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<document id="EDC5A1F490CD0BF1BF3564C755C22BF8" ID-DOI="10.11646/phytotaxa.372.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13728728" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725682108736" checkinUser="felipe" docAuthor="Do, Hai Van, Nguyen, Mai Thi, Nguyen, Huong Thi Thanh, Choudhary, Ritesh Kumar &amp; Deng, Yunfei" docDate="2018" docId="C10F30128070590CFF6857739E5F21D8" docLanguage="en" docName="phytotaxa.372.2.3.pdf" docOrigin="Phytotaxa 372 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.372.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phlogacanthus paniculatus J. B. Imlay 1939" docType="treatment" docVersion="2" lastPageNumber="168" masterDocId="3D36486A8071590DFFE054479D1D2753" masterDocTitle="An updated taxonomy of the genus Phlogacanthus (Acanthaceae) in Vietnam" masterLastPageNumber="175" masterPageNumber="167" pageNumber="168" updateTime="1725693357352" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D9D806C5306D43BFA3D1F40925FE65A1">An updated taxonomy of the genus Phlogacanthus (Acanthaceae) in Vietnam</mods:title>
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<mods:namePart id="87FCC645B3C7E58F3E39B8938D839FC2">Do, Hai Van</mods:namePart>
<mods:affiliation id="CACFEC57243EB02395E272DB39ED1134">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="AAEF2A1DF2AC0F74B1F149C0F15C4D25" type="email">dovanhaiiebr@gmail.com</mods:nameIdentifier>
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<mods:namePart id="91835F6AD1F681E6755D5154CEEDAEA0">Nguyen, Mai Thi</mods:namePart>
<mods:affiliation id="868DA8400C5ABAA2927865D5A70E47AB">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="2BBD2D0202520F44ACC0E5FEB16AAB6B">Nguyen, Huong Thi Thanh</mods:namePart>
<mods:affiliation id="3B0477E466572BD0F9F5DA7FE1CFA697">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="F3E2D81E5B9694600CEA0CF4BE6FFC3D">Choudhary, Ritesh Kumar</mods:namePart>
<mods:affiliation id="DA0361869B874A0EFE1CF0654D049454">Biodiversity &amp; Palaeobiology Group, Agharkar Research Institute, G. G. Agarkar Road, Pune- 411004, India</mods:affiliation>
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<mods:namePart id="D9CDC92E27B6949A2E7B403C3855A866">Deng, Yunfei</mods:namePart>
<mods:affiliation id="C89E95865CC071628FF56D0A18A09F3C">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China.</mods:affiliation>
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<mods:title id="180B341DFDC4A63198AA2FBA4736B803">Phytotaxa</mods:title>
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<mods:date id="363C191A21023086923FDC528030D221">2018</mods:date>
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<mods:number id="50DB4621C1489EF891DC9CA1ABC12E4C">372</mods:number>
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<treatment id="C10F30128070590CFF6857739E5F21D8" LSID="urn:lsid:plazi:treatment:C10F30128070590CFF6857739E5F21D8" httpUri="http://treatment.plazi.org/id/C10F30128070590CFF6857739E5F21D8" lastPageNumber="168" pageId="1" pageNumber="168">
<subSubSection id="01BCD28F8070590CFF6857739ECC241C" box="[136,977,820,847]" pageId="1" pageNumber="168" type="nomenclature">
<paragraph id="491981048070590CFF6857739ECC241C" blockId="1.[136,977,820,847]" box="[136,977,820,847]" pageId="1" pageNumber="168">
<heading id="125136688070590CFF6857739ECC241C" box="[136,977,820,847]" fontSize="11" level="2" pageId="1" pageNumber="168" reason="3">
<treatmentCitationGroup id="69B6A62A8070590CFF6857739ECC241C" box="[136,977,820,847]" pageId="1" pageNumber="168">
<treatmentCitation id="C807A7158070590CFF6857739E2E241C" author="Imlay, J. B." box="[136,819,820,847]" page=")" pageId="1" pageNumber="168" year="1939">
<taxonomicName id="8EA6FA878070590CFF6857739E2E241C" ID-CoL="4GHF9" authority="(T. Anderson) Imlay (1939: 128)" authorityName="J. B. Imlay" authorityPageNumber="128" authorityYear="1939" baseAuthorityName="T. Anderson" baseAuthorityYear="1867" box="[136,819,820,847]" class="Magnoliopsida" family="Acanthaceae" genus="Phlogacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168070590CFF6857739CA2241C" bold="true" box="[136,447,820,847]" italics="true" pageId="1" pageNumber="168">Phlogacanthus paniculatus</emphasis>
(T. Anderson)
<bibRefCitation id="2D37FCF58070590CFD8557719E2E241C" author="Imlay, J. B." box="[613,819,822,847]" pageId="1" pageNumber="168" pagination="109 - 150" refId="ref4223" refString="Imlay, J. B. (1939) Contributions to the flora of Siam, Additamentum LI, new and renamed Siamense Acanthaceae. Bulletin of Miscellaneous Information Kew 3: 109 - 150. https: // doi. org / 10.2307 / 4111677" type="journal article" year="1939">Imlay (1939: 128)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="7BD25D168070590CFCDD57719ECC241C" bold="true" box="[829,977,822,847]" pageId="1" pageNumber="168">
(
<figureCitation id="D19D9D818070590CFCA557719E89241C" box="[837,916,822,847]" captionStart="FIGURE 1" captionStartId="2.[136,229,1569,1590]" captionTargetBox="[168,1426,186,1524]" captionTargetId="figure-15@2.[158,1430,171,1541]" captionTargetPageId="2" captionText="FIGURE 1. Phlogacanthus paniculatus (T. Anderson) J.B. Imlay: A. Flowering and fruiting twig; B. Calyx and bracteoles; C. Corolla (open); D. Ovary and style; E. Fruit (Open) (Illustration drawn by Mrs Le Kim Chi, from D.V. Hai 14, HN)." figureDoi="http://doi.org/10.5281/zenodo.13728730" httpUri="https://zenodo.org/record/13728730/files/figure.png" pageId="1" pageNumber="168">Figs. 1</figureCitation>
&amp;
<figureCitation id="D19D9D818070590CFC5A57719ED4241C" box="[954,969,822,847]" captionStart="FIGURE 2" captionStartId="3.[136,229,1530,1551]" captionTargetBox="[151,1435,190,1505]" captionTargetId="figure-14@3.[151,1436,190,1505]" captionTargetPageId="3" captionText="FIGURE 2. Phlogacanthus paniculatus (T. Anderson) J.B. Imlay: A. Flowering and fruiting twig; B. Flower (front view); C. Calyx and flower; D. Calyx and fruit (Photos by D.V. Hai)." figureDoi="http://doi.org/10.5281/zenodo.13728732" httpUri="https://zenodo.org/record/13728732/files/figure.png" pageId="1" pageNumber="168">2</figureCitation>
)
</emphasis>
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</heading>
</paragraph>
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<treatmentCitationGroup id="69B6A62A8070590CFF68571B9F962422" box="[136,651,860,881]" pageId="1" pageNumber="168">
=
<treatmentCitation id="C807A7158070590CFF7C571B9F9A2422" author="Anderson, T." box="[156,647,860,881]" page=")" pageId="1" pageNumber="168" year="1867">
<taxonomicName id="8EA6FA878070590CFF7C571B9F9A2422" ID-CoL="33P7W" authority="T. Anderson (1867: 458)" authorityName="T. Anderson" authorityPageNumber="458" authorityYear="1867" box="[156,647,860,881]" class="Magnoliopsida" family="Acanthaceae" genus="Cystacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168070590CFF7C571B9C8E2422" box="[156,403,860,881]" italics="true" pageId="1" pageNumber="168">Cystacanthus paniculatus</emphasis>
T.
<bibRefCitation id="2D37FCF58070590CFE50571B9F9A2422" author="Anderson, T." box="[432,647,860,881]" pageId="1" pageNumber="168" pagination="425 - 526" refId="ref3754" refString="Anderson, T. (1867) An enumeration of the Indian species of Acanthaceae. Journal of the Linnean Society, Botany 9: 425 - 526." type="journal article" year="1867">Anderson (1867: 458)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="01BCD28F8070590CFF6857C79F8022DC" pageId="1" pageNumber="168" type="materials_examined">
<paragraph id="491981048070590CFF6857C79FA824C6" blockId="1.[136,693,860,917]" box="[136,693,896,917]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF6857C79DC724C6" bold="true" box="[136,218,896,917]" pageId="1" pageNumber="168">
<typeStatus id="961D3FA68070590CFF6857C79DA024C6" box="[136,189,896,917]" pageId="1" pageNumber="168">Type</typeStatus>
:—
</emphasis>
<collectingCountry id="31B1C1948070590CFF3A57C79C4624C6" box="[218,347,896,917]" name="Myanmar" pageId="1" pageNumber="168">MYANMAR</collectingCountry>
. Moulmein,
<emphasis id="7BD25D168070590CFE3457C79F2324C6" box="[468,574,896,917]" italics="true" pageId="1" pageNumber="168">
<collectionCode id="2FB719C18070590CFE3457C79CFF24C6" box="[468,482,896,917]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="168">T</collectionCode>
.Lobb s. n
</emphasis>
. (
<collectionCode id="2FB719C18070590CFDB157C79F9824C6" box="[593,645,896,917]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="1" pageNumber="168" type="Herbarium">CAL</collectionCode>
,
<collectionCode id="2FB719C18070590CFD6E57C79FB824C6" box="[654,677,896,917]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="168" type="Herbarium">K</collectionCode>
!).
</paragraph>
<paragraph id="491981048070590CFF68578198BB2238" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
Shrubs to
<quantity id="8E5E2CE18070590CFF1D57819C34248C" box="[253,297,966,991]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="168" unit="m" value="1.0">1 m</quantity>
tall. Stems 4-angled, stout, glabrous; young branches winged. Petiole
<quantity id="8E5E2CE18070590CFBAE578199A6248C" box="[1102,1211,966,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="168" unit="cm" value="1.35" valueMax="1.7" valueMin="1.0">1–1.7 cm</quantity>
; leaf blade oblong to oblong-lanceolate, 6–12 ×
<quantity id="8E5E2CE18070590CFE5957AD9F262350" box="[441,571,1002,1027]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.5" metricValueMin="1.5" pageId="1" pageNumber="168" unit="cm" value="3.0" valueMax="4.5" valueMin="1.5">1.5–4.5 cm</quantity>
, abaxially pubescent along veins, adaxially glabrous, secondary veins 6–8 on each side of midvein, base attenuate and decurrent onto petiole, margin undulate to crenulate, apex acuminate. Thyrses terminal, to
<quantity id="8E5E2CE18070590CFEF750759C7E2318" box="[279,355,1074,1099]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="168" unit="cm" value="13.0">13 cm</quantity>
, many-flowered; rachis densely villous with gland-tipped trichomes; bracts oblong–lanceolate, shorter than calyx, gland-tipped pubescent, caducous; bracteoles similar to bracts. Pedicel
<quantity id="8E5E2CE18070590CFB4B50119813233C" box="[1195,1294,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="168" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, gland-tipped pubescent. Calyx
<quantity id="8E5E2CE18070590CFEB0503D9CA023C0" box="[336,445,1146,1171]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="1" pageNumber="168" unit="mm" value="11.0" valueMax="14.0" valueMin="8.0">8–14 mm</quantity>
, outside gland-tipped pubescent, inside slightly pubescent; lobes lanceolate, 7–13 ×
<quantity id="8E5E2CE18070590CFA8C503D9DAB23E4" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="168" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
, equal, apex acute. Corolla white or light purple,
<quantity id="8E5E2CE18070590CFD3A50D99E5823E4" box="[730,837,1182,1207]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="168" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
; bent to 90º and abruptly inflated at middle, ventricose, outside gland-tipped pubescent, inside glabrous with purple stripes; limb 2-lipped; lower lip 3-lobed with lobes orbicular, subequal, patent; upper lip 2-lobed with lobes orbicular, equal. Stamens two, included; filaments barbate; anther thecae oblong, ca.
<quantity id="8E5E2CE18070590CFE42514D9CF92270" box="[418,484,1290,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="168" unit="mm" value="1.0">1 mm</quantity>
, connective barbate; staminodes very small. Ovary pubescent, with 4 ovules per locule; style
<quantity id="8E5E2CE18070590CFF2351699C5F2214" box="[195,322,1326,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="1" pageNumber="168" unit="cm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 cm</quantity>
, glabrous or pubescent; stigma slightly 2-cleft or entire. Capsule greyish brown, clavate, ca.
<quantity id="8E5E2CE18070590CFADE516998BA2214" box="[1342,1447,1326,1351]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="168" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
, pubescent, 8-seeded. Seeds ovate in outline, ca. 3.5 ×
<quantity id="8E5E2CE18070590CFD0451159E202238" box="[740,829,1362,1387]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="168" unit="mm" value="3.5">3.5 mm</quantity>
; densely shortly hairy; hairs standing out when wetted.
</paragraph>
<paragraph id="491981048070590CFF5D51319F8022DC" blockId="1.[136,1452,966,1675]" box="[189,669,1398,1423]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF5D51319CC222DC" bold="true" box="[189,479,1398,1423]" pageId="1" pageNumber="168">Flowering and Fruiting:</emphasis>
June to October.
</paragraph>
</subSubSection>
<subSubSection id="01BCD28F8070590CFF5D51DD9C9F2284" pageId="1" pageNumber="168" type="biology_ecology">
<paragraph id="491981048070590CFF5D51DD9C9F2284" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF5D51DD9CA722E0" bold="true" box="[189,442,1434,1459]" pageId="1" pageNumber="168">Ecology and habitat:</emphasis>
The species usually grows in the understorey of primary broad-leaved evergreen closed forest, in humid areas.
</paragraph>
</subSubSection>
<subSubSection id="01BCD28F8070590CFF5D51A59E5F21D8" pageId="1" pageNumber="168" type="distribution">
<paragraph id="491981048070590CFF5D51A59DE6214C" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF5D51A59C4422A8" bold="true" box="[189,345,1506,1531]" pageId="1" pageNumber="168">Distribution:</emphasis>
<collectingCountry id="31B1C1948070590CFE8551A59CD922A8" box="[357,452,1506,1531]" name="Vietnam" pageId="1" pageNumber="168">Vietnam</collectingCountry>
(
<collectingRegion id="8B624FE68070590CFE3751A59FC522A8" box="[471,728,1506,1531]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="168">Quang Binh Province</collectingRegion>
,
<collectingRegion id="8B624FE68070590CFD0951A59EC622A8" box="[745,987,1506,1531]" country="Vietnam" name="Quang Ninh" pageId="1" pageNumber="168">Quang Ninh District</collectingRegion>
, Ba Ren farm forest);
<collectingCountry id="31B1C1948070590CFB0D51A5982D22A8" box="[1261,1328,1506,1531]" name="China" pageId="1" pageNumber="168">China</collectingCountry>
(
<collectingRegion id="8B624FE68070590CFAA451A598BD22A8" box="[1348,1440,1506,1531]" country="China" name="Yunnan" pageId="1" pageNumber="168">Yunnan</collectingRegion>
),
<collectingCountry id="31B1C1948070590CFF6852419DEA214C" box="[136,247,1542,1567]" name="Myanmar" pageId="1" pageNumber="168">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="491981048070590CFF5D526D9E5F21D8" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF5D526D9CDC2110" bold="true" box="[189,449,1578,1603]" pageId="1" pageNumber="168">Specimens examined:</emphasis>
<materialsCitation id="F9CE8B598070590CFE29526D9DF42134" collectingDate="1981-10-24" collectionCode="HN" collectorName="D. D. Huyen" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
<emphasis id="7BD25D168070590CFE29526D9F4A2110" bold="true" box="[457,599,1578,1603]" pageId="1" pageNumber="168">
<collectingCountry id="31B1C1948070590CFE29526D9F4F2110" box="[457,594,1578,1603]" name="Vietnam" pageId="1" pageNumber="168">VIETNAM</collectingCountry>
.
</emphasis>
<collectingRegion id="8B624FE68070590CFD80526D9E452110" box="[608,856,1578,1603]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="168">Quang Binh Province</collectingRegion>
,
<collectingRegion id="8B624FE68070590CFC85526D994D2110" box="[869,1104,1578,1603]" country="Vietnam" name="Quang Ninh" pageId="1" pageNumber="168">Quang Ninh District</collectingRegion>
,
<date id="3D18A7C48070590CFBBD526D98032110" box="[1117,1310,1578,1603]" pageId="1" pageNumber="168" value="1981-10-24">
<collectingDate id="2D5C5E2C8070590CFBBD526D98032110" box="[1117,1310,1578,1603]" pageId="1" pageNumber="168" value="1981-10-24">24 October 1981</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFACC526D9DB92134" italics="true" pageId="1" pageNumber="168">
<collectorName id="E453E4D28070590CFACC526D98B62110" box="[1324,1451,1578,1603]" pageId="1" pageNumber="168">D.D.Huyen</collectorName>
88
</emphasis>
(
<collectionCode id="2FB719C18070590CFF5452099DFF2134" box="[180,226,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFF1652099E072134" box="[246,794,1614,1639]" collectingDate="2008-08-12" collectionCode="V, HN" collectorName="Hai" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFE7752099F512134" box="[407,588,1614,1639]" pageId="1" pageNumber="168" value="2008-08-12">
<collectingDate id="2D5C5E2C8070590CFE7752099F512134" box="[407,588,1614,1639]" pageId="1" pageNumber="168" value="2008-08-12">12 August 2008</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFDB752099FC92134" box="[599,724,1614,1639]" italics="true" pageId="1" pageNumber="168">
D.
<collectionCode id="2FB719C18070590CFD9052099F9F2134" box="[624,642,1614,1639]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="168" type="Museum">V</collectionCode>
.
<collectorName id="E453E4D28070590CFD6552099FAC2134" box="[645,689,1614,1639]" pageId="1" pageNumber="168">Hai</collectorName>
14
</emphasis>
(
<collectionCode id="2FB719C18070590CFD0452099E0F2134" box="[740,786,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFCC65209987B2134" box="[806,1382,1614,1639]" collectingDate="1981-08-22" collectionCode="T, K, HN" collectorName="Lien" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFC27520999662134" box="[967,1147,1614,1639]" pageId="1" pageNumber="168" value="1981-08-22">
<collectingDate id="2D5C5E2C8070590CFC27520999662134" box="[967,1147,1614,1639]" pageId="1" pageNumber="168" value="1981-08-22">22 August 1981</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFB685209983D2134" box="[1160,1312,1614,1639]" italics="true" pageId="1" pageNumber="168">
<collectionCode id="2FB719C18070590CFB68520999872134" box="[1160,1178,1614,1639]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="168">T</collectionCode>
.
<collectionCode id="2FB719C18070590CFB7D520999AE2134" box="[1181,1203,1614,1639]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="168" type="Herbarium">K</collectionCode>
.
<collectorName id="E453E4D28070590CFB5D520999F22134" box="[1213,1263,1614,1639]" pageId="1" pageNumber="168">Lien</collectorName>
385
</emphasis>
(
<collectionCode id="2FB719C18070590CFAD0520998432134" box="[1328,1374,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFA9252099F8921D8" collectingDate="2003-06-07" collectionCode="N, Q" collectorName="Binh &amp; D. D. Cuong" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFF0852359C6E21D8" box="[232,371,1650,1675]" pageId="1" pageNumber="168" value="2003-06-07">
<collectingDate id="2D5C5E2C8070590CFF0852359C6E21D8" box="[232,371,1650,1675]" pageId="1" pageNumber="168" value="2003-06-07">7 June 2003</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFE9E52359CF321D8" box="[382,494,1650,1675]" italics="true" pageId="1" pageNumber="168">
<collectionCode id="2FB719C18070590CFE9E52359C8A21D8" box="[382,407,1650,1675]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="168" type="Herbarium">N</collectionCode>
.
<collectionCode id="2FB719C18070590CFE7A52359CB221D8" box="[410,431,1650,1675]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="1" pageNumber="168" type="Herbarium">Q</collectionCode>
.
<collectorName id="E453E4D28070590CFE5952359CF321D8" box="[441,494,1650,1675]" pageId="1" pageNumber="168">Binh</collectorName>
</emphasis>
&amp;
<collectorName id="E453E4D28070590CFDF252359F8921D8" box="[530,660,1650,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFDF252359F8921D8" box="[530,660,1650,1675]" italics="true" pageId="1" pageNumber="168">D.D.Cuong</emphasis>
</collectorName>
</materialsCitation>
<materialsCitation id="F9CE8B598070590CFD7A52359E2321D8" box="[666,830,1650,1675]" collectionCode="HN" country="Vietnam" location="VN" pageId="1" pageNumber="168" specimenCount="1">
<emphasis id="7BD25D168070590CFD7A52359FE421D8" box="[666,761,1650,1675]" italics="true" pageId="1" pageNumber="168">
<collectingCountry id="31B1C1948070590CFD7A52359FA321D8" box="[666,702,1650,1675]" name="Vietnam" pageId="1" pageNumber="168">VN</collectingCountry>
1112
</emphasis>
(
<collectionCode id="2FB719C18070590CFCE852359E2B21D8" box="[776,822,1650,1675]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

301
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View file

@ -0,0 +1,301 @@
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<mods:title id="D6479D9C596BF143C86283C598A9CD76">An updated taxonomy of the genus Phlogacanthus (Acanthaceae) in Vietnam</mods:title>
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<taxonomicName id="8EA6FA878070590CFF6852F09E652180" ID-CoL="6VC8T" authority="(Benoist) D. V. Hai, Y. F. Deng, R. K. Choudhary" authorityName="D. V. Hai, Y. F. Deng, R. K. Choudhary" authorityYear="2018" baseAuthorityName="D. V. Hai, Y. F. Deng, R. K. Choudhary" baseAuthorityYear="1928" box="[136,888,1719,1747]" class="Magnoliopsida" family="Acanthaceae" genus="Phlogacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="datii" status="comb. nov.">
<emphasis id="7BD25D168070590CFF6852F09C6D2181" bold="true" box="[136,368,1719,1746]" italics="true" pageId="1" pageNumber="168">Phlogacanthus datii</emphasis>
(Benoist) D.V.Hai, Y.F.Deng, R.K.Choudhary
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(
<figureCitation id="D19D9D818070590CFBF252FD997F2180" box="[1042,1122,1722,1747]" captionStart="FIGURE 3" captionStartId="4.[136,229,1689,1710]" captionTargetBox="[175,1417,198,1661]" captionTargetId="figure-15@4.[151,1436,190,1664]" captionTargetPageId="4" captionText="FIGURE 3. Phlogacanthus datii (Benoist) D.V. Hai, Y.F. Deng, R.K.Choudhary:A. Flowering and fruiting twig; B. Inflorescence (a part); C. Open corolla showing stamens and staminodes; D. Opened fruits showing seeds. Drawn by Mrs Le Kim Chi, from D.V. Hai 05, HN)." figureDoi="http://doi.org/10.5281/zenodo.13728734" httpUri="https://zenodo.org/record/13728734/files/figure.png" pageId="1" pageNumber="168">Figs. 3</figureCitation>
&amp;
<figureCitation id="D19D9D818070590CFB6752FD998B2180" box="[1159,1174,1722,1747]" captionStart="FIGURE 4" captionStartId="5.[136,229,1953,1974]" captionTargetBox="[183,1404,190,1929]" captionTargetId="figure-14@5.[183,1405,190,1929]" captionTargetPageId="5" captionText="FIGURE 4. Phlogacanthus datii (Benoist) D.V. Hai, Y.F. Deng, R.K.Choudhary: A. Flowering twig; B. Inflorescence; C. Part of the inflorescence; D. Flower showing calyx; E. Bud; F. Flower; G. Flower (open); H. Calyx; I. Stamens; J. Ovary and style (Photos by D.V. Hai)." figureDoi="http://doi.org/10.5281/zenodo.13728736" httpUri="https://zenodo.org/record/13728736/files/figure.png" pageId="1" pageNumber="168">4</figureCitation>
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<emphasis id="7BD25D168070590CFF6852A79DE821A6" bold="true" box="[136,245,1760,1781]" pageId="1" pageNumber="168">Basionym:</emphasis>
<taxonomicName id="8EA6FA878070590CFF1B52A79F6A21A6" ID-CoL="33P7T" authority="Benoist (1928: 908)" authorityName="Benoist" authorityPageNumber="908" authorityYear="1928" box="[251,631,1760,1781]" class="Magnoliopsida" family="Acanthaceae" genus="Cystacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="7BD25D168070590CFF1B52A79CB221A6" box="[251,431,1760,1781]" italics="true" pageId="1" pageNumber="168">Cystacanthus datii</emphasis>
<bibRefCitation id="2D37FCF58070590CFE5552A79F6A21A6" author="Benoist, R." box="[437,631,1760,1781]" pageId="1" pageNumber="168" refId="ref3783" refString="https: // doi. org / 10.1111 / j. 1095 - 8339.1867. tb 01308. x Benoist, R. (1928) Acanthacees nouvelles asiatiques. Bulletin de la Societe Botanique de France 74: 907 - 912." type="book" year="1928">Benoist (1928: 908)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="491981048070590CFF6853439847204A" blockId="1.[136,1370,1760,1817]" box="[136,1370,1796,1817]" pageId="1" pageNumber="168">
<materialsCitation id="F9CE8B598070590CFF6853439847204A" box="[136,1370,1796,1817]" country="Vietnam" location="VIETNAM" pageId="1" pageNumber="168" specimenCount="2" typeStatus="lectotype">
<typeStatus id="961D3FA68070590CFF6853439DF5204A" box="[136,232,1796,1817]" pageId="1" pageNumber="168" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="31B1C1948070590CFE5153439F38204A" box="[433,549,1796,1817]" name="Vietnam" pageId="1" pageNumber="168">VIETNAM</collectingCountry>
. Lac Tho, 1892,
<emphasis id="7BD25D168070590CFD2553439E3F204A" box="[709,802,1796,1817]" italics="true" pageId="1" pageNumber="168">Bon 5301</emphasis>
(P-03578821!;
<typeStatus id="961D3FA68070590CFC5A53439926204A" box="[954,1083,1796,1817]" pageId="1" pageNumber="168" type="isolectotype">isolectotypes</typeStatus>
: P-03578822!; P-03578825!).
</materialsCitation>
</paragraph>
<paragraph id="491981048070590CFF68530D9FDC2F44" blockId="1.[136,1452,1866,2071]" pageId="1" pageNumber="168">
Notes: When Benoist (1927: 908) described this species, he cited two collections in the protologue, i.e.
<emphasis id="7BD25D168070590CFAF0530D98732030" box="[1296,1390,1866,1891]" italics="true" pageId="1" pageNumber="168">Bon 301</emphasis>
from
<collectingCountry id="31B1C1948070590CFF6853299DCA20D4" box="[136,215,1902,1927]" name="Vietnam" pageId="1" pageNumber="168">Tonkin</collectingCountry>
(present day
<collectingCountry id="31B1C1948070590CFE8B53299CC520D4" box="[363,472,1902,1927]" name="Vietnam" pageId="1" pageNumber="168">Viet Nam</collectingCountry>
) and
<emphasis id="7BD25D168070590CFDF453299F9120D4" box="[532,652,1902,1927]" italics="true" pageId="1" pageNumber="168">Spire 1508</emphasis>
from
<collectingCountry id="31B1C1948070590CFD2E53299E1A20D4" box="[718,775,1902,1927]" name="Laos" pageId="1" pageNumber="168">Laos</collectingCountry>
, without designating a
<typeStatus id="961D3FA68070590CFBE65329997120D4" box="[1030,1132,1902,1927]" pageId="1" pageNumber="168" type="holotype">holotype</typeStatus>
. A search of these specimens revealed that
<specimenCount id="5FA04A8D8070590CFEC153D59CA620F8" box="[289,443,1938,1963]" count="1" pageId="1" pageNumber="168" type="generic">one specimen</specimenCount>
of
<emphasis id="7BD25D168070590CFE0253D59F5C20F8" box="[482,577,1938,1963]" italics="true" pageId="1" pageNumber="168">Bon 301</emphasis>
at P (P-03578821) has branches with open flowers, which matches very well with the diagnostic characters of the species mentioned in the protologue, and it is selected here as the
<typeStatus id="961D3FA68070590CFB1653F19840209C" box="[1270,1373,1974,1999]" pageId="1" pageNumber="168" type="lectotype">lectotype</typeStatus>
for the name
<taxonomicName id="8EA6FA878070590CFF2D539D9C8220A0" ID-CoL="33P7T" authorityName="Benoist" authorityYear="1928" box="[205,415,2010,2035]" class="Magnoliopsida" family="Acanthaceae" genus="Cystacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="7BD25D168070590CFF2D539D9C8220A0" box="[205,415,2010,2035]" italics="true" pageId="1" pageNumber="168">Cystacanthus datii</emphasis>
</taxonomicName>
(
<figureCitation id="D19D9D818070590CFE4F539D9CEF20A0" box="[431,498,2010,2035]" captionStart="FIGURE 5" captionStartId="6.[136,229,1953,1974]" captionTargetBox="[220,1367,190,1929]" captionTargetId="figure-15@6.[220,1368,190,1929]" captionTargetPageId="6" captionText="FIGURE 5. Lectotype of Phlogacanthus datii (P-03578821)." figureDoi="http://doi.org/10.5281/zenodo.13728738" httpUri="https://zenodo.org/record/13728738/files/figure.png" pageId="1" pageNumber="168">Fig. 5</figureCitation>
) according to Art. 9.2, 9.11 &amp; 9.12 of the ICN (
<bibRefCitation id="2D37FCF58070590CFBF4539D99E920A0" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[1044,1268,2010,2035]" pageId="1" pageNumber="168" refId="ref4568" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Glashutten: Koeltz Botanical Books. https: // doi. org / 10.12705 / Code. 2018" type="journal volume" year="2018">
Turland
<emphasis id="7BD25D168070590CFB96539D99B220A0" box="[1142,1199,2010,2035]" italics="true" pageId="1" pageNumber="168">et al.</emphasis>
2018
</bibRefCitation>
). The remaining duplicates are designated here as the
<typeStatus id="961D3FA68070590CFDC753B99FA02F44" box="[551,701,2046,2071]" pageId="1" pageNumber="168" type="isolectotype">isolectotypes</typeStatus>
.
</paragraph>
<caption id="1DD9D18C8073590FFF68526699892109" ID-DOI="http://doi.org/10.5281/zenodo.13728730" ID-Zenodo-Dep="13728730" httpUri="https://zenodo.org/record/13728730/files/figure.png" pageId="2" pageNumber="169" startId="2.[136,229,1569,1590]" targetBox="[168,1426,186,1524]" targetPageId="2" targetType="figure">
<paragraph id="491981048073590FFF68526699892109" blockId="2.[136,1451,1569,1626]" pageId="2" pageNumber="169">
<emphasis id="7BD25D168073590FFF6852669DE42165" bold="true" box="[136,249,1569,1590]" pageId="2" pageNumber="169">FIGURE 1</emphasis>
.
<taxonomicName id="8EA6FA878073590FFEE652669E162165" authority="(T. Anderson) J. B. Imlay" authorityName="J. B. Imlay" authorityYear="1939" baseAuthorityName="T. Anderson" baseAuthorityYear="1867" box="[262,779,1569,1590]" class="Magnoliopsida" family="Acanthaceae" genus="Phlogacanthus" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="169" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168073590FFEE652669F102165" box="[262,525,1569,1590]" italics="true" pageId="2" pageNumber="169">Phlogacanthus paniculatus</emphasis>
(T. Anderson) J.B. Imlay
</taxonomicName>
: A. Flowering and fruiting twig; B. Calyx and bracteoles; C. Corolla (open); D. Ovary and style; E. Fruit (Open) (Illustration drawn by Mrs Le Kim Chi, from
<emphasis id="7BD25D168073590FFC065202994A2109" box="[998,1111,1605,1626]" italics="true" pageId="2" pageNumber="169">D.V. Hai 14</emphasis>
, HN).
</paragraph>
</caption>
<paragraph id="491981048073590FFF5D52CD9ECB2F75" blockId="2.[136,1452,1674,2086]" pageId="2" pageNumber="169">
Shrubs
<quantity id="8E5E2CE18073590FFEF752CD9C5B21F0" box="[279,326,1674,1699]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="169" unit="m" value="1.0">1 m</quantity>
tall, erect. Stems subterete or slightly 4-angled, pubescent, soon glabrescent. Petiole
<quantity id="8E5E2CE18073590FFAD752CD98B521F0" box="[1335,1448,1674,1699]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="169" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
; leaf blade ovate to oblong, 4.5–14 ×
<quantity id="8E5E2CE18073590FFDD252EA9F972195" box="[562,650,1709,1734]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="2" pageNumber="169" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 cm</quantity>
, abaxially glabrous and puberulent along veins, adaxially glabrescent, secondary veins ca. 4 on each side of midvein, base cuneate to rounded, margin entire, apex acuminate to acute. Thyrses paniculate, to
<quantity id="8E5E2CE18073590FFE6D52B49CC4205F" box="[397,473,1779,1804]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="169" unit="cm" value="15.0">15 cm</quantity>
; bracts linear-lanceolate,
<quantity id="8E5E2CE18073590FFD1D52B49E42205F" box="[765,863,1779,1804]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="2" pageNumber="169" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
, densely gland-tipped trichomes; bracteoles similar to bracts but smaller. Pedicel ca.
<quantity id="8E5E2CE18073590FFDE053509F4E2063" box="[512,595,1815,1840]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="169" unit="mm" value="10.0">10 mm</quantity>
, gland-tipped trichomes. Calyx ca.
<quantity id="8E5E2CE18073590FFC07535099512063" box="[999,1100,1815,1840]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="169" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 cm</quantity>
, outside pubescent with gland-tipped trichomes, inside slightly pubescent; lobes linear, 6–9 ×
<quantity id="8E5E2CE18073590FFCA8537D9E962000" box="[840,907,1850,1875]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="169" unit="mm" value="1.0">1 mm</quantity>
, equal, apex acute. Corolla pale yellow with thin to dense purplish dots,
<quantity id="8E5E2CE18073590FFE72531A9CE22025" box="[402,511,1885,1910]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="169" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
, ventricose, incurved, outside gland-tipped pubescent; tube basally ca.
<quantity id="8E5E2CE18073590FFACA531A98702025" box="[1322,1389,1885,1910]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="169" unit="mm" value="2.0">2 mm</quantity>
wide and short then wide at middle; usually bent ca. 90º and abruptly inflated near middle; limb 2-lipped; lower lip 3-lobed, lobes ovate and equal; upper lip 2-lobed; lobes ascending cochlear in bud. Stamens 2; filaments ca.
<quantity id="8E5E2CE18073590FFB0653E4984C20EF" box="[1254,1361,1955,1980]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="169" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
, basally pilose; anther thecae oblong, ca.
<quantity id="8E5E2CE18073590FFE1853809F4A20B3" box="[504,599,1991,2016]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="169" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, connective barbate; staminodes filiform, ca.
<quantity id="8E5E2CE18073590FFBB5538099D420B3" box="[1109,1225,1991,2016]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="169" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
. Ovary ca.
<quantity id="8E5E2CE18073590FFAA8538098BA20B3" box="[1352,1447,1991,2016]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="169" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, oblong, pubescent, with 4 ovules per locule; style ca.
<quantity id="8E5E2CE18073590FFD1653AD9E6E2F50" box="[758,883,2026,2051]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="169" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
, sparsely pubescent or glabrescent; stigma entire. Capsule clavate, ca.
<quantity id="8E5E2CE18073590FFE8D5C4A9CA12F75" box="[365,444,2061,2086]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="169" unit="cm" value="2.5">2.5 cm</quantity>
, pubescent, 8-seeded. Seeds ovate, compressed.
</paragraph>
</subSubSection>
<caption id="1DD9D18C8072590EFF6851BD9F452160" ID-DOI="http://doi.org/10.5281/zenodo.13728732" ID-Zenodo-Dep="13728732" httpUri="https://zenodo.org/record/13728732/files/figure.png" pageId="3" pageNumber="170" startId="3.[136,229,1530,1551]" targetBox="[151,1435,190,1505]" targetPageId="3" targetType="figure">
<paragraph id="491981048072590EFF6851BD9F452160" blockId="3.[136,1452,1530,1587]" pageId="3" pageNumber="170">
<emphasis id="7BD25D168072590EFF6851BD9DE5215C" bold="true" box="[136,248,1530,1551]" pageId="3" pageNumber="170">FIGURE 2</emphasis>
.
<taxonomicName id="8EA6FA878072590EFEE551BD9E18215C" authority="(T. Anderson) J. B. Imlay" authorityName="J. B. Imlay" authorityYear="1939" baseAuthorityName="T. Anderson" baseAuthorityYear="1867" box="[261,773,1530,1551]" class="Magnoliopsida" family="Acanthaceae" genus="Phlogacanthus" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="170" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168072590EFEE551BD9F16215C" box="[261,523,1530,1551]" italics="true" pageId="3" pageNumber="170">Phlogacanthus paniculatus</emphasis>
(T. Anderson) J.B. Imlay
</taxonomicName>
: A. Flowering and fruiting twig; B. Flower (front view); C. Calyx and flower; D. Calyx and fruit (Photos by D.V. Hai).
</paragraph>
</caption>
<subSubSection id="01BCD28F8072590EFF5D52249F91202B" pageId="3" pageNumber="170" type="distribution">
<paragraph id="491981048072590EFF5D522498B121BB" blockId="3.[136,1452,1635,1912]" pageId="3" pageNumber="170">
<emphasis id="7BD25D168072590EFF5D52249C44212F" bold="true" box="[189,345,1635,1660]" pageId="3" pageNumber="170">Distribution:</emphasis>
Laos, Vietnam.
<emphasis id="7BD25D168072590EFF5D52C09C0421F3" bold="true" box="[189,281,1671,1696]" pageId="3" pageNumber="170">Habitat</emphasis>
: Under evergreen broad-leaved forest,
<quantity id="8E5E2CE18072590EFD3152C09E0521F3" box="[721,792,1671,1696]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="170" unit="m" value="200.0">200 m</quantity>
a.s.l.
<emphasis id="7BD25D168072590EFF5D52EC9C242197" bold="true" box="[189,313,1707,1732]" pageId="3" pageNumber="170">Phenology</emphasis>
: Flowering from March to April.
<emphasis id="7BD25D168072590EFF5D52889CDE21BB" bold="true" box="[189,451,1743,1768]" pageId="3" pageNumber="170">Specimens examined:</emphasis>
<collectingCountry id="31B1C1948072590EFE2E52889F4921BB" box="[462,596,1743,1768]" name="Vietnam" pageId="3" pageNumber="170">VIETNAM</collectingCountry>
:
<collectingRegion id="8B624FE68072590EFD8152889E5021BB" box="[609,845,1743,1768]" country="Vietnam" name="Vinh Phuc" pageId="3" pageNumber="170">Vinh Phuc Province</collectingRegion>
: Phuc Yen town,
<date id="3D18A7C48072590EFBFD528899DE21BB" box="[1053,1219,1743,1768]" pageId="3" pageNumber="170" value="2008-04-15">15 April 2008</date>
,
<emphasis id="7BD25D168072590EFB325288984321BB" box="[1234,1374,1743,1768]" italics="true" pageId="3" pageNumber="170">D.V. Hai 05</emphasis>
(HN);
</paragraph>
<paragraph id="491981048072590EFF6852B498B6205F" blockId="3.[136,1452,1635,1912]" box="[136,1451,1778,1804]" pageId="3" pageNumber="170">
<geoCoordinate id="2C92E7C38072590EFF6852B49C36205F" box="[136,299,1778,1804]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="170" precision="1" seconds="37.1" value="21.393639">
21°23
<emphasis id="7BD25D168072590EFF2B52B59DCC205F" bold="true" box="[203,209,1778,1804]" italics="true" pageId="3" pageNumber="170">′</emphasis>
37.1
<emphasis id="7BD25D168072590EFEE252B59C13205F" bold="true" box="[258,270,1778,1804]" italics="true" pageId="3" pageNumber="170">″</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="2C92E7C38072590EFED552B49CFC205F" box="[309,481,1778,1804]" degrees="105" direction="east" minutes="43" orientation="longitude" pageId="3" pageNumber="170" precision="1" seconds="11.6" value="105.71989">
105°43
<emphasis id="7BD25D168072590EFE6652B59C91205F" bold="true" box="[390,396,1778,1804]" italics="true" pageId="3" pageNumber="170">′</emphasis>
11.6
<emphasis id="7BD25D168072590EFE5D52B59CD4205F" bold="true" box="[445,457,1778,1804]" italics="true" pageId="3" pageNumber="170">″</emphasis>
E
</geoCoordinate>
,
<quantity id="8E5E2CE18072590EFE0C52B49F2F205F" box="[492,562,1779,1804]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="3" pageNumber="170" unit="m" value="222.0">222 m</quantity>
a.s.l.;
<materialsCitation id="F9CE8B598072590EFD9A52B499BB205F" box="[634,1190,1779,1804]" collectingDate="2014-03-16" collectionCode="V, T, HN" collectorName="Hai &amp; X. Thanh" pageId="3" pageNumber="170" specimenCount="1">
<date id="3D18A7C48072590EFD9A52B49E3B205F" box="[634,806,1779,1804]" pageId="3" pageNumber="170" value="2014-03-16">
<collectingDate id="2D5C5E2C8072590EFD9A52B49E3B205F" box="[634,806,1779,1804]" pageId="3" pageNumber="170" value="2014-03-16">16 March 2014</collectingDate>
</date>
,
<emphasis id="7BD25D168072590EFCD052B49E8D205F" box="[816,912,1779,1804]" italics="true" pageId="3" pageNumber="170">
D.
<collectionCode id="2FB719C18072590EFCA952B49E41205F" box="[841,860,1779,1804]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="170" type="Museum">V</collectionCode>
.
<collectorName id="E453E4D28072590EFC8652B49E8D205F" box="[870,912,1779,1804]" pageId="3" pageNumber="170">Hai</collectorName>
</emphasis>
,
<emphasis id="7BD25D168072590EFC7D52B4997F205F" box="[925,1122,1779,1804]" italics="true" pageId="3" pageNumber="170">
<collectionCode id="2FB719C18072590EFC7D52B49EB3205F" box="[925,942,1779,1804]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="170">T</collectionCode>
.
<collectorName id="E453E4D28072590EFC5152B4990A205F" box="[945,1047,1779,1804]" pageId="3" pageNumber="170">X. Thanh</collectorName>
HT 01
</emphasis>
(
<collectionCode id="2FB719C18072590EFB9152B49982205F" box="[1137,1183,1779,1804]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="3" pageNumber="170" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598072590EFB5152B498B6205F" box="[1201,1451,1779,1804]" collectingDate="2014-03-16" collectionCode="V, T, HN" collectorName="Hai &amp; X. Thanh" pageId="3" pageNumber="170" specimenCount="1">same locality, 26 April</materialsCitation>
</paragraph>
<paragraph id="491981048072590EFF68535098B62063" blockId="3.[136,1452,1635,1912]" box="[136,1451,1815,1840]" pageId="3" pageNumber="170">
2004,
<emphasis id="7BD25D168072590EFF3053509C832063" box="[208,414,1815,1840]" italics="true" pageId="3" pageNumber="170">V.X. Phuong 7592</emphasis>
(HN).
<collectingRegion id="8B624FE68072590EFE1153509FCD2063" box="[497,720,1815,1840]" country="Vietnam" name="Hoa Binh" pageId="3" pageNumber="170">Hòa Bình Province</collectingRegion>
: Kim Boi District, Thuong Tien Commune,
<date id="3D18A7C48072590EFB2B535098BA2063" box="[1227,1447,1815,1840]" pageId="3" pageNumber="170" value="1999-12-23">23 December 1999</date>
,
</paragraph>
<paragraph id="491981048072590EFF68537C98B62007" blockId="3.[136,1452,1635,1912]" box="[136,1451,1851,1876]" pageId="3" pageNumber="170">
<emphasis id="7BD25D168072590EFF68537C9C0F2007" box="[136,274,1851,1876]" italics="true" pageId="3" pageNumber="170">V.X. Phuong</emphasis>
2537 (HN).
<collectingRegion id="8B624FE68072590EFE40537C9F432007" box="[416,606,1851,1876]" country="Vietnam" name="Son La" pageId="3" pageNumber="170">Son La Province</collectingRegion>
: Moc Chau District, Muong Sang Commune,
<date id="3D18A7C48072590EFBBE537C99E12007" box="[1118,1276,1851,1876]" pageId="3" pageNumber="170" value="1977-04-13">13 April 1977</date>
,
<emphasis id="7BD25D168072590EFAE6537C98B62007" box="[1286,1451,1851,1876]" italics="true" pageId="3" pageNumber="170">H.T. Dung 186</emphasis>
</paragraph>
<paragraph id="491981048072590EFF6853189F91202B" blockId="3.[136,1452,1635,1912]" box="[136,652,1887,1912]" pageId="3" pageNumber="170">
(HN).
<collectingCountry id="31B1C1948072590EFF3153189C03202B" box="[209,286,1887,1912]" name="Laos" pageId="3" pageNumber="170">LAOS</collectingCountry>
:
<collectingRegion id="8B624FE68072590EFEC953189CCA202B" box="[297,471,1887,1912]" country="Laos" name="Louangphabang" pageId="3" pageNumber="170">Luang Prabang</collectingRegion>
,
<emphasis id="7BD25D168072590EFE0153189F4E202B" box="[481,595,1887,1912]" italics="true" pageId="3" pageNumber="170">Spire1508</emphasis>
(P!).
</paragraph>
</subSubSection>
</treatment>
</document>