982</collectingDate>
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,
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collector” (
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)
</materialsCitation>
.
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<typeStatus id="5B1A18CDB91D7844C5DA4398FE8F24D3" box="[245,358,439,465]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
:
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.
<collectorName id="2954C3B9B91D7844C4EA4398FD9A24D2" box="[453,627,438,465]" pageId="14" pageNumber="15">N. Ngaoundere</collectorName>
, lake margin,
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,
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collector (2
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E843F5FCEC2786" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="841EA66FB91D7844C5E843F5FCEC2786" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E843F5FE3D24F6" bold="true" box="[199,468,474,500]" pageId="14" pageNumber="15">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91D7844C10B43F5FB7024F6" box="[1060,1177,474,500]" name="Cameroon" pageId="14" pageNumber="15">Cameroon</collectingCountry>
by the combination of the small size (ca.
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), the metaventral plaques that are very small, very narrow, subcarinate lines, and the aedeagus (
<figureCitation id="1C9ABAEAB91D7844C422400DFE89273F" box="[269,352,546,573]" captionStart="FIGURE 15" captionStartId="40.[151,250,1979,2003]" captionTargetBox="[256,1332,181,1956]" captionTargetId="figure-25@40.[256,1332,181,1956]" captionTargetPageId="40" captionText="FIGURE 15. Hydraena camerocubita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284886" httpUri="https://zenodo.org/record/7284886/files/figure.png" pageId="14" pageNumber="15">Fig. 15</figureCitation>
). The dorsal habitus is similar to that of
<taxonomicName id="43A1DDECB91D7844C60E400CFC06273E" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[801,1007,546,572]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cameroconfinis" status="sp. nov.">
<emphasis id="B6D57A7DB91D7844C60E400CFC06273E" box="[801,1007,546,572]" italics="true" pageId="14" pageNumber="15">H. cameroconfinis</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB91D7844C6D9400DFBDD273E" bold="true" box="[1014,1076,546,572]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="ADE6C706B91D7844C6D9400DFBDD273E" box="[1014,1076,546,572]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but the present species is much smaller (ca. 1.26 vs.
<quantity id="43590B8AB91D7844C4AA4069FE072762" box="[389,494,582,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="15" unit="mm" value="1.66">1.66 mm</quantity>
), and has different ventral proportions (P2/w/l/s ca. 1/0.15/0.15/2 vs. 1/0.25/1.5/1). The aedeagi of the two species are markedly dissimilar.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E840A1FCEC2032" pageId="14" pageNumber="15" type="description">
<paragraph id="841EA66FB91D7844C5E840A1FBD526A6" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E840A1FEB827AA" bold="true" box="[199,337,654,680]" pageId="14" pageNumber="15">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91D7844C48B40A1FDEF27AA" box="[420,518,654,680]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.26/0.52; head width 0.33; pronotum 0.30/0.41, PA 0.33, PB 0.37; elytra 0.76/0.52. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91D7844C107409DFB9727CF" box="[1064,1150,690,717]" captionStart="FIGURE 15" captionStartId="40.[151,250,1979,2003]" captionTargetBox="[256,1332,181,1956]" captionTargetId="figure-25@40.[256,1332,181,1956]" captionTargetPageId="40" captionText="FIGURE 15. Hydraena camerocubita, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284886" httpUri="https://zenodo.org/record/7284886/files/figure.png" pageId="14" pageNumber="15">Fig. 15</figureCitation>
). Dorsum brown to light brown, head and pronotal macula dark brown to black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc slightly smaller than 1xef, on pronotal disc slightly larger than 1xef, interstices ca. 23xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral pronotal foveae very shallow, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB91D7844C5E84181FB452136" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect slightly separately rounded, in posterior aspect margins forming very shallow angle with one another.</paragraph>
<paragraph id="841EA66FB91D7844C5B84610FCEC2032" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.15/0.15/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, subcarinate lines, straight, convergent anteriorly, very weakly if at all raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Pro-, meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E84715FC212056" box="[199,968,1338,1365]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="841EA66FB91D7844C5E84715FC212056" blockId="14.[151,1437,402,1437]" box="[199,968,1338,1365]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E84715FE882056" bold="true" box="[199,353,1338,1364]" pageId="14" pageNumber="15">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91D7844C7E34714FD152057" box="[716,764,1339,1365]" pageId="14" pageNumber="15">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91D7844C6484715FC522056" box="[871,955,1338,1364]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E84771FE6A209E" pageId="14" pageNumber="15" type="etymology">
<paragraph id="841EA66FB91D7844C5E84771FE6A209E" blockId="14.[151,1437,402,1437]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E84771FEA7207A" bold="true" box="[199,334,1374,1400]" pageId="14" pageNumber="15">Etymology.</emphasis>
Named in reference to the known distribution and the arm-like shape of the gonopore bearing process of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,139 @@
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<mods:title id="DED9FB41875ED08A606EA79AA4E1BD35">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B91D7845C5B847C7FEEC263B" ID-DOI="http://doi.org/10.5281/zenodo.7300194" ID-GBIF-Taxon="204205202" ID-Zenodo-Dep="7300194" LSID="urn:lsid:plazi:treatment:0C081779B91D7845C5B847C7FEEC263B" httpUri="http://treatment.plazi.org/id/0C081779B91D7845C5B847C7FEEC263B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="CCBBF5E4B91D7844C5B847C7FD832301" box="[151,618,1512,1539]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="841EA66FB91D7844C5B847C7FD832301" blockId="14.[151,662,1512,1575]" box="[151,618,1512,1539]" pageId="14" pageNumber="15">
<heading id="DF561103B91D7844C5B847C7FD832301" bold="true" box="[151,618,1512,1539]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
<emphasis id="B6D57A7DB91D7844C5B847C7FD832301" bold="true" box="[151,618,1512,1539]" pageId="14" pageNumber="15">
<taxonomicName id="43A1DDECB91D7844C5B847C7FE252301" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,460,1512,1539]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camerocurvata" status="sp. nov.">
<emphasis id="B6D57A7DB91D7844C5B847C7FE252301" bold="true" box="[151,460,1512,1539]" italics="true" pageId="14" pageNumber="15">Hydraena camerocurvata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91D7844C4F447C6FD832301" box="[475,618,1513,1539]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5B84422FD7F2325" box="[151,662,1549,1575]" pageId="14" pageNumber="15" type="description">
<paragraph id="841EA66FB91D7844C5B84422FD7F2325" blockId="14.[151,662,1512,1575]" box="[151,662,1549,1575]" pageId="14" pageNumber="15">
<figureCitation id="1C9ABAEAB91D7844C5B84422FF0D2325" box="[151,228,1549,1575]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="14" pageNumber="15">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91D7844C5DE4422FEE42325" box="[241,269,1549,1575]" captionStart="FIGURE 16" captionStartId="41.[151,250,1995,2019]" captionTargetBox="[232,1360,192,1957]" captionTargetId="figure-15@41.[196,1392,181,1971]" captionTargetPageId="41" captionText="FIGURE 16. Hydraena camerocurvata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284888" httpUri="https://zenodo.org/record/7284888/files/figure.png" pageId="14" pageNumber="15">16</figureCitation>
(habitus), 16 (aedeagus), 40 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5B8447FFD3A238C" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="841EA66FB91D7844C5B8447FFD3A238C" blockId="14.[151,1437,1615,2038]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5B8447FFEAE236B" bold="true" box="[151,327,1615,1642]" pageId="14" pageNumber="15">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91D7844C4624460FF03238C" ID-GBIF-Occurrence="3962178303" collectingDate="1982-01-30" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91D7844C4624460FE5C236B" box="[333,437,1615,1641]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB91D7844C734447FFD7F2368" box="[539,662,1616,1642]" name="Cameroon" pageId="14" pageNumber="15">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B91D7844C78F447FFCCB2368" box="[672,802,1616,1642]" pageId="14" pageNumber="15">N. W. Prov.</collectorName>
,
<quantity id="43590B8AB91D7844C603447FFC8E236B" box="[812,871,1615,1642]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="km" value="7.0">7 km</quantity>
.
<collectorName id="2954C3B9B91D7844C65F447FFC12236B" box="[880,1019,1615,1642]" pageId="14" pageNumber="15">E. Bamenda</collectorName>
, stream,
<date id="F01F80AFB91D7844C1734460FB21236B" box="[1116,1224,1615,1641]" pageId="14" pageNumber="15" value="1982-01-30">
<collectingDate id="E05B7947B91D7844C1734460FB21236B" box="[1116,1224,1615,1641]" pageId="14" pageNumber="15" value="1982-01-30">30.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B91D7844C02E447FFA7A236B" box="[1281,1427,1615,1642]" pageId="14" pageNumber="15">P. D. Perkins</collectorName>
” (
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)
</materialsCitation>
.
<materialsCitation id="34C9AC32B91D7844C5DA445BFD26238C" ID-GBIF-Occurrence="3962178338" box="[245,719,1651,1678]" collectingDate="1982-01-30" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB91D7844C5DA445BFEB2238C" box="[245,347,1652,1678]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
: Same data as holotype (1
<collectionCode id="E2B03EAAB91D7844C7A9445BFD21238C" box="[646,712,1652,1678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="14" pageNumber="15" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E844B8FB25221F" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="841EA66FB91D7844C5E844B8FB25221F" blockId="14.[151,1437,1615,2038]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E844B8FE3D23B3" bold="true" box="[199,468,1687,1713]" pageId="14" pageNumber="15">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91D7844C10B44B7FB7023B0" box="[1060,1177,1688,1714]" name="Cameroon" pageId="14" pageNumber="15">Cameroon</collectingCountry>
by the combination of the ovate body form, the transversely convex pronotum, the posterosubmedial foveae shallow, confluent medially forming U-shaped impression, the very narrow P2, and the aedeagus (
<figureCitation id="1C9ABAEAB91D7844C68944F0FC1023FB" box="[934,1017,1759,1785]" captionStart="FIGURE 16" captionStartId="41.[151,250,1995,2019]" captionTargetBox="[232,1360,192,1957]" captionTargetId="figure-15@41.[196,1392,181,1971]" captionTargetPageId="41" captionText="FIGURE 16. Hydraena camerocurvata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284888" httpUri="https://zenodo.org/record/7284888/files/figure.png" pageId="14" pageNumber="15">Fig. 16</figureCitation>
). Differentiation from other similarly shaped members of Aedeagal Species Group 2 will require careful examination of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7845C5E84508FB7727CF" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="841EA66FB91D7845C5E84508FF3D241F" blockId="14.[151,1437,1615,2038]" lastBlockId="15.[151,1437,151,825]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E84508FEB82243" bold="true" box="[199,337,1831,1857]" pageId="14" pageNumber="15">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91D7844C48B4508FDEF2243" box="[420,518,1831,1857]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.30/0.57; head width 0.34; pronotum 0.31/0.49, PA 0.39, PB 0.41; elytra 0.80/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91D7844C1214564FB882267" box="[1038,1121,1867,1893]" captionStart="FIGURE 16" captionStartId="41.[151,250,1995,2019]" captionTargetBox="[232,1360,192,1957]" captionTargetId="figure-15@41.[196,1392,181,1971]" captionTargetPageId="41" captionText="FIGURE 16. Hydraena camerocurvata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284888" httpUri="https://zenodo.org/record/7284888/files/figure.png" pageId="14" pageNumber="15">Fig. 16</figureCitation>
). Body form ovate. Dorsum generally brown to dark brown, maxillary palpi, labrum and clypeus testaceous, pronotum with diffusely margined weakly defined macula. Dorsum of head and pronotum moderately coarsely, moderately densely punctulate, frons punctures distinctly finer and less dense than pronotal punctures, interstices shining; punctures of frons disc and pronotal disc ca. 12xef, interstices ca. 12xpd; punctures of clypeus fine, moderately dense, interstices shining medially, dull laterally. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum transversely convex, anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae shallow, posterosubmedial foveae shallow, confluent medially forming U-shaped impression, anteromedial fovea moderately deep.
</paragraph>
<paragraph id="841EA66FB91C7845C5E84308FF1824D3" blockId="15.[151,1437,151,825]" pageId="15" pageNumber="16">Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB91C7845C5B843F4FB7727CF" blockId="15.[151,1437,151,825]" pageId="15" pageNumber="16">Venter: Mentum and postmentum sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/3. P1 laminate; median carina sinuate in profile. P2 very narrow, l/w ca. 4/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, very slightly arcuate, tapering anteriorly and slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 3x P2. All tibiae stout. Protibia widened near midlength, very slightly arcuate. Meso- and metatibiae straight. Abdominal apex symmetrical, last tergite rounded, without apicomedian notch, last ventrite emarginate.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E840F9FC2127F3" box="[199,968,726,753]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="841EA66FB91C7845C5E840F9FC2127F3" blockId="15.[151,1437,151,825]" box="[199,968,726,753]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E840F9FE8827F2" bold="true" box="[199,353,726,752]" pageId="15" pageNumber="16">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91C7845C7E340F8FD1527F3" box="[716,764,727,753]" pageId="15" pageNumber="16">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91C7845C64840F8FC5227F3" box="[871,955,727,753]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="15" pageNumber="16">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91C7845C5E840D4FEEC263B" pageId="15" pageNumber="16" type="etymology">
<paragraph id="841EA66FB91C7845C5E840D4FEEC263B" blockId="15.[151,1437,151,825]" pageId="15" pageNumber="16">
<emphasis id="B6D57A7DB91C7845C5E840D4FEA72617" bold="true" box="[199,334,763,789]" pageId="15" pageNumber="16">Etymology.</emphasis>
Named in reference to the known distribution, and to the ovate body form and very curvaceous aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,211 @@
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<mods:title id="75DD2B02FB543DEBCA3202D7E089BAD3">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B91E7844C5B841A5FACF25D6" ID-DOI="http://doi.org/10.5281/zenodo.7300257" ID-GBIF-Taxon="204205206" ID-Zenodo-Dep="7300257" LSID="urn:lsid:plazi:treatment:0C081779B91E7844C5B841A5FACF25D6" httpUri="http://treatment.plazi.org/id/0C081779B91E7844C5B841A5FACF25D6" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<paragraph id="841EA66FB91E7847C5B841A5FD9D26A6" blockId="13.[151,662,906,969]" box="[151,628,906,933]" pageId="13" pageNumber="14">
<heading id="DF561103B91E7847C5B841A5FD9D26A6" bold="true" box="[151,628,906,933]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B6D57A7DB91E7847C5B841A5FD9D26A6" bold="true" box="[151,628,906,933]" pageId="13" pageNumber="14">
<taxonomicName id="43A1DDECB91E7847C5B841A5FE3F26A7" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,470,906,933]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="camerocrebrata" status="sp. nov.">
<emphasis id="B6D57A7DB91E7847C5B841A5FE3F26A7" bold="true" box="[151,470,906,933]" italics="true" pageId="13" pageNumber="14">Hydraena camerocrebrata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91E7847C4CB41A4FD9D26A6" box="[484,628,906,933]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5B84180FD7F26CB" box="[151,662,943,969]" pageId="13" pageNumber="14" type="description">
<paragraph id="841EA66FB91E7847C5B84180FD7F26CB" blockId="13.[151,662,906,969]" box="[151,662,943,969]" pageId="13" pageNumber="14">
<figureCitation id="1C9ABAEAB91E7847C5B84180FF0D26CB" box="[151,228,943,969]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="13" pageNumber="14">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91E7847C5DE4180FEE426CB" box="[241,269,943,969]" captionStart="FIGURE 14" captionStartId="39.[151,250,1921,1945]" captionTargetBox="[220,1357,194,1880]" captionTargetId="figure-15@39.[158,1430,181,1897]" captionTargetPageId="39" captionText="FIGURE 14. Hydraena camerocrebrata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284882" httpUri="https://zenodo.org/record/7284882/files/figure.png" pageId="13" pageNumber="14">14</figureCitation>
(habitus), 14 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5B841D8FC86217F" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="841EA66FB91E7847C5B841D8FC86217F" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="14">
<emphasis id="B6D57A7DB91E7847C5B841D8FEA02112" bold="true" box="[151,329,1014,1041]" pageId="13" pageNumber="14">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91E7847C47E41D8FE892137" ID-GBIF-Occurrence="3962178337" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="13" pageNumber="14" specimenCount="950" typeStatus="holotype">
<typeStatus id="5B1A18CDB91E7847C47E41D8FE502113" box="[337,441,1015,1041]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB91E7847C73441D9FDC02112" bold="true" box="[539,553,1014,1040]" pageId="13" pageNumber="14"></emphasis>
<collectingCountry id="FCB6E6FFB91E7847C70641D8FD772113" box="[553,670,1015,1041]" name="Cameroon" pageId="13" pageNumber="14">Cameroon</collectingCountry>
:
<location id="817EF0B4B91E7847C78141D8FCFC2113" LSID="urn:lsid:plazi:treatment:0C081779B91E7844C5B841A5FACF25D6:817EF0B4B91E7847C78141D8FCFC2113" box="[686,789,1015,1041]" country="Cameroon" municipality="Mt. Febe" name="Yaounde" pageId="13" pageNumber="14">Yaounde</location>
:
<collectingMunicipality id="647A3C15B91E7847C60F41D8FC602113" box="[800,905,1015,1041]" pageId="13" pageNumber="14">Mt. Febe</collectingMunicipality>
,
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<quantity id="43590B8AB91E7847C6B941D9FC0A2113" box="[918,995,1014,1041]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="m" value="950.0">
<elevation id="0F8C415CB91E7847C6B941D9FC0A2113" box="[918,995,1014,1041]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="m" value="950.0">950 m</elevation>
</quantity>
.
</specimenCount>
, hygropetric,
<date id="F01F80AFB91E7847C1AB41D8FB152113" box="[1156,1276,1015,1041]" pageId="13" pageNumber="14" value="1982-04-10">
<collectingDate id="E05B7947B91E7847C1AB41D8FB152113" box="[1156,1276,1015,1041]" pageId="13" pageNumber="14" value="1982-04-10">10.iv.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B91E7847C02641D8FA752113" box="[1289,1436,1015,1041]" pageId="13" pageNumber="14">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB91E7847C43A4634FEB12137" box="[277,344,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<typeStatus id="5B1A18CDB91E7847C4434634FE302137" box="[364,473,1051,1077]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
(120): Same
<materialsCitation id="34C9AC32B91E7847C75C4634FC402137" ID-GBIF-Occurrence="3962178344" box="[627,937,1051,1077]" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
data as holotype (28
<collectionCode id="E2B03EAAB91E7847C64F4634FC4B2137" box="[864,930,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="34C9AC32B91E7847C69A4634FABE2137" ID-GBIF-Occurrence="3962178325" box="[949,1367,1050,1077]" collectingDate="1982-04-07" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
same data except
<date id="F01F80AFB91E7847C1514634FB0F2137" box="[1150,1254,1051,1077]" pageId="13" pageNumber="14" value="1982-04-07">
<collectingDate id="E05B7947B91E7847C1514634FB0F2137" box="[1150,1254,1051,1077]" pageId="13" pageNumber="14" value="1982-04-07">7.iv.1982</collectingDate>
</date>
(5
<collectionCode id="E2B03EAAB91E7847C0224634FAA62137" box="[1293,1359,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="34C9AC32B91E7847C04C4634FDFB215B" ID-GBIF-Occurrence="3962178345" collectingDate="1983-06-18" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
same data except
<date id="F01F80AFB91E7847C4324610FE7C215B" box="[285,405,1087,1113]" pageId="13" pageNumber="14" value="1983-06-18">
<collectingDate id="E05B7947B91E7847C4324610FE7C215B" box="[285,405,1087,1113]" pageId="13" pageNumber="14" value="1983-06-18">18.vi.1983</collectingDate>
</date>
(64
<collectionCode id="E2B03EAAB91E7847C4E74610FDE3215B" box="[456,522,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="34C9AC32B91E7847C7324610FC30215B" ID-GBIF-Occurrence="3962178326" box="[541,985,1087,1113]" collectingDate="1983-06-14" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
same data except
<date id="F01F80AFB91E7847C7CB4610FCB2215B" box="[740,859,1087,1113]" pageId="13" pageNumber="14" value="1983-06-14">
<collectingDate id="E05B7947B91E7847C7CB4610FCB2215B" box="[740,859,1087,1113]" pageId="13" pageNumber="14" value="1983-06-14">14.vi.1983</collectingDate>
</date>
(12
<collectionCode id="E2B03EAAB91E7847C6A04610FC38215B" box="[911,977,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="34C9AC32B91E7847C6CC4610FC82217F" ID-GBIF-Occurrence="3962178321" collectingDate="1982-05-20" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="900" location="Yaounde" municipality="Mt. Mbankolo" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
<location id="817EF0B4B91E7847C6CC4610FBA1215B" LSID="urn:lsid:plazi:treatment:0C081779B91E7844C5B841A5FACF25D6:817EF0B4B91E7847C6CC4610FBA1215B" box="[995,1096,1087,1113]" country="Cameroon" municipality="Mt. Mbankolo" name="Yaounde" pageId="13" pageNumber="14">Yaounde</location>
,
<collectingMunicipality id="647A3C15B91E7847C17B4610FB11215B" box="[1108,1272,1087,1113]" pageId="13" pageNumber="14">Mt. Mbankolo</collectingMunicipality>
,
<quantity id="43590B8AB91E7847C02B4610FAAF215B" box="[1284,1350,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="m" value="900.0">
<elevation id="0F8C415CB91E7847C02B4610FAAF215B" box="[1284,1350,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="m" value="900.0">900m</elevation>
</quantity>
, hygropetric microhabitat,
<date id="F01F80AFB91E7847C456464CFE03217F" box="[377,490,1123,1149]" pageId="13" pageNumber="14" value="1982-05-20">
<collectingDate id="E05B7947B91E7847C456464CFE03217F" box="[377,490,1123,1149]" pageId="13" pageNumber="14" value="1982-05-20">20.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B91E7847C4D9464CFD6F217F" box="[502,646,1123,1149]" pageId="13" pageNumber="14">P. D. Perkins</collectorName>
collector (11
<collectionCode id="E2B03EAAB91E7847C60D464CFC8D217F" box="[802,868,1123,1149]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5E846A9FD952033" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="841EA66FB91E7847C5E846A9FD952033" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="14">
<emphasis id="B6D57A7DB91E7847C5E846A9FE3D21A2" bold="true" box="[199,468,1158,1184]" pageId="13" pageNumber="14">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91E7847C10B46A8FB7021A3" box="[1060,1177,1159,1185]" name="Cameroon" pageId="13" pageNumber="14">Cameroon</collectingCountry>
by the combination of the very densely punctate dorsum, the very narrow P2 (l/w ca. 4/1) and the aedeagus (
<figureCitation id="1C9ABAEAB91E7847C15A4684FB2221C7" box="[1141,1227,1195,1221]" captionStart="FIGURE 14" captionStartId="39.[151,250,1921,1945]" captionTargetBox="[220,1357,194,1880]" captionTargetId="figure-15@39.[158,1430,181,1897]" captionTargetPageId="39" captionText="FIGURE 14. Hydraena camerocrebrata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284882" httpUri="https://zenodo.org/record/7284882/files/figure.png" pageId="13" pageNumber="14">Fig. 14</figureCitation>
). Also refer to the diagnosis of
<taxonomicName id="43A1DDECB91E7847C40A46E0FDF921EA" authorityName="Perkins" authorityYear="2022" box="[293,528,1231,1256]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="camerocompressa">
<emphasis id="B6D57A7DB91E7847C40A46E0FDF921EA" box="[293,528,1231,1256]" italics="true" pageId="13" pageNumber="14">H. camerocompressa</emphasis>
</taxonomicName>
. Although the aedeagi of
<taxonomicName id="43A1DDECB91E7847C60246E0FBE821EB" authorityName="Perkins" authorityYear="2022" box="[813,1025,1231,1257]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB91E7847C60246E0FBE821EB" box="[813,1025,1231,1257]" italics="true" pageId="13" pageNumber="14">H. camerocrebrata</emphasis>
</taxonomicName>
and
<taxonomicName id="43A1DDECB91E7847C11946E0FB0321EB" authorityName="Perkins" authorityYear="2022" box="[1078,1258,1231,1257]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB91E7847C11946E0FB0321EB" box="[1078,1258,1231,1257]" italics="true" pageId="13" pageNumber="14">H. cameropetila</emphasis>
</taxonomicName>
distinctly differ in several characters, a relationship of the two species is indicated by the shapes of the parameres: large, wide and with similar clusters of setae (
<figureCitation id="1C9ABAEAB91E7847C4C84738FDAD2033" box="[487,580,1303,1329]" captionStart="FIGURE 14" captionStartId="39.[151,250,1921,1945]" captionTargetBox="[220,1357,194,1880]" captionTargetId="figure-15@39.[158,1430,181,1897]" captionTargetPageId="39" captionText="FIGURE 14. Hydraena camerocrebrata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284882" httpUri="https://zenodo.org/record/7284882/files/figure.png" pageId="13" pageNumber="14">Figs. 14</figureCitation>
,
<figureCitation id="1C9ABAEAB91E7847C77F4738FD862033" box="[592,623,1303,1329]" captionStart="FIGURE 18" captionStartId="43.[151,250,1945,1969]" captionTargetBox="[206,1394,202,1889]" captionTargetId="figure-15@43.[156,1431,181,1921]" captionTargetPageId="43" captionText="FIGURE 18. Hydraena cameropetila, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284894" httpUri="https://zenodo.org/record/7284894/files/figure.png" pageId="13" pageNumber="14">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5E84715FD8122DF" pageId="13" pageNumber="14" type="description">
<paragraph id="841EA66FB91E7847C5E84715FC97232F" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="14">
<emphasis id="B6D57A7DB91E7847C5E84715FEB82056" bold="true" box="[199,337,1338,1364]" pageId="13" pageNumber="14">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91E7847C48B4714FDEF2057" box="[420,518,1339,1365]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.33/0.57; head width 0.31; pronotum 0.33/0.48, PA 0.41, PB 0.42; elytra 0.85/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91E7847C1064770FB96207B" box="[1065,1151,1375,1401]" captionStart="FIGURE 14" captionStartId="39.[151,250,1921,1945]" captionTargetBox="[220,1357,194,1880]" captionTargetId="figure-15@39.[158,1430,181,1897]" captionTargetPageId="39" captionText="FIGURE 14. Hydraena camerocrebrata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284882" httpUri="https://zenodo.org/record/7284882/files/figure.png" pageId="13" pageNumber="14">Fig. 14</figureCitation>
). Dorsum dark brown to black, pronotum without macula, palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, each puncture with short, distinctive whitish seta, interstices dull; punctures of frons disc and pronotal disc ca. 23xef, interstices ca. 1xpd or less; punctures of clypeus fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep.
</paragraph>
<paragraph id="841EA66FB91E7847C5E84418FEC823E3" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="14">Elytra with summit of rather abrupt posterior declivity at about posterior 1/3; lateral explanate margins narrow to moderately wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 0.5xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB91E7847C5B844C4FD8122DF" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="14">Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, very weakly shining, with posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/6/6. P1 laminate; median carina weakly sinuate in profile. P2 very narrow, l/w ca. 4/1, sides parallel, apex blunt. Plaques narrow, straight or very slightly arcuate, slightly convergent anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at arcuate posterior margin ca. 2.5x P2. Pro- meso- and metatibiae straight, moderately stout.Abdominal apex symmetrical, without apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E842B9FC2125B2" box="[199,968,150,177]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="841EA66FB91D7844C5E842B9FC2125B2" blockId="14.[199,1318,150,213]" box="[199,968,150,177]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E842B9FE8825B2" bold="true" box="[199,353,150,176]" pageId="14" pageNumber="15">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91D7844C7E342B8FD1525B3" box="[716,764,151,177]" pageId="14" pageNumber="15">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91D7844C64842B9FC5225B2" box="[871,955,150,176]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91D7844C5E84295FACF25D6" box="[199,1318,186,213]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="841EA66FB91D7844C5E84295FACF25D6" blockId="14.[199,1318,150,213]" box="[199,1318,186,213]" pageId="14" pageNumber="15">
<emphasis id="B6D57A7DB91D7844C5E84295FEA725D6" bold="true" box="[199,334,186,212]" pageId="14" pageNumber="15">Etymology.</emphasis>
Named in reference to the known distribution and to the very densely punctate dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,170 @@
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<mods:title id="8C628C686CAD2E28CBF1A1CDB01F12A9">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<subSubSection id="CCBBF5E4B91F7846C5B84399FD7324D2" box="[151,666,438,465]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="841EA66FB91F7846C5B84399FD7324D2" blockId="12.[151,666,438,501]" box="[151,666,438,465]" pageId="12" pageNumber="13">
<heading id="DF561103B91F7846C5B84399FD7324D2" bold="true" box="[151,666,438,465]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="4">
<emphasis id="B6D57A7DB91F7846C5B84399FD7324D2" bold="true" box="[151,666,438,465]" pageId="12" pageNumber="13">
<taxonomicName id="43A1DDECB91F7846C5B84399FE1224D3" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,507,438,465]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cameroclandestina" status="sp. nov.">
<emphasis id="B6D57A7DB91F7846C5B84399FE1224D3" bold="true" box="[151,507,438,465]" italics="true" pageId="12" pageNumber="13">Hydraena cameroclandestina</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91F7846C7254398FD7324D2" box="[522,666,438,465]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5B843F4FD7F24F7" box="[151,662,475,501]" pageId="12" pageNumber="13" type="description">
<paragraph id="841EA66FB91F7846C5B843F4FD7F24F7" blockId="12.[151,666,438,501]" box="[151,662,475,501]" pageId="12" pageNumber="13">
<figureCitation id="1C9ABAEAB91F7846C5B843F4FF0D24F7" box="[151,228,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="12" pageNumber="13">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91F7846C5DE43F4FEE424F7" box="[241,269,475,501]" captionStart="FIGURE 12" captionStartId="37.[151,250,1972,1996]" captionTargetBox="[322,1249,197,1928]" captionTargetId="figure-15@37.[264,1323,181,1948]" captionTargetPageId="37" captionText="FIGURE 12. Hydraena cameroclandestina, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284874" httpUri="https://zenodo.org/record/7284874/files/figure.png" pageId="12" pageNumber="13">12</figureCitation>
(habitus), 12 (aedeagus), 40 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5B8400CFD372763" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="841EA66FB91F7846C5B8400CFD372763" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5B8400CFEAE273E" bold="true" box="[151,327,546,573]" pageId="12" pageNumber="13">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91F7846C462400CFF032763" ID-GBIF-Occurrence="3962178316" collectingDate="1982-01-30" collectionCode="MCZ" collectorName=" P. D. Perkins" country="Cameroon" location="Cameroon" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB91F7846C462400CFE5C273F" box="[333,437,547,573]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB91F7846C734400CFD7F273F" box="[539,662,547,573]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B91F7846C78F400CFCCB273F" box="[672,802,547,573]" pageId="12" pageNumber="13">N. W. Prov.</collectorName>
,
<quantity id="43590B8AB91F7846C603400CFC8E273F" box="[812,871,547,573]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="km" value="7.0">7 km</quantity>
.
<collectorName id="2954C3B9B91F7846C65F400CFC12273F" box="[880,1019,547,573]" pageId="12" pageNumber="13">E. Bamenda</collectorName>
, stream,
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<collectingDate id="E05B7947B91F7846C173400CFB21273F" box="[1116,1224,547,573]" pageId="12" pageNumber="13" value="1982-01-30">30.i.1982</collectingDate>
</date>
, leg.
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” (
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)
</materialsCitation>
.
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<typeStatus id="5B1A18CDB91F7846C5DA4068FE8F2763" box="[245,358,583,609]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: Same data as holotype (5
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E84045FD25263B" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="841EA66FB91F7846C5E84045FD25263B" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E84045FE382786" bold="true" box="[199,465,618,644]" pageId="12" pageNumber="13">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91F7846C1274044FB942787" box="[1032,1149,619,645]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
by the combination of the moderately coarsely, very densely punctulate head and pronotum, and the aedeagus (
<figureCitation id="1C9ABAEAB91F7846C16340A0FB4927AB" box="[1100,1184,655,681]" captionStart="FIGURE 12" captionStartId="37.[151,250,1972,1996]" captionTargetBox="[322,1249,197,1928]" captionTargetId="figure-15@37.[264,1323,181,1948]" captionTargetPageId="37" captionText="FIGURE 12. Hydraena cameroclandestina, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284874" httpUri="https://zenodo.org/record/7284874/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
). The dorsal habitus is very similar to that of
<taxonomicName id="43A1DDECB91F7846C4BA409CFD8327CF" authorityName="Perkins" authorityYear="2022" box="[405,618,691,717]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camerocrebrata">
<emphasis id="B6D57A7DB91F7846C4BA409CFD8327CF" box="[405,618,691,717]" italics="true" pageId="12" pageNumber="13">H. camerocrebrata</emphasis>
</taxonomicName>
and
<taxonomicName id="43A1DDECB91F7846C78D409CFCB127CF" authorityName="Perkins" authorityYear="2022" box="[674,856,691,717]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB91F7846C78D409CFCB127CF" box="[674,856,691,717]" italics="true" pageId="12" pageNumber="13">H. cameropetila</emphasis>
</taxonomicName>
; all three species are in Aedeagal Species Group 2.
<taxonomicName id="43A1DDECB91F7846C5B840F8FE6F27F3" authorityName="Perkins" authorityYear="2022" box="[151,390,727,753]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cameroclandestina">
<emphasis id="B6D57A7DB91F7846C5B840F8FE6F27F3" box="[151,390,727,753]" italics="true" pageId="12" pageNumber="13">H.cameroclandestina</emphasis>
</taxonomicName>
is larger in body size than
<taxonomicName id="43A1DDECB91F7846C79840F8FC8427F3" authorityName="Perkins" authorityYear="2022" box="[695,877,727,753]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cameropetila">
<emphasis id="B6D57A7DB91F7846C79840F8FC8427F3" box="[695,877,727,753]" italics="true" pageId="12" pageNumber="13">H. cameropetila</emphasis>
</taxonomicName>
(ca. 1.32 vs.
<quantity id="43590B8AB91F7846C12B40F8FB8427F3" box="[1028,1133,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="13" unit="mm" value="1.2">1.20 mm</quantity>
); also, the P2 is wider than that of the two compared species. Reliable determinations of all three species will require examination of the male genitalia, which distinctly differ from one another.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E8416DFBE620C3" pageId="12" pageNumber="13" type="description">
<paragraph id="841EA66FB91F7846C5E8416DFCDC2137" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E8416DFEB8265E" bold="true" box="[199,337,834,860]" pageId="12" pageNumber="13">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91F7846C48B416CFDEF265F" box="[420,518,835,861]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.32/0.58; head width 0.33; pronotum 0.32/0.47, PA 0.38, PB 0.40; elytra 0.79/0.58. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91F7846C1064148FB962683" box="[1065,1151,871,897]" captionStart="FIGURE 12" captionStartId="37.[151,250,1972,1996]" captionTargetBox="[322,1249,197,1928]" captionTargetId="figure-15@37.[264,1323,181,1948]" captionTargetPageId="37" captionText="FIGURE 12. Hydraena cameroclandestina, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284874" httpUri="https://zenodo.org/record/7284874/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
). Dorsum dark brown to black, pronotum without macula, maxillary palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, each puncture with short, distinctive whitish seta, interstices dull; punctures of frons disc and pronotal disc ca. 2xef, interstices ca. 1xpd or less; punctures of clypeus fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae shallow.
</paragraph>
<paragraph id="841EA66FB91F7846C5E84610FDC021EB" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">Elytra with summit of posterior declivity at about midlength; lateral explanate margins narrow to moderately wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with short and fine conspicuous whitish seta. Intervals not raised, width ca. 0.51xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB91F7846C5B846DCFBE620C3" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">Venter: Mentum and postmentum very densely very finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, parallel, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae moderately stout. Protibia very slightly arcuate. Meso- and metatibiae straight. Abdominal apex symmetrical, with very small and shallow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E847E5FC2120E7" box="[199,968,1482,1509]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="841EA66FB91F7846C5E847E5FC2120E7" blockId="12.[151,1437,546,1581]" box="[199,968,1482,1509]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E847E5FE8820E6" bold="true" box="[199,353,1482,1508]" pageId="12" pageNumber="13">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91F7846C7E347E4FD1520E7" box="[716,764,1483,1509]" pageId="12" pageNumber="13">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91F7846C64847E4FC5220E7" box="[871,955,1483,1509]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="12" pageNumber="13">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E847C0FDA2232F" pageId="12" pageNumber="13" type="etymology">
<paragraph id="841EA66FB91F7846C5E847C0FDA2232F" blockId="12.[151,1437,546,1581]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E847C0FEA7230B" bold="true" box="[199,334,1519,1545]" pageId="12" pageNumber="13">Etymology.</emphasis>
Named in reference to the known distribution, and to the close dorsal similarity to
<taxonomicName id="43A1DDECB91F7846C1D547C0FF3D232F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camerocrebrata" status="sp. nov.">
<emphasis id="B6D57A7DB91F7846C1D547C0FF3D232F" italics="true" pageId="12" pageNumber="13">H. camerocrebrata</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB91F7846C5F4443CFEF3232F" bold="true" box="[219,282,1555,1581]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="ADE6C706B91F7846C5F4443CFEF3232F" box="[219,282,1555,1581]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="43A1DDECB91F7846C47F443CFDEF232F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[336,518,1555,1581]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cameropetila" status="sp. nov.">
<emphasis id="B6D57A7DB91F7846C47F443CFDEF232F" box="[336,518,1555,1581]" italics="true" pageId="12" pageNumber="13">H. cameropetila</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB91F7846C723443CFDA2232F" bold="true" box="[524,587,1555,1581]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="ADE6C706B91F7846C723443CFDA2232F" box="[524,587,1555,1581]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,148 @@
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<mods:title id="027B3846318C164DF837CD99AD76B029">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="0C081779B91F7847C5B84451FB28263B" ID-DOI="http://doi.org/10.5281/zenodo.7300255" ID-GBIF-Taxon="204205205" ID-Zenodo-Dep="7300255" LSID="urn:lsid:plazi:treatment:0C081779B91F7847C5B84451FB28263B" httpUri="http://treatment.plazi.org/id/0C081779B91F7847C5B84451FB28263B" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="CCBBF5E4B91F7846C5B84451FD6A239A" box="[151,643,1662,1689]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="841EA66FB91F7846C5B84451FD6A239A" blockId="12.[151,662,1662,1725]" box="[151,643,1662,1689]" pageId="12" pageNumber="13">
<heading id="DF561103B91F7846C5B84451FD6A239A" bold="true" box="[151,643,1662,1689]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="4">
<emphasis id="B6D57A7DB91F7846C5B84451FD6A239A" bold="true" box="[151,643,1662,1689]" pageId="12" pageNumber="13">
<taxonomicName id="43A1DDECB91F7846C5B84451FE0D239B" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,484,1662,1689]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camerocomplexa" status="sp. nov.">
<emphasis id="B6D57A7DB91F7846C5B84451FE0D239B" bold="true" box="[151,484,1662,1689]" italics="true" pageId="12" pageNumber="13">Hydraena camerocomplexa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B91F7846C4DC4450FD6A239A" box="[499,643,1662,1689]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5B8448CFD7F23BF" box="[151,662,1699,1725]" pageId="12" pageNumber="13" type="description">
<paragraph id="841EA66FB91F7846C5B8448CFD7F23BF" blockId="12.[151,662,1662,1725]" box="[151,662,1699,1725]" pageId="12" pageNumber="13">
<figureCitation id="1C9ABAEAB91F7846C5B8448CFF0D23BF" box="[151,228,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="12" pageNumber="13">Figs. 3</figureCitation>
,
<figureCitation id="1C9ABAEAB91F7846C5DE448CFEE423BF" box="[241,269,1699,1725]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">13</figureCitation>
(habitus), 13 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5B844C4FCA0222B" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="841EA66FB91F7846C5B844C4FCA0222B" blockId="12.[151,1436,1770,2013]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5B844C4FEA02206" bold="true" box="[151,329,1770,1797]" pageId="12" pageNumber="13">Type Material.</emphasis>
<materialsCitation id="34C9AC32B91F7846C47E44C4FEB6222B" ID-GBIF-Occurrence="3962178309" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="12" pageNumber="13" specimenCount="950" typeStatus="holotype">
<typeStatus id="5B1A18CDB91F7846C47E44C4FE502207" box="[337,441,1771,1797]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB91F7846C73444C5FDC02206" bold="true" box="[539,553,1770,1796]" pageId="12" pageNumber="13"></emphasis>
<collectingCountry id="FCB6E6FFB91F7846C70644C4FD772207" box="[553,670,1771,1797]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
:
<location id="817EF0B4B91F7846C78144C4FCFC2207" LSID="urn:lsid:plazi:treatment:0C081779B91F7847C5B84451FB28263B:817EF0B4B91F7846C78144C4FCFC2207" box="[686,789,1771,1797]" country="Cameroon" municipality="Mt. Febe" name="Yaounde" pageId="12" pageNumber="13">Yaounde</location>
:
<collectingMunicipality id="647A3C15B91F7846C60F44C4FC602207" box="[800,905,1771,1797]" pageId="12" pageNumber="13">Mt. Febe</collectingMunicipality>
,
<specimenCount id="92A76DE6B91F7846C6B944C5FC0E2207" box="[918,999,1770,1797]" pageId="12" pageNumber="13" type="generic">
<quantity id="43590B8AB91F7846C6B944C5FC0A2207" box="[918,995,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="13" unit="m" value="950.0">
<elevation id="0F8C415CB91F7846C6B944C5FC0A2207" box="[918,995,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="13" unit="m" value="950.0">950 m</elevation>
</quantity>
.
</specimenCount>
, hygropetric,
<date id="F01F80AFB91F7846C1AB44C4FB152207" box="[1156,1276,1771,1797]" pageId="12" pageNumber="13" value="1982-04-10">
<collectingDate id="E05B7947B91F7846C1AB44C4FB152207" box="[1156,1276,1771,1797]" pageId="12" pageNumber="13" value="1982-04-10">10.iv.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B91F7846C02644C4FA752207" box="[1289,1436,1771,1797]" pageId="12" pageNumber="13">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB91F7846C43A4520FEB1222B" box="[277,344,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B91F7846C4454520FCAC222B" ID-GBIF-Occurrence="3962178342" box="[362,837,1807,1833]" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="12" pageNumber="13" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB91F7846C4454520FE39222B" box="[362,464,1807,1833]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
: Same data as holotype (1
<collectionCode id="E2B03EAAB91F7846C7D44520FCD4222B" box="[763,829,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7846C5E8451DFD562297" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="841EA66FB91F7846C5E8451DFD562297" blockId="12.[151,1436,1770,2013]" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E8451DFE3D224E" bold="true" box="[199,468,1842,1868]" pageId="12" pageNumber="13">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB91F7846C10B451CFB70224F" box="[1060,1177,1843,1869]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
by the combination of the small size (ca.
<quantity id="43590B8AB91F7846C4474578FE392273" box="[360,464,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="13" unit="mm" value="1.16">1.16 mm</quantity>
), the relatively large and wide pronotum (length/width ratio ca. 17/25), the moderately deep pronotal foveae, and the aedeagus (
<figureCitation id="1C9ABAEAB91F7846C7704554FD5B2297" box="[607,690,1915,1941]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91F7847C5E845B1FCEE27F3" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="841EA66FB91F7847C5E845B1FE142467" blockId="12.[151,1436,1770,2013]" lastBlockId="13.[151,1437,151,825]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B6D57A7DB91F7846C5E845B1FEB822BA" bold="true" box="[199,337,1950,1976]" pageId="12" pageNumber="13">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB91F7846C48B45B0FDEF22BB" box="[420,518,1951,1977]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.16/0.54; head width 0.33; pronotum 0.31/0.46, PA 0.39, PB 0.41; elytra 0.81/0.54. Habitus as illustrated (
<figureCitation id="1C9ABAEAB91F7846C12145ECFB8822DF" box="[1038,1121,1987,2013]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">Fig. 13</figureCitation>
). Dorsum light brown, head and pronotal macula darker (
<typeStatus id="5B1A18CDB91E7847C4F842B8FDD225B3" box="[471,571,151,177]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
is teneral), palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum densely moderately finely punctulate, interstices weakly shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 12xpd; punctures of clypeus very fine and dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum relatively large and wide, length/width ratio ca. 17/25. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae shallow, posterosubmedial foveae moderately deep.
</paragraph>
<paragraph id="841EA66FB91E7847C5E84340FCEE27F3" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="14">Elytra with posterior declivity very gradual, summit slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Venter: Mentum and postmentum finely sparsely punctulate, weakly shining. Genae very slightly raised, shining, with weak posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques moderately narrow, straight, slightly convergent anteriorly, weakly raised, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 1.5x P2. Pro- meso- and metatibiae moderately stout, straight. Abdominal apex symmetrical, without apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5E840D5FC212617" box="[199,968,762,789]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="841EA66FB91E7847C5E840D5FC212617" blockId="13.[151,1437,151,825]" box="[199,968,762,789]" pageId="13" pageNumber="14">
<emphasis id="B6D57A7DB91E7847C5E840D5FE882616" bold="true" box="[199,353,762,788]" pageId="13" pageNumber="14">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB91E7847C7E340D4FD152617" box="[716,764,763,789]" pageId="13" pageNumber="14">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB91E7847C64840D4FC522617" box="[871,955,763,789]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="13" pageNumber="14">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B91E7847C5E84130FB28263B" box="[199,1217,799,825]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="841EA66FB91E7847C5E84130FB28263B" blockId="13.[151,1437,151,825]" box="[199,1217,799,825]" pageId="13" pageNumber="14">
<emphasis id="B6D57A7DB91E7847C5E84130FEA7263B" bold="true" box="[199,334,799,825]" pageId="13" pageNumber="14">Etymology.</emphasis>
Named in reference to the known distribution and the very complex aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Guide to the Vascular Flora of the Savannas and Flatwoods of Shaken Creek Preserve and Vicinity (Pender &amp; Onslow Counties, North Carolina, U. S. A.)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thornhill, Robert</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krings, Alexander</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lindbo, David</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stucky, Jon</mods:namePart>
</mods:name>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
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<mods:start>1099</mods:start>
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<treatment LSID="urn:lsid:plazi:treatment:0C08488F2C6CD39C58F65CA1BBBCCDED" httpUri="http://treatment.plazi.org/id/0C08488F2C6CD39C58F65CA1BBBCCDED" lastPageNumber="1099" pageId="0" pageNumber="1099">
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<taxonomicName authority="(L.) Raf. ex DC." class="Magnoliopsida" family="Asteraceae" genus="Erechtites" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Erechtites hieraciifolius" order="Asterales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="species" species="hieraciifolius">Erechtites hieraciifolius (L.) Raf. ex DC.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1099" type="distribution">
<paragraph pageId="0" pageNumber="1099">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1099">Disturbed areas in pine savannas, dry edges of borrow pits, roadsides.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1099" type="notes">
<paragraph pageId="0" pageNumber="1099">Notes</paragraph>
<paragraph pageId="0" pageNumber="1099">
Rare. Late
<normalizedToken originalValue="JulNov">Jul-Nov</normalizedToken>
. Thornhill 938, 1370 (NCSC). Specimens seen in the vicinity: Sandy Run [Hancock]: Taggart SARU 407 (WNC!). [&lt;RAB; =
<taxonomicName basionym-authority="(L.) Raf. ex DC." class="Magnoliopsida" family="Asteraceae" genus="Erechtites" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Erechtites hieraciifolius var. hieraciifolius" order="Asterales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="variety" species="hieraciifolius" variety="hieraciifolius">Erechtites hieraciifolius (L.) Raf. ex DC. var. hieraciifolius</taxonomicName>
sensu FNA; = Weakley]
</paragraph>
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@ -0,0 +1,114 @@
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<mods:namePart>Sanches, Magda</mods:namePart>
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<mods:role>
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</mods:role>
<mods:namePart>Knoff, Marcelo</mods:namePart>
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<mods:role>
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<paragraph pageId="37" pageNumber="38">Taxon classification Animalia Dactylogyridea Dactylogyridae</paragraph>
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<paragraph pageId="37" pageNumber="38">
<taxonomicName authority="Boeger, Domingues &amp; Pavanelli, 1995" authorityName="Boeger, Domingues &amp; Pavanelli" authorityYear="1995" class="Monogenea" family="Dactylogyridae" genus="Rhinoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinoxenus bulbovaginatus" order="Dactylogyridea" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="bulbovaginatus">Rhinoxenus bulbovaginatus Boeger, Domingues &amp; Pavanelli, 1995</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="type host">
<paragraph pageId="37" pageNumber="38">Type host.</paragraph>
<paragraph pageId="37" pageNumber="38">
<taxonomicName class="Actinopterygii" family="Bryconidae" genus="Salminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salminus brasiliensis" order="Characiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="brasiliensis">Salminus brasiliensis</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="infection site">
<paragraph pageId="37" pageNumber="38">Infection site.</paragraph>
<paragraph pageId="37" pageNumber="38">Nasal cavities.</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="type locality">
<paragraph pageId="37" pageNumber="38">Type locality.</paragraph>
<paragraph pageId="37" pageNumber="38">
Brazil,
<normalizedToken originalValue="Paraná">Parana</normalizedToken>
State,
<normalizedToken originalValue="Paraná">Parana</normalizedToken>
River basin near Porto Rico.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="holotype">
<paragraph pageId="37" pageNumber="38">Holotype.</paragraph>
<paragraph pageId="37" pageNumber="38">CHIOC 33269 a.</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="paratypes">
<paragraph pageId="37" pageNumber="38">Paratypes.</paragraph>
<paragraph pageId="37" pageNumber="38">
CHIOC 33269
<normalizedToken originalValue="be">b-e</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="remarks">
<paragraph pageId="37" pageNumber="38">Remarks.</paragraph>
<paragraph pageId="37" pageNumber="38">Other paratypes deposited in the collections of HWML and USNM.</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="reference">
<paragraph pageId="37" pageNumber="38">Reference.</paragraph>
<paragraph pageId="37" pageNumber="38">
<bibRefCitation author="Boeger, WA" journalOrPublisher="Memorias do Instituto Oswaldo Cruz" pageId="64" pageNumber="65" pagination="695 - 698" title="Neotropical Monogenoidea. 24. Rhinoxenusbulbovaginatus n. sp. (Dactylogyridae, Ancyrocephalinae) from the nasal cavity of Salminusmaxillosus (Osteichthyes, Characidae) from the Rio Parana, Parana, Brazil." url="10.1590/S0074-02761995000600007" volume="90" year="1995 a">Boeger et al. (1995a)</bibRefCitation>
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<emphasis id="D78F9D2FD7D3861F14E657749FD4F6A6" italics="true" pageNumber="315">Aproteles bulmerae</emphasis>
Menzies 1977
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<emphasis id="933267721CBB6FB3D5660379AC4809EE" italics="true" pageNumber="315">Aproteles bulmerae</emphasis>
Menzies 1977
</taxonomicName>
,
<bibCitation id="431C9D546B77B2579862B6192178AF8A" author="Menzies" journal="Aust. J. Zool." pageNumber="315" pagination="331" title="Aproteles bulmerae" volume="25" year="1977">
<collectingRegion id="93C701302ECE12B30157348FE87E76B2" country="Cameroon" name="East">Aust</collectingRegion>
. J. Zool., 25: 331
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="427DCDD3766420D3EDC34E53A1FE3F7C" pageNumber="315" type="materials_examined">
<paragraph id="ABBF001D86F652A73FBCBB5B1CF6C27C" pageNumber="315">
<emphasis id="613E545E5E13A8D21D7CB3D30E9AC188" inLineHeading="true" pageNumber="315">Type Locality:</emphasis>
<materialsCitation id="3BF6E2A8CB8FC8C3324CC6E8DA375228" ID-GBIF-Occurrence="4867333421" country="Papua New Guinea" elevation="1.53" pageNumber="315" stateProvince="Chimbu">
<collectingCountry id="9CE55C16ECEA4384D3903B23277E35C2" name="Papua New Guinea">Papua New Guinea</collectingCountry>
,
<collectingRegion id="E85973268A6C03A6F03C1E45372EB455" country="Papua New Guinea" name="Chimbu">Chimbu Prov.</collectingRegion>
,
<quantity id="BA94D58E28965D5E4ACA9CEB76091260" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SE Chuave Govt. Sta.,
<quantity id="F435FE8A6CD0A6A6ED05C80C58CA5868" metricMagnitude="0" metricUnit="m" metricValue="1.53" unit="m" value="1.53">
<elevation id="3CB26A5ED7F1A423F27E2162CC7E680F">1,530 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D912CDC564326F768ADAA605AE6827DE" pageNumber="315" type="vernacular_names">
<paragraph id="77F9111A9F39B104E15636DDA884888F" pageNumber="315">
<emphasis id="3A9F8C051E7246174D5C6CEBCA962317" inLineHeading="true" pageNumber="315">Vernacular Names:</emphasis>
<vernacularName id="C22FF06466487E07A0DA0097BD53D046" language="eng" pageNumber="315">Bulmer's Fruit Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="926B8524D573C98CEF63D9A145D337FA" pageNumber="315" type="distribution">
<paragraph id="DE4221173E22974D39A4698ECD192D1D" pageNumber="315">
<emphasis id="607A5D5E1CC468B9FAE8422A559654AD" inLineHeading="true" pageNumber="315">Distribution:</emphasis>
Mainland
<collectingCountry id="16172A24556AAAA64244B7913FCB7C61" name="Papua New Guinea">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="AB80C22F0AD69AEF62E2EB61C145551F" pageNumber="315" type="conservation">
<paragraph id="EBA99BC8B6505CA75A315AFE59085CC1" pageNumber="315">
<emphasis id="6F7B38CE589AAF5E0C3F781B1C5B46E8" inLineHeading="true" pageNumber="315">Conservation:</emphasis>
<collectingCountry id="6D4CA725EC96A0B616060FA14ACF2C6B" name="United States of America">U.S.</collectingCountry>
ESA Endangered.
<collectionCode id="E431C78693BA42956208724FCC02A61C">IUCN</collectionCode>
/
<collectionCode id="7C08E50E68352BD630ECBA6EAEC41387" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (1992) Endangered: Limited Distribution.
<collectionCode id="5AF8CC21E4DD2DA3C63E5B88B1DCC43F">IUCN</collectionCode>
2003 Critically Endangered.
</paragraph>
</subSubSection>
<subSubSection id="69D18E6B62281F35958FC3457E93090F" pageNumber="315" type="discussion">
<paragraph id="49EAA1F1D2231244A17F93A8DD335A44" pageNumber="315">
<emphasis id="B5B1A2CC7BB52D5C371D58C62911B867" inLineHeading="true" pageNumber="315">Discussion:</emphasis>
Originally described from fossil material, but since found living (
<bibRefCitation id="EC52AED31F5FF78F9E22D941169CA643" author="Flannery, T. F. &amp; L. Seri" refId="ref103837" refString="Flannery, T. F., and L. Seri. 1993. Rediscovery of Aproteles bulmerae (Chiroptera: Pteropodidae). Morphology, ecology and conservation. Mammalia, 57 (1): 19 - 25." year="1993">Flannery and Seri, 1993</bibRefCitation>
;
<bibRefCitation id="457CDFBA68D08BE5B234E3F838648F00" author="Hyndman, D. &amp; J. I. Menzies" refId="ref164986" refString="Hyndman, D., and J. I. Menzies. 1980. Aproteles bulmerae (Chiroptera: Pteropodidae) of New Guinea is not extinct. Journal of Mammalogy, 61: 159 - 160." year="1980">Hyndman and Menzies, 1980</bibRefCitation>
). See also Flannery (1995
<emphasis id="AAD853104CEF516BB09FF781F25C0833" italics="true" pageNumber="315">a</emphasis>
) and
<bibRefCitation id="E0511F5F469F05BE583186DDEC99C939" author="Bonaccorso, F. J." refId="ref35461" refString="Bonaccorso, F. J. 1998. Bats of Papua New Guinea. Conservation International Tropical Field Guide Series, Conservation International, Washington, D. C., 489 pp." year="1998">Bonaccorso (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,72 @@
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<paragraph id="0D60D16A16E0D6BD0DBE5BF4459AD88E" pageNumber="47">
<materialsCitation id="FC8A94FE6149900C185037E5E8016261" ID-GBIF-Occurrence="923904093" country="Brazil" stateProvince="Para">
<collectionCode id="3E5E0B874686F6F35BE2D67D364ACB45" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZ</collectionCode>
7663, unsexed specimen, 154 mm SL,
<date id="616FB076E21356967EB47FC28615E073" month="08" value="1865-08" year="1865">August 1865</date>
,
<normalizedToken id="2F505A1177DCB60E9CC3D1A25EC3FA8A" originalValue="Pará">
<collectingRegion id="5619C23A67CCC6077B810E6848D33855">Para</collectingRegion>
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(
<normalizedToken id="28946F2E907C81DD4CDAF63F858E28ED" originalValue="Belém">Belem</normalizedToken>
),
<collectingCountry id="137BA634DE3EED8DF293A878C43B7830">Brazil</collectingCountry>
(
<geoCoordinate id="2EA5EB3EDBF7F6BCAE454C83000D96FC" direction="south" orientation="latitude" value="-1.45">1°27' S</geoCoordinate>
,
<geoCoordinate id="E4B453CA81FFF85392F052DCF17CD085" direction="west" orientation="longitude" value="-48.483334">48°29' W</geoCoordinate>
)
</materialsCitation>
,
</paragraph>
</subSubSection>
<subSubSection id="62BA7E6D1CF262C0EFD0938CF63A6ED9" type="nomenclature">
<paragraph id="BBDADDCEC95C1FBC7D7D0CD083FC5AB2" pageNumber="47">
<taxonomicName id="50F75ED45C41BC42D80243B9398216ED" ID-CoL="5XG5Y" family="Ariidae" genus="Cathorops" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="47" phylum="Chordata" rank="species" species="spixii">C. spixii</taxonomicName>
;
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,246 @@
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<mods:namePart>Fassbender, Nico</mods:namePart>
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<mods:nameIdentifier type="email">nico@nektonmission.org</mods:nameIdentifier>
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<mods:namePart>Stefanoudis, Paris V</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:role>
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<mods:role>
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<mods:affiliation>University of Cape Town, Rondebosch, Cape Town, South Africa &amp; blue [c] weed, Brest, France</mods:affiliation>
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<mods:role>
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</mods:name>
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<mods:affiliation>OKEANOS / DOP, University of the Azores, Horta, Portugal</mods:affiliation>
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<mods:role>
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<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wagner, Daniel</mods:namePart>
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</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Walton, Rowana</mods:namePart>
<mods:affiliation>James Michel Blue Economy Research Institute, University of Seychelles, Anse Royale, Mahe ́, Seychelles</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Woodall, Lucy C</mods:namePart>
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<mods:affiliation>Department of Zoology, University of Oxford, Oxford, United Kingdom &amp; Nekton Foundation, Oxford, United Kingdom</mods:affiliation>
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<paragraph pageId="0" pageNumber="65970">Materials</paragraph>
<paragraph pageId="0" pageNumber="65970">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="Submersible OR Remotely Operated Vehicle OR SCUBA" collectorName="Taxon, Hydrozoa, Animalia, Cnidaria, Owen, Location, Indian Ocean, Aldabra N, Aldabra W, Alphonse N, Astove W, D'Arros N, Desroches S, Submersible OR Remotely Operated Vehicle OR" country="Seychelles" county="Human" determinerName="Nico Fassbender, Carlos Moura, Paris Stefanoudis" elevation="16" location="Aldabra N 1, Aldabra W 1, Alphonse N 1, Astove W 1, D'Arros N 1, Desroches S 1" municipality="Record Level" pageId="0" pageNumber="65970" specimenCount="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="65970">Type status:</emphasis>
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.
<emphasis bold="true" pageId="0" pageNumber="65970">
<collectorName>Taxon</collectorName>
:
</emphasis>
scientificName:
<collectorName>Hydrozoa</collectorName>
; kingdom:
<collectorName>Animalia</collectorName>
; phylum:
<collectorName>Cnidaria</collectorName>
; class:
<collectorName>Hydrozoa</collectorName>
; scientificNameAuthorship:
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, 1843;
<emphasis bold="true" pageId="0" pageNumber="65970">
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:
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waterBody:
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; country:
<collectingCountry name="Seychelles" pageId="0" pageNumber="65970">Seychelles</collectingCountry>
; locality:
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:AEC4B6B8D72B51BCAEA0472B5AA6DAE4" country="Seychelles" county="Human" municipality="Record Level" name="Aldabra N 1, Aldabra W 1, Alphonse N 1, Astove W 1, D'Arros N 1, Desroches S 1" pageId="0" pageNumber="65970">
<collectorName>Aldabra N</collectorName>
1,
<collectorName>Aldabra W</collectorName>
1,
<collectorName>Alphonse N</collectorName>
1,
<collectorName>Astove W</collectorName>
1,
<collectorName>D'Arros N</collectorName>
1,
<collectorName>Desroches S</collectorName>
1
</location>
; minimumDepthInMeters:
<quantity metricMagnitude="1" metricUnit="m" metricValue="1.55" unit="m" value="15.5">
<elevation metricMagnitude="1" metricUnit="m" metricValue="1.55" unit="m" value="15.5">15.5 m</elevation>
</quantity>
; maximumDepthInMeters:
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.385" unit="m" value="138.5">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.385" unit="m" value="138.5">138.5 m</elevation>
</quantity>
; locationRemarks:
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:346DF062BDDC56D93C93EBDCF9969CC5" country="Seychelles" county="Human" municipality="Record Level" name="First Descent">First Descent</location>
:
<collectingCountry name="Seychelles">Seychelles</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:8D7F0FFECE4D13DF8BE97A7B1DDCBCDA" country="Seychelles" county="Human" municipality="Record Level" name="Expedition">Expedition</location>
;
<emphasis bold="true" pageId="0" pageNumber="65970">
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:84988ABB1D621EFBE6D6B2E033C611A9" country="Seychelles" county="Human" municipality="Record Level" name="Identification">Identification</location>
:
</emphasis>
identifiedBy:
<determinerName pageId="0" pageNumber="65970">
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:DFB389632FCEBA73A69BAC32146B513E" country="Seychelles" county="Human" municipality="Record Level" name="Nico Fassbender">Nico Fassbender</location>
,
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:2898D72808F6E50405A097E8EBDB97F3" country="Seychelles" county="Human" municipality="Record Level" name="Carlos Moura">Carlos Moura</location>
,
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:B522AE8C356AA0FBBA9FE5BF663A0E8B" country="Seychelles" county="Human" municipality="Record Level" name="Paris Stefanoudis">Paris Stefanoudis</location>
</determinerName>
; dateIdentified: 2019, 2020; identificationRemarks: identified only from imagery;
<emphasis bold="true" pageId="0" pageNumber="65970">
<location LSID="urn:lsid:plazi:treatment:0C098E508E3D5218B766C476E2AEF19B:D2F4CB860CFE79170D1D97286B0EEFCE" country="Seychelles" county="Human" municipality="Record Level" name="Event">Event</location>
:
</emphasis>
samplingProtocol:
<collectingMethod pageId="0" pageNumber="65970">
<collectorName>Submersible OR Remotely Operated Vehicle OR</collectorName>
SCUBA
</collectingMethod>
;
<emphasis bold="true" pageId="0" pageNumber="65970">
<collectingMunicipality>Record Level</collectingMunicipality>
:
</emphasis>
basisOfRecord:
<collectingCounty>Human</collectingCounty>
observation
</materialsCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65970" type="notes">
<paragraph pageId="0" pageNumber="65970">Notes</paragraph>
<paragraph pageId="0" pageNumber="65970">
Often colonial, with varied branching forms and dimensions (few millimetres to few tenths of centimetres). Colouration variable. Aggregations of hydroid colonies commonly overgrow rocks and portions of dead corals. Some hydrozoans bear resemblance to octocorals, but the latter have a much more solid structure and tend to be larger. This group likely contains a variety of species that are difficult to identify from video footage; hence, no attempt was made to identify them at a lower taxonomic level. Members of the
<taxonomicName authorityName="Fleming" authorityYear="1828" class="Hydrozoa" family="Milleporidae" kingdom="Animalia" lsidName="" order="Anthoathecata" pageId="0" pageNumber="65970" phylum="Cnidaria" rank="family">Milleporidae</taxonomicName>
,
<taxonomicName authorityName="Gray" authorityYear="1847" class="Hydrozoa" family="Stylasteridae" kingdom="Animalia" lsidName="" order="Anthoathecata" pageId="0" pageNumber="65970" phylum="Cnidaria" rank="family">Stylasteridae</taxonomicName>
and
<taxonomicName authorityName="Stechow" authorityYear="1920" class="Hydrozoa" family="Thyroscyphidae" kingdom="Animalia" lsidName="" order="Leptothecata" pageId="0" pageNumber="65970" phylum="Cnidaria" rank="family">Thyroscyphidae</taxonomicName>
families may be more easily identifiable from video footage (see below). Commonly known as hydroids (Fig.
<figureCitation captionStartId="F6743367" pageId="0" pageNumber="65970">107</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,236 @@
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<mods:title id="FB1EF5038E90039DD7F76E81A87E9421">Fossil flying squirrels (Petauristinae, Sciuridae, Rodentia) from the Yumidong Cave in Wushan County, Chongqing, China</mods:title>
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<mods:namePart id="86CD93581F12FEC7448F511C61A0C1F1">Pang, Li-bo</mods:namePart>
<mods:affiliation id="4B88D11EB7AF2FA6AF0EEA9C35564596">Hebei International Joint Research Center for Paleoanthropology, College of Earth Sciences, Hebei GEO University, Shijiazhuang 050031, Hebei, China</mods:affiliation>
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<mods:namePart id="6A23B2EB8AA0827D71170E78B333F5F4">Chen, Shao-kun</mods:namePart>
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<mods:namePart id="7B426DEB1041EB2C3A949FE553E40E85">Hu, Xin</mods:namePart>
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<mods:namePart id="80AE60A568BC4A6C44404F216F35C6FC">Wu, Yan</mods:namePart>
<mods:affiliation id="EB3D53F9423CB67D99855FC2960EFD84">China Three Gorges Museum, Chongqing 400015, China</mods:affiliation>
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<mods:namePart id="776A2CE08CB403CF8DAB3650A99E7885">Wei, Guang-biao</mods:namePart>
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<mods:date id="8E830E3EB4F768A322376745606D4E53">2024</mods:date>
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<treatment id="0C0A10DB757F55D180CAE1D91267B49C" ID-DOI="http://doi.org/10.5281/zenodo.11236934" ID-Zenodo-Dep="11236934" ID-arpha="0C0A10DB-757F-55D1-80CA-E1D91267B49C" LSID="urn:lsid:plazi:treatment:0C0A10DB757F55D180CAE1D91267B49C" httpUri="http://treatment.plazi.org/id/0C0A10DB757F55D180CAE1D91267B49C">
<subSubSection id="3F0A27376FB05B89851350A1D48C8009" type="nomenclature">
<paragraph id="022E25687BD09511DA4FFF937F83AF5F">
<taxonomicName id="3551C361B4F8A706A58D9CBD437A6364" ID-CoL="64YQ9" authority="(Milne-Edwards, 1867)" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="9885E44698664F6BB32E65E419B1D20F" italics="true">Aeretes melanopterus</emphasis>
(Milne-Edwards, 1867)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C6B3D808A50D4F458DDC3364DF5E3B33" type="description">
<paragraph id="F920B6E0079D8102FD9D5DE4D9D548D5">
<figureCitation id="09452BD9FA22926433D7600A638485F3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Aeretes melanopterus from the Yumidong Cave. A. 12 YMDT 4 ④ 60.54, right DP 4 - M 1, occlusal view; B. 12 YMDT 4 ② 35.81, right P 4, occlusal view; C. 12 YMDT 4 ④ 60.1, left mandible, labial view; D. 12 YMDT 4 ④ 60.2, right p 4 - m 3, occlusal view. Scale bar: 5 mm (A, B, D); 10 mm (C)." figureDoi="10.3897/fr.27.e115693.figure4" httpUri="https://binary.pensoft.net/fig/1053349">Fig. 4</figureCitation>
<xref id="AB5B1FA56316AE0CFEC7EBEB62F08D24" ref-type="supplementary-material" rid="S1">; Suppl. material 1: table S 5</xref>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EISAE" type="material">
<paragraph id="1FD3C728A73676AE6D4A5C5586E9656E">
<heading id="FDF4C118229CAACE3EE90E0F011DACD2" reason="title">Materials.</heading>
</paragraph>
<paragraph id="F6DC99B32C679E66F9C9CF98066CC782">
As in Suppl. material
<xref id="2C0A412549C0CE2C5A1F421D899BE90F" ref-type="supplementary-material" rid="S1">1</xref>
: table S 1, there are one isolated tooth from the layer ②-2 and three maxillary bones and two mandibular bones from the layer ④.
</paragraph>
<caption id="D1F660F55917CF8F04BC4F54C117D130" ID-DOI="10.3897/fr.27.e115693.figure4" ID-arpha="6785F345-AAF5-5F0A-88AC-90A8F1C9D9AE" httpUri="https://binary.pensoft.net/fig/1053349" startId="F4">
<paragraph id="B6719E61C962D3A7C7B00ED9842A9CBC">
<label id="6524FF5FFEF49C6D1AF00B2A378133A7">Figure 4.</label>
</paragraph>
<paragraph id="7C375BE8A1222450850D565C824D11D1">
<taxonomicName id="25243C64B4FCE63B4DD609E381798CF6" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="5446C13B988A0B26470A5EBC3DE95E66" italics="true">Aeretes melanopterus</emphasis>
</taxonomicName>
from the Yumidong Cave.
<emphasis id="C0D90CF8477F26CA7B757DC12EA92DC9" bold="true">A.</emphasis>
12 YMDT 4 ④ 60.54, right DP 4 - M 1, occlusal view;
<emphasis id="547B945C6F66599FA332654D08915565" bold="true">B.</emphasis>
12 YMDT 4 ② 35.81, right P 4, occlusal view;
<emphasis id="AB5F77E5736BEEF86DEC99B7DF9C8B1A" bold="true">C.</emphasis>
12 YMDT 4 ④ 60.1, left mandible, labial view;
<emphasis id="7F1734BC2AA963C1A241AD117BBA2E1A" bold="true">D.</emphasis>
12 YMDT 4 ④ 60.2, right p 4 - m 3, occlusal view. Scale bar: 5 mm (
<emphasis id="3CDE1425529DEE1FB1A3A55647BF7CB2" bold="true">A, B, D</emphasis>
); 10 mm (
<emphasis id="6A21B39A3F66F7041615CEF67A2D03B5" bold="true">C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWTAE" type="description">
<paragraph id="DFDBCFFBC6AC837A802E0B07055F94B3">
<heading id="3A5162FC889E1986A5CC6F65E2C12155" reason="title">Description.</heading>
</paragraph>
<paragraph id="1723F84C70ADA7719F5BF0FB199166F5">The mandible is robust and the diastemal portion is short. The tip of the incisor is slightly higher than the worn surface of the cheek teeth. Its posterior end lies under the posterior root of m 3. The diastema is short and a medium-sized mental foramen is located at the middle part of the buccal side of the diastema. The inferior margin of the horizontal ramus is smoothly curved. A vascular notch is obvious and under the masseter muscle fossa. The masseter muscle fossa is relatively shallow with a weak ridge. Its anterior angle is at the level of the anterior root of m 1. The pterygoid muscle fossa at the lingual side is very wide and deep. Its anterior end is at the level of the posterior part of m 3. The mandibular foramen is large, oval-shaped and positioned more ventrally than the worn surface of the cheek teeth. The angular process is broken, but the preserved part shows it should be almost as wide as the ascending ramus. The upper part of the ascending ramus is badly preserved, but it seems not very high. The condylar process is a short transverse axis and its neck is long. The coronoid process is thin and higher than the condylar process.</paragraph>
<paragraph id="D4E71D965CD471EF3E4EA397B6F0FB78">DP 4 is smaller than P 4 and M 1. Its occlusal outline is triangular. The parastyle is fused with the anteroloph, forming a high and isolated anterolingual corner. Besides the parastyle, the paracone, the metacone and the protocone are all well developed on the crown surface. The protostyle is invisible. The hypocone is very weak, close behind the protocone. The protoloph is short and straight. No obvious protoconule. The metaloph bends back at the metaconule. The enlarged metaconule is connected with the posteroloph by a short ridge, forming a small hypoconule on the posteroloph and dividing the posterior valley into two. The posteroloph is low. The posterolingual flexus is narrow and closed after being moderately worn. The anterior valley and the central valley are short and the anterior one is wider than the central one. There are one large root at the lingual side and two small ones at the buccal side.</paragraph>
<paragraph id="41F558FF0E34FDCDA1FB4E8FB9892A69">P 3 is not preserved. On the basis of the alveolar, it should be single-rooted, not particularly small and visible from the buccal side.</paragraph>
<paragraph id="EAFCE2F6D1DEF749CD9E729251868415">P 4 is molariformed. Its lingual side is much shorter than the buccal side, making its occlusal outline triangular. The lingual wall is wrinkled and higher than the buccal wall. The protocone is small and separated from the developed hypocone by a vertical groove (the anterolingual flexus) which extends to the base of the tooth crown. The anterolingual flexus is shallow, visible after being moderately worn. The posterolingual flexus is quite deep and narrow. There are four transverse ridges on the crown surface (the anteroloph, the protoloph, the metaloph and the posteroloph). The anteroloph is well developed and the parastyle is fused with it. The anteroloph is connected with the protocone at the slightly worn stage, but separated after being moderately worn by the anterolingual flexus. The protoloph is relatively straight. There are two ridges extending from the protoconule, the smaller one backwards and the larger one forwards. The metaconule is marked and the metaloph bends back at the metaconule. There are three small ridges extending backwards from the metaloph and two of them are connected with the posteroloph. The posteroloph is continuous and is connected with the hypocone only after being very deeply worn. Three roots, one larger at the lingual side and two smaller at the buccal side.</paragraph>
<paragraph id="3FDB44D77496C95EB187542FCB7517D5">The occlusal outline of M 1 is close to a square. The occlusal structure is mainly composed of four cusps (the paracone, the metacone, the protocone and the hypocone) and four transverse ridges (the anteroloph, the protoloph, the metaloph and the posteroloph). The anteroloph and the posteroloph are lower than the protoloph and the metaloph. The protocone is not completely separated from the hypocone and both of them are ridge-shaped. The anteroloph and the protoloph converge with the protocone. The metaloph is connected with the hypocone. The paraconule and the metaconule are small. The main structure of the posterior lobe is similar to that of DP 4.</paragraph>
<paragraph id="494F0BDB698FCB36ECE3304CB48D8809">The occlusal outline of p 4 is a trapezoid with a narrow anterior lobe and a wide posterior lobe. The anteroconid is small, located at the anterobuccal side of the metaconid. The metaconid is the highest amongst all cusps of the tooth. The metastylid is barely visible and is separated from the well-developed mesostylid by a deep groove. The protoconid is weaker than the metaconid, but larger than other cusps. The anterobuccal sinusid is shallow and V-shaped. The buccal valley is wide. The mesoconid is obvious. The mesolophid divides the talonid basin into two. The protolophulid is tiny and the extra posterolophid is short. The posterolophid is curved. Two roots.</paragraph>
<paragraph id="50E547E51BF2D881FD488912C6243AEE">The lower molars have similar occlusal structure with p 4. The anterolophid is developed and connected with the anteroconid and the metaconid. The anterobuccal sinusid is much deeper than that of p 4. The mesoconid is connected with the protoconid. The metalophid is more developed than that of p 4. The posterior lobe of m 3 tapers obviously with a weak entoconid. Four roots.</paragraph>
</subSubSection>
<subSubSection id="SECID0EBUAE" type="comparisons">
<paragraph id="B0897718714EDA37F25BD513875E5015">
<heading id="B659FDA8123D25282B5F28D731073847" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="29A1F186F097CDF8DEF0A17696FF1D11">
Based on the dental dimensions (in Suppl. material
<xref id="60FA3D8AAD69CE8026FBD242BCF25F09" ref-type="supplementary-material" rid="S1">1</xref>
: table S 5), the described fossils should be a large species of flying squirrel, which is most likely to belong to either
<taxonomicName id="6D6B3D942B9413A22BECA1541C9501A0" authorityName="Allen" authorityYear="1940" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="0358F6968E1A831A8CCC685F28D824E9" italics="true">Aeretes</emphasis>
</taxonomicName>
or
<taxonomicName id="F56F478653A563757F86DEBD1E4513B4" authorityName="Link" authorityYear="1795" class="Mammalia" family="Sciuridae" genus="Petaurista" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="D2391D1DD6B5F577858E77A11840B308" italics="true">Petaurista</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F52DB12EF144BC953562A1917446D646">
In
<taxonomicName id="74B8D2EF998E5B26E78602644E4DF5D1" authorityName="Link" authorityYear="1795" class="Mammalia" family="Sciuridae" genus="Petaurista" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="78DC30AC790072C027265CB896E9DAC9" italics="true">Petaurista</emphasis>
</taxonomicName>
, M 1 / 2 lacks the hypocone and the lower cheek teeth have marked anterobuccal sinusid, which can be used for generic diagnosis and are obviously different from the features of the described fossils.
</paragraph>
<paragraph id="E8802CA6D24784076927ADB18C45BF9B">
The diagnosis of the genus
<taxonomicName id="44E72906912241C8DE1A98B042F6936E" authorityName="Allen" authorityYear="1940" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="1CCBB9869C78A0ED29900B3EF7938D7F" italics="true">Aeretes</emphasis>
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includes (summarised by
<bibRefCitation id="FC9B4A325BF9ED3E45D5A28D388A8FDB" author="Li CK &amp; Qiu ZD &amp; Tong YS &amp; Wang BY &amp; Wu WY &amp; Li Q &amp; Li Q &amp; Wang Y" journalOrPublisher="Science Press, Beijing" refId="B13" refString="Li CK, Qiu ZD, Tong YS, Wang BY, Wu WY, Li Q, Li Q, Wang Y (2019 a) Palaeovertebrata Sinica. Volume Ⅲ Fascicle 5 (1) Glires Ⅱ: Rodentia . Science Press, Beijing, 545 pp." title="Palaeovertebrata Sinica. Volume Ⅲ Fascicle 5 (1) Glires Ⅱ: Rodentia .">Li CK et al. (2019 a</bibRefCitation>
)): P 3 small, P 4 larger than molars, the hypocone of P 4 - M 2 weak and located close behind the protocone, the protoconule and the metaconule weak, the hypoconule well developed, the posterolingual flexus very deep, the entoconid of lower molars small, the mesostylid developed and separated from the metastylid and entoconid by deep grooves and the mesoconid small. The described fossils resemble
<taxonomicName id="2B214B8A0843DBC46AAF81CE4EA353CF" authorityName="Allen" authorityYear="1940" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="547A8D4097C4FD3CBF0FAB4A886F3867" italics="true">Aeretes</emphasis>
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.
</paragraph>
<paragraph id="0D945A2149C7077CD3AA10A46C3977DC">
There are three species in the genus
<taxonomicName id="A3784CCB96804076B41AF19281B3834E" authorityName="Allen" authorityYear="1940" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="D30CF1088413A7B67F9CFD488AD08B2E" italics="true">Aeretes</emphasis>
</taxonomicName>
,
<taxonomicName id="55CA8459AD4A84E2676BDB1613FAB240" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="premelanopterus">
<emphasis id="548E9FA3F97436E33A2198EB2716A993" italics="true">A. premelanopterus</emphasis>
</taxonomicName>
,
<taxonomicName id="AFA39FE7E0B3F0AF296A68642936BC4D" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="grandidens">
<emphasis id="C80F01BC771BC7CBAA222BE878DA0782" italics="true">A. grandidens</emphasis>
</taxonomicName>
and
<taxonomicName id="AD3EE3B0D7EC6973F8CBC8C294711699" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="29635354EAB10607BEC91492ABADD3D1" italics="true">A. melanopterus</emphasis>
</taxonomicName>
. The primitive
<taxonomicName id="80E45E85983F0B69B3D553925670D4ED" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="premelanopterus">
<emphasis id="D128FA4C8B2AF1B662AC5611B05AA4EA" italics="true">A. premelanopterus</emphasis>
</taxonomicName>
has weak protoconule, but strong metaconule;
<taxonomicName id="D80D851A575DB1ECD10622CC62A5B7EF" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="grandidens">
<emphasis id="FC47A7AA2BC338D6D0E5F2A60EDDEA0C" italics="true">A. grandidens</emphasis>
</taxonomicName>
has stronger protoconule and metaconule than
<taxonomicName id="84449F23D93C24EA5C640273F1908324" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="premelanopterus">
<emphasis id="E5810FE3763F81802376EDDFD02068E2" italics="true">A. premelanopterus</emphasis>
</taxonomicName>
. The protoconule and metaconule of
<taxonomicName id="76E3E96E7735319EF3D93BB03AB1FA0F" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="0746C3A2EEF58B29426ECFD54CDCFB67" italics="true">A. melanopterus</emphasis>
</taxonomicName>
from different regions varies in morphology: specimens from
<collectingRegion id="C3840EC967FD36EAFA0103C63A831397" country="China" name="Chongqing">Chongqing</collectingRegion>
and
<collectingRegion id="157F77BE3D31B12E84C42FB9CBFCFBBD" country="China" name="Guizhou">Guizhou</collectingRegion>
have a very weak protoconule and developed metaconule (
<bibRefCitation id="5BCFC68A6A10F73391A3F33162A3D9A3" author="Zheng" firstAuthor="Zheng" journalOrPublisher="Science Press, Beijing" refId="B32" refString="Zheng SH (1993) Quaternary rodents of Sichuan-Guizhou area, China. Science Press, Beijing, 270 pp." title="Quaternary rodents of Sichuan-Guizhou area, China." year="1993">Zheng 1993</bibRefCitation>
), while specimens from
<collectingRegion id="8EDB4D1D7BD29BF685002764C723F785" country="China" name="Beijing">Beijing</collectingRegion>
have a marked protoconule and metaconule (
<bibRefCitation id="4BBFB6531A5A33B1C10BB3175E5F76D8" DOI="10.1016/j.geobios.2006.04.006" author="Tong" firstAuthor="Tong" journalOrPublisher="Geobios" pagination="219-230" refId="B23" refString="Tong HW (2007) Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China. Geobios 40: 219230. https://doi.org/10.1016/j.geobios.2006.04.006" title="Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China." volume="40" year="2007">Tong 2007</bibRefCitation>
). The Yumidong specimens are most similar to
<taxonomicName id="B1CDC3FAAE70FD57C7676EC303566D39" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="E6A194E81728EC2567D944D08301E097" italics="true">A. melanopterus</emphasis>
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from Beijng. Furthermore, comparing the Yumidong specimens with
<taxonomicName id="0C97A0B70169705C1254A53A9A813730" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="D2B0651F896269FC7FB82BB7C1225269" italics="true">A. melanopterus</emphasis>
</taxonomicName>
from the Baotansi Cave in
<collectingRegion id="DE4AA9E94FD4AA1A17046B5C528B57D5" country="China" name="Chongqing">Chongqing</collectingRegion>
, there is another obvious dental morphological difference: the lingual vertical groove of P 4 is clear on the Yumidong specimens, but unclear on the Baotansi specimens.
</paragraph>
<paragraph id="7FB83722CAD1868DF4C157D352ACC68B">
Compared with the
<taxonomicName id="4F4A1986B3D7998237BC40E9426800DC" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="203CF609EC7F081A03E60B44E08A2EB6" italics="true">Aeretes melanopterus</emphasis>
</taxonomicName>
from the Tianyuan Cave and extant specimens (based on
<bibRefCitation id="3D2D575D255960AE4EC700C714B98996" DOI="10.1016/j.geobios.2006.04.006" author="Tong HW" firstAuthor="Tong" journalOrPublisher="Geobios" pagination="219-230" refId="B23" refString="Tong HW (2007) Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China. Geobios 40: 219230. https://doi.org/10.1016/j.geobios.2006.04.006" title="Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China." volume="40" year="2007">Tong (2007)</bibRefCitation>
), the morphological structure of the described specimens falls within their variable range. Based on dimensions, the described specimens are much smaller than the Tianyuan Cave specimens and fall within the variable range of the living species. It is reasonable to attribute these fossils to
<taxonomicName id="D820C36B5EC3321DC28D74787DA24714" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1867" class="Mammalia" family="Sciuridae" genus="Aeretes" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="melanopterus">
<emphasis id="C3054AC8F11E6AE02117A020F3D3B01A" italics="true">A. melanopterus</emphasis>
</taxonomicName>
.
<bibRefCitation id="4EFCFDBDF035E078663931BF9DC165E3" DOI="10.1016/j.geobios.2006.04.006" author="Tong HW" firstAuthor="Tong" journalOrPublisher="Geobios" pagination="219-230" refId="B23" refString="Tong HW (2007) Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China. Geobios 40: 219230. https://doi.org/10.1016/j.geobios.2006.04.006" title="Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China." volume="40" year="2007">Tong (2007)</bibRefCitation>
mentioned the diagnosis of this species, including: cheek teeth subhypsodont, enamel slightly rugose, the lingual wall of upper cheek teeth higher than the buccal wall, the posterolingual flexus very narrow and deep.
</paragraph>
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<taxonomicName id="E9ADACE890C66F9014D849E029D6C158" ID-CoL="6N48R" ID-ENA="1217963" authority="Wesmael, 1848" authorityName="Wesmael" authorityYear="1848" class="Insecta" family="Ichneumonidae" genus="Ichneumon" kingdom="Animalia" lsidName="Ichneumon tuberculipes" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="tuberculipes">Ichneumon tuberculipes Wesmael, 1848</taxonomicName>
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<taxonomicName id="600A56435AB616D93C45BB0968868A9E" class="Insecta" family="Ichneumonidae" genus="Ethelurgina" kingdom="Animalia" lsidName="Ethelurgina cuneatus" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="cuneatus">cuneatus</taxonomicName>
Tischbein, 1876
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<taxonomicName id="EF168C8C2A6E58D7E4240A33B173999C" class="Insecta" family="Ichneumonidae" genus="Ethelurgina" kingdom="Animalia" lsidName="Ethelurgina limbatus" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="limbatus">limbatus</taxonomicName>
Tischbein, 1879
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Tischbein, 1879
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Schmiedeknecht, 1930
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<treatment ID-GBIF-Taxon="189599956" LSID="urn:lsid:zoobank.org:act:35B7C13E-13A9-4F89-9291-C9EC65A432F8" httpUri="http://treatment.plazi.org/id/0C0B26B5A29A80A7728539C16A989B96" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/35B7C13E-13A9-4F89-9291-C9EC65A432F8" authority="Filander" authorityName="Filander" authorityYear="2021" family="Dendrophylliidae" genus="Ednapsammia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ednapsammia columnapriva" order="Scleractinia" pageId="0" pageNumber="1" rank="species" species="columnapriva" status="sp. nov.">Ednapsammia columnapriva Filander</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A, B Dendrophyllia sp. 1 (SAMC _ A 090158, off Knysna, 333 m A close-up calicular view B lateral view C, G Ednapsammia columnapriva sp. nov. C, D (SAM _ H 1441, off Mossel Bay, 212 m) C calicular view D lateral view E-G (SAMC _ A 090159, off Gouritsmond, 333 m) E calicular view F lateral view G calicular view H Enallopsammia pusilla (DSCS-INV 238, off Gouritsmond, 333 m): lateral view. I, J Enallopsammia rostrata (SAMC _ A 073270, off Port St Johns, 200 m) I close up calicular view J lateral view K, L Endopachys grayi (DEFF _ NANSEN-INV 32, off Shaka's Rock, 185 m) K calicular view L lateral view M, N Endopsammia philippensis (DIIIb 1, off Durban, 442 m) M calicular view N lateral view O, P Heteropsammia cochlea (EIe 1, locality data unknown) O calicular view P lateral view. Scale bars: 10 mm (A-C, E-I, K-P); 100 mm (D, J)." figureDoi="10.3897/zookeys.1066.69697.figure9" httpUri="https://binary.pensoft.net/fig/604112" pageId="0" pageNumber="1">Fig. 9C-G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Off Knysna, South Africa (RV
<normalizedToken originalValue="Algoa">'Algoa'</normalizedToken>
stn. DCS13:
<geoCoordinate degrees="35" direction="south" minutes="07" orientation="latitude" precision="1" seconds="11.34" value="-35.119816">35°07'11.34&quot;S</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="02" orientation="longitude" precision="1" seconds="41.91" value="23.044975">23°02'41.91&quot;E</geoCoordinate>
); 333 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
- SAMC_A090159 (
<specimenCount type="generic">1 specimen</specimenCount>
): Southern margin,
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.16" unit="km" value="116.0">116 km</quantity>
from Gouritsmond/off Goukamma Estuary,
<geoCoordinate degrees="35" direction="south" minutes="07" orientation="latitude" precision="1" seconds="11.34" value="-35.119816">35°07'11.34&quot;S</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="02" orientation="longitude" precision="1" seconds="41.91" value="23.044975">23°02'41.91&quot;E</geoCoordinate>
;
<specimenCount type="generic">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.3299999999999996" unit="m" value="333.0">333 m</quantity>
.
</specimenCount>
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
-DSCS-INV 226 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 227 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 229 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 231 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 232 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 235 (
<specimenCount type="generic">1 specimen</specimenCount>
), DSCS-INV 238 (
<specimenCount type="generic">1 specimen</specimenCount>
), SAMC_A090149 (
<specimenCount type="generic">1 specimen</specimenCount>
): Southern margin,
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.16" unit="km" value="116.0">116 km</quantity>
from Gouritsmond/off Goukamma Estuary,
<geoCoordinate degrees="35" direction="south" minutes="07" orientation="latitude" precision="1" seconds="11.34" value="-35.119816">35°07'11.34&quot;S</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="02" orientation="longitude" precision="1" seconds="41.91" value="23.044975">23°02'41.91&quot;E</geoCoordinate>
;
<specimenCount type="generic">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.3299999999999996" unit="m" value="333.0">333 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="imagery data">
<paragraph pageId="0" pageNumber="1">Imagery data.</paragraph>
<paragraph pageId="0" pageNumber="1">
SAM_H1441: Southern margin, 3 km from Mosselbaai/11 km off Hartenbos Estuary,
<geoCoordinate degrees="34" direction="south" minutes="10" orientation="latitude" precision="1" seconds="59.99" value="-34.18333">34°10'59.99&quot;S</geoCoordinate>
,
<geoCoordinate degrees="22" direction="east" minutes="10" orientation="longitude" precision="1" seconds="00.00" value="22.166666">22°10'00.00&quot;E</geoCoordinate>
; 216 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species name
<taxonomicName authorityName="Filander" authorityYear="2021" family="Dendrophylliidae" genus="Ednapsammia" kingdom="Animalia" lsidName="Ednapsammia columnapriva" order="Scleractinia" pageId="0" pageNumber="1" rank="species" species="columnapriva">
<emphasis italics="true" pageId="0" pageNumber="1">Ednapsammia columnapriva</emphasis>
</taxonomicName>
(derived from Latin
<emphasis italics="true" pageId="0" pageNumber="1">columna</emphasis>
meaning pillar and
<emphasis italics="true" pageId="0" pageNumber="1">privus</emphasis>
meaning deprived of) alludes to the lack of columella.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Corallum dendroid formed by extra-tentacular budding from base and from theca of parent corallite. Holotype consists of 19 corallites, and 54.7 mm in H. Calice circular to elliptical (GCD:LCD = 1.0-1.1), calicular margin finely laciniate. Thin epitheca, predominantly porous near calicular margin. Costae thick, equal in width, granular, extending to base, and separated by deep intercostal striae. Corallum white.</paragraph>
<paragraph pageId="0" pageNumber="1">Septa hexamerally arranged in three cycles, last cycle being incomplete, according to the formula: S1&gt; S2&gt; S3 (≤ 21 septa). S1 extend to fossa, sometimes almost meeting opposite septa. S2 ~ 1/3 smaller than S1. Third septal cycle incomplete, usually with only one S3 per half-system. S31/3 the width of S2. Primary and secondary septa cycles (S1-2) with straight and vertical axial margins, whilst S3 bears slightly laciniate axial margins. Fossa deep, columella absent.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Regional: Southern margin of South Africa, extending from off Mosselbaai towards Gouritsmond; 216-333 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Examined specimens are easily distinguished from other colonial dendrophylliids by their septa being hexameral and arranged normally (no
<normalizedToken originalValue="Pourtalès">Pourtales</normalizedToken>
plan), in three incomplete cycles. Despite that the imaged specimen (SAM_H1441 [in part: 1 specimen]) was not traceable in the Iziko Museums collection. This record nonetheless confirms that such a form was historically collected off the southern margin of South Africa.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="0C23131C9377B3AA2D17D5B20C52441E" ID-CoL="6HMDN" authority="Griffiths, 1984" authorityName="Griffiths" authorityYear="1984" class="Insecta" family="Braconidae" genus="Exotela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exotela viciae" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="viciae">Exotela viciae Griffiths, 1984</taxonomicName>
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<paragraph id="85C106F91D6DF6B9D485936F4CC937F7" pageId="0" pageNumber="8151">England</paragraph>
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<paragraph id="EDC6EC1FE693D0B108A8ED687A44E387" pageId="0" pageNumber="8151">Notes</paragraph>
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added by
<bibRefCitation id="369B3E9581BBDB283CA58876549A6238" author="Griffiths, G. C. D." journalOrPublisher="Beitraege zur Entomologie" pageId="0" pageNumber="8151" pagination="343 - 362" title="The Alysiinae (Hym. Braconidae) parasites of the Agromyzidae (Diptera). VII. Supplement" volume="34" year="1984">Griffiths (1984)</bibRefCitation>
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†Family
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Kirejtshuk, 1994: 57 [stem: Parandrex-]. Type genus:
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Martynov, 1926. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Parandrexe-).
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<mods:namePart>Wood, John R. I.</mods:namePart>
<mods:affiliation>Department of Plant Sciences, South Parks Road, University of Oxford, OX 1 3 RB, UK &amp; Honorary Research Associate, Royal Botanic Gardens, Kew</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Williams, Bethany R. M.</mods:namePart>
<mods:affiliation>Department of Plant Sciences, South Parks Road, University of Oxford, OX 1 3 RB, UK &amp; Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mitchell, Thomas C.</mods:namePart>
<mods:affiliation>Plant Biodiversity Research, Technische Universitaet Muenchen, Maximus-von-Imhof Forum 2, 85354 Freising, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carine, Mark A.</mods:namePart>
<mods:affiliation>Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harris, David J.</mods:namePart>
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<mods:affiliation>Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scotland, Robert W.</mods:namePart>
<mods:affiliation>Department of Plant Sciences, South Parks Road, University of Oxford, OX 1 3 RB, UK</mods:affiliation>
<mods:nameIdentifier type="email">robert.scotland@plants.ox.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-06-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
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<mods:start>1</mods:start>
<mods:end>282</mods:end>
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<mods:identifier type="Pensoft-UUID">E76E3938E216FF804849B803C469FE14</mods:identifier>
<mods:identifier type="Zenodo-Dep">576310</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182228175" LSID="urn:lsid:plazi:treatment:0C0BCFB50AD85FCB9FA3073CBF1CC66E" httpUri="http://treatment.plazi.org/id/0C0BCFB50AD85FCB9FA3073CBF1CC66E" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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17.
<taxonomicName LSID="0C0BCFB5-0AD8-5FCB-9FA3-073CBF1CC66E" authority="Boiss. &amp; Hausskn., Fl. Orient. [Boissier] 4: 104. 1875. (Boissier 1875 b: 104)." authorityName="Boiss. &amp; Hausskn., Fl. Orient. [Boissier] 4: 104. 1875. (Boissier" authorityYear="1875" class="Magnoliopsida" family="Convolvulaceae" genus="Convolvulus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Convolvulus germaniciae" order="Solanales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="germaniciae">Convolvulus germaniciae Boiss. &amp; Hausskn., Fl. Orient. [Boissier] 4: 104. 1875. (Boissier 1875b: 104).</taxonomicName>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 1 - 7 Convolvulus galaticus 1 leaves 2 bracteole 3 outer epal 4 middle sepal 5 inner sepal 6 stamen 7 ovary and style 1 from Siehe 182 (W) 2 - 7 from Bourgeau 171 (W) 8 - 13 Convolvulus germaniciae 8 leaf 9 bracteole 10 outer sepal 11 middle sepal 12 inner sepal 13 ovary and style. From Haussknecht s. n. (W) 14 - 22 Convolvulus coelesyriacus 14 leaves 15 bracteole 16 outer sepal 17 middle sepal 18 inner sepal 19 stamen 20 ovary and style 21 capsule 22 seed 14 from Davis 2979 (E) 15 - 20 from Davis 3033 (E) 21 - 22 from Meyer &amp; Dinsmore 3619 (L) 23 - 30 Convolvulus althaeoides subsp. althaeoides 23 leaves 24 bracteole 25 outer sepal 26 inner sepal 27 stamen 28 ovary and style 29 capsule 30 seed 23 - 28 from van Soest 131 (L) 29 - 30 from Boulos s. n. (CAIM) 31 - 38 Convolvulus pitardii 31 leaf and flower showing short peduncle and bracteoles 32 outer sepal 33 middle sepal 34 inner sepal 35 stamen 36 ovary and style 37 capsule 38 seed 31 from Souvage 2412 (RAB) 32 - 36 from Souvage 2413 (RAB) 37 - 38 from Souvage 14933 (RAB) 39 - 47 Convolvulus glaouorum 39 leaves 40 bracteole 41 outer sepal 42 middle sepal 43 inner sepal 44 stamen 45 ovary and style 46 capsule 47 seed. From Sauvage &amp; Vindt 2412 (RAB)." pageId="60" pageNumber="61">Figure 5, t. 8-13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="type">
<paragraph pageId="61" pageNumber="62">
<pageBreakToken pageId="61" pageNumber="62" start="start">Type</pageBreakToken>
.
</paragraph>
<paragraph pageId="61" pageNumber="62">
TURKEY, Marash,
<emphasis italics="true" pageId="61" pageNumber="62">Haussknecht</emphasis>
s.n. (holotype G; isotypes JE, W!).
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="description">
<paragraph pageId="61" pageNumber="62">Description.</paragraph>
<paragraph pageId="61" pageNumber="62">
Similar in overall morphology to
<taxonomicName authorityName="Rostan ex Choisy, Prodr. [A. P. de Candolle] 9: 408. 1845. (Choisy" authorityYear="1845" class="Magnoliopsida" family="Convolvulaceae" genus="Convolvulus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Convolvulus galaticus" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="galaticus">
<emphasis italics="true" pageId="61" pageNumber="62">Convolvulus galaticus</emphasis>
</taxonomicName>
but differing as follows: plant pilose with spreading hairs, leaves obscurely sinuate-margined but not crenate-lobed as commonly in
<taxonomicName authorityName="Rostan ex Choisy, Prodr. [A. P. de Candolle] 9: 408. 1845. (Choisy" authorityYear="1845" class="Magnoliopsida" family="Convolvulaceae" genus="Convolvulus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Convolvulus galaticus" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="galaticus">
<emphasis italics="true" pageId="61" pageNumber="62">Convolvulus galaticus</emphasis>
</taxonomicName>
, flowers mostly paired, the inflorescence commonly reflexed, sepals 7-9
<normalizedToken originalValue="×">x</normalizedToken>
3.5-5 mm, broadly elliptic, bicoloured, the apical part terminating in a distinct broad-based mucro; corolla white to pale pink, the ovary always hirsute, style pubescent below, divided 7-7.5 mm above base, stigmas 2-2.5 mm. Capsule pilose; seeds hirsute. [
<bibRefCitation author="Sa'ad, F" journalOrPublisher="Bronder Offset, Rotterdam" pageId="210" pageNumber="211" pagination="1 - 287" refId="B314" refString="Sa'ad, F, 1967. The Convolvulus species of the Canary Isles, the Mediterranean region, and the Near and Middle East. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversite Utrecht. Bronder Offset, Rotterdam: 1 - 287" title="The Convolvulus species of the Canary Isles, the Mediterranean region, and the Near and Middle East. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversite Utrecht" year="1967">
<normalizedToken originalValue="Saad">Sa'ad</normalizedToken>
1967
</bibRefCitation>
: 230;
<bibRefCitation author="Aykurt, C" journalOrPublisher="Biological Diversity and Conservation" pageId="198" pageNumber="199" pagination="55 - 59" refId="B17" refString="Aykurt, C, Suembuel, H, 2011c. Rediscovery of Convolvulus germanicae Boiss. &amp; Hausskn. (Convolvulaceae). Biological Diversity and Conservation 4: 55 - 59" title="Rediscovery of Convolvulus germanicae Boiss. &amp; Hausskn. (Convolvulaceae)." volume="4" year="2011 c">
Aykurt and
<normalizedToken originalValue="Sümbül">Suembuel</normalizedToken>
2011c
</bibRefCitation>
(photo, plate and full description)]
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="distribution">
<paragraph pageId="61" pageNumber="62">Distribution.</paragraph>
<paragraph pageId="61" pageNumber="62">
Endemic to Turkey. Previously known only from the type collection but rediscovered in 2008 (
<bibRefCitation author="Aykurt, C" journalOrPublisher="Biological Diversity and Conservation" pageId="198" pageNumber="199" pagination="55 - 59" refId="B17" refString="Aykurt, C, Suembuel, H, 2011c. Rediscovery of Convolvulus germanicae Boiss. &amp; Hausskn. (Convolvulaceae). Biological Diversity and Conservation 4: 55 - 59" title="Rediscovery of Convolvulus germanicae Boiss. &amp; Hausskn. (Convolvulaceae)." volume="4" year="2011 c">
Aykurt and
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2011c
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). It is clearly very rare.
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<taxonomicName id="1351C2FC93451B598CD94AEEAF1425FE" ID-CoL="8G5WR" authority="Mulvey, 1978 *" class="Enoplea" family="Mononchidae" genus="Coomansus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coomansus fletcherensis" order="Mononchida" pageId="0" pageNumber="1165" phylum="Nematoda" rank="species" species="fletcherensis">Coomansus fletcherensis Mulvey, 1978*</taxonomicName>
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<paragraph id="DE77B0E15A58D91E7F279372CC8D5D2A" pageId="0" pageNumber="1165">Notes</paragraph>
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Nunavut, Canada (
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).
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P.
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alis superioribus testaceo-fuscis immaculatis.
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Europa.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Genitalia of paratype ♂ (RMCA) of H. bonapartei sp. n." figureDoi="10.3897/JHR.41.8099.figure8" httpUri="https://binary.pensoft.net/fig/36029" pageId="3" pageNumber="34">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Wings of Helenanomalon species a fore and hind wings of paratype ♂ (BMNH) of H. bonapartei sp. n. b fore wing of holotype ♀ of H. ashmolei sp. n. c hind wing of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure9" httpUri="https://binary.pensoft.net/fig/36030" pageId="3" pageNumber="34">, 9a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="34" type="description">
<paragraph pageId="3" pageNumber="34">Description.</paragraph>
<paragraph pageId="3" pageNumber="34">
Male. Fore wing length 6.7-7.9 mm in ♂. Antenna with 33 (1 specimen)-36 (2 specimens) flagellomeres; all flagellomeres longer than wide and about equal width; antennal flagellum 1.2
<normalizedToken originalValue="×">x</normalizedToken>
length of fore wing (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype ♂ of Helenanomalon bonapartei sp. n.; scale bar = 10 mm." figureDoi="10.3897/JHR.41.8099.figure1" httpUri="https://binary.pensoft.net/fig/36022" pageId="3" pageNumber="34">1</figureCitation>
); scape 2.3
<normalizedToken originalValue="×">x</normalizedToken>
pedicel; distance between eyes ventrally c.0.8
<normalizedToken originalValue="×">x</normalizedToken>
distance dorsally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Face of a holotype ♂ of H. bonapartei sp. n. and b holotype ♀ of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure3" httpUri="https://binary.pensoft.net/fig/36024" pageId="3" pageNumber="34">3a</figureCitation>
); mandible more weakly twisted than in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="H. ashmolei" order="Hymenoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ashmolei">
<emphasis italics="true" pageId="3" pageNumber="34">H. ashmolei</emphasis>
</taxonomicName>
sp. n., upper tooth 2.0
<normalizedToken originalValue="×">x</normalizedToken>
lower tooth. Head and mesosoma covered in closely spaced, silvery setae, but sparser than in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="H. ashmolei" order="Hymenoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ashmolei">
<emphasis italics="true" pageId="3" pageNumber="34">H. ashmolei</emphasis>
</taxonomicName>
sp. n. (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mesoscutum of a holotype ♂ of H. bonapartei sp. n. and b holotype ♀ of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure4" httpUri="https://binary.pensoft.net/fig/36025" pageId="3" pageNumber="34">4a</figureCitation>
); metasoma, beyond first tergite, covered in slightly sparser setae; notauli strongly impressed and narrow medially but faint on anterior slope and posteriorly disappearing in rugosity (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mesoscutum of a holotype ♂ of H. bonapartei sp. n. and b holotype ♀ of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure4" httpUri="https://binary.pensoft.net/fig/36025" pageId="3" pageNumber="34">4a</figureCitation>
). Head rugose, matt, pronotum rugose-punctate laterally, dorsally punctate, mesoscutum punctate, rugose posteriorly, mesopleuron punctate, transversely striate dorsally, mesosoma with shiny interspaces between obvious punctures. Hind trochanter dorsally 3.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as trochantellus, ventrally 1.0
<normalizedToken originalValue="×">x</normalizedToken>
; hind tarsal claws abruptly curved, fully pectinate (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hind tarsal claw of a holotype ♂ of H. bonapartei sp. n. and b holotype ♀ of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure5" httpUri="https://binary.pensoft.net/fig/36026" pageId="3" pageNumber="34">5a</figureCitation>
). First sternite without convexity sub-basally. Fore wing (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Wings of Helenanomalon species a fore and hind wings of paratype ♂ (BMNH) of H. bonapartei sp. n. b fore wing of holotype ♀ of H. ashmolei sp. n. c hind wing of H. ashmolei sp. n." figureDoi="10.3897/JHR.41.8099.figure9" httpUri="https://binary.pensoft.net/fig/36030" pageId="3" pageNumber="34">9a</figureCitation>
) with
<emphasis italics="true" pageId="3" pageNumber="34">cu-a</emphasis>
slightly distal to
<emphasis italics="true" pageId="3" pageNumber="34">Rs&amp;M</emphasis>
; hind wing with 9-10 hamuli on vein
<emphasis italics="true" pageId="3" pageNumber="34">R</emphasis>
1; distal abscissa of
<emphasis italics="true" pageId="3" pageNumber="34">Cu</emphasis>
1 with distal section reaching wing membrane, proximally distant from
<emphasis italics="true" pageId="3" pageNumber="34">cu-a</emphasis>
+ basal section of
<emphasis italics="true" pageId="3" pageNumber="34">Cu</emphasis>
1 (nervellus). Petiolar index 2.68; fore wing indices: CI 0.82; BI 1.09; DBI 0.49; ICI 1.23; MI 1.60; hind wing index: RI 1.25.
</paragraph>
<caption ID-Zenodo-Dep="960893" doi="10.3897/JHR.41.8099.figure1" httpUri="https://binary.pensoft.net/fig/36022" pageId="3" pageNumber="34" start="Figure 1" startId="F1">
<paragraph pageId="3" pageNumber="34">
<emphasis bold="true" pageId="3" pageNumber="34">Figure 1.</emphasis>
Holotype ♂ of
<taxonomicName authorityName="Broad" authorityYear="2014" class="Insecta" family="Ichneumonidae" genus="Helenanomalon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helenanomalon bonapartei" order="Hymenoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="bonapartei">
<emphasis italics="true" pageId="3" pageNumber="34">Helenanomalon bonapartei</emphasis>
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sp. n.; scale bar = 10 mm.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="34">
Colour. Red, paler than in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="H. ashmolei" order="Hymenoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ashmolei">
<emphasis italics="true" pageId="3" pageNumber="34">H. ashmolei</emphasis>
</taxonomicName>
sp. n.; face, inner orbits, underside of scape bright yellow; frons and antennal flagellum black; mesosoma red with black on middle of median lobes of mesoscutum, mesoscutum posteriorly, mesopleuron dorsally, ventrally, mesosternum, propodeum dorsally, metasternum, upper division of metapleuron, postscutellum. Scutellum (mesoscutellum) black anteriorly, medially, ivory or yellow posteriorly and laterally. Following parts ivory: palps, pronotum dorso-posteriorly, ventro-posteriorly, propleural lobe, subalar prominence, fore and mid coxae and trochanters, fore trochantellus, mid trochantellus ventrally, hind trochanter and trochantellus ventrally. Hind coxa apically, dorsally, trochanter and trochantellus, femur basally, tibia and tarsus dark brown/black. First and second metasomal tergites black, remainder of metasoma and legs red.
</paragraph>
<paragraph pageId="3" pageNumber="34">Female: unknown.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="34" type="material examined">
<paragraph pageId="3" pageNumber="34">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="34">
Holotype ♂ 'St Helena: High Peak Malaise trap: xii.2005-1.2006 S
<geoCoordinate degrees="15" direction="west" minutes="58.7" orientation="longitude" precision="92" value="-15.978333">15°58.7' W</geoCoordinate>
5°44.0' c.752m UTM:02/068 18 82/1903', 'St Helena Peaks Project N.P. Ashmole, M. Ashmole, H. Mendel &amp; E. Thorpe BMNH(E) 2006-9', specimen number BMNH(E) #1022370 (BMNH).
</paragraph>
<paragraph pageId="3" pageNumber="34">
Paratypes: 1 ♂ 'St Helena: Peak Dale gumwoods: 24.i.2006: off foliage (2096)', 'St Helena Peaks Project N.P. Ashmole, M. Ashmole, H. Mendel &amp; E. Thorpe BMNH(E) 2006-9' (RMCA); 1♂ 'St Helena Knollcombes 16.x.1957 C.R.
<normalizedToken originalValue="Wallace">Wallace'</normalizedToken>
, specimen number BMNH(E) #1022371 (BMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="34" type="etymology">
<paragraph pageId="3" pageNumber="34">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="34">Named after the most famous inhabitant of St Helena, Napoleon Bonaparte, who was exiled there from 1815 until his death in 1821.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,93 @@
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<emphasis id="B6D0EAE4FF40FF8AF8F27A85EAF2FA6F" bold="true" box="[242,470,1501,1524]" italics="true" pageId="11" pageNumber="608">Dendroaspis viridis</emphasis>
(Hallowell, 1844)
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<emphasis id="B6D0EAE4FF40FF8AF8D3797BEAADF9AD" box="[211,393,1570,1591]" italics="true" pageId="11" pageNumber="608">Dendroaspis viridis ¢</emphasis>
<bibRefCitation id="E0354B07FF40FF8AF990797BE919F9AC" author="VILLIERS A." box="[400,573,1571,1591]" pageId="11" pageNumber="608" pagination="1 - 155" refId="ref21777" refString="VILLIERS A. 1950. - La collection de serpents de l'IFAN. Institut franCais d'Afrique noire, Dakar, Catalogues 6: 1 - 155." type="journal article" year="1950">Villiers 1950: 128</bibRefCitation>
</treatmentCitation>
,
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<bibRefCitation id="E0354B07FF40FF8AFA47797BE99FF9AC" author="VILLIERS A." box="[583,699,1571,1591]" pageId="11" pageNumber="608" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">1954: 1244</bibRefCitation>
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</taxonomicName>
. —
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<bibRefCitation id="E0354B07FF40FF8AFAE1797BE8E5F9AC" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[737,961,1571,1591]" pageId="11" pageNumber="608" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF40FF8AFB1C797BE86CF9AC" box="[796,840,1571,1591]" italics="true" pageId="11" pageNumber="608">et al.</emphasis>
1954b: 400
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="841B36F6FF40FF8AF8D37938EFB0F952" blockId="11.[211,807,1632,1765]" lastBlockId="11.[839,1434,1632,1737]" pageId="11" pageNumber="608">
MATÉRIEL EXAMINÉ. — [14 exemplaires]:
<collectingCountry id="FCB37666FF40FF8AFA8D7938E9EAF9EF" box="[653,718,1632,1652]" name="Guinea-Bissau" pageId="11" pageNumber="608">Guinée</collectingCountry>
: Bossou (brousse secondaire), MNHN 1951.64 (remis coll. IFAN), coll. IFAN 53.8.78 (
<bibRefCitation id="E0354B07FF40FF8AF96179C0E938F937" author="VILLIERS A." box="[353,540,1688,1708]" pageId="11" pageNumber="608" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1244</bibRefCitation>
); Diécké, MNHN 1943.148 (absent: probablement remis coll. IFAN);
<collectingRegion id="4660F814FF40FF8AFA7F79EDE999F952" box="[639,701,1717,1737]" country="Liberia" name="Nimba" pageId="11" pageNumber="608">Nimba</collectingRegion>
(alt.
<quantity id="435C9B13FF40FF8AFAFA79EDEA36F97E" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="11" pageNumber="608" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500- 700 m</quantity>
), coll. IFAN 46.1.30 (
<bibRefCitation id="E0354B07FF40FF8AF9D97989E9A2F97E" author="VILLIERS A." box="[473,646,1745,1765]" pageId="11" pageNumber="608" pagination="1 - 155" refId="ref21777" refString="VILLIERS A. 1950. - La collection de serpents de l'IFAN. Institut franCais d'Afrique noire, Dakar, Catalogues 6: 1 - 155." type="journal article" year="1950">Villiers 1950: 128</bibRefCitation>
); Nzo (champs), MNHN 1951.62, 1951-63 (remis coll. IFAN); Ziéla, MNHN 1951.61; Près de Ziéla, coll. IFAN 53.8.86 (
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Angel
<emphasis id="B6D0EAE4FF40FF8AFD207924EE69F90B" box="[1312,1357,1660,1680]" italics="true" pageId="11" pageNumber="608">et al.</emphasis>
1954b: 400
</bibRefCitation>
).
<collectingCountry id="FCB37666FF40FF8AFB8679C0E8E1F937" box="[902,965,1688,1708]" name="Liberia" pageId="11" pageNumber="608">Liberia</collectingCountry>
:
<collectingRegion id="4660F814FF40FF8AFBDE79C0EF3AF937" box="[990,1054,1688,1708]" country="Liberia" name="Nimba" pageId="11" pageNumber="608">Nimba</collectingRegion>
, MNHN 1986.1614, 1986.1620-1621, 1986.1661, 1986.1689, 1986.1798.
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<mods:date id="DDFAE94CE6FA3F99749F6E3EB08BA481">2003</mods:date>
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<mods:start id="AF54B8890A624C07436A5F20506BE773">597</mods:start>
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<paragraph id="841B36F6FF44FF8EF8F27CAEE959FB94" blockId="15.[242,637,1014,1039]" box="[242,637,1014,1039]" pageId="15" pageNumber="612">
<heading id="DF53819AFF44FF8EF8F27CAEE959FB94" box="[242,637,1014,1039]" fontSize="10" level="2" pageId="15" pageNumber="612" reason="2">
<taxonomicName id="43A44D75FF44FF8EF8F27CAEE959FB94" authority="(Fischer, 1856)" baseAuthorityName="Fischer" baseAuthorityYear="1856" box="[242,637,1014,1039]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="612" phylum="Chordata" rank="species" species="pulverulenta">
<emphasis id="B6D0EAE4FF44FF8EF8F27CAEEAF3FB96" bold="true" box="[242,471,1014,1037]" italics="true" pageId="15" pageNumber="612">Boiga pulverulenta</emphasis>
(Fischer, 1856)
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<paragraph id="841B36F6FF44FF8EFB477EF6EF02FDF7" blockId="15.[839,1434,430,620]" pageId="15" pageNumber="612">
1943.129; Ziéla (forêt primaire, alt.
<quantity id="435C9B13FF44FF8EFC9B7EF6EFFDFE59" box="[1179,1241,430,450]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="612" unit="m" value="500.0">500 m</quantity>
), MNHN 1951.53 et 1951-54 (remis coll. IFAN), coll. IFAN 53.8.37 (
<bibRefCitation id="E0354B07FF44FF8EFD157E92E852FE60" author="VILLIERS A." pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1243</bibRefCitation>
;
<bibRefCitation id="E0354B07FF44FF8EFB867EBFEF7DFE60" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[902,1113,487,507]" pageId="15" pageNumber="612" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF44FF8EFBBF7EBFE8CDFE60" box="[959,1001,487,507]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
1954b: 398
</bibRefCitation>
;
<bibRefCitation id="E0354B07FF44FF8EFC6A7EBFEFCCFE60" author="WALLACH V." box="[1130,1256,487,507]" pageId="15" pageNumber="612" pagination="95 - 99" refId="ref21842" refString="WALLACH V. 1994. - Aparallactus lineatus (Peters) and Aparallactus niger Boulenger: two valid species from West Africa. Journal of Herpetology 28: 95 - 99." type="journal article" year="1994">Wallach 1994</bibRefCitation>
) et 53.8.64 (
<bibRefCitation id="E0354B07FF44FF8EFD587EBFE893FD8C" author="VILLIERS A." pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1243</bibRefCitation>
;
<bibRefCitation id="E0354B07FF44FF8EFBC97D5BEF68FD8C" author="WALLACH V." box="[969,1100,515,535]" pageId="15" pageNumber="612" pagination="95 - 99" refId="ref21842" refString="WALLACH V. 1994. - Aparallactus lineatus (Peters) and Aparallactus niger Boulenger: two valid species from West Africa. Journal of Herpetology 28: 95 - 99." type="journal article" year="1994">Wallach 1994</bibRefCitation>
), MNHN 1962.385, 1962.387-390; Zouguépo, MNHN 1962.392.
<collectingCountry id="FCB37666FF44FF8EFC967D78EFF1FDAF" box="[1174,1237,544,564]" name="Liberia" pageId="15" pageNumber="612">Liberia</collectingCountry>
:
<collectingRegion id="4660F814FF44FF8EFCF97D78EE1DFDAF" box="[1273,1337,544,564]" country="Liberia" name="Nimba" pageId="15" pageNumber="612">Nimba</collectingRegion>
, MNHN 1986.1865, 1986.1866 (échangé avec le Muséum de
<collectingRegion id="4660F814FF44FF8EFD3C7D64EEB2FDCB" box="[1340,1430,572,592]" country="Zimbabwe" name="Bulawayo" pageId="15" pageNumber="612">Bulawayo</collectingRegion>
,
<collectingCountry id="FCB37666FF44FF8EFB477D00E883FDF7" box="[839,935,600,620]" name="Zimbabwe" pageId="15" pageNumber="612">Zimbabwé</collectingCountry>
), 1986.1867.
</paragraph>
</subSubSection>
<subSubSection id="CCBE657DFF44FF8EFB477C66EFD1FC3C" pageId="15" pageNumber="612" type="description">
<paragraph id="841B36F6FF44FF8EFB477C66EFD1FC3C" blockId="15.[839,1434,830,935]" pageId="15" pageNumber="612">
Nzo (forêt)];
<collectingRegion id="4660F814FF44FF8EFBCB7C66EF2FFCC9" box="[971,1035,830,850]" country="Liberia" name="Nimba" pageId="15" pageNumber="612">Nimba</collectingRegion>
, coll. IFAN 53.9.134 (
<bibRefCitation id="E0354B07FF44FF8EFCD27C66EEA3FCC9" author="VILLIERS A." box="[1234,1415,830,850]" pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1241</bibRefCitation>
); Ziéla, MNHN 1962.355, 1962.404, coll. IFAN 53.8.27 et 53.8.29 (
<bibRefCitation id="E0354B07FF44FF8EFB4D7C2FEF21FC10" author="VILLIERS A." box="[845,1029,887,907]" pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1241</bibRefCitation>
;
<bibRefCitation id="E0354B07FF44FF8EFC177C2FEFD1FC10" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[1047,1269,887,907]" pageId="15" pageNumber="612" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF44FF8EFC517C2FEF59FC10" box="[1105,1149,887,907]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
1954b: 394
</bibRefCitation>
).
<collectingCountry id="FCB37666FF44FF8EFD057C2FEE60FC10" box="[1285,1348,887,907]" name="Liberia" pageId="15" pageNumber="612">Liberia</collectingCountry>
:
<collectingRegion id="4660F814FF44FF8EFD567C2FEEB2FC10" box="[1366,1430,887,907]" country="Liberia" name="Nimba" pageId="15" pageNumber="612">Nimba</collectingRegion>
, (
<quantity id="435C9B13FF44FF8EFB4E7CCBE8AFFC3C" box="[846,907,915,935]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="612" unit="m" value="500.0">500 m</quantity>
) MNHN 1986.1831, 1986.1885-1887.
</paragraph>
</subSubSection>
<subSubSection id="CCBE657DFF44FF8EF8D37B64E802FB79" pageId="15" pageNumber="612" type="reference_group">
<paragraph id="841B36F6FF44FF8EF8D37B64E85EFBCB" blockId="15.[211,890,1083,1104]" box="[211,890,1083,1104]" pageId="15" pageNumber="612">
<treatmentCitationGroup id="A4B411D8FF44FF8EF8D37B64E85EFBCB" box="[211,890,1083,1104]" pageId="15" pageNumber="612">
<treatmentCitation id="050510E7FF44FF8EF8D37B64E961FBCB" author="VILLIERS A." box="[211,581,1083,1104]" page="1241" pageId="15" pageNumber="612" year="1954">
<taxonomicName id="43A44D75FF44FF8EF8D37B64E961FBCB" authority="¢ Villiers 1954: 1241" authorityName="¢ Villiers 1954:" authorityPageNumber="1241" authorityYear="1241" box="[211,581,1083,1104]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="612" phylum="Chordata" rank="species" species="pulverulenta">
<emphasis id="B6D0EAE4FF44FF8EF8D37B64EAA1FBD4" box="[211,389,1083,1104]" italics="true" pageId="15" pageNumber="612">Boiga pulverulenta ¢</emphasis>
<bibRefCitation id="E0354B07FF44FF8EF98C7B64E961FBCB" author="VILLIERS A." box="[396,581,1084,1104]" pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. —
<treatmentCitation id="050510E7FF44FF8EFA6C7B64E852FBCB" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[620,886,1084,1104]" page="394" pageId="15" pageNumber="612" year="1954">
<bibRefCitation id="E0354B07FF44FF8EFA6C7B64E852FBCB" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[620,886,1084,1104]" pageId="15" pageNumber="612" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF44FF8EFAA67B64E9F7FBCB" box="[678,723,1084,1104]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
1954b: 394-395
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="841B36F6FF44FF8EF8D37B21E802FB79" blockId="15.[211,806,1145,1250]" pageId="15" pageNumber="612">
MATÉRIEL EXAMINÉ. — [13 exemplaires]:
<collectingCountry id="FCB37666FF44FF8EFA8E7B21E9EBFB16" box="[654,719,1145,1165]" name="Guinea-Bissau" pageId="15" pageNumber="612">Guinée</collectingCountry>
: Diécké (forêt), MNHN 1943.100 (absent), coll. IFAN 53.8.75 (
<bibRefCitation id="E0354B07FF44FF8EFAE47BCDEA76FB5D" author="VILLIERS A." pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1241</bibRefCitation>
);
<collectingRegion id="4660F814FF44FF8EF9777BEAEA91FB5D" box="[375,437,1202,1222]" country="Liberia" name="Nimba" pageId="15" pageNumber="612">Nimba</collectingRegion>
(forêt primaire), MNHN 1951.46, 1962.354; Zouguépo, MNHN 1962.353.
<collectingCountry id="FCB37666FF44FF8EFA857B96E9E0FB79" box="[645,708,1230,1250]" name="Liberia" pageId="15" pageNumber="612">Liberia</collectingCountry>
:
<collectingRegion id="4660F814FF44FF8EFAE27B96E806FB79" box="[738,802,1230,1250]" country="Liberia" name="Nimba" pageId="15" pageNumber="612">Nimba</collectingRegion>
,
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,77 @@
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<paragraph id="841B36F6FF44FF8EF8F27DE4E957FD4F" blockId="15.[242,627,700,724]" box="[242,627,700,724]" pageId="15" pageNumber="612">
<heading id="DF53819AFF44FF8EF8F27DE4E957FD4F" box="[242,627,700,724]" fontSize="10" level="2" pageId="15" pageNumber="612" reason="2">
<taxonomicName id="43A44D75FF44FF8EF8F27DE4E957FD4F" authority="(Hallowell, 1844)" baseAuthorityName="Hallowell" baseAuthorityYear="1844" box="[242,627,700,724]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="612" phylum="Chordata" rank="species" species="blandingii">
<emphasis id="B6D0EAE4FF44FF8EF8F27DE4EA92FD48" bold="true" box="[242,438,700,723]" italics="true" pageId="15" pageNumber="612">Boiga blandingii</emphasis>
(Hallowell, 1844)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBE657DFF44FF8EF8D37C59E802FC3C" pageId="15" pageNumber="612" type="reference_group">
<paragraph id="841B36F6FF44FF8EF8D37C59E81DFC8E" blockId="15.[211,825,768,789]" box="[211,825,768,789]" pageId="15" pageNumber="612">
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<treatmentCitation id="050510E7FF44FF8EF8D37C59E90AFC8E" author="VILLIERS A." box="[211,558,768,789]" page="1241" pageId="15" pageNumber="612" year="1954">
<taxonomicName id="43A44D75FF44FF8EF8D37C59E90AFC8E" authority="¢ Villiers 1954: 1241" authorityName="¢ Villiers 1954:" authorityPageNumber="1241" authorityYear="1241" box="[211,558,768,789]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="612" phylum="Chordata" rank="species" species="blandingii">
<emphasis id="B6D0EAE4FF44FF8EF8D37C59EA4AFC8F" box="[211,366,768,789]" italics="true" pageId="15" pageNumber="612">Boiga blandingii ¢</emphasis>
<bibRefCitation id="E0354B07FF44FF8EF9757C59E90AFC8E" author="VILLIERS A." box="[373,558,769,789]" pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. —
<treatmentCitation id="050510E7FF44FF8EFA557C59E810FC8E" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[597,820,769,789]" page="394" pageId="15" pageNumber="612" year="1954">
<bibRefCitation id="E0354B07FF44FF8EFA557C59E810FC8E" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[597,820,769,789]" pageId="15" pageNumber="612" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF44FF8EFA907C59E998FC8E" box="[656,700,769,789]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
1954b: 394
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841B36F6FF44FF8EF8D37C66E802FC3C" blockId="15.[211,806,830,935]" pageId="15" pageNumber="612">
MATÉRIEL EXAMINÉ. — [10 exemplaires]:
<collectingCountry id="FCB37666FF44FF8EFA8F7C66E9F4FCC9" box="[655,720,830,850]" name="Guinea-Bissau" pageId="15" pageNumber="612">Guinée</collectingCountry>
: Nimba (forêt), MNHN 1943.101 [
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Angel
<emphasis id="B6D0EAE4FF44FF8EFA137C02E967FCF5" box="[531,579,858,878]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
(1954b: 394)
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donnent Nimba, en forêt comme localité de collecte pour ce spécimen, le catalogue MNHN mentionne ? et les fiches MNHN région de
</paragraph>
</subSubSection>
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View file

@ -0,0 +1,89 @@
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<taxonomicName id="43A44D75FF44FF8EF8F27E74E95BFEDF" authority="Boulenger, 1897" authorityName="Boulenger" authorityYear="1897" box="[242,639,300,324]" class="Reptilia" family="Lamprophiidae" genus="Aparallactus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="612" phylum="Chordata" rank="species" species="niger">
<emphasis id="B6D0EAE4FF44FF8EF8F27E74EAE9FED8" bold="true" box="[242,461,300,323]" italics="true" pageId="15" pageNumber="612">Aparallactus niger</emphasis>
Boulenger, 1897
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<emphasis id="B6D0EAE4FF44FF8EF8D37E29EAA6FE1F" box="[211,386,368,389]" italics="true" pageId="15" pageNumber="612">Aparallactus niger ¢</emphasis>
<bibRefCitation id="E0354B07FF44FF8EF9897E29E912FE1E" author="VILLIERS A." box="[393,566,369,389]" pageId="15" pageNumber="612" pagination="1 - 155" refId="ref21777" refString="VILLIERS A. 1950. - La collection de serpents de l'IFAN. Institut franCais d'Afrique noire, Dakar, Catalogues 6: 1 - 155." type="journal article" year="1950">Villiers 1950: 111</bibRefCitation>
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,
<treatmentCitation id="050510E7FF44FF8EFA407E29E990FE1E" author="VILLIERS A." box="[576,692,369,389]" page="1243" pageId="15" pageNumber="612" year="1954">
<bibRefCitation id="E0354B07FF44FF8EFA407E29E990FE1E" author="VILLIERS A." box="[576,692,369,389]" pageId="15" pageNumber="612" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">1954: 1243</bibRefCitation>
</treatmentCitation>
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. —
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<bibRefCitation id="E0354B07FF44FF8EFADA7E29E8C2FE1E" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[730,998,369,389]" pageId="15" pageNumber="612" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF44FF8EFB157E29E865FE1E" box="[789,833,369,389]" italics="true" pageId="15" pageNumber="612">et al.</emphasis>
1954b: 398-399
</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="E0354B07FF44FF8EFC0C7E29EFABFE1E" author="WALLACH V." box="[1036,1167,369,389]" pageId="15" pageNumber="612" pagination="95 - 99" refId="ref21842" refString="WALLACH V. 1994. - Aparallactus lineatus (Peters) and Aparallactus niger Boulenger: two valid species from West Africa. Journal of Herpetology 28: 95 - 99." type="journal article" year="1994">Wallach 1994</bibRefCitation>
.
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MATÉRIEL EXAMINÉ. — [22 exemplaires]:
<collectingCountry id="FCB37666FF44FF8EFA8A7EF6E9EFFE59" box="[650,715,430,450]" name="Guinea-Bissau" pageId="15" pageNumber="612">Guinée</collectingCountry>
: Nimba, coll. IFAN 46.1.I (sic) (
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), 46.1.5 et 46.1.6 (
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111;
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), (région de Nzo, forêt) MNHN 1943.131; camp derrière la forêt de Niomou, MNHN 1962.386; Nion (forêt, alt.
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), MNHN 1943.130; Nzo (brousse secondaire, alt.
<quantity id="435C9B13FF44FF8EF93C7D64EA5EFDCB" box="[316,378,572,592]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="612" unit="m" value="500.0">500 m</quantity>
), MNHN 1943.132; Sérengbara, MNHN 1962.391; Yalanzou (forêt secondaire, alt.
<quantity id="435C9B13FF44FF8EFA887D00E9E2FDF7" box="[648,710,600,620]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="612" unit="m" value="500.0">500 m</quantity>
), MNHN
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,102 @@
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<emphasis id="B6D0EAE4FF46FF8CF8F27C75EA87FCDF" bold="true" box="[242,419,813,836]" italics="true" pageId="13" pageNumber="610">Bitis nasicornis</emphasis>
(Shaw, 1802)
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<subSubSection id="CCBE657DFF46FF8CF8D37C2BE803FBAE" pageId="13" pageNumber="610" type="reference_group">
<paragraph id="841B36F6FF46FF8CF8D37C2BE80BFC1C" blockId="13.[211,815,882,903]" box="[211,815,882,903]" pageId="13" pageNumber="610">
<treatmentCitationGroup id="A4B411D8FF46FF8CF8D37C2BE80BFC1C" box="[211,815,882,903]" pageId="13" pageNumber="610">
<treatmentCitation id="050510E7FF46FF8CF8D37C2BE901FC1C" author="VILLIERS A." box="[211,549,882,903]" page="1247" pageId="13" pageNumber="610" year="1954">
<taxonomicName id="43A44D75FF46FF8CF8D37C2BE901FC1C" authority="¢ Villiers 1954: 1247" authorityName="¢ Villiers 1954:" authorityPageNumber="1247" authorityYear="1247" box="[211,549,882,903]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="species" species="nasicornis">
<emphasis id="B6D0EAE4FF46FF8CF8D37C2BEA41FC1D" box="[211,357,882,903]" italics="true" pageId="13" pageNumber="610">Bitis nasicornis ¢</emphasis>
<bibRefCitation id="E0354B07FF46FF8CF96B7C2BE901FC1C" author="VILLIERS A." box="[363,549,883,903]" pageId="13" pageNumber="610" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1247</bibRefCitation>
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. —
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<bibRefCitation id="E0354B07FF46FF8CFA4B7C2BE80FFC1C" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[587,811,883,903]" pageId="13" pageNumber="610" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF46FF8CFA867C2BE996FC1C" box="[646,690,883,903]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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.
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MATÉRIEL EXAMINÉ. — [19 exemplaires]:
<collectingCountry id="FCB37666FF46FF8CFA887CE8E9EDFC5F" box="[648,713,944,964]" name="Guinea-Bissau" pageId="13" pageNumber="610">Guinée</collectingCountry>
: Bossou, (
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Angel
<emphasis id="B6D0EAE4FF46FF8CF9187C94EA6CFC7B" box="[280,328,972,992]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
</bibRefCitation>
); mont Nimba, coll. IFAN 53.8.107 (
<bibRefCitation id="E0354B07FF46FF8CF8D97CB1EA8DFC66" author="VILLIERS A." box="[217,425,1001,1021]" pageId="13" pageNumber="610" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1247</bibRefCitation>
); Nimba (prairie daltitude),
<specimenCount id="92A2FD7FFF46FF8CFAE77CB1E806FC66" box="[743,802,1001,1021]" pageId="13" pageNumber="610" type="generic">1 ex.</specimenCount>
, alt.
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(
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Angel
<emphasis id="B6D0EAE4FF46FF8CF9837B5DEA8AFB82" box="[387,430,1029,1049]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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Angel
<emphasis id="B6D0EAE4FF46FF8CFAAF7B5DE9FFFB82" box="[687,731,1029,1049]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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); Nzo (
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Angel
<emphasis id="B6D0EAE4FF46FF8CF9897B79EA9DFBAE" box="[393,441,1057,1077]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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<emphasis id="B6D0EAE4FF46FF8CFAF77B79E803FBAE" box="[759,807,1057,1077]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
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View file

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<emphasis id="B6D0EAE4FF46FF8CF8F27E74EA8EFED8" bold="true" box="[242,426,300,323]" italics="true" pageId="13" pageNumber="610">Bitis rhinoceros</emphasis>
(Schlegel, 1855)
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<taxonomicName id="43A44D75FF46FF8CF8D37E29E93FFE1E" authority="¢ Villiers 1954: 1247" authorityName="¢ Villiers 1954:" authorityPageNumber="1247" authorityYear="1247" box="[211,539,368,389]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="species" species="gabonica">
<emphasis id="B6D0EAE4FF46FF8CF8D37E29EA7FFE1F" box="[211,347,368,389]" italics="true" pageId="13" pageNumber="610">Bitis gabonica ¢</emphasis>
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. —
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Angel
<emphasis id="B6D0EAE4FF46FF8CFA7C7E29E98DFE1E" box="[636,681,369,389]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401-402
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MATÉRIEL EXAMINÉ. — [15 exemplaires]:
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: Camp 3 (prairie, alt.
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), (
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Angel
<emphasis id="B6D0EAE4FF46FF8CF9E17E92E929FE45" box="[481,525,458,478]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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; dépôt non localisé); route entre Mifergui et Sempéré (alt.
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), MNHN 1994.8752 (tête seulement); Ziéla, coll. IFAN 53.8.44 (
<bibRefCitation id="E0354B07FF46FF8CF8D97D78EAB4FDAF" author="VILLIERS A." box="[217,400,544,564]" pageId="13" pageNumber="610" pagination="1234 - 1247" refId="ref21804" refString="VILLIERS A. 1954. - La collection de Serpents de l'I. F. A. N (Acquisitions 1953). Bulletin de l'Institut franCais d'Afrique noire (A) 16: 1234 - 1247." type="journal article" year="1954">Villiers 1954: 1247</bibRefCitation>
;
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Angel
<emphasis id="B6D0EAE4FF46FF8CF9DD7D78E92DFDAF" box="[477,521,544,564]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
1954b: 401
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).
<collectingCountry id="FCB37666FF46FF8CFA917D78E9F4FDAF" box="[657,720,544,564]" name="Liberia" pageId="13" pageNumber="610">Liberia</collectingCountry>
:
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, MNHN 1986.1627, 1986.1673, 1986.1690, 1986.1794-1795, 1986.1808, 1986.1826 et 1994.1216 [autrefois 1986.1826A], 1990.4760-4762; mare après le mont Richard-Molard (alt.
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), MNHN 1990.5077.
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REMARQUE. — Le travail de biologie moléculaire réalisé par
<bibRefCitation id="E0354B07FF46FF8CFB537D3AEF28FDE2" author="LENK P. &amp; HERRMANN H. - W. &amp; JOGER U. &amp; WINK M." box="[851,1036,610,633]" pageId="13" pageNumber="610" pagination="31 - 38" refId="ref21204" refString="LENK P., HERRMANN H. - W., JOGER U. &amp; WINK M. 1999. - Phylogeny and taxonomic subdivision of Bitis (Reptilia: Viperidae) based on molecular evidence. Kaupia 8: 31 - 38." type="journal article" year="1999">
Lenk
<emphasis id="B6D0EAE4FF46FF8CFB8E7D3AE89BFDE2" box="[910,959,610,633]" italics="true" pageId="13" pageNumber="610">et al.</emphasis>
(1999)
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montre que les deux sous-espèces
<emphasis id="B6D0EAE4FF46FF8CFD787D3AEA03FD00" italics="true" pageId="13" pageNumber="610">
B. g.
<taxonomicName id="43A44D75FF46FF8CF8D37DDCEA03FD00" authorityName="¢ Villiers 1954:" authorityYear="1247" box="[211,295,644,667]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="species" species="gabonica">gabonica</taxonomicName>
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et
<taxonomicName id="43A44D75FF46FF8CF9487DDCE904FD00" baseAuthorityName="Schlegel" baseAuthorityYear="1855" box="[328,544,644,667]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="subSpecies" species="gabonica" subSpecies="rhinoceros">
<emphasis id="B6D0EAE4FF46FF8CF9487DDCE904FD00" box="[328,544,644,667]" italics="true" pageId="13" pageNumber="610">B. gabonica rhinoceros</emphasis>
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représentent deux taxa distincts de rang spécifique. En effet, les divergences génétiques que mettent en évidence ces résultats suggèrent un rang taxonomique équivalent pour les trois taxa
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B. g.
<taxonomicName id="43A44D75FF46FF8CFD2C7DFEEEA4FD26" authorityName="¢ Villiers 1954:" authorityYear="1247" box="[1324,1408,678,701]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="species" species="gabonica">gabonica</taxonomicName>
</emphasis>
,
<taxonomicName id="43A44D75FF46FF8CFD907DFEEAB5FD7B" baseAuthorityName="Schlegel" baseAuthorityYear="1855" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="subSpecies" species="gabonica" subSpecies="rhinoceros">
<emphasis id="B6D0EAE4FF46FF8CFD907DFEEAB5FD7B" italics="true" pageId="13" pageNumber="610">B. gabonica rhinoceros</emphasis>
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et
<taxonomicName id="43A44D75FF46FF8CF9B27D91E908FD7B" baseAuthorityName="Shaw" baseAuthorityYear="1802" box="[434,556,713,736]" class="Reptilia" family="Viperidae" genus="Bitis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="610" phylum="Chordata" rank="species" species="nasicornis">
<emphasis id="B6D0EAE4FF46FF8CF9B27D91E908FD7B" box="[434,556,713,736]" italics="true" pageId="13" pageNumber="610">B. nasicornis</emphasis>
</taxonomicName>
.
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<taxonomicName id="43A44D75FF47FF8DF8CC7E74E96AFEDF" authority="Hallowell, 1857" authorityName="Hallowell" authorityYear="1857" box="[204,590,300,324]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="609" phylum="Chordata" rank="species" species="melanoleuca">
<emphasis id="B6D0EAE4FF47FF8DF8CC7E74EA85FED8" bold="true" box="[204,417,300,323]" italics="true" pageId="12" pageNumber="609">Naja melanoleuca</emphasis>
Hallowell, 1857
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCBE657DFF47FF8DF8AD7E29E824FDAF" pageId="12" pageNumber="609" type="reference_group">
<paragraph id="841B36F6FF47FF8DF8AD7E29E83CFE1E" blockId="12.[173,792,300,389]" box="[173,792,368,389]" pageId="12" pageNumber="609">
<treatmentCitationGroup id="A4B411D8FF47FF8DF8AD7E29E83CFE1E" box="[173,792,368,389]" pageId="12" pageNumber="609">
<treatmentCitation id="050510E7FF47FF8DF8AD7E29E92AFE1E" author="VILLIERS A." box="[173,526,368,389]" page="126" pageId="12" pageNumber="609" year="1950">
<taxonomicName id="43A44D75FF47FF8DF8AD7E29E92AFE1E" authority="¢ Villiers 1950: 126" authorityName="¢ Villiers" authorityPageNumber="126" authorityYear="1950" box="[173,526,368,389]" class="Reptilia" family="Elapidae" genus="Naja" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="609" phylum="Chordata" rank="species" species="melanoleuca">
<emphasis id="B6D0EAE4FF47FF8DF8AD7E29EA7EFE1F" box="[173,346,368,389]" italics="true" pageId="12" pageNumber="609">Naja melanoleuca ¢</emphasis>
<bibRefCitation id="E0354B07FF47FF8DF9607E29E92AFE1E" author="VILLIERS A." box="[352,526,369,389]" pageId="12" pageNumber="609" pagination="1 - 155" refId="ref21777" refString="VILLIERS A. 1950. - La collection de serpents de l'IFAN. Institut franCais d'Afrique noire, Dakar, Catalogues 6: 1 - 155." type="journal article" year="1950">Villiers 1950: 126</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. —
<treatmentCitation id="050510E7FF47FF8DFA357E29E830FE1E" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[565,788,369,389]" page="400" pageId="12" pageNumber="609" year="1954">
<bibRefCitation id="E0354B07FF47FF8DFA357E29E830FE1E" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[565,788,369,389]" pageId="12" pageNumber="609" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF47FF8DFA6F7E29E9B8FE1E" box="[623,668,369,389]" italics="true" pageId="12" pageNumber="609">et al.</emphasis>
1954b: 400
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="841B36F6FF47FF8DF8AD7EF6E824FDAF" blockId="12.[173,769,430,564]" pageId="12" pageNumber="609">
MATÉRIEL EXAMINÉ. — [13 exemplaires]:
<collectingCountry id="FCB37666FF47FF8DFA647EF6E981FE59" box="[612,677,430,450]" name="Guinea-Bissau" pageId="12" pageNumber="609">Guinée</collectingCountry>
: Diécké, spécimen cité par
<bibRefCitation id="E0354B07FF47FF8DF9617E92E979FE45" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." box="[353,605,458,478]" pageId="12" pageNumber="609" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF47FF8DF9A07E92EAF4FE45" box="[416,464,458,478]" italics="true" pageId="12" pageNumber="609">et al.</emphasis>
(1954b: 400)
</bibRefCitation>
mais dépôt non localisé; Gama, près du village: spécimen cité par
<bibRefCitation id="E0354B07FF47FF8DFA957EBFEA0FFD8C" author="ANGEL F. &amp; GUIBE J. &amp; LAMOTTE M. &amp; ROY R." pageId="12" pageNumber="609" pagination="381 - 402" refId="ref20075" refString="ANGEL F., GUIBE J., LAMOTTE M. &amp; ROY R. 1954 b. - Serpents, in La reserve naturelle integrale du mont Nimba, II. Memoires de l'Institut FranCais d'Afrique Noire 40: 381 - 402." type="journal article" year="1954">
Angel
<emphasis id="B6D0EAE4FF47FF8DFAD27EBFE825FE60" box="[722,769,487,507]" italics="true" pageId="12" pageNumber="609">et al.</emphasis>
(1954b: 400)
</bibRefCitation>
mais dépôt non localisé, MNHN 1951.161; Nimba, coll. IFAN 46.1.20 (
<bibRefCitation id="E0354B07FF47FF8DF9787D78E915FDAF" author="VILLIERS A." box="[376,561,544,564]" pageId="12" pageNumber="609" pagination="1 - 155" refId="ref21777" refString="VILLIERS A. 1950. - La collection de serpents de l'IFAN. Institut franCais d'Afrique noire, Dakar, Catalogues 6: 1 - 155." type="journal article" year="1950">Villiers 1950: 126</bibRefCitation>
); bords du Zougué,
</paragraph>
</subSubSection>
</treatment>
</document>

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